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<document id="F0D682C6DA85DEBDA7148CB551D30068" ID-DOI="10.11646/zootaxa.3846.4.2" ID-GBIF-Dataset="ea136b1c-dc96-4c41-b078-01ca553453a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="252519" ID-ZooBank="71A80CC3-CDC1-4291-863B-FA7501D60AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467899762" checkinUser="plazi" docAuthor="Câmara, J. T. &amp; Rafael, J. A." docDate="2014" docId="03F6943FE050FFBAC7C7FABAAA6212BA" docLanguage="en" docName="zt03846p546.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opeatocerata tanimboca Câmara &amp; Rafael, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="541" masterDocId="FFCFEC47E075FF9DC750FFEAA8211077" masterDocTitle="Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)" masterLastPageNumber="546" masterPageNumber="502" pageNumber="539" updateTime="1698305230595" updateUser="plazi">
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<mods:title id="6CF976673755BDEEACBC9C45A5A8692B">Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)</mods:title>
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<mods:namePart id="349C4FE13CE14C4705610E2F46E7B93C">Câmara, J. T.</mods:namePart>
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<mods:namePart id="219B0063FB994DA98C8C2953AE009FAC">Rafael, J. A.</mods:namePart>
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<treatment id="03F6943FE050FFBAC7C7FABAAA6212BA" ID-DOI="http://doi.org/10.5281/zenodo.3502123" ID-GBIF-Taxon="119514720" ID-Zenodo-Dep="3502123" LSID="urn:lsid:plazi:treatment:03F6943FE050FFBAC7C7FABAAA6212BA" httpUri="http://treatment.plazi.org/id/03F6943FE050FFBAC7C7FABAAA6212BA" lastPageId="39" lastPageNumber="541" pageId="37" pageNumber="539">
<subSubSection id="C34576A2E050FFB8C7C7FABAA96215FB" pageId="37" pageNumber="539" type="nomenclature">
<paragraph id="8BE02529E050FFB8C7C7FABAAA03151D" blockId="37.[151,546,1360,1420]" box="[151,546,1360,1386]" pageId="37" pageNumber="539">
<heading id="D0A89245E050FFB8C7C7FABAAA03151D" bold="true" box="[151,546,1360,1386]" fontSize="11" level="1" pageId="37" pageNumber="539" reason="1">
<emphasis id="B92BF93BE050FFB8C7C7FABAAA03151D" bold="true" box="[151,546,1360,1386]" pageId="37" pageNumber="539">
<taxonomicName id="4C5F5EAAE050FFB8C7C7FABAA99F151D" ID-CoL="8D3LS" box="[151,446,1360,1386]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="539" phylum="Arthropoda" rank="species" species="tanimboca" status="sp. nov.">
<emphasis id="B92BF93BE050FFB8C7C7FABAA99F151D" bold="true" box="[151,446,1360,1386]" italics="true" pageId="37" pageNumber="539">Opeatocerata tanimboca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E050FFB8C695FABAAA03151D" box="[453,546,1360,1386]" pageId="37" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE02529E050FFB8C7C7FA99A96215FB" blockId="37.[151,546,1360,1420]" box="[151,323,1395,1420]" pageId="37" pageNumber="539">
(
<figureCitation id="136439ACE050FFB8C7CFFA99A91B15FB" box="[159,314,1395,1420]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Figs 185194</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E050FFB8C7C7FA51AC7E1580" pageId="37" pageNumber="539" type="diagnosis">
<paragraph id="8BE02529E050FFB8C7C7FA51AC7E1580" blockId="37.[151,1436,1467,2032]" pageId="37" pageNumber="539">
<emphasis id="B92BF93BE050FFB8C7C7FA51A92B15A3" bold="true" box="[151,266,1467,1492]" pageId="37" pageNumber="539">Diagnosis</emphasis>
. Tergites 36 with paramedian suboval black spots (
<figureCitation id="136439ACE050FFB8C424FA51AC3815A4" box="[884,1049,1467,1492]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Figs 185, 188</figureCitation>
); epandrium ventral lobe slightly bilobate apex (
<figureCitation id="136439ACE050FFB8C66CFA35A9BE1580" box="[316,415,1503,1528]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 189</figureCitation>
); phallus subcylindrical with dorsal subapical spine (
<figureCitation id="136439ACE050FFB8C4A0FA35AC731580" box="[1008,1106,1503,1528]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 193</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E050FFBAC797F9E9AC481137" lastPageId="39" lastPageNumber="541" pageId="37" pageNumber="539" type="description">
<paragraph id="8BE02529E050FFBAC797F9E9A953116A" blockId="37.[151,1436,1467,2032]" lastBlockId="39.[151,1437,151,717]" lastPageId="39" lastPageNumber="541" pageId="37" pageNumber="539">
<emphasis id="B92BF93BE050FFB8C797F9E9A970166B" bold="true" box="[199,337,1539,1564]" pageId="37" pageNumber="539">Description</emphasis>
.
<emphasis id="B92BF93BE050FFB8C60FF9E9A9BA166B" bold="true" box="[351,411,1539,1564]" pageId="37" pageNumber="539">Male</emphasis>
(
<figureCitation id="136439ACE050FFB8C6FBF9E9AA2C166B" box="[427,525,1539,1564]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 185</figureCitation>
).
<emphasis id="B92BF93BE050FFB8C570F9E9AA41166B" bold="true" box="[544,608,1539,1564]" pageId="37" pageNumber="539">Head</emphasis>
: Face parallel-sided, dark brown with gray pruinescence visible in ventral view, about
<date id="FFE103E9E050FFB8C672F9CDA9641648" box="[290,325,1575,1599]" pageId="37" pageNumber="539">4X</date>
longer than lower width. Ocellar tubercle protuberant, dark brown with gray pruinescence; ocelli brown. Scape, pedicel and postpedicel brown with black bristles; postpedicel about 1.5X longer than pedicel; stylus about
<date id="FFE103E9E050FFB8C78FF985A92316F0" box="[223,258,1647,1671]" pageId="37" pageNumber="539">3X</date>
longer than postpedicel. Proboscis dark yellow, shorter than head height.
<emphasis id="B92BF93BE050FFB8C331F985AC9B16FF" bold="true" box="[1121,1210,1647,1672]" pageId="37" pageNumber="539">Thorax</emphasis>
yellow, shiny (
<figureCitation id="136439ACE050FFB8C23EF985A8E416DB" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 185</figureCitation>
). Legs yellow, except for all tarsomeres 45 black; mid, hind tibia and hind tarsomere 1 with anterodorsal, posterodorsal, anteroventral and posteroventral rows of longer bristles.
<emphasis id="B92BF93BE050FFB8C48DF95DAC3F16A7" bold="true" box="[989,1054,1719,1744]" pageId="37" pageNumber="539">Wing</emphasis>
(
<figureCitation id="136439ACE050FFB8C363F95DACB616A7" box="[1075,1175,1719,1744]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 186</figureCitation>
) hyaline, pterostigma, brown, conspicuous, about 2.7X longer than wide. Halter light yellow.
<emphasis id="B92BF93BE050FFB8C4ECF931AC111683" bold="true" box="[956,1072,1755,1780]" pageId="37" pageNumber="539">Abdomen</emphasis>
dark yellow, shiny; tergites 36 with paramedian suboval black spots (
<figureCitation id="136439ACE050FFB8C504F915AADB1760" box="[596,762,1791,1816]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Figs 185, 188</figureCitation>
); tergite 8 divided in two subrectangular plates; sternite 8 divided in two subtriangular plates.
<emphasis id="B92BF93BE050FFB8C562F8C9AA94174B" bold="true" box="[562,693,1827,1852]" pageId="37" pageNumber="539">Terminalia</emphasis>
: Anterior cercus with proclinate dorsal projection in lateral view (
<figureCitation id="136439ACE050FFB8C7CFF8ADA9271728" box="[159,262,1863,1888]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Figs 187</figureCitation>
, 1189), posterior margin truncate and with a median sinus in dorsal view (
<figureCitation id="136439ACE050FFB8C335F8ADACE81728" box="[1125,1225,1863,1888]" captionStart="FIGURE 211" captionStartId="42.[151,250,1955,1977]" captionTargetBox="[235,1368,193,1934]" captionTargetId="figure@42.[216,1371,193,1934]" captionTargetPageId="42" captionText="FIGURE 211. Geographic distribution of species of Opeatocerata." httpUri="https://zenodo.org/record/252546/files/figure.png" pageId="37" pageNumber="539">Fig. 211</figureCitation>
); descendant plate subtrapezoidal in posterior view (
<figureCitation id="136439ACE050FFB8C546F881AA5817F4" box="[534,633,1899,1924]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 191</figureCitation>
). Posterior cercus with a rounded apex in lateral view (
<figureCitation id="136439ACE050FFB8C3A4F881ADB517F4" box="[1268,1428,1899,1924]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Figs 187, 188</figureCitation>
) and with a dorsal submedian subtriangular projection in dorsal view (
<figureCitation id="136439ACE050FFB8C4E2F865AC3417D0" box="[946,1045,1935,1960]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 191</figureCitation>
). Hypoproct subrounded in lateral view (
<figureCitation id="136439ACE050FFB8C78CF859A91B17BC" box="[220,314,1971,1996]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 189</figureCitation>
) and comma-shaped in posterior view (
<figureCitation id="136439ACE050FFB8C5A3F859AB7017BC" box="[755,849,1971,1996]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 191</figureCitation>
). Epandrium ventral lobe with slightly bilobate apex (
<figureCitation id="136439ACE050FFB8C7CFF83DA9231798" box="[159,258,2007,2032]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 189</figureCitation>
); without spiniform bristles, longer bristles apically on ventral lobe (
<figureCitation id="136439ACE050FFB8C340F83DAC521798" box="[1040,1139,2007,2032]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="37" pageNumber="539">Fig. 189</figureCitation>
). Hypandrium wider than long, with concave apex (
<figureCitation id="136439ACE052FFBAC692FF7DAA0710C7" box="[450,550,151,176]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 194</figureCitation>
), without long bristles. Phallus subcylindrical, longer than hypandrium, with dorsal subapical spine (
<figureCitation id="136439ACE052FFBAC6F1FF56AA2510A3" box="[417,516,188,213]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 193</figureCitation>
). Subepandrial sclerite wider than long, U-shaped (
<figureCitation id="136439ACE052FFBAC31AFF56AC8C10A3" box="[1098,1197,188,213]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 192</figureCitation>
). Bacilliform sclerite with folds and longer than subepandrial sclerite (
<figureCitation id="136439ACE052FFBAC585FF35AB1A108F" box="[725,827,223,248]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 192</figureCitation>
).
<emphasis id="B92BF93BE052FFBAC402FF35AC3A108F" bold="true" box="[850,1051,223,248]" pageId="39" pageNumber="541">Specimen length</emphasis>
:
<quantity id="4CA788CCE052FFBAC37EFF0AACAA1080" box="[1070,1163,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="39" pageNumber="541" unit="mm" value="3.2">3.2 mm</quantity>
;
<emphasis id="B92BF93BE052FFBAC3CAFF35AD08108F" bold="true" box="[1178,1321,223,248]" pageId="39" pageNumber="541">wing length</emphasis>
:
<quantity id="4CA788CCE052FFBAC26BFF0AADB91080" box="[1339,1432,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="39" pageNumber="541" unit="mm" value="3.3">3.3 mm</quantity>
.
<emphasis id="B92BF93BE052FFBAC7C7FEEEA8CF116A" bold="true" box="[151,238,260,285]" pageId="39" pageNumber="541">Female</emphasis>
. Unknown.
</paragraph>
<caption id="DF2075A1E053FFBBC7C7F86BACCC1784" httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="38" pageNumber="540" targetBox="[157,1431,193,1908]" targetPageId="38">
<paragraph id="8BE02529E053FFBBC7C7F86BACCC1784" blockId="38.[151,1436,1921,2035]" pageId="38" pageNumber="540">
<emphasis id="B92BF93BE053FFBBC7C7F86BA94617E1" bold="true" box="[151,359,1921,1944]" pageId="38" pageNumber="540">FIGURES 185194</emphasis>
.
<taxonomicName id="4C5F5EAAE053FFBBC624F868AA5017E0" box="[372,625,1921,1943]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="540" phylum="Arthropoda" rank="species" species="tanimboca" status="sp. nov.">
<emphasis id="B92BF93BE053FFBBC624F868AA5017E0" box="[372,625,1921,1943]" italics="true" pageId="38" pageNumber="540">Opeatocerata tanimboca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E053FFBBC528F868AAE217E0" box="[632,707,1922,1943]" pageId="38" pageNumber="540" rank="species">
<emphasis id="B92BF93BE053FFBBC528F868AAE217E0" bold="true" box="[632,707,1922,1943]" pageId="38" pageNumber="540">sp.nov.</emphasis>
</taxonomicNameLabel>
, holotype ♂.
<emphasis id="B92BF93BE053FFBBC41FF86BAB5417E1" bold="true" box="[847,885,1921,1942]" pageId="38" pageNumber="540">185</emphasis>
.
<emphasis id="B92BF93BE053FFBBC4D3F86BABEF17E0" box="[899,974,1921,1943]" italics="true" pageId="38" pageNumber="540">habitus</emphasis>
, lateral view;
<emphasis id="B92BF93BE053FFBBC330F86BACA717E1" bold="true" box="[1120,1158,1921,1942]" pageId="38" pageNumber="540">186</emphasis>
. wing;
<emphasis id="B92BF93BE053FFBBC384F86BACDB17E1" bold="true" box="[1236,1274,1921,1942]" pageId="38" pageNumber="540">187</emphasis>
. abdomen from tergite 6 to terminalia, lateral view;
<emphasis id="B92BF93BE053FFBBC6AEF84AAA0517C2" bold="true" box="[510,548,1952,1973]" pageId="38" pageNumber="540">188</emphasis>
. tergites 18, dorsal view;
<emphasis id="B92BF93BE053FFBBC463F84AAB7817C2" bold="true" box="[819,857,1952,1973]" pageId="38" pageNumber="540">189</emphasis>
. terminalia, lateral view;
<emphasis id="B92BF93BE053FFBBC30BF84AACA017C2" bold="true" box="[1115,1153,1952,1973]" pageId="38" pageNumber="540">190</emphasis>
. lobes of cerci, dorsal view;
<emphasis id="B92BF93BE053FFBBC7C7F855A89C17A3" bold="true" box="[151,189,1983,2004]" pageId="38" pageNumber="540">191</emphasis>
. descendant plate of anterior cercus and hypoproct, posterior view;
<emphasis id="B92BF93BE053FFBBC4DFF855AB9417A3" bold="true" box="[911,949,1983,2004]" pageId="38" pageNumber="540">192</emphasis>
. subepandrial sclerite and bacilliform sclerite, dorsal view;
<emphasis id="B92BF93BE053FFBBC64AF837A9611785" bold="true" box="[282,320,2013,2034]" pageId="38" pageNumber="540">193</emphasis>
. ejaculatory apodeme, phallus and hypandrium, lateral view;
<emphasis id="B92BF93BE053FFBBC4E0F837ABF71785" bold="true" box="[944,982,2013,2034]" pageId="38" pageNumber="540">194</emphasis>
. hypandrium, ventral view.
</paragraph>
</caption>
<paragraph id="8BE02529E052FFBAC797FECDAC481137" blockId="39.[151,1437,151,717]" box="[199,1129,295,320]" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC797FECDA9DC1137" bold="true" box="[199,509,295,320]" pageId="39" pageNumber="541">Geographical distribution</emphasis>
.
<collectingCountry id="F34865B9E052FFBAC55BFECDAA581137" box="[523,633,295,320]" name="Colombia" pageId="39" pageNumber="541">Colombia</collectingCountry>
(Leticia) and
<collectingCountry id="F34865B9E052FFBAC447FECDAB7A1137" box="[791,859,295,320]" name="Brazil" pageId="39" pageNumber="541">Brazil</collectingCountry>
(Amazonas) (
<figureCitation id="136439ACE052FFBAC4ABFECDAC7D1137" box="[1019,1116,295,320]" captionStart="FIGURE 211" captionStartId="42.[151,250,1955,1977]" captionTargetBox="[235,1368,193,1934]" captionTargetId="figure@42.[216,1371,193,1934]" captionTargetPageId="42" captionText="FIGURE 211. Geographic distribution of species of Opeatocerata." httpUri="https://zenodo.org/record/252546/files/figure.png" pageId="39" pageNumber="541">Fig. 211</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E052FFBAC797FEA6AD0011A7" pageId="39" pageNumber="541" type="materials_examined">
<paragraph id="8BE02529E052FFBAC797FEA6ACAD11DB" blockId="39.[151,1437,151,717]" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC797FEA6A94F1112" bold="true" box="[199,366,332,357]" pageId="39" pageNumber="541">
<typeStatus id="54E49B8BE052FFBAC797FEA6A9231112" box="[199,258,332,357]" pageId="39" pageNumber="541">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54E49B8BE052FFBAC62CFEA6AA3A1112" box="[380,539,332,357]" pageId="39" pageNumber="541" type="holotype">
<emphasis id="B92BF93BE052FFBAC62CFEA6AA3A1112" bold="true" box="[380,539,332,357]" pageId="39" pageNumber="541">HOLOTYPE</emphasis>
</typeStatus>
♂, labelled:
<collectingCountry id="F34865B9E052FFBAC5FEFEA6AB3E1112" box="[686,799,332,357]" name="Colombia" pageId="39" pageNumber="541">Colombia</collectingCountry>
, AMZ [Amazonas], Letícia. Reserva
<taxonomicName id="4C5F5EAAE052FFBAC395FEA6AD651112" box="[1221,1348,332,357]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="541" phylum="Arthropoda" rank="species" species="tanimboca" status="sp. nov.">Tanimboca</taxonomicName>
, Km 11 via Tarapacá.
<geoCoordinate id="EE6B43EEE052FFBAC662FE85A95B11FF" box="[306,378,367,392]" direction="south" orientation="latitude" pageId="39" pageNumber="541" precision="925" value="-4.2166667">4º13S</geoCoordinate>
;
<geoCoordinate id="EE6B43EEE052FFBAC62FFE85A9CB11FF" box="[383,490,367,392]" direction="west" orientation="longitude" pageId="39" pageNumber="541" precision="925" value="-69.933334">69º56W</geoCoordinate>
.
<date id="FFE103E9E052FFBAC6A5FE85AAB611FF" box="[501,663,367,392]" pageId="39" pageNumber="541" value="2011-04-24" valueMax="2011-04-29" valueMin="2011-04-24">2429.iv.2011</date>
. Arm[adilha] Luz. Antonio Agudelo
<emphasis id="B92BF93BE052FFBAC364FE85AC7F11FF" box="[1076,1118,367,392]" italics="true" pageId="39" pageNumber="541">leg.</emphasis>
(
<collectionCode id="ED4EBDECE052FFBAC33CFE9AAC9F11FF" box="[1132,1214,368,392]" pageId="39" pageNumber="541">CEUA</collectionCode>
).
<typeStatus id="54E49B8BE052FFBAC381FE85AD1F11FF" box="[1233,1342,367,392]" pageId="39" pageNumber="541" type="paratype">
<emphasis id="B92BF93BE052FFBAC381FE85AD1F11FF" bold="true" box="[1233,1342,367,392]" pageId="39" pageNumber="541">Paratype</emphasis>
</typeStatus>
:
<collectingCountry id="F34865B9E052FFBAC21CFE85ADB411FF" box="[1356,1429,367,392]" name="Brazil" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC21CFE85ADB411FF" bold="true" box="[1356,1429,367,392]" pageId="39" pageNumber="541">Brazil</emphasis>
</collectingCountry>
,
<emphasis id="B92BF93BE052FFBAC7C7FE7EA93211DA" bold="true" box="[151,275,404,429]" pageId="39" pageNumber="541">Amazonas</emphasis>
. Tabatinga,
<date id="FFE103E9E052FFBAC6CDFE7EAA1211DA" box="[413,563,404,429]" pageId="39" pageNumber="541" value="1992-01-13" valueMax="1992-01-17" valueMin="1992-01-13">1317.i.1992</date>
, Arm[adilha] Malaise, J. Vidal &amp; Lílian (
<specimenCount id="9D59EEA0E052FFBAC355FE7EAC1111DB" box="[1029,1072,404,428]" pageId="39" pageNumber="541" type="generic">1 ♂</specimenCount>
,
<collectionCode id="ED4EBDECE052FFBAC36BFE7EACA111DB" box="[1083,1152,404,428]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="39" pageNumber="541">INPA</collectionCode>
).
</paragraph>
<paragraph id="8BE02529E052FFBAC797FE5DAD0011A7" blockId="39.[151,1437,151,717]" box="[199,1313,439,464]" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC797FE5DA99311A7" bold="true" box="[199,434,439,464]" pageId="39" pageNumber="541">
<typeStatus id="54E49B8BE052FFBAC797FE5DA91511A7" box="[199,308,439,464]" pageId="39" pageNumber="541" type="holotype">Holotype</typeStatus>
condition.
</emphasis>
Good; left wing mounted on microslide; abdomen in microtube with glycerin.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E052FFBAC797FE36AAB9126F" pageId="39" pageNumber="541" type="etymology">
<paragraph id="8BE02529E052FFBAC797FE36AAB9126F" blockId="39.[151,1437,151,717]" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC797FE36A9671182" bold="true" box="[199,326,476,501]" pageId="39" pageNumber="541">Etymology</emphasis>
. The specific epithet is a noun in apposition and refers to Reserva
<taxonomicName id="4C5F5EAAE052FFBAC36FFE36AC9F1182" box="[1087,1214,476,501]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="541" phylum="Arthropoda" rank="species" species="tanimboca" status="sp. nov.">Tanimboca</taxonomicName>
, Leticia,
<collectingCountry id="F34865B9E052FFBAC277FE36ADB91182" box="[1319,1432,476,501]" name="Colombia" pageId="39" pageNumber="541">Colombia</collectingCountry>
, the locality where the
<typeStatus id="54E49B8BE052FFBAC6C0FE15A9D3126F" box="[400,498,511,536]" pageId="39" pageNumber="541" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E052FFBAC797FDCEAA6212BA" pageId="39" pageNumber="541" type="discussion">
<paragraph id="8BE02529E052FFBAC797FDCEAA6212BA" blockId="39.[151,1437,151,717]" pageId="39" pageNumber="541">
<emphasis id="B92BF93BE052FFBAC797FDCEA915124A" bold="true" box="[199,308,548,573]" pageId="39" pageNumber="541">Remarks</emphasis>
.
<taxonomicName id="4C5F5EAAE052FFBAC611FDCFAA79124A" box="[321,600,548,573]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="541" phylum="Arthropoda" rank="species" species="tanimboca" status="sp. nov.">
<emphasis id="B92BF93BE052FFBAC611FDCFAA79124A" box="[321,600,548,573]" italics="true" pageId="39" pageNumber="541">Opeatocerata tanimboca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E052FFBAC50FFDCEAA98124A" box="[607,697,548,573]" pageId="39" pageNumber="541" rank="species">
<emphasis id="B92BF93BE052FFBAC50FFDCEAA98124A" bold="true" box="[607,697,548,573]" pageId="39" pageNumber="541">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from other species in the tergites with paramedian suboval spots (
<figureCitation id="136439ACE052FFBAC7B5FDADA9681217" box="[229,329,583,608]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 188</figureCitation>
) (in other species it is subtriangular or subrectangular), anterior cercus with descendant plate subtrapezoidal (
<figureCitation id="136439ACE052FFBAC61EFD86A99312F3" box="[334,434,620,645]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 191</figureCitation>
) (in other species this plate is either subtriangular, subrectangular, with lateroventral expansions or hourglass-shaped) and phallus with dorsal subapical spine (
<figureCitation id="136439ACE052FFBAC4B8FD65AC6B12DF" box="[1000,1098,655,680]" captionStart="FIGURES 185 194" captionStartId="38.[151,264,1921,1944]" captionTargetBox="[157,1431,193,1908]" captionTargetId="figure@38.[155,1432,193,1918]" captionTargetPageId="38" captionText="FIGURES 185 194. Opeatocerata tanimboca sp. nov., holotype ♂. 185. habitus, lateral view; 186. wing; 187. abdomen from tergite 6 to terminalia, lateral view; 188. tergites 1 8, dorsal view; 189. terminalia, lateral view; 190. lobes of cerci, dorsal view; 191. descendant plate of anterior cercus and hypoproct, posterior view; 192. subepandrial sclerite and bacilliform sclerite, dorsal view; 193. ejaculatory apodeme, phallus and hypandrium, lateral view; 194. hypandrium, ventral view." httpUri="https://zenodo.org/record/252543/files/figure.png" pageId="39" pageNumber="541">Fig. 193</figureCitation>
) (in other species there is no dorsal subapical spine on the phallus).
</paragraph>
</subSubSection>
</treatment>
</document>