307 lines
43 KiB
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307 lines
43 KiB
XML
<document id="40658A32734AEE3B2C1A676852633562" ID="10.11646/zootaxa.4137.4.6" ID-DOI="10.11646/zootaxa.4137.4.6" ID-GBIF-Dataset="d84bf5c7-4a51-441e-a1d2-00f109129808" ID-ISSN="1175-5326" ID-Zenodo-Dep="265422" ID-ZooBank="AEC399C6-02FC-4747-B497-68095084B1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468394324568" checkinUser="plazi" docAuthor="Garbelotto, Thereza De A., Campos, Luiz A. & Grazia, Jocelia" docDate="2016" docId="03F55D2AFF90103EFF6BC423E9D4FB97" docLanguage="en" docName="zootaxa.4137.4.6.pdf" docOrigin="Zootaxa 4137 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stapecolis Garbelotto & Campos, gen. nov." docType="treatment" docUuid="2A44EF61-AFE0-4359-9166-C9EBABE2C98A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="549" masterDocId="FFCC2552FF91103AFFFCC17CEC48FFF4" masterDocTitle="Stapecolis, new genus of Ochlerini (Hemiptera: Pentatomidae: Discocephalinae)" masterLastPageNumber="552" masterPageNumber="545" pageNumber="546" updateTime="1698364656965" updateUser="plazi">
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<mods:mods id="5F33E43C441CA0244F60F4131154A167" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4FD19210A29E78331C0A59B80BAD6801">
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<mods:title id="D6280E95F4AFA219EFF7C2D05E7637B0">Stapecolis, new genus of Ochlerini (Hemiptera: Pentatomidae: Discocephalinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="11FDD634B67F5797C3AC4B7A6075E23C" type="personal">
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<mods:role id="92091D799B89A94FFF28012485D8484F">
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<mods:roleTerm id="4248D03AF6569684FF622B89A9B913B2">Author</mods:roleTerm>
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<mods:namePart id="A2C4770B77F715BE135865E4D0E1C046">Garbelotto, Thereza De A.</mods:namePart>
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<mods:affiliation id="614CE4D6E72B2E92F8C5576ABC809624">Universidade Federal do Rio Grande do Sul, Dep. Zoologia, Av. Bento Gonçalves 9500, 91501 - 970, Prédio 43435, Porto Alegre RS</mods:affiliation>
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<mods:nameIdentifier id="85DB1AE1A6764B1C09D1EA540EBC617C" type="email">therezagarbelotto@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="DE9CA5A9BFD175E05E59B9105638E718" type="personal">
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<mods:role id="3DD1D13EB3944F191CB5097011174EAC">
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<mods:roleTerm id="6913782177C422C016FE15DD9B05FE4F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4EA0DBCBEC05AEBD9B4794ED054BACB7">Campos, Luiz A.</mods:namePart>
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<mods:affiliation id="32B9EA5485B2D70F0E4A67E90B07C230">Universidade Federal do Rio Grande do Sul, Dep. Zoologia, Av. Bento Gonçalves 9500, 91501 - 970, Prédio 43435, Porto Alegre RS</mods:affiliation>
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<mods:nameIdentifier id="0C4790E21EF0BBED8182190AAEFB7A57" type="email">luiz.campos@ufrgs.br</mods:nameIdentifier>
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</mods:name>
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<mods:name id="353A3EE8A2C56421F6A85A9693969B0C" type="personal">
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<mods:roleTerm id="C3F2F8F6279B38651E544359A16BA647">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AC165086D239A0EFA57726D212BB7378">Grazia, Jocelia</mods:namePart>
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<mods:affiliation id="F88B70FAD7F1A79EB4252FD158C54A6B">Universidade Federal do Rio Grande do Sul, Dep. Zoologia, Av. Bento Gonçalves 9500, 91501 - 970, Prédio 43435, Porto Alegre RS</mods:affiliation>
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<mods:nameIdentifier id="CA6EABA27494047A49D6F270CB79C084" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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<mods:typeOfResource id="C733BBBF3203FEC41932E221587C2D16">text</mods:typeOfResource>
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<mods:titleInfo id="C7666C5496A1948E1C43D48CAD0ACBD8">
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<mods:title id="7EDE71B829836E7BF51E608E77088050">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FE44FA05DEF847993C1B9D54D88D38E8">
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<mods:date id="DA4282E3743C0D9B48A20B7A1CF9BBE2">2016</mods:date>
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<mods:detail id="3785D353350C1B384D964BE014B0BCC4" type="pubDate">
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<mods:number id="C007AD87BD064A9BD3C824A575782563">2016-07-13</mods:number>
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<mods:detail id="B9FB66F743EBE3E422969825339A5DB9" type="volume">
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<mods:number id="0BAB0A6D13697FBE9B81FADF5544E8B1">4137</mods:number>
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<mods:detail id="64ACABE5148F34540ED0FD4FC4BE8E42" type="issue">
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<mods:number id="9A32E28F5A25291AF8B16F0CD1683578">4</mods:number>
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<mods:classification id="12ACFB4AC0B908B3005017D275A0B621">journal article</mods:classification>
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<mods:identifier id="DE6AD406223FDDB7BA1EB1F9C0524097" type="DOI">10.11646/zootaxa.4137.4.6</mods:identifier>
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<mods:identifier id="11ACBA010EF3D64AFFD0D49205D6A7CA" type="GBIF-Dataset">d84bf5c7-4a51-441e-a1d2-00f109129808</mods:identifier>
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<mods:identifier id="E2CB7D7F88E787079D80DB818A4D4A3E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D3B81E9A92F1565D42501951893A26A8" type="Zenodo-Dep">265422</mods:identifier>
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<mods:identifier id="CE30E7FE218F7020EA80BE68FFEE6F4D" type="ZooBank">AEC399C6-02FC-4747-B497-68095084B1E7</mods:identifier>
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</mods:mods>
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<treatment id="03F55D2AFF90103EFF6BC423E9D4FB97" ID-DOI="http://doi.org/10.5281/zenodo.6068949" ID-GBIF-Taxon="164613087" ID-Zenodo-Dep="6068949" LSID="urn:lsid:zoobank.org:act:2A44EF61-AFE0-4359-9166-C9EBABE2C98A" httpUri="http://treatment.plazi.org/id/03F55D2AFF90103EFF6BC423E9D4FB97" lastPageId="4" lastPageNumber="549" pageId="1" pageNumber="546">
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<subSubSection id="C346BFB7FF90103BFF6BC423EEEAFA8E" box="[151,674,1375,1402]" pageId="1" pageNumber="546" type="nomenclature">
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<paragraph id="8BE3EC3CFF90103BFF6BC423EEEAFA8E" blockId="1.[151,674,1375,1402]" box="[151,674,1375,1402]" pageId="1" pageNumber="546">
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<heading id="D0AB5B50FF90103BFF6BC423EEEAFA8E" bold="true" box="[151,674,1375,1402]" fontSize="11" level="1" pageId="1" pageNumber="546" reason="1">
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<emphasis id="B928302EFF90103BFF6BC423EEEAFA8E" bold="true" box="[151,674,1375,1402]" pageId="1" pageNumber="546">
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||
<taxonomicName id="4C5C97BFFF90103BFF6BC423EE64FA8E" authority="Garbelotto & Campos" authorityName="Garbelotto & Campos" box="[151,556,1375,1402]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B928302EFF90103BFF6BC423ED47FA8D" bold="true" box="[151,271,1375,1401]" italics="true" pageId="1" pageNumber="546">Stapecolis</emphasis>
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Garbelotto & Campos
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</taxonomicName>
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<taxonomicNameLabel id="A21B8D55FF90103BFDC8C41CEEEAFA8E" box="[564,674,1376,1402]" pageId="1" pageNumber="546" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C346BFB7FF90103BFF6BC4DBEFFEFA48" box="[151,950,1446,1469]" pageId="1" pageNumber="546" type="type_taxon">
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||
<paragraph id="8BE3EC3CFF90103BFF6BC4DBEFFEFA48" blockId="1.[151,950,1446,1469]" box="[151,950,1446,1469]" pageId="1" pageNumber="546">
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<typeStatus id="54E7529EFF90103BFF6BC4DBEC81FA48" box="[151,201,1447,1468]" pageId="1" pageNumber="546">Type</typeStatus>
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species:
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<taxonomicName id="4C5C97BFFF90103BFEDAC4DAEEE5FA48" authority="Garbelotto & Campos" authorityName="Garbelotto & Campos" box="[294,685,1446,1469]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="546" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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<emphasis id="B928302EFF90103BFEDAC4DAED8FFA49" bold="true" box="[294,455,1446,1469]" italics="true" pageId="1" pageNumber="546">Stapecolis latus</emphasis>
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Garbelotto & Campos
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</taxonomicName>
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<emphasis id="B928302EFF90103BFD4FC4DBEF4BFA48" bold="true" box="[691,771,1447,1468]" pageId="1" pageNumber="546">
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<taxonomicNameLabel id="A21B8D55FF90103BFD4FC4DBEF4BFA48" box="[691,771,1447,1468]" pageId="1" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C346BFB7FF90103BFF6BC495ED9FF9D2" pageId="1" pageNumber="546" type="etymology">
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<paragraph id="8BE3EC3CFF90103BFF6BC495ED9FF9D2" blockId="1.[151,1437,1513,1970]" pageId="1" pageNumber="546">
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<emphasis id="B928302EFF90103BFF6BC495ED56F9F6" bold="true" box="[151,286,1513,1538]" pageId="1" pageNumber="546">Etymology.</emphasis>
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The name refers to the shape of the ventral rim opening of the male pygophore. Latin:
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<emphasis id="B928302EFF90103BFAEAC496E914F9F5" box="[1302,1372,1514,1537]" italics="true" pageId="1" pageNumber="546">stapes</emphasis>
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(m.), stirrup;
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<emphasis id="B928302EFF90103BFF13C771ED6CF9D1" box="[239,292,1549,1573]" italics="true" pageId="1" pageNumber="546">colis</emphasis>
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, male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C346BFB7FF901039FF3BC74DE90AFDEC" lastPageId="3" lastPageNumber="548" pageId="1" pageNumber="546" type="description">
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<paragraph id="8BE3EC3CFF90103BFF3BC74DECACF966" blockId="1.[151,1437,1513,1970]" pageId="1" pageNumber="546">
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<emphasis id="B928302EFF90103BFF3BC74DED11F9BE" bold="true" box="[199,345,1585,1610]" pageId="1" pageNumber="546">Description.</emphasis>
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Body elongated, general color brown to black with black punctures. Corium with 1+1 yellow spots at apex of radial vein (
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<figureCitation id="1367F0B9FF90103BFE2CC729EE74F99A" box="[464,572,1621,1646]" captionStart="FIGURES 2 – 3. 1" captionStartId="2.[151,264,1101,1124]" captionTargetBox="[163,1432,197,1073]" captionTargetId="figure@2.[151,1436,193,1080]" captionTargetPageId="2" captionText="FIGURES 2 – 3. 1, Stapecolis latus sp. nov., male holotype; 2, Stapecolis bimaculatus sp. nov., male holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/265424/files/figure.png" pageId="1" pageNumber="546">Figs. 2–3</figureCitation>
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). Antennae five-segmented. Ventral rim of pygophore subrectangular (
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<figureCitation id="1367F0B9FF90103BFAB3C729EC9EF966" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="1" pageNumber="546">Figs. 5 and 7</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE3EC3CFF90103BFF3BC7E1E879F846" blockId="1.[151,1437,1513,1970]" pageId="1" pageNumber="546">Head wider than long, declivous; punctures confluent forming wrinkles from base of clypeus to posterior margin of head. Mandibular plates longer than clypeus, apart at apex, this rounded; lateral margins sinuous and reflected. Vertex weakly tumescent. Anteocular process truncated and not projected. Eyes on a short peduncle. Antennal tubercles with lateral spine, visible from above. Antennae five-segmented, segment II the shortest. Ventrally on head 1+1 longitudinal carinae posterior to the eyes, concolorous with the surrounding surface. Bucculae elevated, evanescent posteriorly, not attaining posterior margin of head, with triangular anterior tooth, and narrowed on anterior third. Rostrum not reaching anterior margin of urosternite V; first segment attaining prosternum; insertion of first segment of labium posterior to anterior limit of eyes.</paragraph>
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<caption id="DF23BCB4FF931038FF6BC531EC9AFB76" httpUri="https://zenodo.org/record/265424/files/figure.png" pageId="2" pageNumber="547" targetBox="[163,1432,197,1073]" targetPageId="2">
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<paragraph id="8BE3EC3CFF931038FF6BC531EC9AFB76" blockId="2.[151,1437,1101,1154]" pageId="2" pageNumber="547">
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<emphasis id="B928302EFF931038FF6BC531ED75FB97" bold="true" box="[151,317,1101,1124]" pageId="2" pageNumber="547">FIGURES 2–3.</emphasis>
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1,
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<taxonomicName id="4C5C97BFFF931038FE9EC532EE4CFB90" box="[354,516,1102,1124]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="547" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
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<emphasis id="B928302EFF931038FE9EC532EE4CFB90" box="[354,516,1102,1124]" italics="true" pageId="2" pageNumber="547">Stapecolis latus</emphasis>
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</taxonomicName>
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<emphasis id="B928302EFF931038FDF1C532EE28FB97" bold="true" box="[525,608,1102,1123]" pageId="2" pageNumber="547">
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<taxonomicNameLabel id="A21B8D55FF931038FDF1C532EE28FB97" box="[525,608,1102,1123]" pageId="2" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male holotype; 2,
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<taxonomicName id="4C5C97BFFF931038FCCCC532E854FB90" box="[816,1052,1102,1124]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="547" phylum="Arthropoda" rank="species" species="bimaculatus" status="sp. nov.">
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<emphasis id="B928302EFF931038FCCCC532E854FB90" box="[816,1052,1102,1124]" italics="true" pageId="2" pageNumber="547">Stapecolis bimaculatus</emphasis>
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</taxonomicName>
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<emphasis id="B928302EFF931038FBD9C532E830FB97" bold="true" box="[1061,1144,1102,1123]" pageId="2" pageNumber="547">
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<taxonomicNameLabel id="A21B8D55FF931038FBD9C532E830FB97" box="[1061,1144,1102,1123]" pageId="2" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male holotype. Scale bars: 1mm.
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</paragraph>
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</caption>
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<paragraph id="8BE3EC3CFF931038FF3BC5D7E86DF9DF" blockId="2.[151,1437,1194,2011]" pageId="2" pageNumber="547">Pronotum wider than long, punctures confluent forming wrinkles on posterior half; anterior half declivous. Cicatrices flat. Anterior margin concave on the median third. Anterior angle of pronotum developed as acute tooth, projected laterally by the ocellus diameter. Anterolateral margins of pronotum crenulated; posterior margin subrectilinear; posterolateral margins slightly concave. Scutellum longer than wide, reaching connexival segment V; punctures confluent, forming wrinkles on scutellum disc. Corium longer than scutellum, reaching connexival segment VI; punctures dense and not confluent; 1+1 yellow maculae at apex of radial vein. Hemelytral membrane not surpassing apex of abdomen, with nine veins. Pro-, meso- and metathorax black, punctures not confluent. Prosternum sulcate, meso- and metasternum with median carina along entire length. Mesopleural evaporatorium black, occupying a thin area along the posterior margin of mesopleurae; metapleural evaporatorium black, occupying half the width of metapleurae, lateral margin subrectilinear. Peritreme in spout. Tibiae sulcate dorsally; third segment of metatarsi dorsally flat in males and dorsally sulcated in females.</paragraph>
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<paragraph id="8BE3EC3CFF931038FF3BC74AEDF7F94E" blockId="2.[151,1437,1194,2011]" pageId="2" pageNumber="547">Connexivum exposed from segments III to VIII; punctures smaller than in the remaining dorsal surface of the body, narrow lateral strip without punctures. Abdomen with punctures inconspicuous or absent on disc and external to trichobothria. Trichobothria lateral to imaginary line tangential to spiracles. Spiracles black, somewhat elevated, visible on sternites II–VII.</paragraph>
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<paragraph id="8BE3EC3CFF931038FF3BC7BAEEB9F8F7" blockId="2.[151,1437,1194,2011]" pageId="2" pageNumber="547">Male. Antennae: third segment the widest, twice wider than the next segment. Articular membrane along the posterior margin of urotergite VII expanded medially.</paragraph>
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<paragraph id="8BE3EC3CFF931039FF3BC672E8B6FE7C" blockId="2.[151,1437,1194,2011]" lastBlockId="3.[151,1437,151,536]" lastPageId="3" lastPageNumber="548" pageId="2" pageNumber="547">
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Genitalia. Pygophore subquadrangular (
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<figureCitation id="1367F0B9FF931038FD71C672EEB4F8D3" box="[653,764,1806,1831]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="2" pageNumber="547">Figs. 4–7</figureCitation>
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). Posterolateral angles rounded, narrowed by the foliaceus projection of the inferior layer of ventral rim. Dorsal rim concave, with 1+1 expansions laterad to segment X (
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<figureCitation id="1367F0B9FF931038FA98C64EEC8CF89B" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="2" pageNumber="547">Figs. 4, 6</figureCitation>
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; dr, pa). Inferior layer of ventral rim with 1+1 foliaceus projection over genital cup, directed dorso-anteriorly, parallel to frontal plane of pygophore, their apex surface with a small spinose carina lateral to segment X (
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<figureCitation id="1367F0B9FF931038FABBC606ECEFF843" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="2" pageNumber="547">Figs. 4, 6</figureCitation>
|
||
; pil, ca); 1+1 subrectangular expansion, below the ventral opening of ventral rim, originated on the lateral margin of foliaceus projection of inferior layer of ventral rim (
|
||
<figureCitation id="1367F0B9FF931038FCE9C6BEEFCEF82F" box="[789,902,1986,2011]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="2" pageNumber="547">Figs. 4, 6</figureCitation>
|
||
; rp); outline of ventral opening of ventral rim projected and inferior layer of ventral rim with median conical projection, continued laterally and directed to posterolateral angles (
|
||
<figureCitation id="1367F0B9FF921039FE69C1C0EE41FF20" box="[405,521,188,213]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="3" pageNumber="548">Figs. 5, 7</figureCitation>
|
||
; cp). Median margin of ventral rim excavated, forming a rectangular opening where the parameres project over (
|
||
<figureCitation id="1367F0B9FF921039FDD9C1A3EEDDFF03" box="[549,661,223,248]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="3" pageNumber="548">Figs. 5, 7</figureCitation>
|
||
; vr, par); surface between ventral opening and posterolateral angles excavated. Segment X subtrapezoidal not exceeding the posterior limit of inferior layer of ventral rim; basal third membranous, apex narrowed and surrounded by fringe of bristles; lateral margin sinuous with 1+1 median tuft of bristles (
|
||
<figureCitation id="1367F0B9FF921039FF04C030ED2DFE91" box="[248,357,332,357]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="3" pageNumber="548">Figs. 4, 6</figureCitation>
|
||
; X); anal opening apical and genital opening ventral as longitudinal slit (
|
||
<figureCitation id="1367F0B9FF921039FB58C030E959FE90" box="[1188,1297,332,357]" captionStart="FIGURES 4 – 7" captionStartId="3.[151,264,1903,1926]" captionTargetBox="[168,1426,600,1871]" captionTargetId="figure@3.[150,1436,589,1882]" captionTargetPageId="3" captionText="FIGURES 4 – 7. Stapecolis, male genitalia. Stapecolis latus sp. nov.: 3, dorsal view; 4, ventral view; Stapecolis bimaculatus sp. nov.: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="3" pageNumber="548">Figs. 5, 7</figureCitation>
|
||
). Parameres ventral to segment X, projected over ventral opening of ventral rim, apex rounded with long bristles.
|
||
</paragraph>
|
||
<paragraph id="8BE3EC3CFF921039FF3BC0E8EF21FE59" blockId="3.[151,1437,151,536]" box="[199,873,404,429]" pageId="3" pageNumber="548">Female. Antennae: third segment as wide as the subsequent.</paragraph>
|
||
<paragraph id="8BE3EC3CFF921039FF3BC0CBE90AFDEC" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="548">
|
||
Genitalia. Gonocoxites 8 longer than wide, tumescent on disc (
|
||
<figureCitation id="1367F0B9FF921039FC79C0CBEF81FE24" box="[901,969,439,464]" captionStart="FIGURE 8" captionStartId="5.[151,250,1539,1561]" captionTargetBox="[162,1428,633,1511]" captionTargetId="figure@5.[151,1435,625,1518]" captionTargetPageId="5" captionText="FIGURE 8. Stapecolis latus sp. nov., female genitalia. Abbreviation: gc 8, gonocoxites 8; la 8, laterotergites 8; la 9, laterotergites 9; VII, sternite VII. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265426/files/figure.png" pageId="3" pageNumber="548">Fig. 8</figureCitation>
|
||
; gc8). Laterotergites 8 triangular, spiracle visible and posterior to lateral angle of gonocoxites 8 (
|
||
<figureCitation id="1367F0B9FF921039FD08C0A0EF7FFE00" box="[756,823,476,501]" captionStart="FIGURE 8" captionStartId="5.[151,250,1539,1561]" captionTargetBox="[162,1428,633,1511]" captionTargetId="figure@5.[151,1435,625,1518]" captionTargetPageId="5" captionText="FIGURE 8. Stapecolis latus sp. nov., female genitalia. Abbreviation: gc 8, gonocoxites 8; la 8, laterotergites 8; la 9, laterotergites 9; VII, sternite VII. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265426/files/figure.png" pageId="3" pageNumber="548">Fig. 8</figureCitation>
|
||
; la8). Laterotergites 9 digitiform, slightly depressed on middle; apex truncated, not surpassing laterotergites 8 (
|
||
<figureCitation id="1367F0B9FF921039FCFFC083EF00FDEC" box="[771,840,511,536]" captionStart="FIGURE 8" captionStartId="5.[151,250,1539,1561]" captionTargetBox="[162,1428,633,1511]" captionTargetId="figure@5.[151,1435,625,1518]" captionTargetPageId="5" captionText="FIGURE 8. Stapecolis latus sp. nov., female genitalia. Abbreviation: gc 8, gonocoxites 8; la 8, laterotergites 8; la 9, laterotergites 9; VII, sternite VII. Scale bar: 1 mm." httpUri="https://zenodo.org/record/265426/files/figure.png" pageId="3" pageNumber="548">Fig. 8</figureCitation>
|
||
; la9). Segment X covered by laterotergites 9.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF23BCB4FF921039FF6BC613ED48F816" httpUri="https://zenodo.org/record/265425/files/figure.png" pageId="3" pageNumber="548" targetBox="[168,1426,600,1871]" targetPageId="3">
|
||
<paragraph id="8BE3EC3CFF921039FF6BC613ED48F816" blockId="3.[151,1436,1903,2018]" pageId="3" pageNumber="548">
|
||
<emphasis id="B928302EFF921039FF6BC613ED72F871" bold="true" box="[151,314,1903,1926]" pageId="3" pageNumber="548">FIGURES 4–7.</emphasis>
|
||
<emphasis id="B928302EFF921039FEBCC60CEDE5F872" box="[320,429,1904,1926]" italics="true" pageId="3" pageNumber="548">
|
||
<taxonomicName id="4C5C97BFFF921039FEBCC60CEDE2F872" box="[320,426,1904,1926]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="551" phylum="Arthropoda" rank="genus" status="gen. nov.">Stapecolis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male genitalia.
|
||
<taxonomicName id="4C5C97BFFF921039FDB2C60CEEA4F872" box="[590,748,1904,1926]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
|
||
<emphasis id="B928302EFF921039FDB2C60CEEA4F872" box="[590,748,1904,1926]" italics="true" pageId="3" pageNumber="548">Stapecolis latus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B8D55FF921039FD0EC60CEF0AF871" box="[754,834,1904,1925]" pageId="3" pageNumber="548" rank="species">
|
||
<emphasis id="B928302EFF921039FD0EC60CEF0AF871" bold="true" box="[754,834,1904,1925]" pageId="3" pageNumber="548">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 3, dorsal view; 4, ventral view;
|
||
<taxonomicName id="4C5C97BFFF921039FB73C60CE930F872" box="[1167,1400,1904,1926]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="548" phylum="Arthropoda" rank="species" species="bimaculatus" status="sp. nov.">
|
||
<emphasis id="B928302EFF921039FB73C60CE930F872" box="[1167,1400,1904,1926]" italics="true" pageId="3" pageNumber="548">Stapecolis bimaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF921039FA81C60CEC8BF850" bold="true" pageId="3" pageNumber="548">
|
||
<taxonomicNameLabel id="A21B8D55FF921039FA81C60CEC8BF850" pageId="3" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 5, dorsal view; 6, ventral view. Abbreviations: ca, carina; cp, conical projection; dr, dorsal rim; pa, posterolateral angle; par, paramere; pil, projection of inferior layer of ventral rim; rp, rectangular projection; vr, ventral rim; X, segment X. Scale bar: 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C346BFB7FF95103EFF3BC1EBEE0CFF44" box="[199,580,151,176]" pageId="4" pageNumber="549" type="distribution">
|
||
<paragraph id="8BE3EC3CFF95103EFF3BC1EBEE0CFF44" blockId="4.[151,1437,151,861]" box="[199,580,151,176]" pageId="4" pageNumber="549">
|
||
<emphasis id="B928302EFF95103EFF3BC1EBED28FF44" bold="true" box="[199,352,151,176]" pageId="4" pageNumber="549">Distribution.</emphasis>
|
||
<collectingCountry id="F34BACACFF95103EFE9BC1E4ED8BFF5B" box="[359,451,152,175]" name="Panama" pageId="4" pageNumber="549">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34BACACFF95103EFE32C1EBEE08FF44" box="[462,576,151,176]" name="Colombia" pageId="4" pageNumber="549">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346BFB7FF95103EFF3BC1C0EEC1FC3E" pageId="4" pageNumber="549" type="discussion">
|
||
<paragraph id="8BE3EC3CFF95103EFF3BC1C0EFB1FE59" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="549">
|
||
<emphasis id="B928302EFF95103EFF3BC1C0ED07FF21" bold="true" box="[199,335,188,213]" pageId="4" pageNumber="549">Comments.</emphasis>
|
||
<taxonomicName id="4C5C97BFFF95103EFEA9C1C0ED8FFF20" box="[341,455,188,212]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B928302EFF95103EFEA9C1C0ED8FFF20" box="[341,455,188,212]" italics="true" pageId="4" pageNumber="549">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFE32C1C0EE7FFF21" bold="true" box="[462,567,188,213]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFE32C1C0EE7FFF21" box="[462,567,188,213]" pageId="4" pageNumber="549" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is proposed based on the clade H (
|
||
<figureCitation id="1367F0B9FF95103EFC40C1C0EFB4FF20" box="[956,1020,188,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1178,1200]" captionTargetBox="[160,1424,599,1149]" captionTargetId="figure@1.[151,1437,591,1156]" captionTargetPageId="1" captionText="FIGURE 1. Relationships of the clade F (Herrichela clade of Ochlerini), modified from Garbelotto et al. (2013). In red, Stapecolis gen. nov.; in blue its sister group." httpUri="https://zenodo.org/record/265423/files/figure.png" pageId="4" pageNumber="549">Fig. 1</figureCitation>
|
||
) of the phylogeny of the
|
||
<taxonomicName id="4C5C97BFFF95103EFAE5C1C0E9D4FF20" box="[1305,1436,188,212]" class="Insecta" family="Pentatomidae" genus="Herrichella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B928302EFF95103EFAE5C1C0E9D4FF20" box="[1305,1436,188,212]" italics="true" pageId="4" pageNumber="549">Herrichella</emphasis>
|
||
</taxonomicName>
|
||
clade (
|
||
<bibRefCitation id="EFCD91CDFF95103EFF1EC1A3EDA5FF0C" author="Garbelotto" box="[226,493,223,248]" pageId="4" pageNumber="549" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2013) Cladistics and revision of Alitocoris with considerations on the phylogeny of the Herrichella clade (Hemiptera, Pentatomidae, Discocephalinae, Ochlerini). Zoological Journal of the Linnean Society, 168, 452 - 472. http: // dx. doi. org / 10.1111 / zoj. 12032" type="journal article" year="2013">
|
||
Garbelotto
|
||
<emphasis id="B928302EFF95103EFE99C19DEDEFFF0C" box="[357,423,224,248]" italics="true" pageId="4" pageNumber="549">et. al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). The genus is supported by 10 synapomorphies, including both somatic and male genital morphology; the following six are highlighted because they are also diagnostic characters: the anterolateral margins of pronotum crenulated, and the pygophore structures—dorsal rim with 1+1 expansions laterad to segment X; apex of the foliaceus expansions of the inferior layer of ventral rim with a small spinose carina; the absence of lobular expansions on superior layer of ventral rim; presence of subrectangular expansions below the ventral opening of ventral rim; and the subrectangular ventral opening of pygophore.
|
||
</paragraph>
|
||
<paragraph id="8BE3EC3CFF95103EFF3BC0CBEE9EFD5C" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="549">
|
||
Although only the presence of the spinose carina apical on the foliaceus expansions is unique in Ochlerini, the closely related genera (
|
||
<figureCitation id="1367F0B9FF95103EFE57C0A0EDB8FE00" box="[427,496,476,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1178,1200]" captionTargetBox="[160,1424,599,1149]" captionTargetId="figure@1.[151,1437,591,1156]" captionTargetPageId="1" captionText="FIGURE 1. Relationships of the clade F (Herrichela clade of Ochlerini), modified from Garbelotto et al. (2013). In red, Stapecolis gen. nov.; in blue its sister group." httpUri="https://zenodo.org/record/265423/files/figure.png" pageId="4" pageNumber="549">Fig. 1</figureCitation>
|
||
) have different states for the characters listed above, e.g. dorsal rim with 2+2 expansions laterad to segment X, the presence of lobular expansions on superior layer of ventral rim, and the ventral opening of pygophore not subrectangular (varying in each genera—see
|
||
<bibRefCitation id="EFCD91CDFF95103EFBF5C358E8B3FDC9" author="Garbelotto" box="[1033,1275,548,573]" pageId="4" pageNumber="549" refString="Garbelotto, T. A., Campos, L. A. & Grazia, J. (2011) A new synonymy in Alitocoris Sailer (Hemiptera, Pentatomidae, Discocephalinae). Revista Brasileira de Entomologia, 55 (1), 138 - 140. http: // dx. doi. org / 10.1590 / S 0085 - 56262011000100023" type="journal article" year="2011">
|
||
Garbelotto
|
||
<emphasis id="B928302EFF95103EFB7BC359E8F7FDC8" box="[1159,1215,548,572]" italics="true" pageId="4" pageNumber="549">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
—figs 3–4, 7– 8; 2013—fig 9B; 2014—figs 5, 8). Besides the synapomorphies,
|
||
<taxonomicName id="4C5C97BFFF95103EFC96C334EF94FD94" box="[874,988,584,608]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B928302EFF95103EFC96C334EF94FD94" box="[874,988,584,608]" italics="true" pageId="4" pageNumber="549">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFC1FC33BE803FD94" bold="true" box="[995,1099,583,608]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFC1FC33BE803FD94" box="[995,1099,583,608]" pageId="4" pageNumber="549" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has other particularities in the female genitalia as the gonocoxites 8 with calluses on disc closer to its posterior margin and this projected over the laterotergites 9, and the yellow maculae on corium.
|
||
</paragraph>
|
||
<paragraph id="8BE3EC3CFF95103EFF3BC3C8E807FCA8" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="549">
|
||
The sister-group of
|
||
<taxonomicName id="4C5C97BFFF95103EFE55C3C8EE53FD38" box="[425,539,692,716]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B928302EFF95103EFE55C3C8EE53FD38" box="[425,539,692,716]" italics="true" pageId="4" pageNumber="549">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFDDFC3C8EEC6FD39" bold="true" box="[547,654,692,717]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFDDFC3C8EEC6FD39" box="[547,654,692,717]" pageId="4" pageNumber="549" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the clade I (
|
||
<figureCitation id="1367F0B9FF95103EFCC6C3C8EF34FD38" box="[826,892,692,717]" captionStart="FIGURE 1" captionStartId="1.[151,250,1178,1200]" captionTargetBox="[160,1424,599,1149]" captionTargetId="figure@1.[151,1437,591,1156]" captionTargetPageId="1" captionText="FIGURE 1. Relationships of the clade F (Herrichela clade of Ochlerini), modified from Garbelotto et al. (2013). In red, Stapecolis gen. nov.; in blue its sister group." httpUri="https://zenodo.org/record/265423/files/figure.png" pageId="4" pageNumber="549">Fig. 1</figureCitation>
|
||
) that includes
|
||
<taxonomicName id="4C5C97BFFF95103EFBD5C3C8E91AFD39" authority="Sailer" authorityName="Sailer" box="[1065,1362,692,717]" class="Insecta" family="Pentatomidae" genus="Alitocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="B928302EFF95103EFBD5C3C8E940FD38" box="[1065,1288,692,716]" italics="true" pageId="4" pageNumber="549">Alitocoris brunneus</emphasis>
|
||
Sailer
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B928302EFF95103EFA98C3C8ECB7FD04" italics="true" pageId="4" pageNumber="549">sedis mutabilis</emphasis>
|
||
) and other two new species (to be described elsewere). Species of clade I are supported by female characters: the sutural margins of gonocoxites 8 tumescent and its callosities projected to the lateral angles—two characters states absent in
|
||
<taxonomicName id="4C5C97BFFF95103EFE3AC25CEE70FCCC" box="[454,568,800,824]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B928302EFF95103EFE3AC25CEE70FCCC" box="[454,568,800,824]" italics="true" pageId="4" pageNumber="549">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFDBCC263EEE3FCCC" bold="true" box="[576,683,799,824]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFDBCC263EEE3FCCC" box="[576,683,799,824]" pageId="4" pageNumber="549" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. This sister-group relationship (
|
||
<figureCitation id="1367F0B9FF95103EFBEBC263E811FCCC" box="[1047,1113,799,824]" captionStart="FIGURE 1" captionStartId="1.[151,250,1178,1200]" captionTargetBox="[160,1424,599,1149]" captionTargetId="figure@1.[151,1437,591,1156]" captionTargetPageId="1" captionText="FIGURE 1. Relationships of the clade F (Herrichela clade of Ochlerini), modified from Garbelotto et al. (2013). In red, Stapecolis gen. nov.; in blue its sister group." httpUri="https://zenodo.org/record/265423/files/figure.png" pageId="4" pageNumber="549">Fig. 1</figureCitation>
|
||
) is supported by the second antennal segment shorter than the first and the presence of a carina on gonocoxites 9.
|
||
</paragraph>
|
||
<paragraph id="8BE3EC3CFF95103EFF6BC2CCEEC1FC3E" blockId="4.[151,649,943,970]" box="[151,649,943,970]" pageId="4" pageNumber="549">
|
||
<heading id="D0AB5B50FF95103EFF6BC2CCEEC1FC3E" bold="true" box="[151,649,943,970]" fontSize="11" level="1" pageId="4" pageNumber="549" reason="1">
|
||
<emphasis id="B928302EFF95103EFF6BC2CCEEC1FC3E" bold="true" box="[151,649,943,970]" pageId="4" pageNumber="549">
|
||
Key to the species of
|
||
<taxonomicName id="4C5C97BFFF95103EFE60C2D3EE5DFC3D" box="[412,533,943,969]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B928302EFF95103EFE60C2D3EE5DFC3D" bold="true" box="[412,533,943,969]" italics="true" pageId="4" pageNumber="549">Stapecolis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFDE0C2CCEEC1FC3E" box="[540,649,944,970]" pageId="4" pageNumber="549" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346BFB7FF95103EFF6BC289E9D4FB97" pageId="4" pageNumber="549" type="key">
|
||
<keyStep id="30A8F499FF95103EFF6BC289E9D4FB97" pageId="4" pageNumber="549">
|
||
<paragraph id="8BE3EC3CFF95103EFF6BC289E9D3FBDD" blockId="4.[151,1436,1013,1124]" pageId="4" pageNumber="549">
|
||
<keyLead id="30AD4F09FF95103EFF6BC289E9D3FBDD" pageId="4" pageNumber="549">
|
||
1 Yellow maculae at apex of radial vein of corium equal to ocellus diameter; scutellum with 1+1 spots on fovea.......................................................................................
|
||
<taxonomicName id="4C5C97BFFF95103EFB99C56FE8B3FBDD" box="[1125,1275,1043,1065]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
|
||
<emphasis id="B928302EFF95103EFB99C56FE8B3FBDD" box="[1125,1275,1043,1065]" italics="true" pageId="4" pageNumber="549">Stapecolis latus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFAFDC56FE905FBDD" bold="true" box="[1281,1357,1043,1065]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFAFDC56FE905FBDD" box="[1281,1357,1043,1065]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1367F0B9FF95103EFAA7C56FE9DDFBDD" box="[1371,1429,1043,1065]" captionStart="FIGURES 2 – 3. 1" captionStartId="2.[151,264,1101,1124]" captionTargetBox="[163,1432,197,1073]" captionTargetId="figure@2.[151,1436,193,1080]" captionTargetPageId="2" captionText="FIGURES 2 – 3. 1, Stapecolis latus sp. nov., male holotype; 2, Stapecolis bimaculatus sp. nov., male holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/265424/files/figure.png" pageId="4" pageNumber="549">Fig. 2</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE3EC3CFF95103EFF6BC54DE9D4FB97" blockId="4.[151,1436,1013,1124]" pageId="4" pageNumber="549">
|
||
<keyLead id="30AD4F09FF95103EFF6BC54DE9D4FB97" pageId="4" pageNumber="549">
|
||
- Yellow maculae at apex of radial vein of corium equal to eye diameter; scutellum without spots on fovea.....................................................................................
|
||
<taxonomicName id="4C5C97BFFF95103EFBE2C531E8B3FB97" box="[1054,1275,1101,1123]" class="Insecta" family="Pentatomidae" genus="Stapecolis" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="549" phylum="Arthropoda" rank="species" species="bimaculatus" status="sp. nov.">
|
||
<emphasis id="B928302EFF95103EFBE2C531E8B3FB97" box="[1054,1275,1101,1123]" italics="true" pageId="4" pageNumber="549">Stapecolis bimaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B928302EFF95103EFAFDC532E905FB90" bold="true" box="[1281,1357,1102,1124]" pageId="4" pageNumber="549">
|
||
<taxonomicNameLabel id="A21B8D55FF95103EFAFDC532E905FB90" box="[1281,1357,1102,1124]" pageId="4" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1367F0B9FF95103EFAA7C531E9DCFB97" box="[1371,1428,1101,1123]" captionStart="FIGURES 2 – 3. 1" captionStartId="2.[151,264,1101,1124]" captionTargetBox="[163,1432,197,1073]" captionTargetId="figure@2.[151,1436,193,1080]" captionTargetPageId="2" captionText="FIGURES 2 – 3. 1, Stapecolis latus sp. nov., male holotype; 2, Stapecolis bimaculatus sp. nov., male holotype. Scale bars: 1 mm." httpUri="https://zenodo.org/record/265424/files/figure.png" pageId="4" pageNumber="549">Fig. 3</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |