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<document id="F8D22D341F29403AF0F89544500451E2" ID-DOI="10.37520/aemnp.2023.014" ID-ISSN="1804-6487" ID-Zenodo-Dep="10134093" ID-ZooBank="D982BDAC-CB44-41BE-A35C-92D630F96BF7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699897763845" checkinUser="felipe" docAuthor="Bilton, David T." docDate="2023" docId="03F6746EFFD4B62FFF45F88BFA7FB431" docLanguage="en" docName="ActaEntMusNatPra.63.2.249-264.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.014" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Helophorus Fabricius 1775" docType="key" docVersion="2" lastPageNumber="262" masterDocId="FFCF0C16FFD9B622FFE4FFB6FFE2B204" masterDocTitle="Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)" masterLastPageNumber="264" masterPageNumber="249" pageNumber="262" updateTime="1700044289635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="2A21DFB790A9FB8C8509378445119C17">Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)</mods:title>
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<mods:namePart id="C40E6E3EC04A36E1DFC2E989E4C5C4AC">Bilton, David T.</mods:namePart>
<mods:affiliation id="D7A6A9D74D4448C1DA6F1F4B841D5556">Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus Plymouth PL &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
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<mods:title id="677CBC657CB49AC8136A589152AC6664">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="2D961A3DB2307717A21DD87F3BBD48AA">2023</mods:date>
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<mods:number id="17133BF5724FD0AC951D366AD9DEEC21">2023-09-10</mods:number>
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<mods:identifier id="23FFD5C33F813AA6001CA8566068CA50" type="DOI">10.37520/aemnp.2023.014</mods:identifier>
<mods:identifier id="239625BD1C4E3944B2036A5931A3BF7B" type="ISSN">1804-6487</mods:identifier>
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<treatment id="03F6746EFFD4B62FFF45F88BFA7FB431" LSID="urn:lsid:plazi:treatment:03F6746EFFD4B62FFF45F88BFA7FB431" httpUri="http://treatment.plazi.org/id/03F6746EFFD4B62FFF45F88BFA7FB431" lastPageNumber="262" pageId="13" pageNumber="262">
<subSubSection id="C34596F3FFD4B62FFF45F88BFD0FB553" box="[161,749,1852,1879]" pageId="13" pageNumber="262" type="nomenclature">
<paragraph id="8BE0C578FFD4B62FFF45F88BFD0FB553" blockId="13.[161,749,1852,1879]" box="[161,749,1852,1879]" pageId="13" pageNumber="262">
<heading id="D0A87214FFD4B62FFF45F88BFD0FB553" bold="true" box="[161,749,1852,1879]" centered="true" fontSize="11" level="1" pageId="13" pageNumber="262" reason="1">
<emphasis id="B92B196AFFD4B62FFF45F88BFD0FB553" bold="true" box="[161,749,1852,1879]" pageId="13" pageNumber="262">
Key to the Afrotropical species of
<taxonomicName id="4C5FBEFBFFD4B62FFDBAF88AFD0FB553" ID-CoL="8VRQF" authorityName="Fabricius" authorityYear="1775" box="[606,749,1852,1879]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B92B196AFFD4B62FFDBAF88AFD0FB553" bold="true" box="[606,749,1852,1879]" italics="true" pageId="13" pageNumber="262">Helophorus</emphasis>
</taxonomicName>
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<subSubSection id="C34596F3FFD4B62FFF73F8D8FA7FB431" pageId="13" pageNumber="262" type="key">
<keyStep id="30ABDDDDFFD4B62FFF73F8D8FD15B5A1" pageId="13" pageNumber="262">
<paragraph id="8BE0C578FFD4B62FFF73F8D8FD15B581" blockId="13.[151,760,1902,2053]" box="[151,759,1902,1925]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFF73F8D8FD15B581" box="[151,759,1902,1925]" pageId="13" pageNumber="262">1 Antennae 9-segmented. ............................................ 2</keyLead>
</paragraph>
<paragraph id="8BE0C578FFD4B62FFF73F838FD15B5A1" blockId="13.[151,760,1902,2053]" box="[151,759,1934,1957]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFF73F838FD15B5A1" box="[151,759,1934,1957]" pageId="13" pageNumber="262">– Antennae 8-segmented. ............................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ABDDDDFFD4B62FFF73F818FA7EB391" pageId="13" pageNumber="262">
<paragraph id="8BE0C578FFD4B62FFF73F818FA7EB311" blockId="13.[151,760,1902,2053]" lastBlockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFF73F818FA7EB311" pageId="13" pageNumber="262">
2 Pronotal internal and middle intervals smooth, granulation reduced to pores and setae over most of surface (
<figureCitation id="1364D9FDFFD4B62FFF59F858FEF7BA01" box="[189,277,2030,2053]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1D</figureCitation>
&amp;
<figureCitation id="1364D9FDFFD4B62FFEDEF858FEBEBA01" box="[314,348,2030,2053]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3C</figureCitation>
); median lobe of aedeagus with struts as long as tube (
<figureCitation id="1364D9FDFFD4B62FFBF8FF08FB95B2D1" box="[1052,1143,190,213]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">Fig. 3G</figureCitation>
); habitus relatively robust (
<figureCitation id="1364D9FDFFD4B62FFC86FF68FC54B2F1" box="[866,950,222,245]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Fig. 1D</figureCitation>
), body length 3.45–4.0 mm. .......................... ...................................
<taxonomicName id="4C5FBEFBFFD4B62FFB82FF48FA7EB311" ID-CoL="3KHJ3" authority="Orchymont, 1947" authorityName="Orchymont" authorityYear="1947" baseAuthorityName="Orchymont" baseAuthorityYear="1948" box="[1126,1436,253,277]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="cooperi">
<emphasis id="B92B196AFFD4B62FFB82FF48FB3BB311" bold="true" box="[1126,1241,253,277]" italics="true" pageId="13" pageNumber="262">H. cooperi</emphasis>
Orchymont, 1947
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE0C578FFD4B62FFCD8FEA8FA7EB391" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FEA8FA7EB391" pageId="13" pageNumber="262">
– Pronotal internal and middle intervals with evident granulation over most of surface (
<figureCitation id="1364D9FDFFD4B62FFB32FE88FA8DB351" box="[1238,1391,318,341]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1A &amp; G</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFA99FE88FC68B371" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3A &amp; F</figureCitation>
); median lobe of aedeagus with struts distinctly shorter than tube. ...................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ABDDDDFFD4B62FFCD8FE28FA7FB111" pageId="13" pageNumber="262">
<paragraph id="8BE0C578FFD4B62FFCD8FE28FA7EB051" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FE28FA7EB051" pageId="13" pageNumber="262">
3 Granulation on pronotal internal and middle intervals weaker (
<figureCitation id="1364D9FDFFD4B62FFC58FE08FBF8B3D1" box="[956,1050,446,469]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1A</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFBC9FE08FBB3B3D1" box="[1069,1105,446,469]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3A</figureCitation>
); prontoal submedian groove more strongly angled medially (
<figureCitation id="1364D9FDFFD4B62FFB29FE68FAC8B3F1" box="[1229,1322,478,501]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">Fig. 3A</figureCitation>
); parameres relatively straight, with evenly pointed tips (
<figureCitation id="1364D9FDFFD4B62FFA89FE48FC7CB031" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="13" pageNumber="262">Figs 2A–B</figureCitation>
); body size smaller,
<quantity id="4CA7689DFFD4B62FFB6DFDA8FACEB031" box="[1161,1324,542,565]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="13" pageNumber="262" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.60–3.50 mm</quantity>
; aedeagus smaller,
<quantity id="4CA7689DFFD4B62FFC51FD88FBF5B051" box="[949,1047,574,597]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="262" unit="mm" value="0.45">0.45 mm</quantity>
. ...
<taxonomicName id="4C5FBEFBFFD4B62FFBA6FD88FA7EB051" ID-CoL="6LMZX" authority="Regimbart, 1907" authorityName="Regimbart" authorityYear="1907" box="[1090,1436,573,597]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="B92B196AFFD4B62FFBA6FD88FB00B051" bold="true" box="[1090,1250,573,597]" italics="true" pageId="13" pageNumber="262">H. aethiopicus</emphasis>
<bibRefCitation id="EFCEB889FFD4B62FFB0DFD88FA7EB051" author="REGIMBART M." box="[1257,1436,574,597]" pageId="13" pageNumber="262" pagination="126 - 128" refId="ref16606" refString="REGIMBART M. 1907: Collections recueillis par M. Maurice de Rothschild dans l'Afrique orientale. Insectes coleopteres: Dytiscides et hydrophilides. Bulletin du Museum National d'Histoire Naturelle Paris 13: 126 - 128." type="journal article" year="1907">Régimbart, 1907</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BE0C578FFD4B62FFCD8FDE8FA7FB111" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FDE8FA7FB111" pageId="13" pageNumber="262">
– Granulation on pronotal internal and middle intervals stronger (
<figureCitation id="1364D9FDFFD4B62FFC21FDC8FBC1B091" box="[965,1059,638,661]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1G</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFBD0FDC8FBB1B091" box="[1076,1107,638,661]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3F</figureCitation>
); pronotal submedian groove more weakly angled medially (
<figureCitation id="1364D9FDFFD4B62FFB20FD28FAFFB0B1" box="[1220,1309,670,693]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">Fig. 3F</figureCitation>
); paramere tips more broadly rounded (
<figureCitation id="1364D9FDFFD4B62FFB61FD08FB3AB0D1" box="[1157,1240,702,725]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">Fig. 3L</figureCitation>
); body size larger,
<quantity id="4CA7689DFFD4B62FFCBEFD68FC1AB0F1" box="[858,1016,734,757]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.85" metricValueMin="3.15" pageId="13" pageNumber="262" unit="mm" value="3.5" valueMax="3.85" valueMin="3.15">3.15–3.85 mm</quantity>
; aedeagus larger,
<quantity id="4CA7689DFFD4B62FFB51FD68FAFAB0F1" box="[1205,1304,734,757]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="262" unit="mm" value="0.55">0.55 mm</quantity>
. .................. ................................................
<emphasis id="B92B196AFFD4B62FFB5DFD48FA7FB111" bold="true" box="[1209,1437,765,789]" pageId="13" pageNumber="262">
<taxonomicName id="4C5FBEFBFFD4B62FFB5DFD48FAA3B111" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1209,1345,765,789]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="simiensis" status="sp. nov.">
<emphasis id="B92B196AFFD4B62FFB5DFD48FAA3B111" bold="true" box="[1209,1345,765,789]" italics="true" pageId="13" pageNumber="262">H. simiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFD4B62FFAA3FD48FA7FB111" box="[1351,1437,766,789]" pageId="13" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30ABDDDDFFD4B62FFCD8FCA8FA7EB631" pageId="13" pageNumber="262">
<paragraph id="8BE0C578FFD4B62FFCD8FCA8FA7FB1B1" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FCA8FA7FB1B1" pageId="13" pageNumber="262">
4 Aedeagus very large,
<quantity id="4CA7689DFFD4B62FFBADFCA8FB0BB131" box="[1097,1257,798,821]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.5" metricValueMin="7.8" pageId="13" pageNumber="262" unit="mm" value="0.815" valueMax="0.85" valueMin="0.78">0.78–0.85 mm</quantity>
, parameres with acuminate tips (
<figureCitation id="1364D9FDFFD4B62FFBE0FC88FBB8B151" box="[1028,1114,830,853]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="13" pageNumber="262">Fig. 2K</figureCitation>
); pronotal intervals raised and granulate (
<figureCitation id="1364D9FDFFD4B62FFC2EFCE8FBF9B171" box="[970,1051,862,885]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1F</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFBC1FCE8FBA4B171" box="[1061,1094,862,885]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3E</figureCitation>
); body robust (
<figureCitation id="1364D9FDFFD4B62FFB00FCE8FAD2B171" box="[1252,1328,862,885]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Fig. 1F</figureCitation>
) and relatively large,
<quantity id="4CA7689DFFD4B62FFC04FCC8FB9CB191" box="[992,1150,894,917]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.55" metricValueMin="3.25" pageId="13" pageNumber="262" unit="mm" value="3.9" valueMax="4.55" valueMin="3.25">3.25–4.55 mm</quantity>
; species from East African Rift. ................................
<emphasis id="B92B196AFFD4B62FFB8BFC28FA7FB1B1" bold="true" box="[1135,1437,925,949]" pageId="13" pageNumber="262">
<taxonomicName id="4C5FBEFBFFD4B62FFB8BFC28FAA2B1B1" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1135,1344,925,949]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="nyandaruaensis" status="sp. nov.">
<emphasis id="B92B196AFFD4B62FFB8BFC28FAA2B1B1" bold="true" box="[1135,1344,925,949]" italics="true" pageId="13" pageNumber="262">H. nyandaruaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFD4B62FFAA3FC28FA7FB1B1" box="[1351,1437,926,949]" pageId="13" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BE0C578FFD4B62FFCD8FC08FA7EB631" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FC08FA7EB631" pageId="13" pageNumber="262">
– Aedeagus smaller, maximum length
<quantity id="4CA7689DFFD4B62FFB3FFC08FAD9B1D1" box="[1243,1339,958,981]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="13" pageNumber="262" unit="mm" value="0.66">0.66 mm</quantity>
, paramere tips various, but not strongly acuminate (
<figureCitation id="1364D9FDFFD4B62FFAC3FC68FC6BB611" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="13" pageNumber="262">Figs 2C–F, H–J</figureCitation>
); pronotal intervals flatter (
<figureCitation id="1364D9FDFFD4B62FFB56FC48FAA8B611" box="[1202,1354,1022,1045]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1B–C, E</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFAB3FC48FA7AB611" box="[1367,1432,1022,1045]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3B, E</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFCBEFBA8FC7BB631" box="[858,921,1054,1077]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">4C–F</figureCitation>
). ....................................................................... 5
</keyLead>
</paragraph>
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<keyStep id="30ABDDDDFFD4B62FFCD8FB88FA7FB431" pageId="13" pageNumber="262">
<paragraph id="8BE0C578FFD4B62FFCD8FB88FA7EB731" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FB88FA7EB731" pageId="13" pageNumber="262">
5 Pronotal submedian groove broader, granulation on middle and internal intervals weaker (
<figureCitation id="1364D9FDFFD4B62FFB17FBE8FA88B671" box="[1267,1386,1118,1141]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1B–C</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFA92FBE8FA7AB671" box="[1398,1432,1118,1141]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3B</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFCBEFBC8FC79B691" box="[858,923,1150,1173]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">4C, E</figureCitation>
); walls of pronotal grooves straighter and sharper (
<figureCitation id="1364D9FDFFD4B62FFC6FFB28FBE0B6B1" box="[907,1026,1182,1205]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">Figs 4C, E</figureCitation>
); punctures of elytral striae larger and intervals narrower and more arched apically (
<figureCitation id="1364D9FDFFD4B62FFADEFB08FA72B6D1" box="[1338,1424,1214,1237]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">Fig. 4G</figureCitation>
); aedeagus smaller,
<quantity id="4CA7689DFFD4B62FFBF8FB68FB5AB6F1" box="[1052,1208,1246,1269]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="13" pageNumber="262" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
, parameres with relatively narrow apices (
<figureCitation id="1364D9FDFFD4B62FFBA4FB48FB54B711" box="[1088,1206,1278,1301]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="13" pageNumber="262">Figs 2C–F</figureCitation>
). ............................... ......................
<taxonomicName id="4C5FBEFBFFD4B62FFBEBFAA8FA7EB731" ID-CoL="3KHLV" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" box="[1039,1436,1309,1333]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="aethiops">
<emphasis id="B92B196AFFD4B62FFBEBFAA8FB6FB731" bold="true" box="[1039,1165,1309,1333]" italics="true" pageId="13" pageNumber="262">H. aethiops</emphasis>
J.
<bibRefCitation id="EFCEB889FFD4B62FFB4FFAA8FA7EB731" author="BALFOUR-BROWNE J." box="[1195,1436,1310,1333]" pageId="13" pageNumber="262" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BE0C578FFD4B62FFCD8FA88FA7FB431" blockId="13.[828,1437,190,1589]" pageId="13" pageNumber="262">
<keyLead id="30AE664DFFD4B62FFCD8FA88FA7FB431" pageId="13" pageNumber="262">
– Pronotal submedian groove narrower, granulation on middle and internal intervals stronger (
<figureCitation id="1364D9FDFFD4B62FFAE8FAE8FA87B771" box="[1292,1381,1374,1397]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="13" pageNumber="262">Figs 1E</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFA91FAE8FA7AB771" box="[1397,1432,1374,1397]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="13" pageNumber="262">3D</figureCitation>
,
<figureCitation id="1364D9FDFFD4B62FFCBEFAC8FC7FB791" box="[858,925,1406,1429]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">4D, F</figureCitation>
); walls of pronotal grooves more rounded and indistinct (
<figureCitation id="1364D9FDFFD4B62FFC37FA28FBBBB7B1" box="[979,1113,1438,1461]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">Figs 4D, F</figureCitation>
); punctures of elytral striae smaller and intervals broader and flatter apically (
<figureCitation id="1364D9FDFFD4B62FFA95FA08FC9DB7F1" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E &amp; G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F &amp; H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="13" pageNumber="262">Fig. 4H</figureCitation>
); aedeagus larger,
<quantity id="4CA7689DFFD4B62FFBA5FA68FB3CB7F1" box="[1089,1246,1502,1525]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" metricValueMax="6.6" metricValueMin="5.8" pageId="13" pageNumber="262" unit="mm" value="0.62" valueMax="0.66" valueMin="0.58">0.58–0.66 mm</quantity>
, parameres usually with relatively broad apices (
<figureCitation id="1364D9FDFFD4B62FFB4FFA48FAFCB411" box="[1195,1310,1534,1557]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="13" pageNumber="262">Figs 2H–J</figureCitation>
). ................ ......................................
<emphasis id="B92B196AFFD4B62FFB9CF9A8FA7FB431" bold="true" box="[1144,1437,1565,1589]" pageId="13" pageNumber="262">
<taxonomicName id="4C5FBEFBFFD4B62FFB9CF9A8FAA2B431" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1144,1344,1565,1589]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="262" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
<emphasis id="B92B196AFFD4B62FFB9CF9A8FAA2B431" bold="true" box="[1144,1344,1565,1589]" italics="true" pageId="13" pageNumber="262">H. brumopluvialis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218A411FFD4B62FFAA3F9A8FA7FB431" box="[1351,1437,1566,1589]" pageId="13" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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