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<mods:title id="6DABCEDA2E2C9B7ED5BCF5DD71D94E29">Revision of the genus Glyptoxanthus A. Milne-Edwards, 1879, and establishment of Glyptoxanthinae nov. subfam. (Crustacea: Decapoda: Brachyura: Xanthidae)</mods:title>
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<taxonomicName id="4C514D532671FFC243B8FE96FCD4FE9C" ID-CoL="3GJHF" authority="A. Milne-Edwards, 1878" authorityName="A. Milne-Edwards" authorityYear="1878" box="[151,802,331,358]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosus">
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<emphasis id="B925EAC22671FFC243B8FE96FE21FE9F" bold="true" box="[151,471,332,358]" italics="true" pageId="7" pageNumber="36">Glyptoxanthus cavernosus</emphasis>
(
<bibRefCitation id="EFC04B212671FFC242C8FE91FCEFFE9C" author="Milne-Edwards" box="[487,793,331,357]" pageId="7" pageNumber="36" refString="Milne-Edwards, A. (1878) Description de quelques especes nouvelles de Crustaces provenant du voyage aux iles Cap-Vert de MM. Bouvier et de Cessac. Bulletin de la Societe philomatique (Paris), (ser. 7), 2, 225 - 232." type="journal article" year="1878">A. Milne-Edwards, 1878</bibRefCitation>
)
</emphasis>
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<paragraph id="8BEE36D02671FFC243B8FEB4FEC4FE7F" blockId="7.[151,802,331,390]" box="[151,306,366,390]" pageId="7" pageNumber="36">
(
<figureCitation id="136A2A552671FFC243B0FEB4FF18FE7F" box="[159,238,366,390]" captionStart="FIGURE 3" captionStartId="7.[151,250,1934,1957]" captionTargetBox="[151,1436,1060,1913]" captionTargetId="figure@7.[151,1436,1060,1914]" captionTargetPageId="7" captionText="FIGURE 3. Glyptoxanthus cavernosus (A. Milne-Edwards, 1878), male, 49.6 × 35.8 mm (MNHN-B 8329), Cape Verde: A, dorsal view; B, anterior view; C, ventral view; D, left chela, external view." httpUri="https://zenodo.org/record/207313/files/figure.png" pageId="7" pageNumber="36">Figs. 3</figureCitation>
,
<figureCitation id="136A2A552671FFC243D5FEB4FEE1FE7F" box="[250,279,366,390]" captionStart="FIGURE 10" captionStartId="18.[151,250,1623,1646]" captionTargetBox="[179,1416,213,1568]" captionTargetId="figure@18.[151,1436,193,1602]" captionTargetPageId="18" captionText="FIGURE 10. Glyptoxanthus spp. gonopods (G 1). A, G. angolensis (Brito Capello, 1866); B, G. cavernosus (A. Milne-Edwards, 1878); C, G. erosus (Stimpson, 1859); E, G. c o r ro s u s (H. Mile Edwards, 1834); E, G. labyrinthicus (Stimpson, 1860), (LACM CR 1935.075.3); F, G. meandricus (Lockington, 1877), neotype (LACM CR 1940.65); G, G. vermiculatus (Lamarck, 1818). (A C, D, G, after Guinot 1979)." httpUri="https://zenodo.org/record/207320/files/figure.png" pageId="7" pageNumber="36">10</figureCitation>
B)
</paragraph>
</subSubSection>
<subSubSection id="C34B655B2671FFC243B8FE6DFAB7FD9F" pageId="7" pageNumber="36" type="reference_group">
<paragraph id="8BEE36D02671FFC243B8FE6DFB6CFE35" blockId="7.[151,1436,438,614]" box="[151,1178,438,461]" pageId="7" pageNumber="36">
<treatmentCitationGroup id="AB4111FE2671FFC243B8FE6DFB6CFE35" box="[151,1178,438,461]" pageId="7" pageNumber="36">
<taxonomicName id="4C514D532671FFC243B8FE6DFD77FE35" ID-CoL="9RZS" authority="A. Milne-Edwards, 1878: 226" authorityName="A. Milne-Edwards" authorityPageNumber="226" authorityYear="1878" box="[151,641,438,461]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosa">
<emphasis id="B925EAC22671FFC243B8FE6DFEBAFE34" box="[151,332,439,461]" italics="true" pageId="7" pageNumber="36">Actaea cavernosa</emphasis>
<treatmentCitation id="0AF010C12671FFC2427DFE6DFD77FE35" author="Milne-Edwards" box="[338,641,438,460]" page="226" pageId="7" pageNumber="36" year="1878">
<bibRefCitation id="EFC04B212671FFC2427DFE6DFDA7FE35" author="Milne-Edwards" box="[338,593,438,460]" pageId="7" pageNumber="36" refString="Milne-Edwards, A. (1878) Description de quelques especes nouvelles de Crustaces provenant du voyage aux iles Cap-Vert de MM. Bouvier et de Cessac. Bulletin de la Societe philomatique (Paris), (ser. 7), 2, 225 - 232." type="journal article" year="1878">A. Milne-Edwards, 1878</bibRefCitation>
: 226
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</taxonomicName>
(
<typeStatus id="54EA88722671FFC241A1FE6DFD4DFE35" box="[654,699,439,460]" pageId="7" pageNumber="36">type</typeStatus>
locality:
<collectingCountry id="F34676402671FFC24034FE6DFC66FE35" box="[795,912,438,460]" name="Cape Verde" pageId="7" pageNumber="36">Cape Verde</collectingCountry>
Is.). —
<treatmentCitation id="0AF010C12671FFC240F1FE6CFB60FE35" author="Monod" box="[990,1174,438,460]" page="523" pageId="7" pageNumber="36" year="1933">
<bibRefCitation id="EFC04B212671FFC240F1FE6CFB95FE35" author="Monod" box="[990,1123,438,460]" pageId="7" pageNumber="36" refString="Monod, T. (1933) Sur quelques Crustaces de l'Afrique occidentale (liste des Decapodes mauritaniens et des Xanthides ouestafricains). Bulletin du Comite d'Etudes Historiques et Scientifiques de l'Afrique Occidentale Francaise, 15, 456 - 548." type="journal article" year="1933">Monod 1933</bibRefCitation>
: 523
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BEE36D02671FFC243B8FE0FFC2CFE13" blockId="7.[151,1436,438,614]" box="[151,986,469,491]" pageId="7" pageNumber="36">
<treatmentCitationGroup id="AB4111FE2671FFC243B8FE0FFC2CFE13" box="[151,986,469,491]" pageId="7" pageNumber="36">
<taxonomicName id="4C514D532671FFC243B8FE0FFDDCFE13" ID-CoL="993FR" authority="Osorio 1906: 149" authorityName="Osorio" authorityPageNumber="149" authorityYear="1906" box="[151,554,469,491]" class="Malacostraca" family="Xanthidae" genus="Xantho" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="vermiculatus">
<emphasis id="B925EAC22671FFC243B8FE0FFE9FFE12" box="[151,361,469,491]" italics="true" pageId="7" pageNumber="36">Xantho vermiculatus</emphasis>
,
<treatmentCitation id="0AF010C12671FFC24259FE0FFDDCFE13" author="Osorio" box="[374,554,469,491]" page="149" pageId="7" pageNumber="36" year="1906">
<bibRefCitation id="EFC04B212671FFC24259FE0FFE01FE12" author="Osorio" box="[374,503,469,491]" pageId="7" pageNumber="36" refString="Osorio, B. (1906) Uma nova lista de Crustaceos Africanos. Jornal de Sciencias mathematicas, physicas e naturaes (Lisboa), ser. 2, 7 (27), 149, 150." type="journal article" year="1906">Osorio 1906</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
. Not
<taxonomicName id="4C514D532671FFC2414FFE0CFC23FE13" authority="Lamarck, 1818" authorityName="Lamarck" authorityYear="1818" box="[608,981,469,491]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="vermiculatus">
<emphasis id="B925EAC22671FFC2414FFE0CFCC2FE12" box="[608,820,469,491]" italics="true" pageId="7" pageNumber="36">Cancer vermiculatus</emphasis>
<bibRefCitation id="EFC04B212671FFC24015FE0FFC23FE13" author="Lamarck" box="[826,981,469,491]" pageId="7" pageNumber="36" refString="Lamarck, J. B. P. A. (1818) Histoire naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin, l'exposition des principes fondamentaux de la zoologie. Volume 5: 1 - 612." type="journal article" year="1818">Lamarck, 1818</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BEE36D02671FFC243B8FE2EFBDEFDF0" blockId="7.[151,1436,438,614]" box="[151,1064,499,522]" pageId="7" pageNumber="36">
<treatmentCitationGroup id="AB4111FE2671FFC243B8FE2EFBDEFDF0" box="[151,1064,499,522]" pageId="7" pageNumber="36">
<taxonomicName id="4C514D532671FFC243B8FE2EFD4DFDF0" ID-CoL="9RZS" authority="Monod 1956: 298" authorityName="Monod" authorityPageNumber="298" authorityYear="1956" box="[151,699,499,522]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosa" subGenus="Glyptoxanthus">
<emphasis id="B925EAC22671FFC243B8FE2EFF2BFDF0" box="[151,221,500,521]" italics="true" pageId="7" pageNumber="36">Actaea</emphasis>
(
<emphasis id="B925EAC22671FFC243C3FE2EFE76FDF3" box="[236,384,500,522]" italics="true" pageId="7" pageNumber="36">Glyptoxanthus</emphasis>
)
<emphasis id="B925EAC22671FFC242A1FE2FFE00FDF3" box="[398,502,501,522]" italics="true" pageId="7" pageNumber="36">cavernosa</emphasis>
,
<treatmentCitation id="0AF010C12671FFC2412CFE29FD4DFDF0" author="Monod" box="[515,699,499,521]" page="298" pageId="7" pageNumber="36" year="1956">
<bibRefCitation id="EFC04B212671FFC2412CFE29FD7FFDF0" author="Monod" box="[515,649,499,521]" pageId="7" pageNumber="36" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africaines. Memoires de l'Institut Fondamental d'Afrique Noire, 45, 1 - 674." type="journal article" year="1956">Monod 1956</bibRefCitation>
: 298
</treatmentCitation>
</taxonomicName>
, fig. 362. —
<treatmentCitation id="0AF010C12671FFC24015FE29FC1CFDF0" author="Serene" box="[826,1002,499,521]" page="198" pageId="7" pageNumber="36" year="1961">
<bibRefCitation id="EFC04B212671FFC24015FE29FC4CFDF0" author="Serene" box="[826,954,499,521]" pageId="7" pageNumber="36" refString="Serene, R. (1961) Les especes indo-pacifiques d' Actea et celles des genres Pseudactea et Banareia (1 e partie). Bulletin de la Societe zoologique de France, 86 (2 - 3), 195 - 212." type="journal article" year="1961">Serène 1961</bibRefCitation>
: 198
</treatmentCitation>
(list).
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</paragraph>
<paragraph id="8BEE36D02671FFC243B8FDC8FC98FDD1" blockId="7.[151,1436,438,614]" box="[151,878,530,552]" pageId="7" pageNumber="36">
<treatmentCitationGroup id="AB4111FE2671FFC243B8FDC8FC98FDD1" box="[151,878,530,552]" pageId="7" pageNumber="36">
<taxonomicName id="4C514D532671FFC243B8FDC8FDC9FDD1" ID-CoL="9RZS" authority="Balss 1921: 61" authorityName="Balss" authorityPageNumber="61" authorityYear="1921" box="[151,575,530,552]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosa">
<emphasis id="B925EAC22671FFC243B8FDC8FE6FFDD1" box="[151,409,530,552]" italics="true" pageId="7" pageNumber="36">Glyptoxanthus cavernosa</emphasis>
,
<treatmentCitation id="0AF010C12671FFC2428AFDC8FDC9FDD1" author="Balss" box="[421,575,530,552]" page="61" pageId="7" pageNumber="36" year="1921">
<bibRefCitation id="EFC04B212671FFC2428AFDC8FDEFFDD1" author="Balss" box="[421,537,530,552]" pageId="7" pageNumber="36" refString="Balss, H. (1921) Crustacea VI: Decapoda Anomura (Paguridae) und Brachyura (Dromiacea bis Brachygnatha). In: Michaelsen, W. (Ed.) Beitrage zur Kenntnis der Meeresfauna Westafrikas, III, Lief. 2, Hamburg. pp. 37 - 68." type="journal article" year="1921">Balss 1921</bibRefCitation>
: 61
</treatmentCitation>
</taxonomicName>
. —
<treatmentCitation id="0AF010C12671FFC2414DFDC9FC9CFDD1" author="Forest" box="[610,874,530,552]" page="79" pageId="7" pageNumber="36" year="1966">
<bibRefCitation id="EFC04B212671FFC2414DFDC9FCB2FDD1" author="Forest" box="[610,836,530,552]" pageId="7" pageNumber="36" refString="Forest, J. &amp; Guinot, D. (1966) Crustaces Decapodes: Brachyoures. In: Campagne de la Calypso dans le golfe de Guinee et aux iles Principe, Sao Tome et Annobon (1956), 16. Annales de l'Institut Oceanographique (Monaco), 44, 23 - 124." type="journal article" year="1966">Forest &amp; Guinot 1966</bibRefCitation>
: 79
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36D02671FFC243B8FDEBFAB7FD9F" blockId="7.[151,1436,438,614]" pageId="7" pageNumber="36">
<treatmentCitationGroup id="AB4111FE2671FFC243B8FDEBFAB7FD9F" pageId="7" pageNumber="36">
<taxonomicName id="4C514D532671FFC243B8FDEBFD92FDBE" ID-CoL="3GJHF" authority="Bouvier 1922: 66" authorityName="Bouvier" authorityPageNumber="66" authorityYear="1922" box="[151,612,561,583]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="B925EAC22671FFC243B8FDEBFE55FDBE" box="[151,419,561,583]" italics="true" pageId="7" pageNumber="36">Glyptoxanthus cavernosus</emphasis>
,
<treatmentCitation id="0AF010C12671FFC2429FFDEBFD92FDBE" author="Bouvier" box="[432,612,561,583]" page="66" pageId="7" pageNumber="36" year="1922">
<bibRefCitation id="EFC04B212671FFC2429FFDEBFDC9FDBE" author="Bouvier" box="[432,575,561,583]" pageId="7" pageNumber="36" refString="Bouvier, E. L. (1922) Observations complementaires sur les Crustaces Decapodes (abstraction fait des Carides) provenant des campagnes de S. A. S. le Prince de Monaco. In: Resultats Campagnes Scientifiques du Prince Albert de Monaco, fasc. LXII, 106 pp., 6 pls." type="book" year="1922">Bouvier 1922</bibRefCitation>
: 66
</treatmentCitation>
</taxonomicName>
, pl. 6 fig. 3, 5. —
<treatmentCitation id="0AF010C12671FFC24035FDEBFC26FDBE" author="Guinot" box="[794,976,561,583]" page="551" pageId="7" pageNumber="36" year="1967">
<bibRefCitation id="EFC04B212671FFC24035FDEBFC6BFDBE" author="Guinot" box="[794,925,561,583]" pageId="7" pageNumber="36" refString="Guinot, D. (1967) Recherches preliminaires sur les groupements naturels chez les Crustaces Decapodes Brachyoures. III. A propos des affinites des genre Dairoides Stebbing et Daira de Haan. Bulletin du Museum national d'Histoire naurelle, 2 e serie, 39 (3), 540 - 563." type="journal article" year="1967">Guinot 1967</bibRefCitation>
: 551
</treatmentCitation>
, 553, 556, fig. 31; 1971: 1073; 1979: 68, fig. 18C, pl. 6 fig. 5. —
<treatmentCitation id="0AF010C12671FFC242A2FD8AFD39FD9F" author="Manning" box="[397,719,592,614]" page="135" pageId="7" pageNumber="36" year="1981">
<bibRefCitation id="EFC04B212671FFC242A2FD8AFD6AFD9C" author="Manning" box="[397,668,592,614]" pageId="7" pageNumber="36" refString="Manning, R. B. &amp; Holthuis, L. B. (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, No. 306, i - xii, 1 - 379." type="book chapter" year="1981">Manning &amp; Holthuis 1981</bibRefCitation>
: 135
</treatmentCitation>
. —
<treatmentCitation id="0AF010C12671FFC241DCFD8AFBEAFD9C" author="d'Udekem" box="[755,1052,592,614]" page="229" pageId="7" pageNumber="36" year="1999">
<bibRefCitation id="EFC04B212671FFC241DCFD8AFC1CFD9C" author="d'Udekem" box="[755,1002,592,614]" pageId="7" pageNumber="36" refString="d'Udekem d'Acoz, C. (1999) Inventaire et distribution des Crustaces Decapodes de l'Atlantique nord-oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Patrimonies Naturels (M. N. H. N. / S. P. N.), 40, 1 - 383." type="journal article" year="1999">dUdekem dAcoz 1999</bibRefCitation>
: 229
</treatmentCitation>
. —Ng
<emphasis id="B925EAC22671FFC2474BFD8BFB67FD9F" box="[1124,1169,592,614]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2008: 199 (list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B655B2671FFC243B8FD48FCBAFC17" pageId="7" pageNumber="36" type="materials_examined">
<paragraph id="8BEE36D02671FFC243B8FD48FC90FC5F" blockId="7.[151,1436,658,1007]" pageId="7" pageNumber="36">
<emphasis id="B925EAC22671FFC243B8FD48FDB4FD52" bold="true" box="[151,578,658,683]" pageId="7" pageNumber="36">
Material examined.
<collectingCountry id="F34676402671FFC242A7FD48FDE2FD52" box="[392,532,658,683]" name="Cape Verde" pageId="7" pageNumber="36">Cape Verde</collectingCountry>
Is.:
</emphasis>
<specimenCount id="9D57FD592671FFC24164FD48FD6FFD53" box="[587,665,658,682]" pageId="7" pageNumber="36" type="male">1 male</specimenCount>
, 49.6
<emphasis id="B925EAC22671FFC241F1FD49FD1BFD53" box="[734,749,659,682]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC241DAFD48FCACFD53" box="[757,858,658,682]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="7" pageNumber="36" unit="mm" value="35.8">35.8 mm</quantity>
(MNHN-B8329), Stn 148, 10
<quantity id="4CA99B352671FFC24794FD48FB01FD53" box="[1211,1271,658,682]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="36" unit="m" value="20.0">20 m</quantity>
,
<emphasis id="B925EAC22671FFC2462BFD49FAABFD52" box="[1284,1373,659,683]" italics="true" pageId="7" pageNumber="36">Sylvana</emphasis>
, coll. Le Comte J. Polignac,
<date id="FFEF10102671FFC242BBFD6CFDE8FD37" box="[404,542,694,719]" pageId="7" pageNumber="36" value="1913-05-08">8 May 1913</date>
;
<specimenCount id="9D57FD592671FFC24106FD6CFD81FD37" box="[553,631,694,718]" pageId="7" pageNumber="36" type="male">1 male</specimenCount>
, 26.7
<emphasis id="B925EAC22671FFC24195FD6DFD3FFD37" box="[698,713,695,718]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC241FFFD6CFCC0FD37" box="[720,822,694,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="7" pageNumber="36" unit="mm" value="19.1">19.1 mm</quantity>
,
<specimenCount id="9D57FD592671FFC2406EFD6CFC53FD37" box="[833,933,694,718]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 24.9
<emphasis id="B925EAC22671FFC240C8FD6DFC00FD37" box="[999,1014,695,718]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC240D2FD6CFB97FD37" box="[1021,1121,694,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="7" pageNumber="36" unit="mm" value="17.9">17.9 mm</quantity>
(MNHN-B8330), La Praya,
<quantity id="4CA99B352671FFC243B8FD00FF08FD0B" box="[151,254,730,754]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="7" pageNumber="36" unit="m" value="20.0" valueMax="30.0" valueMin="10.0">1030 m</quantity>
,
<taxonomicName id="4C514D532671FFC24223FD01FE4BFD0B" box="[268,445,730,755]" class="Ostracoda" family="Incertae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myodocopida" pageId="7" pageNumber="36" phylum="Arthropoda" rank="family">
<emphasis id="B925EAC22671FFC24223FD01FE84FD0A" box="[268,370,731,755]" italics="true" pageId="7" pageNumber="36">Talisman</emphasis>
, 1883
</taxonomicName>
;
<specimenCount id="9D57FD592671FFC242E5FD00FDEDFD0B" box="[458,539,730,754]" pageId="7" pageNumber="36" type="male">1 male</specimenCount>
, 21.7
<emphasis id="B925EAC22671FFC2414CFD01FD84FD0B" box="[611,626,731,754]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC24154FD00FD15FD0B" box="[635,739,730,754]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="7" pageNumber="36" unit="mm" value="15.5">15.5 mm</quantity>
,
<specimenCount id="9D57FD592671FFC241DEFD00FCA1FD0B" box="[753,855,730,754]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 27.2
<emphasis id="B925EAC22671FFC240B0FD01FC58FD0B" box="[927,942,731,754]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC24098FD00FBEBFD0B" box="[951,1053,730,754]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="7" pageNumber="36" unit="mm" value="19.7">19.7 mm</quantity>
(RMNH-D40250), stn CANCAP 7.D15, scuba, Baia de Ribeirinha, W coast of
<collectingCountry id="F34676402671FFC2418AFD24FCC7FCEF" box="[677,817,766,790]" name="Saint Vincent and the Grenadines" pageId="7" pageNumber="36">São Vicente</collectingCountry>
, coll.
<date id="FFEF10102671FFC24058FD25FBF4FCEF" box="[887,1026,766,790]" pageId="7" pageNumber="36" value="1986-09-07">7 Sep. 1986</date>
;
<specimenCount id="9D57FD592671FFC24720FD24FBA9FCEF" box="[1039,1119,766,790]" pageId="7" pageNumber="36" type="male">1 male</specimenCount>
, 24.7
<emphasis id="B925EAC22671FFC2478AFD25FB42FCEF" box="[1189,1204,767,790]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC24792FD24FAD3FCEF" box="[1213,1317,766,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="7" pageNumber="36" unit="mm" value="17.8">17.8 mm</quantity>
,
<specimenCount id="9D57FD592671FFC2461DFD24FA6EFCEF" box="[1330,1432,766,790]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 26.7
<emphasis id="B925EAC22671FFC243FCFCF9FF14FCC3" box="[211,226,803,826]" italics="true" pageId="7" pageNumber="36">×</emphasis>
19.0 mm (RMNH-D40251), stn CANCAP 7.K18, snorkelling, intertidal to shallow sublittoral, shallow lagoon with corals, SE coast of Sal, coll.
<date id="FFEF10102671FFC2415FFC9CFCF9FCA7" box="[624,783,838,863]" pageId="7" pageNumber="36" value="1986-08-29">29 Aug. 1986</date>
;
<specimenCount id="9D57FD592671FFC24032FC9CFC74FCA7" box="[797,898,838,862]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 13.7
<emphasis id="B925EAC22671FFC240E7FC9DFC21FCA7" box="[968,983,839,862]" italics="true" pageId="7" pageNumber="36">×</emphasis>
<quantity id="4CA99B352671FFC240CFFC9CFBCEFCA7" box="[992,1080,838,862]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="7" pageNumber="36" unit="mm" value="9.7">9.7 mm</quantity>
(RMNH-D40253),
<quantity id="4CA99B352671FFC24636FC9CFAA0FCA7" box="[1305,1366,838,862]" metricMagnitude="1" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="36" unit="m" value="42.0">42 m</quantity>
, grab, SW of São Tiago, coll.
<date id="FFEF10102671FFC2428AFCB0FDB3FC7B" box="[421,581,874,899]" pageId="7" pageNumber="36" value="1986-08-20">20 Aug. 1986</date>
;
<specimenCount id="9D57FD592671FFC2417DFCB0FD41FC7B" box="[594,695,874,898]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 15.0 ×
<quantity id="4CA99B352671FFC24039FCB0FC8AFC7B" box="[790,892,874,898]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="7" pageNumber="36" unit="mm" value="10.8">10.8 mm</quantity>
(RMNH-D40254), stn CANCAP 6.D11, scuba, Baia de São Pedro, SW coast of
<collectingCountry id="F34676402671FFC2412DFC54FD7AFC5F" box="[514,652,910,934]" name="Saint Vincent and the Grenadines" pageId="7" pageNumber="36">São Vicente</collectingCountry>
, coll.
<date id="FFEF10102671FFC241E0FC54FC94FC5F" box="[719,866,910,934]" pageId="7" pageNumber="36" value="1982-06-20">20 Jun. 1982</date>
.
</paragraph>
<paragraph id="8BEE36D02671FFC243EAFC68FCBAFC17" blockId="7.[151,1436,658,1007]" pageId="7" pageNumber="36">
<emphasis id="B925EAC22671FFC243EAFC68FEB8FC32" bold="true" box="[197,334,946,971]" pageId="7" pageNumber="36">Canary Is.:</emphasis>
<specimenCount id="9D57FD592671FFC24279FC68FE4AFC33" box="[342,444,946,970]" pageId="7" pageNumber="36" type="female">1 female</specimenCount>
, 19.8
<quantity id="4CA99B352671FFC2412DFC68FD9EFC33" box="[514,616,946,970]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="7" pageNumber="36" unit="mm" value="13.8">13.8 mm</quantity>
(RMNH-D40252), stn CANCAP 6.K01, rocky littoral, tidal pools, near los Christianos, SW Coast of Tenerife, coll. 27,
<date id="FFEF10102671FFC24180FC0CFCB1FC17" box="[687,839,982,1007]" pageId="7" pageNumber="36" value="1982-05-30">30 May 1982</date>
.
</paragraph>
</subSubSection>
<caption id="DF2E66582671FFC243B8F854FC7DF83A" httpUri="https://zenodo.org/record/207313/files/figure.png" pageId="7" pageNumber="36" targetBox="[151,1436,1060,1913]" targetPageId="7">
<paragraph id="8BEE36D02671FFC243B8F854FC7DF83A" blockId="7.[151,1436,1934,1987]" pageId="7" pageNumber="36">
<emphasis id="B925EAC22671FFC243B8F854FEE3F85D" bold="true" box="[151,277,1934,1957]" pageId="7" pageNumber="36">FIGURE 3.</emphasis>
<taxonomicName id="4C514D532671FFC24232F855FCB1F85C" authority="A. Milne-Edwards, 1878" authorityName="A. Milne-Edwards" authorityYear="1878" box="[285,839,1935,1957]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="B925EAC22671FFC24232F855FDDDF85C" box="[285,555,1935,1957]" italics="true" pageId="7" pageNumber="36">Glyptoxanthus cavernosus</emphasis>
(A. Milne-Edwards, 1878)
</taxonomicName>
, male, 49.6
<emphasis id="B925EAC22671FFC240E7F84AFC20F85C" box="[968,982,1936,1957]" italics="true" pageId="7" pageNumber="36">×</emphasis>
35.8 mm (MNHN-B8329), Cape Verde: A, dorsal view; B, anterior view; C, ventral view; D, left chela, external view.
</paragraph>
</caption>
<subSubSection id="C34B655B267EFFCD43E8FF4DFAC7FE55" pageId="8" pageNumber="37" type="diagnosis">
<paragraph id="8BEE36D0267EFFCD43E8FF4DFAC7FE55" blockId="8.[151,1436,151,860]" pageId="8" pageNumber="37">
<emphasis id="B925EAC2267EFFCD43E8FF4DFEB7FF49" bold="true" box="[199,321,151,176]" pageId="8" pageNumber="37">Diagnosis.</emphasis>
Carapace transversely ovate, width-to-length ratio about 1.4; carapace vermiculations thick, petaloid, like a pavement of lobules; lobules smooth without traces of individual granules; furrows between lobules deep, relatively narrow, setose; 2M completely divided longitudinally, inner branch separated from 1M by shallow transverse furrow; 3M independent from 2M, posterior region bridged medially with 4M; 2L, 3L, and 4L distinct, 5L fused with 6L; 1P distinct, independent, without any smaller cavities. Front quadrilobate, with deep lateral notches. Anterolateral margin with 4 feeble lobes. External surfaces of thoracic sternum, pereopods and abdomen with similar sculpturing as carapace. Margins of ambulatory legs fringed with long setae. G1 long, slender, bluntly tipped; apex devoid of any setae, but studded with spiniform granules; G2 about one-fourth length of G1.
</paragraph>
</subSubSection>
<subSubSection id="C34B655B267EFFCD43E8FE62FD51FCA5" pageId="8" pageNumber="37" type="discussion">
<paragraph id="8BEE36D0267EFFCD43E8FE62FB1EFD7D" blockId="8.[151,1436,151,860]" pageId="8" pageNumber="37">
<emphasis id="B925EAC2267EFFCD43E8FE62FECDFE29" bold="true" box="[199,315,440,464]" pageId="8" pageNumber="37">Remarks.</emphasis>
Alphonse
<bibRefCitation id="EFC04B21267EFFCD4294FE6DFD48FE36" author="Milne-Edwards" box="[443,702,439,463]" pageId="8" pageNumber="37" refString="Milne-Edwards, A. (1878) Description de quelques especes nouvelles de Crustaces provenant du voyage aux iles Cap-Vert de MM. Bouvier et de Cessac. Bulletin de la Societe philomatique (Paris), (ser. 7), 2, 225 - 232." type="journal article" year="1878">Milne-Edwards (1878)</bibRefCitation>
described
<taxonomicName id="4C514D53267EFFCD4010FE62FBFCFE29" box="[831,1034,440,464]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="cavernosa">
<emphasis id="B925EAC2267EFFCD4010FE62FBFCFE29" box="[831,1034,440,464]" italics="true" pageId="8" pageNumber="37">Actaea cavernosa</emphasis>
</taxonomicName>
from the
<collectingCountry id="F3467640267EFFCD47ACFE6DFAFEFE36" box="[1155,1288,439,463]" name="Cape Verde" pageId="8" pageNumber="37">Cape Verde</collectingCountry>
Islands, and later transferred it to his new genus,
<taxonomicName id="4C514D53267EFFCD4101FE06FD27FE0D" box="[558,721,476,500]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC2267EFFCD4101FE06FD27FE0D" box="[558,721,476,500]" italics="true" pageId="8" pageNumber="37">Glyptoxanthus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C514D53267EFFCD41F0FE06FBF1FE0D" box="[735,1031,476,500]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="B925EAC2267EFFCD41F0FE06FBF1FE0D" box="[735,1031,476,500]" italics="true" pageId="8" pageNumber="37">Glyptoxanthus cavernosus</emphasis>
</taxonomicName>
is easily distinguished from its congeners by the sculpturing on the dorsal surface of its carapace: in particular, the completely independent 3M region and the closely packed, non-elongate lobules and vermiculations, which have distinctly petaloid outlines. In fact, these vermiculations resemble the closely packed granules of some species of
<taxonomicName id="4C514D53267EFFCD473AFD92FB95FD99" box="[1045,1123,584,608]" class="Malacostraca" family="Xanthidae" genus="Actaea" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC2267EFFCD473AFD92FB95FD99" box="[1045,1123,584,608]" italics="true" pageId="8" pageNumber="37">Actaea</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC04B21267EFFCD478DFD9DFAC2FDA6" author="Guinot" box="[1186,1332,583,607]" pageId="8" pageNumber="37" refString="Guinot, D. (1979) Donnees nouvelles sur la morphologie, la phylogenese et la taxonomie des Crustaces Decapodes Brachyoures. Memoires du Museum national d'Histoire naturelle (Paris), (A) Zoologie, 112, 1 - 354, pls. 1 - 27." type="journal article" year="1979">Guinot 1979</bibRefCitation>
: fig. 10), and also of some
<taxonomicName id="4C514D53267EFFCD4274FDB6FE27FD7D" box="[347,465,620,644]" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC2267EFFCD4274FDB6FE27FD7D" box="[347,465,620,644]" italics="true" pageId="8" pageNumber="37">Euxanthus</emphasis>
</taxonomicName>
such as
<taxonomicName id="4C514D53267EFFCD411EFDB7FC89FD7D" authority="Rathbun, 1911" authorityName="Rathbun" authorityYear="1911" box="[561,895,620,644]" class="Malacostraca" family="Xanthidae" genus="Euxanthus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="boletarius">
<emphasis id="B925EAC2267EFFCD411EFDB7FD34FD7D" box="[561,706,620,644]" italics="true" pageId="8" pageNumber="37">E. boletarius</emphasis>
(
<bibRefCitation id="EFC04B21267EFFCD41FFFDB6FC80FD7D" author="Rathbun" box="[720,886,620,644]" pageId="8" pageNumber="37" refString="Rathbun, M. J. (1911) Marine Brachyura. In: The Percy Sladen Trust Expedition to the Indian Ocean in 1905 under the leadership of Mr. J. Stanley Gardiner. Vol. III. No. IX. Transactions of the Linnaean Society of London, Zoology, 14 (2), 191 - 261, pls. 15 - 20." type="journal article" year="1911">Rathbun, 1911</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="EFC04B21267EFFCD4095FDB6FBBFFD7D" author="Serene" box="[954,1097,620,644]" pageId="8" pageNumber="37" refString="Serene, R. (1984) Crustaces Decapodes Brachyoures de l'ocean Indien occidental et de la mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24, 1 - 400, Pls. 1 - 48." type="journal article" year="1984">Serène 1984</bibRefCitation>
: pl. 11 fig. E).
</paragraph>
<paragraph id="8BEE36D0267EFFCD43B8FD55FC69FD16" blockId="8.[151,1436,151,860]" pageId="8" pageNumber="37">
According to
<bibRefCitation id="EFC04B21267EFFCD4200FD55FE25FD5E" author="Monod" box="[303,467,655,679]" pageId="8" pageNumber="37" refString="Monod, T. (1956) Hippidea et Brachyura ouest-africaines. Memoires de l'Institut Fondamental d'Afrique Noire, 45, 1 - 674." type="journal article" year="1956">Monod (1956)</bibRefCitation>
, the
<typeStatus id="54EA8872267EFFCD4128FD4AFDC1FD51" box="[519,567,656,680]" pageId="8" pageNumber="37">type</typeStatus>
specimen was deposited in the MNHN, Paris, but it could not be found. However, this is a distinctive species that was well described and we have no doubt over the identity of our present material, most of which is from the
<typeStatus id="54EA8872267EFFCD4107FD02FDAEFD09" box="[552,600,728,752]" pageId="8" pageNumber="37">type</typeStatus>
locality,
<collectingCountry id="F3467640267EFFCD41EFFD0DFCB7FD16" box="[704,833,727,751]" name="Cape Verde" pageId="8" pageNumber="37">Cape Verde</collectingCountry>
Islands.
</paragraph>
<paragraph id="8BEE36D0267EFFCD43EAFD26FD51FCA5" blockId="8.[151,1436,151,860]" pageId="8" pageNumber="37">
<emphasis id="B925EAC2267EFFCD43EAFD26FD53FCED" bold="true" box="[197,677,763,788]" pageId="8" pageNumber="37">Ecology and geographical distribution.</emphasis>
<taxonomicName id="4C514D53267EFFCD419DFD26FC16FCED" box="[690,992,764,788]" class="Malacostraca" family="Xanthidae" genus="Glyptoxanthus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="cavernosus">
<emphasis id="B925EAC2267EFFCD419DFD26FC16FCED" box="[690,992,764,788]" italics="true" pageId="8" pageNumber="37">Glyptoxanthus cavernosus</emphasis>
</taxonomicName>
is only known from the
<collectingCountry id="F3467640267EFFCD463AFD26FA6AFCED" box="[1301,1436,764,788]" name="Cape Verde" pageId="8" pageNumber="37">Cape Verde</collectingCountry>
Islands and from the Canary Islands, off the northwestern coast of Africa, and has been collected from intertidal to shallow subtidal zones, amidst rocks and coral.
</paragraph>
</subSubSection>
</treatment>
</document>