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<document id="E7B96DE948DFCF30828AEADEAD2D3FB6" ID-CLB-Dataset="27734" ID-DOI="10.11646/zootaxa.4532.4.4" ID-GBIF-Dataset="4809e341-99ce-476f-a3a8-5229de62f5eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615629" ID-ZooBank="A35A92D5-2628-4203-8C94-DDBFFFCDC79E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788686464" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. &amp; Telnov, Dmitry" docDate="2018" docId="03F887BAFFAAFFDFFF37FC7D1C19FA21" docLanguage="en" docName="zootaxa.4532.4.4.pdf" docOrigin="Zootaxa 4532 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Distylopyga beccalonii Anisyutkin &amp; Telnov 2018, sp. nov." docType="treatment" docUuid="D2BE030B-0DE1-4573-9FC5-FD2336C5EECD" docUuidSource="ZooBank" docVersion="9" lastPageNumber="533" masterDocId="FFC1FFC2FFA2FFD5FFA0FFB4184CFE79" masterDocTitle="New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)" masterLastPageNumber="538" masterPageNumber="523" pageNumber="531" updateTime="1698715127610" updateUser="ExternalLinkService">
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<mods:title id="F516322130AD800412D3CE64E1035761">New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)</mods:title>
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<mods:namePart id="30C1A35B9DB2F517B9E794A6A1FF297E">Anisyutkin, Leonid N.</mods:namePart>
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<mods:namePart id="82BA51A57E3384099097CEEB9DA49D15">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="AC24F609580CFA086E1B36420FDE45A1">text</mods:typeOfResource>
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<mods:title id="C21CC89D2DB60E606817DC81AA3C72A7">Zootaxa</mods:title>
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<mods:date id="B4AD3504C62EE7C14BC6BAB191832CA4">2018</mods:date>
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<mods:number id="164A51482D5A9CBD340798537E49B48E">2018-12-20</mods:number>
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<mods:number id="E7DF776228B6C7CAFAA6FA4AEBCA8D97">4532</mods:number>
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<mods:identifier id="72E8329C8E9D994B6D11C6795CFBC5E9" type="DOI">10.11646/zootaxa.4532.4.4</mods:identifier>
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<treatment id="03F887BAFFAAFFDFFF37FC7D1C19FA21" ID-DOI="http://doi.org/10.5281/zenodo.5976584" ID-GBIF-Taxon="154521467" ID-Zenodo-Dep="5976584" LSID="urn:lsid:zoobank.org:act:D2BE030B-0DE1-4573-9FC5-FD2336C5EECD" httpUri="http://treatment.plazi.org/id/03F887BAFFAAFFDFFF37FC7D1C19FA21" lastPageId="10" lastPageNumber="533" pageId="8" pageNumber="531">
<subSubSection id="C34B6527FFAAFFDDFF37FC7D1AE9FD9A" box="[151,677,969,995]" pageId="8" pageNumber="531" type="nomenclature">
<paragraph id="8BEE36ACFFAAFFDDFF37FC7D1AE9FD9A" blockId="8.[151,1028,969,1064]" box="[151,677,969,995]" pageId="8" pageNumber="531">
<heading id="D0A681C0FFAAFFDDFF37FC7D1AE9FD9A" bold="true" box="[151,677,969,995]" fontSize="11" level="1" pageId="8" pageNumber="531" reason="1">
<emphasis id="B925EABEFFAAFFDDFF37FC7D1AE9FD9A" bold="true" box="[151,677,969,995]" pageId="8" pageNumber="531">
<taxonomicName id="4C514D2FFFAAFFDDFF37FC7D1A77FD9A" ID-CoL="8N97J" authority="Anisyutkin" authorityName="Anisyutkin &amp; Telnov" authorityYear="2018" box="[151,571,969,995]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="531" phylum="Arthropoda" rank="species" species="beccalonii" status="sp. nov.">
<emphasis id="B925EABEFFAAFFDDFF37FC7D19E4FD9A" bold="true" box="[151,424,969,995]" italics="true" pageId="8" pageNumber="531">Distylopyga beccalonii</emphasis>
Anisyutkin
</taxonomicName>
,
<taxonomicNameLabel id="A21657C5FFAAFFDDFDE8FC7D1AE9FD9A" box="[584,677,969,995]" pageId="8" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34B6527FFAAFFDDFF37FC581C48FA51" pageId="8" pageNumber="531" type="description">
<paragraph id="8BEE36ACFFAAFFDDFF37FC581987FA7C" blockId="8.[151,1028,969,1064]" box="[151,459,1004,1029]" pageId="8" pageNumber="531">
(
<figureCitation id="136A2A29FFAAFFDDFF3FFC5818A6FA7D" box="[159,234,1004,1029]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="8" pageNumber="531">Figs. 3</figureCitation>
EH, 4AN, 5AE)
</paragraph>
<paragraph id="8BEE36ACFFAAFFDDFF37FBBB1C48FA51" blockId="8.[151,1028,969,1064]" box="[151,1028,1039,1064]" pageId="8" pageNumber="531">
Zoobank: http://zoobank.org/
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</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFAAFFDDFF37FBE31A0EFAA5" pageId="8" pageNumber="531" type="materials_examined">
<paragraph id="8BEE36ACFFAAFFDDFF37FBE31A0EFAA5" blockId="8.[151,1438,1111,2037]" pageId="8" pageNumber="531">
<emphasis id="B925EABEFFAAFFDDFF37FBE318B3FA09" bold="true" box="[151,255,1111,1136]" pageId="8" pageNumber="531">Material</emphasis>
:
<materialsCitation id="3B393CF1FFAAFFDDFEB1FBE31A0EFAA5" ID-GBIF-Occurrence="2120066958" collectingDate="1996-07-30" collectingDateMax="1996-08-05" collectingDateMin="1996-07-30" collectionCode="ZIN" collectorName="A. Gorochov" country="Thailand" elevation="400" location="National Park Kaeng Krachan" pageId="8" pageNumber="531" specimenCount="5" specimenCount-female="5" stateProvince="Phetchaburi" typeStatus="holotype">
<typeStatus id="54EA880EFFAAFFDDFEB1FBE31931FA09" box="[273,381,1111,1136]" pageId="8" pageNumber="531" type="holotype">Holotype</typeStatus>
. male,
<collectingCountry id="F346763CFFAAFFDDFE72FBEC1A2AFA09" box="[466,614,1112,1136]" name="Thailand" pageId="8" pageNumber="531">THAILAND</collectingCountry>
, prov.
<collectingRegion id="4995F84EFFAAFFDDFD17FBE31B0CFA09" box="[695,832,1111,1136]" country="Thailand" name="Phetchaburi" pageId="8" pageNumber="531">Phetchaburi</collectingRegion>
,
<quantity id="4CA99B49FFAAFFDDFCEEFBE31BD4FA09" box="[846,920,1111,1136]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="531" unit="km" value="50.0">50 km</quantity>
SW town
<collectingRegion id="4995F84EFFAAFFDDFBB8FBE31CEDFA09" box="[1048,1185,1111,1136]" country="Thailand" name="Phetchaburi" pageId="8" pageNumber="531">Phetchaburi</collectingRegion>
, environs of
<location id="8E8E6077FFAAFFDDFA9CFBE319C1FAEC" LSID="urn:lsid:plazi:treatment:03F887BAFFAAFFDFFF37FC7D1C19FA21:8E8E6077FFAAFFDDFA9CFBE319C1FAEC" country="Thailand" name="National Park Kaeng Krachan" pageId="8" pageNumber="531" stateProvince="Phetchaburi">National Park Kaeng Krachan</location>
, forest near reservoir, h=
<quantity id="4CA99B49FFAAFFDDFD17FBC81B4AFAED" box="[695,774,1148,1173]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="531" unit="m" value="400.0">
<elevation id="007CD19FFFAAFFDDFD17FBC81B4AFAED" box="[695,774,1148,1173]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="531" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="FFEF106CFFAAFFDDFCB7FBC81B9EFAEC" box="[791,978,1148,1173]" pageId="8" pageNumber="531" value="1996-07-30" valueMax="1996-07-31" valueMin="1996-07-30">
<collectingDate id="EFABE984FFAAFFDDFCB7FBC81B9EFAEC" box="[791,978,1148,1173]" pageId="8" pageNumber="531" value="1996-07-30" valueMax="1996-07-31" valueMin="1996-07-30">3031 July1996</collectingDate>
</date>
, leg.
<collectorName id="26A4537AFFAAFFDDFBB8FBC81CE3FAEC" box="[1048,1199,1148,1173]" pageId="8" pageNumber="531">A. Gorochov</collectorName>
(
<collectionCode id="ED40AE69FFAAFFDDFB63FBC81CBBFAED" box="[1219,1271,1148,1172]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="531" type="Museum">ZIN</collectionCode>
);
<typeStatus id="54EA880EFFAAFFDDFAB0FBC91D33FAEC" box="[1296,1407,1149,1173]" pageId="8" pageNumber="531" type="paratype">paratypes</typeStatus>
.
<specimenCount id="9D57FD25FFAAFFDDFA2EFBC818BDFAC1" pageId="8" pageNumber="531" type="female">3 females</specimenCount>
, 1 larva, same data as holotype, but
<date id="FFEF106CFFAAFFDDFD00FB141BE2FAC1" box="[672,942,1183,1208]" pageId="8" pageNumber="531" value="1996-07-30" valueMax="1996-08-01" valueMin="1996-07-30">
<collectingDate id="EFABE984FFAAFFDDFD00FB141BE2FAC1" box="[672,942,1183,1208]" pageId="8" pageNumber="531" value="1996-07-30" valueMax="1996-08-01" valueMin="1996-07-30">30 July1 August 1996</collectingDate>
</date>
(
<collectionCode id="ED40AE69FFAAFFDDFC60FB141BB8FAC1" box="[960,1012,1184,1208]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="531" type="Museum">ZIN</collectionCode>
);
<typeStatus id="54EA880EFFAAFFDDFBABFB141C36FAC1" box="[1035,1146,1184,1208]" pageId="8" pageNumber="531" type="paratype">paratypes</typeStatus>
.
<specimenCount id="9D57FD25FFAAFFDDFB28FB141CB6FAC1" box="[1160,1274,1183,1208]" pageId="8" pageNumber="531" type="female">2 females</specimenCount>
, same data as
<typeStatus id="54EA880EFFAAFFDDFF37FB7018B0FAA4" box="[151,252,1220,1245]" pageId="8" pageNumber="531" type="holotype">holotype</typeStatus>
, but
<date id="FFEF106CFFAAFFDDFE92FB7019BFFAA4" box="[306,499,1220,1245]" pageId="8" pageNumber="531" value="1996-08-03" valueMax="1996-08-05" valueMin="1996-08-03">
<collectingDate id="EFABE984FFAAFFDDFE92FB7019BFFAA4" box="[306,499,1220,1245]" pageId="8" pageNumber="531" value="1996-08-03" valueMax="1996-08-05" valueMin="1996-08-03">35 August 1996</collectingDate>
</date>
(
<collectionCode id="ED40AE69FFAAFFDDFDA2FB701A7AFAA5" box="[514,566,1220,1244]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="531" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFAAFFDDFF67FB531DD2FB5C" pageId="8" pageNumber="531" type="etymology">
<paragraph id="8BEE36ACFFAAFFDDFF67FB531DD2FB5C" blockId="8.[151,1438,1111,2037]" pageId="8" pageNumber="531">
<emphasis id="B925EABEFFAAFFDDFF67FB53190FFB79" bold="true" box="[199,323,1255,1280]" pageId="8" pageNumber="531">Etymology</emphasis>
. Patronymic. This species is named in honour of Dr. George Beccaloni (the Alfred Russel Wallace Trust,
<collectingCountry id="F346763CFFAAFFDDFF7CFAB819DBFB5C" box="[220,407,1292,1317]" name="United Kingdom" pageId="8" pageNumber="531">United Kingdom</collectingCountry>
), famous specialist in cockroach taxonomy and establisher of Cockroach Species File database.
</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFAAFFDFFF67FA9B1C19FA21" lastPageId="10" lastPageNumber="533" pageId="8" pageNumber="531" type="description">
<paragraph id="8BEE36ACFFAAFFDCFF67FA9B1BECFF46" blockId="8.[151,1438,1111,2037]" lastBlockId="9.[151,1437,151,320]" lastPageId="9" lastPageNumber="532" pageId="8" pageNumber="531">
<emphasis id="B925EABEFFAAFFDDFF67FA9B191DFB31" bold="true" box="[199,337,1327,1352]" pageId="8" pageNumber="531">Description</emphasis>
. Male (the
<typeStatus id="54EA880EFFAAFFDDFE7AFA9B1A0DFB31" box="[474,577,1327,1352]" pageId="8" pageNumber="531" type="holotype">holotype</typeStatus>
). Body generally reddish brown, head slightly darker; eyes black; areas of ocellar spots pale yellowish (
<figureCitation id="136A2A29FFAAFFDDFE7AFAE01A78FB15" box="[474,564,1364,1389]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4A</figureCitation>
); scapus and pedicellum, labrum, anteclypeus, maxillary and labial palps dirty yellow; antennae (scapus and pedicellum excluded), greyish brown; legs yellowish brown. Surfaces smooth and lustrous; punctation very delicate; facial part of head with delicate wrinkles above clypeus (not shown in
<figureCitation id="136A2A29FFAAFFDDFA95FA281DC3FBCD" box="[1333,1423,1436,1461]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4A</figureCitation>
); distal antennomeres dull (approximately from 10th segment). Head rounded at vertex, longer than wide, convex (
<figureCitation id="136A2A29FFAAFFDDFF3FFA5018B1FB85" box="[159,253,1508,1533]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4A</figureCitation>
); epicranial sutures not indicated; eyes small; ocellar spots small, visible only as light spots; distance between eyes 1.7 times eye length; distance between antennal sockets 2.4 times of the scape length (about
<quantity id="4CA99B49FFAAFFDDFA96F9B31DDCF859" box="[1334,1424,1543,1568]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="531" unit="mm" value="0.5">0.5 mm</quantity>
); approximate length ratio of 3rd5th segments of maxillary palpomeres 1.0: 1.0: 1.3. Pronotum campaniform, wider than long, caudall margin slightly angulate (
<figureCitation id="136A2A29FFAAFFDDFD72F9FB1B67F811" box="[722,811,1615,1640]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4B</figureCitation>
). Tegmina reduced to lateral flaps, marginated laterally and truncated apically (
<figureCitation id="136A2A29FFAAFFDDFE3CF9C01A58F8F5" box="[412,532,1652,1677]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4B, C</figureCitation>
). Wings absent. Mesonotum and metanotum transverse, dorsally slightly convex, with posterior margins slightly angulate (
<figureCitation id="136A2A29FFAAFFDDFDDEF9231A90F8C9" box="[638,732,1687,1712]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4B</figureCitation>
). Thoracal tergites and abdominal tergites IIVII marginate laterally (
<figureCitation id="136A2A29FFAAFFDDFEA2F9081930F8AD" box="[258,380,1724,1749]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4B, C</figureCitation>
). Anterior margin of fore femur armed as in the type A, with 1415 spines, including 3 apical ones. Fore tibiae not thickened distally. Tibial spines well developed. Structure of hind tarsi (
<figureCitation id="136A2A29FFAAFFDDFAAFF96B1D15F881" box="[1295,1369,1759,1784]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4</figureCitation>
DF): metatarsus a little longer than other segments combined; all euplantulae small and apical (
<figureCitation id="136A2A29FFAAFFDDFB34F8B01D42F965" box="[1172,1294,1796,1821]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4D, E</figureCitation>
); metatarsus and 2nd metatarsomere with 2 more or less equal rows of spines along its lower margin; 3rd metatarsomere with 2 spines along lower margin; metatarsus and 2nd4th metatarsomeres with 12 &quot;additional spines&quot; bordering euplantulae from inside and outside (
<figureCitation id="136A2A29FFAAFFDDFD93F8DB1A9BF9F1" box="[563,727,1903,1928]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">
Fig. 4D, E,
<emphasis id="B925EABEFFAAFFDDFD15F8C51A9BF9F1" box="[693,727,1905,1928]" italics="true" pageId="8" pageNumber="531">a.s</emphasis>
</figureCitation>
.); claws symmetrical, simple; arolium smaller than half of claw length (
<figureCitation id="136A2A29FFAAFFDDFF4FF8201904F9D5" box="[239,328,1940,1965]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4F</figureCitation>
). Fore- and mid tarsi similar to metatarsi, but with comparatively shorter tarsomeres. Abdomen without visible glandular specializations; posterolateral angles attenuate caudally; tergite VI with caudal margin straight (
<figureCitation id="136A2A29FFAAFFDDFF5EF8681916F98D" box="[254,346,2012,2037]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="8" pageNumber="531">Fig. 4B</figureCitation>
); tergite VII roundly projected caudally. Anal plate (tergite X) short and transverse, medially projected, with very weak triangular median incision on caudal margin (
<figureCitation id="136A2A29FFABFFDCFC7DFF231C76FEC9" box="[989,1082,151,176]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="9" pageNumber="532">Fig. 4G</figureCitation>
). Cerci fusiform and flat, with solidly connected segments (
<figureCitation id="136A2A29FFABFFDCFE49FF081A05FEAD" box="[489,585,188,213]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="9" pageNumber="532">Fig. 4H</figureCitation>
). Paraprocts weakly asymmetrical, sclerotized along caudal margin; pvsclerites well sclerotized, elongated (
<figureCitation id="136A2A29FFABFFDCFDE1FF6B1AD7FE81" box="[577,667,223,248]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="9" pageNumber="532">Fig. 4G</figureCitation>
,
<emphasis id="B925EABEFFABFFDCFD09FF551A85FE81" box="[681,713,225,248]" italics="true" pageId="9" pageNumber="532">pv.</emphasis>
). Hypandrium weakly asymmetrical, wide, antero-lateral parts (lateral sternal apodemes or apophyses) short; caudal margin sinuate between styli (
<figureCitation id="136A2A29FFABFFDCFBE0FEB01CCEFF65" box="[1088,1154,260,285]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="9" pageNumber="532">Fig. 4</figureCitation>
IK); styli asymmetrical: left stylus with 4 small curved spines, right stylus with 2 longer ones.
</paragraph>
<caption id="DF2E6624FFABFFDCFF37F8271B9EF994" httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="9" pageNumber="532" startId="9.[151,249,1939,1961]" targetBox="[191,1409,352,1920]" targetPageId="9">
<paragraph id="8BEE36ACFFABFFDCFF37F8271B9EF994" blockId="9.[151,1436,1939,2030]" pageId="9" pageNumber="532">
<emphasis id="B925EABEFFABFFDCFF37F8271958F9D0" bold="true" box="[151,276,1939,1961]" pageId="9" pageNumber="532">FIGURE 5.</emphasis>
<taxonomicName id="4C514D2FFFABFFDCFEBCF8271AC8F9D0" authority="Anisyutkin" authorityName="Anisyutkin &amp; Telnov" authorityYear="2018" box="[284,644,1939,1961]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="9" pageNumber="532" phylum="Arthropoda" rank="species" species="beccalonii" status="sp. nov.">
<emphasis id="B925EABEFFABFFDCFEBCF8271A45F9D0" box="[284,521,1939,1961]" italics="true" pageId="9" pageNumber="532">Distylopyga beccalonii</emphasis>
Anisyutkin
</taxonomicName>
,
<emphasis id="B925EABEFFABFFDCFD30F8271AAEF9D0" bold="true" box="[656,738,1939,1961]" pageId="9" pageNumber="532">
<taxonomicNameLabel id="A21657C5FFABFFDCFD30F8271AAEF9D0" box="[656,738,1939,1961]" pageId="9" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations:
<emphasis id="B925EABEFFABFFDCFE94F86D1AF6F994" box="[308,698,2007,2030]" italics="true" pageId="9" pageNumber="532">a.p., a.t., R1G, R1Hi., R1Hout., R2, R3</emphasis>
—see text. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BEE36ACFFA8FFDFFF67FF231981FCFC" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="533">
Male genitalia (
<figureCitation id="136A2A29FFA8FFDFFEDDFF231987FEC9" box="[381,459,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Figs. 3</figureCitation>
EH, 5AE): left phallomere (
<figureCitation id="136A2A29FFA8FFDFFC8DFF231B3CFEC9" box="[813,880,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Fig. 3</figureCitation>
EH) with sclerite L4C (L2D) long and narrow, caudally divided into two partly membranous parts, outer one terminating in oppositely directed sharp teeth (
<figureCitation id="136A2A29FFA8FFDFFACFFF08196DFE81" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">
Fig. 3E, G,
<emphasis id="B925EABEFFA8FFDFFF48FF55196DFE81" box="[232,289,225,248]" italics="true" pageId="10" pageNumber="533">a.c.p</emphasis>
</figureCitation>
.); sclerite L3 (L3d) curved in apical part, with small cranially directed tooth; sclerite L4F, possibly L4E+L4F, large, with indistinct margins (
<figureCitation id="136A2A29FFA8FFDFFDD6FEB01A82FF65" box="[630,718,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Fig. 3F</figureCitation>
); sclerite L4D (L
<date id="FFEF106CFFA8FFDFFC39FEB01BF9FF65" box="[921,949,260,284]" pageId="10" pageNumber="533">3v</date>
) elongated (
<figureCitation id="136A2A29FFA8FFDFFBE6FEB01C8DFF65" box="[1094,1217,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Fig. 3F, G</figureCitation>
); sclerite L2 (L
<date id="FFEF106CFFA8FFDFFAD7FEB01DD8FF65" box="[1399,1428,260,284]" pageId="10" pageNumber="533">2v</date>
) long, narrowed in cranial part (
<figureCitation id="136A2A29FFA8FFDFFE4FFE931A2AFF39" box="[495,614,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Fig. 3G, H</figureCitation>
), terminating in 2 acute caudal processes: inner process short and apically forked, outer—long and slender (
<figureCitation id="136A2A29FFA8FFDFFDA8FEF81A9CFF1D" box="[520,720,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">
Fig. 3E, G, H,
<emphasis id="B925EABEFFA8FFDFFD09FEF91A9CFF1D" box="[681,720,333,356]" italics="true" pageId="10" pageNumber="533">a.p</emphasis>
</figureCitation>
.); &quot;additional elongated sclerite&quot;, probably separated part of L2, situated on inner side of phallomere dorsally to L2 (
<figureCitation id="136A2A29FFA8FFDFFD4EFEDB1BC4FFF1" box="[750,904,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">
Fig. 3G,
<emphasis id="B925EABEFFA8FFDFFCF3FEC51BC4FFF1" box="[851,904,369,392]" italics="true" pageId="10" pageNumber="533">a.e.s</emphasis>
</figureCitation>
.), terminating long and slender caudal process (
<figureCitation id="136A2A29FFA8FFDFFF3FFE2018ACFFD5" box="[159,224,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Fig. 3</figureCitation>
EF,
<emphasis id="B925EABEFFA8FFDFFEBDFE201919FFD5" box="[285,341,404,428]" italics="true" pageId="10" pageNumber="533">f.c.p.</emphasis>
) and subtriangular membranous lobe (
<figureCitation id="136A2A29FFA8FFDFFCA8FE201BFFFFD5" box="[776,947,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">
Fig. 3G, H,
<emphasis id="B925EABEFFA8FFDFFC2DFE201BFFFFD5" box="[909,947,404,428]" italics="true" pageId="10" pageNumber="533">m.l</emphasis>
</figureCitation>
.); &quot;additional elongated sclerite&quot; connected with complicate sclerite with dorso-caudal projection (
<figureCitation id="136A2A29FFA8FFDFFD5EFE031BD6FFA9" box="[766,922,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">
Fig. 3G,
<emphasis id="B925EABEFFA8FFDFFCC0FE0C1BD6FFA9" box="[864,922,440,464]" italics="true" pageId="10" pageNumber="533">d.c.p</emphasis>
</figureCitation>
.); sclerite L1 absent. Ventral phallomere L4G (VP) short and wide (
<figureCitation id="136A2A29FFA8FFDFFE26FE6819B2FF8D" box="[390,510,476,501]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">Fig. 5D, E</figureCitation>
). Right phallomere complex in shape, as in
<figureCitation id="136A2A29FFA8FFDFFC46FE681C69FF8C" box="[998,1061,476,501]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">Fig. 5</figureCitation>
AC; basal sclerite (R2) small, as compared with other sclerites of right phallomere; sclerite R1H divided into 2 branches: inner branches apically forked (
<figureCitation id="136A2A29FFA8FFDFFF50FD901924FC45" box="[240,360,548,573]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">
Fig. 5A,
<emphasis id="B925EABEFFA8FFDFFEF6FD901924FC45" box="[342,360,548,572]" italics="true" pageId="10" pageNumber="533">R</emphasis>
</figureCitation>
<figureCitation id="136A2A29FFA8FFDFFEC8FD90193FFC45" box="[360,371,548,572]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="10" pageNumber="533">1</figureCitation>
Hi), outer with apex bent down (
<figureCitation id="136A2A29FFA8FFDFFD46FD901B13FC45" box="[742,863,548,573]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">
Fig. 5A,
<emphasis id="B925EABEFFA8FFDFFCECFD901B13FC45" box="[844,863,548,572]" italics="true" pageId="10" pageNumber="533">R</emphasis>
</figureCitation>
<figureCitation id="136A2A29FFA8FFDFFCFFFD901B26FC45" box="[863,874,548,572]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="10" pageNumber="533">1</figureCitation>
Hout.), additional long spine-like process situated above outer branch (
<figureCitation id="136A2A29FFA8FFDFFE21FDF31A43FC19" box="[385,527,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">
Fig. 5A,
<emphasis id="B925EABEFFA8FFDFFE48FDFD1A43FC19" box="[488,527,585,608]" italics="true" pageId="10" pageNumber="533">a.p</emphasis>
</figureCitation>
.); sclerite R1G subtriangular in shape, with blunt apical tooth (
<figureCitation id="136A2A29FFA8FFDFFB7EFDF31DCBFC19" box="[1246,1415,583,608]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="10" pageNumber="533">
Fig. 5A, B,
<emphasis id="B925EABEFFA8FFDFFAC7FDFD1DCBFC19" box="[1383,1415,585,608]" italics="true" pageId="10" pageNumber="533">a.t</emphasis>
</figureCitation>
.); sclerite R3 large, plate-like.
</paragraph>
<paragraph id="8BEE36ACFFA8FFDFFF67FD3B1DC3FD41" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="533">
<emphasis id="B925EABEFFA8FFDFFF67FD3B1952FCD1" bold="true" box="[199,286,655,680]" pageId="10" pageNumber="533">Female</emphasis>
. Similar to male. Maxillary and labial palps sometimes grey, legs—reddish brown. Head as in
<figureCitation id="136A2A29FFA8FFDFFAE1FD3B1DDBFCD1" box="[1345,1431,655,680]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="10" pageNumber="533">Fig. 4L</figureCitation>
; distance between eyes 1.61.7 times eye length; distance between antennal sockets 2.12.5 times of the scape length (about
<quantity id="4CA99B49FFA8FFDFFE97FD631988FC89" box="[311,452,727,752]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="533" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
); approximate length ratio of 3rd5th segments of maxillary palps 1.0: 1.0: 1.31.4. Thoracal and abdominal tergites as in
<figureCitation id="136A2A29FFA8FFDFFDE7FD481AEAFD6D" box="[583,678,764,789]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="10" pageNumber="533">
Fig.
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. Anterior margin of fore femur armed as in the
<typeStatus id="54EA880EFFA8FFDFFB62FD491CBEFD6C" box="[1218,1266,765,789]" pageId="10" pageNumber="533">type</typeStatus>
A, with 1415 spines, including 23 apical ones. Abdominal apex as in
<figureCitation id="136A2A29FFA8FFDFFCB4FCAB1B3DFD41" box="[788,881,799,824]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="10" pageNumber="533">
Fig.
<specimenCount id="9D57FD25FFA8FFDFFCE7FC941B3DFD41" box="[839,881,800,824]" pageId="10" pageNumber="533" type="generic">4M</specimenCount>
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. Genital plate transverse, valves large (
<figureCitation id="136A2A29FFA8FFDFFA88FCAB1DCEFD41" box="[1320,1410,799,824]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="10" pageNumber="533">Fig. 4N</figureCitation>
).
</paragraph>
<paragraph id="8BEE36ACFFA8FFDFFF67FCF018A5FDF9" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="533">
<emphasis id="B925EABEFFA8FFDFFF67FCF0195CFD24" bold="true" box="[199,272,836,861]" pageId="10" pageNumber="533">Larva</emphasis>
. Similar to adult male, but paler: antennae, anteclypeus, maxillary and labial palps and partly legs yellow.
</paragraph>
<paragraph id="8BEE36ACFFA8FFDFFF67FC381BE0FA69" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="533">
<emphasis id="B925EABEFFA8FFDFFF67FC38193AFDDC" bold="true" box="[199,374,908,933]" pageId="10" pageNumber="533">Measurements</emphasis>
(mm). Head length: male 2.5, female 2.83.0; head width: male 2.3, female 2.42.6; pronotum length: male 3.3, female 3.43.7; pronotum width: male 4.5, female 4.44.7; mesonotum length: male 1.7, female 1.82.0; tegmina length: male 1.9, female 1.62.0; tegmina width: male 1.1, female 0.91.1; metanotum length: male 1.2, female 1.31.6; metanotum width: male 5.1, female 5.05.4.
</paragraph>
<paragraph id="8BEE36ACFFA8FFDFFF67FBA81C19FA21" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="533">
<emphasis id="B925EABEFFA8FFDFFF67FBA81916FA4C" bold="true" box="[199,346,1052,1077]" pageId="10" pageNumber="533">Comparison</emphasis>
. The new species differs from
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<emphasis id="B925EABEFFA8FFDFFD76FBA91AA6FA4D" box="[726,746,1053,1076]" italics="true" pageId="10" pageNumber="533">D</emphasis>
.
<emphasis id="B925EABEFFA8FFDFFD5DFBA91B00FA4D" box="[765,844,1053,1076]" italics="true" pageId="10" pageNumber="533">semoni</emphasis>
</taxonomicName>
in generally paler body (black in
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<emphasis id="B925EABEFFA8FFDFFB4DFBA81CB7FA4A" box="[1261,1275,1052,1075]" italics="true" pageId="10" pageNumber="533">S</emphasis>
.
<emphasis id="B925EABEFFA8FFDFFAAEFBA91D11FA4D" box="[1294,1373,1053,1076]" italics="true" pageId="10" pageNumber="533">semoni</emphasis>
</taxonomicName>
) and distinctly angulate caudal margin of metanotum (
<figureCitation id="136A2A29FFA8FFDFFD1EFB8B1B58FA21" box="[702,788,1087,1112]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="10" pageNumber="533">Fig. 4B</figureCitation>
) (emarginated in
<taxonomicName id="4C514D2FFFA8FFDFFC7AFBF41C09FA2E" baseAuthorityName="Krauss" baseAuthorityYear="1903" box="[986,1093,1088,1111]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="533" phylum="Arthropoda" rank="species" species="semoni">
<emphasis id="B925EABEFFA8FFDFFC7AFBF41BA4FA2E" box="[986,1000,1088,1111]" italics="true" pageId="10" pageNumber="533">S</emphasis>
.
<emphasis id="B925EABEFFA8FFDFFC56FBF41C09FA2E" box="[1014,1093,1088,1111]" italics="true" pageId="10" pageNumber="533">semoni</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>