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<document id="CE694CE1CD6F7D518EFAC85664F48FBD" ID-DOI="10.11646/zootaxa.5228.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7532249" ID-ZooBank="E3FA5ED0-A556-41BA-81D9-4FF1CAED6117" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673571303360" checkinUser="plazi" docAuthor="Nearns, Eugenio H., Swift, Ian P. &amp; Santos-Silva, Antonio" docDate="2023" docId="03F6827C2801FF97FF0415FEFCCFFC97" docLanguage="en" docName="zootaxa.5228.2.3.pdf" docOrigin="Zootaxa 5228 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Methia enigma Martins 1981" docType="treatment" docVersion="3" lastPageNumber="140" masterDocId="FFCFFA042800FF94FF93107CFFDFFFCA" masterDocTitle="Three new species of longhorned beetles (Coleoptera: Cerambycidae) with notes on additional taxa" masterLastPageNumber="156" masterPageNumber="137" pageNumber="138" updateTime="1699446576877" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="72C21A169ABD5860F9F82761CB0C2C42">Three new species of longhorned beetles (Coleoptera: Cerambycidae) with notes on additional taxa</mods:title>
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<mods:namePart id="5CC23A966471A3C19E3050E96AB7C87E">Nearns, Eugenio H.</mods:namePart>
<mods:affiliation id="B48F6CA02D5F6709AA656D28FE90E63D">National Museum of Natural History, Smithsonian Institution, Washington, D. C. 20560, U. S. A.</mods:affiliation>
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<mods:namePart id="BAD262338FF65556FBBD72D9B9431D0F">Swift, Ian P.</mods:namePart>
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<mods:namePart id="E81C23A3EB1EDE5EF8A009E3807B32A3">Santos-Silva, Antonio</mods:namePart>
<mods:affiliation id="4AC4E0E701D5D939E5A1C8A050D10D69">Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil.</mods:affiliation>
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<taxonomicName id="4C5F48E92801FF95FF0415FEFDDAFA56" ID-CoL="6RFJW" authority="Martins, 1981" authorityName="Martins" authorityYear="1981" box="[151,517,1410,1437]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FF0415FEFDDAFA56" bold="true" box="[151,517,1410,1437]" pageId="1" pageNumber="138">
<emphasis id="B92BEF782801FF95FF0415FEFE90FA57" bold="true" box="[151,335,1410,1437]" italics="true" pageId="1" pageNumber="138">Methia enigma</emphasis>
<bibRefCitation id="EFCE4E9B2801FF95FEC515FEFDDAFA56" author="Martins, U. R." box="[342,517,1410,1436]" pageId="1" pageNumber="138" pagination="9 - 17" refId="ref11439" refString="Martins, U. R. (1981) Revis &quot; o do genero Atenizus; os generos Tessaropa e Methia na America do Sul e descric &quot; o de novos taxa em Methiini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 25 (1), 9 - 17." type="journal article" year="1981">Martins, 1981</bibRefCitation>
</emphasis>
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<paragraph id="8BE0336A2801FF95FF0415DBFEC3FA0B" blockId="1.[151,517,1410,1473]" box="[151,284,1447,1473]" pageId="1" pageNumber="138">
<figureCitation id="13642FEF2801FF95FF0415DBFEC3FA0B" box="[151,284,1447,1473]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[192,1396,181,1919]" captionTargetId="figure-21@2.[192,1396,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–7. Methia spp. 1–2) Methia enigma Martins, 1981, paratype male: 1) Dorsal habitus; 2) Eye. 3–4) Methia argentina Bruch, 1918, male from Argentina (Catamarca): 3) Dorsal habitus; 4) Eye. 5) Methia argentina, female from Paraguay (Boquerón): Dorsal habitus; 6) Ventral habitus; 7) Eye." figureDoi="http://doi.org/10.5281/zenodo.7532251" httpUri="https://zenodo.org/record/7532251/files/figure.png" pageId="1" pageNumber="138">Figures 1–2</figureCitation>
</paragraph>
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<subSubSection id="C34560E12801FF95FF041597FAA1F9E8" pageId="1" pageNumber="138" type="reference_group">
<paragraph id="8BE0336A2801FF95FF041597FAA1F9E8" blockId="1.[151,1437,1515,1570]" pageId="1" pageNumber="138">
<treatmentCitationGroup id="AB4F14442801FF95FF041597FAA1F9E8" pageId="1" pageNumber="138">
<treatmentCitation id="0AFE157B2801FF95FF041597FE2FF9C9" author="Martins, U. R." box="[151,496,1515,1539]" page="12" pageId="1" pageNumber="138" year="1981">
<taxonomicName id="4C5F48E92801FF95FF041597FE2FF9C9" ID-CoL="6RFJW" authority="Martins, 1981: 12" authorityName="Martins" authorityPageNumber="12" authorityYear="1981" box="[151,496,1515,1539]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FF041597FEEEF9C9" box="[151,305,1515,1539]" italics="true" pageId="1" pageNumber="138">Methia enigma</emphasis>
<bibRefCitation id="EFCE4E9B2801FF95FEA41597FE2FF9C9" author="Martins, U. R." box="[311,496,1515,1539]" pageId="1" pageNumber="138" pagination="9 - 17" refId="ref11439" refString="Martins, U. R. (1981) Revis &quot; o do genero Atenizus; os generos Tessaropa e Methia na America do Sul e descric &quot; o de novos taxa em Methiini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 25 (1), 9 - 17." type="journal article" year="1981">Martins, 1981: 12</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AFE157B2801FF95FE681597FD75F9C9" author="Monne, M. A." box="[507,682,1515,1539]" page="25" pageId="1" pageNumber="138" year="1993">
<bibRefCitation id="EFCE4E9B2801FF95FE681597FD75F9C9" author="Monne, M. A." box="[507,682,1515,1539]" pageId="1" pageNumber="138" refId="ref11800" refString="Monne, M. A. (1993) Catalogue of the Cerambycidae (Coleoptera) of the Western Hemisphere. Part I. Subfamily Cerambycinae: Tribes Erlandiini, Smodicini, Oemini, Methiini, Xystrocerini, Dodecosini, Opsimini, Achrysonini and Pleiarthrocerini. Sociedade Brasileira de Entomologia, S &quot; o Paulo, 76 pp." type="book" year="1993">Monné, 1993: 25</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0AFE157B2801FF95FD671597FBC5F9C9" author="Monne, M. A. &amp; Giesbert, E. F." box="[756,1050,1515,1539]" page="30" pageId="1" pageNumber="138" year="1994">
<bibRefCitation id="EFCE4E9B2801FF95FD671597FBC5F9C9" author="Monne, M. A. &amp; Giesbert, E. F." box="[756,1050,1515,1539]" pageId="1" pageNumber="138" refId="ref12145" refString="Monne, M. A. &amp; Giesbert, E. F. (1994) Checklist of the Cerambycidae and Disteniidae (Coleoptera) of the Western Hemisphere. Wolfsgarden Books, Burbank, 409 pp." type="book" year="1994">Monné &amp; Giesbert, 1994: 30</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0AFE157B2801FF95FB091597FF2FF9E8" author="Martins, U. R. &amp; Galileo, M. H. M." page="172" pageId="1" pageNumber="138" year="1997">
<bibRefCitation id="EFCE4E9B2801FF95FB091597FF2FF9E8" author="Martins, U. R. &amp; Galileo, M. H. M." pageId="1" pageNumber="138" refId="ref11493" refString="Martins, U. R. &amp; Galileo, M. H. M. (1997) Tribo Methiini. In: Martins, U. R (org.), Cerambycidae Sul-Americanos (Coleoptera). Vol. 1. Sociedade Brasileira de Entomologia, S &quot; o Paulo, pp. 159 - 179." type="book" year="1997">Martins &amp; Galileo, 1997: 172</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFE157B2801FF95FF681676FE6EF9E8" author="Di Iorio, O." box="[251,433,1546,1570]" page="18" pageId="1" pageNumber="138" year="2004">
<bibRefCitation id="EFCE4E9B2801FF95FF681676FE6EF9E8" author="Di Iorio, O." box="[251,433,1546,1570]" pageId="1" pageNumber="138" pagination="9 - 65" refId="ref10917" refString="Di Iorio, O. (2004) Aporte al catalogo de Cerambycidae del Paraguay (Insecta, Coleoptera). Parte IV. Addenda a Bosq (Partes I y II) y Viana (Parte III). Boletin del Museo Nacional de Historia Natural del Paraguay, 15 (1 - 2), 9 - 65." type="journal article" year="2004">Di Iorio, 2004: 18</bibRefCitation>
</treatmentCitation>
(distr.);
<treatmentCitation id="0AFE157B2801FF95FD9B1676FCF5F9E8" author="Martins, U. R. &amp; Galileo, M. H. M." box="[520,810,1546,1570]" page="12" pageId="1" pageNumber="138" year="2004">
<bibRefCitation id="EFCE4E9B2801FF95FD9B1676FCF5F9E8" author="Martins, U. R. &amp; Galileo, M. H. M." box="[520,810,1546,1570]" pageId="1" pageNumber="138" refId="ref11552" refString="Martins, U. R. &amp; Galileo, M. H. M. (2004) Cerambycidae sul-americanos. Suplemento 1. Argania, Barcelona, 147 pp." type="book" year="2004">Martins &amp; Galileo, 2004: 12</bibRefCitation>
</treatmentCitation>
, 102;
<treatmentCitation id="0AFE157B2801FF95FCFB1676FBF1F9E8" author="Monne, M. A." box="[872,1070,1546,1570]" page="395" pageId="1" pageNumber="138" year="2005">
<bibRefCitation id="EFCE4E9B2801FF95FCFB1676FBF1F9E8" author="Monne, M. A." box="[872,1070,1546,1570]" pageId="1" pageNumber="138" pagination="1 - 765" refId="ref11951" refString="Monne, M. A. (2005 a) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical Region. Part I. Subfamily Cerambycinae. Zootaxa, 946 (1), 1 - 765. https: // doi. org / 10.11646 / zootaxa. 946.1.1" type="journal article" year="2005">Monné, 2005a: 395</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="0AFE157B2801FF95FBE51676FAE3F9E8" author="Monne, M. A." box="[1142,1340,1546,1570]" page="475" pageId="1" pageNumber="138" year="2022">
<bibRefCitation id="EFCE4E9B2801FF95FBE51676FAE3F9E8" author="Monne, M. A." box="[1142,1340,1546,1570]" pageId="1" pageNumber="138" refId="ref12053" refString="Monne, M. A. (2022 a) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical region. Part II. Subfamily Cerambycinae. Available from: https: // cerambycids. com / catalog / (accessed 27 September 2022)" type="url" year="2022">Monné, 2022a: 475</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34560E12801FF97FF04162CFD15FD63" lastPageId="3" lastPageNumber="140" pageId="1" pageNumber="138" type="discussion">
<paragraph id="8BE0336A2801FF95FF04162CFE6EF8D4" blockId="1.[151,1437,1616,2039]" pageId="1" pageNumber="138">
<emphasis id="B92BEF782801FF95FF04162CFED4F9A0" bold="true" box="[151,267,1616,1642]" pageId="1" pageNumber="138">Remarks.</emphasis>
According to
<bibRefCitation id="EFCE4E9B2801FF95FE23162CFD86F9A0" author="Martins, U. R." box="[432,601,1616,1642]" pageId="1" pageNumber="138" pagination="9 - 17" refId="ref11439" refString="Martins, U. R. (1981) Revis &quot; o do genero Atenizus; os generos Tessaropa e Methia na America do Sul e descric &quot; o de novos taxa em Methiini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 25 (1), 9 - 17." type="journal article" year="1981">Martins (1981)</bibRefCitation>
(translated): “This species [
<taxonomicName id="4C5F48E92801FF95FC09162CFB9CF9A0" authorityName="Martins" authorityYear="1981" box="[922,1091,1616,1642]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FC09162CFB9CF9A0" box="[922,1091,1616,1642]" italics="true" pageId="1" pageNumber="138">Methia enigma</emphasis>
</taxonomicName>
] is very close to
<taxonomicName id="4C5F48E92801FF95FA99162DFA43F9A0" authorityName="Bruch" authorityYear="1918" box="[1290,1436,1617,1642]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782801FF95FA99162DFA43F9A0" box="[1290,1436,1617,1642]" italics="true" pageId="1" pageNumber="138">M. argentina</emphasis>
</taxonomicName>
[
<figureCitation id="13642FEF2801FF95FF0D1608FED3F944" box="[158,268,1652,1678]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[192,1396,181,1919]" captionTargetId="figure-21@2.[192,1396,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–7. Methia spp. 1–2) Methia enigma Martins, 1981, paratype male: 1) Dorsal habitus; 2) Eye. 3–4) Methia argentina Bruch, 1918, male from Argentina (Catamarca): 3) Dorsal habitus; 4) Eye. 5) Methia argentina, female from Paraguay (Boquerón): Dorsal habitus; 6) Ventral habitus; 7) Eye." figureDoi="http://doi.org/10.5281/zenodo.7532251" httpUri="https://zenodo.org/record/7532251/files/figure.png" pageId="1" pageNumber="138">Figs. 3–7</figureCitation>
] and
<taxonomicName id="4C5F48E92801FF95FEDE1609FDDDF944" baseAuthorityName="Gounelle" baseAuthorityYear="1913" box="[333,514,1652,1678]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="tubuliventris">
<emphasis id="B92BEF782801FF95FEDE1609FDDDF944" box="[333,514,1652,1678]" italics="true" pageId="1" pageNumber="138">M. tubuliventris</emphasis>
</taxonomicName>
due to the cuneiform appearance of the elytra. It differs from both by presenting the upper eye lobes contiguous.” This feature was used in Martin’s key to species of the genus (translated): “1. Upper eye lobes contiguous,” leading to
<taxonomicName id="4C5F48E92801FF95FDD716C1FD63F91C" authorityName="Martins" authorityYear="1981" box="[580,700,1725,1750]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FDD716C1FD63F91C" box="[580,700,1725,1750]" italics="true" pageId="1" pageNumber="138">M. enigma</emphasis>
</taxonomicName>
; “Upper eye lobes very close but not in contiguous (the distance between them is subequal to the diameter of two ommatidia)”, leading to
<taxonomicName id="4C5F48E92801FF95FC41169DFB2AF930" authority="Bruch, 1918" authorityName="Bruch" authorityYear="1918" box="[978,1269,1760,1786]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782801FF95FC41169DFBBCF930" box="[978,1123,1761,1786]" italics="true" pageId="1" pageNumber="138">M. argentina</emphasis>
Bruch, 1918
</taxonomicName>
and
<taxonomicName id="4C5F48E92801FF95FABF169DFE72F8D4" authority="(Gounelle, 1913)" baseAuthorityName="Gounelle" baseAuthorityYear="1913" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="tubuliventris">
<emphasis id="B92BEF782801FF95FABF169DFF3CF8D4" italics="true" pageId="1" pageNumber="138">M. tubuliventris</emphasis>
(Gounelle, 1913)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE0336A2801FF97FF541754FB6BFF33" blockId="1.[151,1437,1616,2039]" lastBlockId="3.[151,1437,151,861]" lastPageId="3" lastPageNumber="140" pageId="1" pageNumber="138">
<bibRefCitation id="EFCE4E9B2801FF95FF541754FE37F888" author="Martins, U. R. &amp; Galileo, M. H. M." box="[199,488,1832,1858]" pageId="1" pageNumber="138" refId="ref11493" refString="Martins, U. R. &amp; Galileo, M. H. M. (1997) Tribo Methiini. In: Martins, U. R (org.), Cerambycidae Sul-Americanos (Coleoptera). Vol. 1. Sociedade Brasileira de Entomologia, S &quot; o Paulo, pp. 159 - 179." type="book" year="1997">Martins &amp; Galileo (1997)</bibRefCitation>
reported (translated): “3(2). Upper eye lobes contiguous; elytra brownish, unicolorous or slightly more yellowish basally,” leading to
<taxonomicName id="4C5F48E92801FF95FD461731FC92F8AC" authorityName="Martins" authorityYear="1981" box="[725,845,1869,1894]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FD461731FC92F8AC" box="[725,845,1869,1894]" italics="true" pageId="1" pageNumber="138">M. enigma</emphasis>
</taxonomicName>
; Upper eye lobes separated by a distance equivalent to a row of ommatidia; elytra yellowish with brownish apical third,” leading to
<taxonomicName id="4C5F48E92801FF95FB85170DFB77F840" authorityName="Bruch" authorityYear="1918" box="[1046,1192,1905,1930]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782801FF95FB85170DFB77F840" box="[1046,1192,1905,1930]" italics="true" pageId="1" pageNumber="138">M. argentina</emphasis>
</taxonomicName>
; and “
<taxonomicName id="4C5F48E92801FF95FB60170CFA43F840" authorityName="Martins" authorityYear="1981" box="[1267,1436,1904,1930]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FB60170CFA43F840" box="[1267,1436,1904,1930]" italics="true" pageId="1" pageNumber="138">Methia enigma</emphasis>
</taxonomicName>
resembles the males of
<taxonomicName id="4C5F48E92801FF95FE0917E9FDF5F864" authorityName="Bruch" authorityYear="1918" box="[410,554,1941,1966]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782801FF95FE0917E9FDF5F864" box="[410,554,1941,1966]" italics="true" pageId="1" pageNumber="138">M. argentina</emphasis>
</taxonomicName>
by the mesonotum without sulcus, and the elytra reduced in length, in this case, equal to 1.88 times the humeral width. In
<taxonomicName id="4C5F48E92801FF95FDF917C5FD24F818" authorityName="Bruch" authorityYear="1918" box="[618,763,1977,2002]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782801FF95FDF917C5FD24F818" box="[618,763,1977,2002]" italics="true" pageId="1" pageNumber="138">M. argentina</emphasis>
</taxonomicName>
, the elytra are also very short but different in color; they are yellowish in the two anterior thirds and reddish in the apical third. Furthermore, in
<taxonomicName id="4C5F48E92801FF95FBD117A1FB66F83C" authorityName="Martins" authorityYear="1981" box="[1090,1209,2013,2038]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="138" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782801FF95FBD117A1FB66F83C" box="[1090,1209,2013,2038]" italics="true" pageId="1" pageNumber="138">M. enigma</emphasis>
</taxonomicName>
the upper eye lobes are contiguous in great extension and in
<taxonomicName id="4C5F48E92803FF97FDF410EBFD25FF7A" authorityName="Bruch" authorityYear="1918" box="[615,762,151,176]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782803FF97FDF410EBFD25FF7A" box="[615,762,151,176]" italics="true" pageId="3" pageNumber="140">M. argentina</emphasis>
</taxonomicName>
they are separated by a distance corresponding to a row of ommatidia. The sulcus of the metatibia occupies, in the middle, only one-fifth of the total length in
<taxonomicName id="4C5F48E92803FF97FB4810C7FA8EFF1E" authorityName="Martins" authorityYear="1981" box="[1243,1361,187,212]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782803FF97FB4810C7FA8EFF1E" box="[1243,1361,187,212]" italics="true" pageId="3" pageNumber="140">M. enigma</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C5F48E92803FF97FF2710A3FE9AFF32" authorityName="Bruch" authorityYear="1918" box="[180,325,223,248]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782803FF97FF2710A3FE9AFF32" box="[180,325,223,248]" italics="true" pageId="3" pageNumber="140">M. argentina</emphasis>
</taxonomicName>
, it occupies the entire central region, except for the basal and apical quarters.”
</paragraph>
<caption id="DF2063E22802FF96FF0B17EBFD1AF832" ID-DOI="http://doi.org/10.5281/zenodo.7532251" ID-Zenodo-Dep="7532251" httpUri="https://zenodo.org/record/7532251/files/figure.png" pageId="2" pageNumber="139" startId="2.[152,270,1943,1967]" targetBox="[192,1396,181,1919]" targetPageId="2">
<paragraph id="8BE0336A2802FF96FF0B17EBFD1AF832" blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="139">
<emphasis id="B92BEF782802FF96FF0B17EBFE9DF865" bold="true" box="[152,322,1943,1967]" pageId="2" pageNumber="139">FIGURES 1–7.</emphasis>
<taxonomicName id="4C5F48E92802FF96FED817E4FE08F865" box="[331,471,1943,1968]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B92BEF782802FF96FED817E4FE4CF87A" box="[331,403,1944,1968]" italics="true" pageId="2" pageNumber="139">Methia</emphasis>
spp.
<emphasis id="B92BEF782802FF96FE5F17EBFE08F865" bold="true" box="[460,471,1943,1967]" pageId="2" pageNumber="139">1</emphasis>
</taxonomicName>
<emphasis id="B92BEF782802FF96FE4417EBFE24F865" bold="true" box="[471,507,1943,1967]" pageId="2" pageNumber="139">–2)</emphasis>
<taxonomicName id="4C5F48E92802FF96FD9017E4FCE8F87A" authority="Martins, 1981" authorityName="Martins" authorityYear="1981" box="[515,823,1944,1968]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782802FF96FD9017E4FD42F865" box="[515,669,1944,1968]" italics="true" pageId="2" pageNumber="139">Methia enigma</emphasis>
<bibRefCitation id="EFCE4E9B2802FF96FD3617E4FCE8F87A" author="Martins, U. R." box="[677,823,1944,1968]" pageId="2" pageNumber="139" pagination="9 - 17" refId="ref11439" refString="Martins, U. R. (1981) Revis &quot; o do genero Atenizus; os generos Tessaropa e Methia na America do Sul e descric &quot; o de novos taxa em Methiini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 25 (1), 9 - 17." type="journal article" year="1981">Martins, 1981</bibRefCitation>
</taxonomicName>
, paratype male:
<emphasis id="B92BEF782802FF96FC7017EBFC26F865" bold="true" box="[995,1017,1943,1967]" pageId="2" pageNumber="139">1)</emphasis>
Dorsal habitus;
<emphasis id="B92BEF782802FF96FB3717EBFB65F865" bold="true" box="[1188,1210,1943,1967]" pageId="2" pageNumber="139">2)</emphasis>
Eye.
<emphasis id="B92BEF782802FF96FB6417EBFAF9F865" bold="true" box="[1271,1318,1943,1967]" pageId="2" pageNumber="139">3–4)</emphasis>
<taxonomicName id="4C5F48E92802FF96FABD17E4FEB6F81E" authority="Bruch, 1918" authorityName="Bruch" authorityYear="1918" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782802FF96FABD17E4FF3CF819" italics="true" pageId="2" pageNumber="139">Methia argentina</emphasis>
Bruch, 1918
</taxonomicName>
, male from Argentina (Catamarca):
<emphasis id="B92BEF782802FF96FD4417C7FD32F819" bold="true" box="[727,749,1979,2003]" pageId="2" pageNumber="139">3)</emphasis>
Dorsal habitus;
<emphasis id="B92BEF782802FF96FC0017C7FC76F819" bold="true" box="[915,937,1979,2003]" pageId="2" pageNumber="139">4)</emphasis>
Eye.
<emphasis id="B92BEF782802FF96FC7017C7FC27F819" bold="true" box="[995,1016,1979,2003]" pageId="2" pageNumber="139">5)</emphasis>
<taxonomicName id="4C5F48E92802FF96FC6C17C0FB70F819" authorityName="Bruch" authorityYear="1918" box="[1023,1199,1980,2004]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="139" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782802FF96FC6C17C0FB70F819" box="[1023,1199,1980,2004]" italics="true" pageId="2" pageNumber="139">Methia argentina</emphasis>
</taxonomicName>
, female from Paraguay (Boquerón): Dorsal habitus;
<emphasis id="B92BEF782802FF96FE2917A3FE10F83D" bold="true" box="[442,463,2015,2039]" pageId="2" pageNumber="139">6)</emphasis>
Ventral habitus;
<emphasis id="B92BEF782802FF96FDE817A3FD4FF83D" bold="true" box="[635,656,2015,2039]" pageId="2" pageNumber="139">7)</emphasis>
Eye.
</paragraph>
</caption>
<paragraph id="8BE0336A2803FF97FF54117FFCE3FDF7" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="140">
Despite all these claims about the differences between the two species, it is probable that they are synonyms. The distance between upper eye lobes in specimens from
<collectingCountry id="F34873FA2803FF97FCB3115BFC57FE8B" box="[800,904,295,321]" name="Paraguay" pageId="3" pageNumber="140">Paraguay</collectingCountry>
and
<collectingCountry id="F34873FA2803FF97FC2D115BFBF2FE8B" box="[958,1069,295,321]" name="Argentina" pageId="3" pageNumber="140">Argentina</collectingCountry>
are variable, from contiguous to very slightly separated and, when contiguous only a small area of each lobe touches (as in the types of
<taxonomicName id="4C5F48E92803FF97FA871137FA54FEAE" authorityName="Martins" authorityYear="1981" box="[1300,1419,331,356]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782803FF97FA871137FA54FEAE" box="[1300,1419,331,356]" italics="true" pageId="3" pageNumber="140">M. enigma</emphasis>
</taxonomicName>
). The color is also somewhat variable in the specimens from
<collectingCountry id="F34873FA2803FF97FCB01113FC4DFE43" box="[803,914,367,393]" name="Argentina" pageId="3" pageNumber="140">Argentina</collectingCountry>
and
<collectingCountry id="F34873FA2803FF97FC551113FBF1FE43" box="[966,1070,367,393]" name="Paraguay" pageId="3" pageNumber="140">Paraguay</collectingCountry>
and can be equal in both species. The
<typeStatus id="54E48DC82803FF97FF5A11EFFEEBFE67" box="[201,308,403,429]" pageId="3" pageNumber="140" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C5F48E92803FF97FECA11EFFE0FFE66" authorityName="Martins" authorityYear="1981" box="[345,464,403,428]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782803FF97FECA11EFFE0FFE66" box="[345,464,403,428]" italics="true" pageId="3" pageNumber="140">M. enigma</emphasis>
</taxonomicName>
have the length of the sulcus of the metatibiae as in males of
<taxonomicName id="4C5F48E92803FF97FBEF11EFFAD2FE66" authorityName="Bruch" authorityYear="1918" box="[1148,1293,403,428]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782803FF97FBEF11EFFAD2FE66" box="[1148,1293,403,428]" italics="true" pageId="3" pageNumber="140">M. argentina</emphasis>
</taxonomicName>
, and not different as reported by
<bibRefCitation id="EFCE4E9B2803FF97FE1111CBFD7DFE1B" author="Martins, U. R. &amp; Galileo, M. H. M." box="[386,674,439,465]" pageId="3" pageNumber="140" refId="ref11493" refString="Martins, U. R. &amp; Galileo, M. H. M. (1997) Tribo Methiini. In: Martins, U. R (org.), Cerambycidae Sul-Americanos (Coleoptera). Vol. 1. Sociedade Brasileira de Entomologia, S &quot; o Paulo, pp. 159 - 179." type="book" year="1997">Martins &amp; Galileo (1997)</bibRefCitation>
. The only reliable difference found by us in the specimens deposited at MZSP collection, is the number of ommatidia between the upper and lower eye lobes: one row in specimens from
<collectingCountry id="F34873FA2803FF97FF041183FED6FDD3" box="[151,265,511,537]" name="Argentina" pageId="3" pageNumber="140">Argentina</collectingCountry>
, identified as
<taxonomicName id="4C5F48E92803FF97FE3A1183FDE4FDD2" authorityName="Bruch" authorityYear="1918" box="[425,571,511,536]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782803FF97FE3A1183FDE4FDD2" box="[425,571,511,536]" italics="true" pageId="3" pageNumber="140">M. argentina</emphasis>
</taxonomicName>
(
<figureCitation id="13642FEF2803FF97FDDF1183FD66FDD3" box="[588,697,511,537]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[192,1396,181,1919]" captionTargetId="figure-21@2.[192,1396,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–7. Methia spp. 1–2) Methia enigma Martins, 1981, paratype male: 1) Dorsal habitus; 2) Eye. 3–4) Methia argentina Bruch, 1918, male from Argentina (Catamarca): 3) Dorsal habitus; 4) Eye. 5) Methia argentina, female from Paraguay (Boquerón): Dorsal habitus; 6) Ventral habitus; 7) Eye." figureDoi="http://doi.org/10.5281/zenodo.7532251" httpUri="https://zenodo.org/record/7532251/files/figure.png" pageId="3" pageNumber="140">Figs. 4, 7</figureCitation>
); two rows in specimens from
<collectingCountry id="F34873FA2803FF97FB881183FB5BFDD3" box="[1051,1156,511,537]" name="Paraguay" pageId="3" pageNumber="140">Paraguay</collectingCountry>
, identified as
<taxonomicName id="4C5F48E92803FF97FAB01183FA43FDD2" authorityName="Martins" authorityYear="1981" box="[1315,1436,511,536]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782803FF97FAB01183FA43FDD2" box="[1315,1436,511,536]" italics="true" pageId="3" pageNumber="140">M. enigma</emphasis>
</taxonomicName>
(
<figureCitation id="13642FEF2803FF97FF0C125FFF3AFDF7" box="[159,229,547,573]" captionStart="FIGURES 1–7" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[192,1396,181,1919]" captionTargetId="figure-21@2.[192,1396,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–7. Methia spp. 1–2) Methia enigma Martins, 1981, paratype male: 1) Dorsal habitus; 2) Eye. 3–4) Methia argentina Bruch, 1918, male from Argentina (Catamarca): 3) Dorsal habitus; 4) Eye. 5) Methia argentina, female from Paraguay (Boquerón): Dorsal habitus; 6) Ventral habitus; 7) Eye." figureDoi="http://doi.org/10.5281/zenodo.7532251" httpUri="https://zenodo.org/record/7532251/files/figure.png" pageId="3" pageNumber="140">Fig. 2</figureCitation>
). See photographs of these species on
<bibRefCitation id="EFCE4E9B2803FF97FD07125FFCE7FDF7" author="Bezark, L. G." box="[660,824,547,573]" pageId="3" pageNumber="140" refId="ref10341" refString="Bezark, L. G. (2022) A photographic Catalog of the Cerambycidae of the World. New World Cerambycidae Catalog. Available from: http: // bezbycids. com / (accessed 27 September 2022)" type="url" year="2022">Bezark (2022)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE0336A2803FF97FF54123BFD15FD63" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="140">
The study of a larger series of specimens from
<collectingCountry id="F34873FA2803FF97FD5B123BFCE8FDAB" box="[712,823,583,609]" name="Argentina" pageId="3" pageNumber="140">Argentina</collectingCountry>
and
<collectingCountry id="F34873FA2803FF97FCF8123BFC0CFDAB" box="[875,979,583,609]" name="Paraguay" pageId="3" pageNumber="140">Paraguay</collectingCountry>
is required to corroborate this difference. For now, we consider the two species as different, but point out that the key from
<bibRefCitation id="EFCE4E9B2803FF97FBD01217FB32FD4F" author="Martins, U. R." box="[1091,1261,619,645]" pageId="3" pageNumber="140" pagination="9 - 17" refId="ref11439" refString="Martins, U. R. (1981) Revis &quot; o do genero Atenizus; os generos Tessaropa e Methia na America do Sul e descric &quot; o de novos taxa em Methiini (Coleoptera, Cerambycidae). Revista Brasileira de Entomologia, 25 (1), 9 - 17." type="journal article" year="1981">Martins (1981)</bibRefCitation>
and
<bibRefCitation id="EFCE4E9B2803FF97FAB41217FEE4FD63" author="Martins, U. R. &amp; Galileo, M. H. M." pageId="3" pageNumber="140" refId="ref11493" refString="Martins, U. R. &amp; Galileo, M. H. M. (1997) Tribo Methiini. In: Martins, U. R (org.), Cerambycidae Sul-Americanos (Coleoptera). Vol. 1. Sociedade Brasileira de Entomologia, S &quot; o Paulo, pp. 159 - 179." type="book" year="1997">Martins &amp; Galileo (1997)</bibRefCitation>
does not adequately separate them.
</paragraph>
</subSubSection>
<subSubSection id="C34560E12803FF97FF5412CEFCCFFC97" pageId="3" pageNumber="140" type="materials_examined">
<paragraph id="8BE0336A2803FF97FF5412CEFD5AFCDF" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="140">
<emphasis id="B92BEF782803FF97FF5412CEFE6BFD06" bold="true" box="[199,436,690,716]" pageId="3" pageNumber="140">Material examined.</emphasis>
<taxonomicName id="4C5F48E92803FF97FE2C12CFFD5AFD06" authorityName="Bruch" authorityYear="1918" box="[447,645,691,717]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="B92BEF782803FF97FE2C12CFFD5AFD06" box="[447,645,691,717]" italics="true" pageId="3" pageNumber="140">Methia argentina</emphasis>
</taxonomicName>
—
<materialsCitation id="3B3739372803FF97FD3212CFFD5BFD3B" collectingDate="1926-01" collectionCode="MZSP" collectorName="La Cienaga &amp; Weiser Woltrs" country="Argentina" determinerName="By Bruch" location="Belen" municipality="La Cienaga" pageId="3" pageNumber="140" specimenCount="2" specimenCount-male="2" stateProvince="Catamarca">
<collectingCountry id="F34873FA2803FF97FD3212CFFC98FD07" box="[673,839,691,717]" name="Argentina" pageId="3" pageNumber="140">ARGENTINA</collectingCountry>
,
<emphasis id="B92BEF782803FF97FCC512CFFC0CFD07" box="[854,979,691,717]" italics="true" pageId="3" pageNumber="140">
<collectingRegion id="499BFD882803FF97FCC512CFFC0CFD07" box="[854,979,691,717]" country="Argentina" name="Catamarca" pageId="3" pageNumber="140">Catamarca</collectingRegion>
</emphasis>
:
<collectorName id="26AA56BC2803FF97FC7612CFFBB6FD07" box="[997,1129,691,717]" pageId="3" pageNumber="140">
<collectingMunicipality id="6B84A9102803FF97FC7612CFFBB6FD07" box="[997,1129,691,717]" pageId="3" pageNumber="140">La Cienaga</collectingMunicipality>
</collectorName>
(
<location id="8E8065B12803FF97FBEF12CFFB1DFD07" LSID="urn:lsid:plazi:treatment:03F6827C2801FF97FF0415FEFCCFFC97:8E8065B12803FF97FBEF12CFFB1DFD07" box="[1148,1218,691,717]" country="Argentina" municipality="La Cienaga" name="Belen" pageId="3" pageNumber="140" stateProvince="Catamarca">Belén</location>
),
<specimenCount id="9D59F8E32803FF97FB4912CFFAECFD07" box="[1242,1331,691,717]" pageId="3" pageNumber="140" type="male">2 males</specimenCount>
(det.
<determinerName id="7E3CDFAE2803FF97FAEF12CFFF3FFD3B" pageId="3" pageNumber="140">By Bruch</determinerName>
),
<date id="FFE115AA2803FF97FF6712ABFEE0FD3B" box="[244,319,727,753]" pageId="3" pageNumber="140" value="1926-01">
<collectingDate id="EFA5EC422803FF97FF6712ABFEE0FD3B" box="[244,319,727,753]" pageId="3" pageNumber="140" value="1926-01">I.1926</collectingDate>
</date>
,
<collectorName id="26AA56BC2803FF97FEDF12ABFE34FD3B" box="[332,491,727,753]" pageId="3" pageNumber="140">Weiser Woltrs</collectorName>
leg. (
<collectionCode id="ED4EABAF2803FF97FDBE12ABFDA3FD3B" box="[557,636,727,753]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="140" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3739372803FF97FD0312ABFD5EFCDF" collectingDate="1990-10-23" collectionCode="MZSP" collectorName="Est. Agropil S. A. &amp; C. Aguilar" country="Paraguay" location="Boqueron" pageId="3" pageNumber="140" specimenCount="1" specimenCount-female="1" stateProvince="Boqueron">
<collectingCountry id="F34873FA2803FF97FD0312ABFCFAFD3B" box="[656,805,727,753]" name="Paraguay" pageId="3" pageNumber="140">PARAGUAY</collectingCountry>
(
<emphasis id="B92BEF782803FF97FCA412ABFBFAFD3B" bold="true" box="[823,1061,727,753]" pageId="3" pageNumber="140">new country record</emphasis>
),
<emphasis id="B92BEF782803FF97FBAE12ABFB74FD3B" box="[1085,1195,727,753]" italics="true" pageId="3" pageNumber="140">
<collectingRegion id="499BFD882803FF97FBAE12ABFB74FD3B" box="[1085,1195,727,753]" country="Paraguay" name="Boqueron" pageId="3" pageNumber="140">Boquerón</collectingRegion>
</emphasis>
:
<collectorName id="26AA56BC2803FF97FB2F12ABFA5FFD3B" box="[1212,1408,727,753]" pageId="3" pageNumber="140">Est. Agropil S.A.</collectorName>
,
<specimenCount id="9D59F8E32803FF97FA1D12ABFF39FCDF" pageId="3" pageNumber="140" type="female">1 female</specimenCount>
,
<date id="FFE115AA2803FF97FF621287FEB5FCDF" box="[241,362,763,789]" pageId="3" pageNumber="140" value="1990-10-23">
<collectingDate id="EFA5EC422803FF97FF621287FEB5FCDF" box="[241,362,763,789]" pageId="3" pageNumber="140" value="1990-10-23">23.X.1990</collectingDate>
</date>
,
<collectorName id="26AA56BC2803FF97FEE51287FE34FCDF" box="[374,491,763,789]" pageId="3" pageNumber="140">C. Aguilar</collectorName>
leg. (
<collectionCode id="ED4EABAF2803FF97FDBA1287FDA7FCDF" box="[553,632,763,789]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="140" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE0336A2803FF97FF541363FCCFFC97" blockId="3.[151,1437,151,861]" pageId="3" pageNumber="140">
<taxonomicName id="4C5F48E92803FF97FF541363FEB0FCF2" authorityName="Martins" authorityYear="1981" box="[199,367,799,825]" class="Insecta" family="Cerambycidae" genus="Methia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="140" phylum="Arthropoda" rank="species" species="enigma">
<emphasis id="B92BEF782803FF97FF541363FEB0FCF2" box="[199,367,799,825]" italics="true" pageId="3" pageNumber="140">Methia enigma</emphasis>
</taxonomicName>
—
<materialsCitation id="3B3739372803FF97FE181363FCD3FC97" collectionCode="MZSP" country="Paraguay" location="Chaco Territory" pageId="3" pageNumber="140" specimenCount="2" stateProvince="Alto Paraguay" typeStatus="paratype">
<collectingCountry id="F34873FA2803FF97FE181363FDFDFCF3" box="[395,546,799,825]" name="Paraguay" pageId="3" pageNumber="140">PARAGUAY</collectingCountry>
,
<emphasis id="B92BEF782803FF97FDBE1363FD0FFCF2" box="[557,720,799,825]" italics="true" pageId="3" pageNumber="140">
<collectingRegion id="499BFD882803FF97FDBE1363FD0FFCF2" box="[557,720,799,825]" country="Paraguay" name="Alto Paraguay" pageId="3" pageNumber="140">Alto Paraguay</collectingRegion>
</emphasis>
:
<location id="8E8065B12803FF97FD4C1363FC4EFCF3" LSID="urn:lsid:plazi:treatment:03F6827C2801FF97FF0415FEFCCFFC97:8E8065B12803FF97FD4C1363FC4EFCF3" box="[735,913,799,825]" country="Paraguay" name="Chaco Territory" pageId="3" pageNumber="140" stateProvince="Alto Paraguay">Chaco Territory</location>
, Abt. [about]
<locationDeviation id="9A522A482803FF97FBB81362FA5CFCF3" box="[1067,1411,798,825]" location="Puerto Casado" pageId="3" pageNumber="140">
<quantity id="4CA79E8F2803FF97FBB81362FB46FCF3" box="[1067,1177,798,825]" metricMagnitude="5" metricUnit="m" metricValue="2.414016" pageId="3" pageNumber="140" unit="mi" value="150.0">150 miles</quantity>
W of Puerto Casado
</locationDeviation>
,
<specimenCount id="9D59F8E32803FF97FA1D1363FEDDFC97" pageId="3" pageNumber="140" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
male, no date and collector indicated (
<collectionCode id="ED4EABAF2803FF97FD26133FFCDBFC97" box="[693,772,835,861]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="140" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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