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<document id="2328233076BAB112BD4FA4403A2ECEF5" ID-CLB-Dataset="27734" ID-DOI="10.11646/zootaxa.4532.4.4" ID-GBIF-Dataset="4809e341-99ce-476f-a3a8-5229de62f5eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615629" ID-ZooBank="A35A92D5-2628-4203-8C94-DDBFFFCDC79E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788686464" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. &amp; Telnov, Dmitry" docDate="2018" docId="03F887BAFFA0FFD6FF37FD7B19C0FB9C" docLanguage="en" docName="zootaxa.4532.4.4.pdf" docOrigin="Zootaxa 4532 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Duchailluia ivindo Anisyutkin &amp; Telnov 2018, sp. nov." docType="treatment" docUuid="62E3B78F-963C-4A80-A4A9-D6C5A37A06BE" docUuidSource="ZooBank" docVersion="9" lastPageNumber="526" masterDocId="FFC1FFC2FFA2FFD5FFA0FFB4184CFE79" masterDocTitle="New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)" masterLastPageNumber="538" masterPageNumber="523" pageNumber="525" updateTime="1698715127610" updateUser="ExternalLinkService">
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<mods:title id="526DF55E921FF939F2C5C66157BE39B4">New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)</mods:title>
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<mods:name id="61EE90059E4ABDE72A591BA4E5807352" type="personal">
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<mods:namePart id="AABA99B074288698C1FBE1187A71826B">Anisyutkin, Leonid N.</mods:namePart>
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<mods:namePart id="E09E183561936ED34C4EA6DDB3E15D98">Telnov, Dmitry</mods:namePart>
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<mods:typeOfResource id="1CC0DEE39447CF552B604565DFB2B5FD">text</mods:typeOfResource>
<mods:relatedItem id="FA6672D946D24D7F009294D13B1FCFFB" type="host">
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<mods:title id="3194A9D5772B7F1672FA83F5ACB581BE">Zootaxa</mods:title>
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<mods:date id="AF5E3E572076A55C6DE9600629036A7E">2018</mods:date>
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<mods:number id="C30EFACE0E8BAB139C56853247FD75A8">2018-12-20</mods:number>
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<mods:number id="67F6223632066BAA37C077903BF949AA">4532</mods:number>
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<mods:identifier id="AD47B5D8D86191B07219807493D119FD" type="DOI">10.11646/zootaxa.4532.4.4</mods:identifier>
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<treatment id="03F887BAFFA0FFD6FF37FD7B19C0FB9C" ID-DOI="http://doi.org/10.5281/zenodo.5976572" ID-GBIF-Taxon="154521473" ID-Zenodo-Dep="5976572" LSID="urn:lsid:zoobank.org:act:62E3B78F-963C-4A80-A4A9-D6C5A37A06BE" httpUri="http://treatment.plazi.org/id/03F887BAFFA0FFD6FF37FD7B19C0FB9C" lastPageId="3" lastPageNumber="526" pageId="2" pageNumber="525">
<subSubSection id="C34B6527FFA0FFD7FF37FD7B1A34FC93" box="[151,632,719,746]" pageId="2" pageNumber="525" type="nomenclature">
<paragraph id="8BEE36ACFFA0FFD7FF37FD7B1A34FC93" blockId="2.[151,1028,719,815]" box="[151,632,719,746]" pageId="2" pageNumber="525">
<heading id="D0A681C0FFA0FFD7FF37FD7B1A34FC93" bold="true" box="[151,632,719,746]" fontSize="11" level="1" pageId="2" pageNumber="525" reason="1">
<emphasis id="B925EABEFFA0FFD7FF37FD7B1A34FC93" bold="true" box="[151,632,719,746]" pageId="2" pageNumber="525">
<taxonomicName id="4C514D2FFFA0FFD7FF37FD7B1A41FC93" ID-CoL="8N97K" authority="Anisyutkin" authorityName="Anisyutkin &amp; Telnov" authorityYear="2018" box="[151,525,719,746]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="525" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
<emphasis id="B925EABEFFA0FFD7FF37FD7B1936FC90" bold="true" box="[151,378,719,745]" italics="true" pageId="2" pageNumber="525">Duchailluia ivindo</emphasis>
Anisyutkin
</taxonomicName>
,
<taxonomicNameLabel id="A21657C5FFA0FFD7FDBAFD641A34FC93" box="[538,632,720,746]" pageId="2" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34B6527FFA0FFD7FF37FD461C48FD56" pageId="2" pageNumber="525" type="description">
<paragraph id="8BEE36ACFFA0FFD7FF37FD46199DFD72" blockId="2.[151,1028,719,815]" box="[151,465,754,779]" pageId="2" pageNumber="525">
(
<figureCitation id="136A2A29FFA0FFD7FF3FFD4618ABFD72" box="[159,231,754,779]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Figs. 1</figureCitation>
AK, 2AD, 3AD)
</paragraph>
<paragraph id="8BEE36ACFFA0FFD7FF37FCA21C48FD56" blockId="2.[151,1028,719,815]" box="[151,1028,790,815]" pageId="2" pageNumber="525">
Zoobank: http://zoobank.org/
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</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFA0FFD7FF37FCEA1C0EFD9A" pageId="2" pageNumber="525" type="materials_examined">
<paragraph id="8BEE36ACFFA0FFD7FF37FCEA1C0EFD9A" blockId="2.[151,1437,862,2009]" pageId="2" pageNumber="525">
<emphasis id="B925EABEFFA0FFD7FF37FCEA18B3FD0E" bold="true" box="[151,255,862,887]" pageId="2" pageNumber="525">Material</emphasis>
:
<materialsCitation id="3B393CF1FFA0FFD7FEB3FCEA1C0EFD9A" ID-GBIF-Occurrence="2120080606" collectingDate="2016-06-14" collectingDateMax="2016-06-15" collectingDateMin="2016-06-14" collectionCode="NME, ZIN" collectorName="Ivindo NP &amp; D. Telnov" country="Gabon" elevation="510" latitude="0.5144445" location="Ogooue-lvindo" longLatPrecision="21" longitude="12.805834" municipality="Makokou" pageId="2" pageNumber="525" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54EA880EFFA0FFD7FEB3FCEA1933FD0E" box="[275,383,862,887]" pageId="2" pageNumber="525" type="holotype">Holotype</typeStatus>
. male,
<collectingCountry id="F346763CFFA0FFD7FE77FCEB1A77FD0E" box="[471,571,863,887]" name="Gabon" pageId="2" pageNumber="525">GABON</collectingCountry>
E, prov.
<location id="8E8E6077FFA0FFD7FD0FFCEA1B17FD0E" LSID="urn:lsid:plazi:treatment:03F887BAFFA0FFD6FF37FD7B19C0FB9C:8E8E6077FFA0FFD7FD0FFCEA1B17FD0E" box="[687,859,862,887]" country="Gabon" latitude="0.5144445" longLatPrecision="21" longitude="12.805834" municipality="Makokou" name="Ogooue-lvindo" pageId="2" pageNumber="525">Ogooué-lvindo</location>
,
<collectingMunicipality id="6B8AACD6FFA0FFD7FCCCFCEA1B9BFD0E" box="[876,983,862,887]" pageId="2" pageNumber="525">Makokou</collectingMunicipality>
<quantity id="4CA99B49FFA0FFD7FC43FCEA1C71FD0E" box="[995,1085,862,887]" metricMagnitude="3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="2" pageNumber="525" unit="km" value="7.0" valueMax="9.0" valueMin="5.0">59 km</quantity>
SW,
<collectorName id="26A4537AFFA0FFD7FB23FCEA1CB0FD0F" box="[1155,1276,862,887]" pageId="2" pageNumber="525">Ivindo NP</collectorName>
, lpassa forest research station (
<geoCoordinate id="EE65506BFFA0FFD7FEFCFC361998FDE2" box="[348,468,898,923]" direction="north" orientation="latitude" pageId="2" pageNumber="525" precision="15" value="0.5144445">0°30'52''N</geoCoordinate>
,
<geoCoordinate id="EE65506BFFA0FFD7FE42FC371A29FDE2" box="[482,613,899,923]" direction="east" orientation="longitude" pageId="2" pageNumber="525" precision="15" value="12.805834">12°48'21''E</geoCoordinate>
),
<quantity id="4CA99B49FFA0FFD7FDDDFC371A99FDE2" box="[637,725,898,923]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="525" unit="km" value="3.5" valueMax="4.0" valueMin="3.0">34 km</quantity>
around, h=
<quantity id="4CA99B49FFA0FFD7FCFAFC361BACFDE3" box="[858,992,898,923]" metricMagnitude="2" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="2" pageNumber="525" unit="m" value="510.0" valueMax="540.0" valueMin="480.0">
<elevation id="007CD19FFFA0FFD7FCFAFC361BACFDE3" box="[858,992,898,923]" metricMagnitude="2" metricUnit="m" metricValue="5.1" metricValueMax="5.4" metricValueMin="4.8" pageId="2" pageNumber="525" unit="m" value="510.0" valueMax="540.0" valueMin="480.0">480540 m</elevation>
</quantity>
,
<date id="FFEF106CFFA0FFD7FC4FFC361CF4FDE2" box="[1007,1208,898,923]" pageId="2" pageNumber="525" value="2016-06-14" valueMax="2016-06-15" valueMin="2016-06-14">
<collectingDate id="EFABE984FFA0FFD7FC4FFC361CF4FDE2" box="[1007,1208,898,923]" pageId="2" pageNumber="525" value="2016-06-14" valueMax="2016-06-15" valueMin="2016-06-14">1415 June 2016</collectingDate>
</date>
, secondary lowland rainforest, night collecting from decaying wood, leg.
<collectorName id="26A4537AFFA0FFD7FCAEFC131BCEFDC6" box="[782,898,934,959]" pageId="2" pageNumber="525">D. Telnov</collectorName>
(
<collectionCode id="ED40AE69FFA0FFD7FC36FC131B96FDC6" box="[918,986,935,959]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="2" pageNumber="525">NME</collectionCode>
), genital complex in prep.
<date id="FFEF106CFFA0FFD7FABDFC121D3FFDC6" box="[1309,1395,934,959]" pageId="2" pageNumber="525" value="17-05-25">250517</date>
/02;
<typeStatus id="54EA880EFFA0FFD7FF37FC7F18B7FD9A" box="[151,251,971,995]" pageId="2" pageNumber="525" type="paratype">paratype</typeStatus>
.
<specimenCount id="9D57FD25FFA0FFD7FEA5FC7F191FFD9A" box="[261,339,970,995]" pageId="2" pageNumber="525" type="male">1 male</specimenCount>
, same data as holotype (
<collectionCode id="ED40AE69FFA0FFD7FDC4FC7F1AD4FD9A" box="[612,664,971,995]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="525" type="Museum">ZIN</collectionCode>
), genital complex in prep.
<date id="FFEF106CFFA0FFD7FC63FC7E1C55FD9A" box="[963,1049,970,995]" pageId="2" pageNumber="525" value="17-05-25">250517</date>
/02.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFA0FFD7FF67FC5A1A21FA52" pageId="2" pageNumber="525" type="etymology">
<paragraph id="8BEE36ACFFA0FFD7FF67FC5A1A21FA52" blockId="2.[151,1437,862,2009]" pageId="2" pageNumber="525">
<emphasis id="B925EABEFFA0FFD7FF67FC5A1902FA7E" bold="true" box="[199,334,1006,1031]" pageId="2" pageNumber="525">Etymology.</emphasis>
Toponymic. The new species is named after the Ivindo River, the main river in E
<collectingCountry id="F346763CFFA0FFD7FABEFC5A1D25FA7E" box="[1310,1385,1006,1031]" name="Gabon" pageId="2" pageNumber="525">Gabon</collectingCountry>
and Ivindo National park. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34B6527FFA0FFD6FF67FB8219C0FB9C" lastPageId="3" lastPageNumber="526" pageId="2" pageNumber="525" type="description">
<paragraph id="8BEE36ACFFA0FFD7FF67FB8219FCF9CC" blockId="2.[151,1437,862,2009]" pageId="2" pageNumber="525">
<emphasis id="B925EABEFFA0FFD7FF67FB82191DFA36" bold="true" box="[199,337,1078,1103]" pageId="2" pageNumber="525">Description</emphasis>
. Male (the
<typeStatus id="54EA880EFFA0FFD7FE75FB821A70FA36" box="[469,572,1078,1103]" pageId="2" pageNumber="525" type="holotype">holotype</typeStatus>
). Body generally reddish brown; eyes black; areas of ocellar spots yellowish (
<figureCitation id="136A2A29FFA0FFD7FF3FFBEE18B6FA0A" box="[159,250,1114,1139]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1A</figureCitation>
); proximal parts of antennae, maxillary and labial palps, and legs darker, nearly black. Surfaces smooth and lustrous; punctation virtually invisible; facial part of head with very delicate wrinkles above clypeus (not shown in
<figureCitation id="136A2A29FFA0FFD7FEA3FB161916FAC2" box="[259,346,1186,1211]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1A</figureCitation>
); antennae dull in distal parts (approximately from 15th segment). Head rounded at vertex, longer than wide, convex (
<figureCitation id="136A2A29FFA0FFD7FED6FB72199CFAA6" box="[374,464,1222,1247]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1A</figureCitation>
); epicranial sutures not indicated; eyes small; ocellar spots visible only as light spots; distance between eyes 1.5 times eye length; distance between antennal sockets 2.1 times of the scape length (about
<quantity id="4CA99B49FFA0FFD7FF37FABB18BEFB5E" box="[151,242,1295,1319]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="525" unit="mm" value="0.8">0.8 mm</quantity>
); approximate length ratio of 3rd5th maxillary palpomeres 1.1: 1.0: 1.0. Pronotum campaniform, wider than long, widely rounded along anterolateral margins, caudal margin slightly angulate (
<figureCitation id="136A2A29FFA0FFD7FBD0FA861C8AFB32" box="[1136,1222,1330,1355]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1B</figureCitation>
). Tegmina reduced to lateral flaps, marginated laterally and truncated apically (
<figureCitation id="136A2A29FFA0FFD7FCE4FAE21BD1FB16" box="[836,925,1366,1391]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1B</figureCitation>
). Wings absent. Mesonotum and metanotum transverse, slightly convex dorsally, with very delicate ridges; posterior margins weakly angulate (
<figureCitation id="136A2A29FFA0FFD7FA90FACE1DC1FBEA" box="[1328,1421,1402,1427]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1B</figureCitation>
). Thoracal tergites and abdominal tergites IIVII marginate laterally (
<figureCitation id="136A2A29FFA0FFD7FC03FA2A1C6CFBCE" box="[931,1056,1438,1463]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1B, C</figureCitation>
, not shown in
<figureCitation id="136A2A29FFA0FFD7FB6DFA2A1D6AFBCE" box="[1229,1318,1438,1463]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1C</figureCitation>
). Anterior margin of fore femur armed as in the type A, with 1415 spines, including 23 apical ones. Fore tibiae not thickened distally. Tibial spines well developed. Structure of hind tarsi (
<figureCitation id="136A2A29FFA0FFD7FC61FA521C4CFB86" box="[961,1024,1510,1535]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1</figureCitation>
DF): metatarsus a little longer than other segments combined, with small and apical euplantula (
<figureCitation id="136A2A29FFA0FFD7FC98F9B91BE3F85F" box="[824,943,1549,1574]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1D, E</figureCitation>
); metatarsus and 2
<superScript id="7C249BE4FFA0FFD7FBDEF9BE1CC2F861" attach="left" box="[1150,1166,1546,1560]" fontSize="6" pageId="2" pageNumber="525">nd</superScript>
segment with 2 more or less equal rows of spines along its lower margins; large euplantulae of 2
<superScript id="7C249BE4FFA0FFD7FC49F9841BB4F847" attach="none" box="[1001,1016,1584,1598]" fontSize="6" pageId="2" pageNumber="525">nd</superScript>
-4
<superScript id="7C249BE4FFA0FFD7FBB0F9841C50F847" attach="left" box="[1040,1052,1584,1598]" fontSize="6" pageId="2" pageNumber="525">th</superScript>
segments; metatarsus and 2
<superScript id="7C249BE4FFA0FFD7FAC4F9841D3FF847" attach="none" box="[1380,1395,1584,1598]" fontSize="6" pageId="2" pageNumber="525">nd</superScript>
4
<superScript id="7C249BE4FFA0FFD7FA30F9841DD0F847" attach="none" box="[1424,1436,1584,1598]" fontSize="6" pageId="2" pageNumber="525">th</superScript>
segments with pair of apical &quot;additional spines&quot; bordering euplantulae laterally (
<figureCitation id="136A2A29FFA0FFD7FB81F9EC1CEAF809" box="[1057,1190,1624,1649]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">
Fig. 1E,
<emphasis id="B925EABEFFA0FFD7FB24F9ED1CEAF809" box="[1156,1190,1625,1648]" italics="true" pageId="2" pageNumber="525">a.s</emphasis>
</figureCitation>
.); claws symmetrical, simple; arolium about half of claw length (
<figureCitation id="136A2A29FFA0FFD7FD30F9C81B5CF8EC" box="[656,784,1660,1685]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1D, F</figureCitation>
). Fore- and mid tarsi similar to metatarsi, but segments comparatively shorter. Abdomen without visible glandular specializations; posterolateral angles attenuate caudally; tergite VI with caudal margin sinusoidally curved (
<figureCitation id="136A2A29FFA0FFD7FD41F9701B76F8A4" box="[737,826,1732,1757]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1C</figureCitation>
); tergite VII projected caudally (
<figureCitation id="136A2A29FFA0FFD7FB15F9701D43F8A4" box="[1205,1295,1732,1757]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1C</figureCitation>
). Anal plate (tergite X) medially projected, with weak triangular median incision on caudal margin (
<figureCitation id="136A2A29FFA0FFD7FBCAF95C1CA9F979" box="[1130,1253,1768,1793]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1C, G</figureCitation>
). Cerci fusiform and flat, with segments solidly connected (
<figureCitation id="136A2A29FFA0FFD7FDDCF8B81AB5F95C" box="[636,761,1804,1829]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1C, G</figureCitation>
). Paraprocts weakly asymmetrical, sclerotized along caudal margin (
<figureCitation id="136A2A29FFA0FFD7FEA1F8841929F931" box="[257,357,1840,1865]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1H</figureCitation>
); pvsclerites well sclerotized, elongated not shown in
<figureCitation id="136A2A29FFA0FFD7FB90F8841CD8F931" box="[1072,1172,1840,1865]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1H</figureCitation>
. Hypandrium weakly asymmetrical, wide, antero-lateral parts (lateral sternal apodemes or apophyses) comparatively short; caudal margin sinuate between styli (
<figureCitation id="136A2A29FFA0FFD7FE4CF8CC1A75F9E9" box="[492,569,1912,1937]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">
Fig.
<date id="FFEF106CFFA0FFD7FD81F8CC1A75F9E9" box="[545,569,1912,1936]" pageId="2" pageNumber="525">1I</date>
</figureCitation>
); styli nearly symmetrical (
<figureCitation id="136A2A29FFA0FFD7FCD3F8CC1BFFF9E9" box="[883,947,1912,1937]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1</figureCitation>
IK), slightly curved down (
<figureCitation id="136A2A29FFA0FFD7FB5AF8CC1D1FF9E9" box="[1274,1363,1912,1937]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="2" pageNumber="525">Fig. 1K</figureCitation>
), with simple lobe at inner side.
</paragraph>
<paragraph id="8BEE36ACFFA0FFD6FF67F8741B32FCFC" blockId="2.[151,1437,862,2009]" lastBlockId="3.[151,1437,151,1509]" lastPageId="3" lastPageNumber="526" pageId="2" pageNumber="525">
Male genitalia (
<figureCitation id="136A2A29FFA0FFD7FED7F8741988F9A1" box="[375,452,1984,2009]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="2" pageNumber="525">Figs. 2</figureCitation>
AD, 3AD): left phallomere (
<figureCitation id="136A2A29FFA0FFD7FCBFF8741B2CF9A1" box="[799,864,1984,2009]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="2" pageNumber="525">Fig. 3</figureCitation>
AD) with sclerite L4C (L2D—here and below the terminology by
<bibRefCitation id="EFC04B5DFFA1FFD6FEF4FF231A67FEC9" author="Grandcolas, P." box="[340,555,151,176]" pageId="3" pageNumber="526" pagination="508 - 527" refId="ref7294" refString="Grandcolas, P. (1996) The phylogeny of cockroach families: a cladistic appraisal of morpho-anatomical data. Canadian Journal of Zoology, 74 (3), 508 - 527. https: // doi. org / 10.1139 / z 96 - 059" type="journal article" year="1996">Grandcolas (1996)</bibRefCitation>
is given in the parentheses) large, caudally divided into two parts, which terminating in apically curved upward processes (
<figureCitation id="136A2A29FFA1FFD6FD73FF081B59FEAD" box="[723,789,188,213]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3</figureCitation>
AC,
<emphasis id="B925EABEFFA1FFD6FCFCFF091BD6FEAD" box="[860,922,189,212]" italics="true" pageId="3" pageNumber="526">a.c.p.</emphasis>
); sclerite L3 (L3d) contracted and curved in apical part, with small directed upward tooth; sclerite L4F, possibly L4E+L4F, large, plate-like, occupying lower half of outer (
<figureCitation id="136A2A29FFA1FFD6FE93FEB019C6FF65" box="[307,394,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3B</figureCitation>
) and partly lower (
<figureCitation id="136A2A29FFA1FFD6FDC4FEB01AF0FF65" box="[612,700,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3D</figureCitation>
) sides of phallomere; sclerite L4D (L
<date id="FFEF106CFFA1FFD6FBCAFEB01CCAFF65" box="[1130,1158,260,284]" pageId="3" pageNumber="526">3v</date>
) elongated (
<figureCitation id="136A2A29FFA1FFD6FAB3FEB01DC2FF65" box="[1299,1422,260,285]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3B, C</figureCitation>
); sclerite L2 (L
<date id="FFEF106CFFA1FFD6FEE0FE9C1911FF39" box="[320,349,296,320]" pageId="3" pageNumber="526">2v</date>
) elongated, occupying ventral and lower half of inner sides of phallomere (
<figureCitation id="136A2A29FFA1FFD6FAACFE931D19FF39" box="[1292,1365,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3</figureCitation>
BD), terminating in acute caudal process (
<figureCitation id="136A2A29FFA1FFD6FD92FEF81AF7FF1D" box="[562,699,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">
Fig. 3B,
<emphasis id="B925EABEFFA1FFD6FD35FEF91AF7FF1D" box="[661,699,333,356]" italics="true" pageId="3" pageNumber="526">a.p</emphasis>
</figureCitation>
.); large &quot;additional elongated sclerite&quot;, probably separated part of L2, situated on inner side of phallomere dorsally to L2 (
<figureCitation id="136A2A29FFA1FFD6FCAEFEDB1BEBFFF1" box="[782,935,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">
Fig. 3C,
<emphasis id="B925EABEFFA1FFD6FCD2FEC51BEBFFF1" box="[882,935,369,392]" italics="true" pageId="3" pageNumber="526">a.e.s</emphasis>
</figureCitation>
.), terminating in forked caudal process (
<figureCitation id="136A2A29FFA1FFD6FACFFEDB18EFFFD5" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">Fig. 3</figureCitation>
AD,
<emphasis id="B925EABEFFA1FFD6FF4AFE20196EFFD5" box="[234,290,404,428]" italics="true" pageId="3" pageNumber="526">f.c.p.</emphasis>
) and rounded membranous lobe (
<figureCitation id="136A2A29FFA1FFD6FD3EFE201B79FFD5" box="[670,821,404,429]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">
Fig. 3C,
<emphasis id="B925EABEFFA1FFD6FCA2FE201B79FFD5" box="[770,821,404,428]" italics="true" pageId="3" pageNumber="526">r.m.l</emphasis>
</figureCitation>
.); this sclerite connected with slightly sclerotized lobe on upper half of inner side of phallomere (
<figureCitation id="136A2A29FFA1FFD6FD34FE031B50FFA9" box="[660,796,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="3" pageNumber="526">
Fig. 3C,
<emphasis id="B925EABEFFA1FFD6FD5FFE0C1B50FFA9" box="[767,796,440,464]" italics="true" pageId="3" pageNumber="526">s.l</emphasis>
</figureCitation>
.); sclerite L1 absent, substituted by membranous lobe. Ventral phallomere L4G (VP) as in
<figureCitation id="136A2A29FFA1FFD6FD93FE681AC1FF8D" box="[563,653,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="3" pageNumber="526">Fig. 2D</figureCitation>
. Right phallomere complex in shape, as in
<figureCitation id="136A2A29FFA1FFD6FB20FE681C8FFF8D" box="[1152,1219,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="3" pageNumber="526">Fig. 2</figureCitation>
AC; basal sclerite (R2) small, as compared with other sclerites of right phallomere, and transverse; sclerite R1H long and curved, apically forked; sclerite R1G subtriangular in shape, with two apical teeth (
<figureCitation id="136A2A29FFA1FFD6FC7BFD901C33FC45" box="[987,1151,548,573]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="3" pageNumber="526">
Fig. 2A, B,
<emphasis id="B925EABEFFA1FFD6FBFFFD911C33FC45" box="[1119,1151,549,572]" italics="true" pageId="3" pageNumber="526">a.t</emphasis>
</figureCitation>
.) and dorsal setal brushes (
<figureCitation id="136A2A29FFA1FFD6FF3FFDF31908FC19" box="[159,324,583,608]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="3" pageNumber="526">
Fig. 2A, C,
<emphasis id="B925EABEFFA1FFD6FE98FDFC1908FC19" box="[312,324,584,608]" italics="true" pageId="3" pageNumber="526">s</emphasis>
</figureCitation>
.br.); sclerite R3 plate-like. Elongated lobe densely covered with bristles situated under right paraproct (
<figureCitation id="136A2A29FFA1FFD6FEB0FDD819CCFCFD" box="[272,384,620,645]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">
Fig. 1H,
<emphasis id="B925EABEFFA1FFD6FED4FDD819CCFCFD" box="[372,384,620,644]" italics="true" pageId="3" pageNumber="526">s</emphasis>
</figureCitation>
.lobe.), it not connected with genital sclerites.
</paragraph>
<paragraph id="8BEE36ACFFA1FFD6FF67FD3B1A9EFCB5" blockId="3.[151,1437,151,1509]" pageId="3" pageNumber="526">
<emphasis id="B925EABEFFA1FFD6FF67FD3B190CFCD1" bold="true" box="[199,320,655,680]" pageId="3" pageNumber="526">Variations</emphasis>
. Male
<typeStatus id="54EA880EFFA1FFD6FE33FD2419BFFCD1" box="[403,499,656,680]" pageId="3" pageNumber="526" type="paratype">paratype</typeStatus>
generally similar to the
<typeStatus id="54EA880EFFA1FFD6FCA8FD3B1B21FCD1" box="[776,877,655,680]" pageId="3" pageNumber="526" type="holotype">holotype</typeStatus>
, but slightly smaller. Approximate length ratio of 3rd5th segments of maxillary palps 1.2: 1.0: 1.1.
</paragraph>
<paragraph id="8BEE36ACFFA1FFD6FF67FD6319E6FC89" blockId="3.[151,1437,151,1509]" box="[199,426,727,752]" pageId="3" pageNumber="526">Female is unknown.</paragraph>
<paragraph id="8BEE36ACFFA1FFD6FF67FD481C16FD24" blockId="3.[151,1437,151,1509]" pageId="3" pageNumber="526">
<emphasis id="B925EABEFFA1FFD6FF67FD48193AFD6C" bold="true" box="[199,374,764,789]" pageId="3" pageNumber="526">Measurements</emphasis>
(mm). Head length: 3.94.2 (4.2), width 3.33.7 (3.7), pronotum length 4.55.0 (5.0), width 6.47.2 (7.2), mesonotum length 2.02.2 (2.2), tegmina length 2.83.0 (3.0), width 1.5 (1.5), metanotum length 1.92.2 (1.9), width 7.37.9 (7.9). Measurements in parenthesis are those of
<typeStatus id="54EA880EFFA1FFD6FC51FCF01C1BFD24" box="[1009,1111,836,861]" pageId="3" pageNumber="526" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BEE36ACFFA1FFD6FF67FCD31CCAFB49" blockId="3.[151,1437,151,1509]" pageId="3" pageNumber="526">
<emphasis id="B925EABEFFA1FFD6FF67FCD31916FDF9" bold="true" box="[199,346,871,896]" pageId="3" pageNumber="526">Comparison</emphasis>
.
<bibRefCitation id="EFC04B5DFFA1FFD6FEC9FCD31A67FDF9" author="Beccaloni, G. W." box="[361,555,871,896]" pageId="3" pageNumber="526" refId="ref6988" refString="Beccaloni, G. W. (2014) Cockroach Species File Online. Version 5.0 / 5.0. World Wide Web electronic publication. Available from: http: // Cockroach. SpeciesFile. org (accessed 24 April 2018)" type="url" year="2014">Beccaloni (2014)</bibRefCitation>
listed 9 African species in the genus
<taxonomicName id="4C514D2FFFA1FFD6FC6EFCDC1C18FDF9" authorityName="Rehn" authorityYear="1933" box="[974,1108,872,896]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABEFFA1FFD6FC6EFCDC1C18FDF9" box="[974,1108,872,896]" italics="true" pageId="3" pageNumber="526">Duchailluia</emphasis>
</taxonomicName>
:
<taxonomicName id="4C514D2FFFA1FFD6FBC3FCDD1896FDDD" authority="(Gerstaecker, 1883)" authorityName="Gerstaecker" authorityYear="1883" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" class="Insecta" family="Blattidae" genus="Periplaneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="B925EABEFFA1FFD6FBC3FCDD1C3BFDF9" box="[1123,1143,873,896]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FB25FCDC1D4CFDF9" box="[1157,1280,872,896]" italics="true" pageId="3" pageNumber="526">anthracina</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FAAEFCD3189EFDDD" author="Gerstaecker, A." pageId="3" pageNumber="526" pagination="1 - 64" refId="ref7249" refString="Gerstaecker, A. (1883) Beitrag zur Kenntnis der Orthopteren-Fauna Guinea's, nach den von R. Buchholz wahrend der Jahre 1872 bis 1875 daselbst gesammelten Arten. Mittheilungen des naturwissenschaftlichen Vereins fur Neuvorpommern und Rugen, 14, 1 - 64." type="journal article" year="1883">Gerstaecker, 1883</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FF49FC391A75FDDD" authority="(Shelford, 1908)" authorityName="Shelford" authorityYear="1908" baseAuthorityName="Shelford" baseAuthorityYear="1908" box="[233,569,908,933]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="B925EABEFFA1FFD6FF49FC3918B1FDDD" box="[233,253,909,932]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FEAFFC38193DFDDD" box="[271,369,908,932]" italics="true" pageId="3" pageNumber="526">assimilis</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FE22FC381A7DFDDD" author="Shelford, R." box="[386,561,908,933]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford, 1908</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FDE9FC391BF2FDDD" authority="(Shelford, 1911 c)" authorityName="Shelford" authorityYear="1911" baseAuthorityName="Shelford" baseAuthorityYear="1911" box="[585,958,908,933]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B925EABEFFA1FFD6FDE9FC391A11FDDD" box="[585,605,909,932]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FDCEFC391AA7FDDD" box="[622,747,909,932]" italics="true" pageId="3" pageNumber="526">congoensis</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FD5CFC381BFBFDDD" author="Shelford, R." box="[764,951,908,933]" pageId="3" pageNumber="526" pagination="198 - 203" refId="ref8141" refString="Shelford, R. (1911 c) New Blattidae collected by Dr. Sheffield Neave in the Katanga Region of Congo. Revue Zoologique Africaine, 1 (2), 198 - 203." type="journal article" year="1911">Shelford, 1911c</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FC6DFC391D50FDDD" authority="(Shelford, 1908)" authorityName="Shelford" authorityYear="1908" baseAuthorityName="Shelford" baseAuthorityYear="1908" box="[973,1308,908,933]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="furcifera">
<emphasis id="B925EABEFFA1FFD6FC6DFC391BADFDDD" box="[973,993,909,932]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FC53FC381C18FDDD" box="[1011,1108,908,932]" italics="true" pageId="3" pageNumber="526">furcifera</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FBC6FC381D58FDDD" author="Shelford, R." box="[1126,1300,908,933]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford, 1908</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FA8CFC39193AFDB1" authority="(Gerstaecker, 1883)" authorityName="Gerstaecker" authorityYear="1883" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="B925EABEFFA1FFD6FA8CFC391D0CFDDD" box="[1324,1344,909,932]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FAF1FC391DD0FDDD" box="[1361,1436,909,932]" italics="true" pageId="3" pageNumber="526">manca</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FF3EFC1B1922FDB1" author="Gerstaecker, A." box="[158,366,943,968]" pageId="3" pageNumber="526" pagination="1 - 64" refId="ref7249" refString="Gerstaecker, A. (1883) Beitrag zur Kenntnis der Orthopteren-Fauna Guinea's, nach den von R. Buchholz wahrend der Jahre 1872 bis 1875 daselbst gesammelten Arten. Mittheilungen des naturwissenschaftlichen Vereins fur Neuvorpommern und Rugen, 14, 1 - 64." type="journal article" year="1883">Gerstaecker, 1883</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FE23FC051A94FDB1" authority="(Shelford, 1908)" authorityName="Shelford" authorityYear="1908" baseAuthorityName="Shelford" baseAuthorityYear="1908" box="[387,728,943,968]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B925EABEFFA1FFD6FE23FC0519DBFDB1" box="[387,407,945,968]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FE05FC041A59FDBE" box="[421,533,944,967]" italics="true" pageId="3" pageNumber="526">nigerrima</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FD84FC1B1A83FDB1" author="Shelford, R." box="[548,719,943,968]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford, 1908</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FD44FC051C5AFDB1" authority="(Jolivet, 1954)" authorityName="Jolivet" authorityYear="1954" baseAuthorityName="Jolivet" baseAuthorityYear="1954" box="[740,1046,943,968]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="shelfordi">
<emphasis id="B925EABEFFA1FFD6FD44FC051AB4FDB1" box="[740,760,945,968]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FCA6FC041B24FDB1" box="[774,872,944,968]" italics="true" pageId="3" pageNumber="526">shelfordi</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FCD8FC1B1C43FDB1" author="Jolivet, Y." box="[888,1039,943,968]" pageId="3" pageNumber="526" pagination="3 - 39" refId="ref7341" refString="Jolivet, Y. (1954) Dictyoptera (Blattodea). In: Adam, W., Janssens, A., Van Meel, L. &amp; Verheyen, R. Eds., Exploration du Parc National de l'Upemba. Vol. 29. Institut des parcs nationaux du Congo et du Rwanda, Bruxelles, pp. 3 - 39." type="book chapter" year="1954">Jolivet, 1954</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FB82FC051D20FDB1" authority="(Krauss, 1890)" authorityName="Krauss" authorityYear="1890" baseAuthorityName="Krauss" baseAuthorityYear="1890" box="[1058,1388,944,968]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="spinulifera">
<emphasis id="B925EABEFFA1FFD6FB82FC051C7AFDB1" box="[1058,1078,945,968]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FBE5FC041CF2FDB1" box="[1093,1214,944,968]" italics="true" pageId="3" pageNumber="526">spinulifera</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FB6DFC041D28FDB1" author="Krauss, H." box="[1229,1380,944,968]" pageId="3" pageNumber="526" pagination="647 - 668" refId="ref7498" refString="Krauss, H. (1890) Beitrag zur Kenntnis westafrikanischer Orthopteren. 2. Orthopteren der Guinea-Inseln Sao Thome und Rolas, gesammelt von Prof. Dr. Richard Graeff. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 5, 647 - 668." type="journal article" year="1890">Krauss, 1890</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C514D2FFFA1FFD6FF37FC6119B9FD95" authority="(Shelford, 1911 a)" authorityName="Shelford" authorityYear="1911" baseAuthorityName="Shelford" baseAuthorityYear="1911" box="[151,501,980,1005]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="togoensis">
<emphasis id="B925EABEFFA1FFD6FF37FC6118E7FD95" box="[151,171,981,1004]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FF1AFC611968FD95" box="[186,292,981,1004]" italics="true" pageId="3" pageNumber="526">togoensis</emphasis>
(
<bibRefCitation id="EFC04B5DFFA1FFD6FE93FC6019A0FD95" author="Shelford, R." box="[307,492,980,1005]" pageId="3" pageNumber="526" pagination="1 - 13" refId="ref8059" refString="Shelford, R. (1911 a) Descriptions of some new Species of Blattidae. Annals and Magazine of Natural History, Series 8, 8 (43), 1 - 13. https: // doi. org / 10.1080 / 00222931108692992" type="journal article" year="1911">Shelford, 1911a</bibRefCitation>
)
</taxonomicName>
. The new species readily differs from
<taxonomicName id="4C514D2FFFA1FFD6FC0AFC611C7CFD95" baseAuthorityName="Shelford" baseAuthorityYear="1908" box="[938,1072,980,1004]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="B925EABEFFA1FFD6FC0AFC611BF2FD95" box="[938,958,981,1004]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FC6EFC601C7CFD95" box="[974,1072,980,1004]" italics="true" pageId="3" pageNumber="526">assimilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FB9FFC611C88FD95" baseAuthorityName="Shelford" baseAuthorityYear="1908" box="[1087,1220,980,1004]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="furcifera">
<emphasis id="B925EABEFFA1FFD6FB9FFC611C1FFD95" box="[1087,1107,981,1004]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FBC3FC601C88FD95" box="[1123,1220,980,1004]" italics="true" pageId="3" pageNumber="526">furcifera</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D2FFFA1FFD6FB73FC611D0DFD95" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" box="[1235,1345,981,1004]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="B925EABEFFA1FFD6FB73FC611CABFD95" box="[1235,1255,981,1004]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FB56FC611D0DFD95" box="[1270,1345,981,1004]" italics="true" pageId="3" pageNumber="526">manca</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C514D2FFFA1FFD6FA21FC61194BFA76" baseAuthorityName="Shelford" baseAuthorityYear="1908" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B925EABEFFA1FFD6FA21FC611DD9FD95" box="[1409,1429,981,1004]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FF37FC4C194BFA76" box="[151,263,1016,1039]" italics="true" pageId="3" pageNumber="526">nigerrima</emphasis>
</taxonomicName>
in shape of anal plate and hypandrium (compare
<figureCitation id="136A2A29FFA1FFD6FC9FFC431B31FA69" box="[831,893,1015,1040]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1</figureCitation>
GK and
<figureCitation id="136A2A29FFA1FFD6FC52FC431C10FA69" box="[1010,1116,1015,1040]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,249,1823,1845]" captionStartId-1="5.[151,249,1936,1958]" captionStartId-2="6.[151,249,1878,1900]" captionStartId-3="7.[151,249,1811,1833]" captionTargetBox-0="[188,1396,214,1803]" captionTargetBox-1="[207,1407,214,1901]" captionTargetBox-2="[230,1361,198,1854]" captionTargetBox-3="[219,1368,193,1791]" captionTargetId-0="figure@4.[173,1417,195,1805]" captionTargetId-1="figure@5.[185,1409,193,1921]" captionTargetId-2="figure@6.[225,1367,188,1868]" captionTargetId-3="figure@7.[212,1380,183,1798]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." captionText-1="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." captionText-2="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. AD—Duchailluia ivindo sp. nov.; EH—Distylopyga beccalonii sp. nov.; IL—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = AD; b = EH; c = I L." captionText-3="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (AK) and female, paratype (LN). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: IX—1st10th abdominal tergites respectively, a.s., par., pv. see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri-0="https://zenodo.org/record/2615631/files/figure.png" httpUri-1="https://zenodo.org/record/2615633/files/figure.png" httpUri-2="https://zenodo.org/record/2615637/files/figure.png" httpUri-3="https://zenodo.org/record/2615639/files/figure.png" pageId="3" pageNumber="526">
Figs.
<quantity id="4CA99B49FFA1FFD6FB91FC4C1C10FA69" box="[1073,1116,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="3" pageNumber="526" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">14</quantity>
</figureCitation>
in
<bibRefCitation id="EFC04B5DFFA1FFD6FB22FC431D6BFA69" author="Shelford, R." box="[1154,1319,1015,1040]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford 1908</bibRefCitation>
). From
<taxonomicName id="4C514D2FFFA1FFD6FA21FC4D195EFA4D" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" class="Insecta" family="Blattidae" genus="Periplaneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="B925EABEFFA1FFD6FA21FC4D1DD9FA69" box="[1409,1429,1017,1040]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FF37FBA8195EFA4D" box="[151,274,1052,1076]" italics="true" pageId="3" pageNumber="526">anthracina</emphasis>
</taxonomicName>
the new species differs in shape of caudal margin of anal plate, which is weakly emarginated (
<figureCitation id="136A2A29FFA1FFD6FA95FBA81DC1FA4D" box="[1333,1421,1052,1077]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1G</figureCitation>
), not &quot;produced to a point&quot;, as indicated for
<taxonomicName id="4C514D2FFFA1FFD6FD20FBF51B53FA21" baseAuthorityName="Gerstaecker" baseAuthorityYear="1883" box="[640,799,1088,1112]" class="Insecta" family="Blattidae" genus="Periplaneta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="B925EABEFFA1FFD6FD20FBF51AD8FA21" box="[640,660,1089,1112]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FD04FBF41B53FA21" box="[676,799,1088,1112]" italics="true" pageId="3" pageNumber="526">anthracina</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B5DFFA1FFD6FCECFB8B1C00FA21" author="Shelford, R." box="[844,1100,1087,1112]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford (1908, p. 32)</bibRefCitation>
.
<taxonomicName id="4C514D2FFFA1FFD6FBF8FBF41D2CFA21" baseAuthorityName="Krauss" baseAuthorityYear="1890" box="[1112,1376,1088,1112]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="spinulifera">
<emphasis id="B925EABEFFA1FFD6FBF8FBF41D2CFA21" box="[1112,1376,1088,1112]" italics="true" pageId="3" pageNumber="526">Duchailluia spinulifera</emphasis>
</taxonomicName>
does not have bifurcate styli (
<bibRefCitation id="EFC04B5DFFA1FFD6FE05FBD01A05FA05" author="Shelford, R." box="[421,585,1124,1149]" pageId="3" pageNumber="526" pagination="27 - 38" refId="ref8024" refString="Shelford, R. (1908) On a small collection of Blattidae in the Naturhistorischen Museum zu Wiesbaden. Jahrbucher des Nassauisshen Vereins fur Naturkunde in Wiesbaden, 61, 27 - 38." type="journal article" year="1908">Shelford 1908</bibRefCitation>
), contrary to
<taxonomicName id="4C514D2FFFA1FFD6FD7DFBD11B09FA05" authority="Anisyutkin &amp; Telnov, 2018" authorityName="Anisyutkin &amp; Telnov" authorityYear="2018" box="[733,837,1124,1148]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="ivindo" status="sp. nov.">
<emphasis id="B925EABEFFA1FFD6FD7DFBD11ABDFA05" box="[733,753,1125,1148]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FD5FFBD01B09FA05" box="[767,837,1124,1148]" italics="true" pageId="3" pageNumber="526">ivindo</emphasis>
</taxonomicName>
<emphasis id="B925EABEFFA1FFD6FCECFBD01BE9FA04" bold="true" box="[844,933,1124,1149]" pageId="3" pageNumber="526">
<taxonomicNameLabel id="A21657C5FFA1FFD6FCECFBD01BE9FA04" box="[844,933,1124,1149]" pageId="3" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
From
<taxonomicName id="4C514D2FFFA1FFD6FC4FFBD11C37FA05" baseAuthorityName="Shelford" baseAuthorityYear="1911" box="[1007,1147,1125,1148]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="togoensis">
<emphasis id="B925EABEFFA1FFD6FC4FFBD11C4FFA05" box="[1007,1027,1125,1148]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FBB1FBD11C37FA05" box="[1041,1147,1125,1148]" italics="true" pageId="3" pageNumber="526">togoensis</emphasis>
</taxonomicName>
the new species differs in shape of caudal margins of anal plate, hypandrium and inner branch of styles (compare
<figureCitation id="136A2A29FFA1FFD6FBD7FB331C80FAD9" box="[1143,1228,1159,1184]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1G</figureCitation>
, IK and
<figureCitation id="136A2A29FFA1FFD6FA9CFB331D33FAD9" box="[1340,1407,1159,1184]" captionStart="FIGURE 2" captionStartId="5.[151,249,1936,1958]" captionTargetBox="[207,1407,214,1901]" captionTargetId="figure@5.[185,1409,193,1921]" captionTargetPageId="5" captionText="FIGURE 2. Duchailluia ivindo Anisyutkin, sp. nov., male genitalia, holotype. AC—right phallomere from above (A), below (B) and outside (C); D—ventral phallomere from below. Dotted areas shows membranous parts. Abbreviations: a.t., R1G, R1H, R2, R3, s.br.—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615633/files/figure.png" pageId="3" pageNumber="526">
Fig.
<quantity id="4CA99B49FFA1FFD6FAD1FB3C1D33FAD9" box="[1393,1407,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="526" unit="in" value="2.0">2</quantity>
</figureCitation>
in
<bibRefCitation id="EFC04B5DFFA1FFD6FF37FB181909FABD" author="Shelford, R." box="[151,325,1196,1221]" pageId="3" pageNumber="526" pagination="1 - 13" refId="ref8059" refString="Shelford, R. (1911 a) Descriptions of some new Species of Blattidae. Annals and Magazine of Natural History, Series 8, 8 (43), 1 - 13. https: // doi. org / 10.1080 / 00222931108692992" type="journal article" year="1911">Shelford 1911a</bibRefCitation>
). From
<taxonomicName id="4C514D2FFFA1FFD6FE3BFB191A75FABD" baseAuthorityName="Shelford" baseAuthorityYear="1911" box="[411,569,1197,1220]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B925EABEFFA1FFD6FE3BFB1919E3FABD" box="[411,431,1197,1220]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FE1DFB191A75FABD" box="[445,569,1197,1220]" italics="true" pageId="3" pageNumber="526">congoensis</emphasis>
</taxonomicName>
the new species differs in shape of tegmina, which is elongated (
<figureCitation id="136A2A29FFA1FFD6FAADFB181D2EFABD" box="[1293,1378,1196,1221]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1B</figureCitation>
), not &quot;broader at base than long&quot;, as indicated for
<taxonomicName id="4C514D2FFFA1FFD6FD3DFB651B72FA9E" baseAuthorityName="Shelford" baseAuthorityYear="1911" box="[669,830,1232,1256]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="congoensis">
<emphasis id="B925EABEFFA1FFD6FD3DFB651AFDFA91" box="[669,689,1233,1256]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FD62FB641B72FA9E" box="[706,830,1232,1255]" italics="true" pageId="3" pageNumber="526">congoensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B5DFFA1FFD6FCCDFB7B1C6AFA91" author="Shelford, R." box="[877,1062,1231,1256]" pageId="3" pageNumber="526" pagination="1 - 13" refId="ref8059" refString="Shelford, R. (1911 a) Descriptions of some new Species of Blattidae. Annals and Magazine of Natural History, Series 8, 8 (43), 1 - 13. https: // doi. org / 10.1080 / 00222931108692992" type="journal article" year="1911">Shelford (1911a</bibRefCitation>
, p. 201) and nearly symmetrical styles (
<figureCitation id="136A2A29FFA1FFD6FF47FB40196BFB75" box="[231,295,1268,1293]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1</figureCitation>
IK). From
<taxonomicName id="4C514D2FFFA1FFD6FE15FB411A77FB75" baseAuthorityName="Jolivet" baseAuthorityYear="1954" box="[437,571,1268,1292]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="shelfordi">
<emphasis id="B925EABEFFA1FFD6FE15FB411985FB75" box="[437,457,1269,1292]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FE78FB401A77FB75" box="[472,571,1268,1292]" italics="true" pageId="3" pageNumber="526">shelfordi</emphasis>
</taxonomicName>
the new species differs in more light colour and distinctly protruded caudal margin of pronotum (
<figureCitation id="136A2A29FFA1FFD6FE27FAA31991FB49" box="[391,477,1303,1328]" captionStart="FIGURE 1" captionStartId="4.[151,249,1823,1845]" captionTargetBox="[188,1396,214,1803]" captionTargetId="figure@4.[173,1417,195,1805]" captionTargetPageId="4" captionText="FIGURE 1. Duchailluia ivindo Anisyutkin, sp. nov., male, holotype. A—facial part of head; B—head, thorax and first abdominal tergite from above; C—abdomen (with exception of first abdominal tergite) from above; D—left hind tarsus from outside; E—1st4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above; H—abdominal apex from below, hypandrium and genitalia removed; I—hypandrium from below; J—caudal margin of hypandrium from above; K—left stylus from outside. Dotted areas shows membranous parts. Dotted line in Fig. 1A delimits areas of ocellar spots. Abbreviations: IVII—1st7th abdominal tergites respectively, a.s., par., s.lobe.—see text. Scale bar 1 mm: a = A; b = B, C; c = D; d = E; e = F; f = G; g = H; h = I; i = J, K." httpUri="https://zenodo.org/record/2615631/files/figure.png" pageId="3" pageNumber="526">Fig. 1B</figureCitation>
), not straight, as indicated for
<taxonomicName id="4C514D2FFFA1FFD6FC94FAAD1BF4FB49" baseAuthorityName="Jolivet" baseAuthorityYear="1954" box="[820,952,1304,1328]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="shelfordi">
<emphasis id="B925EABEFFA1FFD6FC94FAAD1B04FB49" box="[820,840,1305,1328]" italics="true" pageId="3" pageNumber="526">D</emphasis>
.
<emphasis id="B925EABEFFA1FFD6FCF6FAAC1BF4FB49" box="[854,952,1304,1328]" italics="true" pageId="3" pageNumber="526">shelfordi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B5DFFA1FFD6FC43FAA31CCEFB49" author="Jolivet, Y." box="[995,1154,1303,1328]" pageId="3" pageNumber="526" pagination="3 - 39" refId="ref7341" refString="Jolivet, Y. (1954) Dictyoptera (Blattodea). In: Adam, W., Janssens, A., Van Meel, L. &amp; Verheyen, R. Eds., Exploration du Parc National de l'Upemba. Vol. 29. Institut des parcs nationaux du Congo et du Rwanda, Bruxelles, pp. 3 - 39." type="book chapter" year="1954">Jolivet (1954)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEE36ACFFA1FFD6FF67FA8819C0FB9C" blockId="3.[151,1437,151,1509]" pageId="3" pageNumber="526">
<emphasis id="B925EABEFFA1FFD6FF67FA8818B1FB2C" bold="true" box="[199,253,1340,1365]" pageId="3" pageNumber="526">Note</emphasis>
. The field research was carried out in
<date id="FFEF106CFFA1FFD6FD12FA891B6FFB2C" box="[690,803,1340,1365]" pageId="3" pageNumber="526" value="2016-06">
June
<quantity id="4CA99B49FFA1FFD6FD4DFA881B6FFB2C" box="[749,803,1340,1365]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="3" pageNumber="526" unit="in" value="2016.0">2016</quantity>
</date>
in a dry season. Described material comes from an oldgrowth secondary lowland rainforest. Specimens were sampled from upper surface of a large fallen tree (trunk) at morning time (collecting area was partly in shade; at midday this area is fully sun exposed). The trunk was lying horizontally on the ground, with bark partly strongly attached to the trunk, partly loose. Specimens sampled from underside of the trunk.
</paragraph>
</subSubSection>
</treatment>
</document>