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<mods:title id="9CFFC56767B9D6C17BC43C3777D73164">A new species of Uropeltis Cuvier, 1829 (Serpentes: Uropeltidae) from the eastern escarpment of the southern Western Ghats, India</mods:title>
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<mods:affiliation id="401A2DD1DBFA022FE58DDC08A06B4B7F">Centre for Research in Emerging Tropical Diseases, Department of Zoology, University of Calicut, Thenhipalam, Kerala, 673635, India.</mods:affiliation>
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<mods:namePart id="4C887EC459AE52F4B155DB30C96A2CBD">Deepak, V.</mods:namePart>
<mods:affiliation id="07BB7067B9CC575648B3C7ABD9DE1086">Natural History Museum, London SW &amp; School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne NE 1 7 RU, UK.</mods:affiliation>
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<mods:affiliation id="C25502C6D0B0E569830F4223FB5321AB">Bd, Uk. &amp; Wildlife Institute of India, Chandrabani, P. O. 18, Dehradun, India.</mods:affiliation>
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<mods:namePart id="4F50CA92053F85924F0B00A6A79B1DC0">Narayanan, Surya</mods:namePart>
<mods:affiliation id="2A34CFB76FA48D0420C87795DCD9A1D5">Natural History Museum, London SW &amp; SM Sehgal Centre for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka- 560064, India. &amp; Natural History Museum, London SW</mods:affiliation>
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<taxonomicName id="4C514D190B3FFFF1CAD7E3BAFE06FF34" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[151,455,223,250]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B3FFFF1CAD7E3BAFE06FF34" bold="true" box="[151,455,223,250]" italics="true" pageId="3" pageNumber="336">Uropeltis caudomaculata</emphasis>
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<figureCitation id="136A2A1F0B3FFFF1CAD7E242FEC3FE8F" box="[151,258,295,321]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[151,250,1641,1665]" captionStartId-1="5.[151,250,1942,1966]" captionStartId-2="6.[151,250,1104,1128]" captionStartId-3="7.[151,250,1983,2007]" captionStartId-4="8.[151,250,1943,1967]" captionStartId-5="10.[151,250,1922,1946]" captionTargetBox-0="[155,1431,454,1611]" captionTargetBox-1="[151,1435,181,1918]" captionTargetBox-2="[159,1436,181,1073]" captionTargetBox-3="[151,1435,309,1959]" captionTargetBox-4="[179,1408,181,1919]" captionTargetBox-5="[159,1428,1031,1891]" captionTargetId-0="figure-111@4.[151,1435,450,1615]" captionTargetId-1="figure-17@5.[151,1435,181,1918]" captionTargetId-2="figure-18@6.[151,1436,181,1081]" captionTargetId-3="figure-46@7.[151,1435,309,1959]" captionTargetId-4="figure-18@8.[179,1408,181,1919]" captionTargetId-5="figure-414@10.[151,1436,1029,1899]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="10" captionText-0="FIGURE 1. Maximum Likelihood (ML) phylogeny for selected samples of Uropeltis. Numbers on internal branches are Bayesian posterior probability/ML bootstrap support values greater than 0.9/60. See Table 1 for data sources." captionText-1="FIGURE 2. Holotype of Uropeltis caudomaculata sp. nov. (BMNH 1955.1.2.69). Scale bar increments in millimetres. Photographs by Kevin Webb." captionText-2="FIGURE 3. Map indicating localities for Uropeltis pulneyensis and U. caudomaculata sp. nov. Locality symbols with a black dot inside are associated with voucher specimens; those symbols without a black dot indicate where specimens have been observed but not collected. The imprecise type locality for (the only known specimen of) U. guentheri comb. nov. is ca. 1,500 m, possibly in or close to Meghamalai (“Cumbum Hills”)." captionText-3="FIGURE 4. The three most-recently collected paratypes of Uropeltis caudomaculata sp. nov. in approximately dorsal (left column) and ventral (right column) views. A &amp; B, BNHS 3765, C &amp; D, BNHS 3763, and E &amp; F, BNHS 3764." captionText-4="FIGURE 5. Uropeltis caudomaculata sp. nov. in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S." captionText-5="FIGURE 6. Variation in numbers of ventral and subcaudal shields in females (circles) and males (triangles) of Uropeltis pulneyensis (25 females, 19 males), U. caudomaculata sp. nov. (6 females, 2 males) and the holotype of U. guentheri comb. nov. (one male). See Tables 3 and 4 for sources of data." figureDoi-0="http://doi.org/10.5281/zenodo.12732827" figureDoi-1="http://doi.org/10.5281/zenodo.12732831" figureDoi-2="http://doi.org/10.5281/zenodo.12732833" figureDoi-3="http://doi.org/10.5281/zenodo.12732839" figureDoi-4="http://doi.org/10.5281/zenodo.12732841" figureDoi-5="http://doi.org/10.5281/zenodo.12732845" httpUri-0="https://zenodo.org/record/12732827/files/figure.png" httpUri-1="https://zenodo.org/record/12732831/files/figure.png" httpUri-2="https://zenodo.org/record/12732833/files/figure.png" httpUri-3="https://zenodo.org/record/12732839/files/figure.png" httpUri-4="https://zenodo.org/record/12732841/files/figure.png" httpUri-5="https://zenodo.org/record/12732845/files/figure.png" pageId="3" pageNumber="336">Figs. 16</figureCitation>
,
<tableCitation id="C6D303210B3FFFF1CB4DE242FE49FE8F" box="[269,392,295,321]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="2.[151,239,151,175]" captionStartId-1="2.[151,239,1253,1277]" captionStartId-2="11.[151,239,467,493]" captionStartId-3="13.[151,244,150,176]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="11" captionTargetPageId-3="13" captionText-0="TABLE 1. Sequences (with GenBank accession codes) of mitochondrial genes for species of the Indian endemic genus Uropeltis used in molecular phylogenetic analyses. Sequences newly reported in this study highlighted in bold. See Sampaio et al. (2022) for further details. Samples are listed in the order in which they appear (top to bottom) in Fig. 1. States abbreviated as KL (Kerala) and TN (Tamil Nadu)." captionText-1="TABLE 2. Uncorrectedp-distances (%) for intra-andinterspecific comparisonsfor selectedUropeltisfor two mitochondrial genes (16s, nd4). Values are means, with ranges in parentheses, and number of pairwise comparisons in square brackets." captionText-2="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." captionText-3="TABLE 4. Sex, total length (TL) and tail length (tL) (in mm) and scalation data for 42 referred specimens of Uropeltis pulneyensis. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). Blank cells indicate data not recorded. See Appendix 1 for specimen localities." httpUri-0="http://table.plazi.org/id/DF2E66120B3EFFF0CAD7E3F2FD34FED5" httpUri-1="http://table.plazi.org/id/DF2E66120B3EFFF0CAD7E780FA5BFAEF" httpUri-2="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" httpUri-3="http://table.plazi.org/id/DF2E66120B31FFFFCAD7E3F3FB62FF37" pageId="3" pageNumber="336" tableUuid-0="DF2E66120B3EFFF0CAD7E3F2FD34FED5" tableUuid-1="DF2E66120B3EFFF0CAD7E780FA5BFAEF" tableUuid-2="DF2E66120B37FFF9CAD7E2B6F884FDDF" tableUuid-3="DF2E66120B31FFFFCAD7E3F3FB62FF37">Tables 14</tableCitation>
, Appendix 2
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<emphasis id="B925EA880B3FFFF1CAD7E20AFEF7FE47" bold="true" box="[151,310,367,393]" pageId="3" pageNumber="336">Chresonymy</emphasis>
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<taxonomicName id="4C514D190B3FFFF1CAD7E2D5FE42FE06" baseAuthorityName="Beddome" baseAuthorityYear="1877" box="[151,387,432,456]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="dindigalensis">
<emphasis id="B925EA880B3FFFF1CAD7E2D5FE42FE06" box="[151,387,432,456]" italics="true" pageId="3" pageNumber="336">Uropeltis dindigalensis</emphasis>
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:
<bibRefCitation id="EFC04B6B0B3FFFF1CBD1E2D5FD9EFE06" author="Ganesh, S. R. &amp; Asokan, S. &amp; Kannan, P." box="[401,607,432,456]" pageId="3" pageNumber="336" pagination="1 - 9" refId="ref11149" refString="Ganesh, S. R., Asokan, S. &amp; Kannan, P. (2008) A preliminary survey of amphibians and reptiles in Cardamom Hills, Western Ghats, Tamil Nadu. Cobra, 2 (2), 1 - 9." type="journal article" year="2008">
Ganesh
<emphasis id="B925EA880B3FFFF1CBA3E2D5FDD7FE06" box="[483,534,432,456]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
(2008)
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<taxonomicName id="4C514D190B3FFFF1CAD7E2ABFE65FE28" baseAuthorityName="Beddome" baseAuthorityYear="1877" box="[151,420,462,486]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="dindigalensis">
<emphasis id="B925EA880B3FFFF1CAD7E2ABFF37FE28" box="[151,246,462,486]" italics="true" pageId="3" pageNumber="336">Uropeltis</emphasis>
cf.
<emphasis id="B925EA880B3FFFF1CB5DE2ABFE65FE28" box="[285,420,462,486]" italics="true" pageId="3" pageNumber="336">dindigalensis</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFC04B6B0B3FFFF1CBF1E2ABFD3BFE28" author="Chandramouli, S. R. &amp; Ganesh, S. R." box="[433,762,462,486]" pageId="3" pageNumber="336" pagination="72 - 85" refId="ref10808" refString="Chandramouli, S. R. &amp; Ganesh, S. R. (2010) Herpetofauna of the Southern Western Ghats - reinvestigated after decades. Taprobanica, 2 (2), 72 - 85. https: // doi. org / 10.47605 / tapro. v 2 i 2.30" type="journal article" year="2010">Chandramouli &amp; Ganesh (2010)</bibRefCitation>
,
<bibRefCitation id="EFC04B6B0B3FFFF1C945E2ABFC15FE28" author="Ganesh, S. R. &amp; Bhupathy, S. &amp; David, P. &amp; Sathishkumar, N. &amp; Srinivas, G." box="[773,980,462,486]" pageId="3" pageNumber="336" pagination="53 - 64" refId="ref11196" refString="Ganesh, S. R., Bhupathy, S., David, P., Sathishkumar, N. &amp; Srinivas, G. (2014) Snake fauna of high wavy mountains, Western Ghats, India: species richness, status, and distribution pattern. Russian Journal of Herpetology, 21 (1), 53 - 64." type="journal article" year="2014">
Ganesh
<emphasis id="B925EA880B3FFFF1C917E2AAFC45FE28" box="[855,900,462,486]" italics="true" pageId="3" pageNumber="336">et al</emphasis>
. (2014)
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</paragraph>
<paragraph id="8BEE369A0B3FFFF1CAD7E288FF36FDEA" blockId="3.[151,1437,432,578]" pageId="3" pageNumber="336">
<taxonomicName id="4C514D190B3FFFF1CAD7E288FEB5FDCB" authority="(Beddome, 1863)" box="[151,372,493,517]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B3FFFF1CAD7E288FEB5FDCB" box="[151,372,493,517]" italics="true" pageId="3" pageNumber="336">Uropeltis pulneyensis</emphasis>
</taxonomicName>
:
<bibRefCitation id="EFC04B6B0B3FFFF1CBC2E288FDB6FDCB" author="Hutton, A. F. &amp; David, P." box="[386,631,493,517]" pageId="3" pageNumber="336" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David (2008)</bibRefCitation>
,
<bibRefCitation id="EFC04B6B0B3FFFF1C8C3E288FCB3FDCB" author="Chaitanya, R. &amp; Khandekar, A. &amp; Caleb, D. &amp; Mukherjee, N. &amp; Ghosh, A. &amp; Giri, V. B." box="[643,882,493,517]" pageId="3" pageNumber="336" pagination="21 - 37" refId="ref10737" refString="Chaitanya, R., Khandekar, A., Caleb, D., Mukherjee, N., Ghosh, A. &amp; Giri, V. B. (2018) Herpetofauna of the Meghamalai Wildlife Sanctuary, Southern Western Ghats, India: An updated checklist with annotations on taxonomy and nomenclature. Journal of the Bombay Natural History Society, 115, 21 - 37." type="journal article" year="2018">
Chaitanya
<emphasis id="B925EA880B3FFFF1C8B1E28BFCE4FDCB" box="[753,805,493,517]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
(2018)
</bibRefCitation>
,
<bibRefCitation id="EFC04B6B0B3FFFF1C93EE288FB8DFDCB" author="Deepak, V. &amp; Narayanan, S. &amp; Das, S. &amp; Rajkumar, K. P. &amp; Pal, S. &amp; Gerard, J. D. &amp; Gower, D. J." box="[894,1100,493,517]" pageId="3" pageNumber="336" refId="ref10961" refString="Deepak, V., Narayanan, S., Das, S., Rajkumar, K. P., Pal, S., Gerard, J. D. &amp; Gower, D. J. (2022) Snakes and lizards of the sky islands of the Western Ghats. Ashoka Trust for Research in Ecology and the Environment, Bengaluru, 114 pp." type="book" year="2022">
Deepak
<emphasis id="B925EA880B3FFFF1C993E28BFBC6FDCB" box="[979,1031,493,517]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
(2022
</bibRefCitation>
, in part),
<bibRefCitation id="EFC04B6B0B3FFFF1CEF0E288FABCFDCB" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." box="[1200,1405,493,517]" pageId="3" pageNumber="336" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B3FFFF1CF44E28BFAF8FDCB" box="[1284,1337,493,517]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
(2023
</bibRefCitation>
, in part)
</paragraph>
<paragraph id="8BEE369A0B3FFFF1CAD7E14FFDAEFD8C" blockId="3.[151,1437,432,578]" box="[151,623,554,578]" pageId="3" pageNumber="336">
<taxonomicName id="4C514D190B3FFFF1CAD7E14FFE52FD8C" authority="(Beddome, 1863)" box="[151,403,554,578]" class="Reptilia" family="Uropeltidae" genus="Uropeltis" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B3FFFF1CAD7E14FFF37FD8C" box="[151,246,554,578]" italics="true" pageId="3" pageNumber="336">Uropeltis</emphasis>
cf.
<emphasis id="B925EA880B3FFFF1CB5DE14FFE52FD8C" box="[285,403,554,578]" italics="true" pageId="3" pageNumber="336">pulneyensis</emphasis>
</taxonomicName>
: Ganesh
<emphasis id="B925EA880B3FFFF1CBB3E14EFDE7FD8C" box="[499,550,554,578]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
(2022)
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B3FFFF1CAD7E114FCBCFC64" pageId="3" pageNumber="336" type="materials_examined">
<paragraph id="8BEE369A0B3FFFF1CAD7E114FC2DFD39" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">
<materialsCitation id="3B393CC70B3FFFF1CAD7E114FC29FD39" collectionCode="BMNH" collectorName="Angus F. Hutton" country="India" elevation="1219" location="High Wavy Mountains" pageId="3" pageNumber="336" specimenCode="BMNH 1955.1.2.69" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis id="B925EA880B3FFFF1CAD7E114FEC5FD45" bold="true" box="[151,260,625,651]" pageId="3" pageNumber="336">
<typeStatus id="54EA88380B3FFFF1CAD7E114FEC5FD45" box="[151,260,625,651]" pageId="3" pageNumber="336" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="136A2A1F0B3FFFF1CB5AE115FEA2FD45" box="[282,355,624,651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1942,1966]" captionTargetBox="[151,1435,181,1918]" captionTargetId="figure-17@5.[151,1435,181,1918]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Uropeltis caudomaculata sp. nov. (BMNH 1955.1.2.69). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732831" httpUri="https://zenodo.org/record/12732831/files/figure.png" pageId="3" pageNumber="336">Fig. 2</figureCitation>
).
<specimenCode id="DBF79EE10B3FFFF1CBC0E114FDA8FD45" box="[384,617,624,651]" pageId="3" pageNumber="336">BMNH 1955.1.2.69</specimenCode>
,
<specimenCount id="9D57FD130B3FFFF1C83BE115FD0AFD44" box="[635,715,624,650]" count="1" pageId="3" pageNumber="336" type="female">female</specimenCount>
. Collected from “High Wavys” (=
<location id="8E8E60410B3FFFF1CE35E115FABEFD44" LSID="urn:lsid:plazi:treatment:03F8878C0B3FFFE0CAD7E3BAFCEAF9E3:8E8E60410B3FFFF1CE35E115FABEFD44" box="[1141,1407,624,651]" country="India" name="High Wavy Mountains" pageId="3" pageNumber="336" stateProvince="Tamil Nadu">High Wavy Mountains</location>
= Meghamalai),
<collectingRegion id="4995F8780B3FFFF1CB7EE1F1FE0BFD60" box="[318,458,660,686]" country="India" name="Tamil Nadu" pageId="3" pageNumber="336">Tamil Nadu</collectingRegion>
,
<collectingCountry id="F346760A0B3FFFF1CB9AE1F1FDD5FD60" box="[474,532,660,686]" name="India" pageId="3" pageNumber="336">India</collectingCountry>
(
<figureCitation id="136A2A1F0B3FFFF1C867E1F1FDAEFD61" box="[551,623,660,687]" captionStart="FIGURE 3" captionStartId="6.[151,250,1104,1128]" captionTargetBox="[159,1436,181,1073]" captionTargetId="figure-18@6.[151,1436,181,1081]" captionTargetPageId="6" captionText="FIGURE 3. Map indicating localities for Uropeltis pulneyensis and U. caudomaculata sp. nov. Locality symbols with a black dot inside are associated with voucher specimens; those symbols without a black dot indicate where specimens have been observed but not collected. The imprecise type locality for (the only known specimen of) U. guentheri comb. nov. is ca. 1,500 m, possibly in or close to Meghamalai (“Cumbum Hills”)." figureDoi="http://doi.org/10.5281/zenodo.12732833" httpUri="https://zenodo.org/record/12732833/files/figure.png" pageId="3" pageNumber="336">Fig. 3</figureCitation>
), 3,0005,000 feet elevation (
<quantity id="4CA99B7F0B3FFFF1C991E1F1FBADFD61" box="[977,1132,660,687]" metricMagnitude="3" metricUnit="m" metricValue="1.219" metricValueMax="1.524" metricValueMin="0.914" pageId="3" pageNumber="336" unit="m" value="1219.0" valueMax="1524.0" valueMin="914.0">
<elevation id="007CD1A90B3FFFF1C991E1F1FBADFD61" box="[977,1132,660,687]" metricMagnitude="3" metricUnit="m" metricValue="1.219" metricValueMax="1.524" metricValueMin="0.914" pageId="3" pageNumber="336" unit="m" value="1219.0" valueMax="1524.0" valueMin="914.0">9141,524 m</elevation>
</quantity>
) by
<collectorName id="26A4534C0B3FFFF1CEE5E1F0FAAAFD61" box="[1189,1387,661,687]" pageId="3" pageNumber="336">Angus F. Hutton</collectorName>
. No collection date recorded in catalogue, but specimen accessioned into the London collection on
<date id="FFEF105A0B3FFFF1CEF7E1DCFAE2FD1D" box="[1207,1315,697,723]" pageId="3" pageNumber="336">2 January</date>
, 1955, and was probably collected between 1946 and 1948 (see
<bibRefCitation id="EFC04B6B0B3FFFF1C8A6E1B8FC21FD39" author="Hutton, A. F. &amp; David, P." box="[742,992,732,759]" pageId="3" pageNumber="336" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David 2008</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE369A0B3FFFF1CA87E064FCBCFC64" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">
<emphasis id="B925EA880B3FFFF1CA87E064FEFEFCD5" bold="true" box="[199,319,769,795]" pageId="3" pageNumber="336">
<typeStatus id="54EA88380B3FFFF1CA87E064FEFEFCD5" box="[199,319,769,795]" pageId="3" pageNumber="336" type="paratype">Paratypes</typeStatus>
</emphasis>
(n = 7).
<collectionCode id="ED40AE5F0B3FFFF1CBF0E064FDC5FCD5" box="[432,516,769,795]" pageId="3" pageNumber="336">BMNH</collectionCode>
1955.1.2.68 (female), 1955.1.2.70 (male), 1955.1.2.71 (male), and 1955.1.2.72 (female), same data as for
<typeStatus id="54EA88380B3FFFF1CB8BE041FDF0FCF0" box="[459,561,804,830]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
. BNHS 3765 (
<figureCitation id="136A2A1F0B3FFFF1C89BE041FCEDFCF1" box="[731,812,804,831]" captionStart="FIGURE 4" captionStartId="7.[151,250,1983,2007]" captionTargetBox="[151,1435,309,1959]" captionTargetId="figure-46@7.[151,1435,309,1959]" captionTargetPageId="7" captionText="FIGURE 4. The three most-recently collected paratypes of Uropeltis caudomaculata sp. nov. in approximately dorsal (left column) and ventral (right column) views. A &amp; B, BNHS 3765, C &amp; D, BNHS 3763, and E &amp; F, BNHS 3764." figureDoi="http://doi.org/10.5281/zenodo.12732839" httpUri="https://zenodo.org/record/12732839/files/figure.png" pageId="3" pageNumber="336">Figs. 4</figureCitation>
,
<figureCitation id="136A2A1F0B3FFFF1C97AE041FC88FCF1" box="[826,841,804,831]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Uropeltis caudomaculata sp. nov. in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S." figureDoi="http://doi.org/10.5281/zenodo.12732841" httpUri="https://zenodo.org/record/12732841/files/figure.png" pageId="3" pageNumber="336">5</figureCitation>
), female, collected from Manalar, Meghmalai Tiger Reserve,
<collectingRegion id="4995F8780B3FFFF1CABFE02DFE45FCAC" box="[255,388,840,866]" country="India" name="Tamil Nadu" pageId="3" pageNumber="336">Tamil Nadu</collectingRegion>
(
<geoCoordinate id="EE65505D0B3FFFF1CBD3E02DFDE6FCAD" box="[403,551,840,867]" degrees="9.621255" direction="north" orientation="latitude" pageId="3" pageNumber="336" precision="1" value="9.621255">9.621255° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B3FFFF1C872E02CFD0EFCAD" box="[562,719,841,867]" degrees="77.340202" direction="east" orientation="longitude" pageId="3" pageNumber="336" precision="1" value="77.3402">77.340202° E</geoCoordinate>
,
<quantity id="4CA99B7F0B3FFFF1C89BE02DFCF1FCAD" box="[731,816,840,867]" metricMagnitude="3" metricUnit="m" metricValue="1.541" pageId="3" pageNumber="336" unit="m" value="1541.0">1,541m</quantity>
asl) by J.D. Gerard on
<date id="FFEF105A0B3FFFF1CE77E02CFB16FCAD" box="[1079,1239,840,867]" pageId="3" pageNumber="336" value="2021-07-24">
24
<superScript id="7C249BD20B3FFFF1CE13E02DFBA1FC98" attach="left" box="[1107,1120,840,854]" fontSize="6" pageId="3" pageNumber="336">th</superScript>
July 2021
</date>
. BNHS 3763 and BNHS 3764 (
<figureCitation id="136A2A1F0B3FFFF1CB6FE009FEB3FC49" box="[303,370,876,903]" captionStart="FIGURE 4" captionStartId="7.[151,250,1983,2007]" captionTargetBox="[151,1435,309,1959]" captionTargetId="figure-46@7.[151,1435,309,1959]" captionTargetPageId="7" captionText="FIGURE 4. The three most-recently collected paratypes of Uropeltis caudomaculata sp. nov. in approximately dorsal (left column) and ventral (right column) views. A &amp; B, BNHS 3765, C &amp; D, BNHS 3763, and E &amp; F, BNHS 3764." figureDoi="http://doi.org/10.5281/zenodo.12732839" httpUri="https://zenodo.org/record/12732839/files/figure.png" pageId="3" pageNumber="336">Fig. 4</figureCitation>
), female, collected from Yellapetty, Munnar,
<collectingRegion id="4995F8780B3FFFF1C929E009FC75FC48" box="[873,948,876,902]" country="India" name="Kerala" pageId="3" pageNumber="336">Kerala</collectingRegion>
(
<geoCoordinate id="EE65505D0B3FFFF1C981E009FBA0FC49" box="[961,1121,876,903]" degrees="10.119513" direction="north" orientation="latitude" pageId="3" pageNumber="336" precision="1" value="10.119513">10.119513° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B3FFFF1CE2CE009FAC6FC49" box="[1132,1287,876,903]" degrees="77.206169" direction="east" orientation="longitude" pageId="3" pageNumber="336" precision="1" value="77.20617">77.206169° E</geoCoordinate>
,
<quantity id="4CA99B7F0B3FFFF1CF53E009FAAFFC49" box="[1299,1390,876,903]" metricMagnitude="3" metricUnit="m" metricValue="1.952" pageId="3" pageNumber="336" unit="m" value="1952.0">1,952 m</quantity>
asl) by S. Das on
<date id="FFEF105A0B3FFFF1CB6DE0F4FE32FC65" box="[301,499,912,939]" pageId="3" pageNumber="336" value="2022-02-07">
7
<superScript id="7C249BD20B3FFFF1CB7BE0F5FE89FC50" attach="left" box="[315,328,912,926]" fontSize="6" pageId="3" pageNumber="336">th</superScript>
February 2022
</date>
and
<date id="FFEF105A0B3FFFF1C86AE0F5FD25FC65" box="[554,740,912,939]" pageId="3" pageNumber="336" value="2023-03-15">
15
<superScript id="7C249BD20B3FFFF1C805E0F5FD93FC50" attach="left" box="[581,594,912,926]" fontSize="6" pageId="3" pageNumber="336">th</superScript>
March 2023
</date>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B3FFFF1CA87E0D1FE1EFB21" pageId="3" pageNumber="336" type="diagnosis">
<paragraph id="8BEE369A0B3FFFF1CA87E0D1FE1EFB21" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">
<emphasis id="B925EA880B3FFFF1CA87E0D1FE83FC00" bold="true" box="[199,322,948,974]" pageId="3" pageNumber="336">Diagnosis.</emphasis>
<taxonomicName id="4C514D190B3FFFF1CB08E0D1FDA4FC00" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[328,613,948,974]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B3FFFF1CB08E0D1FDA4FC00" box="[328,613,948,974]" italics="true" pageId="3" pageNumber="336">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B3FFFF1C82CE0D0FD04FC01" bold="true" box="[620,709,949,975]" pageId="3" pageNumber="336">
<taxonomicNameLabel id="A21657F30B3FFFF1C82CE0D0FD04FC01" box="[620,709,949,975]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other congeners except
<taxonomicName id="4C514D190B3FFFF1CEC6E0D0FAEAFC00" box="[1158,1323,948,974]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B3FFFF1CEC6E0D0FAEAFC00" box="[1158,1323,948,974]" italics="true" pageId="3" pageNumber="336">U. pulneyensis</emphasis>
</taxonomicName>
and some individuals of
<taxonomicName id="4C514D190B3FFFF1CB7BE0BCFE0EFC3C" box="[315,463,984,1010]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="bhupathyi">
<emphasis id="B925EA880B3FFFF1CB7BE0BCFE0EFC3C" box="[315,463,984,1010]" italics="true" pageId="3" pageNumber="336">U. bhupathyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D190B3FFFF1C847E0BCFD40FC3C" box="[519,641,984,1010]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B925EA880B3FFFF1C847E0BCFD40FC3C" box="[519,641,984,1010]" italics="true" pageId="3" pageNumber="336">U. grandis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC04B6B0B3FFFF1C8FEE0BDFCACFC3D" author="Jins, V. J. &amp; Sampaio, F. L. &amp; Gower, D. J." box="[702,877,984,1011]" pageId="3" pageNumber="336" pagination="401 - 422" refId="ref11605" refString="Jins, V. J., Sampaio, F. L. &amp; Gower, D. J. (2018) A new species of Uropeltis Cuvier, 1829 (Serpentes: Uropeltidae) from the Anaikatty Hills of the Western Ghats of India. Zootaxa, 4415 (3), 401 - 422. https: // doi. org / 10.11646 / zootaxa. 4415.3.1" type="journal article" year="2018">
Jins
<emphasis id="B925EA880B3FFFF1C8B2E0BCFCEAFC3C" box="[754,811,984,1010]" italics="true" pageId="3" pageNumber="336">et al.</emphasis>
2018
</bibRefCitation>
) in having a rostral scale that extends posteriorly to prevent midline contact between the opposite nasal scales. The new species differs from
<taxonomicName id="4C514D190B3FFFF1CEF9E098FAA0FBD8" box="[1209,1377,1020,1046]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B3FFFF1CEF9E098FAA0FBD8" box="[1209,1377,1020,1046]" italics="true" pageId="3" pageNumber="336">U. pulneyensis</emphasis>
</taxonomicName>
(and its current junior synonym
<taxonomicName id="4C514D190B3FFFF1CB91E745FCA2FBF5" authority="Beddome, 1878" authorityName="Beddome" authorityYear="1878" box="[465,867,1056,1083]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B3FFFF1CB91E745FD65FBF4" box="[465,676,1056,1082]" italics="true" pageId="3" pageNumber="336">Silybura guentheri</emphasis>
<bibRefCitation id="EFC04B6B0B3FFFF1C8EEE745FCA2FBF5" author="Beddome, R. H." box="[686,867,1056,1083]" pageId="3" pageNumber="336" pagination="154 - 155" refId="ref10438" refString="Beddome, R. H. (1878) Descriptions of new Uropeltidae from southern India, with remarks on some previously described species. Proceedings of the Zoological Society of London, 46, 154 - 155. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07946. x" type="journal article" year="1878">Beddome, 1878</bibRefCitation>
</taxonomicName>
) in having more ventral scales (among examined specimens, 185187 and 187197 versus 156175 and
<quantity id="4CA99B7F0B3FFFF1C94CE721FC4DFB90" box="[780,908,1092,1118]" metricMagnitude="0" metricUnit="m" metricValue="4.4196" metricValueMax="4.6228" metricValueMin="4.2164" pageId="3" pageNumber="336" unit="in" value="174.0" valueMax="182.0" valueMin="166.0">166182 in</quantity>
males and females, respectively:
<tableCitation id="C6D303210B3FFFF1CF4CE721FAAAFB91" box="[1292,1387,1092,1119]" captionStart="TABLE 3" captionStartId="11.[151,239,467,493]" captionTargetPageId="11" captionText="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="3" pageNumber="336" tableUuid="DF2E66120B37FFF9CAD7E2B6F884FDDF">Tables 3</tableCitation>
and
<tableCitation id="C6D303210B3FFFF1CAD7E70CFF69FB4D" box="[151,168,1129,1155]" captionStart="TABLE 4" captionStartId="13.[151,244,150,176]" captionTargetPageId="13" captionText="TABLE 4. Sex, total length (TL) and tail length (tL) (in mm) and scalation data for 42 referred specimens of Uropeltis pulneyensis. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). Blank cells indicate data not recorded. See Appendix 1 for specimen localities." httpUri="http://table.plazi.org/id/DF2E66120B31FFFFCAD7E3F3FB62FF37" pageId="3" pageNumber="336" tableUuid="DF2E66120B31FFFFCAD7E3F3FB62FF37">4</tableCitation>
).
<taxonomicName id="4C514D190B3FFFF1CAFCE70DFE18FB4C" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[188,473,1128,1154]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B3FFFF1CAFCE70DFE18FB4C" box="[188,473,1128,1154]" italics="true" pageId="3" pageNumber="336">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B3FFFF1CBA0E70CFDF8FB4D" bold="true" box="[480,569,1129,1155]" pageId="3" pageNumber="336">
<taxonomicNameLabel id="A21657F30B3FFFF1CBA0E70CFDF8FB4D" box="[480,569,1129,1155]" pageId="3" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C514D190B3FFFF1C88DE70CFC84FB4C" box="[717,837,1128,1154]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B925EA880B3FFFF1C88DE70CFC84FB4C" box="[717,837,1128,1154]" italics="true" pageId="3" pageNumber="336">U. grandis</emphasis>
</taxonomicName>
in having 17 versus 19 dorsal scale rows at midbody, and in having fewer versus more than 190 ventrals. The new species differs from
<taxonomicName id="4C514D190B3FFFF1CE5FE7E8FB70FB68" box="[1055,1201,1164,1190]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="3" pageNumber="336" phylum="Chordata" rank="species" species="bhupathyi">
<emphasis id="B925EA880B3FFFF1CE5FE7E8FB70FB68" box="[1055,1201,1164,1190]" italics="true" pageId="3" pageNumber="336">U. bhupathyi</emphasis>
</taxonomicName>
in having fewer than 190 versus more than 200 ventrals, and in having a much less distinct tail shield, corresponding to Smiths (1943) shield
<typeStatus id="54EA88380B3FFFF1CAA1E7B0FED8FB21" box="[225,281,1237,1263]" pageId="3" pageNumber="336">Type</typeStatus>
III versus
<typeStatus id="54EA88380B3FFFF1CBD0E7B0FE09FB21" box="[400,456,1237,1263]" pageId="3" pageNumber="336">Type</typeStatus>
I.
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B3FFFFBCA87E79DFF37F979" lastPageId="9" lastPageNumber="342" pageId="3" pageNumber="336" type="description">
<paragraph id="8BEE369A0B3FFFF1CA87E79DFC48FAB0" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">
<emphasis id="B925EA880B3FFFF1CA87E79DFE21FADD" bold="true" box="[199,480,1272,1299]" pageId="3" pageNumber="336">
Description of
<typeStatus id="54EA88380B3FFFF1CB39E79CFE21FADD" box="[377,480,1273,1299]" pageId="3" pageNumber="336" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="136A2A1F0B3FFFF1CBB1E79DFDF4FADD" box="[497,565,1272,1299]" captionStart="FIGURE 2" captionStartId="5.[151,250,1942,1966]" captionTargetBox="[151,1435,181,1918]" captionTargetId="figure-17@5.[151,1435,181,1918]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Uropeltis caudomaculata sp. nov. (BMNH 1955.1.2.69). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732831" httpUri="https://zenodo.org/record/12732831/files/figure.png" pageId="3" pageNumber="336">Fig. 2</figureCitation>
). See
<tableCitation id="C6D303210B3FFFF1C83DE79DFD10FADD" box="[637,721,1272,1299]" captionStart="TABLE 3" captionStartId="11.[151,239,467,493]" captionTargetPageId="11" captionText="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="3" pageNumber="336" tableUuid="DF2E66120B37FFF9CAD7E2B6F884FDDF">Table 3</tableCitation>
for morphometric and meristic data. Female with large (ca.
<quantity id="4CA99B7F0B3FFFF1CFC0E79CFF02FAF9" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="336" unit="mm" value="19.0">19 mm</quantity>
long), yolky ova in oviducts. Generally good condition; longitudinal incision into coelom on right of midventer, ca.
<quantity id="4CA99B7F0B3FFFF1CA82E624FED5FA95" box="[194,276,1345,1371]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="336" unit="mm" value="22.0">22 mm</quantity>
long, extending anteriorly from
<quantity id="4CA99B7F0B3FFFF1C8D4E625FD3AFA95" box="[660,763,1344,1371]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="336" unit="mm" value="12.5">12.5 mm</quantity>
anterior to vent. The specimen smells of wintergreen oil, sometimes added as a bitterant to surgical spirit or rubbing alcohol.
</paragraph>
<paragraph id="8BEE369A0B3FFFF1CA87E6EDFE2AF9C0" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">
Body subcylindrical, venter slightly flattened; tapering over c.
<quantity id="4CA99B7F0B3FFFF1C9CEE6ECFC20FA6D" box="[910,993,1417,1443]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="336" unit="mm" value="10.0">10 mm</quantity>
immediately behind head; very gently tapering posteriorly up to vent, more abruptly on tail. Head strongly tapering in dorsal view, sides straight to slightly convex, pointed; tapering also in lateral view, gently with straight edges to anterior margin of prefrontal then much more abruptly to tip of rostral.
</paragraph>
<paragraph id="8BEE369A0B3FFFF1CA87E57DFAA0F870" blockId="3.[151,1437,624,1983]" pageId="3" pageNumber="336">Rostral rounded anteriorly; longer than wide in dorsal view, about as long as deep in lateral view; at widest slightly in front of and below nostril. Rostral approximately twice as long (in dorsal view) than rostral-frontal gap. Frontal six-sided; lateralmost edges in contact with oculars strongly diverging anteriorly. Frontal much longer, wider than rostral. Paired nasals not in contact behind rostral. Left prefrontal overlaps right along midline. External naris small, subcircular, slightly countersunk within small depression, located in anteroventral corner of undivided nasal shield. Nasal shield subequal in length and height, contacts first and second supralabials (SLs). Prefrontal about as wide as long, much shorter than frontal, contacts SL2 and SL3. Supralabials four, SL1 smallest, making least contribution to margin of mouth; SL2 larger than SL1, smaller than SL3; SL4 much the largest and longest. Ocular contacts SL3 and SL4. Eye small but distinct, diameter approximately 0.4 times length of ocular shield, located near anteroventral corner of ocular; bulges very slightly from ocular surface, pupil appears subcircular; eyes directed antero- (and very slightly dorso-) laterally. Seven maxillary and eight mandibular teeth on each side. Teeth simple, pointed, distinctly retrorse, straight to slightly posteriorly curved distally; evenly spaced, smallest posteriorly.</paragraph>
<paragraph id="8BEE369A0B38FFF6CA87E3F2FD59FEAB" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="337">Paired parietals slightly shorter than frontal, posteriorly broadly rounded. Opposite parietals in brief midline contact, left overlapping right. Parietals much longer than wide, wider than frontal and rostral. Each parietal contacts four scales other than head shields. Infralabials (ILs) 3,3, IL2 slightly longer than IL1, more notably longer than IL3. Left and right first infralabials briefly in midline contact, separating small, slightly prominent mental from first midline ventral scale. First and second ventrals (in contact) and third longer than wide, fourth about as long as wide, fifth and subsequent ventrals wider than long.</paragraph>
<caption id="DF2E66120B38FFF6CAD7E50CFB28F96B" ID-DOI="http://doi.org/10.5281/zenodo.12732827" ID-Zenodo-Dep="12732827" httpUri="https://zenodo.org/record/12732827/files/figure.png" pageId="4" pageNumber="337" startId="4.[151,250,1641,1665]" targetBox="[155,1431,454,1611]" targetPageId="4" targetType="figure">
<paragraph id="8BEE369A0B38FFF6CAD7E50CFB28F96B" blockId="4.[151,1436,1641,1702]" pageId="4" pageNumber="337">
<emphasis id="B925EA880B38FFF6CAD7E50CFED6F94F" bold="true" box="[151,279,1641,1665]" pageId="4" pageNumber="337">FIGURE 1.</emphasis>
Maximum Likelihood (ML) phylogeny for selected samples of
<emphasis id="B925EA880B38FFF6C986E50FFBEDF94C" box="[966,1068,1642,1666]" italics="true" pageId="4" pageNumber="337">
<taxonomicName id="4C514D190B38FFF6C986E50FFBE9F94C" authorityName="Cuvier" authorityYear="1829" box="[966,1064,1642,1666]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="4" pageNumber="337" phylum="Chordata" rank="genus">Uropeltis</taxonomicName>
.
</emphasis>
Numbers on internal branches are Bayesian posterior probability/ML bootstrap support values greater than 0.9/60. See Table 1 for data sources.
</paragraph>
</caption>
<caption id="DF2E66120B39FFF7CAD7E4F3FE03F81C" ID-DOI="http://doi.org/10.5281/zenodo.12732831" ID-Zenodo-Dep="12732831" httpUri="https://zenodo.org/record/12732831/files/figure.png" pageId="5" pageNumber="338" startId="5.[151,250,1942,1966]" targetBox="[151,1435,181,1918]" targetPageId="5" targetType="figure">
<paragraph id="8BEE369A0B39FFF7CAD7E4F3FE03F81C" blockId="5.[151,1437,1942,2002]" pageId="5" pageNumber="338">
<emphasis id="B925EA880B39FFF7CAD7E4F3FED8F860" bold="true" box="[151,281,1942,1966]" pageId="5" pageNumber="338">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C514D190B39FFF7CBF0E4F3FD76F860" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[432,695,1942,1966]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="5" pageNumber="338" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B39FFF7CBF0E4F3FD76F860" box="[432,695,1942,1966]" italics="true" pageId="5" pageNumber="338">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B39FFF7C883E4F2FCD8F861" bold="true" box="[707,793,1943,1967]" pageId="5" pageNumber="338">
<taxonomicNameLabel id="A21657F30B39FFF7C883E4F2FCD8F861" box="[707,793,1943,1967]" pageId="5" pageNumber="338" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(BMNH 1955.1.2.69). Scale bar increments in millimetres. Photographs by Kevin Webb.
</paragraph>
</caption>
<caption id="DF2E66120B3AFFF4CAD7E735FD27FB1B" ID-DOI="http://doi.org/10.5281/zenodo.12732833" ID-Zenodo-Dep="12732833" httpUri="https://zenodo.org/record/12732833/files/figure.png" pageId="6" pageNumber="339" startId="6.[151,250,1104,1128]" targetBox="[159,1436,181,1073]" targetPageId="6" targetType="figure">
<paragraph id="8BEE369A0B3AFFF4CAD7E735FD27FB1B" blockId="6.[151,1436,1104,1237]" pageId="6" pageNumber="339">
<emphasis id="B925EA880B3AFFF4CAD7E735FED2FBA6" bold="true" box="[151,275,1104,1128]" pageId="6" pageNumber="339">FIGURE 3.</emphasis>
Map indicating localities for
<taxonomicName id="4C514D190B3AFFF4C802E734FCDCFBA7" box="[578,797,1105,1129]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="6" pageNumber="339" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B3AFFF4C802E734FCDCFBA7" box="[578,797,1105,1129]" italics="true" pageId="6" pageNumber="339">Uropeltis pulneyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D190B3AFFF4C90FE734FBCAFBA7" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[847,1035,1105,1129]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="6" pageNumber="339" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B3AFFF4C90FE734FBCAFBA7" box="[847,1035,1105,1129]" italics="true" pageId="6" pageNumber="339">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B3AFFF4CE51E734FBA3FBA7" bold="true" box="[1041,1122,1105,1129]" pageId="6" pageNumber="339">
<taxonomicNameLabel id="A21657F30B3AFFF4CE51E734FBA3FBA7" box="[1041,1122,1105,1129]" pageId="6" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Locality symbols with a black dot inside are associated with voucher specimens; those symbols without a black dot indicate where specimens have been observed but not collected. The imprecise type locality for (the only known specimen of)
<taxonomicName id="4C514D190B3AFFF4CE66E7FCFB66FB7F" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" baseAuthorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" baseAuthorityYear="1878" box="[1062,1191,1177,1201]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="6" pageNumber="339" phylum="Chordata" rank="species" species="guentheri" status="comb. nov.">
<emphasis id="B925EA880B3AFFF4CE66E7FCFB66FB7F" box="[1062,1191,1177,1201]" italics="true" pageId="6" pageNumber="339">U. guentheri</emphasis>
</taxonomicName>
<emphasis id="B925EA880B3AFFF4CEEDE7FCFAE0FB7F" bold="true" box="[1197,1313,1177,1201]" pageId="6" pageNumber="339">
<taxonomicNameLabel id="A21657F30B3AFFF4CEEDE7FCFAE0FB7F" box="[1197,1313,1177,1201]" pageId="6" pageNumber="339" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
is ca. 1,500 m, possibly in or close to Meghamalai (“Cumbum Hills”).
</paragraph>
</caption>
<paragraph id="8BEE369A0B3AFFF4CA87E642FCCBF9F3" blockId="6.[151,1437,1319,1598]" pageId="6" pageNumber="339">
Head and body scales macroscopically smooth, lacking keels, except on far posterior of tail. Body scales generally evenly sized on dorsum and along body except for those involved in dorsal scale row reductions. Midline ventral scales between mental and anal of even size though anterior- and posteriormost ones gradually narrow, posteriormost ventral with V-shaped posterior margin (gently rounded in preceding ventrals). Ventrals 187, though could be interpreted as 188 if the small posteriormost element on the left side is counted. Ventrals at midbody approximately 1.6 times as broad as exposed part of adjacent first dorsal scale row. Dorsal scale rows 19 immediately behind head, reducing to 17 by level with 31
<superScript id="7C249BD20B3AFFF4C963E69AFCEFF9C3" attach="left" box="[803,814,1535,1549]" fontSize="6" pageId="6" pageNumber="339">st</superScript>
ventral, maintained thereafter along most of body with minor fluctuation on right between 50
<superScript id="7C249BD20B3AFFF4C801E547FD8FF9FE" attach="left" box="[577,590,1570,1584]" fontSize="6" pageId="6" pageNumber="339">th</superScript>
and 60
<superScript id="7C249BD20B3AFFF4C8E0E547FD6CF9FE" attach="left" box="[672,685,1570,1584]" fontSize="6" pageId="6" pageNumber="339">th</superScript>
ventral:
</paragraph>
<paragraph id="8BEE369A0B3AFFF4CB46E50EFD58F900" blockId="6.[199,880,1643,1742]" pageId="6" pageNumber="339">4+5 (31) +4 (50), 4+5 (53), +4 (57), -4 (58) 19 --------------- 17 ---------------------------------------------- 17 4+5 (28) [b3, s4: 53]</paragraph>
<paragraph id="8BEE369A0B3AFFF4CA87E59EFEF4F84F" blockId="6.[151,1437,1787,2029]" pageId="6" pageNumber="339">Dorsal scale rows 13 at base of tail. Paired anal scales (right overlying left) considerably larger than posteriormost ventrals and subcaudals. Distal margin of each anal overlaps three other scales in addition to anteriormost subcaudals. Subcaudals 8,7. Posteriormost tail scales (most notably on dorsum) bear few, very short and low, subparallel ridgelike tubercles.</paragraph>
<paragraph id="8BEE369A0B3AFFF5CA87E4EEFBEDFF1B" blockId="6.[151,1437,1787,2029]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="340" pageId="6" pageNumber="339">
Tail shield not apparent; terminus most closely matching
<typeStatus id="54EA88380B3AFFF4C917E4EEFC4EF86B" box="[855,911,1931,1957]" pageId="6" pageNumber="339">Type</typeStatus>
III among Smiths (1943) states for
<taxonomicName id="4C514D190B3AFFF4CF73E4EEFA5DF86B" authorityName="Cuvier" authorityYear="1829" box="[1331,1436,1931,1957]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="6" pageNumber="339" phylum="Chordata" rank="genus">
<emphasis id="B925EA880B3AFFF4CF73E4EEFA5DF86B" box="[1331,1436,1931,1957]" italics="true" pageId="6" pageNumber="339">Uropeltis</emphasis>
</taxonomicName>
tails (also Pyron
<emphasis id="B925EA880B3AFFF4CB1BE4D5FE57F807" box="[347,406,1967,1993]" italics="true" pageId="6" pageNumber="339">et al.</emphasis>
s [2016]
<typeStatus id="54EA88380B3AFFF4C845E4CAFDFCF807" box="[517,573,1967,1993]" pageId="6" pageNumber="339">Type</typeStatus>
III). Terminal scute longer and taller than wide, surrounded by nine scales including last subcaudals; ventrally convex, smooth; dorsally concave shortly anterior to terminus; laterally with few, small, low tubercles. Posterior edge of terminal scute ends in pair of narrowly separated (
<quantity id="4CA99B7F0B3BFFF5CEE6E3F2FAC0FF7F" box="[1190,1281,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="340" unit="mm" value="0.7">0.7 mm</quantity>
), paramedian, short, inconspicuous points; these not disposed on thin horizontal plate-like ridge.
</paragraph>
<caption id="DF2E66120B3BFFF5CAD7E4DAFB33F835" ID-DOI="http://doi.org/10.5281/zenodo.12732839" ID-Zenodo-Dep="12732839" httpUri="https://zenodo.org/record/12732839/files/figure.png" pageId="7" pageNumber="340" startId="7.[151,250,1983,2007]" targetBox="[151,1435,309,1959]" targetPageId="7" targetType="figure">
<paragraph id="8BEE369A0B3BFFF5CAD7E4DAFB33F835" blockId="7.[151,1436,1983,2043]" pageId="7" pageNumber="340">
<emphasis id="B925EA880B3BFFF5CAD7E4DAFED4F819" bold="true" box="[151,277,1983,2007]" pageId="7" pageNumber="340">FIGURE 4.</emphasis>
The three most-recently collected paratypes of
<taxonomicName id="4C514D190B3BFFF5C94AE4DAFBCFF819" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[778,1038,1983,2007]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="7" pageNumber="340" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B3BFFF5C94AE4DAFBCFF819" box="[778,1038,1983,2007]" italics="true" pageId="7" pageNumber="340">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B3BFFF5CE57E4DAFBABF819" bold="true" box="[1047,1130,1983,2007]" pageId="7" pageNumber="340">
<taxonomicNameLabel id="A21657F30B3BFFF5CE57E4DAFBABF819" box="[1047,1130,1983,2007]" pageId="7" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in approximately dorsal (left column) and ventral (right column) views. A &amp; B, BNHS 3765, C &amp; D, BNHS 3763, and E &amp; F, BNHS 3764.
</paragraph>
</caption>
<caption id="DF2E66120B34FFFACAD7E4F2FD8DF836" ID-DOI="http://doi.org/10.5281/zenodo.12732841" ID-Zenodo-Dep="12732841" httpUri="https://zenodo.org/record/12732841/files/figure.png" pageId="8" pageNumber="341" startId="8.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="8" targetType="figure">
<paragraph id="8BEE369A0B34FFFACAD7E4F2FD8DF836" blockId="8.[151,1437,1943,2040]" pageId="8" pageNumber="341">
<emphasis id="B925EA880B34FFFACAD7E4F2FED6F87E" bold="true" box="[151,279,1943,1968]" pageId="8" pageNumber="341">FIGURE 5.</emphasis>
<taxonomicName id="4C514D190B34FFFACB61E4F2FDE4F861" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[289,549,1943,1967]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="8" pageNumber="341" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B34FFFACB61E4F2FDE4F861" box="[289,549,1943,1967]" italics="true" pageId="8" pageNumber="341">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B34FFFAC870E4FDFD45F87E" bold="true" box="[560,644,1944,1968]" pageId="8" pageNumber="341">
<taxonomicNameLabel id="A21657F30B34FFFAC870E4FDFD45F87E" box="[560,644,1944,1968]" pageId="8" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S.
</paragraph>
</caption>
<paragraph id="8BEE369A0B35FFFBCA87E3F2FAD6FDF5" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
In alcohol, venter dark (blackish) brown with mostly asymmetrical and somewhat irregularly spaced pale yellow crossbars, approximately one (to two) scales long, mostly extending from midline across first two to three dorsal scale rows; anteriorly these cross bars do not encroach onto ventrals or first dorsal scale rows, instead on dorsal scale rows 45, 35, 24 or 23 (from anterior to posterior). On body, 27 and 25 pale cross bars on left and right, respectively. Posteriormost cross bar level with and encroaching slightly onto lateralmost edges of anal shields. Some ventral pale bars, pale flecking on ventrals and on dorsal scale rows 13 but overall bicoloured; mostly dark below, pale above. Many scales with pale yellow to off-white posterolateral edges. Pale dorsum in the form of an irregularly edged but large (more than 50% of area of each scale) mottled pale tan spot on every scale of (9)11 dorsalmost scale rows, except for anteriormost
<quantity id="4CA99B7F0B35FFFBC8E4E2D0FD32FE1E" box="[676,755,437,464]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="342" unit="mm" value="60.0">60 mm</quantity>
and posteriormost
<quantity id="4CA99B7F0B35FFFBC988E2D3FBD8FE1E" box="[968,1049,438,464]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="342" unit="mm" value="30.0">30 mm</quantity>
. Tail dark brown except for yellow on posteriormost two thirds of terminal scute. Scales not notably iridescent. Underside of head brown except for pale yellow flecks on first infralabials and anterior of mental shield. Dorsally, head dark olive-brown; diffuse pale tip to rostral; large pale yellow spots covering posterior half of each posteriormost (fourth) supralabial.
</paragraph>
<paragraph id="8BEE369A0B35FFFBCA87E120FA54FB9B" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E120FE1CFD91" bold="true" box="[199,477,581,607]" pageId="9" pageNumber="342">
Variation in
<typeStatus id="54EA88380B35FFFBCB20E120FE19FD91" box="[352,472,581,607]" pageId="9" pageNumber="342" type="paratype">paratypes</typeStatus>
.
</emphasis>
Some morphological data and images presented in
<figureCitation id="136A2A1F0B35FFFBCE6BE120FBB1FD91" box="[1067,1136,581,607]" captionStart="FIGURE 4" captionStartId="7.[151,250,1983,2007]" captionTargetBox="[151,1435,309,1959]" captionTargetId="figure-46@7.[151,1435,309,1959]" captionTargetPageId="7" captionText="FIGURE 4. The three most-recently collected paratypes of Uropeltis caudomaculata sp. nov. in approximately dorsal (left column) and ventral (right column) views. A &amp; B, BNHS 3765, C &amp; D, BNHS 3763, and E &amp; F, BNHS 3764." figureDoi="http://doi.org/10.5281/zenodo.12732839" httpUri="https://zenodo.org/record/12732839/files/figure.png" pageId="9" pageNumber="342">Fig. 4</figureCitation>
,
<tableCitation id="C6D303210B35FFFBCE3DE120FB13FD91" box="[1149,1234,581,607]" captionStart="TABLE 3" captionStartId="11.[151,239,467,493]" captionTargetPageId="11" captionText="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="9" pageNumber="342" tableUuid="DF2E66120B37FFF9CAD7E2B6F884FDDF">Table 3</tableCitation>
, and Appendix 2. Among the four
<collectionCode id="ED40AE5F0B35FFFBCB15E10CFE68FD4D" box="[341,425,617,643]" pageId="9" pageNumber="342">BMNH</collectionCode>
specimens, 1955.1.2.68 and more notably 1955.1.2.71 and 72 are somewhat dehydrated. There are longitudinal incisions in the tails of 1955.1.2.70 and 72, and a long ventral incision into the coelom of 1955.1.2.68, which also has some skin on the left side of the head reflected by dissection.
<collectionCode id="ED40AE5F0B35FFFBCEC9E1D4FB1CFD05" box="[1161,1245,689,715]" pageId="9" pageNumber="342">BMNH</collectionCode>
1955.1.2.70 was preserved close to ecdysis.
<collectionCode id="ED40AE5F0B35FFFBCB8FE1B0FDE2FD21" box="[463,547,725,751]" pageId="9" pageNumber="342">BMNH</collectionCode>
1955.1.2.70 and 71 both have small subrectangular supernumerary shields on the temporal region of the head region—between the fourth supralabial and parietal, immediately behind the ocular. In association with this scalation pattern, the parietal shields in these
<specimenCount id="9D57FD130B35FFFBC9D7E078FBFDFCF8" box="[919,1084,796,823]" count="2" pageId="9" pageNumber="342" type="generic">two specimens</specimenCount>
contact 4,5 and 5,6 (rather than 4,4) other head shields, respectively. In
<collectionCode id="ED40AE5F0B35FFFBC81CE024FD71FC95" box="[604,688,833,859]" pageId="9" pageNumber="342">BMNH</collectionCode>
1955.1.2.68 the anal shields are fused, though at the distal edge the right appears to slightly overlap the left (as in all other specimens). In
<collectionCode id="ED40AE5F0B35FFFBC99EE000FBF3FCB1" box="[990,1074,869,895]" pageId="9" pageNumber="342">BMNH</collectionCode>
1955.1.2.71 the right prefrontal overlaps the left rather than vice versa. In 1955.1.2.68 three of the subcaudals are unpaired (fused). As with the
<typeStatus id="54EA88380B35FFFBCAD7E0C9FF3DFC08" box="[151,252,940,966]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
, the female
<collectionCode id="ED40AE5F0B35FFFBCBC7E0C9FE1AFC08" box="[391,475,940,966]" pageId="9" pageNumber="342">BMNH</collectionCode>
<typeStatus id="54EA88380B35FFFBCBA4E0C9FD8EFC08" box="[484,591,940,966]" pageId="9" pageNumber="342" type="paratype">paratypes</typeStatus>
have some low and inconspicuous, longitudinal ridges on some of the tail scales, especially more dorsally and posteriorly (not on the subcaudals). These ridges are slightly more prominent in the male
<collectionCode id="ED40AE5F0B35FFFBCB59E091FEACFBC0" box="[281,365,1012,1038]" pageId="9" pageNumber="342">BMNH</collectionCode>
<typeStatus id="54EA88380B35FFFBCB34E091FE23FBC0" box="[372,482,1012,1038]" pageId="9" pageNumber="342" type="paratype">paratypes</typeStatus>
, more so in 1955.1.2.71 than in 1955.1.2.70, and are more widespread, occurring also on the anal shields and a few of the posteriormost ventrals. The pale mottled spots on the dorsalmost 911 scales are more or less conspicuous; as conspicuous as in hlotype in 1955.1.2.71 and less conspicuous in1955.1.2.68 and 72.
</paragraph>
<paragraph id="8BEE369A0B35FFFBCA87E73AFE4EFA72" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
Among the BNHS
<typeStatus id="54EA88380B35FFFBCBE0E705FDCFFBB4" box="[416,526,1120,1146]" pageId="9" pageNumber="342" type="paratype">paratypes</typeStatus>
, BNHS 3765 is slightly dehydrated, with one ventral incision into the coelom at midbody (extending across six ventral shields) and one close to the anus (extending across four ventrals). BNHS 3763 is well preserved, with a coelomic incision at midbody covering six ventrals. BNHS 3764 is slightly dehydrated, with one coelomic incision anterior to midbody (15 ventrals) and one posterior to midbody (8 ventrals). The right parietal in 3763 contacts only three head shields. The parietals are posteriorly narrow in 3765 and 3763. The third ventral is much shorter and smaller than the first two in 3763. The last (posteriormost) ventral shield is divided in 3763, with one paramedian scale on the right and two small scales on the left. The tips of the points on the terminal scute are blunt in BNHS 3763 and 3764. BNHS 3765 resembles the
<typeStatus id="54EA88380B35FFFBC9D6E63FFC39FABA" box="[918,1016,1370,1396]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
in its colour pattern, but BNHS 3763 and 3764 differ in having a lateral yellow stripe (instead of yellow spots) from the head to anterior of the body and on the base of the tail.
</paragraph>
<paragraph id="8BEE369A0B35FFFBCA87E6A0FB23F985" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E6A0FEACFA11" bold="true" box="[199,365,1477,1503]" pageId="9" pageNumber="342">Colour in life.</emphasis>
See
<figureCitation id="136A2A1F0B35FFFBCBE4E6A3FE26FA2E" box="[420,487,1477,1504]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Uropeltis caudomaculata sp. nov. in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S." figureDoi="http://doi.org/10.5281/zenodo.12732841" httpUri="https://zenodo.org/record/12732841/files/figure.png" pageId="9" pageNumber="342">Fig. 5</figureCitation>
. The darker background is generally dark brown. The pale spots and mottling on the uppermost 911 dorsal scale rows are olive-green to pale yellow. The larger pale spots (or stripes) on and behind the lateral surface of the head, along the ventrolateral and lateral aspect of the body, and on the lateral surface of (at least the base of) the tail are bright yellow, more vivid than the more diffuse dorsal pale markings.
</paragraph>
<paragraph id="8BEE369A0B35FFFBCA87E530FF37F979" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E530FE77F9A1" bold="true" box="[199,438,1621,1647]" pageId="9" pageNumber="342">Sexual dimorphism.</emphasis>
See
<figureCitation id="136A2A1F0B35FFFBCBABE530FDEEF9A1" box="[491,559,1621,1647]" captionStart="FIGURE 6" captionStartId="10.[151,250,1922,1946]" captionTargetBox="[159,1428,1031,1891]" captionTargetId="figure-414@10.[151,1436,1029,1899]" captionTargetPageId="10" captionText="FIGURE 6. Variation in numbers of ventral and subcaudal shields in females (circles) and males (triangles) of Uropeltis pulneyensis (25 females, 19 males), U. caudomaculata sp. nov. (6 females, 2 males) and the holotype of U. guentheri comb. nov. (one male). See Tables 3 and 4 for sources of data." figureDoi="http://doi.org/10.5281/zenodo.12732845" httpUri="https://zenodo.org/record/12732845/files/figure.png" pageId="9" pageNumber="342">Fig. 6</figureCitation>
. The available sample size is too small to confirm probable sexual dimorphism in numbers of ventral shields, but the data do show that males have relatively longer tails with more subcaudals than females.
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B35FFFBCA87E5A4FAB2F8EC" pageId="9" pageNumber="342" type="etymology">
<paragraph id="8BEE369A0B35FFFBCA87E5A4FAB2F8EC" blockId="9.[151,1437,151,2043]" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E5A4FE8FF915" bold="true" box="[199,334,1729,1755]" pageId="9" pageNumber="342">Etymology.</emphasis>
The name of the new species is in reference to the lateral yellow spot on each side of the base of the tail (in specimens from Meghamalai), rather than a stripe as occurs in many congeners, including, typically, its closest known relative,
<taxonomicName id="4C514D190B35FFFBCBE0E46CFD84F8EC" box="[416,581,1800,1826]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="9" pageNumber="342" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B35FFFBCBE0E46CFD84F8EC" box="[416,581,1800,1826]" italics="true" pageId="9" pageNumber="342">U. pulneyensis</emphasis>
</taxonomicName>
. For nomenclatural purposes, the species epithet is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B35FFFBCA87E449FB8EF888" box="[199,1103,1836,1863]" pageId="9" pageNumber="342" type="vernacular_names">
<paragraph id="8BEE369A0B35FFFBCA87E449FB8EF888" blockId="9.[151,1437,151,2043]" box="[199,1103,1836,1863]" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E449FDE3F888" bold="true" box="[199,546,1836,1862]" pageId="9" pageNumber="342">Suggested vernacular names.</emphasis>
Tailspot uropeltis or Tailspot shieldtail (English).
</paragraph>
</subSubSection>
<subSubSection id="C34B65110B35FFF8CA87E435FBE3FC07" lastPageId="10" lastPageNumber="343" pageId="9" pageNumber="342" type="distribution">
<paragraph id="8BEE369A0B35FFF8CA87E435FBC6FED3" blockId="9.[151,1437,151,2043]" lastBlockId="10.[151,1437,151,969]" lastPageId="10" lastPageNumber="343" pageId="9" pageNumber="342">
<emphasis id="B925EA880B35FFFBCA87E435FD5FF8A4" bold="true" box="[199,670,1872,1898]" pageId="9" pageNumber="342">Distribution, habitat and conservation.</emphasis>
The new species is known only from three localities, Meghamalai Tiger Reserve in
<collectingRegion id="4995F8780B35FFFBCB20E411FDE9F841" box="[352,552,1908,1935]" country="India" name="Tamil Nadu" pageId="9" pageNumber="342">Tamil Nadu state</collectingRegion>
, Periyar Tiger Reserve, and Yellapetty, Munnar in
<collectingRegion id="4995F8780B35FFFBCE37E411FB3EF841" box="[1143,1279,1908,1935]" country="India" name="Kerala" pageId="9" pageNumber="342">Kerala state</collectingRegion>
(
<figureCitation id="136A2A1F0B35FFFBCF51E411FA9BF840" box="[1297,1370,1908,1934]" captionStart="FIGURE 2" captionStartId="5.[151,250,1942,1966]" captionTargetBox="[151,1435,181,1918]" captionTargetId="figure-17@5.[151,1435,181,1918]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Uropeltis caudomaculata sp. nov. (BMNH 1955.1.2.69). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732831" httpUri="https://zenodo.org/record/12732831/files/figure.png" pageId="9" pageNumber="342">Fig. 2</figureCitation>
). The
<typeStatus id="54EA88380B35FFFBCAD7E4FDFF38F87C" box="[151,249,1944,1970]" pageId="9" pageNumber="342" type="holotype">holotype</typeStatus>
and four
<collectionCode id="ED40AE5F0B35FFFBCB27E4FCFE7AF87D" box="[359,443,1945,1971]" pageId="9" pageNumber="342">BMNH</collectionCode>
<typeStatus id="54EA88380B35FFFBCB83E4FCFDEFF87D" box="[451,558,1945,1971]" pageId="9" pageNumber="342" type="paratype">paratypes</typeStatus>
were probably collected between 1946 and 1948 (see
<bibRefCitation id="EFC04B6B0B35FFFBCED5E4FCFA51F87C" author="Hutton, A. F. &amp; David, P." box="[1173,1424,1944,1971]" pageId="9" pageNumber="342" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David 2008</bibRefCitation>
), according to the
<collectionCode id="ED40AE5F0B35FFFBCB1CE4D8FE71F819" box="[348,432,1981,2007]" pageId="9" pageNumber="342">BMNH</collectionCode>
catalogue from
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<geoCoordinate id="EE65505D0B35FFFBC830E4D9FD6EF819" box="[624,687,1980,2007]" degrees="3,000" direction="north" orientation="latitude" pageId="9" pageNumber="342" precision="55" value="3.0">3,000</geoCoordinate>
<geoCoordinate id="EE65505D0B35FFFBC8EFE4D9FD3DF819" box="[687,764,1980,2007]" degrees="5,000" direction="west" orientation="longitude" pageId="9" pageNumber="342" precision="55" value="-5.0">5,000</geoCoordinate>
feet
</quantity>
(
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) elevation (but according to
<bibRefCitation id="EFC04B6B0B35FFFBCF6EE4D8FEF5F834" author="Hutton, A. F. &amp; David, P." pageId="9" pageNumber="342" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David [2008]</bibRefCitation>
,
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). It is not possible to be certain that the snakes from Meghamalai reported as
<taxonomicName id="4C514D190B35FFF8CFC1E484FEDBFF7F" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" lastPageId="10" lastPageNumber="343" pageId="9" pageNumber="342" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B35FFF8CFC1E484FEDBFF7F" italics="true" lastPageId="10" lastPageNumber="343" pageId="9" pageNumber="342">U. pulneyensis</emphasis>
</taxonomicName>
by Hutton (1949) are actually
<taxonomicName id="4C514D190B36FFF8C83BE3F2FC8DFF7F" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[635,844,151,177]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B36FFF8C83BE3F2FC8DFF7F" box="[635,844,151,177]" italics="true" pageId="10" pageNumber="343">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B36FFF8C914E3F2FC6EFF7F" bold="true" box="[852,943,151,177]" pageId="10" pageNumber="343">
<taxonomicNameLabel id="A21657F30B36FFF8C914E3F2FC6EFF7F" box="[852,943,151,177]" pageId="10" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because no scalation or colour pattern data were presented. However, Hutton (1949) reported the species identified by him as
<taxonomicName id="4C514D190B36FFF8CE03E3DEFB2BFF1B" box="[1091,1258,187,213]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B36FFF8CE03E3DEFB2BFF1B" box="[1091,1258,187,213]" italics="true" pageId="10" pageNumber="343">U. pulneyensis</emphasis>
</taxonomicName>
to be relatively common, to be encountered on roads, in tea nurseries and under or in decaying wood and heaps of stones, and to be viviparous, giving birth to 8 to 17 young of approximately
<quantity id="4CA99B7F0B36FFF8C96CE266FCBAFED3" box="[812,891,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="343" unit="mm" value="60.0">60 mm</quantity>
total length.
</paragraph>
<paragraph id="8BEE369A0B36FFF8CA87E242FEB8FD3F" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="343">
<typeStatus id="54EA88380B36FFF8CA87E242FEE8FE8F" box="[199,297,295,321]" pageId="10" pageNumber="343" type="paratype">Paratype</typeStatus>
BNHS 3765 was a roadkill specimen collected from the Manalar region of Meghamalai (
<figureCitation id="136A2A1F0B36FFF8CF06E242FA4FFE8F" box="[1350,1422,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1854,1878]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure-18@14.[151,1435,181,1830]" captionTargetPageId="14" captionText="FIGURE 7. Habitats of recent localities for Uropeltis caudomaculata sp. nov. A. Locality where paratype BNHS 3765 was collected, at Manalar, Meghamalai, Tamil Nadu. B. General habitat close to where paratypes BNHS 3763 and 3764 were collected; Yellapetty, Munnar, Kerala. Photographs by JDG (A) and SD (B)." figureDoi="http://doi.org/10.5281/zenodo.12732849" httpUri="https://zenodo.org/record/12732849/files/figure.png" pageId="10" pageNumber="343">Fig. 7</figureCitation>
), bordered by tea plantations, with sparsely distributed windbreak trees and the Manalar catchment on the other side. Three other (uncollected) roadkill individuals were sighted by JDG during the day between tea plantation and forest or reservoir in June and July, 2021 near Meghamalai village (
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,
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;
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,
<geoCoordinate id="EE65505D0B36FFF8CF68E2F6FF6AFE1F" degrees="77.36677" direction="east" orientation="longitude" pageId="10" pageNumber="343" precision="1" value="77.36677">77.36677° E</geoCoordinate>
;
<geoCoordinate id="EE65505D0B36FFF8CAFAE2D3FE80FE1F" box="[186,321,438,465]" degrees="9.61945" direction="north" orientation="latitude" pageId="10" pageNumber="343" precision="1" value="9.61945">9.61945° N</geoCoordinate>
,
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, all at ca.
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elevation). Four uncollected individuals were found by SD and colleagues while rolling logs within Periyar Tiger Reserve (
<geoCoordinate id="EE65505D0B36FFF8C906E2BFFC1CFE3B" box="[838,989,474,501]" degrees="9.587788" direction="north" orientation="latitude" pageId="10" pageNumber="343" precision="1" value="9.587788">9.587788° N</geoCoordinate>
,
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,
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) on
<date id="FFEF105A0B36FFF8CF5EE2BEFF12FDD7" pageId="10" pageNumber="343" value="2017-07-22" valueMax="2017-07-23" valueMin="2017-07-22">2223 July 2017</date>
. One of these individuals is illustrated in
<figureCitation id="136A2A1F0B36FFF8C8E5E29AFD28FDD7" box="[677,745,510,537]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Uropeltis caudomaculata sp. nov. in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S." figureDoi="http://doi.org/10.5281/zenodo.12732841" httpUri="https://zenodo.org/record/12732841/files/figure.png" pageId="10" pageNumber="343">Fig. 5</figureCitation>
. Ventrals and other scales of this individual were not counted in the field but from photographs it is a female with approximately 7,7 subcaudal scales and has 200 middorsal scales (compared with 198 and ca.
<quantity id="4CA99B7F0B36FFF8C865E122FDADFDAF" box="[549,620,583,609]" metricMagnitude="0" metricUnit="m" metricValue="4.7244" pageId="10" pageNumber="343" unit="in" value="186.0">186 in</quantity>
the female
<typeStatus id="54EA88380B36FFF8C8B2E122FC9EFDAF" box="[754,863,583,609]" pageId="10" pageNumber="343" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="4C514D190B36FFF8C9C6E122FB97FDAF" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[902,1110,583,609]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B36FFF8C9C6E122FB97FDAF" box="[902,1110,583,609]" italics="true" pageId="10" pageNumber="343">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B36FFF8CE1DE122FB79FDAF" bold="true" box="[1117,1208,583,609]" pageId="10" pageNumber="343">
<taxonomicNameLabel id="A21657F30B36FFF8CE1DE122FB79FDAF" box="[1117,1208,583,609]" pageId="10" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C514D190B36FFF8CEB0E122FA57FDAF" box="[1264,1430,583,609]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B36FFF8CEB0E122FA57FDAF" box="[1264,1430,583,609]" italics="true" pageId="10" pageNumber="343">U. pulneyensis</emphasis>
</taxonomicName>
, respectively).
<typeStatus id="54EA88380B36FFF8CB79E10EFE67FD4B" box="[313,422,619,645]" pageId="10" pageNumber="343" type="paratype">Paratypes</typeStatus>
BNHS 3763 &amp; 3764 were collected from secondary forest in the Yellapetty area at an elevation of ca.
<quantity id="4CA99B7F0B36FFF8CB0AE1EBFE68FD67" box="[330,425,654,681]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="10" pageNumber="343" unit="m" value="1950.0">1,950 m</quantity>
. Both individuals were collected during the day under logs within the forest area. This site is surrounded by tea plantations on one side and hill slope on the other. The general habitat (
<figureCitation id="136A2A1F0B36FFF8CEEFE1D6FB32FD03" box="[1199,1267,691,717]" captionStart="FIGURE 7" captionStartId="14.[151,250,1854,1878]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure-18@14.[151,1435,181,1830]" captionTargetPageId="14" captionText="FIGURE 7. Habitats of recent localities for Uropeltis caudomaculata sp. nov. A. Locality where paratype BNHS 3765 was collected, at Manalar, Meghamalai, Tamil Nadu. B. General habitat close to where paratypes BNHS 3763 and 3764 were collected; Yellapetty, Munnar, Kerala. Photographs by JDG (A) and SD (B)." figureDoi="http://doi.org/10.5281/zenodo.12732849" httpUri="https://zenodo.org/record/12732849/files/figure.png" pageId="10" pageNumber="343">Fig. 7</figureCitation>
) of this area is shola-forest mosaic.
</paragraph>
<paragraph id="8BEE369A0B36FFF8CA87E19EFBE3FC07" blockId="10.[151,1437,151,969]" pageId="10" pageNumber="343">
<taxonomicName id="4C514D190B36FFF8CA87E19EFE23FCDB" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[199,482,763,789]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B36FFF8CA87E19EFE23FCDB" box="[199,482,763,789]" italics="true" pageId="10" pageNumber="343">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B36FFF8CBA7E19EFDFEFCDB" bold="true" box="[487,575,763,789]" pageId="10" pageNumber="343">
<taxonomicNameLabel id="A21657F30B36FFF8CBA7E19EFDFEFCDB" box="[487,575,763,789]" pageId="10" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from multiple localities up to ca.
<quantity id="4CA99B7F0B36FFF8CE58E19EFB9CFCDB" box="[1048,1117,763,789]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="343" unit="km" value="60.0">60 km</quantity>
apart, and though threatened at least locally by road traffic it appears able to tolerate at least some human-mediated environmental disturbance. However, there is a notable intervening gap between two clusters of locality records (
<figureCitation id="136A2A1F0B36FFF8CE2AE026FB6EFC93" box="[1130,1199,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1104,1128]" captionTargetBox="[159,1436,181,1073]" captionTargetId="figure-18@6.[151,1436,181,1081]" captionTargetPageId="6" captionText="FIGURE 3. Map indicating localities for Uropeltis pulneyensis and U. caudomaculata sp. nov. Locality symbols with a black dot inside are associated with voucher specimens; those symbols without a black dot indicate where specimens have been observed but not collected. The imprecise type locality for (the only known specimen of) U. guentheri comb. nov. is ca. 1,500 m, possibly in or close to Meghamalai (“Cumbum Hills”)." figureDoi="http://doi.org/10.5281/zenodo.12732833" httpUri="https://zenodo.org/record/12732833/files/figure.png" pageId="10" pageNumber="343">Fig. 3</figureCitation>
), only a single point locality is known at the northernmost edge of its recorded range, and very little is known about its ecological tolerances or preferences. Thus, we suggest that for the present, the species is likely to qualify for Data Deficient status under IUCN Red List criteria (
<bibRefCitation id="EFC04B6B0B36FFF8C875E0CAFBD7FC07" author="IUCN Species Survival Commission" box="[565,1046,943,969]" pageId="10" pageNumber="343" refId="ref11575" refString="IUCN Species Survival Commission (2012) IUCN Red List categories and criteria, version 3.1. 2 nd Edition. IUCN, Gland and Cambridge, 32 pp." type="book" year="2012">IUCN Species Survival Commission 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2E66120B36FFF8CAD7E4E7FD0BF82D" ID-DOI="http://doi.org/10.5281/zenodo.12732845" ID-Zenodo-Dep="12732845" httpUri="https://zenodo.org/record/12732845/files/figure.png" pageId="10" pageNumber="343" startId="10.[151,250,1922,1946]" targetBox="[159,1428,1031,1891]" targetPageId="10" targetType="figure">
<paragraph id="8BEE369A0B36FFF8CAD7E4E7FD0BF82D" blockId="10.[151,1437,1922,2019]" pageId="10" pageNumber="343">
<emphasis id="B925EA880B36FFF8CAD7E4E7FED6F855" bold="true" box="[151,279,1922,1947]" pageId="10" pageNumber="343">FIGURE 6.</emphasis>
Variation in numbers of ventral and subcaudal shields in females (circles) and males (triangles) of
<taxonomicName id="4C514D190B36FFF8CF7DE4E6FECCF871" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B36FFF8CF7DE4E6FECCF871" italics="true" pageId="10" pageNumber="343">Uropeltis pulneyensis</emphasis>
</taxonomicName>
(25 females, 19 males),
<taxonomicName id="4C514D190B36FFF8C84DE4C2FD08F871" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[525,713,1959,1983]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B36FFF8C84DE4C2FD08F871" box="[525,713,1959,1983]" italics="true" pageId="10" pageNumber="343">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B36FFF8C891E4C2FCE3F871" bold="true" box="[721,802,1959,1983]" pageId="10" pageNumber="343">
<taxonomicNameLabel id="A21657F30B36FFF8C891E4C2FCE3F871" box="[721,802,1959,1983]" pageId="10" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(6 females, 2 males) and the holotype of
<taxonomicName id="4C514D190B36FFF8CE92E4C2FA92F871" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" baseAuthorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" baseAuthorityYear="1878" box="[1234,1363,1959,1983]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="10" pageNumber="343" phylum="Chordata" rank="species" species="guentheri" status="comb. nov.">
<emphasis id="B925EA880B36FFF8CE92E4C2FA92F871" box="[1234,1363,1959,1983]" italics="true" pageId="10" pageNumber="343">U. guentheri</emphasis>
</taxonomicName>
<emphasis id="B925EA880B36FFF8CF1BE4C2FF02F82D" bold="true" pageId="10" pageNumber="343">
<taxonomicNameLabel id="A21657F30B36FFF8CF1BE4C2FF02F82D" pageId="10" pageNumber="343" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(one male). See Tables 3 and 4 for sources of data.
</paragraph>
</caption>
<caption id="DF2E66120B37FFF9CAD7E2B6F884FDDF" ID-Table-UUID="DF2E66120B37FFF9CAD7E2B6F884FDDF" httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="11" pageNumber="344" startId="11.[151,239,467,493]" targetBox="[162,2023,545,1670]" targetIsTable="true" targetPageId="11" targetType="table">
<paragraph id="8BEE369A0B37FFF9CAD7E2B6F884FDDF" blockId="11.[151,2037,467,529]" pageId="11" pageNumber="344">
<emphasis id="B925EA880B37FFF9CAD7E2B6FEC9FE23" bold="true" box="[151,264,467,493]" pageId="11" pageNumber="344">TABLE 3.</emphasis>
Morphometric (in mm) and meristic data for holotype (*) and paratypes of
<taxonomicName id="4C514D190B37FFF9CE5EE2B6FAEAFE23" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[1054,1323,467,493]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B37FFF9CE5EE2B6FAEAFE23" box="[1054,1323,467,493]" italics="true" pageId="11" pageNumber="344">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B37FFF9CF71E2B6FA44FE23" bold="true" box="[1329,1413,467,493]" pageId="11" pageNumber="344">
<taxonomicNameLabel id="A21657F30B37FFF9CF71E2B6FA44FE23" box="[1329,1413,467,493]" pageId="11" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the lectotype (†) and a paralectotype (#) of
<taxonomicName id="4C514D190B37FFF9CD13E2B6F82FFE23" box="[1875,2030,467,493]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B37FFF9CD13E2B6F82FFE23" box="[1875,2030,467,493]" italics="true" pageId="11" pageNumber="344">U. pulneyensis</emphasis>
</taxonomicName>
, and the holotype of
<taxonomicName id="4C514D190B37FFF9CB2CE292FE32FDDF" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" baseAuthorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" baseAuthorityYear="1878" box="[364,499,503,529]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="guentheri" status="comb. nov.">
<emphasis id="B925EA880B37FFF9CB2CE292FE32FDDF" box="[364,499,503,529]" italics="true" pageId="11" pageNumber="344">U. guentheri</emphasis>
</taxonomicName>
<emphasis id="B925EA880B37FFF9CBBAE292FDB5FDDF" bold="true" box="[506,628,503,529]" pageId="11" pageNumber="344">
<taxonomicNameLabel id="A21657F30B37FFF9CBBAE292FDB5FDDF" box="[506,628,503,529]" pageId="11" pageNumber="344" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(‡). Values in square brackets are those reported by
<bibRefCitation id="EFC04B6B0B37FFF9CEE3E292FA60FDDF" author="Hutton, A. F. &amp; David, P." box="[1187,1441,503,529]" pageId="11" pageNumber="344" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David (2008)</bibRefCitation>
. Blank cells indicate data not recorded.
</paragraph>
</caption>
<paragraph id="8BEE369A0B37FFF9C92AE144F872F948" pageId="11" pageNumber="344">
<table id="F951C43A0B37000DCAE2E144F826F948" box="[162,2023,545,1670]" colsContinueIn="12.[151,2012,954,1278]" gridcols="12" gridrows="23" pageId="11" pageNumber="344">
<tr id="356134D80B37000DCAE2E144F826FDF4" box="[162,2023,545,570]" gridrow="0" pageId="11" pageNumber="344" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76B05DA40B37000DC971E144FB10FDF4" box="[817,1233,545,570]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="11" pageNumber="344">
<taxonomicName id="4C514D190B37FFF9C92AE144FBAAFDF4" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[874,1131,545,570]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B37FFF9C92AE144FBAAFDF4" box="[874,1131,545,570]" italics="true" pageId="11" pageNumber="344">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B37FFF9CE32E144FB03FDF4" bold="true" box="[1138,1218,545,570]" pageId="11" pageNumber="344">
<taxonomicNameLabel id="A21657F30B37FFF9CE32E144FB03FDF4" box="[1138,1218,545,570]" pageId="11" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76B05DA40B37000DCC30E144F88AFDF4" box="[1648,1867,545,570]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="11" pageNumber="344">
<taxonomicName id="4C514D190B37FFF9CCCFE144F8E5FDF4" box="[1679,1828,545,570]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B37FFF9CCCFE144F8E5FDF4" box="[1679,1828,545,570]" italics="true" pageId="11" pageNumber="344">U. pulneyensis</emphasis>
</taxonomicName>
</th>
<th id="76B05DA40B37000DCD2DE144F826FDF4" box="[1901,2023,545,570]" gridcol="11" gridrow="0" pageId="11" pageNumber="344">
<taxonomicName id="4C514D190B37FFF9CD2EE144F826FDF4" authority="* BMNH" box="[1902,2023,545,570]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="11" pageNumber="344" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B37FFF9CD2EE144F826FDF4" box="[1902,2023,545,570]" italics="true" pageId="11" pageNumber="344">U. guentheri</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356134D80B37000DCAE2E137F826FDA5" box="[162,2023,594,619]" gridrow="1" pageId="11" pageNumber="344" rowspan-0="1">
<td id="76B05DA40B37000DC84AE137FD43FDA5" box="[522,642,594,619]" gridcol="1" gridrow="1" pageId="11" pageNumber="344">*BMNH</td>
<td id="76B05DA40B37000DC8DEE137FCD7FDA5" box="[670,790,594,619]" gridcol="2" gridrow="1" pageId="11" pageNumber="344">BMNH</td>
<td id="76B05DA40B37000DC971E137FC6BFDA5" box="[817,938,594,619]" gridcol="3" gridrow="1" pageId="11" pageNumber="344">BMNH</td>
<td id="76B05DA40B37000DC985E137FBFFFDA5" box="[965,1086,594,619]" gridcol="4" gridrow="1" pageId="11" pageNumber="344">BMNH</td>
<td id="76B05DA40B37000DCE19E137FB10FDA5" box="[1113,1233,594,619]" gridcol="5" gridrow="1" pageId="11" pageNumber="344">BMNH</td>
<td id="76B05DA40B37000DCEB1E137FAFCFDA5" box="[1265,1341,594,619]" gridcol="6" gridrow="1" pageId="11" pageNumber="344">BNHS</td>
<td id="76B05DA40B37000DCF22E137FA6FFDA5" box="[1378,1454,594,619]" gridcol="7" gridrow="1" pageId="11" pageNumber="344">BNHS</td>
<td id="76B05DA40B37000DCF92E137F9DEFDA5" box="[1490,1567,594,619]" gridcol="8" gridrow="1" pageId="11" pageNumber="344">BNHS</td>
<td id="76B05DA40B37000DCC30E137F909FDA5" box="[1648,1736,594,619]" gridcol="9" gridrow="1" pageId="11" pageNumber="344">†BMNH</td>
<td id="76B05DA40B37000DCCAFE137F88AFDA5" box="[1775,1867,594,619]" gridcol="10" gridrow="1" pageId="11" pageNumber="344">#MNHN</td>
<td id="76B05DA40B37000DCD2DE137F826FDA5" box="[1901,2023,594,619]" gridcol="11" gridrow="1" pageId="11" pageNumber="344">‡BMNH</td>
</tr>
<tr id="356134D80B37000DCAE2E1E1F826FD53" box="[162,2023,644,669]" gridrow="2" pageId="11" pageNumber="344" rowspan-0="1">
<td id="76B05DA40B37000DC84AE1E1FD43FD53" box="[522,642,644,669]" gridcol="1" gridrow="2" pageId="11" pageNumber="344">1955.1.2.69</td>
<td id="76B05DA40B37000DC8DEE1E1FCD7FD53" box="[670,790,644,669]" gridcol="2" gridrow="2" pageId="11" pageNumber="344">1955.1.2.68</td>
<td id="76B05DA40B37000DC971E1E1FC6BFD53" box="[817,938,644,669]" gridcol="3" gridrow="2" pageId="11" pageNumber="344">1955.1.2.70</td>
<td id="76B05DA40B37000DC985E1E1FBFFFD53" box="[965,1086,644,669]" gridcol="4" gridrow="2" pageId="11" pageNumber="344">1955.1.2.71</td>
<td id="76B05DA40B37000DCE19E1E1FB10FD53" box="[1113,1233,644,669]" gridcol="5" gridrow="2" pageId="11" pageNumber="344">1955.1.2.72</td>
<td id="76B05DA40B37000DCEB1E1E1FAFCFD53" box="[1265,1341,644,669]" gridcol="6" gridrow="2" pageId="11" pageNumber="344">3765</td>
<td id="76B05DA40B37000DCF22E1E1FA6FFD53" box="[1378,1454,644,669]" gridcol="7" gridrow="2" pageId="11" pageNumber="344">3763</td>
<td id="76B05DA40B37000DCF92E1E1F9DEFD53" box="[1490,1567,644,669]" gridcol="8" gridrow="2" pageId="11" pageNumber="344">3764</td>
<td id="76B05DA40B37000DCC30E1E1F909FD53" box="[1648,1736,644,669]" gridcol="9" gridrow="2" pageId="11" pageNumber="344">64.3.9.1</td>
<td id="76B05DA40B37000DCCAFE1E1F88AFD53" box="[1775,1867,644,669]" gridcol="10" gridrow="2" pageId="11" pageNumber="344">1895.78</td>
<td id="76B05DA40B37000DCD2DE1E1F826FD53" box="[1901,2023,644,669]" gridcol="11" gridrow="2" pageId="11" pageNumber="344">83.1.12.34</td>
</tr>
<tr id="356134D80B37000DCAE2E1D3F826FD01" box="[162,2023,694,719]" gridrow="3" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E1D3FE30FD01" box="[162,497,694,719]" gridcol="0" gridrow="3" pageId="11" pageNumber="344">Sex</th>
<td id="76B05DA40B37000DC84AE1D3FD43FD01" box="[522,642,694,719]" gridcol="1" gridrow="3" pageId="11" pageNumber="344">F [M]</td>
<td id="76B05DA40B37000DC8DEE1D3FCD7FD01" box="[670,790,694,719]" gridcol="2" gridrow="3" pageId="11" pageNumber="344">F [M]</td>
<td id="76B05DA40B37000DC971E1D3FC6BFD01" box="[817,938,694,719]" gridcol="3" gridrow="3" pageId="11" pageNumber="344">M [F]</td>
<td id="76B05DA40B37000DC985E1D3FBFFFD01" box="[965,1086,694,719]" gridcol="4" gridrow="3" pageId="11" pageNumber="344">M [F]</td>
<td id="76B05DA40B37000DCE19E1D3FB10FD01" box="[1113,1233,694,719]" gridcol="5" gridrow="3" pageId="11" pageNumber="344">F [M]</td>
<td id="76B05DA40B37000DCEB1E1D3FAFCFD01" box="[1265,1341,694,719]" gridcol="6" gridrow="3" pageId="11" pageNumber="344">F</td>
<td id="76B05DA40B37000DCF22E1D3FA6FFD01" box="[1378,1454,694,719]" gridcol="7" gridrow="3" pageId="11" pageNumber="344">F</td>
<td id="76B05DA40B37000DCF92E1D3F9DEFD01" box="[1490,1567,694,719]" gridcol="8" gridrow="3" pageId="11" pageNumber="344">F</td>
<td id="76B05DA40B37000DCC30E1D3F909FD01" box="[1648,1736,694,719]" gridcol="9" gridrow="3" pageId="11" pageNumber="344">F</td>
<td id="76B05DA40B37000DCCAFE1D3F88AFD01" box="[1775,1867,694,719]" gridcol="10" gridrow="3" pageId="11" pageNumber="344">M</td>
<td id="76B05DA40B37000DCD2DE1D3F826FD01" box="[1901,2023,694,719]" gridcol="11" gridrow="3" pageId="11" pageNumber="344">M</td>
</tr>
<tr id="356134D80B37000DCAE2E18DF826FCCF" box="[162,2023,744,769]" gridrow="4" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E18DFE30FCCF" box="[162,497,744,769]" gridcol="0" gridrow="4" pageId="11" pageNumber="344">Total length</th>
<td id="76B05DA40B37000DC84AE18DFD43FCCF" box="[522,642,744,769]" gridcol="1" gridrow="4" pageId="11" pageNumber="344">245 [198]</td>
<td id="76B05DA40B37000DC8DEE18DFCD7FCCF" box="[670,790,744,769]" gridcol="2" gridrow="4" pageId="11" pageNumber="344">375 [215.5]</td>
<td id="76B05DA40B37000DC971E18DFC6BFCCF" box="[817,938,744,769]" gridcol="3" gridrow="4" pageId="11" pageNumber="344">213 [266]</td>
<td id="76B05DA40B37000DC985E18DFBFFFCCF" box="[965,1086,744,769]" gridcol="4" gridrow="4" pageId="11" pageNumber="344">212 [235]</td>
<td id="76B05DA40B37000DCE19E18DFB10FCCF" box="[1113,1233,744,769]" gridcol="5" gridrow="4" pageId="11" pageNumber="344">266 [209.5]</td>
<td id="76B05DA40B37000DCEB1E18DFAFCFCCF" box="[1265,1341,744,769]" gridcol="6" gridrow="4" pageId="11" pageNumber="344">320</td>
<td id="76B05DA40B37000DCF22E18DFA6FFCCF" box="[1378,1454,744,769]" gridcol="7" gridrow="4" pageId="11" pageNumber="344">305</td>
<td id="76B05DA40B37000DCF92E18DF9DEFCCF" box="[1490,1567,744,769]" gridcol="8" gridrow="4" pageId="11" pageNumber="344">242</td>
<td id="76B05DA40B37000DCC30E18DF909FCCF" box="[1648,1736,744,769]" gridcol="9" gridrow="4" pageId="11" pageNumber="344">215</td>
<td id="76B05DA40B37000DCCAFE18DF88AFCCF" box="[1775,1867,744,769]" gridcol="10" gridrow="4" pageId="11" pageNumber="344">356</td>
<td id="76B05DA40B37000DCD2DE18DF826FCCF" box="[1901,2023,744,769]" gridcol="11" gridrow="4" pageId="11" pageNumber="344">259</td>
</tr>
<tr id="356134D80B37000DCAE2E07FF826FCFD" box="[162,2023,794,819]" gridrow="5" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E07FFE30FCFD" box="[162,497,794,819]" gridcol="0" gridrow="5" pageId="11" pageNumber="344">Tail length</th>
<td id="76B05DA40B37000DC84AE07FFD43FCFD" box="[522,642,794,819]" gridcol="1" gridrow="5" pageId="11" pageNumber="344">7.6 [10]</td>
<td id="76B05DA40B37000DC8DEE07FFCD7FCFD" box="[670,790,794,819]" gridcol="2" gridrow="5" pageId="11" pageNumber="344">8.7 [10.5]</td>
<td id="76B05DA40B37000DC971E07FFC6BFCFD" box="[817,938,794,819]" gridcol="3" gridrow="5" pageId="11" pageNumber="344">10.9 [8]</td>
<td id="76B05DA40B37000DC985E07FFBFFFCFD" box="[965,1086,794,819]" gridcol="4" gridrow="5" pageId="11" pageNumber="344">10.2 [8]</td>
<td id="76B05DA40B37000DCE19E07FFB10FCFD" box="[1113,1233,794,819]" gridcol="5" gridrow="5" pageId="11" pageNumber="344">7.3 [10.5]</td>
<td id="76B05DA40B37000DCEB1E07FFAFCFCFD" box="[1265,1341,794,819]" gridcol="6" gridrow="5" pageId="11" pageNumber="344">8.9</td>
<td id="76B05DA40B37000DCF22E07FFA6FFCFD" box="[1378,1454,794,819]" gridcol="7" gridrow="5" pageId="11" pageNumber="344">9.2</td>
<td id="76B05DA40B37000DCF92E07FF9DEFCFD" box="[1490,1567,794,819]" gridcol="8" gridrow="5" pageId="11" pageNumber="344">6.9</td>
<td id="76B05DA40B37000DCC30E07FF909FCFD" box="[1648,1736,794,819]" gridcol="9" gridrow="5" pageId="11" pageNumber="344">5.8</td>
<td id="76B05DA40B37000DCCAFE07FF88AFCFD" box="[1775,1867,794,819]" gridcol="10" gridrow="5" pageId="11" pageNumber="344">17.5</td>
<td id="76B05DA40B37000DCD2DE07FF826FCFD" box="[1901,2023,794,819]" gridcol="11" gridrow="5" pageId="11" pageNumber="344">13.4</td>
</tr>
<tr id="356134D80B37000DCAE2E029F826FCAB" box="[162,2023,844,869]" gridrow="6" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E029FE30FCAB" box="[162,497,844,869]" gridcol="0" gridrow="6" pageId="11" pageNumber="344">Midbody circumference</th>
<td id="76B05DA40B37000DC84AE029FD43FCAB" box="[522,642,844,869]" gridcol="1" gridrow="6" pageId="11" pageNumber="344">22</td>
<td id="76B05DA40B37000DC8DEE029FCD7FCAB" box="[670,790,844,869]" gridcol="2" gridrow="6" pageId="11" pageNumber="344">32</td>
<td id="76B05DA40B37000DC971E029FC6BFCAB" box="[817,938,844,869]" gridcol="3" gridrow="6" pageId="11" pageNumber="344">20</td>
<td id="76B05DA40B37000DC985E029FBFFFCAB" box="[965,1086,844,869]" gridcol="4" gridrow="6" pageId="11" pageNumber="344">19</td>
<td id="76B05DA40B37000DCE19E029FB10FCAB" box="[1113,1233,844,869]" gridcol="5" gridrow="6" pageId="11" pageNumber="344">20</td>
<td id="76B05DA40B37000DCEB1E029FAFCFCAB" box="[1265,1341,844,869]" gridcol="6" gridrow="6" pageId="11" pageNumber="344">25</td>
<td id="76B05DA40B37000DCF22E029FA6FFCAB" box="[1378,1454,844,869]" gridcol="7" gridrow="6" pageId="11" pageNumber="344">23</td>
<td id="76B05DA40B37000DCF92E029F9DEFCAB" box="[1490,1567,844,869]" gridcol="8" gridrow="6" pageId="11" pageNumber="344">14</td>
<td id="76B05DA40B37000DCC30E029F909FCAB" box="[1648,1736,844,869]" gridcol="9" gridrow="6" pageId="11" pageNumber="344">21</td>
<td id="76B05DA40B37000DCCAFE029F88AFCAB" box="[1775,1867,844,869]" gridcol="10" gridrow="6" pageId="11" pageNumber="344">35</td>
<td id="76B05DA40B37000DCD2DE029F826FCAB" box="[1901,2023,844,869]" gridcol="11" gridrow="6" pageId="11" pageNumber="344">18</td>
</tr>
<tr id="356134D80B37000DCAE2E01BF826FC59" box="[162,2023,894,919]" gridrow="7" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E01BFE30FC59" box="[162,497,894,919]" gridcol="0" gridrow="7" pageId="11" pageNumber="344">Midbody width</th>
<td id="76B05DA40B37000DC84AE01BFD43FC59" box="[522,642,894,919]" gridcol="1" gridrow="7" pageId="11" pageNumber="344">7.1</td>
<td id="76B05DA40B37000DC8DEE01BFCD7FC59" box="[670,790,894,919]" gridcol="2" gridrow="7" pageId="11" pageNumber="344">10.3</td>
<td id="76B05DA40B37000DC971E01BFC6BFC59" box="[817,938,894,919]" gridcol="3" gridrow="7" pageId="11" pageNumber="344">6.3</td>
<td id="76B05DA40B37000DC985E01BFBFFFC59" box="[965,1086,894,919]" gridcol="4" gridrow="7" pageId="11" pageNumber="344">5.9</td>
<td id="76B05DA40B37000DCE19E01BFB10FC59" box="[1113,1233,894,919]" gridcol="5" gridrow="7" pageId="11" pageNumber="344">7.3</td>
<td id="76B05DA40B37000DCEB1E01BFAFCFC59" box="[1265,1341,894,919]" gridcol="6" gridrow="7" pageId="11" pageNumber="344">6.1</td>
<td id="76B05DA40B37000DCF22E01BFA6FFC59" box="[1378,1454,894,919]" gridcol="7" gridrow="7" pageId="11" pageNumber="344">7.3</td>
<td id="76B05DA40B37000DCF92E01BF9DEFC59" box="[1490,1567,894,919]" gridcol="8" gridrow="7" pageId="11" pageNumber="344">4.3</td>
<td id="76B05DA40B37000DCC30E01BF909FC59" box="[1648,1736,894,919]" gridcol="9" gridrow="7" pageId="11" pageNumber="344">6.9</td>
<td id="76B05DA40B37000DCCAFE01BF88AFC59" box="[1775,1867,894,919]" gridcol="10" gridrow="7" pageId="11" pageNumber="344">11.0</td>
<td id="76B05DA40B37000DCD2DE01BF826FC59" box="[1901,2023,894,919]" gridcol="11" gridrow="7" pageId="11" pageNumber="344">6.1</td>
</tr>
<tr id="356134D80B37000DCAE2E0D6F826FC02" box="[162,2023,947,972]" gridrow="8" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E0D6FE30FC02" box="[162,497,947,972]" gridcol="0" gridrow="8" pageId="11" pageNumber="344">Head length</th>
<td id="76B05DA40B37000DC84AE0D6FD43FC02" box="[522,642,947,972]" gridcol="1" gridrow="8" pageId="11" pageNumber="344">5.9</td>
<td id="76B05DA40B37000DC8DEE0D6FCD7FC02" box="[670,790,947,972]" gridcol="2" gridrow="8" pageId="11" pageNumber="344">7.7</td>
<td id="76B05DA40B37000DC971E0D6FC6BFC02" box="[817,938,947,972]" gridcol="3" gridrow="8" pageId="11" pageNumber="344">5.4</td>
<td id="76B05DA40B37000DC985E0D6FBFFFC02" box="[965,1086,947,972]" gridcol="4" gridrow="8" pageId="11" pageNumber="344">5.3</td>
<td id="76B05DA40B37000DCE19E0D6FB10FC02" box="[1113,1233,947,972]" gridcol="5" gridrow="8" pageId="11" pageNumber="344">6.1</td>
<td id="76B05DA40B37000DCEB1E0D6FAFCFC02" box="[1265,1341,947,972]" gridcol="6" gridrow="8" pageId="11" pageNumber="344">6.4</td>
<td id="76B05DA40B37000DCF22E0D6FA6FFC02" box="[1378,1454,947,972]" gridcol="7" gridrow="8" pageId="11" pageNumber="344">7.4</td>
<td id="76B05DA40B37000DCF92E0D6F9DEFC02" box="[1490,1567,947,972]" gridcol="8" gridrow="8" pageId="11" pageNumber="344">5.6</td>
<td id="76B05DA40B37000DCC30E0D6F909FC02" box="[1648,1736,947,972]" gridcol="9" gridrow="8" pageId="11" pageNumber="344">5.7</td>
<td id="76B05DA40B37000DCCAFE0D6F88AFC02" box="[1775,1867,947,972]" gridcol="10" gridrow="8" pageId="11" pageNumber="344">9.2</td>
<td id="76B05DA40B37000DCD2DE0D6F826FC02" box="[1901,2023,947,972]" gridcol="11" gridrow="8" pageId="11" pageNumber="344">5.9</td>
</tr>
<tr id="356134D80B37000DCAE2E080F826FC30" box="[162,2023,997,1022]" gridrow="9" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E080FE30FC30" box="[162,497,997,1022]" gridcol="0" gridrow="9" pageId="11" pageNumber="344">Head width</th>
<td id="76B05DA40B37000DC84AE080FD43FC30" box="[522,642,997,1022]" gridcol="1" gridrow="9" pageId="11" pageNumber="344">4.3</td>
<td id="76B05DA40B37000DC8DEE080FCD7FC30" box="[670,790,997,1022]" gridcol="2" gridrow="9" pageId="11" pageNumber="344">6.7</td>
<td id="76B05DA40B37000DC971E080FC6BFC30" box="[817,938,997,1022]" gridcol="3" gridrow="9" pageId="11" pageNumber="344">4.1</td>
<td id="76B05DA40B37000DC985E080FBFFFC30" box="[965,1086,997,1022]" gridcol="4" gridrow="9" pageId="11" pageNumber="344">3.5</td>
<td id="76B05DA40B37000DCE19E080FB10FC30" box="[1113,1233,997,1022]" gridcol="5" gridrow="9" pageId="11" pageNumber="344">4.1</td>
<td id="76B05DA40B37000DCEB1E080FAFCFC30" box="[1265,1341,997,1022]" gridcol="6" gridrow="9" pageId="11" pageNumber="344">4.2</td>
<td id="76B05DA40B37000DCF22E080FA6FFC30" box="[1378,1454,997,1022]" gridcol="7" gridrow="9" pageId="11" pageNumber="344">5.1</td>
<td id="76B05DA40B37000DCF92E080F9DEFC30" box="[1490,1567,997,1022]" gridcol="8" gridrow="9" pageId="11" pageNumber="344">3.4</td>
<td id="76B05DA40B37000DCC30E080F909FC30" box="[1648,1736,997,1022]" gridcol="9" gridrow="9" pageId="11" pageNumber="344">4.0</td>
<td id="76B05DA40B37000DCCAFE080F88AFC30" box="[1775,1867,997,1022]" gridcol="10" gridrow="9" pageId="11" pageNumber="344">6.5</td>
<td id="76B05DA40B37000DCD2DE080F826FC30" box="[1901,2023,997,1022]" gridcol="11" gridrow="9" pageId="11" pageNumber="344">4.5</td>
</tr>
<tr id="356134D80B37000DCAE2E773F826FBE1" box="[162,2023,1046,1071]" gridrow="10" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E773FE30FBE1" box="[162,497,1046,1071]" gridcol="0" gridrow="10" pageId="11" pageNumber="344">Supralabials; infralabials (left, right)</th>
<td id="76B05DA40B37000DC84AE773FD43FBE1" box="[522,642,1046,1071]" gridcol="1" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DC8DEE773FCD7FBE1" box="[670,790,1046,1071]" gridcol="2" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DC971E773FC6BFBE1" box="[817,938,1046,1071]" gridcol="3" gridrow="10" pageId="11" pageNumber="344">4,5; 3,3</td>
<td id="76B05DA40B37000DC985E773FBFFFBE1" box="[965,1086,1046,1071]" gridcol="4" gridrow="10" pageId="11" pageNumber="344">4,4; 4,4</td>
<td id="76B05DA40B37000DCE19E773FB10FBE1" box="[1113,1233,1046,1071]" gridcol="5" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DCEB1E773FAFCFBE1" box="[1265,1341,1046,1071]" gridcol="6" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DCF22E773FA6FFBE1" box="[1378,1454,1046,1071]" gridcol="7" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DCF92E773F9DEFBE1" box="[1490,1567,1046,1071]" gridcol="8" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DCC30E773F909FBE1" box="[1648,1736,1046,1071]" gridcol="9" gridrow="10" pageId="11" pageNumber="344">4,4; 2,3</td>
<td id="76B05DA40B37000DCCAFE773F88AFBE1" box="[1775,1867,1046,1071]" gridcol="10" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
<td id="76B05DA40B37000DCD2DE773F826FBE1" box="[1901,2023,1046,1071]" gridcol="11" gridrow="10" pageId="11" pageNumber="344">4,4; 3,3</td>
</tr>
<tr id="356134D80B37000DCAE2E72DF826FBAF" box="[162,2023,1096,1121]" gridrow="11" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E72DFE30FBAF" box="[162,497,1096,1121]" gridcol="0" gridrow="11" pageId="11" pageNumber="344">Ventrals</th>
<td id="76B05DA40B37000DC84AE72DFD43FBAF" box="[522,642,1096,1121]" gridcol="1" gridrow="11" pageId="11" pageNumber="344">187 [173]</td>
<td id="76B05DA40B37000DC8DEE72DFCD7FBAF" box="[670,790,1096,1121]" gridcol="2" gridrow="11" pageId="11" pageNumber="344">197 [173]</td>
<td id="76B05DA40B37000DC971E72DFC6BFBAF" box="[817,938,1096,1121]" gridcol="3" gridrow="11" pageId="11" pageNumber="344">185 [180]</td>
<td id="76B05DA40B37000DC985E72DFBFFFBAF" box="[965,1086,1096,1121]" gridcol="4" gridrow="11" pageId="11" pageNumber="344">187 [179]</td>
<td id="76B05DA40B37000DCE19E72DFB10FBAF" box="[1113,1233,1096,1121]" gridcol="5" gridrow="11" pageId="11" pageNumber="344">188 [176]</td>
<td id="76B05DA40B37000DCEB1E72DFAFCFBAF" box="[1265,1341,1096,1121]" gridcol="6" gridrow="11" pageId="11" pageNumber="344">187</td>
<td id="76B05DA40B37000DCF22E72DFA6FFBAF" box="[1378,1454,1096,1121]" gridcol="7" gridrow="11" pageId="11" pageNumber="344">187</td>
<td id="76B05DA40B37000DCF92E72DF9DEFBAF" box="[1490,1567,1096,1121]" gridcol="8" gridrow="11" pageId="11" pageNumber="344">188</td>
<td id="76B05DA40B37000DCC30E72DF909FBAF" box="[1648,1736,1096,1121]" gridcol="9" gridrow="11" pageId="11" pageNumber="344">177</td>
<td id="76B05DA40B37000DCCAFE72DF88AFBAF" box="[1775,1867,1096,1121]" gridcol="10" gridrow="11" pageId="11" pageNumber="344">175</td>
<td id="76B05DA40B37000DCD2DE72DF826FBAF" box="[1901,2023,1096,1121]" gridcol="11" gridrow="11" pageId="11" pageNumber="344">167</td>
</tr>
<tr id="356134D80B37000DCAE2E71FF826FB5D" box="[162,2023,1146,1171]" gridrow="12" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E71FFE30FB5D" box="[162,497,1146,1171]" gridcol="0" gridrow="12" pageId="11" pageNumber="344">Subcaudals (rows unpaired)</th>
<td id="76B05DA40B37000DC84AE71FFD43FB5D" box="[522,642,1146,1171]" gridcol="1" gridrow="12" pageId="11" pageNumber="344">8,7 (0) [12]</td>
<td id="76B05DA40B37000DC8DEE71FFCD7FB5D" box="[670,790,1146,1171]" gridcol="2" gridrow="12" pageId="11" pageNumber="344">7,7 (3) [11]</td>
<td id="76B05DA40B37000DC971E71FFC6BFB5D" box="[817,938,1146,1171]" gridcol="3" gridrow="12" pageId="11" pageNumber="344">12,12 (0) [7]</td>
<td id="76B05DA40B37000DC985E71FFBFFFB5D" box="[965,1086,1146,1171]" gridcol="4" gridrow="12" pageId="11" pageNumber="344">12,12 (0) [7]</td>
<td id="76B05DA40B37000DCE19E71FFB10FB5D" box="[1113,1233,1146,1171]" gridcol="5" gridrow="12" pageId="11" pageNumber="344">8,7 (0) [12]</td>
<td id="76B05DA40B37000DCEB1E71FFAFCFB5D" box="[1265,1341,1146,1171]" gridcol="6" gridrow="12" pageId="11" pageNumber="344">8,7 (0)</td>
<td id="76B05DA40B37000DCF22E71FFA6FFB5D" box="[1378,1454,1146,1171]" gridcol="7" gridrow="12" pageId="11" pageNumber="344">7,7 (0)</td>
<td id="76B05DA40B37000DCF92E71FF9DEFB5D" box="[1490,1567,1146,1171]" gridcol="8" gridrow="12" pageId="11" pageNumber="344">7,7 (0)</td>
<td id="76B05DA40B37000DCC30E71FF909FB5D" box="[1648,1736,1146,1171]" gridcol="9" gridrow="12" pageId="11" pageNumber="344">6,7 (0)</td>
<td id="76B05DA40B37000DCCAFE71FF88AFB5D" box="[1775,1867,1146,1171]" gridcol="10" gridrow="12" pageId="11" pageNumber="344">12,11 (0)</td>
<td id="76B05DA40B37000DCD2DE71FF826FB5D" box="[1901,2023,1146,1171]" gridcol="11" gridrow="12" pageId="11" pageNumber="344">12,12 (0)</td>
</tr>
<tr id="356134D80B37000DCAE2E7C9F826FB0B" box="[162,2023,1196,1221]" gridrow="13" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E7C9FE30FB0B" box="[162,497,1196,1221]" gridcol="0" gridrow="13" pageId="11" pageNumber="344">Scales around base of terminal scute</th>
<td id="76B05DA40B37000DC84AE7C9FD43FB0B" box="[522,642,1196,1221]" gridcol="1" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DC8DEE7C9FCD7FB0B" box="[670,790,1196,1221]" gridcol="2" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DC971E7C9FC6BFB0B" box="[817,938,1196,1221]" gridcol="3" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DC985E7C9FBFFFB0B" box="[965,1086,1196,1221]" gridcol="4" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DCE19E7C9FB10FB0B" box="[1113,1233,1196,1221]" gridcol="5" gridrow="13" pageId="11" pageNumber="344">8</td>
<td id="76B05DA40B37000DCEB1E7C9FAFCFB0B" box="[1265,1341,1196,1221]" gridcol="6" gridrow="13" pageId="11" pageNumber="344">8</td>
<td id="76B05DA40B37000DCF22E7C9FA6FFB0B" box="[1378,1454,1196,1221]" gridcol="7" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DCF92E7C9F9DEFB0B" box="[1490,1567,1196,1221]" gridcol="8" gridrow="13" pageId="11" pageNumber="344">9</td>
<td id="76B05DA40B37000DCC30E7C9F909FB0B" box="[1648,1736,1196,1221]" gridcol="9" gridrow="13" pageId="11" pageNumber="344">8</td>
<td id="76B05DA40B37000DCCAFE7C9F88AFB0B" box="[1775,1867,1196,1221]" gridcol="10" gridrow="13" pageId="11" pageNumber="344">8</td>
<td id="76B05DA40B37000DCD2DE7C9F826FB0B" box="[1901,2023,1196,1221]" gridcol="11" gridrow="13" pageId="11" pageNumber="344">8</td>
</tr>
<tr id="356134D80B37000DCAE2E7BBF826FB39" box="[162,2023,1246,1271]" gridrow="14" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E7BBFE30FB39" box="[162,497,1246,1271]" gridcol="0" gridrow="14" pageId="11" pageNumber="344">Rostral length</th>
<td id="76B05DA40B37000DC84AE7BBFD43FB39" box="[522,642,1246,1271]" gridcol="1" gridrow="14" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DC8DEE7BBFCD7FB39" box="[670,790,1246,1271]" gridcol="2" gridrow="14" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DC971E7BBFC6BFB39" box="[817,938,1246,1271]" gridcol="3" gridrow="14" pageId="11" pageNumber="344">1.5</td>
<td id="76B05DA40B37000DC985E7BBFBFFFB39" box="[965,1086,1246,1271]" gridcol="4" gridrow="14" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCE19E7BBFB10FB39" box="[1113,1233,1246,1271]" gridcol="5" gridrow="14" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCEB1E7BBFAFCFB39" box="[1265,1341,1246,1271]" gridcol="6" gridrow="14" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCF22E7BBFA6FFB39" box="[1378,1454,1246,1271]" gridcol="7" gridrow="14" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DCF92E7BBF9DEFB39" box="[1490,1567,1246,1271]" gridcol="8" gridrow="14" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCC30E7BBF909FB39" box="[1648,1736,1246,1271]" gridcol="9" gridrow="14" pageId="11" pageNumber="344">1.4</td>
<td id="76B05DA40B37000DCCAFE7BBF88AFB39" box="[1775,1867,1246,1271]" gridcol="10" gridrow="14" pageId="11" pageNumber="344">2.4</td>
<td id="76B05DA40B37000DCD2DE7BBF826FB39" box="[1901,2023,1246,1271]" gridcol="11" gridrow="14" pageId="11" pageNumber="344">1.6</td>
</tr>
<tr id="356134D80B37000DCAE2E675F826FAE7" box="[162,2023,1296,1321]" gridrow="15" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E675FE30FAE7" box="[162,497,1296,1321]" gridcol="0" gridrow="15" pageId="11" pageNumber="344">Rostral-frontal gap</th>
<td id="76B05DA40B37000DC84AE675FD43FAE7" box="[522,642,1296,1321]" gridcol="1" gridrow="15" pageId="11" pageNumber="344">0.6</td>
<td id="76B05DA40B37000DC8DEE675FCD7FAE7" box="[670,790,1296,1321]" gridcol="2" gridrow="15" pageId="11" pageNumber="344">0.9</td>
<td id="76B05DA40B37000DC971E675FC6BFAE7" box="[817,938,1296,1321]" gridcol="3" gridrow="15" pageId="11" pageNumber="344">0.8</td>
<td id="76B05DA40B37000DC985E675FBFFFAE7" box="[965,1086,1296,1321]" gridcol="4" gridrow="15" pageId="11" pageNumber="344">0.6</td>
<td id="76B05DA40B37000DCE19E675FB10FAE7" box="[1113,1233,1296,1321]" gridcol="5" gridrow="15" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DCEB1E675FAFCFAE7" box="[1265,1341,1296,1321]" gridcol="6" gridrow="15" pageId="11" pageNumber="344">0.6</td>
<td id="76B05DA40B37000DCF22E675FA6FFAE7" box="[1378,1454,1296,1321]" gridcol="7" gridrow="15" pageId="11" pageNumber="344">0.9</td>
<td id="76B05DA40B37000DCF92E675F9DEFAE7" box="[1490,1567,1296,1321]" gridcol="8" gridrow="15" pageId="11" pageNumber="344">0.5</td>
<td id="76B05DA40B37000DCC30E675F909FAE7" box="[1648,1736,1296,1321]" gridcol="9" gridrow="15" pageId="11" pageNumber="344">0.8</td>
<td id="76B05DA40B37000DCCAFE675F88AFAE7" box="[1775,1867,1296,1321]" gridcol="10" gridrow="15" pageId="11" pageNumber="344">1.1</td>
<td id="76B05DA40B37000DCD2DE675F826FAE7" box="[1901,2023,1296,1321]" gridcol="11" gridrow="15" pageId="11" pageNumber="344">0.6</td>
</tr>
<tr id="356134D80B37000DCAE2E627F826FA95" box="[162,2023,1346,1371]" gridrow="16" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E627FE30FA95" box="[162,497,1346,1371]" gridcol="0" gridrow="16" pageId="11" pageNumber="344">Frontal length</th>
<td id="76B05DA40B37000DC84AE627FD43FA95" box="[522,642,1346,1371]" gridcol="1" gridrow="16" pageId="11" pageNumber="344">2.7</td>
<td id="76B05DA40B37000DC8DEE627FCD7FA95" box="[670,790,1346,1371]" gridcol="2" gridrow="16" pageId="11" pageNumber="344">3.3</td>
<td id="76B05DA40B37000DC971E627FC6BFA95" box="[817,938,1346,1371]" gridcol="3" gridrow="16" pageId="11" pageNumber="344">2.3</td>
<td id="76B05DA40B37000DC985E627FBFFFA95" box="[965,1086,1346,1371]" gridcol="4" gridrow="16" pageId="11" pageNumber="344">2.3</td>
<td id="76B05DA40B37000DCE19E627FB10FA95" box="[1113,1233,1346,1371]" gridcol="5" gridrow="16" pageId="11" pageNumber="344">2.7</td>
<td id="76B05DA40B37000DCEB1E627FAFCFA95" box="[1265,1341,1346,1371]" gridcol="6" gridrow="16" pageId="11" pageNumber="344">2.6</td>
<td id="76B05DA40B37000DCF22E627FA6FFA95" box="[1378,1454,1346,1371]" gridcol="7" gridrow="16" pageId="11" pageNumber="344">3.0</td>
<td id="76B05DA40B37000DCF92E627F9DEFA95" box="[1490,1567,1346,1371]" gridcol="8" gridrow="16" pageId="11" pageNumber="344">2.3</td>
<td id="76B05DA40B37000DCC30E627F909FA95" box="[1648,1736,1346,1371]" gridcol="9" gridrow="16" pageId="11" pageNumber="344">2.2</td>
<td id="76B05DA40B37000DCCAFE627F88AFA95" box="[1775,1867,1346,1371]" gridcol="10" gridrow="16" pageId="11" pageNumber="344">3.5</td>
<td id="76B05DA40B37000DCD2DE627F826FA95" box="[1901,2023,1346,1371]" gridcol="11" gridrow="16" pageId="11" pageNumber="344">2.7</td>
</tr>
<tr id="356134D80B37000DCAE2E611F826FA43" box="[162,2023,1396,1421]" gridrow="17" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E611FE30FA43" box="[162,497,1396,1421]" gridcol="0" gridrow="17" pageId="11" pageNumber="344">Frontal width</th>
<td id="76B05DA40B37000DC84AE611FD43FA43" box="[522,642,1396,1421]" gridcol="1" gridrow="17" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DC8DEE611FCD7FA43" box="[670,790,1396,1421]" gridcol="2" gridrow="17" pageId="11" pageNumber="344">2.6</td>
<td id="76B05DA40B37000DC971E611FC6BFA43" box="[817,938,1396,1421]" gridcol="3" gridrow="17" pageId="11" pageNumber="344">1.8</td>
<td id="76B05DA40B37000DC985E611FBFFFA43" box="[965,1086,1396,1421]" gridcol="4" gridrow="17" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCE19E611FB10FA43" box="[1113,1233,1396,1421]" gridcol="5" gridrow="17" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DCEB1E611FAFCFA43" box="[1265,1341,1396,1421]" gridcol="6" gridrow="17" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCF22E611FA6FFA43" box="[1378,1454,1396,1421]" gridcol="7" gridrow="17" pageId="11" pageNumber="344">2.4</td>
<td id="76B05DA40B37000DCF92E611F9DEFA43" box="[1490,1567,1396,1421]" gridcol="8" gridrow="17" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCC30E611F909FA43" box="[1648,1736,1396,1421]" gridcol="9" gridrow="17" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCCAFE611F88AFA43" box="[1775,1867,1396,1421]" gridcol="10" gridrow="17" pageId="11" pageNumber="344">2.9</td>
<td id="76B05DA40B37000DCD2DE611F826FA43" box="[1901,2023,1396,1421]" gridcol="11" gridrow="17" pageId="11" pageNumber="344">1.9</td>
</tr>
<tr id="356134D80B37000DCAE2E6C3F826FA71" box="[162,2023,1446,1471]" gridrow="18" pageId="11" pageNumber="344" rowspan-10="1">
<th id="76B05DA40B37000DCAE2E6C3FE30FA71" box="[162,497,1446,1471]" gridcol="0" gridrow="18" pageId="11" pageNumber="344">Eye diameter</th>
<td id="76B05DA40B37000DC84AE6C3FD43FA71" box="[522,642,1446,1471]" gridcol="1" gridrow="18" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DC8DEE6C3FCD7FA71" box="[670,790,1446,1471]" gridcol="2" gridrow="18" pageId="11" pageNumber="344">0.8</td>
<td id="76B05DA40B37000DC971E6C3FC6BFA71" box="[817,938,1446,1471]" gridcol="3" gridrow="18" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DC985E6C3FBFFFA71" box="[965,1086,1446,1471]" gridcol="4" gridrow="18" pageId="11" pageNumber="344">0.6</td>
<td id="76B05DA40B37000DCE19E6C3FB10FA71" box="[1113,1233,1446,1471]" gridcol="5" gridrow="18" pageId="11" pageNumber="344">0.6</td>
<td id="76B05DA40B37000DCEB1E6C3FAFCFA71" box="[1265,1341,1446,1471]" gridcol="6" gridrow="18" pageId="11" pageNumber="344">0.8</td>
<td id="76B05DA40B37000DCF22E6C3FA6FFA71" box="[1378,1454,1446,1471]" gridcol="7" gridrow="18" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DCF92E6C3F9DEFA71" box="[1490,1567,1446,1471]" gridcol="8" gridrow="18" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DCC30E6C3F909FA71" box="[1648,1736,1446,1471]" gridcol="9" gridrow="18" pageId="11" pageNumber="344">0.7</td>
<td id="76B05DA40B37000DCD2DE6C3F826FA71" box="[1901,2023,1446,1471]" gridcol="11" gridrow="18" pageId="11" pageNumber="344">0.9</td>
</tr>
<tr id="356134D80B37000DCAE2E6B2F826FA3E" box="[162,2023,1495,1520]" gridrow="19" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E6B2FE30FA3E" box="[162,497,1495,1520]" gridcol="0" gridrow="19" pageId="11" pageNumber="344">Ocular length</th>
<td id="76B05DA40B37000DC84AE6B2FD43FA3E" box="[522,642,1495,1520]" gridcol="1" gridrow="19" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DC8DEE6B2FCD7FA3E" box="[670,790,1495,1520]" gridcol="2" gridrow="19" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DC971E6B2FC6BFA3E" box="[817,938,1495,1520]" gridcol="3" gridrow="19" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DC985E6B2FBFFFA3E" box="[965,1086,1495,1520]" gridcol="4" gridrow="19" pageId="11" pageNumber="344">1.4</td>
<td id="76B05DA40B37000DCE19E6B2FB10FA3E" box="[1113,1233,1495,1520]" gridcol="5" gridrow="19" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DCEB1E6B2FAFCFA3E" box="[1265,1341,1495,1520]" gridcol="6" gridrow="19" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DCF22E6B2FA6FFA3E" box="[1378,1454,1495,1520]" gridcol="7" gridrow="19" pageId="11" pageNumber="344">1.8</td>
<td id="76B05DA40B37000DCF92E6B2F9DEFA3E" box="[1490,1567,1495,1520]" gridcol="8" gridrow="19" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCC30E6B2F909FA3E" box="[1648,1736,1495,1520]" gridcol="9" gridrow="19" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCCAFE6B2F88AFA3E" box="[1775,1867,1495,1520]" gridcol="10" gridrow="19" pageId="11" pageNumber="344">2.7</td>
<td id="76B05DA40B37000DCD2DE6B2F826FA3E" box="[1901,2023,1495,1520]" gridcol="11" gridrow="19" pageId="11" pageNumber="344">1.8</td>
</tr>
<tr id="356134D80B37000DCAE2E56CF826F9EC" box="[162,2023,1545,1570]" gridrow="20" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E56CFE30F9EC" box="[162,497,1545,1570]" gridcol="0" gridrow="20" pageId="11" pageNumber="344">Terminal scute middorsal length</th>
<td id="76B05DA40B37000DC84AE56CFD43F9EC" box="[522,642,1545,1570]" gridcol="1" gridrow="20" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DC8DEE56CFCD7F9EC" box="[670,790,1545,1570]" gridcol="2" gridrow="20" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DC971E56CFC6BF9EC" box="[817,938,1545,1570]" gridcol="3" gridrow="20" pageId="11" pageNumber="344">1.0</td>
<td id="76B05DA40B37000DC985E56CFBFFF9EC" box="[965,1086,1545,1570]" gridcol="4" gridrow="20" pageId="11" pageNumber="344">1.1</td>
<td id="76B05DA40B37000DCE19E56CFB10F9EC" box="[1113,1233,1545,1570]" gridcol="5" gridrow="20" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCEB1E56CFAFCF9EC" box="[1265,1341,1545,1570]" gridcol="6" gridrow="20" pageId="11" pageNumber="344">1.8</td>
<td id="76B05DA40B37000DCF22E56CFA6FF9EC" box="[1378,1454,1545,1570]" gridcol="7" gridrow="20" pageId="11" pageNumber="344">1.2</td>
<td id="76B05DA40B37000DCF92E56CF9DEF9EC" box="[1490,1567,1545,1570]" gridcol="8" gridrow="20" pageId="11" pageNumber="344">1.1</td>
<td id="76B05DA40B37000DCC30E56CF909F9EC" box="[1648,1736,1545,1570]" gridcol="9" gridrow="20" pageId="11" pageNumber="344">1.0</td>
<td id="76B05DA40B37000DCCAFE56CF88AF9EC" box="[1775,1867,1545,1570]" gridcol="10" gridrow="20" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCD2DE56CF826F9EC" box="[1901,2023,1545,1570]" gridcol="11" gridrow="20" pageId="11" pageNumber="344">1.1</td>
</tr>
<tr id="356134D80B37000DCAE2E55EF826F99A" box="[162,2023,1595,1620]" gridrow="21" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E55EFE30F99A" box="[162,497,1595,1620]" gridcol="0" gridrow="21" pageId="11" pageNumber="344">Terminal scute midventral length</th>
<td id="76B05DA40B37000DC84AE55EFD43F99A" box="[522,642,1595,1620]" gridcol="1" gridrow="21" pageId="11" pageNumber="344">1.3</td>
<td id="76B05DA40B37000DC8DEE55EFCD7F99A" box="[670,790,1595,1620]" gridcol="2" gridrow="21" pageId="11" pageNumber="344">1.4</td>
<td id="76B05DA40B37000DC971E55EFC6BF99A" box="[817,938,1595,1620]" gridcol="3" gridrow="21" pageId="11" pageNumber="344">1.1</td>
<td id="76B05DA40B37000DC985E55EFBFFF99A" box="[965,1086,1595,1620]" gridcol="4" gridrow="21" pageId="11" pageNumber="344">1.2</td>
<td id="76B05DA40B37000DCE19E55EFB10F99A" box="[1113,1233,1595,1620]" gridcol="5" gridrow="21" pageId="11" pageNumber="344">1.1</td>
<td id="76B05DA40B37000DCEB1E55EFAFCF99A" box="[1265,1341,1595,1620]" gridcol="6" gridrow="21" pageId="11" pageNumber="344">1.2</td>
<td id="76B05DA40B37000DCF22E55EFA6FF99A" box="[1378,1454,1595,1620]" gridcol="7" gridrow="21" pageId="11" pageNumber="344">1.8</td>
<td id="76B05DA40B37000DCF92E55EF9DEF99A" box="[1490,1567,1595,1620]" gridcol="8" gridrow="21" pageId="11" pageNumber="344">1.3</td>
<td id="76B05DA40B37000DCC30E55EF909F99A" box="[1648,1736,1595,1620]" gridcol="9" gridrow="21" pageId="11" pageNumber="344">1.3</td>
<td id="76B05DA40B37000DCCAFE55EF88AF99A" box="[1775,1867,1595,1620]" gridcol="10" gridrow="21" pageId="11" pageNumber="344">1.9</td>
<td id="76B05DA40B37000DCD2DE55EF826F99A" box="[1901,2023,1595,1620]" gridcol="11" gridrow="21" pageId="11" pageNumber="344">1.1</td>
</tr>
<tr id="356134D80B37000DCAE2E508F826F948" box="[162,2023,1645,1670]" gridrow="22" pageId="11" pageNumber="344">
<th id="76B05DA40B37000DCAE2E508FE30F948" box="[162,497,1645,1670]" gridcol="0" gridrow="22" pageId="11" pageNumber="344">Terminal scute height</th>
<td id="76B05DA40B37000DC84AE508FD43F948" box="[522,642,1645,1670]" gridcol="1" gridrow="22" pageId="11" pageNumber="344">2.2</td>
<td id="76B05DA40B37000DC8DEE508FCD7F948" box="[670,790,1645,1670]" gridcol="2" gridrow="22" pageId="11" pageNumber="344">3.0</td>
<td id="76B05DA40B37000DC971E508FC6BF948" box="[817,938,1645,1670]" gridcol="3" gridrow="22" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DC985E508FBFFF948" box="[965,1086,1645,1670]" gridcol="4" gridrow="22" pageId="11" pageNumber="344">1.6</td>
<td id="76B05DA40B37000DCE19E508FB10F948" box="[1113,1233,1645,1670]" gridcol="5" gridrow="22" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DCEB1E508FAFCF948" box="[1265,1341,1645,1670]" gridcol="6" gridrow="22" pageId="11" pageNumber="344">2.1</td>
<td id="76B05DA40B37000DCF22E508FA6FF948" box="[1378,1454,1645,1670]" gridcol="7" gridrow="22" pageId="11" pageNumber="344">2.3</td>
<td id="76B05DA40B37000DCF92E508F9DEF948" box="[1490,1567,1645,1670]" gridcol="8" gridrow="22" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DCC30E508F909F948" box="[1648,1736,1645,1670]" gridcol="9" gridrow="22" pageId="11" pageNumber="344">1.7</td>
<td id="76B05DA40B37000DCCAFE508F88AF948" box="[1775,1867,1645,1670]" gridcol="10" gridrow="22" pageId="11" pageNumber="344">2.0</td>
<td id="76B05DA40B37000DCD2DE508F826F948" box="[1901,2023,1645,1670]" gridcol="11" gridrow="22" pageId="11" pageNumber="344">1.7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEE369A0B37FFF9CCF1E5C5F829F977" blockId="11.[1713,2024,1696,1721]" box="[1713,2024,1696,1721]" pageId="11" pageNumber="344">
<tableNote id="76B737140B37FFF9CCF1E5C5F829F977" box="[1713,2024,1696,1721]" pageId="11" pageNumber="344" targetBox="[162,2023,545,1670]" targetPageId="11">......continued on the next page</tableNote>
</paragraph>
<caption id="DF2E66120B30FFFECAD7E0F7FE44FC65" box="[151,389,914,939]" isContinuationCaption="true" pageId="12" pageNumber="345" startId="12.[151,235,914,939]" targetType="table">
<paragraph id="8BEE369A0B30FFFECAD7E0F7FE44FC65" blockId="12.[151,389,914,939]" box="[151,389,914,939]" pageId="12" pageNumber="345">
<emphasis id="B925EA880B30FFFECAD7E0F7FF3FFC65" bold="true" box="[151,254,914,939]" pageId="12" pageNumber="345">TABLE 3</emphasis>
. (Continued)
</paragraph>
</caption>
<paragraph id="8BEE369A0B30FFFEC91FE0DFF866FB30" pageId="12" pageNumber="345">
<table id="F951C43A0B30000DCAD7E0DFF81DFB30" box="[151,2012,954,1278]" colsContinueFrom="11.[162,2023,545,1670]" gridcols="12" gridrows="6" pageId="12" pageNumber="345">
<tr id="356134D80B30000DCAD7E0DFF81DFC1D" box="[151,2012,954,979]" gridrow="0" pageId="12" pageNumber="345" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76B05DA40B30000DC966E0DFFB07FC1D" box="[806,1222,954,979]" colspan="3" colspanRight="2" gridcol="3" gridrow="0" pageId="12" pageNumber="345">
<taxonomicName id="4C514D190B30FFFEC91FE0DFFBA1FC1D" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[863,1120,954,979]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="12" pageNumber="345" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B30FFFEC91FE0DFFBA1FC1D" box="[863,1120,954,979]" italics="true" pageId="12" pageNumber="345">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B30FFFECE27E0DFFB76FC1D" bold="true" box="[1127,1207,954,979]" pageId="12" pageNumber="345">
<taxonomicNameLabel id="A21657F30B30FFFECE27E0DFFB76FC1D" box="[1127,1207,954,979]" pageId="12" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76B05DA40B30000DCC24E0DFF8FFFC1D" box="[1636,1854,954,979]" colspan="2" colspanRight="1" gridcol="9" gridrow="0" pageId="12" pageNumber="345">
<taxonomicName id="4C514D190B30FFFECCC4E0DFF8D8FC1D" box="[1668,1817,954,979]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="12" pageNumber="345" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B30FFFECCC4E0DFF8D8FC1D" box="[1668,1817,954,979]" italics="true" pageId="12" pageNumber="345">U. pulneyensis</emphasis>
</taxonomicName>
</th>
<th id="76B05DA40B30000DCD22E0DFF81DFC1D" box="[1890,2012,954,979]" gridcol="11" gridrow="0" pageId="12" pageNumber="345">
<taxonomicName id="4C514D190B30FFFECD23E0DFF81DFC1D" authority="* BMNH" authorityName="BMNH" box="[1891,2012,954,979]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="12" pageNumber="345" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B30FFFECD23E0DFF81DFC1D" box="[1891,2012,954,979]" italics="true" pageId="12" pageNumber="345">U. guentheri</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="356134D80B30000DCAD7E086F81DFBEA" box="[151,2012,995,1060]" gridrow="1" pageId="12" pageNumber="345" rowspan-0="1">
<td id="76B05DA40B30000DCBBFE086FDB6FBEA" box="[511,631,995,1060]" gridcol="1" gridrow="1" pageId="12" pageNumber="345">*BMNH 1955.1.2.69</td>
<td id="76B05DA40B30000DC8D2E086FCCAFBEA" box="[658,779,995,1060]" gridcol="2" gridrow="1" pageId="12" pageNumber="345">BMNH 1955.1.2.68</td>
<td id="76B05DA40B30000DC966E086FC5FFBEA" box="[806,926,995,1060]" gridcol="3" gridrow="1" pageId="12" pageNumber="345">BMNH 1955.1.2.70</td>
<td id="76B05DA40B30000DC9FAE086FBF2FBEA" box="[954,1075,995,1060]" gridcol="4" gridrow="1" pageId="12" pageNumber="345">BMNH 1955.1.2.71</td>
<td id="76B05DA40B30000DCE0EE086FB07FBEA" box="[1102,1222,995,1060]" gridcol="5" gridrow="1" pageId="12" pageNumber="345">BMNH 1955.1.2.72</td>
<td id="76B05DA40B30000DCEAAE086FAEFFBEA" box="[1258,1326,995,1060]" gridcol="6" gridrow="1" pageId="12" pageNumber="345">BNHS 3765</td>
<td id="76B05DA40B30000DCF1BE086FA5EFBEA" box="[1371,1439,995,1060]" gridcol="7" gridrow="1" pageId="12" pageNumber="345">BNHS 3763</td>
<td id="76B05DA40B30000DCF8BE086F9CEFBEA" box="[1483,1551,995,1060]" gridcol="8" gridrow="1" pageId="12" pageNumber="345">BNHS 3764</td>
<td id="76B05DA40B30000DCC24E086F97CFBEA" box="[1636,1725,995,1060]" gridcol="9" gridrow="1" pageId="12" pageNumber="345">†BMNH 64.3.9.1</td>
<td id="76B05DA40B30000DCCA4E086F8FFFBEA" box="[1764,1854,995,1060]" gridcol="10" gridrow="1" pageId="12" pageNumber="345">#MNHN 1895.78</td>
<td id="76B05DA40B30000DCD22E086F81DFBEA" box="[1890,2012,995,1060]" gridcol="11" gridrow="1" pageId="12" pageNumber="345">‡BMNH 83.1.12.34</td>
</tr>
<tr id="356134D80B30000DCAD7E751F81DFB83" box="[151,2012,1076,1101]" gridrow="2" pageId="12" pageNumber="345">
<th id="76B05DA40B30000DCAD7E751FEB1FB83" box="[151,368,1076,1101]" gridcol="0" gridrow="2" pageId="12" pageNumber="345">Terminal scute width</th>
<td id="76B05DA40B30000DCBBFE751FDB6FB83" box="[511,631,1076,1101]" gridcol="1" gridrow="2" pageId="12" pageNumber="345">2.0</td>
<td id="76B05DA40B30000DC8D2E751FCCAFB83" box="[658,779,1076,1101]" gridcol="2" gridrow="2" pageId="12" pageNumber="345">2.4</td>
<td id="76B05DA40B30000DC966E751FC5FFB83" box="[806,926,1076,1101]" gridcol="3" gridrow="2" pageId="12" pageNumber="345">1.5</td>
<td id="76B05DA40B30000DC9FAE751FBF2FB83" box="[954,1075,1076,1101]" gridcol="4" gridrow="2" pageId="12" pageNumber="345">1.4</td>
<td id="76B05DA40B30000DCE0EE751FB07FB83" box="[1102,1222,1076,1101]" gridcol="5" gridrow="2" pageId="12" pageNumber="345">2.0</td>
<td id="76B05DA40B30000DCEAAE751FAEFFB83" box="[1258,1326,1076,1101]" gridcol="6" gridrow="2" pageId="12" pageNumber="345">1.5</td>
<td id="76B05DA40B30000DCF1BE751FA5EFB83" box="[1371,1439,1076,1101]" gridcol="7" gridrow="2" pageId="12" pageNumber="345">2.2</td>
<td id="76B05DA40B30000DCF8BE751F9CEFB83" box="[1483,1551,1076,1101]" gridcol="8" gridrow="2" pageId="12" pageNumber="345">1.5</td>
<td id="76B05DA40B30000DCC24E751F97CFB83" box="[1636,1725,1076,1101]" gridcol="9" gridrow="2" pageId="12" pageNumber="345">1.6</td>
<td id="76B05DA40B30000DCCA4E751F8FFFB83" box="[1764,1854,1076,1101]" gridcol="10" gridrow="2" pageId="12" pageNumber="345">1.8</td>
<td id="76B05DA40B30000DCD22E751F81DFB83" box="[1890,2012,1076,1101]" gridcol="11" gridrow="2" pageId="12" pageNumber="345">1.7</td>
</tr>
<tr id="356134D80B30000DCAD7E703F81DFB6D" box="[151,2012,1126,1187]" gridrow="3" pageId="12" pageNumber="345">
<th id="76B05DA40B30000DCAD7E703FEB1FB6D" box="[151,368,1126,1187]" gridcol="0" gridrow="3" pageId="12" pageNumber="345">Contact between first infralabials?</th>
<td id="76B05DA40B30000DCBBFE703FDB6FB6D" box="[511,631,1126,1187]" gridcol="1" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DC8D2E703FCCAFB6D" box="[658,779,1126,1187]" gridcol="2" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DC966E703FC5FFB6D" box="[806,926,1126,1187]" gridcol="3" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DC9FAE703FBF2FB6D" box="[954,1075,1126,1187]" gridcol="4" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCE0EE703FB07FB6D" box="[1102,1222,1126,1187]" gridcol="5" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCEAAE703FAEFFB6D" box="[1258,1326,1126,1187]" gridcol="6" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCF1BE703FA5EFB6D" box="[1371,1439,1126,1187]" gridcol="7" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCF8BE703F9CEFB6D" box="[1483,1551,1126,1187]" gridcol="8" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCC24E703F97CFB6D" box="[1636,1725,1126,1187]" gridcol="9" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCCA4E703F8FFFB6D" box="[1764,1854,1126,1187]" gridcol="10" gridrow="3" pageId="12" pageNumber="345">Y</td>
<td id="76B05DA40B30000DCD22E703F81DFB6D" box="[1890,2012,1126,1187]" gridcol="11" gridrow="3" pageId="12" pageNumber="345">N</td>
</tr>
<tr id="356134D80B30000DCAD7E7D6F81DFB02" box="[151,2012,1203,1228]" gridrow="4" pageId="12" pageNumber="345" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76B05DA40B30000DCAD7E7D6FEB1FB02" box="[151,368,1203,1228]" gridcol="0" gridrow="4" pageId="12" pageNumber="345">Maxillary teeth</th>
<td id="76B05DA40B30000DCBBFE7D6FDB6FB02" box="[511,631,1203,1228]" gridcol="1" gridrow="4" pageId="12" pageNumber="345">7,7</td>
<td id="76B05DA40B30000DC9FAE7D6FBF2FB02" box="[954,1075,1203,1228]" gridcol="4" gridrow="4" pageId="12" pageNumber="345">7,7</td>
<td id="76B05DA40B30000DCC24E7D6F97CFB02" box="[1636,1725,1203,1228]" gridcol="9" gridrow="4" pageId="12" pageNumber="345">7,7</td>
<td id="76B05DA40B30000DCD22E7D6F81DFB02" box="[1890,2012,1203,1228]" gridcol="11" gridrow="4" pageId="12" pageNumber="345">7,7</td>
</tr>
<tr id="356134D80B30000DCAD7E780F81DFB30" box="[151,2012,1253,1278]" gridrow="5" pageId="12" pageNumber="345" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76B05DA40B30000DCAD7E780FEB1FB30" box="[151,368,1253,1278]" gridcol="0" gridrow="5" pageId="12" pageNumber="345">Dentary teeth</th>
<td id="76B05DA40B30000DCBBFE780FDB6FB30" box="[511,631,1253,1278]" gridcol="1" gridrow="5" pageId="12" pageNumber="345">8,8</td>
<td id="76B05DA40B30000DC9FAE780FBF2FB30" box="[954,1075,1253,1278]" gridcol="4" gridrow="5" pageId="12" pageNumber="345">8,8</td>
<td id="76B05DA40B30000DCC24E780F97CFB30" box="[1636,1725,1253,1278]" gridcol="9" gridrow="5" pageId="12" pageNumber="345">8,8</td>
<td id="76B05DA40B30000DCD22E780F81DFB30" box="[1890,2012,1253,1278]" gridcol="11" gridrow="5" pageId="12" pageNumber="345">8,8</td>
</tr>
</table>
</paragraph>
<caption id="DF2E66120B31FFFFCAD7E3F3FB62FF37" ID-Table-UUID="DF2E66120B31FFFFCAD7E3F3FB62FF37" httpUri="http://table.plazi.org/id/DF2E66120B31FFFFCAD7E3F3FB62FF37" pageId="13" pageNumber="346" startId="13.[151,244,150,176]" targetBox="[163,1405,265,1979]" targetIsTable="true" targetPageId="13" targetType="table">
<paragraph id="8BEE369A0B31FFFFCAD7E3F3FB62FF37" blockId="13.[151,1436,150,249]" pageId="13" pageNumber="346">
<emphasis id="B925EA880B31FFFFCAD7E3F3FED5FF7E" bold="true" box="[151,276,150,176]" pageId="13" pageNumber="346">TABLE 4.</emphasis>
Sex, total length (TL) and tail length (tL) (in mm) and scalation data for 42 referred specimens of
<taxonomicName id="4C514D190B31FFFFCAD7E3DEFE4FFF1B" box="[151,398,187,213]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="13" pageNumber="346" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B31FFFFCAD7E3DEFE4FFF1B" box="[151,398,187,213]" italics="true" pageId="13" pageNumber="346">Uropeltis pulneyensis</emphasis>
</taxonomicName>
. Subcaudal scales given in left, right order (number of fused or undivided subcaudals in parentheses). Blank cells indicate data not recorded. See Appendix 1 for specimen localities.
</paragraph>
</caption>
<paragraph id="8BEE369A0B31FFFFCAE3E26CFA97F875" pageId="13" pageNumber="346">
<table id="F951C43A0B31000DCAE3E26CFABCF875" box="[163,1405,265,1979]" gridcols="6" gridrows="43" pageId="13" pageNumber="346">
<tr id="356134D80B31000DCAE3E26CFABCFEEF" box="[163,1405,265,289]" gridrow="0" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E26CFE47FEEF" box="[163,390,265,289]" gridcol="0" gridrow="0" pageId="13" pageNumber="346">Specimen</th>
<th id="76B05DA40B31000DCBAEE26CFDD5FEEF" box="[494,532,265,289]" gridcol="1" gridrow="0" pageId="13" pageNumber="346">Sex</th>
<th id="76B05DA40B31000DC8D1E26CFD76FEEF" box="[657,695,265,289]" gridcol="2" gridrow="0" pageId="13" pageNumber="346">TL</th>
<th id="76B05DA40B31000DC91CE26CFC49FEEF" box="[860,904,265,289]" gridcol="3" gridrow="0" pageId="13" pageNumber="346">tL</th>
<th id="76B05DA40B31000DCE40E26CFB4AFEEF" box="[1024,1163,265,289]" gridcol="4" gridrow="0" pageId="13" pageNumber="346">Ventral scales</th>
<th id="76B05DA40B31000DCE91E26CFABCFEEF" box="[1233,1405,265,289]" gridcol="5" gridrow="0" pageId="13" pageNumber="346">Subcaudal scales</th>
</tr>
<tr id="356134D80B31000DCAE3E254FABCFE87" box="[163,1405,305,329]" gridrow="1" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E254FE47FE87" box="[163,390,305,329]" gridcol="0" gridrow="1" pageId="13" pageNumber="346">AMNH 77607</th>
<td id="76B05DA40B31000DCBAEE254FDD5FE87" box="[494,532,305,329]" gridcol="1" gridrow="1" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E254FD76FE87" box="[657,695,305,329]" gridcol="2" gridrow="1" pageId="13" pageNumber="346">126</td>
<td id="76B05DA40B31000DC91CE254FC49FE87" box="[860,904,305,329]" gridcol="3" gridrow="1" pageId="13" pageNumber="346">4.6</td>
<td id="76B05DA40B31000DCE40E254FB4AFE87" box="[1024,1163,305,329]" gridcol="4" gridrow="1" pageId="13" pageNumber="346">180</td>
<td id="76B05DA40B31000DCE91E254FABCFE87" box="[1233,1405,305,329]" gridcol="5" gridrow="1" pageId="13" pageNumber="346">7,7 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E23CFABCFEBF" box="[163,1405,345,369]" gridrow="2" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E23CFE47FEBF" box="[163,390,345,369]" gridcol="0" gridrow="2" pageId="13" pageNumber="346">BMNH 74.4.29.111</th>
<td id="76B05DA40B31000DCBAEE23CFDD5FEBF" box="[494,532,345,369]" gridcol="1" gridrow="2" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E23CFD76FEBF" box="[657,695,345,369]" gridcol="2" gridrow="2" pageId="13" pageNumber="346">155</td>
<td id="76B05DA40B31000DC91CE23CFC49FEBF" box="[860,904,345,369]" gridcol="3" gridrow="2" pageId="13" pageNumber="346">7.7</td>
<td id="76B05DA40B31000DCE40E23CFB4AFEBF" box="[1024,1163,345,369]" gridcol="4" gridrow="2" pageId="13" pageNumber="346">167</td>
<td id="76B05DA40B31000DCE91E23CFABCFEBF" box="[1233,1405,345,369]" gridcol="5" gridrow="2" pageId="13" pageNumber="346">12,13 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E2E7FABCFE54" box="[163,1405,386,410]" gridrow="3" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E2E7FE47FE54" box="[163,390,386,410]" gridcol="0" gridrow="3" pageId="13" pageNumber="346">BMNH 74.4.29.112</th>
<td id="76B05DA40B31000DCBAEE2E7FDD5FE54" box="[494,532,386,410]" gridcol="1" gridrow="3" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E2E7FD76FE54" box="[657,695,386,410]" gridcol="2" gridrow="3" pageId="13" pageNumber="346">204</td>
<td id="76B05DA40B31000DC91CE2E7FC49FE54" box="[860,904,386,410]" gridcol="3" gridrow="3" pageId="13" pageNumber="346">7.1</td>
<td id="76B05DA40B31000DCE40E2E7FB4AFE54" box="[1024,1163,386,410]" gridcol="4" gridrow="3" pageId="13" pageNumber="346">173</td>
<td id="76B05DA40B31000DCE91E2E7FABCFE54" box="[1233,1405,386,410]" gridcol="5" gridrow="3" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E2CFFABCFE0C" box="[163,1405,426,450]" gridrow="4" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E2CFFE47FE0C" box="[163,390,426,450]" gridcol="0" gridrow="4" pageId="13" pageNumber="346">BMNH 74.4.29.113</th>
<td id="76B05DA40B31000DCBAEE2CFFDD5FE0C" box="[494,532,426,450]" gridcol="1" gridrow="4" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E2CFFD76FE0C" box="[657,695,426,450]" gridcol="2" gridrow="4" pageId="13" pageNumber="346">300</td>
<td id="76B05DA40B31000DC91CE2CFFC49FE0C" box="[860,904,426,450]" gridcol="3" gridrow="4" pageId="13" pageNumber="346">10.2</td>
<td id="76B05DA40B31000DCE40E2CFFB4AFE0C" box="[1024,1163,426,450]" gridcol="4" gridrow="4" pageId="13" pageNumber="346">179</td>
<td id="76B05DA40B31000DCE91E2CFFABCFE0C" box="[1233,1405,426,450]" gridcol="5" gridrow="4" pageId="13" pageNumber="346">8,7 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E2B7FABCFE24" box="[163,1405,466,490]" gridrow="5" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E2B7FE47FE24" box="[163,390,466,490]" gridcol="0" gridrow="5" pageId="13" pageNumber="346">BMNH83.1.12.19</th>
<td id="76B05DA40B31000DCBAEE2B7FDD5FE24" box="[494,532,466,490]" gridcol="1" gridrow="5" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E2B7FD76FE24" box="[657,695,466,490]" gridcol="2" gridrow="5" pageId="13" pageNumber="346">323</td>
<td id="76B05DA40B31000DC91CE2B7FC49FE24" box="[860,904,466,490]" gridcol="3" gridrow="5" pageId="13" pageNumber="346">12.0</td>
<td id="76B05DA40B31000DCE40E2B7FB4AFE24" box="[1024,1163,466,490]" gridcol="4" gridrow="5" pageId="13" pageNumber="346">175</td>
<td id="76B05DA40B31000DCE91E2B7FABCFE24" box="[1233,1405,466,490]" gridcol="5" gridrow="5" pageId="13" pageNumber="346">8,7 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E29FFABCFDDC" box="[163,1405,506,530]" gridrow="6" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E29FFE47FDDC" box="[163,390,506,530]" gridcol="0" gridrow="6" pageId="13" pageNumber="346">BMNH 83.1.1.55</th>
<td id="76B05DA40B31000DCBAEE29FFDD5FDDC" box="[494,532,506,530]" gridcol="1" gridrow="6" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E29FFD76FDDC" box="[657,695,506,530]" gridcol="2" gridrow="6" pageId="13" pageNumber="346">376</td>
<td id="76B05DA40B31000DC91CE29FFC49FDDC" box="[860,904,506,530]" gridcol="3" gridrow="6" pageId="13" pageNumber="346">12.7</td>
<td id="76B05DA40B31000DCE40E29FFB4AFDDC" box="[1024,1163,506,530]" gridcol="4" gridrow="6" pageId="13" pageNumber="346">170</td>
<td id="76B05DA40B31000DCE91E29FFABCFDDC" box="[1233,1405,506,530]" gridcol="5" gridrow="6" pageId="13" pageNumber="346">7,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E147FABCFDF4" box="[163,1405,546,570]" gridrow="7" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E147FE47FDF4" box="[163,390,546,570]" gridcol="0" gridrow="7" pageId="13" pageNumber="346">BMNH 1922.3.13.3</th>
<td id="76B05DA40B31000DCBAEE147FDD5FDF4" box="[494,532,546,570]" gridcol="1" gridrow="7" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E147FD76FDF4" box="[657,695,546,570]" gridcol="2" gridrow="7" pageId="13" pageNumber="346">191</td>
<td id="76B05DA40B31000DC91CE147FC49FDF4" box="[860,904,546,570]" gridcol="3" gridrow="7" pageId="13" pageNumber="346">10.5</td>
<td id="76B05DA40B31000DCE40E147FB4AFDF4" box="[1024,1163,546,570]" gridcol="4" gridrow="7" pageId="13" pageNumber="346">159</td>
<td id="76B05DA40B31000DCE91E147FABCFDF4" box="[1233,1405,546,570]" gridcol="5" gridrow="7" pageId="13" pageNumber="346">11,11 (2)</td>
</tr>
<tr id="356134D80B31000DCAE3E12EFABCFDAD" box="[163,1405,587,611]" gridrow="8" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E12EFE47FDAD" box="[163,390,587,611]" gridcol="0" gridrow="8" pageId="13" pageNumber="346">BMNH 1922.3.13.4</th>
<td id="76B05DA40B31000DCBAEE12EFDD5FDAD" box="[494,532,587,611]" gridcol="1" gridrow="8" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E12EFD76FDAD" box="[657,695,587,611]" gridcol="2" gridrow="8" pageId="13" pageNumber="346">149</td>
<td id="76B05DA40B31000DC91CE12EFC49FDAD" box="[860,904,587,611]" gridcol="3" gridrow="8" pageId="13" pageNumber="346">7.6</td>
<td id="76B05DA40B31000DCE40E12EFB4AFDAD" box="[1024,1163,587,611]" gridcol="4" gridrow="8" pageId="13" pageNumber="346">164</td>
<td id="76B05DA40B31000DCE91E12EFABCFDAD" box="[1233,1405,587,611]" gridcol="5" gridrow="8" pageId="13" pageNumber="346">11,11 (2)</td>
</tr>
<tr id="356134D80B31000DCAE3E116FABCFD45" box="[163,1405,627,651]" gridrow="9" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E116FE47FD45" box="[163,390,627,651]" gridcol="0" gridrow="9" pageId="13" pageNumber="346">BMNH 1922.3.13.5</th>
<td id="76B05DA40B31000DCBAEE116FDD5FD45" box="[494,532,627,651]" gridcol="1" gridrow="9" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E116FD76FD45" box="[657,695,627,651]" gridcol="2" gridrow="9" pageId="13" pageNumber="346">196</td>
<td id="76B05DA40B31000DC91CE116FC49FD45" box="[860,904,627,651]" gridcol="3" gridrow="9" pageId="13" pageNumber="346">6.9</td>
<td id="76B05DA40B31000DCE40E116FB4AFD45" box="[1024,1163,627,651]" gridcol="4" gridrow="9" pageId="13" pageNumber="346">166</td>
<td id="76B05DA40B31000DCE91E116FABCFD45" box="[1233,1405,627,651]" gridcol="5" gridrow="9" pageId="13" pageNumber="346">6,6 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E1FEFABCFD7D" box="[163,1405,667,691]" gridrow="10" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E1FEFE47FD7D" box="[163,390,667,691]" gridcol="0" gridrow="10" pageId="13" pageNumber="346">BMNH 1923.10.13.28</th>
<td id="76B05DA40B31000DCBAEE1FEFDD5FD7D" box="[494,532,667,691]" gridcol="1" gridrow="10" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E1FEFD76FD7D" box="[657,695,667,691]" gridcol="2" gridrow="10" pageId="13" pageNumber="346">249</td>
<td id="76B05DA40B31000DC91CE1FEFC49FD7D" box="[860,904,667,691]" gridcol="3" gridrow="10" pageId="13" pageNumber="346">8.3</td>
<td id="76B05DA40B31000DCE40E1FEFB4AFD7D" box="[1024,1163,667,691]" gridcol="4" gridrow="10" pageId="13" pageNumber="346">166</td>
<td id="76B05DA40B31000DCE91E1FEFABCFD7D" box="[1233,1405,667,691]" gridcol="5" gridrow="10" pageId="13" pageNumber="346">8,6 (1)</td>
</tr>
<tr id="356134D80B31000DCAE3E1A6FABCFD15" box="[163,1405,707,731]" gridrow="11" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E1A6FE47FD15" box="[163,390,707,731]" gridcol="0" gridrow="11" pageId="13" pageNumber="346">CAS 244795</th>
<td id="76B05DA40B31000DCBAEE1A6FDD5FD15" box="[494,532,707,731]" gridcol="1" gridrow="11" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DCE40E1A6FB4AFD15" box="[1024,1163,707,731]" gridcol="4" gridrow="11" pageId="13" pageNumber="346">165</td>
<td id="76B05DA40B31000DCE91E1A6FABCFD15" box="[1233,1405,707,731]" gridcol="5" gridrow="11" pageId="13" pageNumber="346">10, 11 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E189FABCFCCA" box="[163,1405,748,772]" gridrow="12" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E189FE47FCCA" box="[163,390,748,772]" gridcol="0" gridrow="12" pageId="13" pageNumber="346">CAS 244809</th>
<td id="76B05DA40B31000DCBAEE189FDD5FCCA" box="[494,532,748,772]" gridcol="1" gridrow="12" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DCE40E189FB4AFCCA" box="[1024,1163,748,772]" gridcol="4" gridrow="12" pageId="13" pageNumber="346">174</td>
<td id="76B05DA40B31000DCE91E189FABCFCCA" box="[1233,1405,748,772]" gridcol="5" gridrow="12" pageId="13" pageNumber="346">11,11 (1)</td>
</tr>
<tr id="356134D80B31000DCAE3E071FABCFCE2" box="[163,1405,788,812]" gridrow="13" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E071FE47FCE2" box="[163,390,788,812]" gridcol="0" gridrow="13" pageId="13" pageNumber="346">CAS 244810</th>
<td id="76B05DA40B31000DCBAEE071FDD5FCE2" box="[494,532,788,812]" gridcol="1" gridrow="13" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DCE40E071FB4AFCE2" box="[1024,1163,788,812]" gridcol="4" gridrow="13" pageId="13" pageNumber="346">176</td>
<td id="76B05DA40B31000DCE91E071FABCFCE2" box="[1233,1405,788,812]" gridcol="5" gridrow="13" pageId="13" pageNumber="346">6,6 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E059FABCFC9A" box="[163,1405,828,852]" gridrow="14" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E059FE47FC9A" box="[163,390,828,852]" gridcol="0" gridrow="14" pageId="13" pageNumber="346">CAS 244811</th>
<td id="76B05DA40B31000DCBAEE059FDD5FC9A" box="[494,532,828,852]" gridcol="1" gridrow="14" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DCE40E059FB4AFC9A" box="[1024,1163,828,852]" gridcol="4" gridrow="14" pageId="13" pageNumber="346">172</td>
<td id="76B05DA40B31000DCE91E059FABCFC9A" box="[1233,1405,828,852]" gridcol="5" gridrow="14" pageId="13" pageNumber="346">11,10 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E001FABCFCB2" box="[163,1405,868,892]" gridrow="15" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E001FE47FCB2" box="[163,390,868,892]" gridcol="0" gridrow="15" pageId="13" pageNumber="346">CAS 244812</th>
<td id="76B05DA40B31000DCBAEE001FDD5FCB2" box="[494,532,868,892]" gridcol="1" gridrow="15" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DCE40E001FB4AFCB2" box="[1024,1163,868,892]" gridcol="4" gridrow="15" pageId="13" pageNumber="346">171</td>
<td id="76B05DA40B31000DCE91E001FABCFCB2" box="[1233,1405,868,892]" gridcol="5" gridrow="15" pageId="13" pageNumber="346">11,12 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E0E9FABCFC6B" box="[163,1405,908,933]" gridrow="16" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E0E9FE47FC6B" box="[163,390,908,933]" gridcol="0" gridrow="16" pageId="13" pageNumber="346">MCZ 1335</th>
<td id="76B05DA40B31000DCBAEE0E9FDD5FC6B" box="[494,532,908,933]" gridcol="1" gridrow="16" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E0E9FD76FC6B" box="[657,695,908,933]" gridcol="2" gridrow="16" pageId="13" pageNumber="346">343</td>
<td id="76B05DA40B31000DC91CE0E9FC49FC6B" box="[860,904,908,933]" gridcol="3" gridrow="16" pageId="13" pageNumber="346">11.6</td>
<td id="76B05DA40B31000DCE40E0E9FB4AFC6B" box="[1024,1163,908,933]" gridcol="4" gridrow="16" pageId="13" pageNumber="346">169</td>
<td id="76B05DA40B31000DCE91E0E9FABCFC6B" box="[1233,1405,908,933]" gridcol="5" gridrow="16" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E0D0FABCFC03" box="[163,1405,949,973]" gridrow="17" pageId="13" pageNumber="346" rowspan-2="1" rowspan-3="1">
<th id="76B05DA40B31000DCAE3E0D0FE47FC03" box="[163,390,949,973]" gridcol="0" gridrow="17" pageId="13" pageNumber="346">MCZ 3870</th>
<td id="76B05DA40B31000DCBAEE0D0FDD5FC03" box="[494,532,949,973]" gridcol="1" gridrow="17" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DCE40E0D0FB4AFC03" box="[1024,1163,949,973]" gridcol="4" gridrow="17" pageId="13" pageNumber="346">166</td>
<td id="76B05DA40B31000DCE91E0D0FABCFC03" box="[1233,1405,949,973]" gridcol="5" gridrow="17" pageId="13" pageNumber="346">12,11 (1)</td>
</tr>
<tr id="356134D80B31000DCAE3E0B8FABCFC3B" box="[163,1405,989,1013]" gridrow="18" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E0B8FE47FC3B" box="[163,390,989,1013]" gridcol="0" gridrow="18" pageId="13" pageNumber="346">MCZ 7773</th>
<td id="76B05DA40B31000DCBAEE0B8FDD5FC3B" box="[494,532,989,1013]" gridcol="1" gridrow="18" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E0B8FD76FC3B" box="[657,695,989,1013]" gridcol="2" gridrow="18" pageId="13" pageNumber="346">229</td>
<td id="76B05DA40B31000DC91CE0B8FC49FC3B" box="[860,904,989,1013]" gridcol="3" gridrow="18" pageId="13" pageNumber="346">9.1</td>
<td id="76B05DA40B31000DCE40E0B8FB4AFC3B" box="[1024,1163,989,1013]" gridcol="4" gridrow="18" pageId="13" pageNumber="346">177</td>
<td id="76B05DA40B31000DCE91E0B8FABCFC3B" box="[1233,1405,989,1013]" gridcol="5" gridrow="18" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E760FABCFBD3" box="[163,1405,1029,1053]" gridrow="19" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E760FE47FBD3" box="[163,390,1029,1053]" gridcol="0" gridrow="19" pageId="13" pageNumber="346">MCZ 33506</th>
<td id="76B05DA40B31000DCBAEE760FDD5FBD3" box="[494,532,1029,1053]" gridcol="1" gridrow="19" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E760FD76FBD3" box="[657,695,1029,1053]" gridcol="2" gridrow="19" pageId="13" pageNumber="346">212</td>
<td id="76B05DA40B31000DC91CE760FC49FBD3" box="[860,904,1029,1053]" gridcol="3" gridrow="19" pageId="13" pageNumber="346">8.6</td>
<td id="76B05DA40B31000DCE40E760FB4AFBD3" box="[1024,1163,1029,1053]" gridcol="4" gridrow="19" pageId="13" pageNumber="346">171</td>
<td id="76B05DA40B31000DCE91E760FABCFBD3" box="[1233,1405,1029,1053]" gridcol="5" gridrow="19" pageId="13" pageNumber="346">7,8 (1)</td>
</tr>
<tr id="356134D80B31000DCAE3E748FABCFB88" box="[163,1405,1069,1094]" gridrow="20" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E748FE47FB88" box="[163,390,1069,1094]" gridcol="0" gridrow="20" pageId="13" pageNumber="346">MCZ 33507</th>
<td id="76B05DA40B31000DCBAEE748FDD5FB88" box="[494,532,1069,1094]" gridcol="1" gridrow="20" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E748FD76FB88" box="[657,695,1069,1094]" gridcol="2" gridrow="20" pageId="13" pageNumber="346">237</td>
<td id="76B05DA40B31000DC91CE748FC49FB88" box="[860,904,1069,1094]" gridcol="3" gridrow="20" pageId="13" pageNumber="346">13.7</td>
<td id="76B05DA40B31000DCE40E748FB4AFB88" box="[1024,1163,1069,1094]" gridcol="4" gridrow="20" pageId="13" pageNumber="346">165</td>
<td id="76B05DA40B31000DCE91E748FABCFB88" box="[1233,1405,1069,1094]" gridcol="5" gridrow="20" pageId="13" pageNumber="346">11,11 (5)</td>
</tr>
<tr id="356134D80B31000DCAE3E733FABCFBA0" box="[163,1405,1110,1134]" gridrow="21" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E733FE47FBA0" box="[163,390,1110,1134]" gridcol="0" gridrow="21" pageId="13" pageNumber="346">MCZ 47042</th>
<td id="76B05DA40B31000DCBAEE733FDD5FBA0" box="[494,532,1110,1134]" gridcol="1" gridrow="21" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E733FD76FBA0" box="[657,695,1110,1134]" gridcol="2" gridrow="21" pageId="13" pageNumber="346">236</td>
<td id="76B05DA40B31000DC91CE733FC49FBA0" box="[860,904,1110,1134]" gridcol="3" gridrow="21" pageId="13" pageNumber="346">7.5</td>
<td id="76B05DA40B31000DCE40E733FB4AFBA0" box="[1024,1163,1110,1134]" gridcol="4" gridrow="21" pageId="13" pageNumber="346">177</td>
<td id="76B05DA40B31000DCE91E733FABCFBA0" box="[1233,1405,1110,1134]" gridcol="5" gridrow="21" pageId="13" pageNumber="346">8,7 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E71BFABCFB58" box="[163,1405,1150,1174]" gridrow="22" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E71BFE47FB58" box="[163,390,1150,1174]" gridcol="0" gridrow="22" pageId="13" pageNumber="346">MCZ 47043</th>
<td id="76B05DA40B31000DCBAEE71BFDD5FB58" box="[494,532,1150,1174]" gridcol="1" gridrow="22" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E71BFD76FB58" box="[657,695,1150,1174]" gridcol="2" gridrow="22" pageId="13" pageNumber="346">235</td>
<td id="76B05DA40B31000DC91CE71BFC49FB58" box="[860,904,1150,1174]" gridcol="3" gridrow="22" pageId="13" pageNumber="346">14.7</td>
<td id="76B05DA40B31000DCE40E71BFB4AFB58" box="[1024,1163,1150,1174]" gridcol="4" gridrow="22" pageId="13" pageNumber="346">166</td>
<td id="76B05DA40B31000DCE91E71BFABCFB58" box="[1233,1405,1150,1174]" gridcol="5" gridrow="22" pageId="13" pageNumber="346">12,12 (2)</td>
</tr>
<tr id="356134D80B31000DCAE3E7C3FABCFB70" box="[163,1405,1190,1214]" gridrow="23" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E7C3FE47FB70" box="[163,390,1190,1214]" gridcol="0" gridrow="23" pageId="13" pageNumber="346">MNHN 1946.44</th>
<td id="76B05DA40B31000DCBAEE7C3FDD5FB70" box="[494,532,1190,1214]" gridcol="1" gridrow="23" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E7C3FD76FB70" box="[657,695,1190,1214]" gridcol="2" gridrow="23" pageId="13" pageNumber="346">244</td>
<td id="76B05DA40B31000DC91CE7C3FC49FB70" box="[860,904,1190,1214]" gridcol="3" gridrow="23" pageId="13" pageNumber="346">13.2</td>
<td id="76B05DA40B31000DCE40E7C3FB4AFB70" box="[1024,1163,1190,1214]" gridcol="4" gridrow="23" pageId="13" pageNumber="346">166</td>
<td id="76B05DA40B31000DCE91E7C3FABCFB70" box="[1233,1405,1190,1214]" gridcol="5" gridrow="23" pageId="13" pageNumber="346">11,11 (2)</td>
</tr>
<tr id="356134D80B31000DCAE3E7ABFABCFB29" box="[163,1405,1230,1255]" gridrow="24" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E7ABFE47FB29" box="[163,390,1230,1255]" gridcol="0" gridrow="24" pageId="13" pageNumber="346">MNHN 1946.46</th>
<td id="76B05DA40B31000DCBAEE7ABFDD5FB29" box="[494,532,1230,1255]" gridcol="1" gridrow="24" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E7ABFD76FB29" box="[657,695,1230,1255]" gridcol="2" gridrow="24" pageId="13" pageNumber="346">218</td>
<td id="76B05DA40B31000DC91CE7ABFC49FB29" box="[860,904,1230,1255]" gridcol="3" gridrow="24" pageId="13" pageNumber="346">7.3</td>
<td id="76B05DA40B31000DCE40E7ABFB4AFB29" box="[1024,1163,1230,1255]" gridcol="4" gridrow="24" pageId="13" pageNumber="346">177</td>
<td id="76B05DA40B31000DCE91E7ABFABCFB29" box="[1233,1405,1230,1255]" gridcol="5" gridrow="24" pageId="13" pageNumber="346">9,9 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E792FABCFAC1" box="[163,1405,1271,1295]" gridrow="25" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E792FE47FAC1" box="[163,390,1271,1295]" gridcol="0" gridrow="25" pageId="13" pageNumber="346">MNHN 1946.47</th>
<td id="76B05DA40B31000DCBAEE792FDD5FAC1" box="[494,532,1271,1295]" gridcol="1" gridrow="25" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E792FD76FAC1" box="[657,695,1271,1295]" gridcol="2" gridrow="25" pageId="13" pageNumber="346">238</td>
<td id="76B05DA40B31000DC91CE792FC49FAC1" box="[860,904,1271,1295]" gridcol="3" gridrow="25" pageId="13" pageNumber="346">7.8</td>
<td id="76B05DA40B31000DCE40E792FB4AFAC1" box="[1024,1163,1271,1295]" gridcol="4" gridrow="25" pageId="13" pageNumber="346">177</td>
<td id="76B05DA40B31000DCE91E792FABCFAC1" box="[1233,1405,1271,1295]" gridcol="5" gridrow="25" pageId="13" pageNumber="346">7,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E67AFABCFAF9" box="[163,1405,1311,1335]" gridrow="26" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E67AFE47FAF9" box="[163,390,1311,1335]" gridcol="0" gridrow="26" pageId="13" pageNumber="346">MNHN 1946.268</th>
<td id="76B05DA40B31000DCBAEE67AFDD5FAF9" box="[494,532,1311,1335]" gridcol="1" gridrow="26" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E67AFD76FAF9" box="[657,695,1311,1335]" gridcol="2" gridrow="26" pageId="13" pageNumber="346">250</td>
<td id="76B05DA40B31000DC91CE67AFC49FAF9" box="[860,904,1311,1335]" gridcol="3" gridrow="26" pageId="13" pageNumber="346">13.4</td>
<td id="76B05DA40B31000DCE40E67AFB4AFAF9" box="[1024,1163,1311,1335]" gridcol="4" gridrow="26" pageId="13" pageNumber="346">162</td>
<td id="76B05DA40B31000DCE91E67AFABCFAF9" box="[1233,1405,1311,1335]" gridcol="5" gridrow="26" pageId="13" pageNumber="346">12,12 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E622FABCFA91" box="[163,1405,1351,1375]" gridrow="27" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E622FE47FA91" box="[163,390,1351,1375]" gridcol="0" gridrow="27" pageId="13" pageNumber="346">MNHN 1946.269</th>
<td id="76B05DA40B31000DCBAEE622FDD5FA91" box="[494,532,1351,1375]" gridcol="1" gridrow="27" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E622FD76FA91" box="[657,695,1351,1375]" gridcol="2" gridrow="27" pageId="13" pageNumber="346">247</td>
<td id="76B05DA40B31000DC91CE622FC49FA91" box="[860,904,1351,1375]" gridcol="3" gridrow="27" pageId="13" pageNumber="346">8.2</td>
<td id="76B05DA40B31000DCE40E622FB4AFA91" box="[1024,1163,1351,1375]" gridcol="4" gridrow="27" pageId="13" pageNumber="346">169</td>
<td id="76B05DA40B31000DCE91E622FABCFA91" box="[1233,1405,1351,1375]" gridcol="5" gridrow="27" pageId="13" pageNumber="346">8,8 (2)</td>
</tr>
<tr id="356134D80B31000DCAE3E60AFABCFA46" box="[163,1405,1391,1416]" gridrow="28" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E60AFE47FA46" box="[163,390,1391,1416]" gridcol="0" gridrow="28" pageId="13" pageNumber="346">MNHN 1948.253</th>
<td id="76B05DA40B31000DCBAEE60AFDD5FA46" box="[494,532,1391,1416]" gridcol="1" gridrow="28" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E60AFD76FA46" box="[657,695,1391,1416]" gridcol="2" gridrow="28" pageId="13" pageNumber="346">239</td>
<td id="76B05DA40B31000DC91CE60AFC49FA46" box="[860,904,1391,1416]" gridcol="3" gridrow="28" pageId="13" pageNumber="346">7.9</td>
<td id="76B05DA40B31000DCE40E60AFB4AFA46" box="[1024,1163,1391,1416]" gridcol="4" gridrow="28" pageId="13" pageNumber="346">179</td>
<td id="76B05DA40B31000DCE91E60AFABCFA46" box="[1233,1405,1391,1416]" gridcol="5" gridrow="28" pageId="13" pageNumber="346">7,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E6FDFABCFA7E" box="[163,1405,1432,1456]" gridrow="29" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E6FDFE47FA7E" box="[163,390,1432,1456]" gridcol="0" gridrow="29" pageId="13" pageNumber="346">MNHN 1948.255</th>
<td id="76B05DA40B31000DCBAEE6FDFDD5FA7E" box="[494,532,1432,1456]" gridcol="1" gridrow="29" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E6FDFD76FA7E" box="[657,695,1432,1456]" gridcol="2" gridrow="29" pageId="13" pageNumber="346">244</td>
<td id="76B05DA40B31000DC91CE6FDFC49FA7E" box="[860,904,1432,1456]" gridcol="3" gridrow="29" pageId="13" pageNumber="346">8.8</td>
<td id="76B05DA40B31000DCE40E6FDFB4AFA7E" box="[1024,1163,1432,1456]" gridcol="4" gridrow="29" pageId="13" pageNumber="346">181</td>
<td id="76B05DA40B31000DCE91E6FDFABCFA7E" box="[1233,1405,1432,1456]" gridcol="5" gridrow="29" pageId="13" pageNumber="346">9,9 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E6A5FABCFA16" box="[163,1405,1472,1496]" gridrow="30" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E6A5FE47FA16" box="[163,390,1472,1496]" gridcol="0" gridrow="30" pageId="13" pageNumber="346">MNHN 1948.256</th>
<td id="76B05DA40B31000DCBAEE6A5FDD5FA16" box="[494,532,1472,1496]" gridcol="1" gridrow="30" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E6A5FD76FA16" box="[657,695,1472,1496]" gridcol="2" gridrow="30" pageId="13" pageNumber="346">187</td>
<td id="76B05DA40B31000DC91CE6A5FC49FA16" box="[860,904,1472,1496]" gridcol="3" gridrow="30" pageId="13" pageNumber="346">10.9</td>
<td id="76B05DA40B31000DCE40E6A5FB4AFA16" box="[1024,1163,1472,1496]" gridcol="4" gridrow="30" pageId="13" pageNumber="346">167</td>
<td id="76B05DA40B31000DCE91E6A5FABCFA16" box="[1233,1405,1472,1496]" gridcol="5" gridrow="30" pageId="13" pageNumber="346">13,14 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E68DFABCF9CE" box="[163,1405,1512,1536]" gridrow="31" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E68DFE47F9CE" box="[163,390,1512,1536]" gridcol="0" gridrow="31" pageId="13" pageNumber="346">MNHN 1948.751</th>
<td id="76B05DA40B31000DCBAEE68DFDD5F9CE" box="[494,532,1512,1536]" gridcol="1" gridrow="31" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E68DFD76F9CE" box="[657,695,1512,1536]" gridcol="2" gridrow="31" pageId="13" pageNumber="346">190</td>
<td id="76B05DA40B31000DC91CE68DFC49F9CE" box="[860,904,1512,1536]" gridcol="3" gridrow="31" pageId="13" pageNumber="346">7.5</td>
<td id="76B05DA40B31000DCE40E68DFB4AF9CE" box="[1024,1163,1512,1536]" gridcol="4" gridrow="31" pageId="13" pageNumber="346">169</td>
<td id="76B05DA40B31000DCE91E68DFABCF9CE" box="[1233,1405,1512,1536]" gridcol="5" gridrow="31" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E575FABCF9E7" box="[163,1405,1552,1577]" gridrow="32" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E575FE47F9E7" box="[163,390,1552,1577]" gridcol="0" gridrow="32" pageId="13" pageNumber="346">MNHN 1948.752</th>
<td id="76B05DA40B31000DCBAEE575FDD5F9E7" box="[494,532,1552,1577]" gridcol="1" gridrow="32" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E575FD76F9E7" box="[657,695,1552,1577]" gridcol="2" gridrow="32" pageId="13" pageNumber="346">252</td>
<td id="76B05DA40B31000DC91CE575FC49F9E7" box="[860,904,1552,1577]" gridcol="3" gridrow="32" pageId="13" pageNumber="346">8.1</td>
<td id="76B05DA40B31000DCE40E575FB4AF9E7" box="[1024,1163,1552,1577]" gridcol="4" gridrow="32" pageId="13" pageNumber="346">180</td>
<td id="76B05DA40B31000DCE91E575FABCF9E7" box="[1233,1405,1552,1577]" gridcol="5" gridrow="32" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E55DFABCF99F" box="[163,1405,1592,1617]" gridrow="33" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E55DFE47F99F" box="[163,390,1592,1617]" gridcol="0" gridrow="33" pageId="13" pageNumber="346">MNHN 1948.753</th>
<td id="76B05DA40B31000DCBAEE55DFDD5F99F" box="[494,532,1592,1617]" gridcol="1" gridrow="33" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E55DFD76F99F" box="[657,695,1592,1617]" gridcol="2" gridrow="33" pageId="13" pageNumber="346">149</td>
<td id="76B05DA40B31000DC91CE55DFC49F99F" box="[860,904,1592,1617]" gridcol="3" gridrow="33" pageId="13" pageNumber="346">7.6</td>
<td id="76B05DA40B31000DCE40E55DFB4AF99F" box="[1024,1163,1592,1617]" gridcol="4" gridrow="33" pageId="13" pageNumber="346">173</td>
<td id="76B05DA40B31000DCE91E55DFABCF99F" box="[1233,1405,1592,1617]" gridcol="5" gridrow="33" pageId="13" pageNumber="346">11,12 (6)</td>
</tr>
<tr id="356134D80B31000DCAE3E504FABCF9B7" box="[163,1405,1633,1657]" gridrow="34" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E504FE47F9B7" box="[163,390,1633,1657]" gridcol="0" gridrow="34" pageId="13" pageNumber="346">MNHN 1948.754</th>
<td id="76B05DA40B31000DCBAEE504FDD5F9B7" box="[494,532,1633,1657]" gridcol="1" gridrow="34" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E504FD76F9B7" box="[657,695,1633,1657]" gridcol="2" gridrow="34" pageId="13" pageNumber="346">239</td>
<td id="76B05DA40B31000DC91CE504FC49F9B7" box="[860,904,1633,1657]" gridcol="3" gridrow="34" pageId="13" pageNumber="346">12.5</td>
<td id="76B05DA40B31000DCE40E504FB4AF9B7" box="[1024,1163,1633,1657]" gridcol="4" gridrow="34" pageId="13" pageNumber="346">156</td>
<td id="76B05DA40B31000DCE91E504FABCF9B7" box="[1233,1405,1633,1657]" gridcol="5" gridrow="34" pageId="13" pageNumber="346">11,11 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E5ECFABCF96F" box="[163,1405,1673,1697]" gridrow="35" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E5ECFE47F96F" box="[163,390,1673,1697]" gridcol="0" gridrow="35" pageId="13" pageNumber="346">MNHN 1948.755</th>
<td id="76B05DA40B31000DCBAEE5ECFDD5F96F" box="[494,532,1673,1697]" gridcol="1" gridrow="35" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E5ECFD76F96F" box="[657,695,1673,1697]" gridcol="2" gridrow="35" pageId="13" pageNumber="346">119</td>
<td id="76B05DA40B31000DC91CE5ECFC49F96F" box="[860,904,1673,1697]" gridcol="3" gridrow="35" pageId="13" pageNumber="346">8.1</td>
<td id="76B05DA40B31000DCE40E5ECFB4AF96F" box="[1024,1163,1673,1697]" gridcol="4" gridrow="35" pageId="13" pageNumber="346">165</td>
<td id="76B05DA40B31000DCE91E5ECFABCF96F" box="[1233,1405,1673,1697]" gridcol="5" gridrow="35" pageId="13" pageNumber="346">13,13 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E5D4FABCF904" box="[163,1405,1713,1738]" gridrow="36" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E5D4FE47F904" box="[163,390,1713,1738]" gridcol="0" gridrow="36" pageId="13" pageNumber="346">MNHN 1948.757</th>
<td id="76B05DA40B31000DCBAEE5D4FDD5F904" box="[494,532,1713,1738]" gridcol="1" gridrow="36" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E5D4FD76F904" box="[657,695,1713,1738]" gridcol="2" gridrow="36" pageId="13" pageNumber="346">256</td>
<td id="76B05DA40B31000DC91CE5D4FC49F904" box="[860,904,1713,1738]" gridcol="3" gridrow="36" pageId="13" pageNumber="346">7.5</td>
<td id="76B05DA40B31000DCE40E5D4FB4AF904" box="[1024,1163,1713,1738]" gridcol="4" gridrow="36" pageId="13" pageNumber="346">171</td>
<td id="76B05DA40B31000DCE91E5D4FABCF904" box="[1233,1405,1713,1738]" gridcol="5" gridrow="36" pageId="13" pageNumber="346">7,7 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E5BCFABCF93C" box="[163,1405,1753,1778]" gridrow="37" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E5BCFE47F93C" box="[163,390,1753,1778]" gridcol="0" gridrow="37" pageId="13" pageNumber="346">MNHN 1948.758</th>
<td id="76B05DA40B31000DCBAEE5BCFDD5F93C" box="[494,532,1753,1778]" gridcol="1" gridrow="37" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E5BCFD76F93C" box="[657,695,1753,1778]" gridcol="2" gridrow="37" pageId="13" pageNumber="346">204</td>
<td id="76B05DA40B31000DC91CE5BCFC49F93C" box="[860,904,1753,1778]" gridcol="3" gridrow="37" pageId="13" pageNumber="346">7.4</td>
<td id="76B05DA40B31000DCE40E5BCFB4AF93C" box="[1024,1163,1753,1778]" gridcol="4" gridrow="37" pageId="13" pageNumber="346">174</td>
<td id="76B05DA40B31000DCE91E5BCFABCF93C" box="[1233,1405,1753,1778]" gridcol="5" gridrow="37" pageId="13" pageNumber="346">7,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E467FABCF8D4" box="[163,1405,1794,1818]" gridrow="38" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E467FE47F8D4" box="[163,390,1794,1818]" gridcol="0" gridrow="38" pageId="13" pageNumber="346">MHNG 844.70</th>
<td id="76B05DA40B31000DCBAEE467FDD5F8D4" box="[494,532,1794,1818]" gridcol="1" gridrow="38" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E467FD76F8D4" box="[657,695,1794,1818]" gridcol="2" gridrow="38" pageId="13" pageNumber="346">245</td>
<td id="76B05DA40B31000DC91CE467FC49F8D4" box="[860,904,1794,1818]" gridcol="3" gridrow="38" pageId="13" pageNumber="346">8.9</td>
<td id="76B05DA40B31000DCE40E467FB4AF8D4" box="[1024,1163,1794,1818]" gridcol="4" gridrow="38" pageId="13" pageNumber="346">175</td>
<td id="76B05DA40B31000DCE91E467FABCF8D4" box="[1233,1405,1794,1818]" gridcol="5" gridrow="38" pageId="13" pageNumber="346">8,8 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E44FFABCF88C" box="[163,1405,1834,1858]" gridrow="39" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E44FFE47F88C" box="[163,390,1834,1858]" gridcol="0" gridrow="39" pageId="13" pageNumber="346">MHNG 2745.088</th>
<td id="76B05DA40B31000DCBAEE44FFDD5F88C" box="[494,532,1834,1858]" gridcol="1" gridrow="39" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E44FFD76F88C" box="[657,695,1834,1858]" gridcol="2" gridrow="39" pageId="13" pageNumber="346">232</td>
<td id="76B05DA40B31000DC91CE44FFC49F88C" box="[860,904,1834,1858]" gridcol="3" gridrow="39" pageId="13" pageNumber="346">11.8</td>
<td id="76B05DA40B31000DCE40E44FFB4AF88C" box="[1024,1163,1834,1858]" gridcol="4" gridrow="39" pageId="13" pageNumber="346">175</td>
<td id="76B05DA40B31000DCE91E44FFABCF88C" box="[1233,1405,1834,1858]" gridcol="5" gridrow="39" pageId="13" pageNumber="346">12,11 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E437FABCF8A4" box="[163,1405,1874,1898]" gridrow="40" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E437FE47F8A4" box="[163,390,1874,1898]" gridcol="0" gridrow="40" pageId="13" pageNumber="346">ZMB 5537</th>
<td id="76B05DA40B31000DCBAEE437FDD5F8A4" box="[494,532,1874,1898]" gridcol="1" gridrow="40" pageId="13" pageNumber="346">F</td>
<td id="76B05DA40B31000DC8D1E437FD76F8A4" box="[657,695,1874,1898]" gridcol="2" gridrow="40" pageId="13" pageNumber="346">259</td>
<td id="76B05DA40B31000DC91CE437FC49F8A4" box="[860,904,1874,1898]" gridcol="3" gridrow="40" pageId="13" pageNumber="346">9.8</td>
<td id="76B05DA40B31000DCE40E437FB4AF8A4" box="[1024,1163,1874,1898]" gridcol="4" gridrow="40" pageId="13" pageNumber="346">182</td>
<td id="76B05DA40B31000DCE91E437FABCF8A4" box="[1233,1405,1874,1898]" gridcol="5" gridrow="40" pageId="13" pageNumber="346">8,9 (0)</td>
</tr>
<tr id="356134D80B31000DCAE3E41FFABCF85D" box="[163,1405,1914,1939]" gridrow="41" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E41FFE47F85D" box="[163,390,1914,1939]" gridcol="0" gridrow="41" pageId="13" pageNumber="346">ZMB 5538</th>
<td id="76B05DA40B31000DCBAEE41FFDD5F85D" box="[494,532,1914,1939]" gridcol="1" gridrow="41" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E41FFD76F85D" box="[657,695,1914,1939]" gridcol="2" gridrow="41" pageId="13" pageNumber="346">231</td>
<td id="76B05DA40B31000DC91CE41FFC49F85D" box="[860,904,1914,1939]" gridcol="3" gridrow="41" pageId="13" pageNumber="346">8.2</td>
<td id="76B05DA40B31000DCE40E41FFB4AF85D" box="[1024,1163,1914,1939]" gridcol="4" gridrow="41" pageId="13" pageNumber="346">177</td>
<td id="76B05DA40B31000DCE91E41FFABCF85D" box="[1233,1405,1914,1939]" gridcol="5" gridrow="41" pageId="13" pageNumber="346">7,7 (1)</td>
</tr>
<tr id="356134D80B31000DCAE3E4C6FABCF875" box="[163,1405,1955,1979]" gridrow="42" pageId="13" pageNumber="346">
<th id="76B05DA40B31000DCAE3E4C6FE47F875" box="[163,390,1955,1979]" gridcol="0" gridrow="42" pageId="13" pageNumber="346">ZMB 10357</th>
<td id="76B05DA40B31000DCBAEE4C6FDD5F875" box="[494,532,1955,1979]" gridcol="1" gridrow="42" pageId="13" pageNumber="346">M</td>
<td id="76B05DA40B31000DC8D1E4C6FD76F875" box="[657,695,1955,1979]" gridcol="2" gridrow="42" pageId="13" pageNumber="346">300</td>
<td id="76B05DA40B31000DC91CE4C6FC49F875" box="[860,904,1955,1979]" gridcol="3" gridrow="42" pageId="13" pageNumber="346">14.1</td>
<td id="76B05DA40B31000DCE40E4C6FB4AF875" box="[1024,1163,1955,1979]" gridcol="4" gridrow="42" pageId="13" pageNumber="346">167</td>
<td id="76B05DA40B31000DCE91E4C6FABCF875" box="[1233,1405,1955,1979]" gridcol="5" gridrow="42" pageId="13" pageNumber="346">13,13 (0)</td>
</tr>
</table>
</paragraph>
<caption id="DF2E66120B32FFFCCAD7E45BFC5BF850" ID-DOI="http://doi.org/10.5281/zenodo.12732849" ID-Zenodo-Dep="12732849" httpUri="https://zenodo.org/record/12732849/files/figure.png" pageId="14" pageNumber="347" startId="14.[151,250,1854,1878]" targetBox="[151,1435,181,1830]" targetPageId="14" targetType="figure">
<paragraph id="8BEE369A0B32FFFCCAD7E45BFC5BF850" blockId="14.[151,1436,1854,1950]" pageId="14" pageNumber="347">
<emphasis id="B925EA880B32FFFCCAD7E45BFED4F898" bold="true" box="[151,277,1854,1878]" pageId="14" pageNumber="347">FIGURE 7.</emphasis>
Habitats of recent localities for
<taxonomicName id="4C514D190B32FFFCC826E45BFCA8F898" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[614,873,1854,1878]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="14" pageNumber="347" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B32FFFCC826E45BFCA8F898" box="[614,873,1854,1878]" italics="true" pageId="14" pageNumber="347">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B32FFFCC932E45BFC05F898" bold="true" box="[882,964,1854,1878]" pageId="14" pageNumber="347">
<taxonomicNameLabel id="A21657F30B32FFFCC932E45BFC05F898" box="[882,964,1854,1878]" pageId="14" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Locality where paratype BNHS 3765 was collected, at Manalar, Meghamalai, Tamil Nadu. B. General habitat close to where paratypes BNHS 3763 and 3764 were collected; Yellapetty, Munnar, Kerala. Photographs by JDG (A) and SD (B).
</paragraph>
</caption>
<caption id="DF2E66120B33FFFDCAD7E4F2FC2EF81D" ID-DOI="http://doi.org/10.5281/zenodo.12732851" ID-Zenodo-Dep="12732851" httpUri="https://zenodo.org/record/12732851/files/figure.png" pageId="15" pageNumber="348" startId="15.[151,250,1943,1967]" targetBox="[303,1283,181,1919]" targetPageId="15" targetType="figure">
<paragraph id="8BEE369A0B33FFFDCAD7E4F2FC2EF81D" blockId="15.[151,1436,1943,2003]" pageId="15" pageNumber="348">
<emphasis id="B925EA880B33FFFDCAD7E4F2FED5F861" bold="true" box="[151,276,1943,1967]" pageId="15" pageNumber="348">FIGURE 8.</emphasis>
Lectotype (by present designation) of
<taxonomicName id="4C514D190B33FFFDC8E7E4F2FB87F861" authority="(Boulenger, 1863)" baseAuthorityName="Boulenger" baseAuthorityYear="1863" box="[679,1094,1943,1967]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="15" pageNumber="348" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B33FFFDC8E7E4F2FC45F861" box="[679,900,1943,1967]" italics="true" pageId="15" pageNumber="348">Uropeltis pulneyensis</emphasis>
(Boulenger, 1863)
</taxonomicName>
(BMNH 1946.1.17.6, previously BMNH 64.3.9.1). Scale bar increments in millimetres. Photographs by Kevin Webb.
</paragraph>
</caption>
<caption id="DF2E66120B2CFFE2CAD7E451FD10F8BF" ID-DOI="http://doi.org/10.5281/zenodo.12732853" ID-Zenodo-Dep="12732853" httpUri="https://zenodo.org/record/12732853/files/figure.png" pageId="16" pageNumber="349" startId="16.[151,250,1844,1868]" targetBox="[151,1436,181,1821]" targetPageId="16" targetType="figure">
<paragraph id="8BEE369A0B2CFFE2CAD7E451FD10F8BF" blockId="16.[151,1436,1844,1905]" pageId="16" pageNumber="349">
<emphasis id="B925EA880B2CFFE2CAD7E451FED3F882" bold="true" box="[151,274,1844,1868]" pageId="16" pageNumber="349">FIGURE 9.</emphasis>
Holotype of
<taxonomicName id="4C514D190B2CFFE2CBD5E450FCCEF883" authority="(Beddome, 1878)" baseAuthorityName="Beddome" baseAuthorityYear="1878" box="[405,783,1845,1869]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="16" pageNumber="349" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2CFFE2CBD5E450FD9BF883" box="[405,602,1845,1869]" italics="true" pageId="16" pageNumber="349">Uropeltis guentheri</emphasis>
(
<bibRefCitation id="EFC04B6B0B2CFFE2C826E450FCC9F883" author="Beddome, R. H." box="[614,776,1845,1869]" pageId="16" pageNumber="349" pagination="154 - 155" refId="ref10438" refString="Beddome, R. H. (1878) Descriptions of new Uropeltidae from southern India, with remarks on some previously described species. Proceedings of the Zoological Society of London, 46, 154 - 155. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07946. x" type="journal article" year="1878">Beddome, 1878</bibRefCitation>
)
</taxonomicName>
(BMNH 1946.1.16.74, previously BMNH 83.1.12.34). Scale bar increments in millimetres. Photographs by Kevin Webb.
</paragraph>
</caption>
<subSubSection id="C34B65110B2DFFE0CA87E3F2FCEAF9E3" lastPageId="18" lastPageNumber="351" pageId="17" pageNumber="350" type="discussion">
<paragraph id="8BEE369A0B2DFFE3CA87E3F2FDB7FCDB" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="350">
<emphasis id="B925EA880B2DFFE3CA87E3F2FEFAFF7F" bold="true" box="[199,315,151,177]" pageId="17" pageNumber="350">Remarks.</emphasis>
<bibRefCitation id="EFC04B6B0B2DFFE3CB01E3F2FE3CFF7F" author="Beddome, R. H." box="[321,509,151,177]" pageId="17" pageNumber="350" pagination="225 - 229" refId="ref10547" refString="Beddome, R. H. (1863) Descriptions of new species of the family Uropeltidae from Southern India, with notes on other littleknown species. Proceedings of the Zoological Society of London, 1863, 225 - 229." type="book chapter" year="1863">Beddome (1863)</bibRefCitation>
did not specify the number of specimens available to him in his original description of
<taxonomicName id="4C514D190B2DFFE3CAF6E3DEFE69FF1B" baseAuthorityName="Boulenger" baseAuthorityYear="1863" box="[182,424,187,213]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CAF6E3DEFE69FF1B" box="[182,424,187,213]" italics="true" pageId="17" pageNumber="350">Uropeltis pulneyensis</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C514D190B2DFFE3CB97E3DEFD16FF1B" authorityName="Beddome" authorityYear="1863" box="[471,727,187,213]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CB97E3DEFD16FF1B" box="[471,727,187,213]" italics="true" pageId="17" pageNumber="350">Plectrurus pulneyensis</emphasis>
</taxonomicName>
). It can be inferred that there were likely at least two (probably at least three) type specimens, given that Beddome reported the number of subcaudals as “male, about 12, female 68”. Subsequent workers have suggested up to
<specimenCount id="9D57FD130B2DFFE3C8F6E266FC93FED3" box="[694,850,259,285]" count="9" pageId="17" pageNumber="350" type="generic" typeStatus="syntype">nine syntypes</specimenCount>
, currently stored in Kolkata (ZSI 4381 &amp; 6948 from “Palni Hills”; 6950 &amp; 697275 from “Madras Presidency”: but see
<bibRefCitation id="EFC04B6B0B2DFFE3C9D6E242FBE4FE8F" author="Gans, C." box="[918,1061,295,321]" pageId="17" pageNumber="350" refId="ref11399" refString="Gans, C. (1966) Liste der Rezenten Amphibien und Reptilien: Uropeltidae. W. de Gruyter, Berlin, 34 pp." type="book" year="1966">Gans [1966]</bibRefCitation>
, who reported that there were no ZSI types of this species), London (
<collectionCode id="ED40AE5F0B2DFFE3C87FE22EFD54FEAB" box="[575,661,331,357]" pageId="17" pageNumber="350">BMNH</collectionCode>
1946.1.17.6, previously
<collectionCode id="ED40AE5F0B2DFFE3C9F8E22EFBCDFEAB" box="[952,1036,331,357]" pageId="17" pageNumber="350">BMNH</collectionCode>
64.3.9.1, “Anamallays” according to accession register but Pulney Hills according to Boulenger 1863:
<figureCitation id="136A2A1F0B2DFFE3C9EAE20AFC2EFE47" box="[938,1007,367,393]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[303,1283,181,1919]" captionTargetId="figure-17@15.[303,1283,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lectotype (by present designation) of Uropeltis pulneyensis (Boulenger, 1863) (BMNH 1946.1.17.6, previously BMNH 64.3.9.1). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732851" httpUri="https://zenodo.org/record/12732851/files/figure.png" pageId="17" pageNumber="350">Fig. 8</figureCitation>
), and Paris (MNHN 1895.78, Pulney Hills) (
<bibRefCitation id="EFC04B6B0B2DFFE3CAA8E2F6FE5AFE63" author="Das, I. &amp; Dattagupta, B. &amp; Gayen, N. C." box="[232,411,403,429]" pageId="17" pageNumber="350" pagination="121 - 172" refId="ref10914" refString="Das, I., Dattagupta, B. &amp; Gayen, N. C. (1998) History and catalogue of reptile types in the collection of the Zoological Survey of India. Journal of South Asian Natural History 3, 121 - 172." type="journal article" year="1998">
Das
<emphasis id="B925EA880B2DFFE3CB5DE2F6FE96FE63" box="[285,343,403,429]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="EFC04B6B0B2DFFE3CBE8E2F6FD70FE63" author="McDiarmid, R. W. &amp; Campbell, J. A. &amp; Tourei, T." box="[424,689,403,429]" pageId="17" pageNumber="350" refId="ref11926" refString="McDiarmid, R. W., Campbell, J. A. &amp; Tourei, T. (1999) Snake Species of the World. A Taxonomic and Geographic Reference. Vol. 1. The Herpetologists' League, Washington, 511 pp." type="book" year="1999">
McDiarmid
<emphasis id="B925EA880B2DFFE3C873E2F6FDACFE63" box="[563,621,403,429]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="EFC04B6B0B2DFFE3C8FEE2F6FC49FE63" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[702,904,403,429]" pageId="17" pageNumber="350" pagination="453 - 506" refId="ref12156" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V. &amp; Somaweera, R. (2016) A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema, 38 (4), 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2" type="journal article" year="2016">
Pyron
<emphasis id="B925EA880B2DFFE3C94AE2F6FC85FE63" box="[778,836,403,429]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2016
</bibRefCitation>
;
<bibRefCitation id="EFC04B6B0B2DFFE3C9D5E2F6FBB0FE63" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." box="[917,1137,403,429]" pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3C9B1E2F6FBEAFE63" box="[1009,1067,403,429]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2023
</bibRefCitation>
). We did not examine ZSI specimens, but MNHN 1895.78 has 12,11 subcaudals and
<collectionCode id="ED40AE5F0B2DFFE3C977E2D2FC4AFE1F" box="[823,907,439,465]" pageId="17" pageNumber="350">BMNH</collectionCode>
1946.1.17.6 has 6,7 subcaudals so both could be from Beddomes type series. The accession date of the
<collectionCode id="ED40AE5F0B2DFFE3C96FE2BEFC42FE3B" box="[815,899,475,501]" pageId="17" pageNumber="350">BMNH</collectionCode>
specimen (
<date id="FFEF105A0B2DFFE3CE45E2BEFB50FE3B" box="[1029,1169,475,501]" pageId="17" pageNumber="350" value="1864-03">March 1864</date>
) is also consistent with this. Given that we have examined it in detail, that it is in good condition (unlike the ZSI specimens:
<bibRefCitation id="EFC04B6B0B2DFFE3CF49E29AFF12FDF3" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3CF24E29AFA5CFDD7" box="[1380,1437,511,537]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2023
</bibRefCitation>
), and colour photographs are provided here (
<figureCitation id="136A2A1F0B2DFFE3C899E146FCDFFDF3" box="[729,798,547,573]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[303,1283,181,1919]" captionTargetId="figure-17@15.[303,1283,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Lectotype (by present designation) of Uropeltis pulneyensis (Boulenger, 1863) (BMNH 1946.1.17.6, previously BMNH 64.3.9.1). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732851" httpUri="https://zenodo.org/record/12732851/files/figure.png" pageId="17" pageNumber="350">Fig. 8</figureCitation>
), we designate
<collectionCode id="ED40AE5F0B2DFFE3C994E146FBE9FDF3" box="[980,1064,547,573]" pageId="17" pageNumber="350">BMNH</collectionCode>
1946.1.17.6 (previously
<collectionCode id="ED40AE5F0B2DFFE3CF08E146FA5DFDF3" box="[1352,1436,547,573]" pageId="17" pageNumber="350">BMNH</collectionCode>
64.3.9.1) as the
<typeStatus id="54EA88380B2DFFE3CB0FE122FE77FDAF" box="[335,438,583,609]" pageId="17" pageNumber="350" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C514D190B2DFFE3CB9FE122FC5EFDAF" authority="Beddome, 1863" authorityName="Beddome" authorityYear="1863" box="[479,927,583,609]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CB9FE122FD20FDAF" box="[479,737,583,609]" italics="true" pageId="17" pageNumber="350">Plectrurus pulneyensis</emphasis>
<bibRefCitation id="EFC04B6B0B2DFFE3C8A9E122FC5EFDAF" author="Beddome, R. H." box="[745,927,583,609]" pageId="17" pageNumber="350" pagination="225 - 229" refId="ref10547" refString="Beddome, R. H. (1863) Descriptions of new species of the family Uropeltidae from Southern India, with notes on other littleknown species. Proceedings of the Zoological Society of London, 1863, 225 - 229." type="book chapter" year="1863">Beddome, 1863</bibRefCitation>
</taxonomicName>
. At least MNHN 1895.78 is a
<typeStatus id="54EA88380B2DFFE3CEBEE122FA59FDAF" box="[1278,1432,583,609]" pageId="17" pageNumber="350" type="paralectotype">paralectotype</typeStatus>
, other
<typeStatus id="54EA88380B2DFFE3CA9AE10EFEBDFD4B" box="[218,380,619,645]" pageId="17" pageNumber="350" type="paralectotype">paralectotypes</typeStatus>
are likely present in the ZSI collection (e.g.,
<bibRefCitation id="EFC04B6B0B2DFFE3C9CCE10EFBA8FD4B" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." box="[908,1129,619,645]" pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3C9A9E10EFBE2FD4B" box="[1001,1059,619,645]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2023
</bibRefCitation>
).
<bibRefCitation id="EFC04B6B0B2DFFE3CE3EE10EFAFCFD4B" author="Beddome, R. H." box="[1150,1341,619,645]" pageId="17" pageNumber="350" pagination="3 - 33" refId="ref10496" refString="Beddome, R. H. (1886) An account of the earth-snakes of the peninsula of India and Ceylon. Annals and Magazine of Natural History, Series 5, 17 (97), 3 - 33. https: // doi. org / 10.1080 / 00222938609460106" type="journal article" year="1886">Beddome (1886)</bibRefCitation>
thought there was no sexual dimorphism in numbers of ventrals in
<taxonomicName id="4C514D190B2DFFE3C969E1EAFC0FFD67" authorityName="Beddome" authorityYear="1863" box="[809,974,655,681]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C969E1EAFC0FFD67" box="[809,974,655,681]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
, but his sample size might not have been large, and examination of more specimens shows that females, on average, have more ventrals as well as nonoverlapping numbers of subcaudals (
<figureCitation id="136A2A1F0B2DFFE3C875E1B2FDBBFD3F" box="[565,634,727,753]" captionStart="FIGURE 6" captionStartId="10.[151,250,1922,1946]" captionTargetBox="[159,1428,1031,1891]" captionTargetId="figure-414@10.[151,1436,1029,1899]" captionTargetPageId="10" captionText="FIGURE 6. Variation in numbers of ventral and subcaudal shields in females (circles) and males (triangles) of Uropeltis pulneyensis (25 females, 19 males), U. caudomaculata sp. nov. (6 females, 2 males) and the holotype of U. guentheri comb. nov. (one male). See Tables 3 and 4 for sources of data." figureDoi="http://doi.org/10.5281/zenodo.12732845" httpUri="https://zenodo.org/record/12732845/files/figure.png" pageId="17" pageNumber="350">Fig. 6</figureCitation>
; see also
<bibRefCitation id="EFC04B6B0B2DFFE3C8A6E1B2FC48FD3F" author="Boettger, O." box="[742,905,727,753]" pageId="17" pageNumber="350" pagination="65 - 164" refId="ref10589" refString="Boettger, O. (1892) Listen von Kriechtieren und Lurchen aus dem tropischen Asien und aus Papuasien. Berichte uber die Tatigkeit des Offenbacher Vereins fur Naturkunde, 29 - 32, 65 - 164." type="book chapter" year="1892">Boettger 1892</bibRefCitation>
, noting that it is not clear how the anteriormost ventral shield was identified in that study).
</paragraph>
<paragraph id="8BEE369A0B2DFFE3CA87E07AFE83FA9B" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="350">
As far as we know,
<taxonomicName id="4C514D190B2DFFE3CBE9E07AFD8EFCF7" authorityName="Beddome" authorityYear="1863" box="[425,591,799,825]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CBE9E07AFD8EFCF7" box="[425,591,799,825]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
occurs only in the Palani Hills, the westernmost specimen records that we know of being Kurduvadi (Kadavari?; ca.
<geoCoordinate id="EE65505D0B2DFFE3C82FE026FD24FC93" box="[623,741,835,861]" degrees="10.216" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="55" value="10.216">10.216° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B2DFFE3C8B0E026FCA3FC93" box="[752,866,835,861]" degrees="77.286" direction="east" orientation="longitude" pageId="17" pageNumber="350" precision="55" value="77.286">77.286° E</geoCoordinate>
, ca.
<quantity id="4CA99B7F0B2DFFE3C9D5E026FC31FC93" box="[917,1008,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="350" unit="m" value="2200.0">2,200 m</quantity>
elevation according to Google Earth), Kukkal (Kookal; ca.
<geoCoordinate id="EE65505D0B2DFFE3CB3FE003FE35FC4F" box="[383,500,870,897]" degrees="10.295" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="55" value="10.295">10.295° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B2DFFE3CBBEE003FDAEFC4F" box="[510,623,870,897]" degrees="77.365" direction="east" orientation="longitude" pageId="17" pageNumber="350" precision="55" value="77.365">77.365° E</geoCoordinate>
, ca.
<quantity id="4CA99B7F0B2DFFE3C8E6E002FCC0FC4F" box="[678,769,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="350" unit="m" value="1800.0">1,800 m</quantity>
elevation according to Google Earth) and Marian Shola (ca.
<geoCoordinate id="EE65505D0B2DFFE3CAD7E0EFFECDFC6B" box="[151,268,906,933]" degrees="10.155" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="55" value="10.155">10.155° N</geoCoordinate>
, 77.77.348° E, ca.
<quantity id="4CA99B7F0B2DFFE3CBA2E0EEFDFCFC6B" box="[482,573,907,933]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="350" unit="m" value="2200.0">2,200 m</quantity>
elevation according to Google Earth). The imprecise locality records of “near Madras” (MCZ 3870, 470424703; AMNH 77607) and “Nilgherries” (ZMB 5538) are here considered questionable.
<bibRefCitation id="EFC04B6B0B2DFFE3CAD7E0B6FE40FC23" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." box="[151,385,979,1005]" pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3CAB4E0B6FEEFFC23" box="[244,302,979,1005]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
(2023)
</bibRefCitation>
additionally listed localities for
<taxonomicName id="4C514D190B2DFFE3C8B5E0B6FC5DFC23" authorityName="Beddome" authorityYear="1863" box="[757,924,979,1005]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C8B5E0B6FC5DFC23" box="[757,924,979,1005]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
in Meghamalai (High Wavy Mountains) and in Nilambur. We have not examined the reported voucher specimens, and
<bibRefCitation id="EFC04B6B0B2DFFE3C995E092FB7AFBDF" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." box="[981,1211,1015,1041]" pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3CE70E092FBA8FBDF" box="[1072,1129,1015,1041]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
(2023)
</bibRefCitation>
presented ranges of scalation data rather than disaggregated for individual specimens. However, photographs of Meghamalai specimens previously published appear to be of
<taxonomicName id="4C514D190B2DFFE3C806E75AFCD9FB97" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[582,792,1087,1113]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3C806E75AFCD9FB97" box="[582,792,1087,1113]" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3C962E75AFCBFFB97" bold="true" box="[802,894,1087,1113]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3C962E75AFCBFFB97" box="[802,894,1087,1113]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
based on colour and number of midline scales (see Ganesh
<emphasis id="B925EA880B2DFFE3CB6AE706FEA4FBB3" box="[298,357,1123,1149]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2022; specimens identified as
<taxonomicName id="4C514D190B2DFFE3C889E706FC57FBB3" authorityName="Beddome" authorityYear="1863" box="[713,918,1123,1149]" class="Squamata" family="Uropeltidae" genus="Plectrurus" isUncertain="true" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C889E706FD25FBB2" box="[713,740,1123,1148]" italics="true" pageId="17" pageNumber="350">U.</emphasis>
cf.
<emphasis id="B925EA880B2DFFE3C953E706FC57FBB3" box="[787,918,1123,1149]" italics="true" pageId="17" pageNumber="350">pulneyensis</emphasis>
</taxonomicName>
). Nilambur (
<geoCoordinate id="EE65505D0B2DFFE3CE6BE706FB0FFBB3" box="[1067,1230,1123,1149]" degrees="11.276944" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="1" value="11.276944">11.276944° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B2DFFE3CE9BE707FAB9FBB3" box="[1243,1400,1122,1149]" degrees="76.225833" direction="east" orientation="longitude" pageId="17" pageNumber="350" precision="1" value="76.22583">76.225833° E</geoCoordinate>
) is highly disjunct from verified localities for both
<taxonomicName id="4C514D190B2DFFE3C8FBE7E2FCA2FB6F" authorityName="Beddome" authorityYear="1863" box="[699,867,1159,1185]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C8FBE7E2FCA2FB6F" box="[699,867,1159,1185]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D190B2DFFE3C9DDE7E2FBAFFB6F" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[925,1134,1159,1185]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3C9DDE7E2FBAFFB6F" box="[925,1134,1159,1185]" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3CE36E7E2FB13FB6F" bold="true" box="[1142,1234,1159,1185]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3CE36E7E2FB13FB6F" box="[1142,1234,1159,1185]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, lying at only ca.
<quantity id="4CA99B7F0B2DFFE3CAD7E7CEFF1EFB0B" box="[151,223,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="350" unit="m" value="400.0">400 m</quantity>
elevation, more than
<quantity id="4CA99B7F0B2DFFE3CB96E7CFFDECFB0B" box="[470,557,1194,1221]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="350" unit="km" value="150.0">150 km</quantity>
north of Munnar on the other side of the lowland Palghat (Palakkad) Gap, so we consider that this locality is doubtful for either species. Four MNHN specimens are reportedly from Kurduvadi (Appendix 1), though it seems extremely unlikely that this is the town in southern
<collectingRegion id="4995F8780B2DFFE3CE68E796FB75FAC3" box="[1064,1204,1267,1293]" country="India" name="Maharashtra" pageId="17" pageNumber="350">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE65505D0B2DFFE3CE83E796FAFBFAC3" box="[1219,1338,1267,1293]" degrees="18.097" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="55" value="18.097">18.097° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B2DFFE3CF04E797FF6AFAFF" degrees="75.425" direction="east" orientation="longitude" pageId="17" pageNumber="350" precision="55" value="75.425">75.425° E</geoCoordinate>
), almost
<quantity id="4CA99B7F0B2DFFE3CB57E672FEAEFAFF" box="[279,367,1303,1329]" metricMagnitude="5" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="350" unit="km" value="900.0">900 km</quantity>
to the north of the Palani Hills at below
<quantity id="4CA99B7F0B2DFFE3C916E672FC61FAFF" box="[854,928,1303,1329]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="350" unit="m" value="600.0">600 m</quantity>
elevation in the Deccan Plateau rather than Western Ghats.
</paragraph>
<paragraph id="8BEE369A0B2DFFE3CA87E63AFCE0F8BF" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="350">
<bibRefCitation id="EFC04B6B0B2DFFE3CA87E63AFE4AFAB7" author="Rajendran, M. V." box="[199,395,1374,1401]" pageId="17" pageNumber="350" refId="ref12233" refString="Rajendran, M. V. (1985) Studies in uropeltid snakes. Madurai Kamaraj University, Madurai, 132 pp." type="book" year="1985">Rajendran (1985)</bibRefCitation>
included a population from Sevenmalai, near Munnar under
<taxonomicName id="4C514D190B2DFFE3CE75E63AFB1BFAB7" authorityName="Beddome" authorityYear="1863" box="[1077,1242,1375,1401]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CE75E63AFB1BFAB7" box="[1077,1242,1375,1401]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
(see also
<bibRefCitation id="EFC04B6B0B2DFFE3CF08E63AFED9FA53" author="Ganesh, S. R. &amp; Adhikari, O. D. &amp; Srikanthan, A. N." pageId="17" pageNumber="350" refId="ref11329" refString="Ganesh, S. R., Adhikari, O. D. &amp; Srikanthan, A. N. (2023) Taxonomy and distribution of two shieldtail snakes Uropeltis pulneyensis (Beddome, 1863) and Uropeltis woodmasoni (Theobald, 1876) (Squamata: Uropeltidae) with redescriptions of type specimens. Journal of the Bombay Natural History Society, 120. [published online]" type="journal volume" year="2023">
Ganesh
<emphasis id="B925EA880B2DFFE3CAD7E6E6FF13FA53" box="[151,210,1411,1437]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
2023
</bibRefCitation>
), but we have seen no specimens of
<taxonomicName id="4C514D190B2DFFE3C886E6E6FCAEFA53" authorityName="Beddome" authorityYear="1863" box="[710,879,1411,1437]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C886E6E6FCAEFA53" box="[710,879,1411,1437]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
from in or very close to Munnar, and all of the
<taxonomicName id="4C514D190B2DFFE3CAD7E6C2FEC1FA0F" authorityName="Cuvier" authorityYear="1829" box="[151,256,1447,1473]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="genus">
<emphasis id="B925EA880B2DFFE3CAD7E6C2FEC1FA0F" box="[151,256,1447,1473]" italics="true" pageId="17" pageNumber="350">Uropeltis</emphasis>
</taxonomicName>
specimens we have seen from that locality that have a tail morphology generally like that of
<taxonomicName id="4C514D190B2DFFE3CEB9E6C2FA5CFA0F" authorityName="Beddome" authorityYear="1863" box="[1273,1437,1447,1473]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CEB9E6C2FA5CFA0F" box="[1273,1437,1447,1473]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
were subsequently identified as
<taxonomicName id="4C514D190B2DFFE3C84DE6AEFC85FA2B" authority="Gower, 2023" authorityName="Gower" authorityYear="2023" box="[525,836,1483,1509]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="tricuspida">
<emphasis id="B925EA880B2DFFE3C84DE6AEFD64FA2B" box="[525,677,1483,1509]" italics="true" pageId="17" pageNumber="350">U. tricuspida</emphasis>
<bibRefCitation id="EFC04B6B0B2DFFE3C8EFE6AEFC85FA2B" author="Gower, D. J." box="[687,836,1483,1509]" pageId="17" pageNumber="350" pagination="103 - 109" refId="ref11425" refString="Gower, D. J. (2023) On the taxonomy of Uropeltis petersi (Beddome, 1878) (Serpentes: Uropeltidae) and description of a new, closely related species from the Western Ghats of India. Zootaxa, 5319 (1), 103 - 109. https: // doi. org / 10.11646 / zootaxa. 5319.1.7" type="journal article" year="2023">Gower, 2023</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFC04B6B0B2DFFE3C916E6AEFC2AFA2B" author="Gower, D. J." box="[854,1003,1483,1509]" pageId="17" pageNumber="350" pagination="103 - 109" refId="ref11425" refString="Gower, D. J. (2023) On the taxonomy of Uropeltis petersi (Beddome, 1878) (Serpentes: Uropeltidae) and description of a new, closely related species from the Western Ghats of India. Zootaxa, 5319 (1), 103 - 109. https: // doi. org / 10.11646 / zootaxa. 5319.1.7" type="journal article" year="2023">Gower 2023</bibRefCitation>
).
<bibRefCitation id="EFC04B6B0B2DFFE3CE41E6AEFB08FA2B" author="Rajendran, M. V." box="[1025,1225,1482,1509]" pageId="17" pageNumber="350" refId="ref12233" refString="Rajendran, M. V. (1985) Studies in uropeltid snakes. Madurai Kamaraj University, Madurai, 132 pp." type="book" year="1985">Rajendran (1985)</bibRefCitation>
also reported
<specimenCount id="9D57FD130B2DFFE3CF34E6AEFEC0F9C7" count="1" pageId="17" pageNumber="350" type="generic">one specimen</specimenCount>
of
<taxonomicName id="4C514D190B2DFFE3CB68E68AFE0FF9C7" authorityName="Beddome" authorityYear="1863" box="[296,462,1519,1545]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CB68E68AFE0FF9C7" box="[296,462,1519,1545]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
from a
<quantity id="4CA99B7F0B2DFFE3C868E68BFDAEF9C7" box="[552,623,1518,1545]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="350" unit="m" value="250.0">250 m</quantity>
elevation site in the Alargarkoyil hills (ca.
<geoCoordinate id="EE65505D0B2DFFE3CE1AE68AFB02F9C7" box="[1114,1219,1519,1545]" degrees="10.12" direction="north" orientation="latitude" pageId="17" pageNumber="350" precision="555" value="10.12">10.12° N</geoCoordinate>
,
<geoCoordinate id="EE65505D0B2DFFE3CE8FE68AFAF4F9C7" box="[1231,1333,1519,1545]" degrees="78.23" direction="east" orientation="longitude" pageId="17" pageNumber="350" precision="555" value="78.23">78.23° E</geoCoordinate>
), a range below
<quantity id="4CA99B7F0B2DFFE3CAA3E576FEEBF9E3" box="[227,298,1555,1581]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="350" unit="m" value="900.0">900 m</quantity>
elevation near Madurai, disjunct from the main ranges of the Western Ghats. We do not know of any museum specimens or published images of uropeltids from the Alargarkoyil Hills and are so unable to comment on the identity of that population. The known distributions of
<taxonomicName id="4C514D190B2DFFE3C920E53EFBC6F9BB" authorityName="Beddome" authorityYear="1863" box="[864,1031,1627,1653]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C920E53EFBC6F9BB" box="[864,1031,1627,1653]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D190B2DFFE3CE02E53EFAD2F9BB" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[1090,1299,1627,1653]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3CE02E53EFAD2F9BB" box="[1090,1299,1627,1653]" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3CF5CE53EFAB6F9BB" bold="true" box="[1308,1399,1627,1653]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3CF5CE53EFAB6F9BB" box="[1308,1399,1627,1653]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
do not overlap (assuming that
<taxonomicName id="4C514D190B2DFFE3CB87E51AFD59F957" box="[455,664,1663,1689]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2DFFE3CB87E51AFD59F957" box="[455,664,1663,1689]" italics="true" pageId="17" pageNumber="350">Silybura guentheri</emphasis>
</taxonomicName>
is not a synonym of
<taxonomicName id="4C514D190B2DFFE3C9C2E51AFBE6F957" authorityName="Beddome" authorityYear="1863" box="[898,1063,1663,1689]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C9C2E51AFBE6F957" box="[898,1063,1663,1689]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
: see below), but the westernmost known localities of
<taxonomicName id="4C514D190B2DFFE3CB38E5C6FC14F973" authority="(Kadavari, Kukkal and Marian Shola)" baseAuthorityName="Kadavari, Kukkal and Marian Shola" box="[376,981,1699,1725]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CB38E5C6FDDEF973" box="[376,543,1699,1725]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
(Kadavari, Kukkal and Marian Shola)
</taxonomicName>
, and northernmost-known locality of
<taxonomicName id="4C514D190B2DFFE3CFC1E5C6FE85F92F" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3CFC1E5C6FE85F92F" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3CB0DE5A2FE68F92F" bold="true" box="[333,425,1735,1761]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3CB0DE5A2FE68F92F" box="[333,425,1735,1761]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Yellapetty, Munnar) are only approximately 14, 16 and
<quantity id="4CA99B7F0B2DFFE3CE7BE5A3FB45F92F" box="[1083,1156,1734,1761]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="350" unit="km" value="25.0">25 km</quantity>
apart, respectively, so it is possible that the two species are sympatric somewhere in this region. In Deepak
<emphasis id="B925EA880B2DFFE3CE0DE58EFB46F8CB" box="[1101,1159,1771,1797]" italics="true" pageId="17" pageNumber="350">et al.</emphasis>
s (2022: 55) map of the distribution of
<taxonomicName id="4C514D190B2DFFE3CB01E46AFE29F8E7" box="[321,488,1807,1833]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CB01E46AFE29F8E7" box="[321,488,1807,1833]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
, the westernmost dot (Munnar) is
<bibRefCitation id="EFC04B6B0B2DFFE3C935E46AFB8FF8E7" author="Rajendran, M. V." box="[885,1102,1806,1833]" pageId="17" pageNumber="350" refId="ref12233" refString="Rajendran, M. V. (1985) Studies in uropeltid snakes. Madurai Kamaraj University, Madurai, 132 pp." type="book" year="1985">Rajendrans (1985)</bibRefCitation>
record from Sevenmalai that possibly pertains instead to
<taxonomicName id="4C514D190B2DFFE3CB8CE456FD9EF883" authorityName="Gower" authorityYear="2023" box="[460,607,1843,1869]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="tricuspida">
<emphasis id="B925EA880B2DFFE3CB8CE456FD9EF883" box="[460,607,1843,1869]" italics="true" pageId="17" pageNumber="350">U. tricuspida</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC04B6B0B2DFFE3C8D9E456FCE4F883" author="Gower, D. J." box="[665,805,1843,1869]" pageId="17" pageNumber="350" pagination="103 - 109" refId="ref11425" refString="Gower, D. J. (2023) On the taxonomy of Uropeltis petersi (Beddome, 1878) (Serpentes: Uropeltidae) and description of a new, closely related species from the Western Ghats of India. Zootaxa, 5319 (1), 103 - 109. https: // doi. org / 10.11646 / zootaxa. 5319.1.7" type="journal article" year="2023">Gower 2023</bibRefCitation>
) and the dot shortly to the east of that (Yellapetty) and in Meghamalai pertain instead to
<taxonomicName id="4C514D190B2DFFE3CBB2E432FD00F8BF" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[498,705,1879,1905]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3CBB2E432FD00F8BF" box="[498,705,1879,1905]" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3C888E432FCE0F8BF" bold="true" box="[712,801,1879,1905]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3C888E432FCE0F8BF" box="[712,801,1879,1905]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEE369A0B2DFFE0CA87E41EFEEDFED3" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1437,151,1581]" lastPageId="18" lastPageNumber="351" pageId="17" pageNumber="350">
<bibRefCitation id="EFC04B6B0B2DFFE3CA87E41EFE0EF85B" author="Hutton, A. F. &amp; David, P." box="[199,463,1915,1941]" pageId="17" pageNumber="350" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David (2008)</bibRefCitation>
identified the
<collectionCode id="ED40AE5F0B2DFFE3C831E41EFD04F85B" box="[625,709,1915,1941]" pageId="17" pageNumber="350">BMNH</collectionCode>
<typeStatus id="54EA88380B2DFFE3C88CE41EFD3DF85B" box="[716,764,1915,1941]" pageId="17" pageNumber="350">type</typeStatus>
specimens of
<taxonomicName id="4C514D190B2DFFE3C9DCE41EFBAAF85B" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[924,1131,1915,1941]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2DFFE3C9DCE41EFBAAF85B" box="[924,1131,1915,1941]" italics="true" pageId="17" pageNumber="350">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2DFFE3CE32E41EFB0AF85B" bold="true" box="[1138,1227,1915,1941]" pageId="17" pageNumber="350">
<taxonomicNameLabel id="A21657F30B2DFFE3CE32E41EFB0AF85B" box="[1138,1227,1915,1941]" pageId="17" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as
<taxonomicName id="4C514D190B2DFFE3CEB0E41EFA57F85B" box="[1264,1430,1915,1941]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3CEB0E41EFA57F85B" box="[1264,1430,1915,1941]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC04B6B0B2DFFE3CAD7E4FAFE27F877" author="Hutton, A. F. &amp; David, P." box="[151,486,1951,1977]" pageId="17" pageNumber="350" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David (2008: 302)</bibRefCitation>
mentioned seven
<collectionCode id="ED40AE5F0B2DFFE3C8FCE4FAFCD1F877" box="[700,784,1951,1977]" pageId="17" pageNumber="350">BMNH</collectionCode>
specimens of
<taxonomicName id="4C514D190B2DFFE3C9FCE4FAFBA5F877" box="[956,1124,1951,1977]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="17" pageNumber="350" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2DFFE3C9FCE4FAFBA5F877" box="[956,1124,1951,1977]" italics="true" pageId="17" pageNumber="350">U. pulneyensis</emphasis>
</taxonomicName>
from Meghamalai, but we assume that this is an inadvertent error because they reported only the same five
<collectionCode id="ED40AE5F0B2DFFE3CE67E4A6FBBAF813" box="[1063,1147,1987,2013]" pageId="17" pageNumber="350">BMNH</collectionCode>
specimens that we report here. The sex, size and scalation data presented by
<bibRefCitation id="EFC04B6B0B2EFFE0C8A9E3F2FC38FF7F" author="Hutton, A. F. &amp; David, P." box="[745,1017,151,177]" pageId="18" pageNumber="351" pagination="299 - 316" refId="ref11522" refString="Hutton, A. F. &amp; David, P. (2008) Note on a collection of snakes from south India, with emphasis on the snake fauna of the Meghamalai Hills (High Wavy Mountains) Journal of the Bombay Natural History Society, 105, 299 - 316." type="journal article" year="2008">Hutton &amp; David (2008)</bibRefCitation>
for these
<specimenCount id="9D57FD130B2EFFE0CE2FE3F2FADBFF7F" box="[1135,1306,151,177]" count="5" pageId="18" pageNumber="351" type="generic">five specimens</specimenCount>
are greatly at odds with our observations (see
<tableCitation id="C6D303210B2EFFE0C86DE3DEFD45FF1B" box="[557,644,187,213]" captionStart="TABLE 3" captionStartId="11.[151,239,467,493]" captionTargetPageId="11" captionText="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="18" pageNumber="351" tableUuid="DF2E66120B37FFF9CAD7E2B6F884FDDF">Table 3</tableCitation>
). We have no explanation for these discrepancies, even allowing for some potential mixing of data and/or specimen tags among the
<specimenCount id="9D57FD130B2EFFE0C936E3BAFBE1FF37" box="[886,1056,223,249]" count="5" pageId="18" pageNumber="351" type="generic">five specimens</specimenCount>
subsequent to Hutton &amp; Davids observations.
</paragraph>
<paragraph id="8BEE369A0B2EFFE0CAB7E242FE36FE3B" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="351">
All of the
<taxonomicName id="4C514D190B2EFFE0CB2DE242FD4AFE8F" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[365,651,295,321]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CB2DE242FD4AFE8F" box="[365,651,295,321]" italics="true" pageId="18" pageNumber="351">Uropeltis caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0C8D3E242FD2CFE8F" bold="true" box="[659,749,295,321]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0C8D3E242FD2CFE8F" box="[659,749,295,321]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens we have examined from Meghamalai differ from
<taxonomicName id="4C514D190B2EFFE0CAD7E22EFEFFFEAB" box="[151,318,331,357]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CAD7E22EFEFFFEAB" box="[151,318,331,357]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
in having yellow spots or blotches rather than stripes on the lateral surface of the base of the tail and on the back of the head and anterior end of the body. The two collected specimens of the new species from Yellapetty near Munnar resemble
<taxonomicName id="4C514D190B2EFFE0C85CE2F6FD02FE63" box="[540,707,403,429]" class="Squamata" family="Uropeltidae" genus="Plectrurus" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0C85CE2F6FD02FE63" box="[540,707,403,429]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
in this feature, but are identified here as
<taxonomicName id="4C514D190B2EFFE0CEE1E2F6FAB3FE63" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[1185,1394,403,429]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CEE1E2F6FAB3FE63" box="[1185,1394,403,429]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0CF3BE2F6FF09FE1F" bold="true" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0CF3BE2F6FF09FE1F" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
because they much more closely match the Meghamalai material in terms of numbers of ventral shields, and in mitochondrial DNA sequences.
</paragraph>
<paragraph id="8BEE369A0B2EFFE0CA87E29AFB3AFB6F" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="351">
<bibRefCitation id="EFC04B6B0B2EFFE0CA87E29AFE43FDD7" author="Beddome, R. H." box="[199,386,511,537]" pageId="18" pageNumber="351" pagination="154 - 155" refId="ref10438" refString="Beddome, R. H. (1878) Descriptions of new Uropeltidae from southern India, with remarks on some previously described species. Proceedings of the Zoological Society of London, 46, 154 - 155. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07946. x" type="journal article" year="1878">Beddome (1878)</bibRefCitation>
decribed
<taxonomicName id="4C514D190B2EFFE0CBABE29AFD7BFDD7" box="[491,698,511,537]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2EFFE0CBABE29AFD7BFDD7" box="[491,698,511,537]" italics="true" pageId="18" pageNumber="351">Silybura guentheri</emphasis>
</taxonomicName>
on the basis of a single specimen (
<collectionCode id="ED40AE5F0B2EFFE0CE6DE29AFB42FDD7" box="[1069,1155,511,537]" pageId="18" pageNumber="351">BMNH</collectionCode>
1946.1.16.74, previously
<collectionCode id="ED40AE5F0B2EFFE0CAD7E146FF2AFDF3" box="[151,235,547,573]" pageId="18" pageNumber="351">BMNH</collectionCode>
83.1.12.34:
<figureCitation id="136A2A1F0B2EFFE0CB36E146FE76FDF3" box="[374,439,547,573]" captionStart="FIGURE 9" captionStartId="16.[151,250,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-18@16.[151,1436,181,1821]" captionTargetPageId="16" captionText="FIGURE 9. Holotype of Uropeltis guentheri (Beddome, 1878) (BMNH 1946.1.16.74, previously BMNH 83.1.12.34). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732853" httpUri="https://zenodo.org/record/12732853/files/figure.png" pageId="18" pageNumber="351">Fig. 9</figureCitation>
) from High Wavy Mountains (Meghamalai). This species was listed as a junior subjective synonym of
<taxonomicName id="4C514D190B2EFFE0CB5FE122FDC7FDAF" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[287,518,583,609]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CB5FE122FDC7FDAF" box="[287,518,583,609]" italics="true" pageId="18" pageNumber="351">Silybura pulneyensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B6B0B2EFFE0C86BE122FD30FDAF" author="Boulenger, G. A." box="[555,753,583,609]" pageId="18" pageNumber="351" refId="ref10627" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.). Vol. I. Taylor &amp; Francis, London, 448 pp." type="book" year="1893">Boulenger (1893)</bibRefCitation>
, which seems to have been followed by subsequent taxonomic works (e.g., Smith 1943;
<bibRefCitation id="EFC04B6B0B2EFFE0CBF1E10EFDECFD4B" author="Gans, C." box="[433,557,619,645]" pageId="18" pageNumber="351" refId="ref11399" refString="Gans, C. (1966) Liste der Rezenten Amphibien und Reptilien: Uropeltidae. W. de Gruyter, Berlin, 34 pp." type="book" year="1966">Gans 1966</bibRefCitation>
;
<bibRefCitation id="EFC04B6B0B2EFFE0C878E10EFCFBFD4B" author="McDiarmid, R. W. &amp; Campbell, J. A. &amp; Tourei, T." box="[568,826,619,645]" pageId="18" pageNumber="351" refId="ref11926" refString="McDiarmid, R. W., Campbell, J. A. &amp; Tourei, T. (1999) Snake Species of the World. A Taxonomic and Geographic Reference. Vol. 1. The Herpetologists' League, Washington, 511 pp." type="book" year="1999">
McDiarmid
<emphasis id="B925EA880B2EFFE0C881E10EFD38FD4B" box="[705,761,619,645]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
1999
</bibRefCitation>
;
<bibRefCitation id="EFC04B6B0B2EFFE0C906E10EFBCBFD4B" author="Pyron, R. A. &amp; Ganesh, S. R. &amp; Sayyed, A. &amp; Sharma, V. &amp; Wallach, V. &amp; Somaweera, R." box="[838,1034,619,645]" pageId="18" pageNumber="351" pagination="453 - 506" refId="ref12156" refString="Pyron, R. A., Ganesh, S. R., Sayyed, A., Sharma, V., Wallach, V. &amp; Somaweera, R. (2016) A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae). Zoosystema, 38 (4), 453 - 506. https: // doi. org / 10.5252 / z 2016 n 4 a 2" type="journal article" year="2016">
Pyron
<emphasis id="B925EA880B2EFFE0C9CFE10EFC06FD4B" box="[911,967,619,645]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
2016
</bibRefCitation>
). The
<typeStatus id="54EA88380B2EFFE0CE0EE10EFB71FD4B" box="[1102,1200,619,645]" pageId="18" pageNumber="351" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C514D190B2EFFE0CE94E10EFA9AFD4B" box="[1236,1371,619,645]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2EFFE0CE94E10EFA9AFD4B" box="[1236,1371,619,645]" italics="true" pageId="18" pageNumber="351">S. guentheri</emphasis>
</taxonomicName>
does, indeed, resemble
<taxonomicName id="4C514D190B2EFFE0CB19E1EAFE3CFD67" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[345,509,655,681]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CB19E1EAFE3CFD67" box="[345,509,655,681]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
in having the nasal shields separated by the rostral, 17 dorsal scale rows at midbody, a tail matching Smiths (1943) Group III, and in being a male with 167 ventral shields and 12,12 subcaudals (
<figureCitation id="136A2A1F0B2EFFE0CF0BE1D6FA4FFD03" box="[1355,1422,691,717]" captionStart="FIGURE 6" captionStartId="10.[151,250,1922,1946]" captionTargetBox="[159,1428,1031,1891]" captionTargetId="figure-414@10.[151,1436,1029,1899]" captionTargetPageId="10" captionText="FIGURE 6. Variation in numbers of ventral and subcaudal shields in females (circles) and males (triangles) of Uropeltis pulneyensis (25 females, 19 males), U. caudomaculata sp. nov. (6 females, 2 males) and the holotype of U. guentheri comb. nov. (one male). See Tables 3 and 4 for sources of data." figureDoi="http://doi.org/10.5281/zenodo.12732845" httpUri="https://zenodo.org/record/12732845/files/figure.png" pageId="18" pageNumber="351">Fig. 6</figureCitation>
). However, as well as originating from Meghamalai versus the Palani Hills, the specimen differs from
<taxonomicName id="4C514D190B2EFFE0CEB8E1B2FA5CFD3F" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1272,1437,727,753]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CEB8E1B2FA5CFD3F" box="[1272,1437,727,753]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
notably in colour, as also noted by
<bibRefCitation id="EFC04B6B0B2EFFE0C865E19EFD1CFCDB" author="Beddome, R. H." box="[549,733,763,789]" pageId="18" pageNumber="351" pagination="154 - 155" refId="ref10438" refString="Beddome, R. H. (1878) Descriptions of new Uropeltidae from southern India, with remarks on some previously described species. Proceedings of the Zoological Society of London, 46, 154 - 155. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 07946. x" type="journal article" year="1878">Beddome (1878</bibRefCitation>
,
<bibRefCitation id="EFC04B6B0B2EFFE0C8A9E19EFCE3FCDB" author="Beddome, R. H." box="[745,802,763,789]" pageId="18" pageNumber="351" pagination="3 - 33" refId="ref10496" refString="Beddome, R. H. (1886) An account of the earth-snakes of the peninsula of India and Ceylon. Annals and Magazine of Natural History, Series 5, 17 (97), 3 - 33. https: // doi. org / 10.1080 / 00222938609460106" type="journal article" year="1886">1886</bibRefCitation>
) and
<bibRefCitation id="EFC04B6B0B2EFFE0C923E19EFBE7FCDB" author="Boulenger, G. A." box="[867,1062,763,789]" pageId="18" pageNumber="351" refId="ref10627" refString="Boulenger, G. A. (1893) Catalogue of the snakes in the British Museum (Nat. Hist.). Vol. I. Taylor &amp; Francis, London, 448 pp." type="book" year="1893">Boulenger (1893)</bibRefCitation>
—having an almost entirely pale venter (
<figureCitation id="136A2A1F0B2EFFE0CAADE07AFEF0FCF7" box="[237,305,799,825]" captionStart="FIGURE 9" captionStartId="16.[151,250,1844,1868]" captionTargetBox="[151,1436,181,1821]" captionTargetId="figure-18@16.[151,1436,181,1821]" captionTargetPageId="16" captionText="FIGURE 9. Holotype of Uropeltis guentheri (Beddome, 1878) (BMNH 1946.1.16.74, previously BMNH 83.1.12.34). Scale bar increments in millimetres. Photographs by Kevin Webb." figureDoi="http://doi.org/10.5281/zenodo.12732853" httpUri="https://zenodo.org/record/12732853/files/figure.png" pageId="18" pageNumber="351">Fig. 9</figureCitation>
), having a stripe (versus spots) on the head and tail (in which it differs from at least the Meghamalai specimens of
<taxonomicName id="4C514D190B2EFFE0CB72E026FDC0FC93" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[306,513,835,861]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CB72E026FDC0FC93" box="[306,513,835,861]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0C848E026FDA3FC93" bold="true" box="[520,610,835,861]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0C848E026FDA3FC93" box="[520,610,835,861]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and with pale spots on the uppermost dorsal scale rows being less dense (more mottled). It is not only colour and number of ventral scales in which the
<taxonomicName id="4C514D190B2EFFE0CE5BE002FB62FC4F" box="[1051,1187,871,897]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2EFFE0CE5BE002FB62FC4F" box="[1051,1187,871,897]" italics="true" pageId="18" pageNumber="351">S. guentheri</emphasis>
</taxonomicName>
<typeStatus id="54EA88380B2EFFE0CEEBE002FACCFC4F" box="[1195,1293,871,897]" pageId="18" pageNumber="351" type="holotype">holotype</typeStatus>
differs from
<taxonomicName id="4C514D190B2EFFE0CAD7E0EEFEABFC6B" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[151,362,907,933]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CAD7E0EEFEABFC6B" box="[151,362,907,933]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0CB33E0EEFE11FC6B" bold="true" box="[371,464,907,933]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0CB33E0EEFE11FC6B" box="[371,464,907,933]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—the former has relatively larger eyes (compared to, for example, the length of the ocular shield:
<tableCitation id="C6D303210B2EFFE0CB79E0CAFE4FFC07" box="[313,398,943,969]" captionStart="TABLE 3" captionStartId="11.[151,239,467,493]" captionTargetPageId="11" captionText="TABLE 3. Morphometric (in mm) and meristic data for holotype (*) and paratypes of Uropeltis caudomaculata sp. nov., the lectotype (†) and a paralectotype (#) of U. pulneyensis, and the holotype of U. guentheri comb. nov. (‡). Values in square brackets are those reported by Hutton &amp; David (2008). Blank cells indicate data not recorded." httpUri="http://table.plazi.org/id/DF2E66120B37FFF9CAD7E2B6F884FDDF" pageId="18" pageNumber="351" tableUuid="DF2E66120B37FFF9CAD7E2B6F884FDDF">Table 3</tableCitation>
), and also lacks midline contact between the opposite first infralabial shields (which make contact in all but one of the known specimens of
<taxonomicName id="4C514D190B2EFFE0C893E0B6FC64FC23" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[723,933,979,1005]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0C893E0B6FC64FC23" box="[723,933,979,1005]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0C9EDE0B6FBC8FC23" bold="true" box="[941,1033,979,1005]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0C9EDE0B6FBC8FC23" box="[941,1033,979,1005]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; rare outliers for this feature occur in
<taxonomicName id="4C514D190B2EFFE0CAF6E092FE9FFBDF" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[182,350,1015,1041]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CAF6E092FE9FFBDF" box="[182,350,1015,1041]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
: present in only 2 out of 25 of the specimens examined here for that feature). Pending further data, we consider
<taxonomicName id="4C514D190B2EFFE0CB24E77EFD81FBFB" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" baseAuthorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" baseAuthorityYear="1878" box="[356,576,1051,1077]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="guentheri" status="comb. nov.">
<emphasis id="B925EA880B2EFFE0CB24E77EFD81FBFB" box="[356,576,1051,1077]" italics="true" pageId="18" pageNumber="351">Uropeltis guentheri</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0C808E77EFD08FBFB" bold="true" box="[584,713,1051,1077]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0C808E77EFD08FBFB" box="[584,713,1051,1077]" pageId="18" pageNumber="351" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
to be a valid species, most similar to
<taxonomicName id="4C514D190B2EFFE0CE37E77EFADCFBFB" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[1143,1309,1051,1077]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CE37E77EFADCFBFB" box="[1143,1309,1051,1077]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
in external morphology but (if historical locality data are correct) occurring in Meghamalai rather than the Palani Hills. Other than finding additional specimens that resemble the only known specimen of
<taxonomicName id="4C514D190B2EFFE0CE4CE706FB5AFBB3" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" baseAuthorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" baseAuthorityYear="1878" box="[1036,1179,1123,1149]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B925EA880B2EFFE0CE4CE706FB5AFBB3" box="[1036,1179,1123,1149]" italics="true" pageId="18" pageNumber="351">U. guentheri</emphasis>
</taxonomicName>
, additional data might come in the form of internal anatomy (including osteology) and/or DNA sequences for the
<typeStatus id="54EA88380B2EFFE0CED2E7E2FB39FB6F" box="[1170,1272,1159,1185]" pageId="18" pageNumber="351" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BEE369A0B2EFFE0CA87E7CEFCEAF9E3" blockId="18.[151,1437,151,1581]" pageId="18" pageNumber="351">
There are, to the best of our knowledge, no other voucher specimens of the new species other than the
<specimenCount id="9D57FD130B2EFFE0CF24E7CEFECDFB27" count="8" pageId="18" pageNumber="351" type="generic">eight specimens</specimenCount>
reported here, though we suspect that several observations of
<taxonomicName id="4C514D190B2EFFE0C9A5E7AAFB79FB27" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[997,1208,1231,1257]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0C9A5E7AAFB79FB27" box="[997,1208,1231,1257]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0CE81E7AAFADFFB27" bold="true" box="[1217,1310,1231,1257]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0CE81E7AAFADFFB27" box="[1217,1310,1231,1257]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have been reported previously under various identifications. Probable observations of
<taxonomicName id="4C514D190B2EFFE0CE56E796FB2DFAC3" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[1046,1260,1267,1293]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CE56E796FB2DFAC3" box="[1046,1260,1267,1293]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0CEB9E796FA98FAC3" bold="true" box="[1273,1369,1267,1293]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0CEB9E796FA98FAC3" box="[1273,1369,1267,1293]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from Meghamalai were reported by
<bibRefCitation id="EFC04B6B0B2EFFE0CBACE672FD11FAFF" author="Ganesh, S. R. &amp; Asokan, S. &amp; Kannan, P." box="[492,720,1303,1329]" pageId="18" pageNumber="351" pagination="1 - 9" refId="ref11149" refString="Ganesh, S. R., Asokan, S. &amp; Kannan, P. (2008) A preliminary survey of amphibians and reptiles in Cardamom Hills, Western Ghats, Tamil Nadu. Cobra, 2 (2), 1 - 9." type="journal article" year="2008">
Ganesh
<emphasis id="B925EA880B2EFFE0C807E672FDBEFAFF" box="[583,639,1303,1329]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2008)
</bibRefCitation>
and
<bibRefCitation id="EFC04B6B0B2EFFE0C945E672FBA6FAFF" author="Chandramouli, S. R. &amp; Ganesh, S. R." box="[773,1127,1303,1329]" pageId="18" pageNumber="351" pagination="72 - 85" refId="ref10808" refString="Chandramouli, S. R. &amp; Ganesh, S. R. (2010) Herpetofauna of the Southern Western Ghats - reinvestigated after decades. Taprobanica, 2 (2), 72 - 85. https: // doi. org / 10.47605 / tapro. v 2 i 2.30" type="journal article" year="2010">Chandramouli &amp; Ganesh (2010</bibRefCitation>
; also
<bibRefCitation id="EFC04B6B0B2EFFE0CEE5E672FAB6FAFF" author="Ganesh, S. R. &amp; Bhupathy, S. &amp; David, P. &amp; Sathishkumar, N. &amp; Srinivas, G." box="[1189,1399,1303,1329]" pageId="18" pageNumber="351" pagination="53 - 64" refId="ref11196" refString="Ganesh, S. R., Bhupathy, S., David, P., Sathishkumar, N. &amp; Srinivas, G. (2014) Snake fauna of high wavy mountains, Western Ghats, India: species richness, status, and distribution pattern. Russian Journal of Herpetology, 21 (1), 53 - 64." type="journal article" year="2014">
Ganesh
<emphasis id="B925EA880B2EFFE0CEBFE672FAF1FAFF" box="[1279,1328,1303,1329]" italics="true" pageId="18" pageNumber="351">et al</emphasis>
. 2014
</bibRefCitation>
) as
<taxonomicName id="4C514D190B2EFFE0CAD7E65EFE8CFA9B" baseAuthorityName="Beddome" baseAuthorityYear="1877" box="[151,333,1339,1365]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="dindigalensis">
<emphasis id="B925EA880B2EFFE0CAD7E65EFE8CFA9B" box="[151,333,1339,1365]" italics="true" pageId="18" pageNumber="351">U. dindigalensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D190B2EFFE0CBC0E65EFD99FA9B" baseAuthorityName="Beddome" baseAuthorityYear="1877" box="[384,600,1339,1365]" class="Squamata" family="Uropeltidae" genus="Uropeltis" isUncertain="true" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="dindigalensis">
<emphasis id="B925EA880B2EFFE0CBC0E65EFE5DFA9A" box="[384,412,1339,1364]" italics="true" pageId="18" pageNumber="351">U.</emphasis>
cf.
<emphasis id="B925EA880B2EFFE0CB83E65EFD99FA9B" box="[451,600,1339,1365]" italics="true" pageId="18" pageNumber="351">dindigalensis</emphasis>
</taxonomicName>
, respectively, and as
<taxonomicName id="4C514D190B2EFFE0C901E65EFC25FA9B" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[833,996,1339,1365]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0C901E65EFC25FA9B" box="[833,996,1339,1365]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC04B6B0B2EFFE0CE4BE65EFACBFA9B" author="Chaitanya, R. &amp; Khandekar, A. &amp; Caleb, D. &amp; Mukherjee, N. &amp; Ghosh, A. &amp; Giri, V. B." box="[1035,1290,1339,1365]" pageId="18" pageNumber="351" pagination="21 - 37" refId="ref10737" refString="Chaitanya, R., Khandekar, A., Caleb, D., Mukherjee, N., Ghosh, A. &amp; Giri, V. B. (2018) Herpetofauna of the Meghamalai Wildlife Sanctuary, Southern Western Ghats, India: An updated checklist with annotations on taxonomy and nomenclature. Journal of the Bombay Natural History Society, 115, 21 - 37." type="journal article" year="2018">
Chaitanya
<emphasis id="B925EA880B2EFFE0CEC2E65EFB79FA9B" box="[1154,1208,1339,1365]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2018)
</bibRefCitation>
, and as
<taxonomicName id="4C514D190B2EFFE0CF1FE65EFEDBFAB7" baseAuthorityName="Beddome" baseAuthorityYear="1863" class="Squamata" family="Uropeltidae" genus="Silybura" isUncertain="true" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CF1FE65EFABBFA9A" box="[1375,1402,1339,1364]" italics="true" pageId="18" pageNumber="351">U.</emphasis>
cf.
<emphasis id="B925EA880B2EFFE0CAD7E63AFEDBFAB7" box="[151,282,1375,1401]" italics="true" pageId="18" pageNumber="351">pulneyensis</emphasis>
</taxonomicName>
by Ganesh
<emphasis id="B925EA880B2EFFE0CBDBE63AFE13FAB7" box="[411,466,1375,1401]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2022). Ganesh
<emphasis id="B925EA880B2EFFE0C8C8E63AFD7EFAB7" box="[648,703,1375,1401]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2022) reported that one uncollected specimen had approximately 190 ventral shields, 17 dorsal scale rows at midbody, and a tail matching Smiths (1943) Group III—data that, along with the colour pattern seen in published photographs (
<bibRefCitation id="EFC04B6B0B2EFFE0C942E6C2FC1BFA0F" author="Ganesh, S. R. &amp; Bhupathy, S. &amp; David, P. &amp; Sathishkumar, N. &amp; Srinivas, G." box="[770,986,1447,1473]" pageId="18" pageNumber="351" pagination="53 - 64" refId="ref11196" refString="Ganesh, S. R., Bhupathy, S., David, P., Sathishkumar, N. &amp; Srinivas, G. (2014) Snake fauna of high wavy mountains, Western Ghats, India: species richness, status, and distribution pattern. Russian Journal of Herpetology, 21 (1), 53 - 64." type="journal article" year="2014">
Ganesh
<emphasis id="B925EA880B2EFFE0C91FE6C2FC56FA0F" box="[863,919,1447,1473]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
2014
</bibRefCitation>
: fig. 6a;
<bibRefCitation id="EFC04B6B0B2EFFE0CE7CE6C2FA59FA0F" author="Chandramouli, S. R. &amp; Ganesh, S. R." box="[1084,1432,1447,1473]" pageId="18" pageNumber="351" pagination="72 - 85" refId="ref10808" refString="Chandramouli, S. R. &amp; Ganesh, S. R. (2010) Herpetofauna of the Southern Western Ghats - reinvestigated after decades. Taprobanica, 2 (2), 72 - 85. https: // doi. org / 10.47605 / tapro. v 2 i 2.30" type="journal article" year="2010">Chandramouli &amp; Ganesh 2010</bibRefCitation>
: fig. 16;
<bibRefCitation id="EFC04B6B0B2EFFE0CAADE6AEFE1CFA2B" author="Chaitanya, R. &amp; Khandekar, A. &amp; Caleb, D. &amp; Mukherjee, N. &amp; Ghosh, A. &amp; Giri, V. B." box="[237,477,1483,1509]" pageId="18" pageNumber="351" pagination="21 - 37" refId="ref10737" refString="Chaitanya, R., Khandekar, A., Caleb, D., Mukherjee, N., Ghosh, A. &amp; Giri, V. B. (2018) Herpetofauna of the Meghamalai Wildlife Sanctuary, Southern Western Ghats, India: An updated checklist with annotations on taxonomy and nomenclature. Journal of the Bombay Natural History Society, 115, 21 - 37." type="journal article" year="2018">
Chaitanya
<emphasis id="B925EA880B2EFFE0CB24E6AEFE5DFA2B" box="[356,412,1483,1509]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
2018
</bibRefCitation>
: fig. 3a), allow us to reasonably confidently identify this as
<taxonomicName id="4C514D190B2EFFE0CE30E6AEFAFFFA2B" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[1136,1342,1483,1509]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CE30E6AEFAFFFA2B" box="[1136,1342,1483,1509]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0CF04E6AEFA5DFA2B" bold="true" box="[1348,1436,1483,1509]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0CF04E6AEFA5DFA2B" box="[1348,1436,1483,1509]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The photograph of
<taxonomicName id="4C514D190B2EFFE0CB2EE68AFDD2F9C7" baseAuthorityName="Beddome" baseAuthorityYear="1863" box="[366,531,1519,1545]" class="Squamata" family="Uropeltidae" genus="Silybura" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="pulneyensis">
<emphasis id="B925EA880B2EFFE0CB2EE68AFDD2F9C7" box="[366,531,1519,1545]" italics="true" pageId="18" pageNumber="351">U. pulneyensis</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFC04B6B0B2EFFE0C8E6E68AFC77F9C7" author="Deepak, V. &amp; Narayanan, S. &amp; Das, S. &amp; Rajkumar, K. P. &amp; Pal, S. &amp; Gerard, J. D. &amp; Gower, D. J." box="[678,950,1518,1545]" pageId="18" pageNumber="351" refId="ref10961" refString="Deepak, V., Narayanan, S., Das, S., Rajkumar, K. P., Pal, S., Gerard, J. D. &amp; Gower, D. J. (2022) Snakes and lizards of the sky islands of the Western Ghats. Ashoka Trust for Research in Ecology and the Environment, Bengaluru, 114 pp." type="book" year="2022">
Deepak
<emphasis id="B925EA880B2EFFE0C942E68AFCFBF9C7" box="[770,826,1519,1545]" italics="true" pageId="18" pageNumber="351">et al.</emphasis>
(2022: 55)
</bibRefCitation>
and reproduced here in
<figureCitation id="136A2A1F0B2EFFE0CE84E68AFAD9F9C7" box="[1220,1304,1518,1545]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@8.[179,1408,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Uropeltis caudomaculata sp. nov. in life. A. Paratype BNHS 3765 from Manalar, Meghamalai, Tamil Nadu; photograph by JDG. B. Uncollected live individual from Periyar Tiger Reserve; photograph by SD. C. Paratype BNHS 3764 from Yellapetty; photograph by Sidharth S." figureDoi="http://doi.org/10.5281/zenodo.12732841" httpUri="https://zenodo.org/record/12732841/files/figure.png" pageId="18" pageNumber="351">Fig. 5A</figureCitation>
is of one of the
<typeStatus id="54EA88380B2EFFE0CA80E576FEEAF9E3" box="[192,299,1555,1581]" pageId="18" pageNumber="351" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C514D190B2EFFE0CB11E576FDE1F9E3" authorityName="Gower &amp; Das &amp; Deepak &amp; Gerard &amp; Narayanan" authorityYear="2024" box="[337,544,1555,1581]" class="Squamata" family="Uropeltidae" genus="Uropeltis" kingdom="Animalia" pageId="18" pageNumber="351" phylum="Chordata" rank="species" species="caudomaculata" status="sp. nov.">
<emphasis id="B925EA880B2EFFE0CB11E576FDE1F9E3" box="[337,544,1555,1581]" italics="true" pageId="18" pageNumber="351">U. caudomaculata</emphasis>
</taxonomicName>
<emphasis id="B925EA880B2EFFE0C867E576FD41F9E3" bold="true" box="[551,640,1555,1581]" pageId="18" pageNumber="351">
<taxonomicNameLabel id="A21657F30B2EFFE0C867E576FD41F9E3" box="[551,640,1555,1581]" pageId="18" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(BNHS 3765).
</paragraph>
</subSubSection>
</treatment>
</document>