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<mods:title id="A5B6C00D36BFAD68AE4D0E2A6468C418">Cavonus Sharp, 1875 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini Pseudoryctina) in Western Australia with description of a new species and the female of Cavonus sculpturatus Blackburn, 1888</mods:title>
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<mods:namePart id="66893A394EA1CF284BB5606BE84C2928">Hutchinson, Paul M.</mods:namePart>
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<mods:affiliation id="8871EB695BEC295C3BFDC379C542EB9F">Department of Agriculture and Food WA (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia. paul. hutchinson @ dpird. wa. gov. au; https: // orcid. org / 0000 - 0001 - 8747 - 1320</mods:affiliation>
<mods:nameIdentifier id="95DC958741EC15C303AB596F2E643889" type="email">paul.hutchinson@dpird.wa.gov.au</mods:nameIdentifier>
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<mods:namePart id="28D8287B83E5A9C21A9A3BC95CAE2C3A">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="3D6225BD15D4293547FDA6827C08E139">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:date id="0D5700C7D4543170020924FB9E825E60">2020</mods:date>
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<paragraph id="8BE39D18FF8BAB74FF12F89DFDF8F88B" blockId="1.[151,629,1887,1950]" box="[151,629,1887,1914]" pageId="1" pageNumber="450">
<heading id="D0AB2A74FF8BAB74FF12F89DFDF8F88B" bold="true" box="[151,629,1887,1914]" fontSize="11" level="1" pageId="1" pageNumber="450" reason="1">
<taxonomicName id="4C5CE69BFF8BAB74FF12F89DFDF8F88B" authority="Blackburn, 1888" authorityName="Blackburn" authorityYear="1888" box="[151,629,1887,1914]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="450" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8BAB74FF12F89DFDF8F88B" bold="true" box="[151,629,1887,1914]" pageId="1" pageNumber="450">
<emphasis id="B928410AFF8BAB74FF12F89DFE13F88B" bold="true" box="[151,414,1887,1914]" italics="true" pageId="1" pageNumber="450">Cavonus sculpturatus</emphasis>
<bibRefCitation id="EFCDE0E9FF8BAB74FE23F89DFDF8F88B" author="Blackburn, T." box="[422,629,1888,1914]" pageId="1" pageNumber="450" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn, 1888</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BE39D18FF8BAB74FF12F879FEC7F86F" blockId="1.[151,629,1887,1950]" box="[151,330,1924,1950]" pageId="1" pageNumber="450">
(
<figureCitation id="1367819DFF8BAB74FF1AF879FE95F86F" box="[159,280,1924,1950]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 310" captionStart-2="FIGURES 1116" captionStartId-0="3.[151,264,1998,2022]" captionStartId-1="4.[151,264,1959,1983]" captionStartId-2="5.[151,264,1884,1908]" captionTargetBox-0="[167,1405,181,1934]" captionTargetBox-1="[182,1374,194,1935]" captionTargetBox-2="[151,1407,238,1861]" captionTargetId-0="figure-19@3.[157,1430,181,1973]" captionTargetId-1="figure-25@4.[182,1405,181,1935]" captionTargetId-2="figure-19@5.[151,1436,181,1861]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 12. Cavonus sculpturatus Blackburn, 1888, holotype male. 1, Dorsum; 2, labels." captionText-1="FIGURES 310. Cavonus sculpturatus Blackburn, 1888, holotype male. 3, Dorsum; 4, venter; 5, lateral; 6, head, lateral; 7, head, dorsal; 8, pygidium; 9, parameres, apices missing; 10, aedeagus, lateral." captionText-2="FIGURES 1116. Cavonus sculpturatus Blackburn, 1888, female (allotype of Epironastes nigrisetosus). 11, Dorsum; 12, venter; 13, lateral; 14, head lateral; 15, head, dorsal; 16, pygidium." figureDoi-0="http://doi.org/10.5281/zenodo.4410081" figureDoi-1="http://doi.org/10.5281/zenodo.4410083" figureDoi-2="http://doi.org/10.5281/zenodo.4410085" httpUri-0="https://zenodo.org/record/4410081/files/figure.png" httpUri-1="https://zenodo.org/record/4410083/files/figure.png" httpUri-2="https://zenodo.org/record/4410085/files/figure.png" pageId="1" pageNumber="450">Figs. 116</figureCitation>
,
<figureCitation id="1367819DFF8BAB74FEA0F879FECFF86F" box="[293,322,1924,1950]" captionStart="FIGURE 28" captionStartId="9.[151,250,1556,1580]" captionTargetBox="[151,1435,849,1533]" captionTargetId="figure-292@9.[151,1435,849,1533]" captionTargetPageId="9" captionText="FIGURE 28. Collection localities of Cavonus fovealacus new species and C. sculpturatus Blackburn, 1888 in Western Australia and South Australia." figureDoi="http://doi.org/10.5281/zenodo.4410091" httpUri="https://zenodo.org/record/4410091/files/figure.png" pageId="1" pageNumber="450">28</figureCitation>
)
</paragraph>
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<paragraph id="8BE39D18FF8BAB77FF12F834FD31FF1D" blockId="1.[151,1436,1992,2017]" lastBlockId="2.[199,1437,151,236]" lastPageId="2" lastPageNumber="451" pageId="1" pageNumber="450">
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<treatmentCitation id="0AFDBB09FF8BAB74FF12F834FDD6F810" author="Blackburn, T." box="[151,603,1992,2017]" page="223" pageId="1" pageNumber="450" year="1888">
<taxonomicName id="4C5CE69BFF8BAB74FF12F834FDD6F810" authority="Blackburn, 1888: 223" authorityName="Blackburn" authorityPageNumber="223" authorityYear="1888" box="[151,603,1992,2017]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="450" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8BAB74FF12F834FEF9F811" box="[151,372,1992,2016]" italics="true" pageId="1" pageNumber="450">Cavonus sculpturatus</emphasis>
<bibRefCitation id="EFCDE0E9FF8BAB74FEFEF835FDD6F810" author="Blackburn, T." box="[379,603,1992,2017]" pageId="1" pageNumber="450" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn, 1888: 223</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AFDBB09FF8BAB74FDE3F834FC9CF810" author="Carne, P. B." box="[614,785,1992,2017]" page="132" pageId="1" pageNumber="450" year="1957">
<bibRefCitation id="EFCDE0E9FF8BAB74FDE3F834FC9CF810" author="Carne, P. B." box="[614,785,1992,2017]" pageId="1" pageNumber="450" refId="ref6492" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957: 132</bibRefCitation>
</treatmentCitation>
, Fig. 335 (scutellum), Fig. 338 (head and pronotum), Figs. 345, 349 (parameres), Map 17; female (as
<taxonomicName id="4C5CE69BFF88AB77FDD0FF6AFCDDFF5F" authorityName="Carne" authorityYear="1957" box="[597,848,151,174]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="nigrisetosus">
<emphasis id="B928410AFF88AB77FDD0FF6AFCDDFF5F" box="[597,848,151,174]" italics="true" pageId="2" pageNumber="451">Epironastes nigrisetosus</emphasis>
</taxonomicName>
<typeStatus id="54E723BAFF88AB77FCDFFF6AFC3CFF5E" box="[858,945,151,175]" pageId="2" pageNumber="451" type="allotype">allotype</typeStatus>
): 126, Fig. 326 (head and pronotum), Map 17;
<treatmentCitation id="0AFDBB09FF88AB77FF42FF48FE2DFF3C" author="Endrodi, S." box="[199,416,181,205]" page="38" pageId="2" pageNumber="451" year="1974">
<bibRefCitation id="EFCDE0E9FF88AB77FF42FF48FE2DFF3C" author="Endrodi, S." box="[199,416,181,205]" pageId="2" pageNumber="451" pagination="17 - 70" refId="ref6646" refString="Endrodi, S. (1974) Monographie der Dynastinae 4. Tribus: Pentodontini von Australien und Neu-Zeeland (Coleoptera). Folia Entomologica Hungarica, Series Nova, 27, 17 - 70." type="journal article" year="1974">Endrődi 1974: 3839</bibRefCitation>
</treatmentCitation>
, Fig. 55 (parameres);
<treatmentCitation id="0AFDBB09FF88AB77FDF8FF48FCB1FF3C" author="Endrodi, S." box="[637,828,181,205]" page="386" pageId="2" pageNumber="451" year="1985">
<bibRefCitation id="EFCDE0E9FF88AB77FDF8FF48FCB1FF3C" author="Endrodi, S." box="[637,828,181,205]" pageId="2" pageNumber="451" refId="ref6682" refString="Endrodi, S. (1985) The Dynastinae of the World. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 386</bibRefCitation>
</treatmentCitation>
, Fig. 1402 (parameres);
<treatmentCitation id="0AFDBB09FF88AB77FBB6FF48FB6DFF3C" author="Krajcik, M." box="[1075,1248,181,205]" page="39" pageId="2" pageNumber="451" year="2005">
<bibRefCitation id="EFCDE0E9FF88AB77FBB6FF48FB6DFF3C" author="Krajcik, M." box="[1075,1248,181,205]" pageId="2" pageNumber="451" pagination="1 - 122" refId="ref6798" refString="Krajcik, M. (2005) Dynastinae of the World, checklist. Animma. X Supplement, 2, 1 - 122." type="journal article" year="2005">Krajcik 2005: 39</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFDBB09FF88AB77FB6FFF48FA15FF3C" author="Krajcik, M." box="[1258,1432,181,205]" page="66" pageId="2" pageNumber="451" year="2012">
<bibRefCitation id="EFCDE0E9FF88AB77FB6FFF48FA15FF3C" author="Krajcik, M." box="[1258,1432,181,205]" pageId="2" pageNumber="451" pagination="1 - 278" refId="ref6823" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. X Supplement, 5, 1 - 278." type="journal article" year="2012">Krajcik 2012: 66</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFDBB09FF88AB77FF42FF29FE13FF1D" author="Dechambre, R. - P." box="[199,414,212,236]" page="70" pageId="2" pageNumber="451" year="2005">
<bibRefCitation id="EFCDE0E9FF88AB77FF42FF29FE13FF1D" author="Dechambre, R. - P." box="[199,414,212,236]" pageId="2" pageNumber="451" refId="ref6583" refString="Dechambre, R. - P. (2005) Les Coleopteres du Monde. Vol. 30. Dynastidae Australiens et Oceaniens. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre 2005: 70</bibRefCitation>
</treatmentCitation>
, unnumbered image (adult).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C346CE93FF88AB77FF12FEE7FBFFFDFC" pageId="2" pageNumber="451" type="materials_examined">
<paragraph id="8BE39D18FF88AB77FF12FEE7FB5AFE8C" blockId="2.[151,1437,282,2037]" pageId="2" pageNumber="451">
<emphasis id="B928410AFF88AB77FF12FEE7FF54FEC5" bold="true" box="[151,217,282,308]" pageId="2" pageNumber="451">Type.</emphasis>
<materialsCitation id="3B349745FF88AB77FF64FEE7FB5AFE8C" ID-GBIF-Occurrence="3032720301" collectionCode="T, NHML" collectorName="Tate" country="Australia" latitude="-30.459" location="Ooldea" longLatPrecision="72" longitude="131.836" pageId="2" pageNumber="451" specimenCount="1" specimenCount-male="1" stateProvince="South Australia" typeStatus="holotype">
<typeStatus id="54E723BAFF88AB77FF64FEE7FEC4FEC5" box="[225,329,282,308]" pageId="2" pageNumber="451" type="holotype">Holotype</typeStatus>
<specimenCount id="9D5A5691FF88AB77FED7FEE6FEEAFEC4" box="[338,359,283,309]" pageId="2" pageNumber="451" type="male"></specimenCount>
(by monotypy):
<taxonomicName id="4C5CE69BFF88AB77FDADFEE7FC0CFEC5" authority=", Blackb." authorityName="Blackb." box="[552,897,282,308]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="sculpturatus">Cavonus sculpturatus, Blackb.</taxonomicName>
//
<collectingCountry id="F34BDD88FF88AB77FC1AFEE7FB8BFEC5" box="[927,1030,282,308]" name="Australia" pageId="2" pageNumber="451">Australia</collectingCountry>
<collectorName id="26A9F8CEFF88AB77FB8BFEE7FA88FEC4" box="[1038,1285,282,309]" pageId="2" pageNumber="451">Blackburn Coll. B.M.</collectorName>
1910236. // 1409 //
<collectorName id="26A9F8CEFF88AB77FF6FFEC2FEAFFEA8" box="[234,290,319,345]" pageId="2" pageNumber="451">Type</collectorName>
//
<collectorName id="26A9F8CEFF88AB77FEB9FEC2FE19FEA8" box="[316,404,319,345]" pageId="2" pageNumber="451">Type H.</collectorName>
<collectionCode id="ED4D05DDFF88AB77FE16FEC2FE2BFEA8" box="[403,422,319,345]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="451">T</collectionCode>
. // mentum on card // genitalia on card [in
<collectionCode id="ED4D05DDFF88AB77FCF3FEC2FC46FEA8" box="[886,971,319,345]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="2" pageNumber="451">NHML</collectionCode>
, examined] (
<figureCitation id="1367819DFF88AB77FBDCFEC3FB49FEA9" box="[1113,1220,318,344]" captionStart="FIGURES 12" captionStartId="3.[151,264,1998,2022]" captionTargetBox="[167,1405,181,1934]" captionTargetId="figure-19@3.[157,1430,181,1973]" captionTargetPageId="3" captionText="FIGURES 12. Cavonus sculpturatus Blackburn, 1888, holotype male. 1, Dorsum; 2, labels." figureDoi="http://doi.org/10.5281/zenodo.4410081" httpUri="https://zenodo.org/record/4410081/files/figure.png" pageId="2" pageNumber="451">Figs. 12</figureCitation>
).
<bibRefCitation id="EFCDE0E9FF88AB77FB52FEC3FA11FEA9" author="Blackburn, T." box="[1239,1436,318,344]" pageId="2" pageNumber="451" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn (1888)</bibRefCitation>
saw a single male collected by
<collectorName id="26A9F8CEFF88AB77FE76FE9EFDAEFE8C" box="[499,547,355,381]" pageId="2" pageNumber="451">Tate</collectorName>
at “Ouldea” [
<location id="8E83CBC3FF88AB77FD39FE9FFC9FFE8D" LSID="urn:lsid:plazi:treatment:03F52C0EFF8BAB73FF12F89DFAA9FD3C:8E83CBC3FF88AB77FD39FE9FFC9FFE8D" box="[700,786,354,380]" country="Australia" latitude="-30.459" longLatPrecision="72" longitude="131.836" name="Ooldea" pageId="2" pageNumber="451" stateProvince="South Australia">Ooldea</location>
,
<collectingRegion id="499853FAFF88AB77FC9BFE9FFC42FE8D" box="[798,975,354,380]" country="Australia" name="South Australia" pageId="2" pageNumber="451">South Australia</collectingRegion>
,
<geoCoordinate id="EE68FBDFFF88AB77FC5CFE9FFBC9FE8C" box="[985,1092,354,381]" degrees="30.459" direction="south" orientation="latitude" pageId="2" pageNumber="451" precision="55" value="-30.459">30.459°S</geoCoordinate>
,
<geoCoordinate id="EE68FBDFFF88AB77FBCAFE9FFB41FE8C" box="[1103,1228,354,381]" degrees="131.836" direction="east" orientation="longitude" pageId="2" pageNumber="451" precision="55" value="131.836">131.836°E</geoCoordinate>
].
</materialsCitation>
</paragraph>
<paragraph id="8BE39D18FF88AB77FF42FE7BFBFFFDFC" blockId="2.[151,1437,282,2037]" pageId="2" pageNumber="451">
<emphasis id="B928410AFF88AB77FF42FE7BFD98FE51" bold="true" box="[199,533,390,416]" pageId="2" pageNumber="451">Other specimens examined.</emphasis>
<materialsCitation id="3B349745FF88AB77FD9BFE7BFA15FE50" ID-GBIF-Occurrence="3032720304" box="[542,1432,390,417]" collectionCode="WAM" location="Forrest" pageId="2" pageNumber="451" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D5A5691FF88AB77FD9BFE7BFDC9FE51" box="[542,580,390,416]" pageId="2" pageNumber="451" type="male">1♂</specimenCount>
:
<taxonomicName id="4C5CE69BFF88AB77FDD6FE7BFC44FE51" authority="P. B. Carne" authorityName="P. B. Carne" box="[595,969,390,417]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="sculpturatus">Cavonus sculpturatus P.B. Carne</taxonomicName>
det. 1952 // 49-1101
<location id="8E83CBC3FF88AB77FB43FE7AFA98FE50" LSID="urn:lsid:plazi:treatment:03F52C0EFF8BAB73FF12F89DFAA9FD3C:8E83CBC3FF88AB77FB43FE7AFA98FE50" box="[1222,1301,391,417]" name="Forrest" pageId="2" pageNumber="451">Forrest</location>
[in
<collectionCode id="ED4D05DDFF88AB77FAC2FE7BFA1FFE50" box="[1351,1426,390,417]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="451" type="Museum">WAM</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="3B349745FF88AB77FF12FE57FE64FE18" ID-GBIF-Occurrence="3032720303" collectingDate="1962-10-28" collectionCode="WAM" collectorName="W. A. M. Reg. No &amp; A. Elliot" country="Australia" location="Forrest" pageId="2" pageNumber="451" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D5A5691FF88AB77FF12FE57FF30FE35" box="[151,189,426,452]" pageId="2" pageNumber="451" type="male">1♂</specimenCount>
:
<taxonomicName id="4C5CE69BFF88AB77FF4CFE57FDF0FE35" authority="Blkb. P. B. Carne" box="[201,637,426,453]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="sculpturatus">Cavonus sculpturatus Blkb. P.B. Carne</taxonomicName>
det. 1978 //
<collectorName id="26A9F8CEFF88AB77FC8FFE57FC44FE34" box="[778,969,426,453]" pageId="2" pageNumber="451">W.A.M. Reg. No</collectorName>
78/124 //
<location id="8E83CBC3FF88AB77FBB9FE56FB06FE34" LSID="urn:lsid:plazi:treatment:03F52C0EFF8BAB73FF12F89DFAA9FD3C:8E83CBC3FF88AB77FBB9FE56FB06FE34" box="[1084,1163,427,453]" country="Australia" name="Forrest" pageId="2" pageNumber="451">Forrest</location>
aerodrome W.
<collectingCountry id="F34BDD88FF88AB77FAB3FE57FA10FE35" box="[1334,1437,426,452]" name="Australia" pageId="2" pageNumber="451">Australia</collectingCountry>
<date id="FFE2BBD8FF88AB77FF12FE33FE8AFE19" box="[151,263,462,488]" pageId="2" pageNumber="451" value="1962-10-28">
<collectingDate id="EFA64230FF88AB77FF12FE33FE8AFE19" box="[151,263,462,488]" pageId="2" pageNumber="451" value="1962-10-28">28.x.1962</collectingDate>
</date>
<collectorName id="26A9F8CEFF88AB77FE89FE33FEE1FE19" box="[268,364,462,488]" pageId="2" pageNumber="451">A. Elliot</collectorName>
[in
<collectionCode id="ED4D05DDFF88AB77FE12FE33FE6FFE18" box="[407,482,462,489]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="451" type="Museum">WAM</collectionCode>
]
</materialsCitation>
;
<specimenCount id="9D5A5691FF88AB77FE71FE33FD95FE19" box="[500,536,462,488]" pageId="2" pageNumber="451" type="female">1♀</specimenCount>
: 49-1104 Forrest //?
<taxonomicName id="4C5CE69BFF88AB77FC83FE33FB98FE19" authorityName="Carne" authorityYear="1957" box="[774,1045,462,488]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="nigrisetosus">Epironastes nigrisetosus</taxonomicName>
mihi P.B. Carne comp. with type //
<materialsCitation id="3B349745FF88AB77FF12FE0FFBE3FDFC" ID-GBIF-Occurrence="3032720302" box="[151,1134,498,525]" collectionCode="WAM" collectorName="M. W. Woodie" country="Australia" pageId="2" pageNumber="451" specimenCount="1" typeStatus="allotype">
<typeStatus id="54E723BAFF88AB77FF12FE0FFF74FDFD" box="[151,249,498,524]" pageId="2" pageNumber="451" type="allotype">Allotype</typeStatus>
(see
<bibRefCitation id="EFCDE0E9FF88AB77FEB1FE0FFE7DFDFC" author="Carne, P. B." box="[308,496,498,525]" pageId="2" pageNumber="451" refId="ref6492" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne 1957: 126</bibRefCitation>
) [in
<collectionCode id="ED4D05DDFF88AB77FDA0FE0FFDE0FDFD" box="[549,621,498,524]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="451" type="Museum">WAM</collectionCode>
; register gives
<collectorName id="26A9F8CEFF88AB77FC90FE0EFC3DFDFD" box="[789,944,498,525]" pageId="2" pageNumber="451">M.W. Woodie</collectorName>
as the collector]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C346CE93FF88AB73FF42FDEBFCB7FEEC" lastPageId="6" lastPageNumber="455" pageId="2" pageNumber="451" type="description">
<paragraph id="8BE39D18FF88AB77FF42FDEBFB21FAF1" blockId="2.[151,1437,282,2037]" pageId="2" pageNumber="451">
<emphasis id="B928410AFF88AB77FF42FDEBFE35FDC1" bold="true" box="[199,440,534,560]" pageId="2" pageNumber="451">Redescription. Male</emphasis>
(
<figureCitation id="1367819DFF88AB77FE40FDEBFD9FFDC1" box="[453,530,534,560]" captionStart="FIGURES 12" captionStartId="3.[151,264,1998,2022]" captionTargetBox="[167,1405,181,1934]" captionTargetId="figure-19@3.[157,1430,181,1973]" captionTargetPageId="3" captionText="FIGURES 12. Cavonus sculpturatus Blackburn, 1888, holotype male. 1, Dorsum; 2, labels." figureDoi="http://doi.org/10.5281/zenodo.4410081" httpUri="https://zenodo.org/record/4410081/files/figure.png" pageId="2" pageNumber="451">Figs. 1</figureCitation>
,
<figureCitation id="1367819DFF88AB77FD9AFDEBFDD5FDC1" box="[543,600,534,560]" captionStart="FIGURES 310" captionStartId="4.[151,264,1959,1983]" captionTargetBox="[182,1374,194,1935]" captionTargetId="figure-25@4.[182,1405,181,1935]" captionTargetPageId="4" captionText="FIGURES 310. Cavonus sculpturatus Blackburn, 1888, holotype male. 3, Dorsum; 4, venter; 5, lateral; 6, head, lateral; 7, head, dorsal; 8, pygidium; 9, parameres, apices missing; 10, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410083" httpUri="https://zenodo.org/record/4410083/files/figure.png" pageId="2" pageNumber="451">310</figureCitation>
). Body black, with brown suffusion on meso- and metalegs and elytra,
<quantity id="4CA430FDFF88AB77FAEEFDEBFF4EFDA4" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="2" pageNumber="451" unit="mm" value="19.6">19.6 mm</quantity>
long,
<quantity id="4CA430FDFF88AB77FE88FDC7FEFCFDA4" box="[269,369,570,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="451" unit="mm" value="11.5">11.5 mm</quantity>
wide at elytra. Mentum setose on disc, compressed near ligula and reflexed; palps short with palpomere 4 cylindrical, basal two-thirds foveate; labrum concealed, medially bearing long yellow setae. Antennae with 10 antennomeres, scape and pedicel setose, club slightly shorter than shaft, ovate and setose on inner face. Clypeus transverse, dorsally subcircular in outline, margins strongly raised, truncate face moderately deep, slightly produced in middle of base and sparsely setose, clypeal surface heavily punctate; clypeofrontal ridge strongly elevated. Frons concentrically rugulose; ocular canthi at angle to clypeus, anterior margin straight and bearing setae, surface rugulose. Pronotum with sides slightly contracted before anterolateral angles, anterolateral angles not at right angles, anterior margin with strong erect process, with longitudinal ridge extending posteriorly into impression, latter defined by rounded lateral elevations, extending over basal convexity as a broad shallow depression, impression and anterior part of pronotum with large shallow incomplete often connected punctures, lateral and basal convexities with light sparse punctation, basal margin trisinuate with medial and paramedial shallow depressions. Scutellum with submarginal garland of punctures. Each elytron with sutural striae linear-punctate, disc with welldefined punctate striae, with occasional random punctures on intervals 2, 4, and 6, surface finely microreticulate, epipleurae with lateral setae extending as far as abdominal sternite 1. Pygidium moderately convex with long brown setae at sides, with shorter setae across base, rugose across basal half, becoming sparsely punctate towards apex, apical ridge very strongly broadened and setose. Postcoxal process tapering, densely setose. Posterior margins of all abdominal sternites setose across middle. Coxae bearing abundant dark-brown vestiture; protibiae broad, evenly and strongly toothed, protarsomere 5 longer than protarsomere 1; metafemora moderately stout, unisulcate; metatibiae stout, length 2x apical width, basal carina bearing long setae; medial carina with long blunt cilia and bristles, spurs subspatulate, smaller slightly curved, larger spur straight. Genitalia of
<typeStatus id="54E723BAFF88AB77FC68FB1BFBC2FAF1" box="[1005,1103,1254,1280]" pageId="2" pageNumber="451" type="holotype">holotype</typeStatus>
broken.
</paragraph>
<paragraph id="8BE39D18FF88AB73FF42FAF7FCB7FEEC" blockId="2.[151,1437,282,2037]" lastBlockId="6.[151,1437,150,717]" lastPageId="6" lastPageNumber="455" pageId="2" pageNumber="451">
<emphasis id="B928410AFF88AB77FF42FAF7FE93FAD5" bold="true" box="[199,286,1290,1316]" pageId="2" pageNumber="451">Female</emphasis>
(
<figureCitation id="1367819DFF88AB77FEA8FAF7FE3FFAD4" box="[301,434,1290,1317]" captionStart="FIGURES 1116" captionStartId="5.[151,264,1884,1908]" captionTargetBox="[151,1407,238,1861]" captionTargetId="figure-19@5.[151,1436,181,1861]" captionTargetPageId="5" captionText="FIGURES 1116. Cavonus sculpturatus Blackburn, 1888, female (allotype of Epironastes nigrisetosus). 11, Dorsum; 12, venter; 13, lateral; 14, head lateral; 15, head, dorsal; 16, pygidium." figureDoi="http://doi.org/10.5281/zenodo.4410085" httpUri="https://zenodo.org/record/4410085/files/figure.png" pageId="2" pageNumber="451">Figs. 1116</figureCitation>
). Body dark brown tending black,
<quantity id="4CA430FDFF88AB77FCBCFAF7FC10FAD4" box="[825,925,1290,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="2" pageNumber="451" unit="mm" value="16.7">16.7 mm</quantity>
long,
<quantity id="4CA430FDFF88AB77FC66FAF7FBB5FAD4" box="[995,1080,1290,1317]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="451" unit="mm" value="9.2">9.2 mm</quantity>
wide at elytra. Mentum wide at base, slightly narrowed to ligula, strongly reflexed, rounded at apex, disc setose; palps short [palpomere 4 of mandibular palp missing], labial palps with apical segments cylindrical; mandibles exposed, evenly rounded. Antennae with 10 antennomeres, scape and pedicel setose, club subovate, slightly shorter than shaft, inner face setose. Labrum concealed, bearing long yellow setae. Clypeus evenly arcuate, subcircular, apex not reflexed, lateral ridges distinct and slightly raised, truncated face not produced medially, sparely setose [setae broken], surface evenly coarsely punctate; clypeofrontal suture slightly raised, transverse-linear. Frons concentrically rugulose; ocular canthi rugulose, at angle to clypeus, anterior margin straight and bearing setae, lateral margin subequal to outline of eye. Pronotum with anterior margin membranous, anterolateral angles slightly acute, basolateral angles rounded, basal margin bisinuate with shallow paramedial depressions, basal ridge complete, widest at midlength, surface distinctly punctate becoming denser and larger at anterior and posterior margins, anterior margin medially with slight swelling preceding a flattened region bearing semi-annulate punctures tending rugulose, midline impunctate, surface slightly convex in lateral view, basal convexity absent. Scutellum bearing a garland of submarginal punctures, disc and base impunctate. Each elytron with sutural striae linear-punctate, disc with deeply impressed punctures forming distinct striae, intervals with random punctures, lateral epipleural setae absent. Pygidium moderately convex, disc concentrically punctate, denser and rugulose across base, finer and sparser at apex, bearing microsetae laterally, apical ridge slightly broadened and punctate medially, glabrous. Postcoxal prosternal process long, narrow, abundantly clothed with long dark brown vestiture, apex tapering posteriorly and glabrous. Abdominal sternites 15 punctate across posterior margin medially, laterally punctures bearing setae over entire surface, posterior margin of sternite 6 densely punctate bearing microsetae with sparse punctures across base. Protibiae stout, tridentate, denticles evenly spaced and broad [left protarsomeres 15 and right protarsomeres 14 missing]; metafemora stout, unisulcate; metatibiae moderately contracted before apex, length about 2.5x width, stout, apex broad and bearing short, fine cilia set about 1 width apart, bicarinate, basal carina bearing long setae, medial carina bearing long cilia and longer bristles, apical spurs stout, subspatulate, slightly curved; metatarsomere 1 moderately unilaterally expanded [left metatarsomeres 25 and right metatarsomeres 35 missing].
</paragraph>
</subSubSection>
<caption id="DF23CD90FF89AB76FF12F833FBC7F817" ID-DOI="http://doi.org/10.5281/zenodo.4410081" ID-Zenodo-Dep="4410081" box="[151,1098,1998,2022]" httpUri="https://zenodo.org/record/4410081/files/figure.png" pageId="3" pageNumber="452" startId="3.[151,264,1998,2022]" targetBox="[167,1405,181,1934]" targetPageId="3">
<paragraph id="8BE39D18FF89AB76FF12F833FBC7F817" blockId="3.[151,1098,1998,2022]" box="[151,1098,1998,2022]" pageId="3" pageNumber="452">
<emphasis id="B928410AFF89AB76FF12F833FEB7F817" bold="true" box="[151,314,1998,2022]" pageId="3" pageNumber="452">FIGURES 12.</emphasis>
<taxonomicName id="4C5CE69BFF89AB76FEC4F833FD42F817" authority="Blackburn, 1888" authorityName="Blackburn" authorityYear="1888" box="[321,719,1998,2022]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF89AB76FEC4F833FD90F817" box="[321,541,1998,2022]" italics="true" pageId="3" pageNumber="452">Cavonus sculpturatus</emphasis>
<bibRefCitation id="EFCDE0E9FF89AB76FDA6F833FD42F817" author="Blackburn, T." box="[547,719,1998,2022]" pageId="3" pageNumber="452" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn, 1888</bibRefCitation>
</taxonomicName>
, holotype male.
<emphasis id="B928410AFF89AB76FCF3F833FC0EF817" bold="true" box="[886,899,1998,2022]" pageId="3" pageNumber="452">1</emphasis>
, Dorsum;
<emphasis id="B928410AFF89AB76FC6BF833FC76F817" bold="true" box="[1006,1019,1998,2022]" pageId="3" pageNumber="452">2</emphasis>
, labels.
</paragraph>
</caption>
<caption id="DF23CD90FF8EAB71FF12F85AFC20F812" ID-DOI="http://doi.org/10.5281/zenodo.4410083" ID-Zenodo-Dep="4410083" httpUri="https://zenodo.org/record/4410083/files/figure.png" pageId="4" pageNumber="453" startId="4.[151,264,1959,1983]" targetBox="[182,1374,194,1935]" targetPageId="4">
<paragraph id="8BE39D18FF8EAB71FF12F85AFC20F812" blockId="4.[151,1436,1959,2019]" pageId="4" pageNumber="453">
<emphasis id="B928410AFF8EAB71FF12F85AFEC4F84E" bold="true" box="[151,329,1959,1983]" pageId="4" pageNumber="453">FIGURES 310.</emphasis>
<taxonomicName id="4C5CE69BFF8EAB71FED4F85AFD68F84E" authority="Blackburn, 1888" authorityName="Blackburn" authorityYear="1888" box="[337,741,1959,1983]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="453" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8EAB71FED4F85AFDA2F84E" box="[337,559,1959,1983]" italics="true" pageId="4" pageNumber="453">Cavonus sculpturatus</emphasis>
<bibRefCitation id="EFCDE0E9FF8EAB71FDBDF85AFD68F84E" author="Blackburn, T." box="[568,741,1959,1983]" pageId="4" pageNumber="453" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn, 1888</bibRefCitation>
</taxonomicName>
, holotype male.
<emphasis id="B928410AFF8EAB71FC17F85AFC12F84E" bold="true" box="[914,927,1959,1983]" pageId="4" pageNumber="453">3</emphasis>
, Dorsum;
<emphasis id="B928410AFF8EAB71FB8BF85AFB96F84E" bold="true" box="[1038,1051,1959,1983]" pageId="4" pageNumber="453">4</emphasis>
, venter;
<emphasis id="B928410AFF8EAB71FBF2F85AFB09F84E" bold="true" box="[1143,1156,1959,1983]" pageId="4" pageNumber="453">5</emphasis>
, lateral;
<emphasis id="B928410AFF8EAB71FB64F85AFB63F84E" bold="true" box="[1249,1262,1959,1983]" pageId="4" pageNumber="453">6</emphasis>
, head, lateral;
<emphasis id="B928410AFF8EAB71FA0CF85AFA1BF84E" bold="true" box="[1417,1430,1959,1983]" pageId="4" pageNumber="453">7</emphasis>
, head, dorsal;
<emphasis id="B928410AFF8EAB71FE9AF836FEA1F812" bold="true" box="[287,300,1995,2019]" pageId="4" pageNumber="453">8</emphasis>
, pygidium;
<emphasis id="B928410AFF8EAB71FE22F836FE39F812" bold="true" box="[423,436,1995,2019]" pageId="4" pageNumber="453">9</emphasis>
, parameres, apices missing;
<emphasis id="B928410AFF8EAB71FD52F836FD7DF812" bold="true" box="[727,752,1995,2019]" pageId="4" pageNumber="453">10</emphasis>
, aedeagus, lateral.
</paragraph>
</caption>
<caption id="DF23CD90FF8FAB70FF12F8A1FC9AF868" ID-DOI="http://doi.org/10.5281/zenodo.4410085" ID-Zenodo-Dep="4410085" httpUri="https://zenodo.org/record/4410085/files/figure.png" pageId="5" pageNumber="454" startId="5.[151,264,1884,1908]" targetBox="[151,1407,238,1861]" targetPageId="5">
<paragraph id="8BE39D18FF8FAB70FF12F8A1FC9AF868" blockId="5.[151,1436,1884,1945]" pageId="5" pageNumber="454">
<emphasis id="B928410AFF8FAB70FF12F8A1FEDEF884" bold="true" box="[151,339,1884,1909]" pageId="5" pageNumber="454">FIGURES 1116.</emphasis>
<taxonomicName id="4C5CE69BFF8FAB70FEDCF8A0FD6BF884" authority="Blackburn, 1888" authorityName="Blackburn" authorityYear="1888" box="[345,742,1885,1909]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8FAB70FEDCF8A0FDB8F884" box="[345,565,1885,1909]" italics="true" pageId="5" pageNumber="454">Cavonus sculpturatus</emphasis>
<bibRefCitation id="EFCDE0E9FF8FAB70FDBEF8A0FD6BF884" author="Blackburn, T." box="[571,742,1885,1909]" pageId="5" pageNumber="454" pagination="177 - 287" refId="ref6419" refString="Blackburn, T. (1888) Further notes on Australian Coleoptera, with descriptions of new species. Transactions of the Royal Society of South Australia, 10, 177 - 287." type="journal article" year="1888">Blackburn, 1888</bibRefCitation>
</taxonomicName>
, female (allotype of
<taxonomicName id="4C5CE69BFF8FAB70FC3DF8A0FB3DF885" authorityName="Carne" authorityYear="1957" box="[952,1200,1885,1908]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="454" phylum="Arthropoda" rank="species" species="nigrisetosus">
<emphasis id="B928410AFF8FAB70FC3DF8A0FB3DF885" box="[952,1200,1885,1908]" italics="true" pageId="5" pageNumber="454">Epironastes nigrisetosus</emphasis>
</taxonomicName>
).
<emphasis id="B928410AFF8FAB70FB43F8A0FB53F884" bold="true" box="[1222,1246,1885,1909]" pageId="5" pageNumber="454">11</emphasis>
, Dorsum;
<emphasis id="B928410AFF8FAB70FACCF8A0FAEFF884" bold="true" box="[1353,1378,1885,1909]" pageId="5" pageNumber="454">12</emphasis>
, venter;
<emphasis id="B928410AFF8FAB70FF3AF87CFF55F868" bold="true" box="[191,216,1921,1945]" pageId="5" pageNumber="454">13</emphasis>
, lateral;
<emphasis id="B928410AFF8FAB70FEB7F87CFEC6F868" bold="true" box="[306,331,1921,1945]" pageId="5" pageNumber="454">14</emphasis>
, head lateral;
<emphasis id="B928410AFF8FAB70FE5EF87CFE79F868" bold="true" box="[475,500,1921,1945]" pageId="5" pageNumber="454">15</emphasis>
, head, dorsal;
<emphasis id="B928410AFF8FAB70FD0CF87CFD2FF868" bold="true" box="[649,674,1921,1945]" pageId="5" pageNumber="454">16</emphasis>
, pygidium.
</paragraph>
</caption>
<subSubSection id="C346CE93FF8CAB73FF42FEDBFAA9FD3C" pageId="6" pageNumber="455" type="discussion">
<paragraph id="8BE39D18FF8CAB73FF42FEDBFEA4FDE8" blockId="6.[151,1437,150,717]" pageId="6" pageNumber="455">
<emphasis id="B928410AFF8CAB73FF42FEDBFEC1FEB1" bold="true" box="[199,332,294,320]" pageId="6" pageNumber="455">Discussion.</emphasis>
Known only from Forrest and Ooldea on the Nullarbor Plain (
<figureCitation id="1367819DFF8CAB73FB94FEDAFBEBFEB0" box="[1041,1126,295,321]" captionStart="FIGURE 28" captionStartId="9.[151,250,1556,1580]" captionTargetBox="[151,1435,849,1533]" captionTargetId="figure-292@9.[151,1435,849,1533]" captionTargetPageId="9" captionText="FIGURE 28. Collection localities of Cavonus fovealacus new species and C. sculpturatus Blackburn, 1888 in Western Australia and South Australia." figureDoi="http://doi.org/10.5281/zenodo.4410091" httpUri="https://zenodo.org/record/4410091/files/figure.png" pageId="6" pageNumber="455">Fig. 28</figureCitation>
). The female specimen that Carne attributed to
<taxonomicName id="4C5CE69BFF8CAB73FEF6FEB6FD97FE95" authorityName="Carne" authorityYear="1957" box="[371,538,331,356]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="nigrisetosus">
<emphasis id="B928410AFF8CAB73FEF6FEB6FD97FE95" box="[371,538,331,356]" italics="true" pageId="6" pageNumber="455">E. nigrisetosus</emphasis>
</taxonomicName>
was collected at Forrest by M.W. Woodie and, along with a male
<taxonomicName id="4C5CE69BFF8CAB73FA8DFEB6FF35FE79" authorityName="Blackburn" authorityYear="1888" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8CAB73FA8DFEB6FF35FE79" italics="true" pageId="6" pageNumber="455">C. sculpturatus</emphasis>
</taxonomicName>
and three other specimens, was donated to the WAM in one lot. The morphological combination of an arcuate clypeal margin and lack of a transverse ridge adjoining antennal angles anterior to clypeofrontal suture separates it from
<taxonomicName id="4C5CE69BFF8CAB73FF56FE4AFED5FE21" authorityName="Carne" authorityYear="1957" box="[211,344,439,464]" class="Insecta" family="Dynastidae" genus="Epironastes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="B928410AFF8CAB73FF56FE4AFED5FE21" box="[211,344,439,464]" italics="true" pageId="6" pageNumber="455">Epironastes</emphasis>
</taxonomicName>
and places it in
<emphasis id="B928410AFF8CAB73FD8CFE4AFDFFFE20" box="[521,626,439,465]" italics="true" pageId="6" pageNumber="455">
<taxonomicName id="4C5CE69BFF8CAB73FD8CFE4AFDE2FE20" authorityName="Sharp" authorityYear="1875" box="[521,623,439,465]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="genus">Cavonus</taxonomicName>
.
</emphasis>
Its collection with males of
<taxonomicName id="4C5CE69BFF8CAB73FC2CFE4AFBDFFE20" authorityName="Blackburn" authorityYear="1888" box="[937,1106,439,465]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8CAB73FC2CFE4AFBDFFE20" box="[937,1106,439,465]" italics="true" pageId="6" pageNumber="455">C. sculpturatus</emphasis>
</taxonomicName>
and sharing the characteristic garland of punctures on the scutellum, similar coloured vestiture and similar size show that it is the female of
<taxonomicName id="4C5CE69BFF8CAB73FA06FE26FEACFDE8" authorityName="Blackburn" authorityYear="1888" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B928410AFF8CAB73FA06FE26FEACFDE8" italics="true" pageId="6" pageNumber="455">C. sculpturatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE39D18FF8CAB73FF42FDDEFDC3FD58" blockId="6.[151,1437,150,717]" pageId="6" pageNumber="455">
Although the
<typeStatus id="54E723BAFF8CAB73FEE1FDDEFE4BFDCC" box="[356,454,547,573]" pageId="6" pageNumber="455" type="holotype">holotype</typeStatus>
has mandibular palpomere 4 missing,
<bibRefCitation id="EFCDE0E9FF8CAB73FC05FDDEFBA4FDCC" author="Carne, P. B." box="[896,1065,546,573]" pageId="6" pageNumber="455" refId="ref6492" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carnes (1957)</bibRefCitation>
generic description for
<taxonomicName id="4C5CE69BFF8CAB73FABFFDDEFA11FDCC" authorityName="Sharp" authorityYear="1875" box="[1338,1436,547,573]" class="Insecta" family="Dynastidae" genus="Cavonus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="B928410AFF8CAB73FABFFDDEFA11FDCC" box="[1338,1436,547,573]" italics="true" pageId="6" pageNumber="455">Cavonus</emphasis>
</taxonomicName>
states palps “cylindrical”. Both males examined from Forrest (WAM) have the mandibular palpomeres foveate on the basal two-thirds. The parameres from the
<typeStatus id="54E723BAFF8CAB73FD25FD96FC8FFD74" box="[672,770,619,645]" pageId="6" pageNumber="455" type="holotype">holotype</typeStatus>
are broken near the apices (
<figureCitation id="1367819DFF8CAB73FBC2FD96FB00FD74" box="[1095,1165,619,645]" captionStart="FIGURES 310" captionStartId="4.[151,264,1959,1983]" captionTargetBox="[182,1374,194,1935]" captionTargetId="figure-25@4.[182,1405,181,1935]" captionTargetPageId="4" captionText="FIGURES 310. Cavonus sculpturatus Blackburn, 1888, holotype male. 3, Dorsum; 4, venter; 5, lateral; 6, head, lateral; 7, head, dorsal; 8, pygidium; 9, parameres, apices missing; 10, aedeagus, lateral." figureDoi="http://doi.org/10.5281/zenodo.4410083" httpUri="https://zenodo.org/record/4410083/files/figure.png" pageId="6" pageNumber="455">Fig. 9</figureCitation>
); complete versions are shown in
<bibRefCitation id="EFCDE0E9FF8CAB73FE81FD72FE1FFD58" author="Carne, P. B." box="[260,402,654,681]" pageId="6" pageNumber="455" refId="ref6492" refString="Carne, P. B. (1957) A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957</bibRefCitation>
, Figs. 345, 349).
</paragraph>
<paragraph id="8BE39D18FF8CAB73FF42FD4EFAA9FD3C" blockId="6.[151,1437,150,717]" box="[199,1316,691,717]" pageId="6" pageNumber="455">We were intrigued with the double irony of the collector Woodie, at Forrest, on the Nullarbor Plain!</paragraph>
</subSubSection>
</treatment>
</document>