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<mods:title id="A405E12E6825C9B7C8B9D808F87891F0">Luciogobius punctilineatus n. sp., a new earthworm goby from southern Japan</mods:title>
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<mods:namePart id="4FD0180735A642D0BACB37A904311FC9">Koreeda, Reo</mods:namePart>
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<mods:affiliation id="6CF89EFEFED846E5DD2CD6944FDE5134">Graduate School of Agriculture, Forestry and Fisheries, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 0065, Japan. https: // orcid. org / 0000 - 0002 - 6977 - 3357</mods:affiliation>
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<taxonomicName id="4C514D00FFB5E93EFF04FA1AFE712681" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[151,481,1497,1524]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="138" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB5E93EFF04FA1AFE712681" bold="true" box="[151,481,1497,1524]" italics="true" pageId="1" pageNumber="138">Luciogobius punctilineatus</emphasis>
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<paragraph id="8BEE3683FFB5E93EFF04FA3DFA84256D" blockId="1.[151,1300,1497,1596]" box="[151,1300,1533,1560]" pageId="1" pageNumber="138">[New English name: Dot-lined Earthworm Goby; new standard Japanese name: Shiranui-mimizuhaze]</paragraph>
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<paragraph id="8BEE3683FFB5E93EFF04F9E2FE60254E" blockId="1.[151,1300,1497,1596]" box="[151,496,1569,1596]" pageId="1" pageNumber="138">
<figureCitation id="136A2A06FFB5E93EFF04F9E2FE8E254E" box="[151,286,1569,1595]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1810,1834]" captionStartId-1="4.[151,250,1727,1751]" captionStartId-2="6.[151,250,1976,2000]" captionStartId-3="8.[151,250,1281,1305]" captionTargetBox-0="[151,1436,179,1785]" captionTargetBox-1="[182,1433,196,1691]" captionTargetBox-2="[178,1410,1041,1953]" captionTargetBox-3="[180,1408,181,1257]" captionTargetId-0="figure-17@3.[151,1436,179,1785]" captionTargetId-1="figure-30@4.[154,1433,181,1704]" captionTargetId-2="figure-197@6.[178,1410,1035,1953]" captionTargetId-3="figure-30@8.[180,1408,181,1257]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." captionText-1="FIGURE 2. Head of the holotype of Luciogobius punctilineatus n. sp., showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively." captionText-2="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." captionText-3="FIGURE 4. Preserved specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL)." figureDoi-0="http://doi.org/10.5281/zenodo.6556882" figureDoi-1="http://doi.org/10.5281/zenodo.6556884" figureDoi-2="http://doi.org/10.5281/zenodo.6556886" figureDoi-3="http://doi.org/10.5281/zenodo.6556890" httpUri-0="https://zenodo.org/record/6556882/files/figure.png" httpUri-1="https://zenodo.org/record/6556884/files/figure.png" httpUri-2="https://zenodo.org/record/6556886/files/figure.png" httpUri-3="https://zenodo.org/record/6556890/files/figure.png" pageId="1" pageNumber="138">Figures 1–4</figureCitation>
, 
<figureCitation id="136A2A06FFB5E93EFEB9F9E2FEDF2549" box="[298,335,1569,1596]" captionStart="FIGURE 5" captionStartId="8.[151,250,1979,2003]" captionTargetBox="[174,1414,1402,1956]" captionTargetId="figure-84@8.[174,1414,1402,1956]" captionTargetPageId="8" captionText="FIGURE 5. Pectoral fin of Luciogobius punctilineatus n. sp. (A: KAUM–I. 157125) and Luciogobius sp. 11 (B: KAUM–I. 159027), showing indentation of fin membrane." figureDoi="http://doi.org/10.5281/zenodo.6556892" httpUri="https://zenodo.org/record/6556892/files/figure.png" pageId="1" pageNumber="138">5A</figureCitation>
, 
<figureCitation id="136A2A06FFB5E93EFEC9F9E2FEFA254E" box="[346,362,1569,1595]" captionStart="FIGURE 6" captionStartId="9.[151,250,805,829]" captionTargetBox="[159,1428,186,776]" captionTargetId="figure-17@9.[154,1433,181,781]" captionTargetPageId="9" captionText="FIGURE 6. Neighbor-joining tree based on 12S ribosomal RNA gene sequences (133 bp) of species of Luciogobius with Inu koma as outgroup. Numbers near branches indicate bootstrap values." figureDoi="http://doi.org/10.5281/zenodo.6556894" httpUri="https://zenodo.org/record/6556894/files/figure.png" pageId="1" pageNumber="138">6</figureCitation>
; 
<tableCitation id="C6D30338FFB5E93EFEE4F9E2FE60254E" box="[375,496,1569,1595]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="5.[151,239,151,175]" captionStartId-1="10.[151,239,151,175]" captionStartId-2="10.[151,239,1050,1074]" captionTargetBox-0="[166,1420,226,1952]" captionTargetBox-1="[151,1411,263,1001]" captionTargetBox-2="[166,1232,1090,1427]" captionTargetPageId-0="5" captionTargetPageId-1="10" captionTargetPageId-2="10" captionText-0="TABLE 1. Counts and proportional measurements of Luciogobius punctilineatus n. sp. (modes and means ± standard deviation shown in parentheses)." captionText-1="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." captionText-2="TABLE 3. Comparisons of selected morphometrics between Luciogobius punctilineatus n. sp. and Luciogobius sp. 11." httpUri-0="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" httpUri-1="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" httpUri-2="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="1" pageNumber="138" tableUuid-0="DF2E660BFFB1E93AFF04FF54FE6623A6" tableUuid-1="DF2E660BFFBEE935FF04FF54FC582382" tableUuid-2="DF2E660BFFBEE935FF04FBD9FA1D2747">Tables 1–3</tableCitation>
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sp.
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: 
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, fig. 8A–E (Shimokoshiki-shima island, Koshiki Islands, 
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, 
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<emphasis id="B925EA91FFB5E93EFF04F960FE8425CE" box="[151,276,1699,1723]" italics="true" pageId="1" pageNumber="138">Luciogobius</emphasis>
sp.
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11 (not of 
<bibRefCitation id="EFC04B72FFB5E93EFE34F960FD0625CE" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[423,662,1699,1723]" pageId="1" pageNumber="138" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB5E93EFD8DF967FDDB25CE" box="[542,587,1699,1723]" italics="true" pageId="1" pageNumber="138">et al</emphasis>
., 2019
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): 
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, fig. 4 (Kaifu, 
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, 
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<typeStatus id="54EA8821FFB5E93EFF04F927FE96258B" box="[151,262,1764,1790]" pageId="1" pageNumber="138" type="holotype">Holotype</typeStatus>
.
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–I. 156161. 
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SL, mouth of 
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, 
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<collectingRegion id="4995F861FFB5E93EFBDAF927FB5B258B" box="[1097,1227,1764,1790]" country="Japan" name="Kagoshima" pageId="1" pageNumber="138">Kagoshima</collectingRegion>
, 
<location id="8E8E6058FFB5E93EFB45F927FF5F2457" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB5E93EFB45F927FF5F2457" country="Japan" county="Furuhashi" municipality="Koreeda" name="southwestern Kyushu" pageId="1" pageNumber="138" stateProvince="Kagoshima">southwestern Kyushu</location>
, 
<collectingCountry id="F3467613FFB5E93EFF49F8CAFE8D2456" box="[218,285,1801,1827]" name="Japan" pageId="1" pageNumber="138">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB5E93EFE10F8CAFE092456" box="[387,409,1801,1827]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="138">R</collectionCode>
. 
<collectingMunicipality id="6B8AACF9FFB5E93EFE37F8CBFD932457" box="[420,515,1800,1826]" pageId="1" pageNumber="138">Koreeda</collectingMunicipality>
&amp; 
<collectionCode id="ED40AE46FFB5E93EFDB5F8CAFDAC2456" box="[550,572,1801,1827]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="138">R</collectionCode>
. 
<collectingCounty id="628F4E0FFFB5E93EFDD4F8CBFD2A2457" box="[583,698,1800,1826]" pageId="1" pageNumber="138">Furuhashi</collectingCounty>
, 
<date id="FFEF1043FFB5E93EFD56F8CBFCDB2457" box="[709,843,1800,1826]" pageId="1" pageNumber="138" value="2021-04-01">
<collectingDate id="EFABE9ABFFB5E93EFD56F8CBFCDB2457" box="[709,843,1800,1826]" pageId="1" pageNumber="138" value="2021-04-01">1 Apr. 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE3683FFB5E93DFF54F8EFFE7822FC" blockId="1.[151,1437,1764,2043]" lastBlockId="2.[151,1437,150,2004]" lastPageId="2" lastPageNumber="139" pageId="1" pageNumber="138">
<materialsCitation id="3B393CDEFFB5E93EFF54F8EFFA08248F" ID-GBIF-Occurrence="3771630309" collectingDate="2021-04-01" collectionCode="KAUM" country="Japan" county="Furuhashi" location="Kyushu" municipality="Shirinashi River" pageId="1" pageNumber="138" specimenCount="20" stateProvince="Kagoshima" typeStatus="paratype">
<emphasis id="B925EA91FFB5E93EFF54F8EFFED62433" bold="true" box="[199,326,1836,1862]" pageId="1" pageNumber="138">
<typeStatus id="54EA8821FFB5E93EFF54F8EFFED12433" box="[199,321,1836,1862]" pageId="1" pageNumber="138" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D57FD0AFFB5E93EFEDDF8EFFE772433" box="[334,487,1836,1862]" pageId="1" pageNumber="138" type="generic">20 specimens</specimenCount>
(
<quantity id="4CA99B66FFB5E93EFE65F8EFFD0C2432" box="[502,668,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.31" metricValueMin="2.09" pageId="1" pageNumber="138" unit="mm" value="27.0" valueMax="33.1" valueMin="20.9">20.9–33.1 mm</quantity>
SL). 
<emphasis id="B925EA91FFB5E93EFD4FF8EFFCD32433" bold="true" box="[732,835,1836,1862]" pageId="1" pageNumber="138">
<location id="8E8E6058FFB5E93EFD4FF8EFFCAE2433" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB5E93EFD4FF8EFFCAE2433" box="[732,830,1836,1862]" municipality="Shirinashi River" name="Kyushu" pageId="1" pageNumber="138">Kyushu</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB5E93EFCD8F8EFFC342433" box="[843,932,1836,1862]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156162. 
<quantity id="4CA99B66FFB5E93EFBBAF8EFFB1E2432" box="[1065,1166,1836,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" pageId="1" pageNumber="138" unit="mm" value="33.1">33.1 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFB56F8EFFA8E2433" box="[1221,1310,1836,1862]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156163. 
<quantity id="4CA99B66FFB5E93EFF04F893FF6C241E" box="[151,252,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" pageId="1" pageNumber="138" unit="mm" value="27.7">27.7 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFEA0F893FE1C241F" box="[307,396,1872,1898]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156164. 
<quantity id="4CA99B66FFB5E93EFD81F893FDE7241E" box="[530,631,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="1" pageNumber="138" unit="mm" value="28.2">28.2 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFD3DF893FC97241F" box="[686,775,1872,1898]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156165. 
<quantity id="4CA99B66FFB5E93EFC1EF893FC62241E" box="[909,1010,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.9899999999999998" pageId="1" pageNumber="138" unit="mm" value="29.9">29.9 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFBBAF893FB12241F" box="[1065,1154,1872,1898]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156166. 
<quantity id="4CA99B66FFB5E93EFA9BF893FAFD241E" box="[1288,1389,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="1" pageNumber="138" unit="mm" value="28.4">28.4 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFF04F8B7FF6024FB" box="[151,240,1908,1934]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 156167. 
<quantity id="4CA99B66FFB5E93EFEE8F8B7FE7224FA" box="[379,482,1908,1935]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="1" pageNumber="138" unit="mm" value="25.1">25.1 mm</quantity>
SL, collected with holotype; 
<collectionCode id="ED40AE46FFB5E93EFCAEF8B7FC0624FB" box="[829,918,1908,1934]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158051, 
<quantity id="4CA99B66FFB5E93EFBB3F8B7FB1724FA" box="[1056,1159,1908,1935]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="1" pageNumber="138" unit="mm" value="27.6">27.6 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFB50F8B7FA8C24FB" box="[1219,1308,1908,1934]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158052, 28.0 mm SL, 
<collectionCode id="ED40AE46FFB5E93EFEA0F85BFE1C24C7" box="[307,396,1944,1970]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158053, 
<quantity id="4CA99B66FFB5E93EFD81F85BFDE724C6" box="[530,631,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="1" pageNumber="138" unit="mm" value="25.8">25.8 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFD3DF85BFC9724C7" box="[686,775,1944,1970]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158054, 
<quantity id="4CA99B66FFB5E93EFC1EF85BFC6224C6" box="[909,1010,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="1" pageNumber="138" unit="mm" value="26.4">26.4 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFBBAF85BFB1224C7" box="[1065,1154,1944,1970]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158055, 
<quantity id="4CA99B66FFB5E93EFA9BF85BFAFD24C6" box="[1288,1389,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="1" pageNumber="138" unit="mm" value="24.5">24.5 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFF04F87FFF6024A3" box="[151,240,1980,2006]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158056, 
<quantity id="4CA99B66FFB5E93EFEE9F87FFE7124A2" box="[378,481,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="1" pageNumber="138" unit="mm" value="24.9">24.9 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFD8EF87FFDE624A3" box="[541,630,1980,2006]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158057, 
<quantity id="4CA99B66FFB5E93EFC93F87FFCF724A2" box="[768,871,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="1" pageNumber="138" unit="mm" value="23.5">23.5 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFC30F87FFC6C24A3" box="[931,1020,1980,2006]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 158058, 
<quantity id="4CA99B66FFB5E93EFB15F87FFB7D24A2" box="[1158,1261,1980,2007]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="138" unit="mm" value="25.4">25.4 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFABBF87FFA1124A3" box="[1320,1409,1980,2006]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 159066, 
<quantity id="4CA99B66FFB5E93EFF6BF823FECC248E" box="[248,348,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="1" pageNumber="138" unit="mm" value="28.6">28.6 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFE03F823FE79248F" box="[400,489,2016,2042]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 159067, 
<quantity id="4CA99B66FFB5E93EFDF8F823FD5E248E" box="[619,718,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="1" pageNumber="138" unit="mm" value="26.7">26.7 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB5E93EFC91F823FCCB248F" box="[770,859,2016,2042]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="1" pageNumber="138">KAUM</collectionCode>
–I. 159068, 
<quantity id="4CA99B66FFB5E93EFC4DF823FBD1248E" box="[990,1089,2016,2043]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="1" pageNumber="138" unit="mm" value="25.5">25.5 mm</quantity>
SL, mouth of 
<collectingMunicipality id="6B8AACF9FFB5E93EFB73F823FA08248F" box="[1248,1432,2016,2042]" pageId="1" pageNumber="138">Shirinashi River</collectingMunicipality>
</materialsCitation>
, 
<materialsCitation id="3B393CDEFFB6E93DFF04FF54FC2323C4" ID-GBIF-Occurrence="3771630307" box="[151,947,151,177]" collectingDate="2021-06-26" collectionCode="R" collectorName="Koreeda" country="Japan" location="Okawa" municipality="Akune" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<location id="8E8E6058FFB6E93DFF04FF54FF7923C4" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFF04FF54FF7923C4" box="[151,233,151,177]" country="Japan" municipality="Akune" name="Okawa" pageId="2" pageNumber="139" stateProvince="Kagoshima">Okawa</location>
, 
<collectingMunicipality id="6B8AACF9FFB6E93DFF67FF54FED223C4" box="[244,322,151,177]" pageId="2" pageNumber="139">Akune</collectingMunicipality>
, 
<collectingRegion id="4995F861FFB6E93DFEDDFF54FE4023C4" box="[334,464,151,177]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFE4EFF54FDB023C4" box="[477,544,151,177]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB6E93DFD1AFF54FD0F23C4" box="[649,671,151,177]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="139">R</collectionCode>
. 
<collectorName id="26A45355FFB6E93DFD38FF54FC9D23C4" box="[683,781,151,177]" pageId="2" pageNumber="139">Koreeda</collectorName>
, 
<date id="FFEF1043FFB6E93DFC8AFF54FC2323C4" box="[793,947,151,177]" pageId="2" pageNumber="139" value="2021-06-26">
<collectingDate id="EFABE9ABFFB6E93DFC8AFF54FC2323C4" box="[793,947,151,177]" pageId="2" pageNumber="139" value="2021-06-26">26 June 2021</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B393CDEFFB6E93DFC2CFF55FB48238C" ID-GBIF-Occurrence="3771630310" collectingDate="2020-10-17" collectionCode="KAUM, R" collectorName="Nakagawa" country="Japan" county="Koshiki Islands" location="Koshiki Islands" municipality="Shimokoshiki-shima island" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<emphasis id="B925EA91FFB6E93DFC2CFF55FB1423C4" bold="true" box="[959,1156,150,177]" pageId="2" pageNumber="139">
<location id="8E8E6058FFB6E93DFC2CFF55FBEF23C4" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFC2CFF55FBEF23C4" box="[959,1151,150,177]" country="Japan" county="Koshiki Islands" municipality="Shimokoshiki-shima island" name="Koshiki Islands" pageId="2" pageNumber="139" stateProvince="Kagoshima">Koshiki Islands</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB6E93DFB1FFF54FB7523C4" box="[1164,1253,151,177]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 147759, 
<quantity id="4CA99B66FFB6E93DFAF8FF54FF5323A0" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="2" pageNumber="139" unit="mm" value="24.7">24.7 mm</quantity>
SL, south of fishing port at east side of 
<collectorName id="26A45355FFB6E93DFD09FF78FCD923A0" box="[666,841,187,213]" hasSafeAffix="true" pageId="2" pageNumber="139">Nagahama Bay</collectorName>
, 
<collectorName id="26A45355FFB6E93DFCC5FF78FC4123A0" box="[854,977,187,213]" pageId="2" pageNumber="139">Nagahama</collectorName>
, 
<collectorName id="26A45355FFB6E93DFC4DFF78FBED23A0" box="[990,1149,187,213]" pageId="2" pageNumber="139">Shimokoshiki</collectorName>
, 
<collectorName id="26A45355FFB6E93DFB19FF78FAAD23A0" box="[1162,1341,187,213]" pageId="2" pageNumber="139">Satsuma-sendai</collectorName>
, 
<collectingMunicipality id="6B8AACF9FFB6E93DFAD9FF78FE15238C" pageId="2" pageNumber="139">Shimokoshiki-shima island</collectingMunicipality>
, 
<collectingCounty id="628F4E0FFFB6E93DFE00FF1CFDD7238C" box="[403,583,223,249]" pageId="2" pageNumber="139">Koshiki Islands</collectingCounty>
, 
<collectingRegion id="4995F861FFB6E93DFDC7FF1CFD46238C" box="[596,726,223,249]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFD77FF1CFCB7238C" box="[740,807,223,249]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB6E93DFC07FF1CFC3A238C" box="[916,938,223,249]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="139">R</collectionCode>
. 
<collectorName id="26A45355FFB6E93DFC24FF1CFBA0238C" box="[951,1072,223,249]" pageId="2" pageNumber="139">Nakagawa</collectorName>
, 
<date id="FFEF1043FFB6E93DFBADFF1CFB48238C" box="[1086,1240,223,249]" pageId="2" pageNumber="139" value="2020-10-17">
<collectingDate id="EFABE9ABFFB6E93DFBADFF1CFB48238C" box="[1086,1240,223,249]" pageId="2" pageNumber="139" value="2020-10-17">17 Oct. 2020</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B393CDEFFB6E93DFB76FF1DFB8B2234" ID-GBIF-Occurrence="3771630301" collectingDate="2020-09-17" collectionCode="KAUM, R" collectorName="Furuhashi" country="Japan" county="Osumi Islands" location="Osumi Islands" municipality="Tanega-shima island" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<emphasis id="B925EA91FFB6E93DFB76FF1DFA0C238C" bold="true" box="[1253,1436,222,249]" pageId="2" pageNumber="139">
<location id="8E8E6058FFB6E93DFB76FF1DFA07238C" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFB76FF1DFA07238C" box="[1253,1431,222,249]" country="Japan" county="Osumi Islands" municipality="Tanega-shima island" name="Osumi Islands" pageId="2" pageNumber="139" stateProvince="Kagoshima">Osumi Islands</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB6E93DFF04FEC0FF602268" box="[151,240,259,285]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 146246, 
<quantity id="4CA99B66FFB6E93DFEE9FEC0FE702268" box="[378,480,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="2" pageNumber="139" unit="mm" value="24.9">24.9 mm</quantity>
SL, north of 
<collectorName id="26A45355FFB6E93DFD12FEC0FCEA2268" box="[641,890,259,285]" hasSafeAffix="true" pageId="2" pageNumber="139">Shojiura Fishing Port</collectorName>
, 
<collectorName id="26A45355FFB6E93DFC14FEC0FC432268" box="[903,979,259,285]" pageId="2" pageNumber="139">Genna</collectorName>
, 
<collectorName id="26A45355FFB6E93DFC72FEC0FB132268" box="[993,1155,259,285]" pageId="2" pageNumber="139">Nishinoomote</collectorName>
, 
<collectorName id="26A45355FFB6E93DFB03FEC0FB622268" box="[1168,1266,259,285]" pageId="2" pageNumber="139">Kumage</collectorName>
, 
<collectingMunicipality id="6B8AACF9FFB6E93DFB6CFEC0FF4D2234" pageId="2" pageNumber="139">Tanega-shima island</collectingMunicipality>
, 
<collectingCounty id="628F4E0FFFB6E93DFF79FEE4FE1F2234" box="[234,399,295,321]" pageId="2" pageNumber="139">Osumi Islands</collectingCounty>
, 
<collectingRegion id="4995F861FFB6E93DFE08FEE4FD8D2234" box="[411,541,295,321]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFDB9FEE4FDFD2234" box="[554,621,295,321]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB6E93DFD4BFEE4FD7E2234" box="[728,750,295,321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="139">R</collectionCode>
. 
<collectorName id="26A45355FFB6E93DFD69FEE4FCFD2234" box="[762,877,295,321]" pageId="2" pageNumber="139">Furuhashi</collectorName>
, 
<date id="FFEF1043FFB6E93DFCEAFEE4FB8B2234" box="[889,1051,295,321]" pageId="2" pageNumber="139" value="2020-09-17">
<collectingDate id="EFABE9ABFFB6E93DFCEAFEE4FB8B2234" box="[889,1051,295,321]" pageId="2" pageNumber="139" value="2020-09-17">17 Sept. 2020</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B393CDEFFB6E93DFBB4FEE4FE7422FC" ID-GBIF-Occurrence="3771630306" collectingDate="2021-12-04" collectionCode="KAUM" collectorName="Kumage-gun &amp; D. Kato" country="Japan" county="Osumi Islands" location="Otoko River" municipality="Yaku-shima island" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<collectionCode id="ED40AE46FFB6E93DFBB4FEE4FB102234" box="[1063,1152,295,321]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 163150, 
<quantity id="4CA99B66FFB6E93DFA94FEE4FAFC2234" box="[1287,1388,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="2" pageNumber="139" unit="mm" value="20.9">20.9 mm</quantity>
SL, mouth of 
<location id="8E8E6058FFB6E93DFE94FE88FE062210" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFE94FE88FE062210" box="[263,406,331,357]" country="Japan" county="Osumi Islands" municipality="Yaku-shima island" name="Otoko River" pageId="2" pageNumber="139" stateProvince="Kagoshima">Otoko River</location>
, 
<location id="8E8E6058FFB6E93DFE30FE88FE6A2210" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFE30FE88FE6A2210" box="[419,506,331,357]" country="Japan" county="Osumi Islands" municipality="Yaku-shima island" name="Koseda" pageId="2" pageNumber="139" stateProvince="Kagoshima">Koseda</location>
, 
<location id="8E8E6058FFB6E93DFD96FE88FD152210" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFD96FE88FD152210" box="[517,645,331,357]" country="Japan" county="Osumi Islands" municipality="Yaku-shima island" name="Yakushima" pageId="2" pageNumber="139" stateProvince="Kagoshima">Yakushima</location>
, 
<collectorName id="26A45355FFB6E93DFD01FE88FCB72210" box="[658,807,331,357]" pageId="2" pageNumber="139">Kumage-gun</collectorName>
, 
<collectingMunicipality id="6B8AACF9FFB6E93DFCA0FE88FB972210" box="[819,1031,331,357]" pageId="2" pageNumber="139">Yaku-shima island</collectingMunicipality>
, 
<collectingCounty id="628F4E0FFFB6E93DFB87FE88FB292210" box="[1044,1209,331,357]" pageId="2" pageNumber="139">Osumi Islands</collectingCounty>
, 
<collectingRegion id="4995F861FFB6E93DFB55FE88FAD82210" box="[1222,1352,331,357]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFAC6FE88FA082210" box="[1365,1432,331,357]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectorName id="26A45355FFB6E93DFF61FEACFEDD22FC" box="[242,333,367,393]" pageId="2" pageNumber="139">D. Kato</collectorName>
, 
<date id="FFEF1043FFB6E93DFECAFEACFE7422FC" box="[345,484,367,393]" pageId="2" pageNumber="139" value="2021-12-04">
<collectingDate id="EFABE9ABFFB6E93DFECAFEACFE7422FC" box="[345,484,367,393]" pageId="2" pageNumber="139" value="2021-12-04">4 Dec. 2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE3683FFB6E93DFF54FE50FBAF21B8" blockId="2.[151,1437,150,2004]" pageId="2" pageNumber="139">
<emphasis id="B925EA91FFB6E93DFF54FE50FE2C22D9" bold="true" box="[199,444,402,429]" pageId="2" pageNumber="139">Non-type specimens.</emphasis>
<materialsCitation id="3B393CDEFFB6E93DFE51FE51FBB4216C" ID-GBIF-Occurrence="3771630303" collectingDate="2021-04-10" collectionCode="KAUM, R" collectorName="H. Saito &amp; Koreeda" country="India" county="Kimotsuki" location="Shikoku" municipality="Minami-osumi" pageId="2" pageNumber="139" specimenCount="15" stateProvince="Kagoshima">
<specimenCount id="9D57FD0AFFB6E93DFE51FE51FDC822D8" box="[450,600,402,429]" pageId="2" pageNumber="139" type="generic">15 specimens</specimenCount>
(
<quantity id="4CA99B66FFB6E93DFDF5FE50FC9922D8" box="[614,777,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.275" metricValueMax="3.34" metricValueMin="1.21" pageId="2" pageNumber="139" unit="mm" value="22.75" valueMax="33.4" valueMin="12.1">12.1–33.4 mm</quantity>
SL). 
<emphasis id="B925EA91FFB6E93DFCD6FE51FC2222D9" bold="true" box="[837,946,402,428]" pageId="2" pageNumber="139">
<location id="8E8E6058FFB6E93DFCD6FE51FC3D22D9" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFCD6FE51FC3D22D9" box="[837,941,402,428]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Shikoku" pageId="2" pageNumber="139" stateProvince="Kagoshima">Shikoku</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB6E93DFC2BFE50FB8122D8" box="[952,1041,403,429]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 159034, 
<quantity id="4CA99B66FFB6E93DFB00FE50FB6522D8" box="[1171,1269,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="2" pageNumber="139" unit="mm" value="24.7">24.7 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFABBFE50FA1122D8" box="[1320,1409,403,429]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 159035, 
<quantity id="4CA99B66FFB6E93DFF68FE74FEF022A4" box="[251,352,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="2" pageNumber="139" unit="mm" value="22.9">22.9 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFE0AFE74FE6222A4" box="[409,498,439,465]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 159036, 
<quantity id="4CA99B66FFB6E93DFDEAFE74FD4E22A4" box="[633,734,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="2" pageNumber="139" unit="mm" value="20.9">20.9 mm</quantity>
SL, mouth of 
<location id="8E8E6058FFB6E93DFC14FE74FB8922A4" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFC14FE74FB8922A4" box="[903,1049,439,465]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Shinjo River" pageId="2" pageNumber="139" stateProvince="Kagoshima">Shinjo River</location>
, 
<location id="8E8E6058FFB6E93DFBB6FE74FBE222A4" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFBB6FE74FBE222A4" box="[1061,1138,439,465]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Susaki" pageId="2" pageNumber="139" stateProvince="Kagoshima">Susaki</location>
, 
<collectingCountry id="F3467613FFB6E93DFB13FE74FB5722A4" box="[1152,1223,439,465]" name="India" pageId="2" pageNumber="139">Kochi</collectingCountry>
, 
<collectingCountry id="F3467613FFB6E93DFB47FE74FA8722A4" box="[1236,1303,439,465]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectorName id="26A45355FFB6E93DFA12FE74FF432280" pageId="2" pageNumber="139">H. Saito</collectorName>
, 22 
<location id="8E8E6058FFB6E93DFE93FE18FEA02280" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFE93FE18FEA02280" box="[256,304,475,501]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="July" pageId="2" pageNumber="139" stateProvince="Kagoshima">July</location>
, 2021. 
<emphasis id="B925EA91FFB6E93DFEECFE18FE762280" bold="true" box="[383,486,475,501]" pageId="2" pageNumber="139">
<location id="8E8E6058FFB6E93DFEECFE18FE712280" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFEECFE18FE712280" box="[383,481,475,501]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Kyushu" pageId="2" pageNumber="139" stateProvince="Kagoshima">Kyushu</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB6E93DFE7FFE18FDD52280" box="[492,581,475,501]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 156187, 
<quantity id="4CA99B66FFB6E93DFD54FE18FCBA2280" box="[711,810,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="139" unit="mm" value="26.3">26.3 mm</quantity>
SL, north of mouth of 
<location id="8E8E6058FFB6E93DFBB4FE18FA942280" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFBB4FE18FA942280" box="[1063,1284,475,501]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Shimadomari River" pageId="2" pageNumber="139" stateProvince="Kagoshima">Shimadomari River</location>
, 
<location id="8E8E6058FFB6E93DFA9DFE18FA082280" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFA9DFE18FA082280" box="[1294,1432,475,501]" country="India" county="Kimotsuki" municipality="Minami-osumi" name="Sataizashiki" pageId="2" pageNumber="139" stateProvince="Kagoshima">Sataizashiki</location>
, 
<collectingMunicipality id="6B8AACF9FFB6E93DFF04FE3CFED0216C" box="[151,320,511,537]" pageId="2" pageNumber="139">Minami-osumi</collectingMunicipality>
, 
<collectingCounty id="628F4E0FFFB6E93DFED8FE3CFE54216C" box="[331,452,511,537]" pageId="2" pageNumber="139">Kimotsuki</collectingCounty>
, 
<collectingRegion id="4995F861FFB6E93DFE5CFE3CFDC1216C" box="[463,593,511,537]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFDCFFE3CFD0F216C" box="[604,671,511,537]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB6E93DFC90FE3CFC89216C" box="[771,793,511,537]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="139">R</collectionCode>
. 
<collectorName id="26A45355FFB6E93DFCB7FE3CFC16216C" box="[804,902,511,537]" pageId="2" pageNumber="139">Koreeda</collectorName>
, 
<date id="FFEF1043FFB6E93DFC03FE3CFBB4216C" box="[912,1060,511,537]" pageId="2" pageNumber="139" value="2021-04-10">
<collectingDate id="EFABE9ABFFB6E93DFC03FE3CFBB4216C" box="[912,1060,511,537]" pageId="2" pageNumber="139" value="2021-04-10">10 Apr. 2021</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B393CDEFFB6E93DFBBCFE3CFA082148" ID-GBIF-Occurrence="3771630302" collectingDate="2021-04-26" collectionCode="KAUM" collectorName="H. Saito &amp; Koreeda" country="India" county="Kimotsuki" location="Shikoku" municipality="Minami-osumi" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<collectionCode id="ED40AE46FFB6E93DFBBCFE3CFB18216C" box="[1071,1160,511,537]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 157124, 
<quantity id="4CA99B66FFB6E93DFA98FE3CFAFF216C" box="[1291,1391,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="2" pageNumber="139" unit="mm" value="33.4">33.4 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFF04FDE0FF602148" box="[151,240,547,573]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 157125, 
<quantity id="4CA99B66FFB6E93DFEE2FDE0FE442148" box="[369,468,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="2" pageNumber="139" unit="mm" value="28.7">28.7 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFD94FDE0FDF02148" box="[519,608,547,573]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 157126, 
<quantity id="4CA99B66FFB6E93DFD72FDE0FCD42148" box="[737,836,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="139" unit="mm" value="26.3">26.3 mm</quantity>
SL, same locality as KAUM–I. 156187, 
<date id="FFEF1043FFB6E93DFA95FDE0FA082148" box="[1286,1432,547,573]" pageId="2" pageNumber="139" value="2021-04-26">
<collectingDate id="EFABE9ABFFB6E93DFA95FDE0FA082148" box="[1286,1432,547,573]" pageId="2" pageNumber="139" value="2021-04-26">26 Apr. 2021</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B393CDEFFB6E93DFF04FD85FBAC21B8" ID-GBIF-Occurrence="3771630308" collectingDate="2020-07-23" collectionCode="KAUM, R" collectorName="Koreeda &amp; N. Shimizu" country="Japan" county="Amamioshima island" location="Amami Islands" municipality="Heda River" pageId="2" pageNumber="139" specimenCount="1" stateProvince="Kagoshima">
<emphasis id="B925EA91FFB6E93DFF04FD85FEC72114" bold="true" box="[151,343,582,609]" pageId="2" pageNumber="139">
<location id="8E8E6058FFB6E93DFF04FD85FEC22114" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB6E93DFF04FD85FEC22114" box="[151,338,582,609]" country="Japan" county="Amamioshima island" municipality="Heda River" name="Amami Islands" pageId="2" pageNumber="139" stateProvince="Kagoshima">Amami Islands</location>
:
</emphasis>
<collectionCode id="ED40AE46FFB6E93DFECCFD84FE282114" box="[351,440,583,609]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145562, 
<quantity id="4CA99B66FFB6E93DFDACFD84FD342114" box="[575,676,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="2" pageNumber="139" unit="mm" value="12.1">12.1 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFD4FFD84FCA52114" box="[732,821,583,609]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145563, 15.0 mm SL, 
<collectionCode id="ED40AE46FFB6E93DFBCBFD84FB212114" box="[1112,1201,583,609]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145564, 
<quantity id="4CA99B66FFB6E93DFAA4FD85FA0D2114" box="[1335,1437,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="2" pageNumber="139" unit="mm" value="25.7">25.7 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFF55FDA8FE8F21F0" box="[198,287,619,645]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145565, 
<quantity id="4CA99B66FFB6E93DFE30FDA8FD9721F0" box="[419,519,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="2" pageNumber="139" unit="mm" value="26.8">26.8 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFDAFFDA8FD0521F0" box="[572,661,619,645]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145566, 24.0 mm SL, 
<collectionCode id="ED40AE46FFB6E93DFC21FDA8FB9B21F0" box="[946,1035,619,645]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145567, 
<quantity id="4CA99B66FFB6E93DFB1CFDA8FB6321F0" box="[1167,1267,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="2" pageNumber="139" unit="mm" value="28.1">28.1 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFABBFDA8FA1121F0" box="[1320,1409,619,645]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145568, 
<quantity id="4CA99B66FFB6E93DFF6AFD4CFECC21DC" box="[249,348,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="2" pageNumber="139" unit="mm" value="24.8">24.8 mm</quantity>
SL, 
<collectionCode id="ED40AE46FFB6E93DFE02FD4CFE7A21DC" box="[401,490,655,681]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="2" pageNumber="139">KAUM</collectionCode>
–I. 145569, 
<quantity id="4CA99B66FFB6E93DFDFEFD4CFD4021DC" box="[621,720,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="2" pageNumber="139" unit="mm" value="18.6">18.6 mm</quantity>
SL, south of mouth of 
<collectingMunicipality id="6B8AACF9FFB6E93DFC41FD4CFBC421DC" box="[978,1108,655,681]" pageId="2" pageNumber="139">Heda River</collectingMunicipality>
, Uken, Oshima-gun, 
<collectingCounty id="628F4E0FFFB6E93DFAACFD4CFEA521B8" pageId="2" pageNumber="139">Amamioshima island</collectingCounty>
, 
<collectingRegion id="4995F861FFB6E93DFED3FD70FE5221B8" box="[320,450,691,717]" country="Japan" name="Kagoshima" pageId="2" pageNumber="139">Kagoshima</collectingRegion>
, 
<collectingCountry id="F3467613FFB6E93DFE5DFD70FD8121B8" box="[462,529,691,717]" name="Japan" pageId="2" pageNumber="139">Japan</collectingCountry>
, coll. by 
<collectionCode id="ED40AE46FFB6E93DFDE5FD70FD1C21B8" box="[630,652,691,717]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="139">R</collectionCode>
. 
<collectorName id="26A45355FFB6E93DFD04FD70FD6621B8" box="[663,758,691,717]" pageId="2" pageNumber="139">Koreeda</collectorName>
&amp; 
<collectorName id="26A45355FFB6E93DFC89FD70FC0C21B8" box="[794,924,691,717]" pageId="2" pageNumber="139">N. Shimizu</collectorName>
, 
<date id="FFEF1043FFB6E93DFC34FD70FBAC21B8" box="[935,1084,691,717]" pageId="2" pageNumber="139" value="2020-07-23">
<collectingDate id="EFABE9ABFFB6E93DFC34FD70FBAC21B8" box="[935,1084,691,717]" pageId="2" pageNumber="139" value="2020-07-23">23 July 2020</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B6508FFB6E93DFF54FD15FD97277A" pageId="2" pageNumber="139" type="diagnosis">
<paragraph id="8BEE3683FFB6E93DFF54FD15FD97277A" blockId="2.[151,1437,150,2004]" pageId="2" pageNumber="139">
<emphasis id="B925EA91FFB6E93DFF54FD15FED12184" bold="true" box="[199,321,726,753]" pageId="2" pageNumber="139">Diagnosis.</emphasis>
A species of 
<taxonomicName id="4C514D00FFB6E93DFE45FD14FDF02184" authorityName="Gill" authorityYear="1859" box="[470,608,727,753]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="B925EA91FFB6E93DFE45FD14FDF02184" box="[470,608,727,753]" italics="true" pageId="2" pageNumber="139">Luciogobius</emphasis>
</taxonomicName>
with the following combination of characters: total second dorsal-fin rays 10–12 (modally 11); total anal-fin rays 12–14 (modally 13); pectoral-fin rays 8–12 (modally 10); vertebrae 16–18 + 22–24 = 39–42 (modally 17 + 23 = 40); pectoral-fin posterior margin slightly concave; pelvic fins united, forming a disc; snout relatively short, length 3.1–4.3% of SL; AAA distance twice body depth at anus, 11.4–16.9% of SL; snout length less than 34.7% of AAA distance; pre-anus length less than 85.5% of pre-anal-fin length; single poorly defined black longitudinal line along mid-lateral body region from behind pectoral fin to caudal-fin base, indistinct anteriorly (line embedded, visible through semi-transparent muscle tissue in fresh or live specimens); black spots forming a single longitudinal row on mid-lateral body surface from behind pectoral fin to caudal-fin base (more distinct in preserved specimens).
</paragraph>
</subSubSection>
<subSubSection id="C34B6508FFB6E938FF54FBDBFA1F2106" lastPageId="7" lastPageNumber="144" pageId="2" pageNumber="139" type="description">
<paragraph id="8BEE3683FFB6E93DFF54FBDBFAC125AC" blockId="2.[151,1437,150,2004]" pageId="2" pageNumber="139">
<emphasis id="B925EA91FFB6E93DFF54FBDBFEC92747" bold="true" box="[199,345,1048,1074]" pageId="2" pageNumber="139">Description.</emphasis>
Counts and measurements given in 
<tableCitation id="C6D30338FFB6E93DFC92FBDBFCF22747" box="[769,866,1048,1074]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1420,226,1952]" captionTargetPageId="5" captionText="TABLE 1. Counts and proportional measurements of Luciogobius punctilineatus n. sp. (modes and means ± standard deviation shown in parentheses)." httpUri="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" pageId="2" pageNumber="139" tableUuid="DF2E660BFFB1E93AFF04FF54FE6623A6">Tables 1</tableCitation>
and 
<tableCitation id="C6D30338FFB6E93DFC0CFBDBFC202747" box="[927,944,1048,1074]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[151,1411,263,1001]" captionTargetPageId="10" captionText="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="2" pageNumber="139" tableUuid="DF2E660BFFBEE935FF04FF54FC582382">2</tableCitation>
. In this description, data for 
<typeStatus id="54EA8821FFB6E93DFA90FBDBFAF52747" box="[1283,1381,1048,1074]" pageId="2" pageNumber="139" type="holotype">holotype</typeStatus>
presented first, followed by data for 
<typeStatus id="54EA8821FFB6E93DFD9DFBFEFDE92722" box="[526,633,1085,1111]" pageId="2" pageNumber="139" type="paratype">paratypes</typeStatus>
in parentheses (if different). Body elongate, slightly compressed. Head depressed. Anterior nostril with short tube reaching to upper lip; posterior nostril horizontally ovate (
<figureCitation id="136A2A06FFB6E93DFA92FBA3FAD6270F" box="[1281,1350,1120,1146]" captionStart="FIGURE 2" captionStartId="4.[151,250,1727,1751]" captionTargetBox="[182,1433,196,1691]" captionTargetId="figure-30@4.[154,1433,181,1704]" captionTargetPageId="4" captionText="FIGURE 2. Head of the holotype of Luciogobius punctilineatus n. sp., showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556884" httpUri="https://zenodo.org/record/6556884/files/figure.png" pageId="2" pageNumber="139">Fig. 2</figureCitation>
). Snout slightly swollen dorsally, especially above nostril. Eye small, rounded, covered with thin skin, located anterodorsally on head; dorsal and posterior surface of skin covering eyes rounded, not projecting posteriorly. Dorsal profile of head between eye and nape swollen (centrally concave) in dorsal view. Interorbital space wider than snout length, slightly swollen anteriorly (or with a transverse ridge anteriorly, rarely completely flat). Single low longitudinal ridge extending from below anterior nostril to behind eye (
<figureCitation id="136A2A06FFB6E93DFCBBFAD1FCFD2659" box="[808,877,1298,1324]" captionStart="FIGURE 2" captionStartId="4.[151,250,1727,1751]" captionTargetBox="[182,1433,196,1691]" captionTargetId="figure-30@4.[154,1433,181,1704]" captionTargetPageId="4" captionText="FIGURE 2. Head of the holotype of Luciogobius punctilineatus n. sp., showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556884" httpUri="https://zenodo.org/record/6556884/files/figure.png" pageId="2" pageNumber="139">Fig. 2</figureCitation>
). Mouth oblique; posterior end of maxilla extending beyond vertical through anterior margin of eye. Pair of symphysial flaps fused anteriorly, rounded. Jaws with bands of 4–6 rows of small conical teeth; outer teeth larger; upper jaw band wider. Tongue free, anteriorly bilobed. Gill opening narrow, extending below upper end of pectoral-fin base to level with posterior end of jaw.Anus located slightly behind midpoint of body. Urogenital papilla just behind anus, small and round (in both sexes; median slit in one mature female 
<typeStatus id="54EA8821FFB6E93DFEE2FA05FE412695" box="[369,465,1478,1504]" pageId="2" pageNumber="139" type="paratype">paratype</typeStatus>
slightly larger than in others). Origin of second dorsal fin posterior to anal-fin origin. Distance between anus and anal-fin origin twice body depth at anus. First and second dorsal-fin rays of second dorsal fin spinous (or only first ray spinous), remaining rays soft, segmented; sixth ray longest (fifth to seventh). First and second anal-fin rays spinous (or only first ray spinous), remaining rays soft, segmented. Pectoral fin small; membranes between adjacent rays moderately incised posteriorly for one-sixth to one-fourth (rarely one-third) of each ray pair; membrane between 2 uppermost and 2 lowermost rays not incised (sometimes incised); all rays lacking serrations; free rays absent. Sucker-shaped pelvic fins small; paired fins fused, frenum with smooth margin. Caudal fin rounded (sometimes slightly elongate). Zygapophyses on anterior base of haemal spines well developed.
</paragraph>
<paragraph id="8BEE3683FFB6E93DFF54F920FD7E24A1" blockId="2.[151,1437,150,2004]" pageId="2" pageNumber="139">
<emphasis id="B925EA91FFB6E93DFF54F920FE4E2588" box="[199,478,1763,1789]" italics="true" pageId="2" pageNumber="139">Cephalic sensory system</emphasis>
: Canals and pores absent. Cephalic sensory papillae (shown in 
<figureCitation id="136A2A06FFB6E93DFB27F920FB682588" box="[1204,1272,1763,1789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1727,1751]" captionTargetBox="[182,1433,196,1691]" captionTargetId="figure-30@4.[154,1433,181,1704]" captionTargetPageId="4" captionText="FIGURE 2. Head of the holotype of Luciogobius punctilineatus n. sp., showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556884" httpUri="https://zenodo.org/record/6556884/files/figure.png" pageId="2" pageNumber="139">Fig. 2</figureCitation>
) in four rows; row 
<emphasis id="B925EA91FFB6E93DFF5AF8C4FF472455" box="[201,215,1799,1824]" italics="true" pageId="2" pageNumber="139">a</emphasis>
located behind eye; row 
<emphasis id="B925EA91FFB6E93DFE63F8C4FE6E2454" box="[496,510,1799,1825]" italics="true" pageId="2" pageNumber="139">b</emphasis>
extending from below anterior nostril to behind eye; row 
<emphasis id="B925EA91FFB6E93DFB16F8C4FB012455" box="[1157,1169,1799,1824]" italics="true" pageId="2" pageNumber="139">c</emphasis>
located at highest point of longitudinal skin ridge on cheek, extending to behind eye; and row 
<emphasis id="B925EA91FFB6E93DFC37F8E9FC222431" box="[932,946,1834,1860]" italics="true" pageId="2" pageNumber="139">d</emphasis>
extending from snout tip to above midpoint of maxilla, along upper margin of maxilla, subsequently curving dorsoposteriorly to middle of head. Row 
<emphasis id="B925EA91FFB6E93DFAD5F88CFAF0241D" box="[1350,1376,1871,1896]" italics="true" pageId="2" pageNumber="139">cp</emphasis>
, longitudinal papillae row on lower margin of cheek swelling between rows 
<emphasis id="B925EA91FFB6E93DFC55F8B0FC4224F9" box="[966,978,1907,1932]" italics="true" pageId="2" pageNumber="139">c</emphasis>
and 
<emphasis id="B925EA91FFB6E93DFB9AF8B1FB8724F9" box="[1033,1047,1906,1932]" italics="true" pageId="2" pageNumber="139">d</emphasis>
. Row 
<emphasis id="B925EA91FFB6E93DFBF1F8B1FBFA24F9" box="[1122,1130,1906,1932]" italics="true" pageId="2" pageNumber="139">f</emphasis>
located behind symphysial flaps. Rows 
<emphasis id="B925EA91FFB6E93DFEB6F854FEA124C5" box="[293,305,1943,1968]" italics="true" pageId="2" pageNumber="139">e</emphasis>
and 
<emphasis id="B925EA91FFB6E93DFEF9F854FEE224C5" box="[362,370,1943,1968]" italics="true" pageId="2" pageNumber="139">i</emphasis>
extending along weak flap on lower margin of lower jaw to lower part of preopercle. Row 
<emphasis id="B925EA91FFB6E93DFA15F854FA0C24C5" box="[1414,1436,1943,1968]" italics="true" pageId="2" pageNumber="139">ot</emphasis>
discontinuous. Rows 
<emphasis id="B925EA91FFB6E93DFE1BF878FE3124A1" box="[392,417,1979,2004]" italics="true" pageId="2" pageNumber="139">os</emphasis>
and 
<emphasis id="B925EA91FFB6E93DFE44F878FE7D24A1" box="[471,493,1979,2004]" italics="true" pageId="2" pageNumber="139">oi</emphasis>
located behind row 
<emphasis id="B925EA91FFB6E93DFD42F878FD7724A1" box="[721,743,1979,2004]" italics="true" pageId="2" pageNumber="139">ot</emphasis>
.
</paragraph>
<caption id="DF2E660BFFB7E93CFF04F8D1FC8D24E2" ID-DOI="http://doi.org/10.5281/zenodo.6556882" ID-Zenodo-Dep="6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="3" pageNumber="140" startId="3.[151,250,1810,1834]" targetBox="[151,1436,179,1785]" targetPageId="3">
<paragraph id="8BEE3683FFB7E93CFF04F8D1FC8D24E2" blockId="3.[151,1437,1810,1943]" pageId="3" pageNumber="140">
<emphasis id="B925EA91FFB7E93CFF04F8D1FE84245E" bold="true" box="[151,276,1810,1835]" pageId="3" pageNumber="140">FIGURE 1.</emphasis>
Fresh specimens of 
<taxonomicName id="4C514D00FFB7E93CFE7BF8D0FD6D245E" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[488,765,1811,1835]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="140" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB7E93CFE7BF8D0FD6D245E" box="[488,765,1811,1835]" italics="true" pageId="3" pageNumber="140">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB7E93CFC90F8D0FCAE245E" bold="true" box="[771,830,1811,1835]" pageId="3" pageNumber="140">
<taxonomicNameLabel id="A21657EAFFB7E93CFC90F8D0FCAE245E" box="[771,830,1811,1835]" pageId="3" pageNumber="140" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively.
</paragraph>
</caption>
<caption id="DF2E660BFFB0E93BFF04F97CFDF82455" ID-DOI="http://doi.org/10.5281/zenodo.6556884" ID-Zenodo-Dep="6556884" httpUri="https://zenodo.org/record/6556884/files/figure.png" pageId="4" pageNumber="141" startId="4.[151,250,1727,1751]" targetBox="[182,1433,196,1691]" targetPageId="4">
<paragraph id="8BEE3683FFB0E93BFF04F97CFDF82455" blockId="4.[151,1437,1727,1824]" pageId="4" pageNumber="141">
<emphasis id="B925EA91FFB0E93BFF04F97CFE8425A2" bold="true" box="[151,276,1727,1751]" pageId="4" pageNumber="141">FIGURE 2.</emphasis>
Head of the holotype of 
<taxonomicName id="4C514D00FFB0E93BFD8BF903FCBC25AD" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[536,812,1728,1752]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="141" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB0E93BFD8BF903FCBC25AD" box="[536,812,1728,1752]" italics="true" pageId="4" pageNumber="141">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB0E93BFCA0F903FCFE25AD" bold="true" box="[819,878,1728,1752]" pageId="4" pageNumber="141">
<taxonomicNameLabel id="A21657EAFFB0E93BFCA0F903FCFE25AD" box="[819,878,1728,1752]" pageId="4" pageNumber="141" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively.
</paragraph>
</caption>
<caption id="DF2E660BFFB1E93AFF04FF54FE6623A6" ID-Table-UUID="DF2E660BFFB1E93AFF04FF54FE6623A6" httpUri="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" pageId="5" pageNumber="142" startId="5.[151,239,151,175]" targetBox="[166,1420,226,1952]" targetIsTable="true" targetPageId="5">
<paragraph id="8BEE3683FFB1E93AFF04FF54FE6623A6" blockId="5.[151,1436,151,211]" pageId="5" pageNumber="142">
<emphasis id="B925EA91FFB1E93AFF04FF54FE9C23DA" bold="true" box="[151,268,151,175]" pageId="5" pageNumber="142">TABLE 1.</emphasis>
Counts and proportional measurements of 
<taxonomicName id="4C514D00FFB1E93AFD76FF54FB9823DA" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[741,1032,151,175]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="142" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB1E93AFD76FF54FB9823DA" box="[741,1032,151,175]" italics="true" pageId="5" pageNumber="142">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB1E93AFB82FF54FBDF23DA" bold="true" box="[1041,1103,151,175]" pageId="5" pageNumber="142">
<taxonomicNameLabel id="A21657EAFFB1E93AFB82FF54FBDF23DA" box="[1041,1103,151,175]" pageId="5" pageNumber="142" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(modes and means ± standard deviation shown in parentheses).
</paragraph>
</caption>
<paragraph id="8BEE3683FFB1E93AFDA4FF20FAF624D5" pageId="5" pageNumber="142">
<table id="F951C423FFB116C0FF35FF21FA1C24D5" box="[166,1420,226,1952]" colsContinueIn="6.[166,1420,201,943]" gridcols="5" gridrows="37" pageId="5" pageNumber="142">
<tr id="356134C1FFB116C0FF35FF21FA1C2389" box="[166,1420,226,252]" gridrow="0" pageId="5" pageNumber="142" rowspan-0="1" rowspan-4="1">
<th id="76B05DBDFFB116C0FDA4FF21FD372389" box="[567,679,226,252]" gridcol="1" gridrow="0" pageId="5" pageNumber="142">Holotype</th>
<th id="76B05DBDFFB116C0FD5BFF21FC0F2389" box="[712,927,226,252]" gridcol="2" gridrow="0" pageId="5" pageNumber="142">Paratypes1</th>
<th id="76B05DBDFFB116C0FC51FF21FB082389" box="[962,1176,226,252]" gridcol="3" gridrow="0" pageId="5" pageNumber="142">Non-types</th>
</tr>
<tr id="356134C1FFB116C0FF35FECFFA1C223D" box="[166,1420,268,328]" gridrow="1" pageId="5" pageNumber="142" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76B05DBDFFB116C0FC51FECFFB08223D" box="[962,1176,268,328]" gridcol="3" gridrow="1" pageId="5" pageNumber="142">Amami-oshima island</td>
<td id="76B05DBDFFB116C0FB25FECFFA1C223D" box="[1206,1420,268,328]" gridcol="4" gridrow="1" pageId="5" pageNumber="142">Kochi Pref. and Osumi Pen.</td>
</tr>
<tr id="356134C1FFB116C0FF35FE9BFA1C2205" box="[166,1420,344,368]" gridrow="2" pageId="5" pageNumber="142" rowspan-0="1" rowspan-1="1">
<td id="76B05DBDFFB116C0FD5BFE9BFC0F2205" box="[712,927,344,368]" gridcol="2" gridrow="2" pageId="5" pageNumber="142">
<emphasis id="B925EA91FFB1E93AFD5BFE9AFD452205" box="[712,725,345,368]" italics="true" pageId="5" pageNumber="142">n</emphasis>
= 18
</td>
<td id="76B05DBDFFB116C0FC51FE9BFB082205" box="[962,1176,344,368]" gridcol="3" gridrow="2" pageId="5" pageNumber="142">
<emphasis id="B925EA91FFB1E93AFC51FE9AFC5F2205" box="[962,975,345,368]" italics="true" pageId="5" pageNumber="142">n</emphasis>
= 8
</td>
<td id="76B05DBDFFB116C0FB25FE9BFA1C2205" box="[1206,1420,344,368]" gridcol="4" gridrow="2" pageId="5" pageNumber="142">
<emphasis id="B925EA91FFB1E93AFB25FE9AFB532205" box="[1206,1219,345,368]" italics="true" pageId="5" pageNumber="142">n</emphasis>
= 7
</td>
</tr>
<tr id="356134C1FFB116C0FF35FE43FA1C22ED" box="[166,1420,384,408]" gridrow="3" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FE43FD9022ED" box="[166,512,384,408]" gridcol="0" gridrow="3" pageId="5" pageNumber="142">Standard length (SL; mm)</th>
<td id="76B05DBDFFB116C0FDA4FE43FD3722ED" box="[567,679,384,408]" gridcol="1" gridrow="3" pageId="5" pageNumber="142">29.8</td>
<td id="76B05DBDFFB116C0FD5BFE43FC0F22ED" box="[712,927,384,408]" gridcol="2" gridrow="3" pageId="5" pageNumber="142">20.9–33.1</td>
<td id="76B05DBDFFB116C0FC51FE43FB0822ED" box="[962,1176,384,408]" gridcol="3" gridrow="3" pageId="5" pageNumber="142">12.1–28.1</td>
<td id="76B05DBDFFB116C0FB25FE43FA1C22ED" box="[1206,1420,384,408]" gridcol="4" gridrow="3" pageId="5" pageNumber="142">20.9–33.4</td>
</tr>
<tr id="356134C1FFB116C0FF35FE6BFA1C22B5" box="[166,1420,424,448]" gridrow="4" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FE6BFA1C22B5" box="[166,1420,424,448]" colspan="5" colspanRight="4" gridcol="0" gridrow="4" pageId="5" pageNumber="142">Counts</th>
</tr>
<tr id="356134C1FFB116C0FF35FE12FA1C229C" box="[166,1420,465,489]" gridrow="5" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FE12FD90229C" box="[166,512,465,489]" gridcol="0" gridrow="5" pageId="5" pageNumber="142">Total dorsal-fin elements</th>
<td id="76B05DBDFFB116C0FDA4FE12FD37229C" box="[567,679,465,489]" gridcol="1" gridrow="5" pageId="5" pageNumber="142">11</td>
<td id="76B05DBDFFB116C0FD5BFE12FC0F229C" box="[712,927,465,489]" gridcol="2" gridrow="5" pageId="5" pageNumber="142">10–12</td>
<td id="76B05DBDFFB116C0FC51FE12FB08229C" box="[962,1176,465,489]" gridcol="3" gridrow="5" pageId="5" pageNumber="142">10–11 (10)</td>
<td id="76B05DBDFFB116C0FB25FE12FA1C229C" box="[1206,1420,465,489]" gridcol="4" gridrow="5" pageId="5" pageNumber="142">10–12 (11)</td>
</tr>
<tr id="356134C1FFB116C0FF35FE3AFA1C2164" box="[166,1420,505,529]" gridrow="6" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FE3AFD902164" box="[166,512,505,529]" gridcol="0" gridrow="6" pageId="5" pageNumber="142">Total anal-fin elements</th>
<td id="76B05DBDFFB116C0FDA4FE3AFD372164" box="[567,679,505,529]" gridcol="1" gridrow="6" pageId="5" pageNumber="142">13</td>
<td id="76B05DBDFFB116C0FD5BFE3AFC0F2164" box="[712,927,505,529]" gridcol="2" gridrow="6" pageId="5" pageNumber="142">12–14</td>
<td id="76B05DBDFFB116C0FC51FE3AFB082164" box="[962,1176,505,529]" gridcol="3" gridrow="6" pageId="5" pageNumber="142">12–14 (12)</td>
<td id="76B05DBDFFB116C0FB25FE3AFA1C2164" box="[1206,1420,505,529]" gridcol="4" gridrow="6" pageId="5" pageNumber="142">12–14 (13)</td>
</tr>
<tr id="356134C1FFB116C0FF35FDE2FA1C214F" box="[166,1420,545,570]" gridrow="7" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FDE2FD90214F" box="[166,512,545,570]" gridcol="0" gridrow="7" pageId="5" pageNumber="142">Pectoral-fin rays</th>
<td id="76B05DBDFFB116C0FDA4FDE2FD37214F" box="[567,679,545,570]" gridcol="1" gridrow="7" pageId="5" pageNumber="142">11</td>
<td id="76B05DBDFFB116C0FD5BFDE2FC0F214F" box="[712,927,545,570]" gridcol="2" gridrow="7" pageId="5" pageNumber="142">9–12</td>
<td id="76B05DBDFFB116C0FC51FDE2FB08214F" box="[962,1176,545,570]" gridcol="3" gridrow="7" pageId="5" pageNumber="142">8–12 (9)</td>
<td id="76B05DBDFFB116C0FB25FDE2FA1C214F" box="[1206,1420,545,570]" gridcol="4" gridrow="7" pageId="5" pageNumber="142">9–11 (10)</td>
</tr>
<tr id="356134C1FFB116C0FF35FD8AFA1C2117" box="[166,1420,585,610]" gridrow="8" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FD8AFD902117" box="[166,512,585,610]" gridcol="0" gridrow="8" pageId="5" pageNumber="142">Caudal-fin segmented rays</th>
<td id="76B05DBDFFB116C0FDA4FD8AFD372117" box="[567,679,585,610]" gridcol="1" gridrow="8" pageId="5" pageNumber="142">9 + 8</td>
<td id="76B05DBDFFB116C0FD5BFD8AFC0F2117" box="[712,927,585,610]" gridcol="2" gridrow="8" pageId="5" pageNumber="142">9–10 + 7–8</td>
<td id="76B05DBDFFB116C0FC51FD8AFB082117" box="[962,1176,585,610]" gridcol="3" gridrow="8" pageId="5" pageNumber="142">9–10 + 7–8</td>
<td id="76B05DBDFFB116C0FB25FD8AFA1C2117" box="[1206,1420,585,610]" gridcol="4" gridrow="8" pageId="5" pageNumber="142">9–10 + 8</td>
</tr>
<tr id="356134C1FFB116C0FF35FDB2FA1C21FF" box="[166,1420,625,650]" gridrow="9" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FDB2FD9021FF" box="[166,512,625,650]" gridcol="0" gridrow="9" pageId="5" pageNumber="142">Pelvic-fin rays</th>
<td id="76B05DBDFFB116C0FDA4FDB2FD3721FF" box="[567,679,625,650]" gridcol="1" gridrow="9" pageId="5" pageNumber="142">I, 5</td>
<td id="76B05DBDFFB116C0FD5BFDB2FC0F21FF" box="[712,927,625,650]" gridcol="2" gridrow="9" pageId="5" pageNumber="142">I, 5</td>
<td id="76B05DBDFFB116C0FC51FDB2FB0821FF" box="[962,1176,625,650]" gridcol="3" gridrow="9" pageId="5" pageNumber="142">I, 5</td>
<td id="76B05DBDFFB116C0FB25FDB2FA1C21FF" box="[1206,1420,625,650]" gridcol="4" gridrow="9" pageId="5" pageNumber="142">I, 5</td>
</tr>
<tr id="356134C1FFB116C0FF35FD59FA1C2037" box="[166,1420,666,834]" gridrow="10" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FD59FD902037" box="[166,512,666,834]" gridcol="0" gridrow="10" pageId="5" pageNumber="142">Vertebrae</th>
<td id="76B05DBDFFB116C0FDA4FD59FD372037" box="[567,679,666,834]" gridcol="1" gridrow="10" pageId="5" pageNumber="142">17 + 23 = 40</td>
<td id="76B05DBDFFB116C0FD5BFD59FC0F2037" box="[712,927,666,834]" gridcol="2" gridrow="10" pageId="5" pageNumber="142">17 + 23 = 40 (8), 17 + 24 = 41 (1), 18 + 22 = 40 (1), 18 + 23 = 41 (6), 18 + 24 = 42 (1)</td>
<td id="76B05DBDFFB116C0FC51FD59FB082037" box="[962,1176,666,834]" gridcol="3" gridrow="10" pageId="5" pageNumber="142">18 + 24 = 42 (6)</td>
<td id="76B05DBDFFB116C0FB25FD59FA1C2037" box="[1206,1420,666,834]" gridcol="4" gridrow="10" pageId="5" pageNumber="142">16 + 23 = 39 (1), 16 + 22 = 39 (2), 17 + 23 = 40 (1), 18 + 23 = 41 (2), 18 + 24 = 42 (1)</td>
</tr>
<tr id="356134C1FFB116C0FF35FC91FA1C20FB" box="[166,1420,850,910]" gridrow="11" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FC91FD9020FB" box="[166,512,850,910]" gridcol="0" gridrow="11" pageId="5" pageNumber="142">Dorsal-fin pterygiophore formula (P-V)</th>
<td id="76B05DBDFFB116C0FDA4FC91FD3720FB" box="[567,679,850,910]" gridcol="1" gridrow="11" pageId="5" pageNumber="142">24</td>
<td id="76B05DBDFFB116C0FD5BFC91FC0F20FB" box="[712,927,850,910]" gridcol="2" gridrow="11" pageId="5" pageNumber="142">23–24-25 (23-24)</td>
<td id="76B05DBDFFB116C0FC51FC91FB0820FB" box="[962,1176,850,910]" gridcol="3" gridrow="11" pageId="5" pageNumber="142">24-25–25-26</td>
<td id="76B05DBDFFB116C0FB25FC91FA1C20FB" box="[1206,1420,850,910]" gridcol="4" gridrow="11" pageId="5" pageNumber="142">22-23–25</td>
</tr>
<tr id="356134C1FFB116C0FF35FC5DFA1C20AF" box="[166,1420,926,986]" gridrow="12" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FC5DFD9020AF" box="[166,512,926,986]" gridcol="0" gridrow="12" pageId="5" pageNumber="142">First anal-fin pterygiophore insertion behind</th>
<td id="76B05DBDFFB116C0FDA4FC5DFD3720AF" box="[567,679,926,986]" gridcol="1" gridrow="12" pageId="5" pageNumber="142">5th haemal spine</td>
<td id="76B05DBDFFB116C0FD5BFC5DFC0F20AF" box="[712,927,926,986]" gridcol="2" gridrow="12" pageId="5" pageNumber="142">4–6th (5th) haemal spine</td>
<td id="76B05DBDFFB116C0FC51FC5DFB0820AF" box="[962,1176,926,986]" gridcol="3" gridrow="12" pageId="5" pageNumber="142">5–6th (5th) haemal spine</td>
<td id="76B05DBDFFB116C0FB25FC5DFA1C20AF" box="[1206,1420,926,986]" gridcol="4" gridrow="12" pageId="5" pageNumber="142">4–6th (5th) haemal spine</td>
</tr>
<tr id="356134C1FFB116C0FF35FC28FA1C2776" box="[166,1420,1003,1027]" gridrow="13" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FC28FA1C2776" box="[166,1420,1003,1027]" colspan="5" colspanRight="4" gridcol="0" gridrow="13" pageId="5" pageNumber="142">Measurements (% SL)</th>
</tr>
<tr id="356134C1FFB116C0FF35FBD0FA1C275E" box="[166,1420,1043,1067]" gridrow="14" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FBD0FD90275E" box="[166,512,1043,1067]" gridcol="0" gridrow="14" pageId="5" pageNumber="142">Head length</th>
<td id="76B05DBDFFB116C0FDA4FBD0FD37275E" box="[567,679,1043,1067]" gridcol="1" gridrow="14" pageId="5" pageNumber="142">15.1</td>
<td id="76B05DBDFFB116C0FD5BFBD0FC0F275E" box="[712,927,1043,1067]" gridcol="2" gridrow="14" pageId="5" pageNumber="142">13.1–14.8 (13.9±0.4)</td>
<td id="76B05DBDFFB116C0FC51FBD0FB08275E" box="[962,1176,1043,1067]" gridcol="3" gridrow="14" pageId="5" pageNumber="142">13.2–16.4 (14.6±1.1)</td>
<td id="76B05DBDFFB116C0FB25FBD0FA1C275E" box="[1206,1420,1043,1067]" gridcol="4" gridrow="14" pageId="5" pageNumber="142">12.7–16.6 (14.8±1.7)</td>
</tr>
<tr id="356134C1FFB116C0FF35FBF8FA1C2726" box="[166,1420,1083,1107]" gridrow="15" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FBF8FD902726" box="[166,512,1083,1107]" gridcol="0" gridrow="15" pageId="5" pageNumber="142">Head depth</th>
<td id="76B05DBDFFB116C0FDA4FBF8FD372726" box="[567,679,1083,1107]" gridcol="1" gridrow="15" pageId="5" pageNumber="142">5.6</td>
<td id="76B05DBDFFB116C0FD5BFBF8FC0F2726" box="[712,927,1083,1107]" gridcol="2" gridrow="15" pageId="5" pageNumber="142">4.8–6.6 (5.6±0.5)</td>
<td id="76B05DBDFFB116C0FC51FBF8FB082726" box="[962,1176,1083,1107]" gridcol="3" gridrow="15" pageId="5" pageNumber="142">5.0–6.0 (5.4±0.4)</td>
<td id="76B05DBDFFB116C0FB25FBF8FA1C2726" box="[1206,1420,1083,1107]" gridcol="4" gridrow="15" pageId="5" pageNumber="142">5.3–7.6 (6.1±0.8)</td>
</tr>
<tr id="356134C1FFB116C0FF35FBA0FA1C270E" box="[166,1420,1123,1147]" gridrow="16" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FBA0FD90270E" box="[166,512,1123,1147]" gridcol="0" gridrow="16" pageId="5" pageNumber="142">Head width</th>
<td id="76B05DBDFFB116C0FDA4FBA0FD37270E" box="[567,679,1123,1147]" gridcol="1" gridrow="16" pageId="5" pageNumber="142">8.1</td>
<td id="76B05DBDFFB116C0FD5BFBA0FC0F270E" box="[712,927,1123,1147]" gridcol="2" gridrow="16" pageId="5" pageNumber="142">6.4–8.2 (7.4±0.5)</td>
<td id="76B05DBDFFB116C0FC51FBA0FB08270E" box="[962,1176,1123,1147]" gridcol="3" gridrow="16" pageId="5" pageNumber="142">5.3–7.1 (6.3±0.6)</td>
<td id="76B05DBDFFB116C0FB25FBA0FA1C270E" box="[1206,1420,1123,1147]" gridcol="4" gridrow="16" pageId="5" pageNumber="142">7.0–9.3 (8.1±0.9)</td>
</tr>
<tr id="356134C1FFB116C0FF35FB48FA1C27D1" box="[166,1420,1163,1188]" gridrow="17" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FB48FD9027D1" box="[166,512,1163,1188]" gridcol="0" gridrow="17" pageId="5" pageNumber="142">Snout length</th>
<td id="76B05DBDFFB116C0FDA4FB48FD3727D1" box="[567,679,1163,1188]" gridcol="1" gridrow="17" pageId="5" pageNumber="142">3.6</td>
<td id="76B05DBDFFB116C0FD5BFB48FC0F27D1" box="[712,927,1163,1188]" gridcol="2" gridrow="17" pageId="5" pageNumber="142">3.1–3.6 (3.4±0.2)</td>
<td id="76B05DBDFFB116C0FC51FB48FB0827D1" box="[962,1176,1163,1188]" gridcol="3" gridrow="17" pageId="5" pageNumber="142">3.1–4.3 (3.7±0.5)</td>
<td id="76B05DBDFFB116C0FB25FB48FA1C27D1" box="[1206,1420,1163,1188]" gridcol="4" gridrow="17" pageId="5" pageNumber="142">3.3–3.8 (3.5±0.2)</td>
</tr>
<tr id="356134C1FFB116C0FF35FB77FA1C27B9" box="[166,1420,1204,1228]" gridrow="18" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FB77FD9027B9" box="[166,512,1204,1228]" gridcol="0" gridrow="18" pageId="5" pageNumber="142">Upper-jaw length</th>
<td id="76B05DBDFFB116C0FDA4FB77FD3727B9" box="[567,679,1204,1228]" gridcol="1" gridrow="18" pageId="5" pageNumber="142">4.8</td>
<td id="76B05DBDFFB116C0FD5BFB77FC0F27B9" box="[712,927,1204,1228]" gridcol="2" gridrow="18" pageId="5" pageNumber="142">3.8–5.2 (4.6±0.3)</td>
<td id="76B05DBDFFB116C0FC51FB77FB0827B9" box="[962,1176,1204,1228]" gridcol="3" gridrow="18" pageId="5" pageNumber="142">4.1–5.2 (4.7±0.4)</td>
<td id="76B05DBDFFB116C0FB25FB77FA1C27B9" box="[1206,1420,1204,1228]" gridcol="4" gridrow="18" pageId="5" pageNumber="142">4.2–5.3 (4.8±0.4)</td>
</tr>
<tr id="356134C1FFB116C0FF35FB1FFA1C2781" box="[166,1420,1244,1268]" gridrow="19" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FB1FFD902781" box="[166,512,1244,1268]" gridcol="0" gridrow="19" pageId="5" pageNumber="142">Eye diameter</th>
<td id="76B05DBDFFB116C0FDA4FB1FFD372781" box="[567,679,1244,1268]" gridcol="1" gridrow="19" pageId="5" pageNumber="142">1.0</td>
<td id="76B05DBDFFB116C0FD5BFB1FFC0F2781" box="[712,927,1244,1268]" gridcol="2" gridrow="19" pageId="5" pageNumber="142">0.9–1.2 (1.1±0.1)</td>
<td id="76B05DBDFFB116C0FC51FB1FFB082781" box="[962,1176,1244,1268]" gridcol="3" gridrow="19" pageId="5" pageNumber="142">0.7–1.6 (1.0±0.3)</td>
<td id="76B05DBDFFB116C0FB25FB1FFA1C2781" box="[1206,1420,1244,1268]" gridcol="4" gridrow="19" pageId="5" pageNumber="142">0.9–1.3 (1.1±0.2)</td>
</tr>
<tr id="356134C1FFB116C0FF35FAC7FA1C2669" box="[166,1420,1284,1308]" gridrow="20" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FAC7FD902669" box="[166,512,1284,1308]" gridcol="0" gridrow="20" pageId="5" pageNumber="142">Interorbital width</th>
<td id="76B05DBDFFB116C0FDA4FAC7FD372669" box="[567,679,1284,1308]" gridcol="1" gridrow="20" pageId="5" pageNumber="142">2.7</td>
<td id="76B05DBDFFB116C0FD5BFAC7FC0F2669" box="[712,927,1284,1308]" gridcol="2" gridrow="20" pageId="5" pageNumber="142">1.8–2.5 (2.1±0.2)</td>
<td id="76B05DBDFFB116C0FC51FAC7FB082669" box="[962,1176,1284,1308]" gridcol="3" gridrow="20" pageId="5" pageNumber="142">1.7–2.3 (2.0±0.2)</td>
<td id="76B05DBDFFB116C0FB25FAC7FA1C2669" box="[1206,1420,1284,1308]" gridcol="4" gridrow="20" pageId="5" pageNumber="142">1.8–3.1 (2.5±0.5)</td>
</tr>
<tr id="356134C1FFB116C0FF35FAEFFA1C2631" box="[166,1420,1324,1348]" gridrow="21" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FAEFFD902631" box="[166,512,1324,1348]" gridcol="0" gridrow="21" pageId="5" pageNumber="142">Body depth at pelvic-fin origin</th>
<td id="76B05DBDFFB116C0FDA4FAEFFD372631" box="[567,679,1324,1348]" gridcol="1" gridrow="21" pageId="5" pageNumber="142">5.9</td>
<td id="76B05DBDFFB116C0FD5BFAEFFC0F2631" box="[712,927,1324,1348]" gridcol="2" gridrow="21" pageId="5" pageNumber="142">4.7–6.5 (5.5±0.5)</td>
<td id="76B05DBDFFB116C0FC51FAEFFB082631" box="[962,1176,1324,1348]" gridcol="3" gridrow="21" pageId="5" pageNumber="142">4.0–5.7 (5.1±0.6)</td>
<td id="76B05DBDFFB116C0FB25FAEFFA1C2631" box="[1206,1420,1324,1348]" gridcol="4" gridrow="21" pageId="5" pageNumber="142">5.1–6.9 (6.1±0.6)</td>
</tr>
<tr id="356134C1FFB116C0FF35FA97FA1C2618" box="[166,1420,1364,1389]" gridrow="22" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FA97FD902618" box="[166,512,1364,1389]" gridcol="0" gridrow="22" pageId="5" pageNumber="142">Body depth at anus</th>
<td id="76B05DBDFFB116C0FDA4FA97FD372618" box="[567,679,1364,1389]" gridcol="1" gridrow="22" pageId="5" pageNumber="142">6.4</td>
<td id="76B05DBDFFB116C0FD5BFA97FC0F2618" box="[712,927,1364,1389]" gridcol="2" gridrow="22" pageId="5" pageNumber="142">5.1–7.3 (6.1±0.7)</td>
<td id="76B05DBDFFB116C0FC51FA97FB082618" box="[962,1176,1364,1389]" gridcol="3" gridrow="22" pageId="5" pageNumber="142">4.9–6.5 (5.4±0.5)</td>
<td id="76B05DBDFFB116C0FB25FA97FA1C2618" box="[1206,1420,1364,1389]" gridcol="4" gridrow="22" pageId="5" pageNumber="142">6.1–7.2 (6.6±0.5)</td>
</tr>
<tr id="356134C1FFB116C0FF35FABEFA1C26E0" box="[166,1420,1405,1429]" gridrow="23" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FABEFD9026E0" box="[166,512,1405,1429]" gridcol="0" gridrow="23" pageId="5" pageNumber="142">Body depth at anal-fin origin</th>
<td id="76B05DBDFFB116C0FDA4FABEFD3726E0" box="[567,679,1405,1429]" gridcol="1" gridrow="23" pageId="5" pageNumber="142">6.8</td>
<td id="76B05DBDFFB116C0FD5BFABEFC0F26E0" box="[712,927,1405,1429]" gridcol="2" gridrow="23" pageId="5" pageNumber="142">5.6–7.5 (6.7±0.6)</td>
<td id="76B05DBDFFB116C0FC51FABEFB0826E0" box="[962,1176,1405,1429]" gridcol="3" gridrow="23" pageId="5" pageNumber="142">5.1–6.7 (5.8±0.6)</td>
<td id="76B05DBDFFB116C0FB25FABEFA1C26E0" box="[1206,1420,1405,1429]" gridcol="4" gridrow="23" pageId="5" pageNumber="142">6.3–8.1 (7.2±0.7)</td>
</tr>
<tr id="356134C1FFB116C0FF35FA66FA1C26C8" box="[166,1420,1445,1469]" gridrow="24" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FA66FD9026C8" box="[166,512,1445,1469]" gridcol="0" gridrow="24" pageId="5" pageNumber="142">Body width</th>
<td id="76B05DBDFFB116C0FDA4FA66FD3726C8" box="[567,679,1445,1469]" gridcol="1" gridrow="24" pageId="5" pageNumber="142">6.0</td>
<td id="76B05DBDFFB116C0FD5BFA66FC0F26C8" box="[712,927,1445,1469]" gridcol="2" gridrow="24" pageId="5" pageNumber="142">5.1–6.6 (5.7±0.4)</td>
<td id="76B05DBDFFB116C0FC51FA66FB0826C8" box="[962,1176,1445,1469]" gridcol="3" gridrow="24" pageId="5" pageNumber="142">3.9–5.7 (4.9±0.6)</td>
<td id="76B05DBDFFB116C0FB25FA66FA1C26C8" box="[1206,1420,1445,1469]" gridcol="4" gridrow="24" pageId="5" pageNumber="142">5.4–6.7 (6.1±0.4)</td>
</tr>
<tr id="356134C1FFB116C0FF35FA0EFA1C2693" box="[166,1420,1485,1510]" gridrow="25" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FA0EFD902693" box="[166,512,1485,1510]" gridcol="0" gridrow="25" pageId="5" pageNumber="142">AAA distance</th>
<td id="76B05DBDFFB116C0FDA4FA0EFD372693" box="[567,679,1485,1510]" gridcol="1" gridrow="25" pageId="5" pageNumber="142">13.1</td>
<td id="76B05DBDFFB116C0FD5BFA0EFC0F2693" box="[712,927,1485,1510]" gridcol="2" gridrow="25" pageId="5" pageNumber="142">11.4–14.6 (12.9±0.9)</td>
<td id="76B05DBDFFB116C0FC51FA0EFB082693" box="[962,1176,1485,1510]" gridcol="3" gridrow="25" pageId="5" pageNumber="142">13.8–16.9 (15.1±1.0)</td>
<td id="76B05DBDFFB116C0FB25FA0EFA1C2693" box="[1206,1420,1485,1510]" gridcol="4" gridrow="25" pageId="5" pageNumber="142">11.7–13.4 (12.5±0.6)</td>
</tr>
<tr id="356134C1FFB116C0FF35FA36FA1C257B" box="[166,1420,1525,1550]" gridrow="26" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35FA36FD90257B" box="[166,512,1525,1550]" gridcol="0" gridrow="26" pageId="5" pageNumber="142">Least caudal-peduncle depth</th>
<td id="76B05DBDFFB116C0FDA4FA36FD37257B" box="[567,679,1525,1550]" gridcol="1" gridrow="26" pageId="5" pageNumber="142">6.1</td>
<td id="76B05DBDFFB116C0FD5BFA36FC0F257B" box="[712,927,1525,1550]" gridcol="2" gridrow="26" pageId="5" pageNumber="142">4.9–6.8 (5.9±0.5)</td>
<td id="76B05DBDFFB116C0FC51FA36FB08257B" box="[962,1176,1525,1550]" gridcol="3" gridrow="26" pageId="5" pageNumber="142">4.4–5.7 (5.3±0.5)</td>
<td id="76B05DBDFFB116C0FB25FA36FA1C257B" box="[1206,1420,1525,1550]" gridcol="4" gridrow="26" pageId="5" pageNumber="142">5.7–7.3 (6.5±0.7)</td>
</tr>
<tr id="356134C1FFB116C0FF35F9DDFA1C2543" box="[166,1420,1566,1590]" gridrow="27" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F9DDFD902543" box="[166,512,1566,1590]" gridcol="0" gridrow="27" pageId="5" pageNumber="142">Maximum caudal-peduncle depth</th>
<td id="76B05DBDFFB116C0FDA4F9DDFD372543" box="[567,679,1566,1590]" gridcol="1" gridrow="27" pageId="5" pageNumber="142">6.4</td>
<td id="76B05DBDFFB116C0FD5BF9DDFC0F2543" box="[712,927,1566,1590]" gridcol="2" gridrow="27" pageId="5" pageNumber="142">5.5–6.7 (6.1±0.4)</td>
<td id="76B05DBDFFB116C0FC51F9DDFB082543" box="[962,1176,1566,1590]" gridcol="3" gridrow="27" pageId="5" pageNumber="142">4.8–5.9 (5.6±0.3)</td>
<td id="76B05DBDFFB116C0FB25F9DDFA1C2543" box="[1206,1420,1566,1590]" gridcol="4" gridrow="27" pageId="5" pageNumber="142">5.5–8.1 (6.8±1.1)</td>
</tr>
<tr id="356134C1FFB116C0FF35F985FA1C252B" box="[166,1420,1606,1630]" gridrow="28" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F985FD90252B" box="[166,512,1606,1630]" gridcol="0" gridrow="28" pageId="5" pageNumber="142">Caudal-peduncle length</th>
<td id="76B05DBDFFB116C0FDA4F985FD37252B" box="[567,679,1606,1630]" gridcol="1" gridrow="28" pageId="5" pageNumber="142">18.2</td>
<td id="76B05DBDFFB116C0FD5BF985FC0F252B" box="[712,927,1606,1630]" gridcol="2" gridrow="28" pageId="5" pageNumber="142">15.8–19.1 (17.6±1.0)</td>
<td id="76B05DBDFFB116C0FC51F985FB08252B" box="[962,1176,1606,1630]" gridcol="3" gridrow="28" pageId="5" pageNumber="142">15.7–18.5 (17.0±1.0)</td>
<td id="76B05DBDFFB116C0FB25F985FA1C252B" box="[1206,1420,1606,1630]" gridcol="4" gridrow="28" pageId="5" pageNumber="142">17.0–18.8 (17.5±0.6)</td>
</tr>
<tr id="356134C1FFB116C0FF35F9ADFA1C25F3" box="[166,1420,1646,1670]" gridrow="29" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F9ADFD9025F3" box="[166,512,1646,1670]" gridcol="0" gridrow="29" pageId="5" pageNumber="142">Pre-anus length</th>
<td id="76B05DBDFFB116C0FDA4F9ADFD3725F3" box="[567,679,1646,1670]" gridcol="1" gridrow="29" pageId="5" pageNumber="142">52.7</td>
<td id="76B05DBDFFB116C0FD5BF9ADFC0F25F3" box="[712,927,1646,1670]" gridcol="2" gridrow="29" pageId="5" pageNumber="142">52.5–56.4 (54.1±0.9)</td>
<td id="76B05DBDFFB116C0FC51F9ADFB0825F3" box="[962,1176,1646,1670]" gridcol="3" gridrow="29" pageId="5" pageNumber="142">52.2–60.3 (55.5±2.9)</td>
<td id="76B05DBDFFB116C0FB25F9ADFA1C25F3" box="[1206,1420,1646,1670]" gridcol="4" gridrow="29" pageId="5" pageNumber="142">52.6–54.9 (53.9±1.0)</td>
</tr>
<tr id="356134C1FFB116C0FF35F955FA1C25DA" box="[166,1420,1686,1711]" gridrow="30" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F955FD9025DA" box="[166,512,1686,1711]" gridcol="0" gridrow="30" pageId="5" pageNumber="142">Pre-second dorsal-fin length</th>
<td id="76B05DBDFFB116C0FDA4F955FD3725DA" box="[567,679,1686,1711]" gridcol="1" gridrow="30" pageId="5" pageNumber="142">70.6</td>
<td id="76B05DBDFFB116C0FD5BF955FC0F25DA" box="[712,927,1686,1711]" gridcol="2" gridrow="30" pageId="5" pageNumber="142">69.2–73.2 (71.2±1.3)</td>
<td id="76B05DBDFFB116C0FC51F955FB0825DA" box="[962,1176,1686,1711]" gridcol="3" gridrow="30" pageId="5" pageNumber="142">71.3–75.7 (73.9±1.4)</td>
<td id="76B05DBDFFB116C0FB25F955FA1C25DA" box="[1206,1420,1686,1711]" gridcol="4" gridrow="30" pageId="5" pageNumber="142">70.2–73.4 (71.8±1.3)</td>
</tr>
<tr id="356134C1FFB116C0FF35F97CFA1C25A2" box="[166,1420,1727,1751]" gridrow="31" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F97CFD9025A2" box="[166,512,1727,1751]" gridcol="0" gridrow="31" pageId="5" pageNumber="142">Pre-anal-fin length</th>
<td id="76B05DBDFFB116C0FDA4F97CFD3725A2" box="[567,679,1727,1751]" gridcol="1" gridrow="31" pageId="5" pageNumber="142">67.3</td>
<td id="76B05DBDFFB116C0FD5BF97CFC0F25A2" box="[712,927,1727,1751]" gridcol="2" gridrow="31" pageId="5" pageNumber="142">66.1–69.8 (68.0±0.9)</td>
<td id="76B05DBDFFB116C0FC51F97CFB0825A2" box="[962,1176,1727,1751]" gridcol="3" gridrow="31" pageId="5" pageNumber="142">66.4–70.8 (69.2±1.4)</td>
<td id="76B05DBDFFB116C0FB25F97CFA1C25A2" box="[1206,1420,1727,1751]" gridcol="4" gridrow="31" pageId="5" pageNumber="142">66.0–71.0 (67.6±2.1)</td>
</tr>
<tr id="356134C1FFB116C0FF35F924FA1C258A" box="[166,1420,1767,1791]" gridrow="32" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F924FD90258A" box="[166,512,1767,1791]" gridcol="0" gridrow="32" pageId="5" pageNumber="142">Pre-pelvic-fin length</th>
<td id="76B05DBDFFB116C0FDA4F924FD37258A" box="[567,679,1767,1791]" gridcol="1" gridrow="32" pageId="5" pageNumber="142">15.5</td>
<td id="76B05DBDFFB116C0FD5BF924FC0F258A" box="[712,927,1767,1791]" gridcol="2" gridrow="32" pageId="5" pageNumber="142">14.2–16.8 (15.4±0.7)</td>
<td id="76B05DBDFFB116C0FC51F924FB08258A" box="[962,1176,1767,1791]" gridcol="3" gridrow="32" pageId="5" pageNumber="142">13.8–18.2 (15.5±1.6)</td>
<td id="76B05DBDFFB116C0FB25F924FA1C258A" box="[1206,1420,1767,1791]" gridcol="4" gridrow="32" pageId="5" pageNumber="142">14.9–18.1 (15.9±1.3)</td>
</tr>
<tr id="356134C1FFB116C0FF35F8CCFA1C2452" box="[166,1420,1807,1831]" gridrow="33" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F8CCFD902452" box="[166,512,1807,1831]" gridcol="0" gridrow="33" pageId="5" pageNumber="142">Second dorsal-fin base length</th>
<td id="76B05DBDFFB116C0FDA4F8CCFD372452" box="[567,679,1807,1831]" gridcol="1" gridrow="33" pageId="5" pageNumber="142">12.4</td>
<td id="76B05DBDFFB116C0FD5BF8CCFC0F2452" box="[712,927,1807,1831]" gridcol="2" gridrow="33" pageId="5" pageNumber="142">10.2–13.2 (12.0±0.8)</td>
<td id="76B05DBDFFB116C0FC51F8CCFB082452" box="[962,1176,1807,1831]" gridcol="3" gridrow="33" pageId="5" pageNumber="142">9.2–11.7 (10.2±1.0)</td>
<td id="76B05DBDFFB116C0FB25F8CCFA1C2452" box="[1206,1420,1807,1831]" gridcol="4" gridrow="33" pageId="5" pageNumber="142">10.5–14.1 (12.5±1.3)</td>
</tr>
<tr id="356134C1FFB116C0FF35F8F4FA1C2425" box="[166,1420,1847,1872]" gridrow="34" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F8F4FD902425" box="[166,512,1847,1872]" gridcol="0" gridrow="34" pageId="5" pageNumber="142">Anal-fin base length</th>
<td id="76B05DBDFFB116C0FDA4F8F4FD372425" box="[567,679,1847,1872]" gridcol="1" gridrow="34" pageId="5" pageNumber="142">15.4</td>
<td id="76B05DBDFFB116C0FD5BF8F4FC0F2425" box="[712,927,1847,1872]" gridcol="2" gridrow="34" pageId="5" pageNumber="142">14.0–17.1 (15.5±0.9)</td>
<td id="76B05DBDFFB116C0FC51F8F4FB082425" box="[962,1176,1847,1872]" gridcol="3" gridrow="34" pageId="5" pageNumber="142">11.9–15.0 (13.7±1.1)</td>
<td id="76B05DBDFFB116C0FB25F8F4FA1C2425" box="[1206,1420,1847,1872]" gridcol="4" gridrow="34" pageId="5" pageNumber="142">14.8–17.1 (16.1±1.0)</td>
</tr>
<tr id="356134C1FFB116C0FF35F8A3FA1C240D" box="[166,1420,1888,1912]" gridrow="35" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F8A3FD90240D" box="[166,512,1888,1912]" gridcol="0" gridrow="35" pageId="5" pageNumber="142">Second dorsal-fin length</th>
<td id="76B05DBDFFB116C0FDA4F8A3FD37240D" box="[567,679,1888,1912]" gridcol="1" gridrow="35" pageId="5" pageNumber="142">4.1</td>
<td id="76B05DBDFFB116C0FD5BF8A3FC0F240D" box="[712,927,1888,1912]" gridcol="2" gridrow="35" pageId="5" pageNumber="142">3.2–5.8 (4.2±0.7)</td>
<td id="76B05DBDFFB116C0FC51F8A3FB08240D" box="[962,1176,1888,1912]" gridcol="3" gridrow="35" pageId="5" pageNumber="142">3.1–4.7 (4.0±0.7)</td>
<td id="76B05DBDFFB116C0FB25F8A3FA1C240D" box="[1206,1420,1888,1912]" gridcol="4" gridrow="35" pageId="5" pageNumber="142">3.8–5.6 (4.6±0.5)</td>
</tr>
<tr id="356134C1FFB116C0FF35F84BFA1C24D5" box="[166,1420,1928,1952]" gridrow="36" pageId="5" pageNumber="142">
<th id="76B05DBDFFB116C0FF35F84BFD9024D5" box="[166,512,1928,1952]" gridcol="0" gridrow="36" pageId="5" pageNumber="142">Anal-fin length</th>
<td id="76B05DBDFFB116C0FDA4F84BFD3724D5" box="[567,679,1928,1952]" gridcol="1" gridrow="36" pageId="5" pageNumber="142">3.5</td>
<td id="76B05DBDFFB116C0FD5BF84BFC0F24D5" box="[712,927,1928,1952]" gridcol="2" gridrow="36" pageId="5" pageNumber="142">3.0–4.7 (3.7±0.5)</td>
<td id="76B05DBDFFB116C0FC51F84BFB0824D5" box="[962,1176,1928,1952]" gridcol="3" gridrow="36" pageId="5" pageNumber="142">2.8–3.9 (3.4±0.5)</td>
<td id="76B05DBDFFB116C0FB25F84BFA1C24D5" box="[1206,1420,1928,1952]" gridcol="4" gridrow="36" pageId="5" pageNumber="142">2.7–5.5 (4.1±1.0)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEE3683FFB1E93AFBCBF872FA1F24BC" blockId="5.[1112,1423,1969,1993]" box="[1112,1423,1969,1993]" pageId="5" pageNumber="142">
<tableNote id="76B7370DFFB1E93AFBCBF872FA1F24BC" box="[1112,1423,1969,1993]" pageId="5" pageNumber="142" targetBox="[166,1420,226,1952]" targetPageId="5">......continued on the next page</tableNote>
</paragraph>
<caption id="DF2E660BFFB2E939FF35FF62FE0C23CC" box="[166,412,161,185]" isContinuationCaption="true" pageId="6" pageNumber="143" startId="6.[166,250,161,185]" targetBox="[166,1420,201,943]" targetIsTable="true" targetPageId="6">
<paragraph id="8BEE3683FFB2E939FF35FF62FE0C23CC" blockId="6.[166,412,161,185]" box="[166,412,161,185]" pageId="6" pageNumber="143">TABLE 1. (Continued)</paragraph>
</caption>
<paragraph id="8BEE3683FFB2E939FDA4FF09FA1C20DA" pageId="6" pageNumber="143">
<table id="F951C423FFB216C0FF35FF0AFA1C20DA" box="[166,1420,201,943]" colsContinueFrom="5.[166,1420,226,1952]" gridcols="5" gridrows="17" pageId="6" pageNumber="143">
<tr id="356134C1FFB216C0FF35FF0AFA1C2396" box="[166,1420,201,227]" gridrow="0" pageId="6" pageNumber="143" rowspan-0="1" rowspan-4="1">
<th id="76B05DBDFFB216C0FDA4FF0AFD052396" box="[567,661,201,227]" gridcol="1" gridrow="0" pageId="6" pageNumber="143">Holotype</th>
<th id="76B05DBDFFB216C0FD5BFF0AFC0F2396" box="[712,927,201,227]" gridcol="2" gridrow="0" pageId="6" pageNumber="143">Paratypes1</th>
<th id="76B05DBDFFB216C0FC51FF0AFB082396" box="[962,1176,201,227]" gridcol="3" gridrow="0" pageId="6" pageNumber="143">Non-types</th>
</tr>
<tr id="356134C1FFB216C0FF35FF31FA1C225B" box="[166,1420,242,302]" gridrow="1" pageId="6" pageNumber="143" rowspan-0="1" rowspan-1="1" rowspan-2="1">
<td id="76B05DBDFFB216C0FC51FF31FB08225B" box="[962,1176,242,302]" gridcol="3" gridrow="1" pageId="6" pageNumber="143">Amami-oshima island</td>
<td id="76B05DBDFFB216C0FB25FF31FA1C225B" box="[1206,1420,242,302]" gridcol="4" gridrow="1" pageId="6" pageNumber="143">Kochi Pref. and Osumi Pen.</td>
</tr>
<tr id="356134C1FFB216C0FF35FEFCFA1C2222" box="[166,1420,319,343]" gridrow="2" pageId="6" pageNumber="143" rowspan-0="1" rowspan-1="1">
<td id="76B05DBDFFB216C0FD5BFEFCFC0F2222" box="[712,927,319,343]" gridcol="2" gridrow="2" pageId="6" pageNumber="143">
<emphasis id="B925EA91FFB2E939FD5BFEFCFD452223" box="[712,725,319,342]" italics="true" pageId="6" pageNumber="143">n</emphasis>
= 18
</td>
<td id="76B05DBDFFB216C0FC51FEFCFB082222" box="[962,1176,319,343]" gridcol="3" gridrow="2" pageId="6" pageNumber="143">
<emphasis id="B925EA91FFB2E939FC51FEFCFC5F2223" box="[962,975,319,342]" italics="true" pageId="6" pageNumber="143">n</emphasis>
= 8
</td>
<td id="76B05DBDFFB216C0FB25FEFCFA1C2222" box="[1206,1420,319,343]" gridcol="4" gridrow="2" pageId="6" pageNumber="143">
<emphasis id="B925EA91FFB2E939FB25FEFCFB532223" box="[1206,1219,319,342]" italics="true" pageId="6" pageNumber="143">n</emphasis>
= 7
</td>
</tr>
<tr id="356134C1FFB216C0FF35FEA4FA1C22F5" box="[166,1420,359,384]" gridrow="3" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FEA4FEF922F5" box="[166,361,359,384]" gridcol="0" gridrow="3" pageId="6" pageNumber="143">Pectoral-fin length</th>
<td id="76B05DBDFFB216C0FDA4FEA4FD0522F5" box="[567,661,359,384]" gridcol="1" gridrow="3" pageId="6" pageNumber="143">6.4</td>
<td id="76B05DBDFFB216C0FD5BFEA4FC0F22F5" box="[712,927,359,384]" gridcol="2" gridrow="3" pageId="6" pageNumber="143">5.0–7.5 (6.3±0.8)</td>
<td id="76B05DBDFFB216C0FC51FEA4FB0822F5" box="[962,1176,359,384]" gridcol="3" gridrow="3" pageId="6" pageNumber="143">4.8–7.6 (6.2±0.9)</td>
<td id="76B05DBDFFB216C0FB25FEA4FA1C22F5" box="[1206,1420,359,384]" gridcol="4" gridrow="3" pageId="6" pageNumber="143">5.8–8.9 (7.3±1.2)</td>
</tr>
<tr id="356134C1FFB216C0FF35FE53FA1C22DD" box="[166,1420,400,424]" gridrow="4" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FE53FEF922DD" box="[166,361,400,424]" gridcol="0" gridrow="4" pageId="6" pageNumber="143">Pelvic-fin length</th>
<td id="76B05DBDFFB216C0FDA4FE53FD0522DD" box="[567,661,400,424]" gridcol="1" gridrow="4" pageId="6" pageNumber="143">2.7</td>
<td id="76B05DBDFFB216C0FD5BFE53FC0F22DD" box="[712,927,400,424]" gridcol="2" gridrow="4" pageId="6" pageNumber="143">2.1–3.3 (2.8±0.3)</td>
<td id="76B05DBDFFB216C0FC51FE53FB0822DD" box="[962,1176,400,424]" gridcol="3" gridrow="4" pageId="6" pageNumber="143">2.1–3.5 (2.7±0.4)</td>
<td id="76B05DBDFFB216C0FB25FE53FA1C22DD" box="[1206,1420,400,424]" gridcol="4" gridrow="4" pageId="6" pageNumber="143">2.7–4.4 (3.4±0.6)</td>
</tr>
<tr id="356134C1FFB216C0FF35FE7BFA1C22A5" box="[166,1420,440,464]" gridrow="5" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FE7BFEF922A5" box="[166,361,440,464]" gridcol="0" gridrow="5" pageId="6" pageNumber="143">Caudal-fin length</th>
<td id="76B05DBDFFB216C0FDA4FE7BFD0522A5" box="[567,661,440,464]" gridcol="1" gridrow="5" pageId="6" pageNumber="143">11.3</td>
<td id="76B05DBDFFB216C0FD5BFE7BFC0F22A5" box="[712,927,440,464]" gridcol="2" gridrow="5" pageId="6" pageNumber="143">9.2–13.3 (11.0±1.0)</td>
<td id="76B05DBDFFB216C0FC51FE7BFB0822A5" box="[962,1176,440,464]" gridcol="3" gridrow="5" pageId="6" pageNumber="143">11.1–15.1 (12.8±1.6)</td>
<td id="76B05DBDFFB216C0FB25FE7BFA1C22A5" box="[1206,1420,440,464]" gridcol="4" gridrow="5" pageId="6" pageNumber="143">9.4–13.8 (12.0±1.5)</td>
</tr>
<tr id="356134C1FFB216C0FF35FE23FA1C228D" box="[166,1420,480,504]" gridrow="6" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FE23FA1C228D" box="[166,1420,480,504]" colspan="5" colspanRight="4" gridcol="0" gridrow="6" pageId="6" pageNumber="143">Measurements (% HL)</th>
</tr>
<tr id="356134C1FFB216C0FF35FDCBFA1C2154" box="[166,1420,520,545]" gridrow="7" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FDCBFEF92154" box="[166,361,520,545]" gridcol="0" gridrow="7" pageId="6" pageNumber="143">Head depth</th>
<td id="76B05DBDFFB216C0FDA4FDCBFD052154" box="[567,661,520,545]" gridcol="1" gridrow="7" pageId="6" pageNumber="143">37.0</td>
<td id="76B05DBDFFB216C0FD5BFDCBFC0F2154" box="[712,927,520,545]" gridcol="2" gridrow="7" pageId="6" pageNumber="143">36.0–46.6 (40.2±3.5)</td>
<td id="76B05DBDFFB216C0FC51FDCBFB082154" box="[962,1176,520,545]" gridcol="3" gridrow="7" pageId="6" pageNumber="143">32.5–40.0 (37.1±2.6)</td>
<td id="76B05DBDFFB216C0FB25FDCBFA1C2154" box="[1206,1420,520,545]" gridcol="4" gridrow="7" pageId="6" pageNumber="143">35.4–46.9 (41.5±3.6)</td>
</tr>
<tr id="356134C1FFB216C0FF35FDF2FA1C213C" box="[166,1420,561,585]" gridrow="8" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FDF2FEF9213C" box="[166,361,561,585]" gridcol="0" gridrow="8" pageId="6" pageNumber="143">Head width</th>
<td id="76B05DBDFFB216C0FDA4FDF2FD05213C" box="[567,661,561,585]" gridcol="1" gridrow="8" pageId="6" pageNumber="143">53.9</td>
<td id="76B05DBDFFB216C0FD5BFDF2FC0F213C" box="[712,927,561,585]" gridcol="2" gridrow="8" pageId="6" pageNumber="143">45.8–62.4 (53.3±4.4)</td>
<td id="76B05DBDFFB216C0FC51FDF2FB08213C" box="[962,1176,561,585]" gridcol="3" gridrow="8" pageId="6" pageNumber="143">40.3–45.7 (43.3±2.1)</td>
<td id="76B05DBDFFB216C0FB25FDF2FA1C213C" box="[1206,1420,561,585]" gridcol="4" gridrow="8" pageId="6" pageNumber="143">51.3–57.8 (54.8±2.6)</td>
</tr>
<tr id="356134C1FFB216C0FF35FD9AFA1C2104" box="[166,1420,601,625]" gridrow="9" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FD9AFEF92104" box="[166,361,601,625]" gridcol="0" gridrow="9" pageId="6" pageNumber="143">Snout length</th>
<td id="76B05DBDFFB216C0FDA4FD9AFD052104" box="[567,661,601,625]" gridcol="1" gridrow="9" pageId="6" pageNumber="143">23.6</td>
<td id="76B05DBDFFB216C0FD5BFD9AFC0F2104" box="[712,927,601,625]" gridcol="2" gridrow="9" pageId="6" pageNumber="143">22.2–26.3 (24.5±1.3)</td>
<td id="76B05DBDFFB216C0FC51FD9AFB082104" box="[962,1176,601,625]" gridcol="3" gridrow="9" pageId="6" pageNumber="143">23.5–27.5 (25.0±1.6)</td>
<td id="76B05DBDFFB216C0FB25FD9AFA1C2104" box="[1206,1420,601,625]" gridcol="4" gridrow="9" pageId="6" pageNumber="143">21.1–26.7 (24.0±2.0)</td>
</tr>
<tr id="356134C1FFB216C0FF35FD42FA1C21EC" box="[166,1420,641,665]" gridrow="10" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FD42FEF921EC" box="[166,361,641,665]" gridcol="0" gridrow="10" pageId="6" pageNumber="143">Upper-jaw length</th>
<td id="76B05DBDFFB216C0FDA4FD42FD0521EC" box="[567,661,641,665]" gridcol="1" gridrow="10" pageId="6" pageNumber="143">31.8</td>
<td id="76B05DBDFFB216C0FD5BFD42FC0F21EC" box="[712,927,641,665]" gridcol="2" gridrow="10" pageId="6" pageNumber="143">29.0–37.6 (32.9±2.2)</td>
<td id="76B05DBDFFB216C0FC51FD42FB0821EC" box="[962,1176,641,665]" gridcol="3" gridrow="10" pageId="6" pageNumber="143">29.2–38.4 (32.4±3.5)</td>
<td id="76B05DBDFFB216C0FB25FD42FA1C21EC" box="[1206,1420,641,665]" gridcol="4" gridrow="10" pageId="6" pageNumber="143">31.2–37.1 (32.7±2.0)</td>
</tr>
<tr id="356134C1FFB216C0FF35FD6AFA1C21B7" box="[166,1420,681,706]" gridrow="11" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FD6AFEF921B7" box="[166,361,681,706]" gridcol="0" gridrow="11" pageId="6" pageNumber="143">Eye diameter</th>
<td id="76B05DBDFFB216C0FDA4FD6AFD0521B7" box="[567,661,681,706]" gridcol="1" gridrow="11" pageId="6" pageNumber="143">6.7</td>
<td id="76B05DBDFFB216C0FD5BFD6AFC0F21B7" box="[712,927,681,706]" gridcol="2" gridrow="11" pageId="6" pageNumber="143">6.5–9.3 (7.7±0.7)</td>
<td id="76B05DBDFFB216C0FC51FD6AFB0821B7" box="[962,1176,681,706]" gridcol="3" gridrow="11" pageId="6" pageNumber="143">5.1–9.5 (7.1±1.6)</td>
<td id="76B05DBDFFB216C0FB25FD6AFA1C21B7" box="[1206,1420,681,706]" gridcol="4" gridrow="11" pageId="6" pageNumber="143">6.3–8.4 (7.6±0.8)</td>
</tr>
<tr id="356134C1FFB216C0FF35FD11FA1C219F" box="[166,1420,722,746]" gridrow="12" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FD11FEF9219F" box="[166,361,722,746]" gridcol="0" gridrow="12" pageId="6" pageNumber="143">Interorbital width</th>
<td id="76B05DBDFFB216C0FDA4FD11FD05219F" box="[567,661,722,746]" gridcol="1" gridrow="12" pageId="6" pageNumber="143">17.8</td>
<td id="76B05DBDFFB216C0FD5BFD11FC0F219F" box="[712,927,722,746]" gridcol="2" gridrow="12" pageId="6" pageNumber="143">13.0–18.2 (15.4±1.6)</td>
<td id="76B05DBDFFB216C0FC51FD11FB08219F" box="[962,1176,722,746]" gridcol="3" gridrow="12" pageId="6" pageNumber="143">11.3–15.7 (13.6±1.4)</td>
<td id="76B05DBDFFB216C0FB25FD11FA1C219F" box="[1206,1420,722,746]" gridcol="4" gridrow="12" pageId="6" pageNumber="143">14.0–18.5 (16.5±1.9)</td>
</tr>
<tr id="356134C1FFB216C0FF35FD39FA1C2067" box="[166,1420,762,786]" gridrow="13" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FD39FA1C2067" box="[166,1420,762,786]" colspan="5" colspanRight="4" gridcol="0" gridrow="13" pageId="6" pageNumber="143">Measurements (% AAA distance)</th>
</tr>
<tr id="356134C1FFB216C0FF35FCE1FA1C204F" box="[166,1420,802,826]" gridrow="14" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FCE1FEF9204F" box="[166,361,802,826]" gridcol="0" gridrow="14" pageId="6" pageNumber="143">Snout length</th>
<td id="76B05DBDFFB216C0FDA4FCE1FD05204F" box="[567,661,802,826]" gridcol="1" gridrow="14" pageId="6" pageNumber="143">27.2</td>
<td id="76B05DBDFFB216C0FD5BFCE1FC0F204F" box="[712,927,802,826]" gridcol="2" gridrow="14" pageId="6" pageNumber="143">22.8–30.3 (26.3±1.7)</td>
<td id="76B05DBDFFB216C0FC51FCE1FB08204F" box="[962,1176,802,826]" gridcol="3" gridrow="14" pageId="6" pageNumber="143">19.2–34.7 (24.8±5.3)</td>
<td id="76B05DBDFFB216C0FB25FCE1FA1C204F" box="[1206,1420,802,826]" gridcol="4" gridrow="14" pageId="6" pageNumber="143">24.9–30.8 (28.2±2.4)</td>
</tr>
<tr id="356134C1FFB216C0FF35FC89FA1C20F3" box="[166,1420,842,902]" gridrow="15" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FC89FA1C20F3" box="[166,1420,842,902]" colspan="5" colspanRight="4" gridcol="0" gridrow="15" pageId="6" pageNumber="143">Measurements (% pre-anal-fin length)</th>
</tr>
<tr id="356134C1FFB216C0FF35FC54FA1C20DA" box="[166,1420,919,943]" gridrow="16" pageId="6" pageNumber="143">
<th id="76B05DBDFFB216C0FF35FC54FEF920DA" box="[166,361,919,943]" gridcol="0" gridrow="16" pageId="6" pageNumber="143">Pre-anus length</th>
<td id="76B05DBDFFB216C0FDA4FC54FD0520DA" box="[567,661,919,943]" gridcol="1" gridrow="16" pageId="6" pageNumber="143">78.3</td>
<td id="76B05DBDFFB216C0FD5BFC54FC0F20DA" box="[712,927,919,943]" gridcol="2" gridrow="16" pageId="6" pageNumber="143">77.6–81.1 (79.5±1.0)</td>
<td id="76B05DBDFFB216C0FC51FC54FB0820DA" box="[962,1176,919,943]" gridcol="3" gridrow="16" pageId="6" pageNumber="143">75.0–85.5 (80.2±3.8)</td>
<td id="76B05DBDFFB216C0FB25FC54FA1C20DA" box="[1206,1420,919,943]" gridcol="4" gridrow="16" pageId="6" pageNumber="143">77.3–82.0 (79.5±1.9)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEE3683FFB2E939FF04FC78FB9920A1" blockId="6.[151,1033,955,981]" box="[151,1033,955,981]" pageId="6" pageNumber="143">
<tableNote id="76B7370DFFB2E939FF04FC78FB9920A1" box="[151,1033,955,981]" pageId="6" pageNumber="143" targetBox="[166,1420,201,943]" targetPageId="6">
<superScript id="7C249BCBFFB2E939FF04FC78FF0F20BC" attach="right" box="[151,159,955,969]" fontSize="6" pageId="6" pageNumber="143">1</superScript>
Excludes two paratypes preserved in 99.5% ethanol (KAUM–I. 158056, 158057).
</tableNote>
</paragraph>
<caption id="DF2E660BFFB2E939FF04F87BFB772481" ID-DOI="http://doi.org/10.5281/zenodo.6556886" ID-Zenodo-Dep="6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="6" pageNumber="143" startId="6.[151,250,1976,2000]" targetBox="[178,1410,1041,1953]" targetPageId="6">
<paragraph id="8BEE3683FFB2E939FF04F87BFB772481" blockId="6.[151,1437,1976,2037]" pageId="6" pageNumber="143">
<emphasis id="B925EA91FFB2E939FF04F87BFE8524A5" bold="true" box="[151,277,1976,2000]" pageId="6" pageNumber="143">FIGURE 3.</emphasis>
Live individuals of 
<taxonomicName id="4C514D00FFB2E939FE79F87BFD6F24A5" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[490,767,1976,2000]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="143" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB2E939FE79F87BFD6F24A5" box="[490,767,1976,2000]" italics="true" pageId="6" pageNumber="143">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB2E939FC94F87AFCD224A4" bold="true" box="[775,834,1977,2001]" pageId="6" pageNumber="143">
<taxonomicNameLabel id="A21657EAFFB2E939FC94F87AFCD224A4" box="[775,834,1977,2001]" pageId="6" pageNumber="143" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124).
</paragraph>
</caption>
<paragraph id="8BEE3683FFB3E938FF54FF54FC07217E" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
<emphasis id="B925EA91FFB3E938FF54FF54FDE523C4" box="[199,629,151,177]" italics="true" pageId="7" pageNumber="144">Coloration (live and fresh specimens.)</emphasis>
Based on 
<figureCitation id="136A2A06FFB3E938FD7EFF54FCC623C4" box="[749,854,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[151,1436,179,1785]" captionTargetId="figure-17@3.[151,1436,179,1785]" captionTargetPageId="3" captionText="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="7" pageNumber="144">Figures 1</figureCitation>
, 
<figureCitation id="136A2A06FFB3E938FCF7FF54FCE223C4" box="[868,882,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[178,1410,1041,1953]" captionTargetId="figure-197@6.[178,1410,1035,1953]" captionTargetPageId="6" captionText="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." figureDoi="http://doi.org/10.5281/zenodo.6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="7" pageNumber="144">3</figureCitation>
and 
<bibRefCitation id="EFC04B72FFB3E938FC39FF54FB6023C4" author="Koreeda, R. &amp; Motomura, H." box="[938,1264,151,177]" pageId="7" pageNumber="144" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura (2021</bibRefCitation>
: fig. 8A–C, E): Body yellowish-green to brown dorsally, whitish ventrally (yellow to orange ventrally in gravid females). A single poorly defined black longitudinal line mid-laterally on body from behind pectoral fin to caudal-fin base, embedded internally and visible through semi-transparent muscle issue. Small black spots forming a single longitudinal row mid-laterally on body surface from behind pectoral fin to caudal-fin base, more distinct posteriorly, discontinuous anteriorly (sometimes indistinct in live specimens). Black to brown spots scattered on dorsal body surface and caudal peduncle. Dorsal-, anal-, pectoral-, and pelvic-fin rays and membranes semitransparent white to pale yellow. Caudal fin semitransparent white, with yellow-margined black rounded blotch basally; posterior margin of black blotch reaching posteriorly to between one-fourth and half of caudal-fin length. In juveniles (12.1–15.0 mm SL), body pale yellow to beige with few pigments; black stripe and dotted line indistinct; caudal-fin blotch reaching posteriorly to less than one-fifth of fin length; yellow margin absent.
</paragraph>
<paragraph id="8BEE3683FFB3E938FF54FDD7FA1F2106" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
<emphasis id="B925EA91FFB3E938FF54FDD7FDDF215B" box="[199,591,532,558]" italics="true" pageId="7" pageNumber="144">Coloration of preserved specimens</emphasis>
. Based on 
<figureCitation id="136A2A06FFB3E938FD5EFDD7FCBB215B" box="[717,811,532,558]" captionStart="FIGURE 4" captionStartId="8.[151,250,1281,1305]" captionTargetBox="[180,1408,181,1257]" captionTargetId="figure-30@8.[180,1408,181,1257]" captionTargetPageId="8" captionText="FIGURE 4. Preserved specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.6556890" httpUri="https://zenodo.org/record/6556890/files/figure.png" pageId="7" pageNumber="144">Figure 4</figureCitation>
and 
<bibRefCitation id="EFC04B72FFB3E938FCF0FDD7FB39215B" author="Koreeda, R. &amp; Motomura, H." box="[867,1193,532,559]" pageId="7" pageNumber="144" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura (2021</bibRefCitation>
: fig. 8D): Body whitish, with a single distinct longitudinal line on mid-lateral surface, and small brown blotches scattered dorsally and on caudal peduncle. Caudal fin with black to brown blotch basally. Single embedded longitudinal line not visible.
</paragraph>
</subSubSection>
<subSubSection id="C34B6508FFB3E938FF54FDBFFCF421CC" pageId="7" pageNumber="144" type="etymology">
<paragraph id="8BEE3683FFB3E938FF54FDBFFCF421CC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
<emphasis id="B925EA91FFB3E938FF54FDBFFEDE21E3" bold="true" box="[199,334,636,662]" pageId="7" pageNumber="144">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C514D00FFB3E938FDA2FDBFFD4021E3" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[561,720,636,662]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FDA2FDBFFD4021E3" box="[561,720,636,662]" italics="true" pageId="7" pageNumber="144">punctilineatus</emphasis>
</taxonomicName>
is derived from Latin meaning “dotted line,” and refers to the single longitudinal dotted line on the lateral surface of the body.
</paragraph>
</subSubSection>
<subSubSection id="C34B6508FFB3E932FF54FD02FEBF2234" lastPageId="13" lastPageNumber="150" pageId="7" pageNumber="144" type="distribution">
<paragraph id="8BEE3683FFB3E938FF54FD02FEC62012" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
<emphasis id="B925EA91FFB3E938FF54FD02FDF021AE" bold="true" box="[199,608,705,731]" pageId="7" pageNumber="144">Distribution and ecological notes.</emphasis>
Currently known from southern 
<collectingCountry id="F3467613FFB3E938FC71FD01FBB521A9" box="[994,1061,706,732]" name="Japan" pageId="7" pageNumber="144">Japan</collectingCountry>
, specimens having been collected from 
<collectingRegion id="4995F861FFB3E938FF6EFD27FEE9218B" box="[253,377,740,766]" country="Japan" name="Tokushima" pageId="7" pageNumber="144">Tokushima</collectingRegion>
(
<bibRefCitation id="EFC04B72FFB3E938FE18FD27FD1B218B" author="Ito, T. &amp; Okumura, D." box="[395,651,740,766]" pageId="7" pageNumber="144" pagination="1 - 7" refId="ref7959" refString="Ito, T. &amp; Okumura, D. (2021) First records of Luciogobius parvulus and Luciogobius sp. 11 sensu Shibukawa et al., 2019 (Teleostei: Gobiidae) from Tokushima Prefecture, Japan. Ichthy, Natural History of Fishes of Japan, 12, 1 - 7. [in Japanese] https: // doi. org / 10.34583 / ichthy. 12.0 _ 1" type="journal article" year="2021">Ito &amp; Okumura, 2021</bibRefCitation>
) and 
<collectingRegion id="4995F861FFB3E938FD42FD27FC85218B" box="[721,789,740,766]" country="Japan" name="Kochi" pageId="7" pageNumber="144">Kochi</collectingRegion>
prefectures (both southern Shikoku), 
<collectingRegion id="4995F861FFB3E938FB42FD27FAC1218B" box="[1233,1361,740,766]" country="Japan" name="Kagoshima" pageId="7" pageNumber="144">Kagoshima</collectingRegion>
mainland (southern Kyushu), Shimokoshiki-shima island (Koshiki Islands, off 
<collectingRegion id="4995F861FFB3E938FC73FCC4FBF02054" box="[992,1120,775,801]" country="Japan" name="Kagoshima" pageId="7" pageNumber="144">Kagoshima</collectingRegion>
mainland, East 
<collectingCountry id="F3467613FFB3E938FA8BFCC4FACB2054" box="[1304,1371,775,801]" name="China" pageId="7" pageNumber="144">China</collectingCountry>
Sea), Tanegashima and Yakushima islands (Osumi Islands, south of Kyushu), and Amami-oshima island (Amami Islands, Ryukyu Islands).
</paragraph>
<paragraph id="8BEE3683FFB3E938FF54FCACFEF12743" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
Most specimens were collected from gravel in the vicinity of freshwater springs near coastal river mouths (
<figureCitation id="136A2A06FFB3E938FAFCFCACFF3720D9" captionStart="FIGURE 7" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[151,1436,889,1844]" captionTargetId="figure-48@9.[151,1436,889,1844]" captionTargetPageId="9" captionText="FIGURE 7. Habitats of Luciogobius punctilineatus n. sp. near (A) the Shirinashi River mouth, western coast of Satsuma Peninsula, and (B) the Shimadomari River mouth, southern coast of Osumi Peninsula. Black arrows indicate collection points, near fresh water springs; blue arrows indicate river mouths." figureDoi="http://doi.org/10.5281/zenodo.6556896" httpUri="https://zenodo.org/record/6556896/files/figure.png" pageId="7" pageNumber="144">Fig. 7</figureCitation>
), but not from the rivers or river mouths themselves. On Amami-oshima island, all specimens were collected from the gravel layer (with a freshwater spring 
<quantity id="4CA99B66FFB3E938FDFAFC77FD3E20BA" box="[617,686,948,975]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="144" unit="cm" value="10.0">10 cm</quantity>
below the surface) on a sandy beach 
<quantity id="4CA99B66FFB3E938FBDCFC77FB0520BA" box="[1103,1173,948,975]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="144" unit="m" value="100.0">100 m</quantity>
west of the Heda River mouth. 
<taxonomicName id="4C514D00FFB3E938FF7CFC14FDB22084" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[239,546,983,1009]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FF7CFC14FDB22084" box="[239,546,983,1009]" italics="true" pageId="7" pageNumber="144">Luciogobius punctilineatus</emphasis>
</taxonomicName>
was abundant in the above-mentioned habitats, often occurring together with 
<taxonomicName id="4C514D00FFB3E938FF04FC3AFECD2766" box="[151,349,1017,1044]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undefined-4">
<emphasis id="B925EA91FFB3E938FF04FC3AFEB12766" box="[151,289,1017,1043]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp. 4
</taxonomicName>
<emphasis id="B925EA91FFB3E938FEF7FC39FE322766" box="[356,418,1018,1043]" italics="true" pageId="7" pageNumber="144">sensu</emphasis>
<bibRefCitation id="EFC04B72FFB3E938FE3AFC3AFDED2766" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." box="[425,637,1017,1044]" pageId="7" pageNumber="144" pagination="162 - 174" refId="ref8331" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55 (2), 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda 
<emphasis id="B925EA91FFB3E938FE6EFC39FDBE2766" box="[509,558,1017,1043]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
., 2008
</bibRefCitation>
, although suitable habitats for 
<taxonomicName id="4C514D00FFB3E938FC46FC39FB012766" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[981,1169,1017,1043]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FC46FC39FB012766" box="[981,1169,1017,1043]" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
were more limited than those of the latter.
</paragraph>
<paragraph id="8BEE3683FFB3E938FF54FBFCFD5627D4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
Juveniles of 
<taxonomicName id="4C514D00FFB3E938FEC0FBFCFD9F272C" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[339,527,1087,1113]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FEC0FBFCFD9F272C" box="[339,527,1087,1113]" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
(12.1–15.0 mm SL; 
<figureCitation id="136A2A06FFB3E938FD67FBFCFCD7272C" box="[756,839,1087,1113]" captionStart="FIGURE 1" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[151,1436,179,1785]" captionTargetId="figure-17@3.[151,1436,179,1785]" captionTargetPageId="3" captionText="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="7" pageNumber="144">Fig. 1C</figureCitation>
) were collected in July on Amami-oshima island, and fully matured females (
<figureCitation id="136A2A06FFB3E938FE0FFBA0FE642708" box="[412,500,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[178,1410,1041,1953]" captionTargetId="figure-197@6.[178,1410,1035,1953]" captionTargetPageId="6" captionText="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." figureDoi="http://doi.org/10.5281/zenodo.6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="7" pageNumber="144">Fig. 3A</figureCitation>
) collected from April to June on 
<collectingRegion id="4995F861FFB3E938FCF4FBA0FC772708" box="[871,999,1123,1149]" country="Japan" name="Kagoshima" pageId="7" pageNumber="144">Kagoshima</collectingRegion>
mainland. The reproductive season for this species is likely to be spring to early summer.
</paragraph>
<paragraph id="8BEE3683FFB3E938FF54FB69FCC526B4" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="144">
<emphasis id="B925EA91FFB3E938FF54FB69FEFD27B1" bold="true" box="[199,365,1194,1220]" pageId="7" pageNumber="144">Comparisons.</emphasis>
<taxonomicName id="4C514D00FFB3E938FEE6FB68FD3727B0" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[373,679,1195,1221]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FEE6FB68FD3727B0" box="[373,679,1195,1221]" italics="true" pageId="7" pageNumber="144">Luciogobius punctilineatus</emphasis>
</taxonomicName>
belongs to the 
<taxonomicName id="4C514D00FFB3E938FCCBFB68FBC827B0" authorityName="Regan" authorityYear="1905" box="[856,1112,1195,1221]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B925EA91FFB3E938FCCBFB68FBC827B0" box="[856,1112,1195,1221]" italics="true" pageId="7" pageNumber="144">Luciogobius elongatus</emphasis>
</taxonomicName>
complex, defined by 
<bibRefCitation id="EFC04B72FFB3E938FAC1FB68FEF7279C" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FF48FB0CFE9D279C" box="[219,269,1231,1257]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
. (2019)
</bibRefCitation>
as having no free pectoral-fin rays, total vertebrae 39–42, the AAA distance greater than body depth at the anus, first to third anal-fin rays unbranched and spine-like, and anteriormost pterygiophore of the anal fin inserted behind the fourth haemal spine. 
<taxonomicName id="4C514D00FFB3E938FD16FAD4FC262644" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[645,950,1303,1329]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FD16FAD4FC262644" box="[645,950,1303,1329]" italics="true" pageId="7" pageNumber="144">Luciogobius punctilineatus</emphasis>
</taxonomicName>
is easily distinguished from all other members of the complex (except 
<taxonomicName id="4C514D00FFB3E938FE5CFAF8FDEF2620" box="[463,639,1339,1365]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FE5CFAF8FDC92620" box="[463,601,1339,1365]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11 
<emphasis id="B925EA91FFB3E938FD36FAF8FD732621" box="[677,739,1339,1364]" italics="true" pageId="7" pageNumber="144">sensu</emphasis>
<bibRefCitation id="EFC04B72FFB3E938FD7AFAF8FC7B2620" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[745,1003,1339,1365]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FCFFFAF8FC0B2620" box="[876,923,1339,1365]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
., 2019
</bibRefCitation>
), by having 10–12 total dorsal-fin rays, 12–14 total anal-fin rays, 8–12 pectoral-fin rays with moderately incised membranes, pelvic fins united, forming a ventral disc, 39–42 total vertebrae, and a single poorly defined longitudinal black line (embedded in muscle tissue) and dotted line (on body surface) mid-laterally along the body.
</paragraph>
<paragraph id="8BEE3683FFB3E935FF54FA08FCA125C0" blockId="7.[151,1437,151,2013]" lastBlockId="10.[151,1437,1511,2042]" lastPageId="10" lastPageNumber="147" pageId="7" pageNumber="144">
<taxonomicName id="4C514D00FFB3E938FF54FA08FEE92690" box="[199,377,1483,1509]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FF54FA08FEC12690" box="[199,337,1483,1509]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (
<figureCitation id="136A2A06FFB3E938FE39FA08FE7D2690" box="[426,493,1483,1509]" captionStart="FIGURE 8" captionStartId="11.[151,250,1149,1173]" captionTargetBox="[151,1437,381,1125]" captionTargetId="figure-111@11.[151,1437,381,1125]" captionTargetPageId="11" captionText="FIGURE 8. Fresh (upper and middle) and preserved (lower) specimens of Luciogobius sp. 11 (KAUM–I. 159027 25.9 mm SL) from Kochi Prefecture." figureDoi="http://doi.org/10.5281/zenodo.6556903" httpUri="https://zenodo.org/record/6556903/files/figure.png" pageId="7" pageNumber="144">Fig. 8</figureCitation>
) is similar to 
<taxonomicName id="4C514D00FFB3E938FD1AFA08FCD52690" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[649,837,1483,1509]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FD1AFA08FCD52690" box="[649,837,1483,1509]" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
in sharing the above-mentioned characters (
<tableCitation id="C6D30338FFB3E938FAA6FA08FA072690" box="[1333,1431,1483,1509]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[151,1411,263,1001]" captionTargetPageId="10" captionText="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FF54FC582382">Tables 2</tableCitation>
, 
<tableCitation id="C6D30338FFB3E938FF04FA2CFF36257C" box="[151,166,1519,1545]" captionStart="TABLE 3" captionStartId="10.[151,239,1050,1074]" captionTargetBox="[166,1232,1090,1427]" captionTargetPageId="10" captionText="TABLE 3. Comparisons of selected morphometrics between Luciogobius punctilineatus n. sp. and Luciogobius sp. 11." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FBD9FA1D2747">3</tableCitation>
), the two species co-occurring at least in 
<collectingRegion id="4995F861FFB3E938FDE4FA2CFCA5257C" box="[631,821,1519,1545]" country="Japan" name="Kochi" pageId="7" pageNumber="144">Kochi Prefecture</collectingRegion>
(
<bibRefCitation id="EFC04B72FFB3E938FCD7FA2CFBDA257C" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[836,1098,1519,1545]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FC5AFA2CFC6A257C" box="[969,1018,1519,1545]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
., 2019
</bibRefCitation>
; 
<bibRefCitation id="EFC04B72FFB3E938FBC5FA2CFA07257C" author="Koreeda, R. &amp; Motomura, H." box="[1110,1431,1519,1545]" pageId="7" pageNumber="144" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
; this study). However, the latter has a greater AAA distance [11.4–16.9% (mean 13.3%) of SL vs. 9.1–10.4% (9.6%) in 
<taxonomicName id="4C514D00FFB3E938FF27F9F4FEF62524" box="[180,358,1591,1617]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FF27F9F4FEAE2524" box="[180,318,1591,1617]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11; 
<figureCitation id="136A2A06FFB3E938FE05F9F4FE7D2524" box="[406,493,1591,1617]" captionStart="FIGURE 9" captionStartId="12.[151,250,1813,1837]" captionTargetBox="[241,1346,191,1781]" captionTargetId="figure-30@12.[214,1374,181,1789]" captionTargetPageId="12" captionText="FIGURE 9. Relationships of (A) snout length, (B) AAA distance (measured between posterior margin of anus and anal-fin origin), (C) PAL (pre-anus length), and (D) pre-anal-fin length as % of standard length, and (E) snout length as % of AAA, and (F) PAL as % of pre-anal-fin length vs. standard length (mm) in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Closed red star, holotype of L. punctilineatus n. sp.; open red stars, paratypes of L. punctilineatus n. sp. from Shimokoshiki-shima island, Tanega-shima island; yellow stars, L. punctilineatus n. sp. from Amami-oshima island; green stars; L. punctilineatus n. sp. from Kochi Prefecture and Osumi Peninsula; blue triangles, Luciogobius sp. 11." figureDoi="http://doi.org/10.5281/zenodo.6556909" httpUri="https://zenodo.org/record/6556909/files/figure.png" pageId="7" pageNumber="144">Fig. 9B</figureCitation>
; 
<tableCitation id="C6D30338FFB3E938FE6AF9F4FDDE2524" box="[505,590,1590,1617]" captionStart="TABLE 3" captionStartId="10.[151,239,1050,1074]" captionTargetBox="[166,1232,1090,1427]" captionTargetPageId="10" captionText="TABLE 3. Comparisons of selected morphometrics between Luciogobius punctilineatus n. sp. and Luciogobius sp. 11." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FBD9FA1D2747">Table 3</tableCitation>
], and shorter snout [19.2–34.7% (26.4%) of AAA vs. 36.7–44.8% (40.9%); 
<figureCitation id="136A2A06FFB3E938FF04F998FF7C2500" box="[151,236,1627,1653]" captionStart="FIGURE 9" captionStartId="12.[151,250,1813,1837]" captionTargetBox="[241,1346,191,1781]" captionTargetId="figure-30@12.[214,1374,181,1789]" captionTargetPageId="12" captionText="FIGURE 9. Relationships of (A) snout length, (B) AAA distance (measured between posterior margin of anus and anal-fin origin), (C) PAL (pre-anus length), and (D) pre-anal-fin length as % of standard length, and (E) snout length as % of AAA, and (F) PAL as % of pre-anal-fin length vs. standard length (mm) in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Closed red star, holotype of L. punctilineatus n. sp.; open red stars, paratypes of L. punctilineatus n. sp. from Shimokoshiki-shima island, Tanega-shima island; yellow stars, L. punctilineatus n. sp. from Amami-oshima island; green stars; L. punctilineatus n. sp. from Kochi Prefecture and Osumi Peninsula; blue triangles, Luciogobius sp. 11." figureDoi="http://doi.org/10.5281/zenodo.6556909" httpUri="https://zenodo.org/record/6556909/files/figure.png" pageId="7" pageNumber="144">Fig. 9E</figureCitation>
; 
<tableCitation id="C6D30338FFB3E938FF65F998FED92500" box="[246,329,1626,1653]" captionStart="TABLE 3" captionStartId="10.[151,239,1050,1074]" captionTargetBox="[166,1232,1090,1427]" captionTargetPageId="10" captionText="TABLE 3. Comparisons of selected morphometrics between Luciogobius punctilineatus n. sp. and Luciogobius sp. 11." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FBD9FA1D2747">Table 3</tableCitation>
] and pre-anus lengths [75.0–82.0% (79.3%) of preanal-fin length (except in juveniles less than 15.0 mm SL) vs. 84.4–88.8% (86.1%); 
<figureCitation id="136A2A06FFB3E938FD8CF9BCFDE425EC" box="[543,628,1663,1689]" captionStart="FIGURE 9" captionStartId="12.[151,250,1813,1837]" captionTargetBox="[241,1346,191,1781]" captionTargetId="figure-30@12.[214,1374,181,1789]" captionTargetPageId="12" captionText="FIGURE 9. Relationships of (A) snout length, (B) AAA distance (measured between posterior margin of anus and anal-fin origin), (C) PAL (pre-anus length), and (D) pre-anal-fin length as % of standard length, and (E) snout length as % of AAA, and (F) PAL as % of pre-anal-fin length vs. standard length (mm) in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Closed red star, holotype of L. punctilineatus n. sp.; open red stars, paratypes of L. punctilineatus n. sp. from Shimokoshiki-shima island, Tanega-shima island; yellow stars, L. punctilineatus n. sp. from Amami-oshima island; green stars; L. punctilineatus n. sp. from Kochi Prefecture and Osumi Peninsula; blue triangles, Luciogobius sp. 11." figureDoi="http://doi.org/10.5281/zenodo.6556909" httpUri="https://zenodo.org/record/6556909/files/figure.png" pageId="7" pageNumber="144">Fig. 9F</figureCitation>
; 
<tableCitation id="C6D30338FFB3E938FD13F9BCFD4725EC" box="[640,727,1662,1689]" captionStart="TABLE 3" captionStartId="10.[151,239,1050,1074]" captionTargetBox="[166,1232,1090,1427]" captionTargetPageId="10" captionText="TABLE 3. Comparisons of selected morphometrics between Luciogobius punctilineatus n. sp. and Luciogobius sp. 11." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FBD9FA1D2747">Table 3</tableCitation>
]. The membranes between adjacent pectoral-fin rays (except 2 uppermost and 2 lowermost rays) are incised posteriorly for one-sixth to one-fourth of each ray pair in 
<taxonomicName id="4C514D00FFB3E938FABDF960FF602594" authorityName="Koreeda &amp; Motomura" authorityYear="2022" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FABDF960FF602594" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
, to one-third to half in 
<taxonomicName id="4C514D00FFB3E938FE68F904FD3D2594" box="[507,685,1735,1761]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FE68F904FD152594" box="[507,645,1735,1761]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (
<bibRefCitation id="EFC04B72FFB3E938FD73F904FC792594" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[736,1001,1735,1761]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FCF5F904FC0F2594" box="[870,927,1735,1761]" italics="true" pageId="7" pageNumber="144">et al.</emphasis>
, 2019
</bibRefCitation>
; 
<bibRefCitation id="EFC04B72FFB3E938FC65F904FAAB2594" author="Koreeda, R. &amp; Motomura, H." box="[1014,1339,1735,1761]" pageId="7" pageNumber="144" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
; 
<figureCitation id="136A2A06FFB3E938FADBF904FA052594" box="[1352,1429,1735,1761]" captionStart="FIGURE 1" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[151,1436,179,1785]" captionTargetId="figure-17@3.[151,1436,179,1785]" captionTargetPageId="3" captionText="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="7" pageNumber="144">Figs. 1</figureCitation>
, 
<figureCitation id="136A2A06FFB3E938FF04F928FF372470" box="[151,167,1771,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1727,1751]" captionTargetBox="[182,1433,196,1691]" captionTargetId="figure-30@4.[154,1433,181,1704]" captionTargetPageId="4" captionText="FIGURE 2. Head of the holotype of Luciogobius punctilineatus n. sp., showing the cephalic sensory system. Black dots and lowercase letters indicate papillae and name of each row, respectively. Arrows, AN, and PN indicate point of gill opening, anterior nostril, and posterior nostril, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556884" httpUri="https://zenodo.org/record/6556884/files/figure.png" pageId="7" pageNumber="144">2</figureCitation>
, 
<figureCitation id="136A2A06FFB3E938FF26F929FF552470" box="[181,197,1770,1797]" captionStart="FIGURE 5" captionStartId="8.[151,250,1979,2003]" captionTargetBox="[174,1414,1402,1956]" captionTargetId="figure-84@8.[174,1414,1402,1956]" captionTargetPageId="8" captionText="FIGURE 5. Pectoral fin of Luciogobius punctilineatus n. sp. (A: KAUM–I. 157125) and Luciogobius sp. 11 (B: KAUM–I. 159027), showing indentation of fin membrane." figureDoi="http://doi.org/10.5281/zenodo.6556892" httpUri="https://zenodo.org/record/6556892/files/figure.png" pageId="7" pageNumber="144">5</figureCitation>
, 
<figureCitation id="136A2A06FFB3E938FF40F928FF722470" box="[211,226,1771,1797]" captionStart="FIGURE 8" captionStartId="11.[151,250,1149,1173]" captionTargetBox="[151,1437,381,1125]" captionTargetId="figure-111@11.[151,1437,381,1125]" captionTargetPageId="11" captionText="FIGURE 8. Fresh (upper and middle) and preserved (lower) specimens of Luciogobius sp. 11 (KAUM–I. 159027 25.9 mm SL) from Kochi Prefecture." figureDoi="http://doi.org/10.5281/zenodo.6556903" httpUri="https://zenodo.org/record/6556903/files/figure.png" pageId="7" pageNumber="144">8</figureCitation>
), and the first anal-fin pterygiophore is usually inserted behind the fifth haemal spine in 
<taxonomicName id="4C514D00FFB3E938FB4DF928FA0D2470" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[1246,1437,1771,1797]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FB4DF928FA0D2470" box="[1246,1437,1771,1797]" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
and behind the fourth spine in 
<taxonomicName id="4C514D00FFB3E938FE60F8CCFD35245C" box="[499,677,1807,1833]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FE60F8CCFDED245C" box="[499,637,1807,1833]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (
<tableCitation id="C6D30338FFB3E938FD44F8CCFCA0245C" box="[727,816,1807,1833]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[151,1411,263,1001]" captionTargetPageId="10" captionText="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FF54FC582382">Table 2</tableCitation>
). The new species further differs from 
<taxonomicName id="4C514D00FFB3E938FB79F8CCFA0C245C" box="[1258,1436,1807,1833]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FB79F8CCFAE4245C" box="[1258,1396,1807,1833]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
<quantity id="4CA99B66FFB3E938FF04F8F0FF402438" box="[151,208,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="7" pageNumber="144" unit="in" value="11.0">11 in</quantity>
having relatively lower numbers of anal-fin rays [12–14 (modally 13) vs. 14–16 (15) in 
<taxonomicName id="4C514D00FFB3E938FB2CF8F0FAE22438" box="[1215,1394,1843,1869]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FB2CF8F0FAD92438" box="[1215,1353,1843,1869]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11; 
<bibRefCitation id="EFC04B72FFB3E938FF04F894FE0D2404" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[151,413,1879,1905]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FE88F894FEDD2404" box="[283,333,1879,1905]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
., 2019
</bibRefCitation>
; 
<tableCitation id="C6D30338FFB3E938FE3BF894FD962404" box="[424,518,1879,1905]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1420,226,1952]" captionTargetPageId="5" captionText="TABLE 1. Counts and proportional measurements of Luciogobius punctilineatus n. sp. (modes and means ± standard deviation shown in parentheses)." httpUri="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" pageId="7" pageNumber="144" tableUuid="DF2E660BFFB1E93AFF04FF54FE6623A6">Tables 1</tableCitation>
, 
<tableCitation id="C6D30338FFB3E938FD86F895FDB52404" box="[533,549,1878,1905]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[151,1411,263,1001]" captionTargetPageId="10" captionText="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="7" pageNumber="144" tableUuid="DF2E660BFFBEE935FF04FF54FC582382">2</tableCitation>
] and vertebrae [16–18 (18) + 22–24 (23) = 39–42 (40) vs. 17–18 (18) + 22–25 (23) = 40–43 (41); 
<bibRefCitation id="EFC04B72FFB3E938FEFAF8B8FDF924E0" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[361,617,1915,1941]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FE78F8B8FD8A24E0" box="[491,538,1915,1941]" italics="true" pageId="7" pageNumber="144">et al</emphasis>
., 2019
</bibRefCitation>
; 
<bibRefCitation id="EFC04B72FFB3E938FDE0F8B8FC2124E0" author="Koreeda, R. &amp; Motomura, H." box="[627,945,1915,1941]" pageId="7" pageNumber="144" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
; 
<tableCitation id="C6D30338FFB3E938FC28F8B8FB8724E0" box="[955,1047,1914,1941]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1420,226,1952]" captionTargetPageId="5" captionText="TABLE 1. Counts and proportional measurements of Luciogobius punctilineatus n. sp. (modes and means ± standard deviation shown in parentheses)." httpUri="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" pageId="7" pageNumber="144" tableUuid="DF2E660BFFB1E93AFF04FF54FE6623A6">Tables 1</tableCitation>
], and a relatively higher number of pectoral-fin rays [8–12 (10) vs. 8–9 (9); 
<bibRefCitation id="EFC04B72FFB3E938FDC8F85CFCCE24CC" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[603,862,1951,1977]" pageId="7" pageNumber="144" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB3E938FD4DF85CFC8624CC" box="[734,790,1951,1977]" italics="true" pageId="7" pageNumber="144">et al.</emphasis>
, 2019
</bibRefCitation>
; 
<tableCitation id="C6D30338FFB3E938FCFAF85CFC5724CC" box="[873,967,1950,1977]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[166,1420,226,1952]" captionTargetPageId="5" captionText="TABLE 1. Counts and proportional measurements of Luciogobius punctilineatus n. sp. (modes and means ± standard deviation shown in parentheses)." httpUri="http://table.plazi.org/id/DF2E660BFFB1E93AFF04FF54FE6623A6" pageId="7" pageNumber="144" tableUuid="DF2E660BFFB1E93AFF04FF54FE6623A6">Tables 1</tableCitation>
]. The anus and anal-fin origin of 
<taxonomicName id="4C514D00FFB3E938FAACF85CFF6F24A8" authorityName="Koreeda &amp; Motomura" authorityYear="2022" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFB3E938FAACF85CFF6F24A8" italics="true" pageId="7" pageNumber="144">L. punctilineatus</emphasis>
</taxonomicName>
are located more anteriorly and posteriorly, respectively, compared with 
<taxonomicName id="4C514D00FFB3E938FBA1F800FB7424A8" box="[1074,1252,1987,2013]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="144" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB3E938FBA1F800FB2C24A8" box="[1074,1212,1987,2013]" italics="true" pageId="7" pageNumber="144">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (
<figureCitation id="136A2A06FFB3E938FA86F800FA0024A8" box="[1301,1424,1987,2013]" captionStart="FIGURE 9" captionStartId="12.[151,250,1813,1837]" captionTargetBox="[241,1346,191,1781]" captionTargetId="figure-30@12.[214,1374,181,1789]" captionTargetPageId="12" captionText="FIGURE 9. Relationships of (A) snout length, (B) AAA distance (measured between posterior margin of anus and anal-fin origin), (C) PAL (pre-anus length), and (D) pre-anal-fin length as % of standard length, and (E) snout length as % of AAA, and (F) PAL as % of pre-anal-fin length vs. standard length (mm) in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Closed red star, holotype of L. punctilineatus n. sp.; open red stars, paratypes of L. punctilineatus n. sp. from Shimokoshiki-shima island, Tanega-shima island; yellow stars, L. punctilineatus n. sp. from Amami-oshima island; green stars; L. punctilineatus n. sp. from Kochi Prefecture and Osumi Peninsula; blue triangles, Luciogobius sp. 11." figureDoi="http://doi.org/10.5281/zenodo.6556909" httpUri="https://zenodo.org/record/6556909/files/figure.png" pageId="7" pageNumber="144">Fig. 9C, D</figureCitation>
). In preserved specimens, a single longitudinal row formed by black spots on the mid-lateral surface of the body is more distinct in 
<taxonomicName id="4C514D00FFBEE935FEF4F9CFFDB42550" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[359,548,1547,1573]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FEF4F9CFFDB42550" box="[359,548,1547,1573]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="4C514D00FFBEE935FDEDF9C8FCA02553" box="[638,816,1547,1574]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FDEDF9C8FC982550" box="[638,776,1547,1573]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (see 
<bibRefCitation id="EFC04B72FFBEE935FC1EF9C8FB5F2550" author="Koreeda, R. &amp; Motomura, H." box="[909,1231,1547,1574]" pageId="10" pageNumber="147" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
: fig. 8; 
<figureCitation id="136A2A06FFBEE935FAB7F9C8FAE22550" box="[1316,1394,1547,1573]" captionStart="FIGURE 4" captionStartId="8.[151,250,1281,1305]" captionTargetBox="[180,1408,181,1257]" captionTargetId="figure-30@8.[180,1408,181,1257]" captionTargetPageId="8" captionText="FIGURE 4. Preserved specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.6556890" httpUri="https://zenodo.org/record/6556890/files/figure.png" pageId="10" pageNumber="147">Figs. 4</figureCitation>
, 
<figureCitation id="136A2A06FFBEE935FAEDF9C8FA1D2550" box="[1406,1421,1547,1573]" captionStart="FIGURE 8" captionStartId="11.[151,250,1149,1173]" captionTargetBox="[151,1437,381,1125]" captionTargetId="figure-111@11.[151,1437,381,1125]" captionTargetPageId="11" captionText="FIGURE 8. Fresh (upper and middle) and preserved (lower) specimens of Luciogobius sp. 11 (KAUM–I. 159027 25.9 mm SL) from Kochi Prefecture." figureDoi="http://doi.org/10.5281/zenodo.6556903" httpUri="https://zenodo.org/record/6556903/files/figure.png" pageId="10" pageNumber="147">8</figureCitation>
), although this difference is unclear in live individuals. 
<taxonomicName id="4C514D00FFBEE935FD7DF9ECFC0F253F" box="[750,927,1583,1610]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FD7DF9ECFCE8253C" box="[750,888,1583,1609]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11 also exhibits sexual dichromatism (yellow males and brown females; 
<bibRefCitation id="EFC04B72FFBEE935FE53F990FD522518" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[448,706,1619,1646]" pageId="10" pageNumber="147" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFBEE935FDD1F997FDE22518" box="[578,626,1619,1645]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2019
</bibRefCitation>
), which is unknown in 
<taxonomicName id="4C514D00FFBEE935FC51F997FBED2518" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[962,1149,1619,1645]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FC51F997FBED2518" box="[962,1149,1619,1645]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
. Accordingly, individuals of 
<taxonomicName id="4C514D00FFBEE935FF27F9BBFEE025E4" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[180,368,1655,1681]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FF27F9BBFEE025E4" box="[180,368,1655,1681]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
are similar to males of 
<taxonomicName id="4C514D00FFBEE935FDE3F9B4FCB125E7" box="[624,801,1655,1682]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FDE3F9B4FD6A25E4" box="[624,762,1655,1681]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
<quantity id="4CA99B66FFBEE935FCB5F9B4FCCD25E4" box="[806,861,1655,1681]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="10" pageNumber="147" unit="in" value="11.0">11 in</quantity>
having a yellow body (
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Shibukawa 
<emphasis id="B925EA91FFBEE935FB71F9BBFA8925E4" box="[1250,1305,1655,1681]" italics="true" pageId="10" pageNumber="147">et al.</emphasis>
, 2019
</bibRefCitation>
: figs. 26D–E, 27D; 
<bibRefCitation id="EFC04B72FFBEE935FEA7F958FDE625C0" author="Koreeda, R. &amp; Motomura, H." box="[308,630,1691,1718]" pageId="10" pageNumber="147" pagination="27 - 52" refId="ref8136" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [in Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
: fig. 8; 
<figureCitation id="136A2A06FFBEE935FD58F958FCB425C0" box="[715,804,1691,1717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[178,1410,1041,1953]" captionTargetId="figure-197@6.[178,1410,1035,1953]" captionTargetPageId="6" captionText="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." figureDoi="http://doi.org/10.5281/zenodo.6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="10" pageNumber="147">Fig. 3A</figureCitation>
).
</paragraph>
<caption id="DF2E660BFFBCE937FF04FAC2FAAC2617" ID-DOI="http://doi.org/10.5281/zenodo.6556890" ID-Zenodo-Dep="6556890" httpUri="https://zenodo.org/record/6556890/files/figure.png" pageId="8" pageNumber="145" startId="8.[151,250,1281,1305]" targetBox="[180,1408,181,1257]" targetPageId="8">
<paragraph id="8BEE3683FFBCE937FF04FAC2FAAC2617" blockId="8.[151,1437,1281,1378]" pageId="8" pageNumber="145">
<emphasis id="B925EA91FFBCE937FF04FAC2FE87266C" bold="true" box="[151,279,1281,1305]" pageId="8" pageNumber="145">FIGURE 4.</emphasis>
Preserved specimens of 
<taxonomicName id="4C514D00FFBCE937FDB7FAC2FCAC266C" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[548,828,1281,1305]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="145" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBCE937FDB7FAC2FCAC266C" box="[548,828,1281,1305]" italics="true" pageId="8" pageNumber="145">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBCE937FCD5FAC1FC13266F" bold="true" box="[838,899,1282,1306]" pageId="8" pageNumber="145">
<taxonomicNameLabel id="A21657EAFFBCE937FCD5FAC1FC13266F" box="[838,899,1282,1306]" pageId="8" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL).
</paragraph>
</caption>
<caption id="DF2E660BFFBCE937FF04F878FDED2482" ID-DOI="http://doi.org/10.5281/zenodo.6556892" ID-Zenodo-Dep="6556892" httpUri="https://zenodo.org/record/6556892/files/figure.png" pageId="8" pageNumber="145" startId="8.[151,250,1979,2003]" targetBox="[174,1414,1402,1956]" targetPageId="8">
<paragraph id="8BEE3683FFBCE937FF04F878FDED2482" blockId="8.[151,1437,1979,2039]" pageId="8" pageNumber="145">
<emphasis id="B925EA91FFBCE937FF04F878FE8524A6" bold="true" box="[151,277,1979,2003]" pageId="8" pageNumber="145">FIGURE 5.</emphasis>
Pectoral fin of 
<taxonomicName id="4C514D00FFBCE937FE29F878FD4024A6" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[442,720,1979,2003]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="145" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBCE937FE29F878FD4024A6" box="[442,720,1979,2003]" italics="true" pageId="8" pageNumber="145">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBCE937FD4BF878FC8324A6" bold="true" box="[728,787,1979,2003]" pageId="8" pageNumber="145">
<taxonomicNameLabel id="A21657EAFFBCE937FD4BF878FC8324A6" box="[728,787,1979,2003]" pageId="8" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A: KAUM–I. 157125) and 
<taxonomicName id="4C514D00FFBCE937FBACF878FB7224A6" box="[1087,1250,1979,2003]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="145" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBCE937FBACF878FB2C24A6" box="[1087,1212,1979,2003]" italics="true" pageId="8" pageNumber="145">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (B: KAUM–I. 159027), showing indentation of fin membrane.
</paragraph>
</caption>
<caption id="DF2E660BFFBDE936FF04FCE6FCC12014" ID-DOI="http://doi.org/10.5281/zenodo.6556894" ID-Zenodo-Dep="6556894" httpUri="https://zenodo.org/record/6556894/files/figure.png" pageId="9" pageNumber="146" startId="9.[151,250,805,829]" targetBox="[159,1428,186,776]" targetPageId="9">
<paragraph id="8BEE3683FFBDE936FF04FCE6FCC12014" blockId="9.[151,1436,805,866]" pageId="9" pageNumber="146">
<emphasis id="B925EA91FFBDE936FF04FCE6FE842048" bold="true" box="[151,276,805,829]" pageId="9" pageNumber="146">FIGURE 6.</emphasis>
Neighbor-joining tree based on 12S ribosomal RNA gene sequences (133 bp) of species of 
<taxonomicName id="4C514D00FFBDE936FB51FCE6FAAF2048" authorityName="Gill" authorityYear="1859" box="[1218,1343,805,829]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="146" phylum="Chordata" rank="genus">
<emphasis id="B925EA91FFBDE936FB51FCE6FAAF2048" box="[1218,1343,805,829]" italics="true" pageId="9" pageNumber="146">Luciogobius</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C514D00FFBDE936FAE9FCE5FF5E2014" authorityName="Snyder" authorityYear="1909" class="Actinopterygii" family="Gobiidae" genus="Inu" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="koma">
<emphasis id="B925EA91FFBDE936FAE9FCE5FF5E2014" italics="true" pageId="9" pageNumber="146">Inu koma</emphasis>
</taxonomicName>
as outgroup. Numbers near branches indicate bootstrap values.
</paragraph>
</caption>
<caption id="DF2E660BFFBDE936FF04F88FFD5124D9" ID-DOI="http://doi.org/10.5281/zenodo.6556896" ID-Zenodo-Dep="6556896" httpUri="https://zenodo.org/record/6556896/files/figure.png" pageId="9" pageNumber="146" startId="9.[151,250,1868,1892]" targetBox="[151,1436,889,1844]" targetPageId="9">
<paragraph id="8BEE3683FFBDE936FF04F88FFD5124D9" blockId="9.[151,1437,1868,1964]" pageId="9" pageNumber="146">
<emphasis id="B925EA91FFBDE936FF04F88FFE822411" bold="true" box="[151,274,1868,1892]" pageId="9" pageNumber="146">FIGURE 7.</emphasis>
Habitats of 
<taxonomicName id="4C514D00FFBDE936FE1FF88FFD0F2411" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[396,671,1868,1892]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="146" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBDE936FE1FF88FFD0F2411" box="[396,671,1868,1892]" italics="true" pageId="9" pageNumber="146">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBDE936FD37F88FFD4C2411" bold="true" box="[676,732,1868,1892]" pageId="9" pageNumber="146">
<taxonomicNameLabel id="A21657EAFFBDE936FD37F88FFD4C2411" box="[676,732,1868,1892]" pageId="9" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
near (A) the Shirinashi River mouth, western coast of Satsuma Peninsula, and (B) the Shimadomari River mouth, southern coast of Osumi Peninsula. Black arrows indicate collection points, near fresh water springs; blue arrows indicate river mouths.
</paragraph>
</caption>
<caption id="DF2E660BFFBEE935FF04FF54FC582382" ID-Table-UUID="DF2E660BFFBEE935FF04FF54FC582382" httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="10" pageNumber="147" startId="10.[151,239,151,175]" targetBox="[151,1411,263,1001]" targetIsTable="true" targetPageId="10">
<paragraph id="8BEE3683FFBEE935FF04FF54FC582382" blockId="10.[151,1437,151,247]" pageId="10" pageNumber="147">
<emphasis id="B925EA91FFBEE935FF04FF54FE9C23DA" bold="true" box="[151,268,151,175]" pageId="10" pageNumber="147">TABLE 2.</emphasis>
Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in 
<taxonomicName id="4C514D00FFBEE935FD9CFF78FCBF23A6" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[527,815,187,211]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBEE935FD9CFF78FCBF23A6" box="[527,815,187,211]" italics="true" pageId="10" pageNumber="147">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBEE935FCA5FF78FCE123A6" bold="true" box="[822,881,187,211]" pageId="10" pageNumber="147">
<taxonomicNameLabel id="A21657EAFFBEE935FCA5FF78FCE123A6" box="[822,881,187,211]" pageId="10" pageNumber="147" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C514D00FFBEE935FC30FF78FBDC23A6" box="[931,1100,187,211]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FC30FF78FBB723A6" box="[931,1063,187,211]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11. Gray shaded values include data from 
<bibRefCitation id="EFC04B72FFBEE935FE96FF1CFD982382" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[261,520,223,247]" pageId="10" pageNumber="147" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFBEE935FE11FF23FE212382" box="[386,433,223,247]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
. (2019)
</bibRefCitation>
. *Including holotype of 
<taxonomicName id="4C514D00FFBEE935FC9EFF23FC512382" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[781,961,223,247]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FC9EFF23FC512382" box="[781,961,223,247]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEE3683FFBEE935FE79FEC4FC46209C" pageId="10" pageNumber="147">
<table id="F951C423FFBE16C0FF04FEC4FA13209C" box="[151,1411,263,1001]" gridcols="11" gridrows="11" pageId="10" pageNumber="147">
<tr id="356134C1FFBE16C0FF04FEC4FA132255" box="[151,1411,263,288]" gridrow="0" pageId="10" pageNumber="147" rowspan-0="1" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FE79FEC4FD162255" box="[490,646,263,288]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="10" pageNumber="147">Dorsal-fin rays</th>
<th id="76B05DBDFFBE16C0FCF9FEC4FC612255" box="[874,1009,263,288]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="10" pageNumber="147">Anal-fin rays</th>
</tr>
<tr id="356134C1FFBE16C0FF04FEECFA13223D" box="[151,1411,303,328]" gridrow="1" pageId="10" pageNumber="147" rowspan-0="1">
<td id="76B05DBDFFBE16C0FE79FEECFD93223D" box="[490,515,303,328]" gridcol="1" gridrow="1" pageId="10" pageNumber="147">10</td>
<td id="76B05DBDFFBE16C0FDD9FEECFD16223D" box="[586,646,303,328]" gridcol="2" gridrow="1" pageId="10" pageNumber="147">11</td>
<td id="76B05DBDFFBE16C0FD39FEECFD54223D" box="[682,708,303,328]" gridcol="3" gridrow="1" pageId="10" pageNumber="147">12</td>
<td id="76B05DBDFFBE16C0FC99FEECFCB4223D" box="[778,804,303,328]" gridcol="4" gridrow="1" pageId="10" pageNumber="147">13</td>
<td id="76B05DBDFFBE16C0FCF9FEECFC14223D" box="[874,900,303,328]" gridcol="5" gridrow="1" pageId="10" pageNumber="147">11</td>
<td id="76B05DBDFFBE16C0FC5AFEECFC61223D" box="[969,1009,303,328]" gridcol="6" gridrow="1" pageId="10" pageNumber="147">12</td>
<td id="76B05DBDFFBE16C0FBBAFEECFBC1223D" box="[1065,1105,303,328]" gridcol="7" gridrow="1" pageId="10" pageNumber="147">13</td>
<td id="76B05DBDFFBE16C0FB1AFEECFB35223D" box="[1161,1189,303,328]" gridcol="8" gridrow="1" pageId="10" pageNumber="147">14</td>
<td id="76B05DBDFFBE16C0FB7AFEECFA95223D" box="[1257,1285,303,328]" gridcol="9" gridrow="1" pageId="10" pageNumber="147">15</td>
<td id="76B05DBDFFBE16C0FADAFEECFA13223D" box="[1353,1411,303,328]" gridcol="10" gridrow="1" pageId="10" pageNumber="147">16</td>
</tr>
<tr id="356134C1FFBE16C0FF04FE9BFA1322ED" box="[151,1411,344,408]" gridrow="2" pageId="10" pageNumber="147" rowspan-10="1" rowspan-4="1" rowspan-5="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FF04FE9BFE1822ED" box="[151,392,344,408]" gridcol="0" gridrow="2" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FE9AFEC02205" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[166,336,344,368]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FF35FE9AFEC02205" box="[166,336,344,368]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
Kochi to Tanegashima
</th>
<td id="76B05DBDFFBE16C0FE79FE9BFD9322ED" box="[490,515,344,408]" gridcol="1" gridrow="2" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FDD9FE9BFD1622ED" box="[586,646,344,408]" gridcol="2" gridrow="2" pageId="10" pageNumber="147">16*</td>
<td id="76B05DBDFFBE16C0FD39FE9BFD5422ED" box="[682,708,344,408]" gridcol="3" gridrow="2" pageId="10" pageNumber="147">5</td>
<td id="76B05DBDFFBE16C0FC5AFE9BFC6122ED" box="[969,1009,344,408]" gridcol="6" gridrow="2" pageId="10" pageNumber="147">2</td>
<td id="76B05DBDFFBE16C0FBBAFE9BFBC122ED" box="[1065,1105,344,408]" gridcol="7" gridrow="2" pageId="10" pageNumber="147">14*</td>
<td id="76B05DBDFFBE16C0FB1AFE9BFB3522ED" box="[1161,1189,344,408]" gridcol="8" gridrow="2" pageId="10" pageNumber="147">9</td>
</tr>
<tr id="356134C1FFBE16C0FF04FE6BFA13229C" box="[151,1411,424,489]" gridrow="3" pageId="10" pageNumber="147" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FF04FE6BFE18229C" box="[151,392,424,489]" gridcol="0" gridrow="3" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FE6AFEC022B5" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[166,336,424,448]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FF35FE6AFEC022B5" box="[166,336,424,448]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
Amami-oshima
</th>
<td id="76B05DBDFFBE16C0FE79FE6BFD93229C" box="[490,515,424,489]" gridcol="1" gridrow="3" pageId="10" pageNumber="147">6</td>
<td id="76B05DBDFFBE16C0FDD9FE6BFD16229C" box="[586,646,424,489]" gridcol="2" gridrow="3" pageId="10" pageNumber="147">2</td>
<td id="76B05DBDFFBE16C0FC5AFE6BFC61229C" box="[969,1009,424,489]" gridcol="6" gridrow="3" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FBBAFE6BFBC1229C" box="[1065,1105,424,489]" gridcol="7" gridrow="3" pageId="10" pageNumber="147">3</td>
<td id="76B05DBDFFBE16C0FB1AFE6BFB35229C" box="[1161,1189,424,489]" gridcol="8" gridrow="3" pageId="10" pageNumber="147">1</td>
</tr>
<tr id="356134C1FFBE16C0FF04FE3AFA13214C" box="[151,1411,505,569]" gridrow="4" pageId="10" pageNumber="147" rowspan-1="1" rowspan-10="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76B05DBDFFBE16C0FF04FE3AFE18214C" box="[151,392,505,569]" gridcol="0" gridrow="4" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FE3AFED72165" box="[166,327,505,529]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FF35FE3AFEB32164" box="[166,291,505,529]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11 Shizuoka and Kochi
</th>
<td id="76B05DBDFFBE16C0FD39FE3AFD54214C" box="[682,708,505,569]" gridcol="3" gridrow="4" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FC99FE3AFCB4214C" box="[778,804,505,569]" gridcol="4" gridrow="4" pageId="10" pageNumber="147">1</td>
<td id="76B05DBDFFBE16C0FB1AFE3AFB35214C" box="[1161,1189,505,569]" gridcol="8" gridrow="4" pageId="10" pageNumber="147">2</td>
<td id="76B05DBDFFBE16C0FB7AFE3AFA95214C" box="[1257,1285,505,569]" gridcol="9" gridrow="4" pageId="10" pageNumber="147">3</td>
</tr>
<tr id="356134C1FFBE16C0FF04FDA9FA1321F7" box="[151,1411,618,642]" gridrow="5" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF04FDA9FA1321F7" box="[151,1411,618,642]" colspan="11" colspanRight="10" gridcol="0" gridrow="5" pageId="10" pageNumber="147">continued.</th>
</tr>
<tr id="356134C1FFBE16C0FF04FD50FA1321BA" box="[151,1411,659,719]" gridrow="6" pageId="10" pageNumber="147" rowspan-0="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="76B05DBDFFBE16C0FE79FD50FD1621BA" box="[490,646,659,719]" colspan="2" colspanRight="1" gridcol="1" gridrow="6" pageId="10" pageNumber="147">Pectral-fin rays</td>
<td id="76B05DBDFFBE16C0FC5AFD50FA1321BA" box="[969,1411,659,719]" colspan="5" colspanRight="4" gridcol="6" gridrow="6" pageId="10" pageNumber="147">First anal-fin pterygiophore inserted behind haemal spine</td>
</tr>
<tr id="356134C1FFBE16C0FF04FD1CFA132182" box="[151,1411,735,759]" gridrow="7" pageId="10" pageNumber="147" rowspan-0="1">
<td id="76B05DBDFFBE16C0FE79FD1CFD932182" box="[490,515,735,759]" gridcol="1" gridrow="7" pageId="10" pageNumber="147">8</td>
<td id="76B05DBDFFBE16C0FDD9FD1CFD162182" box="[586,646,735,759]" gridcol="2" gridrow="7" pageId="10" pageNumber="147">9</td>
<td id="76B05DBDFFBE16C0FD39FD1CFD542182" box="[682,708,735,759]" gridcol="3" gridrow="7" pageId="10" pageNumber="147">10</td>
<td id="76B05DBDFFBE16C0FC99FD1CFCB42182" box="[778,804,735,759]" gridcol="4" gridrow="7" pageId="10" pageNumber="147">11</td>
<td id="76B05DBDFFBE16C0FCF9FD1CFC142182" box="[874,900,735,759]" gridcol="5" gridrow="7" pageId="10" pageNumber="147">12</td>
<td id="76B05DBDFFBE16C0FC5AFD1CFC612182" box="[969,1009,735,759]" gridcol="6" gridrow="7" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FBBAFD1CFBC12182" box="[1065,1105,735,759]" gridcol="7" gridrow="7" pageId="10" pageNumber="147">5</td>
<td id="76B05DBDFFBE16C0FB1AFD1CFB352182" box="[1161,1189,735,759]" gridcol="8" gridrow="7" pageId="10" pageNumber="147">6</td>
<td id="76B05DBDFFBE16C0FB7AFD1CFA952182" box="[1257,1285,735,759]" gridcol="9" gridrow="7" pageId="10" pageNumber="147">7</td>
<td id="76B05DBDFFBE16C0FADAFD1CFA132182" box="[1353,1411,735,759]" gridcol="10" gridrow="7" pageId="10" pageNumber="147">8</td>
</tr>
<tr id="356134C1FFBE16C0FF04FCC4FA13203D" box="[151,1411,775,840]" gridrow="8" pageId="10" pageNumber="147" rowspan-1="1" rowspan-10="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FF04FCC4FE18203D" box="[151,392,775,840]" gridcol="0" gridrow="8" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FCCBFEC0206A" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[166,336,775,799]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FF35FCCBFEC0206A" box="[166,336,775,799]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
Kochi to Tanegashima
</th>
<td id="76B05DBDFFBE16C0FDD9FCC4FD16203D" box="[586,646,775,840]" gridcol="2" gridrow="8" pageId="10" pageNumber="147">6</td>
<td id="76B05DBDFFBE16C0FD39FCC4FD54203D" box="[682,708,775,840]" gridcol="3" gridrow="8" pageId="10" pageNumber="147">9</td>
<td id="76B05DBDFFBE16C0FC99FCC4FCB4203D" box="[778,804,775,840]" gridcol="4" gridrow="8" pageId="10" pageNumber="147">9*</td>
<td id="76B05DBDFFBE16C0FCF9FCC4FC14203D" box="[874,900,775,840]" gridcol="5" gridrow="8" pageId="10" pageNumber="147">1</td>
<td id="76B05DBDFFBE16C0FC5AFCC4FC61203D" box="[969,1009,775,840]" gridcol="6" gridrow="8" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FBBAFCC4FBC1203D" box="[1065,1105,775,840]" gridcol="7" gridrow="8" pageId="10" pageNumber="147">15*</td>
<td id="76B05DBDFFBE16C0FB1AFCC4FB35203D" box="[1161,1189,775,840]" gridcol="8" gridrow="8" pageId="10" pageNumber="147">2</td>
</tr>
<tr id="356134C1FFBE16C0FF04FC9BFA1320ED" box="[151,1411,856,920]" gridrow="9" pageId="10" pageNumber="147" rowspan-10="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FF04FC9BFE1820ED" box="[151,392,856,920]" gridcol="0" gridrow="9" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FC9BFEC02005" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[166,336,856,880]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FF35FC9BFEC02005" box="[166,336,856,880]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
Amami-oshima
</th>
<td id="76B05DBDFFBE16C0FE79FC9BFD9320ED" box="[490,515,856,920]" gridcol="1" gridrow="9" pageId="10" pageNumber="147">1</td>
<td id="76B05DBDFFBE16C0FDD9FC9BFD1620ED" box="[586,646,856,920]" gridcol="2" gridrow="9" pageId="10" pageNumber="147">3</td>
<td id="76B05DBDFFBE16C0FD39FC9BFD5420ED" box="[682,708,856,920]" gridcol="3" gridrow="9" pageId="10" pageNumber="147">2</td>
<td id="76B05DBDFFBE16C0FC99FC9BFCB420ED" box="[778,804,856,920]" gridcol="4" gridrow="9" pageId="10" pageNumber="147">1</td>
<td id="76B05DBDFFBE16C0FBBAFC9BFBC120ED" box="[1065,1105,856,920]" gridcol="7" gridrow="9" pageId="10" pageNumber="147">3</td>
<td id="76B05DBDFFBE16C0FB1AFC9BFB3520ED" box="[1161,1189,856,920]" gridcol="8" gridrow="9" pageId="10" pageNumber="147">2</td>
</tr>
<tr id="356134C1FFBE16C0FF04FC6BFA13209C" box="[151,1411,936,1001]" gridrow="10" pageId="10" pageNumber="147" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76B05DBDFFBE16C0FF04FC6BFE18209C" box="[151,392,936,1001]" gridcol="0" gridrow="10" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF35FC6BFED720B5" box="[166,327,936,960]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FF35FC6BFEB320B5" box="[166,291,936,960]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11 Shizuoka and Kochi
</th>
<td id="76B05DBDFFBE16C0FE79FC6BFD93209C" box="[490,515,936,1001]" gridcol="1" gridrow="10" pageId="10" pageNumber="147">1</td>
<td id="76B05DBDFFBE16C0FDD9FC6BFD16209C" box="[586,646,936,1001]" gridcol="2" gridrow="10" pageId="10" pageNumber="147">4</td>
<td id="76B05DBDFFBE16C0FC5AFC6BFC61209C" box="[969,1009,936,1001]" gridcol="6" gridrow="10" pageId="10" pageNumber="147">4</td>
</tr>
</table>
</paragraph>
<caption id="DF2E660BFFBEE935FF04FBD9FA1D2747" ID-Table-UUID="DF2E660BFFBEE935FF04FBD9FA1D2747" box="[151,1421,1050,1074]" httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FBD9FA1D2747" pageId="10" pageNumber="147" startId="10.[151,239,1050,1074]" targetBox="[166,1232,1090,1427]" targetIsTable="true" targetPageId="10">
<paragraph id="8BEE3683FFBEE935FF04FBD9FA1D2747" blockId="10.[151,1421,1050,1074]" box="[151,1421,1050,1074]" pageId="10" pageNumber="147">
<emphasis id="B925EA91FFBEE935FF04FBD9FE9A2747" bold="true" box="[151,266,1050,1074]" pageId="10" pageNumber="147">TABLE 3.</emphasis>
Comparisons of selected morphometrics between 
<taxonomicName id="4C514D00FFBEE935FCB6FBD9FBD72747" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[805,1095,1050,1074]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBEE935FCB6FBD9FBD72747" box="[805,1095,1050,1074]" italics="true" pageId="10" pageNumber="147">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBEE935FBDDFBD9FB1A2747" bold="true" box="[1102,1162,1050,1074]" pageId="10" pageNumber="147">
<taxonomicNameLabel id="A21657EAFFBEE935FBDDFBD9FB1A2747" box="[1102,1162,1050,1074]" pageId="10" pageNumber="147" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C514D00FFBEE935FB2DFBD9FAF72744" box="[1214,1383,1050,1074]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FB2DFBD9FAD22747" box="[1214,1346,1050,1074]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11.
</paragraph>
</caption>
<paragraph id="8BEE3683FFBEE935FDDEFB81FB40261A" pageId="10" pageNumber="147">
<table id="F951C423FFBE16C0FF35FB81FB4026E6" box="[166,1232,1090,1427]" gridcols="3" gridrows="7" pageId="10" pageNumber="147">
<tr id="356134C1FFBE16C0FF35FB81FB40272F" box="[166,1232,1090,1114]" gridrow="0" pageId="10" pageNumber="147" rowspan-0="1">
<th id="76B05DBDFFBE16C0FDDEFB81FCF0272F" box="[589,864,1090,1114]" gridcol="1" gridrow="0" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FDDEFB81FCF0272F" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[589,864,1090,1114]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FDDEFB81FCF0272F" box="[589,864,1090,1114]" italics="true" pageId="10" pageNumber="147">Luciogobius punctilineatus</emphasis>
</taxonomicName>
</th>
<th id="76B05DBDFFBE16C0FC6FFB81FB40272F" box="[1020,1232,1090,1114]" gridcol="2" gridrow="0" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FC6FFB81FB0D272F" box="[1020,1181,1090,1114]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FC6FFB81FBE9272F" box="[1020,1145,1090,1114]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11
</th>
</tr>
<tr id="356134C1FFBE16C0FF35FBA9FB4027F7" box="[166,1232,1130,1154]" gridrow="1" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FBA9FE3E27F7" box="[166,430,1130,1154]" gridcol="0" gridrow="1" pageId="10" pageNumber="147">Snout length (%SL)</th>
<td id="76B05DBDFFBE16C0FDDEFBA9FCF027F7" box="[589,864,1130,1154]" gridcol="1" gridrow="1" pageId="10" pageNumber="147">3.1–4.3% (3.5%)</td>
<td id="76B05DBDFFBE16C0FC6FFBA9FB4027F7" box="[1020,1232,1130,1154]" gridcol="2" gridrow="1" pageId="10" pageNumber="147">3.7–4.2% (3.9%)</td>
</tr>
<tr id="356134C1FFBE16C0FF35FB51FB4027DE" box="[166,1232,1170,1195]" gridrow="2" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FB51FE3E27DE" box="[166,430,1170,1195]" gridcol="0" gridrow="2" pageId="10" pageNumber="147">AAA distance (%SL)</th>
<td id="76B05DBDFFBE16C0FDDEFB51FCF027DE" box="[589,864,1170,1195]" gridcol="1" gridrow="2" pageId="10" pageNumber="147">11.4–16.9% (13.3%)e</td>
<td id="76B05DBDFFBE16C0FC6FFB51FB4027DE" box="[1020,1232,1170,1195]" gridcol="2" gridrow="2" pageId="10" pageNumber="147">9.1–10.4% (9.6%)</td>
</tr>
<tr id="356134C1FFBE16C0FF35FB78FB4027A6" box="[166,1232,1211,1235]" gridrow="3" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FB78FE3E27A6" box="[166,430,1211,1235]" gridcol="0" gridrow="3" pageId="10" pageNumber="147">Pre-anus length (%SL)</th>
<td id="76B05DBDFFBE16C0FDDEFB78FCF027A6" box="[589,864,1211,1235]" gridcol="1" gridrow="3" pageId="10" pageNumber="147">52.2–60.3% (54.5%)</td>
<td id="76B05DBDFFBE16C0FC6FFB78FB4027A6" box="[1020,1232,1211,1235]" gridcol="2" gridrow="3" pageId="10" pageNumber="147">55.2–57.5% (56.1%)</td>
</tr>
<tr id="356134C1FFBE16C0FF35FB20FB40278E" box="[166,1232,1251,1275]" gridrow="4" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FB20FE3E278E" box="[166,430,1251,1275]" gridcol="0" gridrow="4" pageId="10" pageNumber="147">Pre-anal-fin length (%SL)</th>
<td id="76B05DBDFFBE16C0FDDEFB20FCF0278E" box="[589,864,1251,1275]" gridcol="1" gridrow="4" pageId="10" pageNumber="147">66.0–71.0% (68.2%)</td>
<td id="76B05DBDFFBE16C0FC6FFB20FB40278E" box="[1020,1232,1251,1275]" gridcol="2" gridrow="4" pageId="10" pageNumber="147">64.4–66.3% (65.1%)</td>
</tr>
<tr id="356134C1FFBE16C0FF35FAC8FB402632" box="[166,1232,1291,1351]" gridrow="5" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FAC8FE3E2632" box="[166,430,1291,1351]" gridcol="0" gridrow="5" pageId="10" pageNumber="147">Snout length (%AAA distance)</th>
<td id="76B05DBDFFBE16C0FDDEFAC8FCF02632" box="[589,864,1291,1351]" gridcol="1" gridrow="5" pageId="10" pageNumber="147">19.2–34.7% (26.4%)</td>
<td id="76B05DBDFFBE16C0FC6FFAC8FB402632" box="[1020,1232,1291,1351]" gridcol="2" gridrow="5" pageId="10" pageNumber="147">36.7–44.8% (40.9%)</td>
</tr>
<tr id="356134C1FFBE16C0FF35FA94FB4026E6" box="[166,1232,1367,1427]" gridrow="6" pageId="10" pageNumber="147">
<th id="76B05DBDFFBE16C0FF35FA94FE3E26E6" box="[166,430,1367,1427]" gridcol="0" gridrow="6" pageId="10" pageNumber="147">Pre-anus length (%pre-anal-fin length)</th>
<td id="76B05DBDFFBE16C0FDDEFA94FCF026E6" box="[589,864,1367,1427]" gridcol="1" gridrow="6" pageId="10" pageNumber="147">75.0–82.0% (79.3%)*</td>
<td id="76B05DBDFFBE16C0FC6FFA94FB4026E6" box="[1020,1232,1367,1427]" gridcol="2" gridrow="6" pageId="10" pageNumber="147">84.4–88.8% (86.1%)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEE3683FFBEE935FF04FA62FD4226CC" blockId="10.[151,722,1441,1465]" box="[151,722,1441,1465]" pageId="10" pageNumber="147">
<tableNote id="76B7370DFFBEE935FF04FA62FD4226CC" box="[151,722,1441,1465]" pageId="10" pageNumber="147" targetBox="[166,1232,1090,1427]" targetPageId="10">*excluding data for specimens less than 15.0 mm SL.</tableNote>
</paragraph>
<paragraph id="8BEE3683FFBEE935FF54F903FCB92433" blockId="10.[151,1437,1511,2042]" pageId="10" pageNumber="147">
<bibRefCitation id="EFC04B72FFBEE935FF54F903FE4B25AC" author="Ito, T. &amp; Okumura, D." box="[199,475,1727,1754]" pageId="10" pageNumber="147" pagination="1 - 7" refId="ref7959" refString="Ito, T. &amp; Okumura, D. (2021) First records of Luciogobius parvulus and Luciogobius sp. 11 sensu Shibukawa et al., 2019 (Teleostei: Gobiidae) from Tokushima Prefecture, Japan. Ichthy, Natural History of Fishes of Japan, 12, 1 - 7. [in Japanese] https: // doi. org / 10.34583 / ichthy. 12.0 _ 1" type="journal article" year="2021">Ito and Okumura (2021)</bibRefCitation>
reported a single specimen (
<quantity id="4CA99B66FFBEE935FCB2F97CFC1725AF" box="[801,903,1727,1754]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="10" pageNumber="147" unit="mm" value="20.1">20.1 mm</quantity>
SL) of 
<taxonomicName id="4C514D00FFBEE935FC73F97CFB0325AF" box="[992,1171,1727,1754]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBEE935FC73F97CFBFA25AC" box="[992,1130,1727,1753]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp.
</taxonomicName>
11 
<emphasis id="B925EA91FFBEE935FB2EF903FB6B25AC" box="[1213,1275,1728,1753]" italics="true" pageId="10" pageNumber="147">sensu</emphasis>
<bibRefCitation id="EFC04B72FFBEE935FA97F97CFF6D2588" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." pageId="10" pageNumber="147" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFBEE935FA1BF903FF3D2588" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2019
</bibRefCitation>
from 
<collectingRegion id="4995F861FFBEE935FED5F920FDD22588" box="[326,578,1763,1789]" country="Japan" name="Tokushima" pageId="10" pageNumber="147">Tokushima Prefecture</collectingRegion>
, Shikoku, 
<collectingCountry id="F3467613FFBEE935FD2DF927FC91258B" box="[702,769,1764,1790]" name="Japan" pageId="10" pageNumber="147">Japan</collectingCountry>
. However, based on their description and photograph, the specimen is most likely to have been 
<taxonomicName id="4C514D00FFBEE935FDA9F8CBFD662454" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[570,758,1799,1825]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FDA9F8CBFD662454" box="[570,758,1799,1825]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
, having a relatively greater AAA distance (twice body depth at anus) and relatively shorter snout length (25.4% as HL).
</paragraph>
<paragraph id="8BEE3683FFBEE935FF54F88CFAD024A0" blockId="10.[151,1437,1511,2042]" pageId="10" pageNumber="147">
<taxonomicName id="4C514D00FFBEE935FF54F88CFE00241C" box="[199,400,1871,1898]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undefined-3">
<emphasis id="B925EA91FFBEE935FF54F88CFEC1241C" box="[199,337,1871,1897]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp. 3
</taxonomicName>
<emphasis id="B925EA91FFBEE935FE0AF893FE47241C" box="[409,471,1872,1897]" italics="true" pageId="10" pageNumber="147">sensu</emphasis>
<bibRefCitation id="EFC04B72FFBEE935FE4CF88CFD25241C" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." box="[479,693,1871,1898]" pageId="10" pageNumber="147" pagination="162 - 174" refId="ref8331" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55 (2), 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda 
<emphasis id="B925EA91FFBEE935FDA7F893FDF7241C" box="[564,615,1871,1897]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2008
</bibRefCitation>
is also similar to 
<taxonomicName id="4C514D00FFBEE935FC11F893FBD0241C" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[898,1088,1871,1897]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBEE935FC11F893FBD0241C" box="[898,1088,1871,1897]" italics="true" pageId="10" pageNumber="147">L. punctilineatus</emphasis>
</taxonomicName>
in most meristics, pectoral-fin membrane structure, and coloration (
<bibRefCitation id="EFC04B72FFBEE935FDA5F8B0FC9E24F8" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." box="[566,782,1907,1934]" pageId="10" pageNumber="147" pagination="162 - 174" refId="ref8331" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55 (2), 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda 
<emphasis id="B925EA91FFBEE935FD1FF8B7FD2D24F8" box="[652,701,1907,1933]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2008
</bibRefCitation>
; 
<bibRefCitation id="EFC04B72FFBEE935FC89F8B0FC2224F8" author="Maeda, K." box="[794,946,1907,1933]" pageId="10" pageNumber="147" pagination="1250 - 1251" refId="ref8280" refString="Maeda, K. (2014) Luciogobius sp. 2. In: Okiyama, M. (Ed.), An Atlas of Early Stage Fishes in Japan. Second Edition. Tokai University Press, Hadano, pp. 1250 - 1251. [in Japanese]" type="book chapter" year="2014">Maeda, 2014</bibRefCitation>
). However, the latter is easily distinguished from 
<taxonomicName id="4C514D00FFBEE935FF45F854FE0F24C4" box="[214,415,1943,1970]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undefined-3">
<emphasis id="B925EA91FFBEE935FF45F854FEF024C4" box="[214,352,1943,1969]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp. 3
</taxonomicName>
by vertebral number (16–18 + 22–24 = 39–42 vs. 21–23 + 19–21 = 41–43; 
<bibRefCitation id="EFC04B72FFBEE935FA94F854FF4224A0" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." pageId="10" pageNumber="147" pagination="162 - 174" refId="ref8331" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55 (2), 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda 
<emphasis id="B925EA91FFBEE935FACFF85BFA1E24C4" box="[1372,1422,1943,1969]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2008
</bibRefCitation>
; 
<bibRefCitation id="EFC04B72FFBEE935FF4DF878FEE224A0" author="Maeda, K." box="[222,370,1979,2005]" pageId="10" pageNumber="147" pagination="1250 - 1251" refId="ref8280" refString="Maeda, K. (2014) Luciogobius sp. 2. In: Okiyama, M. (Ed.), An Atlas of Early Stage Fishes in Japan. Second Edition. Tokai University Press, Hadano, pp. 1250 - 1251. [in Japanese]" type="book chapter" year="2014">Maeda, 2014</bibRefCitation>
) and greater AAA distance (about twice that of 
<taxonomicName id="4C514D00FFBEE935FC1AF878FBC224A0" box="[905,1106,1979,2006]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="species" species="undefined-3">
<emphasis id="B925EA91FFBEE935FC1AF878FB8324A0" box="[905,1043,1979,2005]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
sp. 3
</taxonomicName>
; 
<bibRefCitation id="EFC04B72FFBEE935FBCCF878FAA524A0" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." box="[1119,1333,1979,2006]" pageId="10" pageNumber="147" pagination="162 - 174" refId="ref8331" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55 (2), 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda 
<emphasis id="B925EA91FFBEE935FB21F87FFB7324A0" box="[1202,1251,1979,2005]" italics="true" pageId="10" pageNumber="147">et al</emphasis>
., 2008
</bibRefCitation>
).
</paragraph>
<paragraph id="8BEE3683FFBEE934FF54F81CFD362210" blockId="10.[151,1437,1511,2042]" lastBlockId="11.[151,1437,151,357]" lastPageId="11" lastPageNumber="148" pageId="10" pageNumber="147">
The 
<emphasis id="B925EA91FFBEE935FF68F823FE99248C" box="[251,265,2016,2041]" italics="true" pageId="10" pageNumber="147">p</emphasis>
-distances among the four species of 
<taxonomicName id="4C514D00FFBEE935FD3CF81CFCA9248C" authorityName="Gill" authorityYear="1859" box="[687,825,2015,2041]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="genus">
<emphasis id="B925EA91FFBEE935FD3CF81CFCA9248C" box="[687,825,2015,2041]" italics="true" pageId="10" pageNumber="147">Luciogobius</emphasis>
</taxonomicName>
and one species of 
<taxonomicName id="4C514D00FFBEE935FB88F823FBD0248C" authorityName="Snyder" authorityYear="1909" box="[1051,1088,2016,2041]" class="Actinopterygii" family="Gobiidae" genus="Inu" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="147" phylum="Chordata" rank="genus">
<emphasis id="B925EA91FFBEE935FB88F823FBD0248C" box="[1051,1088,2016,2041]" italics="true" pageId="10" pageNumber="147">Inu</emphasis>
</taxonomicName>
, estimated from the partial sequences of the 12S rRNA gene, showed that differences between 
<taxonomicName id="4C514D00FFBFE934FCFEFF54FBB923C4" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[877,1065,151,177]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBFE934FCFEFF54FBB923C4" box="[877,1065,151,177]" italics="true" pageId="11" pageNumber="148">L. punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBFE934FBBCFF54FBFD23C4" bold="true" box="[1071,1133,151,177]" pageId="11" pageNumber="148">
<taxonomicNameLabel id="A21657EAFFBFE934FBBCFF54FBFD23C4" box="[1071,1133,151,177]" pageId="11" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C514D00FFBFE934FB31FF54FA9B23C4" authorityName="Snyder" authorityYear="1909" box="[1186,1291,151,177]" class="Actinopterygii" family="Gobiidae" genus="Inu" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="koma">
<emphasis id="B925EA91FFBFE934FB31FF54FA9B23C4" box="[1186,1291,151,177]" italics="true" pageId="11" pageNumber="148">Inu koma</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C514D00FFBFE934FA8BFF54FA0523C4" authorityName="Di Caporiacco" authorityYear="1948" box="[1304,1429,151,177]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="martellii">
<emphasis id="B925EA91FFBFE934FA8BFF54FA0523C4" box="[1304,1429,151,177]" italics="true" pageId="11" pageNumber="148">L. martellii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C514D00FFBFE934FF04FF78FEB223A0" authorityName="Regan" authorityYear="1905" box="[151,290,187,213]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B925EA91FFBFE934FF04FF78FEB223A0" box="[151,290,187,213]" italics="true" pageId="11" pageNumber="148">L. elongatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C514D00FFBFE934FECDFF78FD8023A0" box="[350,528,187,213]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FECDFF78FE7823A0" box="[350,488,187,213]" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11 were 17.34–18.28%, 5.50–6.32%, 3.91–5.53%, and 1.53–3.90%, respectively (
<tableCitation id="C6D30338FFBFE934FF0CFF1CFF68238C" box="[159,248,223,249]" captionStart="TABLE 4" captionStartId="11.[151,239,1257,1281]" captionTargetPageId="11" captionText="TABLE 4. Genetic distances for 133 base pairs of the mitochondrial 12S rRNA gene between Luciogobius punctilineatus n. sp., Luciogobius sp. 11, L. elongatus, and L. martellii." httpUri="http://table.plazi.org/id/DF2E660BFFBFE934FF04FB2AFD652651" pageId="11" pageNumber="148" tableUuid="DF2E660BFFBFE934FF04FB2AFD652651">Table 4</tableCitation>
). 
<taxonomicName id="4C514D00FFBFE934FE9EFF1CFDAF238C" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[269,575,223,249]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBFE934FE9EFF1CFDAF238C" box="[269,575,223,249]" italics="true" pageId="11" pageNumber="148">Luciogobius punctilineatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C514D00FFBFE934FDEBFF1CFCBB238C" box="[632,811,223,249]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FDEBFF1CFC92238C" box="[632,770,223,249]" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11 were recovered in different groups, although it was supported by a low bootstrap value (
<figureCitation id="136A2A06FFBFE934FDDBFEC0FD00226B" box="[584,656,259,286]" captionStart="FIGURE 6" captionStartId="9.[151,250,805,829]" captionTargetBox="[159,1428,186,776]" captionTargetId="figure-17@9.[154,1433,181,781]" captionTargetPageId="9" captionText="FIGURE 6. Neighbor-joining tree based on 12S ribosomal RNA gene sequences (133 bp) of species of Luciogobius with Inu koma as outgroup. Numbers near branches indicate bootstrap values." figureDoi="http://doi.org/10.5281/zenodo.6556894" httpUri="https://zenodo.org/record/6556894/files/figure.png" pageId="11" pageNumber="148">Fig. 6</figureCitation>
). Although the molecular differences among 
<taxonomicName id="4C514D00FFBFE934FB30FEC0FAC92268" box="[1187,1369,259,285]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatu">
<emphasis id="B925EA91FFBFE934FB30FEC0FAC92268" box="[1187,1369,259,285]" italics="true" pageId="11" pageNumber="148">L. punctilineatu</emphasis>
</taxonomicName>
s, 
<taxonomicName id="4C514D00FFBFE934FAE6FEC0FEBC2234" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FAE6FEC0FE942234" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11, and 
<taxonomicName id="4C514D00FFBFE934FE1FFEE4FD872234" authorityName="Regan" authorityYear="1905" box="[396,535,295,321]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B925EA91FFBFE934FE1FFEE4FD872234" box="[396,535,295,321]" italics="true" pageId="11" pageNumber="148">L. elongatus</emphasis>
</taxonomicName>
were less clear, the species are differentiated from each other by AAA distance, pectoral- and pelvic-fin shapes, and coloration.
</paragraph>
<caption id="DF2E660BFFBFE934FF04FBBEFE1327CC" ID-DOI="http://doi.org/10.5281/zenodo.6556903" ID-Zenodo-Dep="6556903" httpUri="https://zenodo.org/record/6556903/files/figure.png" pageId="11" pageNumber="148" startId="11.[151,250,1149,1173]" targetBox="[151,1437,381,1125]" targetPageId="11">
<paragraph id="8BEE3683FFBFE934FF04FBBEFE1327CC" blockId="11.[151,1436,1149,1209]" pageId="11" pageNumber="148">
<emphasis id="B925EA91FFBFE934FF04FBBEFE8227E0" bold="true" box="[151,274,1149,1173]" pageId="11" pageNumber="148">FIGURE 8.</emphasis>
Fresh (upper and middle) and preserved (lower) specimens of 
<taxonomicName id="4C514D00FFBFE934FC19FBBEFBBB27E1" box="[906,1067,1149,1173]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FC19FBBEFB9727E0" box="[906,1031,1149,1173]" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (KAUM–I. 159027 25.9 mm SL) from Kochi Prefecture.
</paragraph>
</caption>
<caption id="DF2E660BFFBFE934FF04FB2AFD652651" ID-Table-UUID="DF2E660BFFBFE934FF04FB2AFD652651" httpUri="http://table.plazi.org/id/DF2E660BFFBFE934FF04FB2AFD652651" pageId="11" pageNumber="148" startId="11.[151,239,1257,1281]" targetBox="[166,1343,1333,1559]" targetIsTable="true" targetPageId="11">
<paragraph id="8BEE3683FFBFE934FF04FB2AFD652651" blockId="11.[151,1436,1257,1317]" pageId="11" pageNumber="148">
<emphasis id="B925EA91FFBFE934FF04FB2AFE992674" bold="true" box="[151,265,1257,1281]" pageId="11" pageNumber="148">TABLE 4.</emphasis>
Genetic distances for 133 base pairs of the mitochondrial 12S rRNA gene between 
<taxonomicName id="4C514D00FFBFE934FBE8FB2AFA0C2674" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[1147,1436,1257,1281]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFBFE934FBE8FB2AFA0C2674" box="[1147,1436,1257,1281]" italics="true" pageId="11" pageNumber="148">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFBFE934FF04FACEFF432650" bold="true" box="[151,211,1293,1317]" pageId="11" pageNumber="148">
<taxonomicNameLabel id="A21657EAFFBFE934FF04FACEFF432650" box="[151,211,1293,1317]" pageId="11" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C514D00FFBFE934FF72FACEFE1A2651" box="[225,394,1293,1317]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FF72FACEFEF52650" box="[225,357,1293,1317]" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11, 
<taxonomicName id="4C514D00FFBFE934FE24FACDFDAB2650" authorityName="Regan" authorityYear="1905" box="[439,571,1293,1317]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B925EA91FFBFE934FE24FACDFDAB2650" box="[439,571,1293,1317]" italics="true" pageId="11" pageNumber="148">L. elongatus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C514D00FFBFE934FDE5FACDFD7E2650" authorityName="Di Caporiacco" authorityYear="1948" box="[630,750,1293,1317]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="martellii">
<emphasis id="B925EA91FFBFE934FDE5FACDFD7E2650" box="[630,750,1293,1317]" italics="true" pageId="11" pageNumber="148">L. martellii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEE3683FFBFE934FD61FAF6FAAF2562" pageId="11" pageNumber="148">
<table id="F951C423FFBF16C0FF35FAF6FAAF2562" box="[166,1343,1333,1559]" gridcols="6" gridrows="6" pageId="11" pageNumber="148">
<tr id="356134C1FFBF16C0FF35FAF6FAAF2638" box="[166,1343,1333,1357]" gridrow="0" pageId="11" pageNumber="148" rowspan-0="1" rowspan-1="1">
<th id="76B05DBDFFBF16C0FD61FAF6FCC82638" box="[754,856,1333,1357]" gridcol="2" gridrow="0" pageId="11" pageNumber="148">1</th>
<th id="76B05DBDFFBF16C0FC0AFAF6FB872638" box="[921,1047,1333,1357]" gridcol="3" gridrow="0" pageId="11" pageNumber="148">2</th>
<th id="76B05DBDFFBF16C0FBCCFAF6FB082638" box="[1119,1176,1333,1357]" gridcol="4" gridrow="0" pageId="11" pageNumber="148">3</th>
<th id="76B05DBDFFBF16C0FA95FAF6FAAF2638" box="[1286,1343,1333,1357]" gridcol="5" gridrow="0" pageId="11" pageNumber="148">4</th>
</tr>
<tr id="356134C1FFBF16C0FF35FA9EFAAF2603" box="[166,1343,1373,1398]" gridrow="1" pageId="11" pageNumber="148" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76B05DBDFFBF16C0FF35FA9EFF232603" box="[166,179,1373,1398]" gridcol="0" gridrow="1" pageId="11" pageNumber="148">1</th>
<td id="76B05DBDFFBF16C0FE97FA9EFDF32603" box="[260,611,1373,1398]" gridcol="1" gridrow="1" pageId="11" pageNumber="148">
<taxonomicName id="4C514D00FFBFE934FE97FA9EFD882600" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[260,536,1373,1397]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBFE934FE97FA9EFD882600" box="[260,536,1373,1397]" italics="true" pageId="11" pageNumber="148">Luciogobius punctilineatus</emphasis>
</taxonomicName>
(
<emphasis id="B925EA91FFBFE934FDB5FA9DFDA32600" box="[550,563,1374,1397]" italics="true" pageId="11" pageNumber="148">n</emphasis>
= 7)
</td>
<td id="76B05DBDFFBF16C0FD61FA9EFCC82603" box="[754,856,1373,1398]" gridcol="2" gridrow="1" pageId="11" pageNumber="148">0–1.53</td>
</tr>
<tr id="356134C1FFBF16C0FF35FA46FAAF26EB" box="[166,1343,1413,1438]" gridrow="2" pageId="11" pageNumber="148" rowspan-4="1" rowspan-5="1">
<th id="76B05DBDFFBF16C0FF35FA46FF2326EB" box="[166,179,1413,1438]" gridcol="0" gridrow="2" pageId="11" pageNumber="148">2</th>
<td id="76B05DBDFFBF16C0FE97FA46FDF326EB" box="[260,611,1413,1438]" gridcol="1" gridrow="2" pageId="11" pageNumber="148">
<taxonomicName id="4C514D00FFBFE934FE97FA45FE3526E8" box="[260,421,1414,1438]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFBFE934FE97FA45FE1126EB" box="[260,385,1414,1438]" italics="true" pageId="11" pageNumber="148">Luciogobius</emphasis>
sp.
</taxonomicName>
11 (
<emphasis id="B925EA91FFBFE934FE41FA45FE4F26E8" box="[466,479,1414,1437]" italics="true" pageId="11" pageNumber="148">n</emphasis>
= 4)
</td>
<td id="76B05DBDFFBF16C0FD61FA46FCC826EB" box="[754,856,1413,1438]" gridcol="2" gridrow="2" pageId="11" pageNumber="148">1.53–3.90</td>
<td id="76B05DBDFFBF16C0FC0AFA46FB8726EB" box="[921,1047,1413,1438]" gridcol="3" gridrow="2" pageId="11" pageNumber="148">0–0.76</td>
</tr>
<tr id="356134C1FFBF16C0FF35FA6DFAAF26B3" box="[166,1343,1454,1478]" gridrow="3" pageId="11" pageNumber="148" rowspan-5="1">
<th id="76B05DBDFFBF16C0FF35FA6DFF2326B3" box="[166,179,1454,1478]" gridcol="0" gridrow="3" pageId="11" pageNumber="148">3</th>
<td id="76B05DBDFFBF16C0FE97FA6DFDF326B3" box="[260,611,1454,1478]" gridcol="1" gridrow="3" pageId="11" pageNumber="148">
<taxonomicName id="4C514D00FFBFE934FE97FA6DFE7B26B3" authorityName="Regan" authorityYear="1905" box="[260,491,1454,1478]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="B925EA91FFBFE934FE97FA6DFE7B26B3" box="[260,491,1454,1478]" italics="true" pageId="11" pageNumber="148">Luciogobius elongatus</emphasis>
</taxonomicName>
(
<emphasis id="B925EA91FFBFE934FE6AFA6CFD9626B3" box="[505,518,1455,1478]" italics="true" pageId="11" pageNumber="148">n</emphasis>
= 1)
</td>
<td id="76B05DBDFFBF16C0FD61FA6DFCC826B3" box="[754,856,1454,1478]" gridcol="2" gridrow="3" pageId="11" pageNumber="148">3.91–5.53</td>
<td id="76B05DBDFFBF16C0FC0AFA6DFB8726B3" box="[921,1047,1454,1478]" gridcol="3" gridrow="3" pageId="11" pageNumber="148">2.33–3.13</td>
<td id="76B05DBDFFBF16C0FBCCFA6DFB0826B3" box="[1119,1176,1454,1478]" gridcol="4" gridrow="3" pageId="11" pageNumber="148">―</td>
</tr>
<tr id="356134C1FFBF16C0FF35FA15FAAF269B" box="[166,1343,1494,1518]" gridrow="4" pageId="11" pageNumber="148">
<th id="76B05DBDFFBF16C0FF35FA15FF23269B" box="[166,179,1494,1518]" gridcol="0" gridrow="4" pageId="11" pageNumber="148">4</th>
<td id="76B05DBDFFBF16C0FE97FA15FDF3269B" box="[260,611,1494,1518]" gridcol="1" gridrow="4" pageId="11" pageNumber="148">
<taxonomicName id="4C514D00FFBFE934FE97FA15FE4F269B" authorityName="Di Caporiacco" authorityYear="1948" box="[260,479,1494,1518]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="martellii">
<emphasis id="B925EA91FFBFE934FE97FA15FE4F269B" box="[260,479,1494,1518]" italics="true" pageId="11" pageNumber="148">Luciogobius martellii</emphasis>
</taxonomicName>
(
<emphasis id="B925EA91FFBFE934FE7DFA14FE6B269B" box="[494,507,1495,1518]" italics="true" pageId="11" pageNumber="148">n</emphasis>
= 1)
</td>
<td id="76B05DBDFFBF16C0FD61FA15FCC8269B" box="[754,856,1494,1518]" gridcol="2" gridrow="4" pageId="11" pageNumber="148">5.50–6.32</td>
<td id="76B05DBDFFBF16C0FC0AFA15FB87269B" box="[921,1047,1494,1518]" gridcol="3" gridrow="4" pageId="11" pageNumber="148">4.69–5.51</td>
<td id="76B05DBDFFBF16C0FBCCFA15FB08269B" box="[1119,1176,1494,1518]" gridcol="4" gridrow="4" pageId="11" pageNumber="148">5.51</td>
<td id="76B05DBDFFBF16C0FA95FA15FAAF269B" box="[1286,1343,1494,1518]" gridcol="5" gridrow="4" pageId="11" pageNumber="148">―</td>
</tr>
<tr id="356134C1FFBF16C0FF35FA3DFAAF2562" box="[166,1343,1534,1559]" gridrow="5" pageId="11" pageNumber="148">
<th id="76B05DBDFFBF16C0FF35FA3DFF232562" box="[166,179,1534,1559]" gridcol="0" gridrow="5" pageId="11" pageNumber="148">5</th>
<td id="76B05DBDFFBF16C0FE97FA3DFDF32562" box="[260,611,1534,1559]" gridcol="1" gridrow="5" pageId="11" pageNumber="148">
<taxonomicName id="4C514D00FFBFE934FE97FA3CFEF32563" authorityName="Snyder" authorityYear="1909" box="[260,355,1534,1558]" class="Actinopterygii" family="Gobiidae" genus="Inu" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="koma">
<emphasis id="B925EA91FFBFE934FE97FA3CFEF32563" box="[260,355,1534,1558]" italics="true" pageId="11" pageNumber="148">Inu koma</emphasis>
</taxonomicName>
(
<emphasis id="B925EA91FFBFE934FEE1FA3CFE062563" box="[370,406,1535,1558]" italics="true" pageId="11" pageNumber="148">n =</emphasis>
1)
</td>
<td id="76B05DBDFFBF16C0FD61FA3DFCC82562" box="[754,856,1534,1559]" gridcol="2" gridrow="5" pageId="11" pageNumber="148">17.3–18.3</td>
<td id="76B05DBDFFBF16C0FC0AFA3DFB872562" box="[921,1047,1534,1559]" gridcol="3" gridrow="5" pageId="11" pageNumber="148">17.34–18.37</td>
<td id="76B05DBDFFBF16C0FBCCFA3DFB082562" box="[1119,1176,1534,1559]" gridcol="4" gridrow="5" pageId="11" pageNumber="148">19.42</td>
<td id="76B05DBDFFBF16C0FA95FA3DFAAF2562" box="[1286,1343,1534,1559]" gridcol="5" gridrow="5" pageId="11" pageNumber="148">19.17</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEE3683FFBFE934FF54F985FD8424D1" blockId="11.[151,1437,1606,1957]" pageId="11" pageNumber="148">
<emphasis id="B925EA91FFBFE934FF54F985FE5F2515" bold="true" box="[199,463,1606,1632]" pageId="11" pageNumber="148">Geographic variation.</emphasis>
Geographic variations in some meristics and coloration were apparent in 
<taxonomicName id="4C514D00FFBFE934FA82F984FF4225F1" authorityName="Koreeda &amp; Motomura" authorityYear="2022" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBFE934FA82F984FF4225F1" italics="true" pageId="11" pageNumber="148">L. punctilineatus</emphasis>
</taxonomicName>
, the dorsal-, anal-, and pectoral-fin rays in specimens from Amami-oshima island tending to be fewer than in specimens from other localities (
<tableCitation id="C6D30338FFBFE934FD91F94DFDC925DD" box="[514,601,1678,1704]" captionStart="TABLE 2" captionStartId="10.[151,239,151,175]" captionTargetBox="[151,1411,263,1001]" captionTargetPageId="10" captionText="TABLE 2. Frequency distribution of dorsal-, anal-, and pectoral-fin ray counts, and insertion of first anal-fin pterygiophore relative to haemal spines in Luciogobius punctilineatus n. sp. and Luciogobius sp. 11. Gray shaded values include data from Shibukawa et al. (2019). *Including holotype of L. punctilineatus." httpUri="http://table.plazi.org/id/DF2E660BFFBEE935FF04FF54FC582382" pageId="11" pageNumber="148" tableUuid="DF2E660BFFBEE935FF04FF54FC582382">Table 2</tableCitation>
). Similarly, vertebral numbers in specimens from Shikoku (
<collectingRegion id="4995F861FFBFE934FB7CF94DFAFD25DD" box="[1263,1389,1678,1704]" country="Japan" name="Tokushima" pageId="11" pageNumber="148">Tokushima</collectingRegion>
and 
<collectingRegion id="4995F861FFBFE934FF04F971FF4B25B9" box="[151,219,1714,1740]" country="Japan" name="Kochi" pageId="11" pageNumber="148">Kochi</collectingRegion>
prefectures) tended to be lower than in specimens from other localities [16–17 (mainly 17) + 22–23 (22) = 
<quantity id="4CA99B66FFBFE934FF04F915FF402585" box="[151,208,1750,1776]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="11" pageNumber="148" unit="in" value="39.0">39 in</quantity>
specimens from Shikoku (
<bibRefCitation id="EFC04B72FFBFE934FE6FF915FC952585" author="Ito, T. &amp; Okumura, D." box="[508,773,1750,1776]" pageId="11" pageNumber="148" pagination="1 - 7" refId="ref7959" refString="Ito, T. &amp; Okumura, D. (2021) First records of Luciogobius parvulus and Luciogobius sp. 11 sensu Shibukawa et al., 2019 (Teleostei: Gobiidae) from Tokushima Prefecture, Japan. Ichthy, Natural History of Fishes of Japan, 12, 1 - 7. [in Japanese] https: // doi. org / 10.34583 / ichthy. 12.0 _ 1" type="journal article" year="2021">Ito and Okumura, 2021</bibRefCitation>
; this study) vs. 17–18 + 22–24 (23) = 40–42 (40–41) in the latter]. Specimens from 
<collectingRegion id="4995F861FFBFE934FE34F939FDF62461" box="[423,614,1786,1812]" country="Japan" name="Kochi" pageId="11" pageNumber="148">Kochi Prefecture</collectingRegion>
and the Osumi Peninsula (
<figureCitation id="136A2A06FFBFE934FC06F939FB892460" box="[917,1049,1786,1813]" captionStart="FIGURE 1" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[151,1436,179,1785]" captionTargetId="figure-17@3.[151,1436,179,1785]" captionTargetPageId="3" captionText="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="11" pageNumber="148">Figs. 1D, E</figureCitation>
, 
<figureCitation id="136A2A06FFBFE934FBB6F939FBD82461" box="[1061,1096,1786,1812]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[178,1410,1041,1953]" captionTargetId="figure-197@6.[178,1410,1035,1953]" captionTargetPageId="6" captionText="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." figureDoi="http://doi.org/10.5281/zenodo.6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="11" pageNumber="148">3C</figureCitation>
, 
<figureCitation id="136A2A06FFBFE934FBC7F939FB072460" box="[1108,1175,1786,1813]" captionStart="FIGURE 4" captionStartId="8.[151,250,1281,1305]" captionTargetBox="[180,1408,181,1257]" captionTargetId="figure-30@8.[180,1408,181,1257]" captionTargetPageId="8" captionText="FIGURE 4. Preserved specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.6556890" httpUri="https://zenodo.org/record/6556890/files/figure.png" pageId="11" pageNumber="148">4D, E</figureCitation>
) had a relatively larger black caudal-fin blotch with a narrower yellowish margin, compared with specimens from other localities (
<figureCitation id="136A2A06FFBFE934FAF7F8DDFF4C2429" captionStart="FIGURE 1" captionStartId="3.[151,250,1810,1834]" captionTargetBox="[151,1436,179,1785]" captionTargetId="figure-17@3.[151,1436,179,1785]" captionTargetPageId="3" captionText="FIGURE 1. Fresh specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL). Dorsal and lateral views taken before and immediately after fixation, respectively." figureDoi="http://doi.org/10.5281/zenodo.6556882" httpUri="https://zenodo.org/record/6556882/files/figure.png" pageId="11" pageNumber="148">Figs. 1A–C</figureCitation>
, 
<figureCitation id="136A2A06FFBFE934FF79F881FEA22428" box="[234,306,1858,1885]" captionStart="FIGURE 3" captionStartId="6.[151,250,1976,2000]" captionTargetBox="[178,1410,1041,1953]" captionTargetId="figure-197@6.[178,1410,1035,1953]" captionTargetPageId="6" captionText="FIGURE 3. Live individuals of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: two specimens of KAUM–I. 158052–158055, B: KAUM–I. 156165) and Osumi Peninsula (C: KAUM–I. 157125, D: KAUM–I. 157124)." figureDoi="http://doi.org/10.5281/zenodo.6556886" httpUri="https://zenodo.org/record/6556886/files/figure.png" pageId="11" pageNumber="148">3A, B</figureCitation>
, 
<figureCitation id="136A2A06FFBFE934FED3F881FE182429" box="[320,392,1858,1884]" captionStart="FIGURE 4" captionStartId="8.[151,250,1281,1305]" captionTargetBox="[180,1408,181,1257]" captionTargetId="figure-30@8.[180,1408,181,1257]" captionTargetPageId="8" captionText="FIGURE 4. Preserved specimens of Luciogobius punctilineatus n. sp. from Satsuma Peninsula (A: Holotype; KAUM–I. 156161. 29.8 mm SL), Amami-oshima island (B: KAUM–I. 145567, 28.1 mm SL, C: KAUM–I. 145562, 12.1 mm SL), Osumi Peninsula (D: KAUM–I. 157125, 28.7 mm SL), and Kochi Prefecture, Shikoku (E: KAUM–I. 159035, 22.9 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.6556890" httpUri="https://zenodo.org/record/6556890/files/figure.png" pageId="11" pageNumber="148">4A–C</figureCitation>
). Although these minor differences are regarded here as representing geographic variations within 
<taxonomicName id="4C514D00FFBFE934FF77F8A4FE3024F5" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[228,416,1894,1920]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="148" phylum="Chordata" rank="species" species="punctilineatus">
<emphasis id="B925EA91FFBFE934FF77F8A4FE3024F5" box="[228,416,1894,1920]" italics="true" pageId="11" pageNumber="148">L. punctilineatus</emphasis>
</taxonomicName>
, specimens from Shikoku, Osumi Peninsula, and Amami-oshima island were excluded from the 
<typeStatus id="54EA8821FFBFE934FF53F848FF6024D0" box="[192,240,1931,1957]" pageId="11" pageNumber="148">type</typeStatus>
series of the new species.
</paragraph>
<caption id="DF2E660BFFB8E933FF04F8D6FC762497" ID-DOI="http://doi.org/10.5281/zenodo.6556909" ID-Zenodo-Dep="6556909" httpUri="https://zenodo.org/record/6556909/files/figure.png" pageId="12" pageNumber="149" startId="12.[151,250,1813,1837]" targetBox="[241,1346,191,1781]" targetPageId="12">
<paragraph id="8BEE3683FFB8E933FF04F8D6FC762497" blockId="12.[151,1437,1813,2018]" pageId="12" pageNumber="149">
<emphasis id="B925EA91FFB8E933FF04F8D6FE832458" bold="true" box="[151,275,1813,1837]" pageId="12" pageNumber="149">FIGURE 9.</emphasis>
Relationships of (A) snout length, (B) AAA distance (measured between posterior margin of anus and anal-fin origin), (C) PAL (pre-anus length), and (D) pre-anal-fin length as % of standard length, and (E) snout length as % of AAA, and (F) PAL as % of pre-anal-fin length vs. standard length (mm) in 
<taxonomicName id="4C514D00FFB8E933FC90F89EFB872400" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[771,1047,1885,1909]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB8E933FC90F89EFB872400" box="[771,1047,1885,1909]" italics="true" pageId="12" pageNumber="149">Luciogobius punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB8E933FB8EF89DFBC72403" bold="true" box="[1053,1111,1886,1910]" pageId="12" pageNumber="149">
<taxonomicNameLabel id="A21657EAFFB8E933FB8EF89DFBC72403" box="[1053,1111,1886,1910]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C514D00FFB8E933FB1AF89EFABA2400" box="[1161,1322,1885,1909]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB8E933FB1AF89EFA962400" box="[1161,1286,1885,1909]" italics="true" pageId="12" pageNumber="149">Luciogobius</emphasis>
sp.
</taxonomicName>
11. Closed red star, holotype of 
<taxonomicName id="4C514D00FFB8E933FEE2F841FD8E24EC" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[369,542,1921,1945]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB8E933FEE2F841FD8E24EC" box="[369,542,1921,1945]" italics="true" pageId="12" pageNumber="149">L. punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB8E933FDB5F841FDF224EF" bold="true" box="[550,610,1922,1946]" pageId="12" pageNumber="149">
<taxonomicNameLabel id="A21657EAFFB8E933FDB5F841FDF224EF" box="[550,610,1922,1946]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; open red stars, paratypes of 
<taxonomicName id="4C514D00FFB8E933FC0BF841FBD524EC" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[920,1093,1921,1945]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB8E933FC0BF841FBD524EC" box="[920,1093,1921,1945]" italics="true" pageId="12" pageNumber="149">L. punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB8E933FBDEF841FB1824EF" bold="true" box="[1101,1160,1922,1946]" pageId="12" pageNumber="149">
<taxonomicNameLabel id="A21657EAFFB8E933FBDEF841FB1824EF" box="[1101,1160,1922,1946]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Shimokoshiki-shima island, Tanega-shima island; yellow stars, 
<taxonomicName id="4C514D00FFB8E933FDD5F865FD6124C8" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[582,753,1957,1981]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB8E933FDD5F865FD6124C8" box="[582,753,1957,1981]" italics="true" pageId="12" pageNumber="149">L. punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB8E933FD64F865FCA024CB" bold="true" box="[759,816,1958,1982]" pageId="12" pageNumber="149">
<taxonomicNameLabel id="A21657EAFFB8E933FD64F865FCA024CB" box="[759,816,1958,1982]" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Amami-oshima island; green stars; 
<taxonomicName id="4C514D00FFB8E933FB44F865FA1224C8" authority="Koreeda &amp; Motomura, 2022" authorityName="Koreeda &amp; Motomura" authorityYear="2022" box="[1239,1410,1957,1981]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="punctilineatus" status="sp. nov.">
<emphasis id="B925EA91FFB8E933FB44F865FA1224C8" box="[1239,1410,1957,1981]" italics="true" pageId="12" pageNumber="149">L. punctilineatus</emphasis>
</taxonomicName>
<emphasis id="B925EA91FFB8E933FA1BF865FF262497" bold="true" pageId="12" pageNumber="149">
<taxonomicNameLabel id="A21657EAFFB8E933FA1BF865FF262497" pageId="12" pageNumber="149" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Kochi Prefecture and Osumi Peninsula; blue triangles, 
<taxonomicName id="4C514D00FFB8E933FCB2F80AFC522494" box="[801,962,1993,2017]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="149" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB8E933FCB2F80AFC0E2494" box="[801,926,1993,2017]" italics="true" pageId="12" pageNumber="149">Luciogobius</emphasis>
sp.
</taxonomicName>
11.
</paragraph>
</caption>
<paragraph id="8BEE3683FFB9E932FF54FF55FEBF2234" blockId="13.[151,1436,150,321]" pageId="13" pageNumber="150">
<emphasis id="B925EA91FFB9E932FF54FF55FDC323C5" bold="true" box="[199,595,150,176]" pageId="13" pageNumber="150">Comparative material examined.</emphasis>
<taxonomicName id="4C514D00FFB9E932FDC9FF54FC9C23C4" box="[602,780,151,177]" class="Actinopterygii" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="150" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B925EA91FFB9E932FDC9FF54FD7423C4" box="[602,740,151,177]" italics="true" pageId="13" pageNumber="150">Luciogobius</emphasis>
sp.
</taxonomicName>
11 
<emphasis id="B925EA91FFB9E932FCA5FF54FCE423C5" box="[822,884,151,176]" italics="true" pageId="13" pageNumber="150">sensu</emphasis>
<bibRefCitation id="EFC04B72FFB9E932FCE8FF54FB1223C4" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[891,1154,151,177]" pageId="13" pageNumber="150" pagination="29 - 96" refId="ref8715" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [in Japanese]" type="journal article" year="2019">
Shibukawa 
<emphasis id="B925EA91FFB9E932FC6CFF54FBA123C4" box="[1023,1073,151,177]" italics="true" pageId="13" pageNumber="150">et al</emphasis>
., 2019
</bibRefCitation>
: 
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<collectionCode id="ED40AE46FFB9E932FB1EFF54FB7623C4" box="[1165,1254,151,177]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="13" pageNumber="150">KAUM</collectionCode>
–I. 142153, 
<quantity id="4CA99B66FFB9E932FAF8FF54FF5323A0" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="13" pageNumber="150" unit="mm" value="29.3">29.3 mm</quantity>
SL, 
<location id="8E8E6058FFB9E932FF6CFF78FEDD23A0" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB9E932FF6CFF78FEDD23A0" box="[255,333,187,213]" country="Japan" municipality="Yaizu" name="Iifuchi" pageId="13" pageNumber="150" stateProvince="Shizuoka">Iifuchi</location>
, 
<collectingMunicipality id="6B8AACF9FFB9E932FEC9FF78FE0D23A0" box="[346,413,187,213]" pageId="13" pageNumber="150">Yaizu</collectingMunicipality>
, 
<collectingRegion id="4995F861FFB9E932FE3FFF78FD8723A0" box="[428,535,187,213]" country="Japan" name="Shizuoka" pageId="13" pageNumber="150">Shizuoka</collectingRegion>
, 
<collectingCountry id="F3467613FFB9E932FDB6FF78FDF823A0" box="[549,616,187,213]" name="Japan" pageId="13" pageNumber="150">Japan</collectingCountry>
, coll. by 
<collectorName id="26A45355FFB9E932FD4BFF78FCE023A0" box="[728,880,187,213]" pageId="13" pageNumber="150">S. Yamashita</collectorName>
, 
<date id="FFEF1043FFB9E932FCEDFF78FB8123A0" box="[894,1041,187,213]" pageId="13" pageNumber="150" value="2020-03-09">
<collectingDate id="EFABE9ABFFB9E932FCEDFF78FB8123A0" box="[894,1041,187,213]" pageId="13" pageNumber="150" value="2020-03-09">9 Mar. 2020</collectingDate>
</date>
</materialsCitation>
; KAUM–I. 145507, 
<quantity id="4CA99B66FFB9E932FA90FF78FAFB23A0" box="[1283,1387,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="13" pageNumber="150" unit="mm" value="26.4">26.4 mm</quantity>
SL, KAUM–I. 145508, 
<quantity id="4CA99B66FFB9E932FEE7FF1CFE47238C" box="[372,471,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="13" pageNumber="150" unit="mm" value="28.6">28.6 mm</quantity>
SL, 
<materialsCitation id="3B393CDEFFB9E932FD9FFF1CFBA3238C" ID-GBIF-Occurrence="3771630311" box="[524,1075,222,249]" collectingDate="2020-03-09" collectionCode="KAUM" collectorName="S. Yamashita" country="Japan" location="Iifuchi" municipality="Yaizu" pageId="13" pageNumber="150" specimenCount="1" stateProvince="Shizuoka">
<collectionCode id="ED40AE46FFB9E932FD9FFF1CFDF5238C" box="[524,613,223,249]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="13" pageNumber="150">KAUM</collectionCode>
–I. 145509, 
<quantity id="4CA99B66FFB9E932FD7BFF1CFCDC238C" box="[744,844,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="13" pageNumber="150" unit="mm" value="28.3">28.3 mm</quantity>
SL, same locality of
</materialsCitation>
<materialsCitation id="3B393CDEFFB9E932FBA9FF1CFEC22268" ID-GBIF-Occurrence="3771630312" collectingDate="2020-03-10" collectionCode="KAUM" country="Japan" location="Yamashita" pageId="13" pageNumber="150" specimenCount="1" stateProvince="Shizuoka">
<collectionCode id="ED40AE46FFB9E932FBA9FF1CFB03238C" box="[1082,1171,223,249]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="13" pageNumber="150">KAUM</collectionCode>
–I. 142153, S. 
<location id="8E8E6058FFB9E932FAA1FF1CFF3E2268" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB9E932FAA1FF1CFF3E2268" country="Japan" name="Yamashita" pageId="13" pageNumber="150" stateProvince="Shizuoka">Yamashita</location>
, 
<date id="FFEF1043FFB9E932FF2AFEC0FEC22268" box="[185,338,259,285]" pageId="13" pageNumber="150" value="2020-03-10">
<collectingDate id="EFABE9ABFFB9E932FF2AFEC0FEC22268" box="[185,338,259,285]" pageId="13" pageNumber="150" value="2020-03-10">10 Mar. 2020</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B393CDEFFB9E932FECFFEC0FEBB2234" ID-GBIF-Occurrence="3771630304" collectingDate="2021-07-13" collectionCode="KAUM" collectorName="H. Saito" country="India" location="Niyodo River" municipality="Haruno" pageId="13" pageNumber="150" specimenCount="1">
<collectionCode id="ED40AE46FFB9E932FECFFEC0FE252268" box="[348,437,259,285]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="13" pageNumber="150">KAUM</collectionCode>
–I. 159027, 
<quantity id="4CA99B66FFB9E932FDABFEC1FD0B2268" box="[568,667,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="13" pageNumber="150" unit="mm" value="25.9">25.9 mm</quantity>
SL, mouth of 
<location id="8E8E6058FFB9E932FCA8FEC0FC452268" LSID="urn:lsid:plazi:treatment:03F88795FFB5E932FF04FA1AFEBF2234:8E8E6058FFB9E932FCA8FEC0FC452268" box="[827,981,259,285]" country="India" municipality="Haruno" name="Niyodo River" pageId="13" pageNumber="150">Niyodo River</location>
, 
<collectingMunicipality id="6B8AACF9FFB9E932FC73FEC0FBA72268" box="[992,1079,259,285]" pageId="13" pageNumber="150">Haruno</collectingMunicipality>
, 
<collectingCountry id="F3467613FFB9E932FBD2FEC0FB182268" box="[1089,1160,259,285]" name="India" pageId="13" pageNumber="150">Kochi</collectingCountry>
, 
<collectingCountry id="F3467613FFB9E932FB00FEC0FB462268" box="[1171,1238,259,285]" name="Japan" pageId="13" pageNumber="150">Japan</collectingCountry>
, coll. by 
<collectorName id="26A45355FFB9E932FAA9FEC0FA082268" box="[1338,1432,259,285]" pageId="13" pageNumber="150">H. Saito</collectorName>
, 
<date id="FFEF1043FFB9E932FF04FEE4FEBB2234" box="[151,299,295,321]" pageId="13" pageNumber="150" value="2021-07-13">
<collectingDate id="EFABE9ABFFB9E932FF04FEE4FEBB2234" box="[151,299,295,321]" pageId="13" pageNumber="150" value="2021-07-13">13 July 2021</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>