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<document id="37B8AAD7C1E96961DC704B3DA6030FF8" ID-CLB-Dataset="57602" ID-DOI="10.5852/ejt.2023.867.2109" ID-GBIF-Dataset="1d2570da-e150-4d75-94d8-bfc5813062bc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7891021" ID-ZooBank="8DEF295C-A8B1-4A6B-B873-B30949F64E07" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1683117081100" checkinUser="felipe" docAuthor="Iniesta, Luiz Felipe Moretti, Bouzan, Rodrigo Salvador &amp; Brescovit, Antonio Domingos" docDate="2023" docId="03F887BA136BB1384D4FFC91FC9456D8" docLanguage="en" docName="ejt-867_1-312.pdf.imf" docOrigin="European Journal of Taxonomy 867 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.867.2109" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pseudonannolene sulcatula Silvestri 1895" docType="treatment" docVersion="7" lastPageNumber="90" masterDocId="FFC1FFC21333B1614F56FFECFF9A553E" masterDocTitle="A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)" masterLastPageNumber="312" masterPageNumber="1" pageNumber="89" updateTime="1699515670513" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="49BC26BE861704455C2B59638A06E501">A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)</mods:title>
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<mods:namePart id="DCDD9B71EEAFF561FB16C863DE76697D">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:affiliation id="771AE218B224DBB6860BADF136148DBC">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="CBD1B6556150B86535CE0F119023FD61" type="email">rodrigobouzan@outlook.com</mods:nameIdentifier>
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<mods:namePart id="58F1D13CF3D17D0705C17659A4692523">Bouzan, Rodrigo Salvador</mods:namePart>
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<mods:affiliation id="5734F51B99FFF3B2CA0D0E5490805235">Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil. &amp; Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="25972D006B5E120D43B8B7594590822F">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="BC6AA39331E1CBD25F7C089D909A3986">Laboratório de Coleções Zoológicas, Instituto Butantan, Avenida Vital Brasil, 1500, 05503 - 090, São Paulo, Brazil.</mods:affiliation>
<mods:nameIdentifier id="02884DB5A86208A2E3175B2EDD491FE1" type="email">rodrigobouzan@outlook.com&amp;antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:title id="AE5C307CFDE7B18373C1485468AF76FF">European Journal of Taxonomy</mods:title>
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<mods:date id="33790D89EDC5F96B2DB0D0C974EFE039">2023</mods:date>
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<treatment id="03F887BA136BB1384D4FFC91FC9456D8" ID-DOI="http://doi.org/10.5281/zenodo.7907941" ID-GBIF-Taxon="209588364" ID-Zenodo-Dep="7907941" LSID="urn:lsid:plazi:treatment:03F887BA136BB1384D4FFC91FC9456D8" httpUri="http://treatment.plazi.org/id/03F887BA136BB1384D4FFC91FC9456D8" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89">
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<taxonomicName id="4C514D2F136BB1394D4FFC91FB8156A7" ID-CoL="4P4ND" authority="Silvestri, 1895" authorityName="Silvestri" authorityYear="1895" box="[537,1051,893,921]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sulcatula">
<emphasis id="B925EABE136BB1394D4FFC91FCF256A6" bold="true" box="[537,872,893,920]" italics="true" pageId="88" pageNumber="89">Pseudonannolene sulcatula</emphasis>
Silvestri, 1895
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<subSubSection id="C34B6527136BB1394DF3FC4CFC145685" box="[677,910,928,955]" pageId="88" pageNumber="89" type="description">
<paragraph id="8BEE36AC136BB1394DF3FC4CFC145685" blockId="88.[537,1051,893,955]" box="[677,910,928,955]" pageId="88" pageNumber="89">
<figureCitation id="136A2A29136BB1394DF3FC4CFC975685" box="[677,781,928,955]" captionStart="Fig" captionStartId="238.[189,232,1217,1243]" captionTargetBox="[189,1398,266,1175]" captionTargetId="figure-13@238.[189,1398,266,1175]" captionTargetPageId="238" captionText="Fig. 131. Pseudonannolene sulcatula Silvestri, 1895, ♀ (NHMD). A. Body rings, in lateral view, and original label. B. Posterior region in lateral view. C. Schematic drawing of right gonopod in anal view (modified from Silvestri 1895b: fig. 14). Abbreviations: see Material and methods. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891324" httpUri="https://zenodo.org/record/7891324/files/figure.png" pageId="88" pageNumber="89">Figs 131</figureCitation>
,
<figureCitation id="136A2A29136BB1394C4CFC4CFCCC5684" box="[794,854,928,954]" captionStart="Fig" captionStartId="279.[189,232,1892,1918]" captionTargetBox="[318,1270,265,1851]" captionTargetId="figure-11@279.[318,1270,265,1851]" captionTargetPageId="279" captionText="Fig. 176. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. pusilla Silvestri, 1895. B. P. bovei Silvestri, 1895. C. P. brevis Silvestri, 1902. D. P. centralis Silvestri, 1902. E. P. rugosetta Silvestri, 1897. F. P. sulcatula Silvestri, 1895. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891437" httpUri="https://zenodo.org/record/7891437/files/figure.png" pageId="88" pageNumber="89">176F</figureCitation>
,
<figureCitation id="136A2A29136BB1394C34FC4DFC145685" box="[866,910,929,955]" captionStart="Fig" captionStartId="291.[189,232,1392,1418]" captionTargetBox="[192,1368,289,1302]" captionTargetId="figure-11@291.[189,1398,265,1329]" captionTargetPageId="291" captionText="Fig. 188. Distribution map of the species Pseudonannolene segmentata Silvestri, 1895, P. silvestris Schubart, 1944, P. spelaea Iniesta &amp; Ferreira, 2013, P. strinatii Mauriès, 1974, and P. sulcatula Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891461" httpUri="https://zenodo.org/record/7891461/files/figure.png" pageId="88" pageNumber="89">188</figureCitation>
</paragraph>
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<paragraph id="8BEE36AC136BB1394FEBFC0DFCDA56C2" blockId="88.[189,832,993,1020]" box="[189,832,993,1020]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="AB411182136BB1394FEBFC0DFCDA56C2" box="[189,832,993,1020]" pageId="88" pageNumber="89">
<treatmentCitation id="0AF010BD136BB1394FEBFC0DFD7856C2" author="Silvestri F." box="[189,738,993,1020]" page="7" pageId="88" pageNumber="89" year="1895">
<taxonomicName id="4C514D2F136BB1394FEBFC0DFD7856C2" ID-CoL="4P4ND" authority="Silvestri, 1895 b: 7" authorityName="Silvestri" authorityPageNumber="7" authorityYear="1895" box="[189,738,993,1020]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sulcatula">
<emphasis id="B925EABE136BB1394FEBFC0DFD9B56C5" box="[189,513,993,1019]" italics="true" pageId="88" pageNumber="89">Pseudonannolene sulcatula</emphasis>
<bibRefCitation id="EFC04B5D136BB1394D5EFC0DFD7856C2" author="Silvestri F." box="[520,738,993,1020]" pageId="88" pageNumber="89" pagination="1 - 12" refId="ref108090" refString="Silvestri F. 1895 b. Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino 10 (203): 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri, 1895b: 7</bibRefCitation>
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, fig. 14.
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</paragraph>
<paragraph id="8BEE36AC136BB1394FEBFBCEFC685103" blockId="88.[189,1010,1058,1085]" box="[189,1010,1058,1085]" pageId="88" pageNumber="89">
<treatmentCitationGroup id="AB411182136BB1394FEBFBCEFC685103" box="[189,1010,1058,1085]" pageId="88" pageNumber="89">
<taxonomicName id="4C514D2F136BB1394FEBFBCEFD9B5102" ID-CoL="4P4ND" authorityName="Silvestri" authorityYear="1895" box="[189,513,1058,1084]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sulcatula">
<emphasis id="B925EABE136BB1394FEBFBCEFD9B5102" box="[189,513,1058,1084]" italics="true" pageId="88" pageNumber="89">Pseudonannolene sulcatula</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AF010BD136BB1394D4BFBCEFC9E5102" author="Viggiani V." box="[541,772,1058,1085]" page="367" pageId="88" pageNumber="89" year="1973">
<bibRefCitation id="EFC04B5D136BB1394D4BFBCEFC9E5102" author="Viggiani V." box="[541,772,1058,1085]" pageId="88" pageNumber="89" pagination="351 - 417" refId="ref108936" refString="Viggiani V. 1973. Le specie descritte da Filippo Silvestri (1873 - 1949). Bollettino del Laboratorio di entomologia agraria &quot; Filippo Silvestri &quot; 30: 351 - 417." type="journal article" year="1973">Viggiani 1973: 367</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF010BD136BB1394C65FBCEFC745103" author="Jeekel C. A. W." box="[819,1006,1058,1085]" page="91" pageId="88" pageNumber="89" year="2004">
<bibRefCitation id="EFC04B5D136BB1394C65FBCEFC745103" author="Jeekel C. A. W." box="[819,1006,1058,1085]" pageId="88" pageNumber="89" pagination="43 - 109" refId="ref104347" refString="Jeekel C. A. W. 2004. A bibliographic catalogue of the &quot; Cambaloidea &quot; (Diplopoda, Spirostreptida). Myriapod Memoranda 7: 43 - 109." type="journal article" year="2004">Jeekel 2004: 91</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C34B6527136BB1394FEBFB8FFBC151CF" pageId="88" pageNumber="89" type="diagnosis">
<paragraph id="8BEE36AC136BB1394FEBFB8FFEAC5143" blockId="88.[189,310,1123,1149]" box="[189,310,1123,1149]" pageId="88" pageNumber="89">
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<emphasis id="B925EABE136BB1394FEBFB8FFEAC5143" bold="true" box="[189,310,1123,1149]" pageId="88" pageNumber="89">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBFB7DFBC151CF" blockId="88.[189,1399,1169,1265]" pageId="88" pageNumber="89">
Males of
<taxonomicName id="4C514D2F136BB1394E7AFB7EFE2F5195" authorityName="Silvestri" authorityYear="1895" box="[300,437,1169,1195]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="sulcatula">
<emphasis id="B925EABE136BB1394E7AFB7EFE2F5195" box="[300,437,1169,1195]" italics="true" pageId="88" pageNumber="89">P. sulcatula</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C514D2F136BB1394DC4FB7EFC865195" authorityName="Schubart" authorityYear="1944" box="[658,796,1169,1195]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="ophiiulus">
<emphasis id="B925EABE136BB1394DC4FB7EFC865195" box="[658,796,1169,1195]" italics="true" pageId="88" pageNumber="89">P. ophiiulus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C514D2F136BB1394C7DFB7EFC3F5195" authorityName="Mauries" authorityYear="1974" box="[811,933,1170,1195]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="strinatii">
<emphasis id="B925EABE136BB1394C7DFB7EFC3F5195" box="[811,933,1170,1195]" italics="true" pageId="88" pageNumber="89">P. strinatii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C514D2F136BB1394CB0FB7EFBE25195" authorityName="Fontanetti" authorityYear="1996" box="[998,1144,1170,1195]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="tocaiensis">
<emphasis id="B925EABE136BB1394CB0FB7EFBE25195" box="[998,1144,1170,1195]" italics="true" pageId="88" pageNumber="89">P. tocaiensis</emphasis>
</taxonomicName>
by having a gonopod with a subtriangular gonocoxa; the internal branch being shovel-shaped, but differing by the absence of a horizontal plate in the internal branch (
<bibRefCitation id="EFC04B5D136BB1394DCFFB3BFCF751CF" author="Silvestri F." box="[665,877,1238,1265]" pageId="88" pageNumber="89" pagination="1 - 12" refId="ref108090" refString="Silvestri F. 1895 b. Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino 10 (203): 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri 1895b: 8</bibRefCitation>
, fig. 14;
<figureCitation id="136A2A29136BB1394C83FB3BFBD451CF" box="[981,1102,1239,1265]" captionStart="Fig" captionStartId="238.[189,232,1217,1243]" captionTargetBox="[189,1398,266,1175]" captionTargetId="figure-13@238.[189,1398,266,1175]" captionTargetPageId="238" captionText="Fig. 131. Pseudonannolene sulcatula Silvestri, 1895, ♀ (NHMD). A. Body rings, in lateral view, and original label. B. Posterior region in lateral view. C. Schematic drawing of right gonopod in anal view (modified from Silvestri 1895b: fig. 14). Abbreviations: see Material and methods. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891324" httpUri="https://zenodo.org/record/7891324/files/figure.png" pageId="88" pageNumber="89">Fig. 131C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B6527136BB1394FEBFAF4FAC95061" box="[189,1363,1304,1376]" pageId="88" pageNumber="89" type="etymology">
<paragraph id="8BEE36AC136BB1394FEBFAF4FEDE500C" blockId="88.[189,324,1304,1330]" box="[189,324,1304,1330]" pageId="88" pageNumber="89">
<heading id="D0A681C0136BB1394FEBFAF4FEDE500C" bold="true" box="[189,324,1304,1330]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="B925EABE136BB1394FEBFAF4FEDE500C" bold="true" box="[189,324,1304,1330]" pageId="88" pageNumber="89">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBFAA9FAC95061" blockId="88.[189,1363,1349,1376]" box="[189,1363,1349,1376]" pageId="88" pageNumber="89">Unspecified in the original description and not related to any morphological structure of the species.</paragraph>
</subSubSection>
<subSubSection id="C34B6527136BB1394FEBFA6AFD4E50CC" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="8BEE36AC136BB1394FEBFA6AFDAF509F" blockId="88.[189,565,1414,1441]" box="[189,565,1414,1441]" pageId="88" pageNumber="89">
<emphasis id="B925EABE136BB1394FEBFA6AFE30509E" bold="true" box="[189,426,1414,1440]" pageId="88" pageNumber="89">Material examined</emphasis>
(total:
<specimenCount id="9D57FD25136BB1394EA9FA6BFDB3509F" box="[511,553,1415,1441]" pageId="88" pageNumber="89" type="female">1 ♀</specimenCount>
)
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBFA59FD4E50CC" blockId="88.[189,1399,1460,1522]" pageId="88" pageNumber="89">
<materialsCitation id="3B393CF1136BB1394FEBFA59FD5750CC" ID-GBIF-Occurrence="4101309406" collectingDate="1979-04-11" collectionCode="NHMD" collectorName="Rosario de la Frontera &amp; Mision Cientifica Danesa" country="Argentina" elevation="200" latitude="-25.800215" location="Salta" longLatPrecision="1" longitude="-64.96783" pageId="88" pageNumber="89" specimenCount="1" specimenCount-female="1" stateProvince="Salta">
<collectingCountry id="F346763C136BB1394FEBFA59FEFC50F1" box="[189,358,1461,1487]" name="Argentina" pageId="88" pageNumber="89">ARGENTINA</collectingCountry>
–
<emphasis id="B925EABE136BB1394EDFFA58FE5350F0" bold="true" box="[393,457,1460,1486]" pageId="88" pageNumber="89">
<collectingRegion id="4995F84E136BB1394EDFFA58FE5350F0" box="[393,457,1460,1486]" country="Argentina" name="Salta" pageId="88" pageNumber="89">Salta</collectingRegion>
</emphasis>
•
<specimenCount id="9D57FD25136BB1394EB2FA59FD8850F1" box="[484,530,1461,1487]" pageId="88" pageNumber="89" type="female">1 ♀</specimenCount>
;
<collectorName id="26A4537A136BB1394D75FA58FCA850F1" box="[547,818,1460,1487]" pageId="88" pageNumber="89">Rosário de la Frontera</collectorName>
; [
<geoCoordinate id="EE65506B136BB1394C10FA58FC4950F1" box="[838,979,1460,1487]" degrees="25.800215" direction="south" orientation="latitude" pageId="88" pageNumber="89" precision="1" value="-25.800215">-25.800215</geoCoordinate>
,
<geoCoordinate id="EE65506B136BB1394CB6FA58FBF750F1" box="[992,1133,1460,1487]" degrees="64.967830" direction="west" orientation="longitude" pageId="88" pageNumber="89" precision="1" value="-64.96783">-64.967830</geoCoordinate>
];
<elevation id="007CD19F136BB1394BD4FA59FA8A50F0" box="[1154,1296,1460,1487]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="88" pageNumber="89" unit="m" value="200.0">
<quantity id="4CA99B49136BB1394BD4FA59FB5450F1" box="[1154,1230,1461,1487]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="88" pageNumber="89" unit="m" value="200.0">200 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEF106C136BB1394A49FA59FF6750CF" pageId="88" pageNumber="89" value="1979-04-11">
<collectingDate id="EFABE984136BB1394A49FA59FF6750CF" pageId="88" pageNumber="89" value="1979-04-11">11 Apr. 1979</collectingDate>
</date>
;
<collectorName id="26A4537A136BB1394E5FFA3BFDA950CC" box="[265,563,1495,1522]" pageId="88" pageNumber="89">Misión Científica Danesa</collectorName>
leg.;
<collectionCode id="ED40AE69136BB1394D25FA34FD5750CC" box="[627,717,1496,1522]" pageId="88" pageNumber="89">NHMD</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B6527136BB1384FEBF9F4FD2E5781" lastPageId="89" lastPageNumber="90" pageId="88" pageNumber="89" type="description">
<paragraph id="8BEE36AC136BB1394FEBF9F4FE09530C" blockId="88.[189,403,1560,1586]" box="[189,403,1560,1586]" pageId="88" pageNumber="89">
<heading id="D0A681C0136BB1394FEBF9F4FE09530C" bold="true" box="[189,403,1560,1586]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="B925EABE136BB1394FEBF9F4FE09530C" bold="true" box="[189,403,1560,1586]" pageId="88" pageNumber="89">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBF9AAFA8F53BD" blockId="88.[189,1399,1606,1667]" pageId="88" pageNumber="89">
Gonopod description adapted from
<bibRefCitation id="EFC04B5D136BB1394D21F9AAFCF2535F" author="Silvestri F." box="[631,872,1606,1633]" pageId="88" pageNumber="89" pagination="1 - 12" refId="ref108090" refString="Silvestri F. 1895 b. Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino 10 (203): 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri (1895b: 7)</bibRefCitation>
to supplement original description and to introduce gonopod terminology; non-sexual characters described based on examined topotype.
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBF946FE1653D9" blockId="88.[189,1398,1705,1767]" pageId="88" pageNumber="89">
<smallCapsWord id="8D08A070136BB1394FEBF946FEF453FC" baselines="1725,1725" box="[189,366,1706,1732]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="88" pageNumber="89">MEASUREMENTS</smallCapsWord>
. 73 body rings (1 apodous + telson). Males: body length ca
<quantity id="4CA99B49136BB1394B78F945FB1E53FA" box="[1070,1156,1705,1732]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="88" pageNumber="89" unit="mm" value="85.0">85 mm</quantity>
; maximum midbody diameter
<quantity id="4CA99B49136BB1394E7DF920FE1253D9" box="[299,392,1740,1767]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="88" pageNumber="89" unit="mm" value="3.5">3.5 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBF8E1FDF55274" blockId="88.[188,1398,1805,1866]" pageId="88" pageNumber="89">
<smallCapsWord id="8D08A070136BB1394FEBF8E1FE935218" baselines="1824,1825" box="[189,265,1805,1831]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="88" pageNumber="89">COLOR</smallCapsWord>
. Body color greyish; prozonites anteriorly greyish; metazonites with a medial greyish band and a posterior whitish; legs lighter brown.
</paragraph>
<paragraph id="8BEE36AC136BB1394FEBF89DFD6052EF" blockId="88.[188,1399,1905,2001]" pageId="88" pageNumber="89">
<smallCapsWord id="8D08A070136BB1394FEBF89DFF6652B7" baselines="1924,1924" box="[189,252,1905,1931]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="88" pageNumber="89">HEAD</smallCapsWord>
. Antennae short, just reaching back to end of ring 5 when extended dorsally; relative antennomere lengths 1&lt;2≈3&gt;4=5≈6&gt;7. Mandibular cardo with ventral margin narrow. Ommatidial cluster well-developed, elliptical; ca 30 ommatidia in 5 rows.
</paragraph>
<paragraph id="8BEE36AC136AB1384FEBFEE5FB2A5454" blockId="89.[187,1399,265,362]" pageId="89" pageNumber="90">
<smallCapsWord id="8D08A070136AB1384FEBFEE5FF64541F" baselines="284,284" box="[189,254,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="89" pageNumber="90">BODY</smallCapsWord>
<smallCapsWord id="8D08A070136AB1384E55FEE2FEDB541F" baselines="284" box="[259,321,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="89" pageNumber="90">RINGS</smallCapsWord>
. Collum with lateral lobes rounded, with ca 7 shallow striae. Very faintly constricted between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae below ozopore. Anterior sterna in midbody rings subrectangular, without transverse striae (
<figureCitation id="136A2A29136AB1384B7AFEA3FB395454" box="[1068,1187,335,362]" captionStart="Fig" captionStartId="279.[189,232,1892,1918]" captionTargetBox="[318,1270,265,1851]" captionTargetId="figure-11@279.[318,1270,265,1851]" captionTargetPageId="279" captionText="Fig. 176. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. pusilla Silvestri, 1895. B. P. bovei Silvestri, 1895. C. P. brevis Silvestri, 1902. D. P. centralis Silvestri, 1902. E. P. rugosetta Silvestri, 1897. F. P. sulcatula Silvestri, 1895. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891437" httpUri="https://zenodo.org/record/7891437/files/figure.png" pageId="89" pageNumber="90">Fig. 176F</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC136AB1384FEBFE7DFBCC5765" blockId="89.[188,1399,401,603]" pageId="89" pageNumber="90">
<smallCapsWord id="8D08A070136AB1384FEBFE7DFEAE5497" baselines="420,420" box="[189,308,401,427]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="89" pageNumber="90">GONOPODS</smallCapsWord>
. Gonocoxa (
<emphasis id="B925EABE136AB1384E9CFE7DFE6E5495" bold="true" box="[458,500,401,427]" italics="true" pageId="89" pageNumber="90">gcx</emphasis>
) subtriangular, progressively less wide (
<figureCitation id="136A2A29136AB1384C9CFE7DFBD85495" box="[970,1090,401,427]" captionStart="Fig" captionStartId="238.[189,232,1217,1243]" captionTargetBox="[189,1398,266,1175]" captionTargetId="figure-13@238.[189,1398,266,1175]" captionTargetPageId="238" captionText="Fig. 131. Pseudonannolene sulcatula Silvestri, 1895, ♀ (NHMD). A. Body rings, in lateral view, and original label. B. Posterior region in lateral view. C. Schematic drawing of right gonopod in anal view (modified from Silvestri 1895b: fig. 14). Abbreviations: see Material and methods. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891324" httpUri="https://zenodo.org/record/7891324/files/figure.png" pageId="89" pageNumber="90">Fig. 131C</figureCitation>
), with the base not arched; antero-posteriorly flattened; with rows of papillae mesally. Seminal groove (
<emphasis id="B925EABE136AB1384B07FE58FBF154F0" bold="true" box="[1105,1131,436,462]" italics="true" pageId="89" pageNumber="90">sg</emphasis>
) not visible. Shoulder absent. Telopodite (
<emphasis id="B925EABE136AB1384EFCFE34FE5B54CC" bold="true" box="[426,449,472,498]" italics="true" pageId="89" pageNumber="90">tp</emphasis>
) almost as wide as
<emphasis id="B925EABE136AB1384DE3FE34FD4554CC" bold="true" box="[693,735,472,498]" italics="true" pageId="89" pageNumber="90">gcx</emphasis>
; solenomere (
<emphasis id="B925EABE136AB1384CDBFE3BFC3B54CC" bold="true" box="[909,929,471,498]" italics="true" pageId="89" pageNumber="90">sl</emphasis>
) with apicomesal process (
<emphasis id="B925EABE136AB1384BBAFE34FABA54CC" bold="true" box="[1260,1312,472,498]" italics="true" pageId="89" pageNumber="90">amp</emphasis>
) short, subtriangular; ectal process (
<emphasis id="B925EABE136AB1384D5DFE17FDBD572B" bold="true" box="[523,551,507,533]" italics="true" pageId="89" pageNumber="90">ep</emphasis>
) inconspicuous, apparently separated from
<emphasis id="B925EABE136AB1384B77FE17FBCF572B" bold="true" box="[1057,1109,507,533]" italics="true" pageId="89" pageNumber="90">amp</emphasis>
by shallow notch;
<emphasis id="B925EABE136AB1384A66FE17FAD0572B" bold="true" box="[1328,1354,507,533]" italics="true" pageId="89" pageNumber="90">sa</emphasis>
not not visible apically. Internal branch (
<emphasis id="B925EABE136AB1384D3FFDF1FD1A5706" bold="true" box="[617,640,541,568]" italics="true" pageId="89" pageNumber="90">ib</emphasis>
) shovel-shaped, narrow; almost not surrounding basally
<emphasis id="B925EABE136AB1384A40FDF2FAB75706" bold="true" box="[1302,1325,542,568]" italics="true" pageId="89" pageNumber="90">tp</emphasis>
; setae restricted to the apical region of
<emphasis id="B925EABE136AB1384D6CFDACFDCB5765" bold="true" box="[570,593,576,603]" italics="true" pageId="89" pageNumber="90">ib</emphasis>
exceeding seminal region of
<emphasis id="B925EABE136AB1384CFDFDACFC255765" bold="true" box="[939,959,576,603]" italics="true" pageId="89" pageNumber="90">sl</emphasis>
(
<figureCitation id="136A2A29136AB1384C99FDADFBD35765" box="[975,1097,577,603]" captionStart="Fig" captionStartId="238.[189,232,1217,1243]" captionTargetBox="[189,1398,266,1175]" captionTargetId="figure-13@238.[189,1398,266,1175]" captionTargetPageId="238" captionText="Fig. 131. Pseudonannolene sulcatula Silvestri, 1895, ♀ (NHMD). A. Body rings, in lateral view, and original label. B. Posterior region in lateral view. C. Schematic drawing of right gonopod in anal view (modified from Silvestri 1895b: fig. 14). Abbreviations: see Material and methods. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891324" httpUri="https://zenodo.org/record/7891324/files/figure.png" pageId="89" pageNumber="90">Fig. 131C</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC136AB1384FEBFD6FFD2E5781" blockId="89.[188,1398,642,704]" pageId="89" pageNumber="90">
<smallCapsWord id="8D08A070136AB1384FEBFD6FFE8E57A5" baselines="662,662" box="[189,276,643,669]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="89" pageNumber="90">VULVAE</smallCapsWord>
. As typical for the genus. Bursa subtriangular, glabrous; internal valve subtriangular; operculum narrow; external valve wide, subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C34B6527136AB1384FEBFD0BFD62560E" box="[189,760,743,816]" pageId="89" pageNumber="90" type="distribution">
<paragraph id="8BEE36AC136AB1384FEBFD0BFECC563F" blockId="89.[189,342,743,769]" box="[189,342,743,769]" pageId="89" pageNumber="90">
<heading id="D0A681C0136AB1384FEBFD0BFECC563F" bold="true" box="[189,342,743,769]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="B925EABE136AB1384FEBFD0BFECC563F" bold="true" box="[189,342,743,769]" pageId="89" pageNumber="90">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC136AB1384FEBFCFAFD62560E" blockId="89.[189,760,789,816]" box="[189,760,789,816]" pageId="89" pageNumber="90">
Known only from northern
<collectingCountry id="F346763C136AB1384D56FCF9FDEF5611" box="[512,629,789,815]" name="Argentina" pageId="89" pageNumber="90">Argentina</collectingCountry>
(
<figureCitation id="136A2A29136AB1384DD2FCF9FD71560E" box="[644,747,789,816]" captionStart="Fig" captionStartId="291.[189,232,1392,1418]" captionTargetBox="[192,1368,289,1302]" captionTargetId="figure-11@291.[189,1398,265,1329]" captionTargetPageId="291" captionText="Fig. 188. Distribution map of the species Pseudonannolene segmentata Silvestri, 1895, P. silvestris Schubart, 1944, P. spelaea Iniesta &amp; Ferreira, 2013, P. strinatii Mauriès, 1974, and P. sulcatula Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891461" httpUri="https://zenodo.org/record/7891461/files/figure.png" pageId="89" pageNumber="90">Fig. 188</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B6527136AB1384FEBFCBBFC9456D8" pageId="89" pageNumber="90" type="discussion">
<paragraph id="8BEE36AC136AB1384FEBFCBBFEDE564F" blockId="89.[189,324,855,881]" box="[189,324,855,881]" pageId="89" pageNumber="90">
<heading id="D0A681C0136AB1384FEBFCBBFEDE564F" bold="true" box="[189,324,855,881]" fontSize="11" level="3" pageId="89" pageNumber="90" reason="3">
<emphasis id="B925EABE136AB1384FEBFCBBFEDE564F" bold="true" box="[189,324,855,881]" pageId="89" pageNumber="90">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC136AB1384FEBFC69FC9456D8" blockId="89.[189,1399,901,998]" pageId="89" pageNumber="90">
The
<typeStatus id="54EA880E136AB1384FA0FC6AFEB3569E" box="[246,297,902,928]" pageId="89" pageNumber="90">type</typeStatus>
material described by
<bibRefCitation id="EFC04B5D136AB1384D14FC69FC91569E" author="Silvestri F." box="[578,779,901,928]" pageId="89" pageNumber="90" pagination="1 - 12" refId="ref108090" refString="Silvestri F. 1895 b. Viaggio del dottor Alfredo Borelli nella Repubblica Argentina e nel Paraguay. XIV. Chilopodi e Diplopodi. Bollettino del musei di zoologia e di anatomia comparata della Reale Universita di Torino 10 (203): 1 - 12. https: // doi. org / 10.5962 / bhl. part. 8048" type="journal article" year="1895">Silvestri (1895b)</bibRefCitation>
and supposedly deposited at the Museo Regionale Scienze Naturali, Torino,
<collectingCountry id="F346763C136AB1384EBAFC44FDB856FC" box="[492,546,936,962]" name="Italy" pageId="89" pageNumber="90">Italy</collectingCountry>
(MRSN) (
<bibRefCitation id="EFC04B5D136AB1384DF6FC44FC1356FD" author="Viggiani V." box="[672,905,936,963]" pageId="89" pageNumber="90" pagination="351 - 417" refId="ref108936" refString="Viggiani V. 1973. Le specie descritte da Filippo Silvestri (1873 - 1949). Bollettino del Laboratorio di entomologia agraria &quot; Filippo Silvestri &quot; 30: 351 - 417." type="journal article" year="1973">Viggiani 1973: 367</bibRefCitation>
), was not found. Nevertheless, a topotype from northern
<collectingCountry id="F346763C136AB1384E31FC20FE4656D8" box="[359,476,972,998]" name="Argentina" pageId="89" pageNumber="90">Argentina</collectingCountry>
was examined (
<figureCitation id="136A2A29136AB1384DCCFC20FC9B56D8" box="[666,769,972,998]" captionStart="Fig" captionStartId="291.[189,232,1392,1418]" captionTargetBox="[192,1368,289,1302]" captionTargetId="figure-11@291.[189,1398,265,1329]" captionTargetPageId="291" captionText="Fig. 188. Distribution map of the species Pseudonannolene segmentata Silvestri, 1895, P. silvestris Schubart, 1944, P. spelaea Iniesta &amp; Ferreira, 2013, P. strinatii Mauriès, 1974, and P. sulcatula Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891461" httpUri="https://zenodo.org/record/7891461/files/figure.png" pageId="89" pageNumber="90">Fig. 188</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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