284 lines
45 KiB
XML
284 lines
45 KiB
XML
<document id="5931272E8F66D21DD65973B3A89ECDEB" ID-DOI="10.5281/zenodo.207632" ID-GBIF-Dataset="e7d0f1a5-538c-4099-8e16-501c6665ea59" ID-ISSN="1175-5326" ID-Zenodo-Dep="207632" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460307744023" checkinUser="plazi" docAuthor="Wilson, George D. F. & Reddy, Yenumula Ranga" docDate="2011" docId="03F50154FA26FFB3FF75FD4A7269FA0D" docLanguage="en" docName="zt02869p053.pdf" docOrigin="Zootaxa 2869" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Andhracoides Wilson & Reddy, 2011, gen. nov." docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="FFCC792CFA22FFB6FFE2FF877136FFCD" masterDocTitle="Andhracoides shabuddin gen. nov., sp. nov., a new phreatoicidean isopod (Crustacea, Hypsimetopidae) from hypogean aquatic habitats in Andhra Pradesh, India" masterLastPageNumber="53" masterPageNumber="37" pageNumber="41" updateTime="1698253599076" updateUser="plazi">
|
||
<mods:mods id="D5F659F5D2D8E7CEB6973530725DEC55" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="12A6E2BEF015288095F8A23F8B40296F">
|
||
<mods:title id="DF60E6DCCB0A309648614E133E1B3E15">Andhracoides shabuddin gen. nov., sp. nov., a new phreatoicidean isopod (Crustacea, Hypsimetopidae) from hypogean aquatic habitats in Andhra Pradesh, India</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7AE4058C609B327247056645F2725FF0" type="personal">
|
||
<mods:role id="22E2A80832268452E8CDC950BC7F0477">
|
||
<mods:roleTerm id="9EFD1D43D8FFFDDA83DF27ADCEA65501">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="049F4E5B3E82F5BB3E007BC6298E887B">Wilson, George D. F.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DB06A90920B69A018EC5B930876BA575" type="personal">
|
||
<mods:role id="980F75A8DE9A50021BDC6FDBC64D058D">
|
||
<mods:roleTerm id="27B22EB10290C2F8A4D2442A04765180">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D12C97A1A4904AC02925892E39009357">Reddy, Yenumula Ranga</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="564D41B8890A5E505090FE6652676A7A">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9A72E513D4A1D7C862C4AEC7CBAC63B0" type="host">
|
||
<mods:titleInfo id="780874F74827EA47C92432F2383BF540">
|
||
<mods:title id="ACB81E623B8ECBDF2A1524CD7DBEA5BE">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="ED92702AF9AA404CAB0766F752767245">
|
||
<mods:date id="45D2C44AB1FB6356CB4DFEBF1DEF9F26">2011</mods:date>
|
||
<mods:detail id="DC12A229F0D74BEF9757A39788FCBC3B" type="volume">
|
||
<mods:number id="1B1439E471BE6389A8F7D332E161CC77">2869</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="046948FA2A97972E86B92DC6F4A52E66" unit="page">
|
||
<mods:start id="24A0B0EBA228DA46797D1A132CDE19A6">37</mods:start>
|
||
<mods:end id="39CDC890FF19CD31DCD33536897D4395">53</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="1E89F94EC038BC0E43CFDE6B856083BB">journal article</mods:classification>
|
||
<mods:identifier id="F4BB8963B9E7945B895643371AC21EB6" type="DOI">10.5281/zenodo.207632</mods:identifier>
|
||
<mods:identifier id="DCD18CFBC7D1E2430814ACA9DCC0B49B" type="GBIF-Dataset">e7d0f1a5-538c-4099-8e16-501c6665ea59</mods:identifier>
|
||
<mods:identifier id="BF37D7BD9D6C60CCC3A072A78D8DAE1D" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="CF3170FB42D12A384179E673AF9869F7" type="Zenodo-Dep">207632</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03F50154FA26FFB3FF75FD4A7269FA0D" ID-DOI="http://doi.org/10.5281/zenodo.6191561" ID-GBIF-Taxon="119400530" ID-Zenodo-Dep="6191561" LSID="urn:lsid:zoobank.org:act:EABF8CB5-D391-404C-9BF0-87C17C46309A" httpUri="http://treatment.plazi.org/id/03F50154FA26FFB3FF75FD4A7269FA0D" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
|
||
<subSubSection id="C346E3C9FA26FFB2FF75FD4A7082FD2A" box="[151,436,717,743]" pageId="4" pageNumber="41" type="nomenclature">
|
||
<paragraph id="8BE3B042FA26FFB2FF75FD4A7082FD2A" blockId="4.[151,436,717,743]" box="[151,436,717,743]" pageId="4" pageNumber="41">
|
||
<heading id="D0AB072EFA26FFB2FF75FD4A7082FD2A" bold="true" box="[151,436,717,743]" fontSize="11" level="1" pageId="4" pageNumber="41" reason="1">
|
||
<emphasis id="B9286C50FA26FFB2FF75FD4A7082FD2A" bold="true" box="[151,436,717,743]" pageId="4" pageNumber="41">
|
||
<taxonomicName id="4C5CCBC1FA26FFB2FF75FD4A7009FD2A" ID-CoL="7NPRD" box="[151,319,717,743]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9286C50FA26FFB2FF75FD4A7009FD2A" bold="true" box="[151,319,717,743]" italics="true" pageId="4" pageNumber="41">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BD12BFA26FFB2FEA4FD4A7082FD2A" box="[326,436,717,743]" pageId="4" pageNumber="41" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346E3C9FA26FFB2FF75FC937542FCE1" box="[151,1140,787,812]" pageId="4" pageNumber="41" type="materials_examined">
|
||
<paragraph id="8BE3B042FA26FFB2FF75FC937542FCE1" blockId="4.[151,1437,787,2036]" box="[151,1140,787,812]" pageId="4" pageNumber="41">
|
||
<emphasis id="B9286C50FA26FFB2FF75FC937004FCE1" bold="true" box="[151,306,787,812]" pageId="4" pageNumber="41">
|
||
<typeStatus id="54E70EE0FA26FFB2FF75FC9371E4FCE1" box="[151,210,788,812]" pageId="4" pageNumber="41">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C5CCBC1FA26FFB2FEDBFC937366FCE1" box="[313,592,788,812]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="shabuddin" status="sp. nov.">
|
||
<emphasis id="B9286C50FA26FFB2FEDBFC937366FCE1" box="[313,592,788,812]" italics="true" pageId="4" pageNumber="41">Andhracoides shabuddin</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21BD12BFA26FFB2FDBCFC937381FCE1" box="[606,695,788,812]" pageId="4" pageNumber="41" rank="species">
|
||
<emphasis id="B9286C50FA26FFB2FDBCFC937381FCE1" bold="true" box="[606,695,788,812]" pageId="4" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, here designated. Currently monotypic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346E3C9FA26FFB2FF27FCBF709CFC71" pageId="4" pageNumber="41" type="etymology">
|
||
<paragraph id="8BE3B042FA26FFB2FF27FCBF709CFC71" blockId="4.[151,1437,787,2036]" pageId="4" pageNumber="41">
|
||
<emphasis id="B9286C50FA26FFB2FF27FCBF707DFC9D" bold="true" box="[197,331,824,848]" pageId="4" pageNumber="41">Etymology.</emphasis>
|
||
The prefix of the generic name, “Andhra” alludes to the Andhra Pradesh State in peninsular
|
||
<collectingCountry id="F34BF0D2FA26FFB2FAB9FCB074AEFC82" box="[1371,1432,823,847]" name="India" pageId="4" pageNumber="41">India</collectingCountry>
|
||
, where the new genus has been found. The Latin suffix for likeness “oides”, an adjective with one termination, with “c” retained, is taken from the hypsimetopid genus name
|
||
<taxonomicName id="4C5CCBC1FA26FFB2FCFAFC077565FC5A" authority="Sayce, 1900" authorityName="Sayce" authorityYear="1900" box="[792,1107,895,920]" class="Malacostraca" family="Hypsimetopidae" genus="Phreatoicoides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA26FFB2FCFAFC0772F7FC55" box="[792,961,896,920]" italics="true" pageId="4" pageNumber="41">Phreatoicoides</emphasis>
|
||
<bibRefCitation id="EFCDCDB3FA26FFB2FC25FCF87565FC5A" author="Sayce" box="[967,1107,895,919]" pageId="4" pageNumber="41" refString="Sayce, O. A. (1900) Phreatoicoides a new genus of freshwater Isopoda. Proceedings of the Royal Society of Victoria, 12, 122 - 138, pls. 110 - 112." type="journal article" year="1900">Sayce, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
. The gender of
|
||
<taxonomicName id="4C5CCBC1FA26FFB2FAE2FC0774AAFC55" box="[1280,1436,896,920]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA26FFB2FAE2FC0774AAFC55" box="[1280,1436,896,920]" italics="true" pageId="4" pageNumber="41">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
is considered masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346E3C9FA26FFB3FF27FC407233FEF2" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41" type="diagnosis">
|
||
<paragraph id="8BE3B042FA26FFB3FF27FC407233FEF2" blockId="4.[151,1437,787,2036]" lastBlockId="5.[151,1436,151,1473]" lastPageId="5" lastPageNumber="42" pageId="4" pageNumber="41">
|
||
<emphasis id="B9286C50FA26FFB2FF27FC407009FC2D" bold="true" box="[197,319,967,992]" pageId="4" pageNumber="41">Diagnosis.</emphasis>
|
||
<emphasis id="B9286C50FA26FFB2FEA8FC4F70B0FC2D" box="[330,390,968,992]" italics="true" pageId="4" pageNumber="41">Head</emphasis>
|
||
. Frontal process above antennula (
|
||
<figureCitation id="1367ACC7FA26FFB2FCC1FC40725FFC12" box="[803,873,967,991]" captionStart="FIGURE 3" captionStartId="7.[151,250,1429,1452]" captionTargetBox="[151,1436,193,1407]" captionTargetId="figure@7.[151,1436,193,1408]" captionTargetPageId="7" captionText="FIGURE 3. Andhracoides shabuddin gen. nov., sp. nov., head. Paratype male, AM P. 81107: A, head, lateral view; B, dorsal view; C, antennula and antenna; D, antennula with enlargements of distal articles; E, paratype female, AM P. 81108, head and pereopod I lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207635/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
A; dorsal margin of antennal notch) not curved.
|
||
<emphasis id="B9286C50FA26FFB2FF75FC6A71D1FBC9" box="[151,231,1005,1028]" italics="true" pageId="4" pageNumber="41">Pereon</emphasis>
|
||
pereonite 1 dorsal margin in lateral view shorter than on pereonite 2 (
|
||
<figureCitation id="1367ACC7FA26FFB2FBF6FC6B756CFBC9" box="[1044,1114,1004,1028]" captionStart="FIGURE 2" captionStartId="6.[151,250,1919,1942]" captionTargetBox="[151,1436,553,1898]" captionTargetId="figure@6.[151,1436,553,1898]" captionTargetPageId="6" captionText="FIGURE 2. Andhracoides shabuddin gen. nov., sp. nov. Holotype male, AM P. 81105: A. lateral view; B, dorsal view; C, head and pereonites 1 – 3; D, pleotelson; E, paratype female, AM P. 81106, lateral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/207634/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
|
||
A). Pereonites
|
||
<quantity id="4CA41DA7FA26FFB2FAE7FC6B7478FBC9" box="[1285,1358,1004,1028]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.778" metricValueMin="0.508" pageId="4" pageNumber="41" unit="in" value="4.5" valueMax="7.0" valueMin="2.0">2–7 in</quantity>
|
||
dorsal view anteriorly longer than wide, decreasing posteriorly to wider than long. Coxal articulation to pereonites 2–4 nearly fused, 5–7 free.
|
||
<emphasis id="B9286C50FA26FFB2FE7FFBB37325FB81" box="[413,531,1076,1100]" italics="true" pageId="4" pageNumber="41">Pleotelson</emphasis>
|
||
shallow (lateral fields sloping), lateral margin in dorsal view linear, broadening posteriorly (
|
||
<figureCitation id="1367ACC7FA26FFB2FEC2FBDF705EFBBD" box="[288,360,1112,1136]" captionStart="FIGURE 2" captionStartId="6.[151,250,1919,1942]" captionTargetBox="[151,1436,553,1898]" captionTargetId="figure@6.[151,1436,553,1898]" captionTargetPageId="6" captionText="FIGURE 2. Andhracoides shabuddin gen. nov., sp. nov. Holotype male, AM P. 81105: A. lateral view; B, dorsal view; C, head and pereonites 1 – 3; D, pleotelson; E, paratype female, AM P. 81106, lateral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/207634/files/figure.png" pageId="4" pageNumber="41">Figs 2</figureCitation>
|
||
D, 9A), dorsal surface in lateral view weakly curving, length substantially greater than width of uropodal insertion, lateral margin with fine setae only; posterolateral margin in lateral view uninterrupted, without distinct inflection differentiating apex, crenate, with 4 teeth, teeth rounded in cross-section (
|
||
<figureCitation id="1367ACC7FA26FFB2FB76FB2775EDFB75" box="[1172,1243,1184,1208]" captionStart="FIGURE 2" captionStartId="6.[151,250,1919,1942]" captionTargetBox="[151,1436,553,1898]" captionTargetId="figure@6.[151,1436,553,1898]" captionTargetPageId="6" captionText="FIGURE 2. Andhracoides shabuddin gen. nov., sp. nov. Holotype male, AM P. 81105: A. lateral view; B, dorsal view; C, head and pereonites 1 – 3; D, pleotelson; E, paratype female, AM P. 81106, lateral view. Scale bars = 5 mm." httpUri="https://zenodo.org/record/207634/files/figure.png" pageId="4" pageNumber="41">Figs 2</figureCitation>
|
||
D, 9D, G); posterior apex in dorsal view projecting posteriorly (not indented), dorsal margin of ventral anal ring projecting posteriorly (
|
||
<figureCitation id="1367ACC7FA26FFB2FF37FB6F702DFACD" box="[213,283,1256,1280]" captionStart="FIGURE 9" captionStartId="13.[151,250,1770,1793]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@13.[151,1436,193,1749]" captionTargetPageId="13" captionText="FIGURE 9. Andhracoides shabuddin gen. nov., sp. nov., pleotelson and uropods. Paratype male, AM P. 81107: A, pleotelson dorsal and lateral views; B, left uropod, dorsal and medial views; C, pleotelson distal margin and anus, ventral view; D, pleotelson distal margin and uropod protopod, dorsal view; E, distoventral margin of left uropod protopod. Paratype female, AM P. 81108: F, pleotelson, dorsal view; G, pleotelson distal margin and uropod protopod, dorsal view; H, pleotelson and right uropod, dorsolateral oblique view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207641/files/figure.png" pageId="4" pageNumber="41">Fig. 9</figureCitation>
|
||
C); dorsal surface without setose tubercles.
|
||
<emphasis id="B9286C50FA26FFB2FCEDFB6F752DFACD" box="[783,1051,1256,1280]" italics="true" pageId="4" pageNumber="41">Antennula and Antenna</emphasis>
|
||
elongate with numerous flagellar articles. Antennular distal articles (
|
||
<figureCitation id="1367ACC7FA26FFB2FDC0FA8B7352FAE9" box="[546,612,1292,1316]" captionStart="FIGURE 3" captionStartId="7.[151,250,1429,1452]" captionTargetBox="[151,1436,193,1407]" captionTargetId="figure@7.[151,1436,193,1408]" captionTargetPageId="7" captionText="FIGURE 3. Andhracoides shabuddin gen. nov., sp. nov., head. Paratype male, AM P. 81107: A, head, lateral view; B, dorsal view; C, antennula and antenna; D, antennula with enlargements of distal articles; E, paratype female, AM P. 81108, head and pereopod I lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207635/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
D) in cross-section circular; terminal article tubular, shorter than penultimate article; penultimate article not distinctly longer than other articles, width approximately subequal to antepenultimate article width. Antenna article 1 absent, article 5 longer than article 4, article 6 subequal to articles 4 and 5 combined (
|
||
<figureCitation id="1367ACC7FA26FFB2FEF5FAFF706DFA5D" box="[279,347,1400,1424]" captionStart="FIGURE 3" captionStartId="7.[151,250,1429,1452]" captionTargetBox="[151,1436,193,1407]" captionTargetId="figure@7.[151,1436,193,1408]" captionTargetPageId="7" captionText="FIGURE 3. Andhracoides shabuddin gen. nov., sp. nov., head. Paratype male, AM P. 81107: A, head, lateral view; B, dorsal view; C, antennula and antenna; D, antennula with enlargements of distal articles; E, paratype female, AM P. 81108, head and pereopod I lateral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207635/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
C, E).
|
||
<emphasis id="B9286C50FA26FFB2FE4BFAFF732BFA5D" box="[425,541,1400,1424]" italics="true" pageId="4" pageNumber="41">Mandible.</emphasis>
|
||
Palp article 3 medial surface naked (
|
||
<figureCitation id="1367ACC7FA26FFB2FC29FAFF7527FA5D" box="[971,1041,1400,1424]" captionStart="FIGURE 4" captionStartId="8.[151,250,1934,1957]" captionTargetBox="[151,1436,805,1912]" captionTargetId="figure@8.[151,1436,805,1913]" captionTargetPageId="8" captionText="FIGURE 4. Andhracoides shabuddin gen. nov., sp. nov. Paratype male, AM P. 81107. Mandible: A, left palp, dorsal view; B, right gnathal tip, medial view; C, left, medial view; D, left gnathal margin, dorsal view; E, right gnathal margin, ventral view; F, paragnath, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/207636/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
|
||
A). Right lacinia mobilis with two dentate plates (smaller plate on anterior surface of larger plate) (
|
||
<figureCitation id="1367ACC7FA26FFB2FC82FA1B7294FA79" box="[864,930,1436,1460]" captionStart="FIGURE 4" captionStartId="8.[151,250,1934,1957]" captionTargetBox="[151,1436,805,1912]" captionTargetId="figure@8.[151,1436,805,1913]" captionTargetPageId="8" captionText="FIGURE 4. Andhracoides shabuddin gen. nov., sp. nov. Paratype male, AM P. 81107. Mandible: A, left palp, dorsal view; B, right gnathal tip, medial view; C, left, medial view; D, left gnathal margin, dorsal view; E, right gnathal margin, ventral view; F, paragnath, ventral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/207636/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
|
||
E). Spine rows with bifurcate spines, forming strongly convex arc in ventral view, protruding medially, basal insertions curving dorsally to posteriorly (
|
||
<figureCitation id="1367ACC7FA26FFB2FADAFA47744CFA15" box="[1336,1402,1472,1496]" captionStart="FIGURE 9" captionStartId="13.[151,250,1770,1793]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@13.[151,1436,193,1749]" captionTargetPageId="13" captionText="FIGURE 9. Andhracoides shabuddin gen. nov., sp. nov., pleotelson and uropods. Paratype male, AM P. 81107: A, pleotelson dorsal and lateral views; B, left uropod, dorsal and medial views; C, pleotelson distal margin and anus, ventral view; D, pleotelson distal margin and uropod protopod, dorsal view; E, distoventral margin of left uropod protopod. Paratype female, AM P. 81108: F, pleotelson, dorsal view; G, pleotelson distal margin and uropod protopod, dorsal view; H, pleotelson and right uropod, dorsolateral oblique view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207641/files/figure.png" pageId="4" pageNumber="41">Fig. 9</figureCitation>
|
||
B– D). Molar process stout, heavily keratinised; complex setulate spines forming posterior row (
|
||
<figureCitation id="1367ACC7FA26FFB2FB4DFA6375C5FA31" box="[1199,1267,1508,1532]" captionStart="FIGURE 9" captionStartId="13.[151,250,1770,1793]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@13.[151,1436,193,1749]" captionTargetPageId="13" captionText="FIGURE 9. Andhracoides shabuddin gen. nov., sp. nov., pleotelson and uropods. Paratype male, AM P. 81107: A, pleotelson dorsal and lateral views; B, left uropod, dorsal and medial views; C, pleotelson distal margin and anus, ventral view; D, pleotelson distal margin and uropod protopod, dorsal view; E, distoventral margin of left uropod protopod. Paratype female, AM P. 81108: F, pleotelson, dorsal view; G, pleotelson distal margin and uropod protopod, dorsal view; H, pleotelson and right uropod, dorsolateral oblique view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207641/files/figure.png" pageId="4" pageNumber="41">Fig. 9</figureCitation>
|
||
D).
|
||
<emphasis id="B9286C50FA26FFB2FAFCFA6374ABFA31" box="[1310,1437,1508,1532]" italics="true" pageId="4" pageNumber="41">Pereopods.</emphasis>
|
||
Pereopod I not sexually dimorphic, male and female pereopods similar; dactylus without distal accessory spines (
|
||
<figureCitation id="1367ACC7FA26FFB2FF7DF9AB71D4F989" box="[159,226,1580,1604]" captionStart="FIGURE 6" captionStartId="10.[151,250,1942,1965]" captionTargetBox="[151,1436,409,1920]" captionTargetId="figure@10.[151,1436,409,1921]" captionTargetPageId="10" captionText="FIGURE 6. Andhracoides shabuddin gen. nov., sp. nov., anterior pereopods. A, paratype female, AM P. 81108, pereopod I, lateral view with enlargement of palm region, medial view. Paratype male, AM P. 81107: B, pereopod I, lateral view with enlargement of palm region; C – E, pereopods II – IV, lateral view with enlargements of dactylus and propodus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207638/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
|
||
A–B); propodus without developed palm, margin convex to straight; without spines or rugose cuticular pad, cuticular fringe weakly developed; with only stout conical simple setae; merus distodorsal margin in cross-section shelf-like and U-shaped. Pereopods II–III (
|
||
<figureCitation id="1367ACC7FA26FFB2FD95F9F3738EF941" box="[631,696,1652,1676]" captionStart="FIGURE 6" captionStartId="10.[151,250,1942,1965]" captionTargetBox="[151,1436,409,1920]" captionTargetId="figure@10.[151,1436,409,1921]" captionTargetPageId="10" captionText="FIGURE 6. Andhracoides shabuddin gen. nov., sp. nov., anterior pereopods. A, paratype female, AM P. 81108, pereopod I, lateral view with enlargement of palm region, medial view. Paratype male, AM P. 81107: B, pereopod I, lateral view with enlargement of palm region; C – E, pereopods II – IV, lateral view with enlargements of dactylus and propodus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207638/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
|
||
C–D) dactylus shorter than propodus, with 1 distal accessory claw and no additional distal spines; propodus without articular plate on posterior side of limb. Pereopods II–IV (
|
||
<figureCitation id="1367ACC7FA26FFB2FA8DF91F7192F919" captionStart="FIGURE 6" captionStartId="10.[151,250,1942,1965]" captionTargetBox="[151,1436,409,1920]" captionTargetId="figure@10.[151,1436,409,1921]" captionTargetPageId="10" captionText="FIGURE 6. Andhracoides shabuddin gen. nov., sp. nov., anterior pereopods. A, paratype female, AM P. 81108, pereopod I, lateral view with enlargement of palm region, medial view. Paratype male, AM P. 81107: B, pereopod I, lateral view with enlargement of palm region; C – E, pereopods II – IV, lateral view with enlargements of dactylus and propodus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207638/files/figure.png" pageId="4" pageNumber="41">Fig. 6</figureCitation>
|
||
C–E) basis lateral face ridge absent; ischium dorsal margin with simple setae, none robust. In male. Pereopod IV simple (not prehensile); propodus articular plate on posterior side of limb absent, basis dorsal ridge in cross-section angular and produced but not forming distinct plate. Pereopods V–VII (
|
||
<figureCitation id="1367ACC7FA26FFB2FC25F883753CF8D1" box="[967,1034,1796,1820]" captionStart="FIGURE 7" captionStartId="11.[151,250,1522,1545]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@11.[151,1436,193,1488]" captionTargetPageId="11" captionText="FIGURE 7. Andhracoides shabuddin gen. nov., sp. nov. Paratype male, AM P. 81107: A, right pereopod V, right lateral view with enlargement of propodus and dactylus; B, pereopod VII, lateral view with enlargement of propodus and dactylus; C, penes, right side, posteromedial view; D – E, appendix masculina, dorsal and ventral views. Scale bars: A – C = 1 mm; D – E = 0.2 mm." httpUri="https://zenodo.org/record/207639/files/figure.png" pageId="4" pageNumber="41">Fig. 7</figureCitation>
|
||
A–B) dactylus with accessory claw ventral to primary claw; propodus articular plate on posterior side of limb absent; basis dorsal ridge angular in cross-section, not distinctly separated from basis shaft, with no large setae; lateral face ventral ridge present, setae absent. Pereopod VII ischium dorsal ridge without flange.
|
||
<emphasis id="B9286C50FA26FFB2FCCAF8F772A6F845" box="[808,912,1904,1928]" italics="true" pageId="4" pageNumber="41">Pleopods</emphasis>
|
||
II–V (
|
||
<figureCitation id="1367ACC7FA26FFB2FC3EF8F7752AF845" box="[988,1052,1904,1928]" captionStart="FIGURE 8" captionStartId="12.[151,250,1985,2008]" captionTargetBox="[151,1436,462,1974]" captionTargetId="figure@12.[151,1436,462,1974]" captionTargetPageId="12" captionText="FIGURE 8. Andhracoides shabuddin gen. nov., sp. nov. Paratype male, AM P. 84983, pleopods, dorsal view. A, pleopod I; B – C, pleopod II, with enlargement of appendix masculina; D – E, pleopods III – V. Scale bar = 1 mm." httpUri="https://zenodo.org/record/207640/files/figure.png" pageId="4" pageNumber="41">Fig. 8</figureCitation>
|
||
B–F) exopod proximal article distolateral lobes shorter than distal article; exopod I lateral proximal lobe absent; endopods unilobed; without setae on margins; protopods medial margin coupling hooks absent. Pleopod II endopod appendix masculina (
|
||
<figureCitation id="1367ACC7FA26FFB2FAF9F83F7457F81D" box="[1307,1377,1976,2000]" captionStart="FIGURE 7" captionStartId="11.[151,250,1522,1545]" captionTargetBox="[151,1436,193,1486]" captionTargetId="figure@11.[151,1436,193,1488]" captionTargetPageId="11" captionText="FIGURE 7. Andhracoides shabuddin gen. nov., sp. nov. Paratype male, AM P. 81107: A, right pereopod V, right lateral view with enlargement of propodus and dactylus; B, pereopod VII, lateral view with enlargement of propodus and dactylus; C, penes, right side, posteromedial view; D – E, appendix masculina, dorsal and ventral views. Scale bars: A – C = 1 mm; D – E = 0.2 mm." httpUri="https://zenodo.org/record/207639/files/figure.png" pageId="4" pageNumber="41">Figs 7</figureCitation>
|
||
D–E, 8C) geniculate, projecting laterally at base of appendage; basal musculature pronounced; proximal half of shaft broadly concave in ventral cross-section, not forming tube; distal tip acutely rounded, margins smooth; medial and lateral margins with stiff elongate setae.
|
||
<emphasis id="B9286C50FA27FFB3FDB9FF3B7387FF19" box="[603,689,188,212]" italics="true" pageId="5" pageNumber="42">Uropod</emphasis>
|
||
(
|
||
<figureCitation id="1367ACC7FA27FFB3FD22FF3B7234FF19" box="[704,770,188,212]" captionStart="FIGURE 9" captionStartId="13.[151,250,1770,1793]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@13.[151,1436,193,1749]" captionTargetPageId="13" captionText="FIGURE 9. Andhracoides shabuddin gen. nov., sp. nov., pleotelson and uropods. Paratype male, AM P. 81107: A, pleotelson dorsal and lateral views; B, left uropod, dorsal and medial views; C, pleotelson distal margin and anus, ventral view; D, pleotelson distal margin and uropod protopod, dorsal view; E, distoventral margin of left uropod protopod. Paratype female, AM P. 81108: F, pleotelson, dorsal view; G, pleotelson distal margin and uropod protopod, dorsal view; H, pleotelson and right uropod, dorsolateral oblique view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207641/files/figure.png" pageId="5" pageNumber="42">Fig. 9</figureCitation>
|
||
A–B, F–H) protopod margin with rounded denticles, dorsomedial ridge in dorsal view parallel to ventral margin, robustly curving to oppose pleotelson posterior margin, distomedial margin without spinose or robust setae. Endopod subequal to protopod length, straight dorsally, dorsal margin without robust setae or spines on dorsal margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C346E3C9FA27FFB3FF27FECB7269FA0D" pageId="5" pageNumber="42" type="discussion">
|
||
<paragraph id="8BE3B042FA27FFB3FF27FECB7321FC91" blockId="5.[151,1436,151,1473]" pageId="5" pageNumber="42">
|
||
<emphasis id="B9286C50FA27FFB3FF27FECB700FFEA9" bold="true" box="[197,313,332,356]" pageId="5" pageNumber="42">Remarks.</emphasis>
|
||
This genus was originally thought to be part of the
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FC99FECB72D8FEA9" box="[891,1006,332,356]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FC99FECB72D8FEA9" box="[891,1006,332,356]" italics="true" pageId="5" pageNumber="42">Nichollsia</emphasis>
|
||
</taxonomicName>
|
||
clade, but its pleotelson is sufficiently divergent that we have assigned it to a new genus in the family
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FCB9FEE875F7FE4A" authority="Nicholls, 1943" authorityName="Nicholls" authorityYear="1943" box="[859,1217,367,391]" class="Malacostraca" family="Hypsimetopidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="family">
|
||
Hypsimetopidae
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FBFBFEE875F7FE4A" author="Nicholls" box="[1049,1217,367,391]" pageId="5" pageNumber="42" refString="Nicholls, G. E. (1943) The Phreatoicoidea. Part I. The Amphisopidae. Papers and Proceedings of the Royal Society of Tasmania, 1942, 1 - 145." type="book chapter" year="1943">Nicholls, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FB2EFEF77409FE45" box="[1228,1343,368,392]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FB2EFEF77409FE45" box="[1228,1343,368,392]" italics="true" pageId="5" pageNumber="42">Nichollsia</emphasis>
|
||
</taxonomicName>
|
||
has two similar described species
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FE5EFE127256FE61" authority="Chopra & Tiwari, 1950" authorityName="Chopra & Tiwari" authorityYear="1950" box="[444,864,404,428]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="kashiense">
|
||
<emphasis id="B9286C50FA27FFB3FE5EFE12737DFE61" box="[444,587,404,428]" italics="true" pageId="5" pageNumber="42">N. kashiense</emphasis>
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FDB6FE137256FE61" author="Chopra" box="[596,864,404,428]" pageId="5" pageNumber="42" refString="Chopra, B. & Tiwari, K. K. (1950) On a new genus of phreatoicid isopod from wells in Banaras. Records of the Indian Museum (Calcutta), 47, 277 - 289, pls. 17 - 20." type="journal article" year="1950">Chopra & Tiwari, 1950</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FC78FE12759CFE61" authority="Tiwari, 1955" authorityName="Tiwari" authorityYear="1955" box="[922,1194,404,428]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="menoni">
|
||
<emphasis id="B9286C50FA27FFB3FC78FE127539FE61" box="[922,1039,405,428]" italics="true" pageId="5" pageNumber="42">N. menoni</emphasis>
|
||
Tiwari, 1955
|
||
</taxonomicName>
|
||
(although
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FAC5FE1274AAFE61" box="[1319,1436,405,428]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="menoni">
|
||
<emphasis id="B9286C50FA27FFB3FAC5FE1274AAFE61" box="[1319,1436,405,428]" italics="true" pageId="5" pageNumber="42">N. menoni</emphasis>
|
||
</taxonomicName>
|
||
would benefit from a revision). Moreover, we have found that the morphology described above applies to several undescribed species of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FE44FE5B7374FE39" box="[422,578,476,500]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9286C50FA27FFB3FE44FE5B7374FE39" box="[422,578,476,500]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BD12BFA27FFB3FDAEFE5B738FFE39" box="[588,697,476,500]" pageId="5" pageNumber="42" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FDAEFE5B738FFE39" bold="true" box="[588,697,476,500]" pageId="5" pageNumber="42">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, found in Andhra Pradesh. The general pleotelson form of this genus, being flattened and broadening posteriorly with a well-developed terminal tip, is unlike any other hypsimetopid, which are either indented or at least transverse in dorsal view. Indeed,
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FBECFDA3759CFDF1" box="[1038,1194,548,572]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FBECFDA3759CFDF1" box="[1038,1194,548,572]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
is unique among the Phreatoicidea in having a pleotelson that is not strongly vaulted, and therefore much more like other isopods in being rather flattened. We do not think, however, that this condition is basal because the pleotelson is similar to that seen in other
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FED4FD0870C7FD6A" box="[310,497,655,679]" class="Malacostraca" family="Hypsimetopidae" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="family">Hypsimetopidae</taxonomicName>
|
||
; for example, the postanal margin, which is present in other phreatoicideans and members of other families, is missing, as is typical for other members of this family. The terminal projection of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FF75FD5F7005FD3D" box="[151,307,728,752]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FF75FD5F7005FD3D" box="[151,307,728,752]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
is an extension of the anal ring and, like many other hypsimetopids, has dentition along the pleotelson posterior margin. Moreover, the males of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FD44FD7B7274FCD9" box="[678,834,764,788]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FD44FD7B7274FCD9" box="[678,834,764,788]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
species lack the enormously elongate uropodal exopods possessed by
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FE96FCA770D1FCF5" box="[372,487,800,824]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FE96FCA770D1FCF5" box="[372,487,800,824]" italics="true" pageId="5" pageNumber="42">Nichollsia</emphasis>
|
||
</taxonomicName>
|
||
adult males. The exopod is reverse to this condition, with the male exopod being shorter than in the female (
|
||
<figureCitation id="1367ACC7FA27FFB3FE26FCC3733FFC91" box="[452,521,836,860]" captionStart="FIGURE 9" captionStartId="13.[151,250,1770,1793]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@13.[151,1436,193,1749]" captionTargetPageId="13" captionText="FIGURE 9. Andhracoides shabuddin gen. nov., sp. nov., pleotelson and uropods. Paratype male, AM P. 81107: A, pleotelson dorsal and lateral views; B, left uropod, dorsal and medial views; C, pleotelson distal margin and anus, ventral view; D, pleotelson distal margin and uropod protopod, dorsal view; E, distoventral margin of left uropod protopod. Paratype female, AM P. 81108: F, pleotelson, dorsal view; G, pleotelson distal margin and uropod protopod, dorsal view; H, pleotelson and right uropod, dorsolateral oblique view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/207641/files/figure.png" pageId="5" pageNumber="42">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE3B042FA27FFB3FF27FCEF7298FBB1" blockId="5.[151,1436,151,1473]" pageId="5" pageNumber="42">
|
||
A phylogenetic analysis of 65 phreatoicidean species and 421 characters (Wilson, in progress; data available on request; see
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FEC7FC0B708BFC69" author="Wilson" box="[293,445,908,932]" pageId="5" pageNumber="42" refString="Wilson, G. D. F. (2008) Gondwanan groundwater: subterranean connections of Australian phreatoicidean isopods (Crustacea) to India and New Zealand. Invertebrate Systematics, 22, 301 - 310." type="journal article" year="2008">Wilson, 2008</bibRefCitation>
|
||
for details) finds that
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FD5DFC0B726DFC69" box="[703,859,908,932]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9286C50FA27FFB3FD5DFC0B726DFC69" box="[703,859,908,932]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BD12BFA27FFB3FC81FC0B72F8FC69" box="[867,974,908,932]" pageId="5" pageNumber="42" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FC81FC0B72F8FC69" bold="true" box="[867,974,908,932]" pageId="5" pageNumber="42">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is sister to
|
||
<emphasis id="B9286C50FA27FFB3FBBEFC0B75E0FC69" box="[1116,1238,908,932]" italics="true" pageId="5" pageNumber="42">
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FBBEFC0B75E4FC69" box="[1116,1234,908,932]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Nichollsia</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
as expected, and this group forms a clade with the two species of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FD58FC377566FC0A" authority="Knott and Halse" authorityName="Knott and Halse" box="[698,1104,943,968]" class="Malacostraca" family="Hypsimetopidae" genus="Pilbarophreatoicus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FD58FC3772A4FC05" box="[698,914,944,968]" italics="true" pageId="5" pageNumber="42">Pilbarophreatoicus</emphasis>
|
||
Knott and Halse
|
||
</taxonomicName>
|
||
(one undescribed). Additionally, the hypsimetopid part of the cladogram is similar to that shown in
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FC2DFC5375F6FC21" author="Wilson" box="[975,1216,980,1004]" pageId="5" pageNumber="42" refString="Wilson, G. D. F. (2008) Gondwanan groundwater: subterranean connections of Australian phreatoicidean isopods (Crustacea) to India and New Zealand. Invertebrate Systematics, 22, 301 - 310." type="journal article" year="2008">Wilson (2008: fig. 4)</bibRefCitation>
|
||
but the position of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FF75FC7F707DFBDD" box="[151,331,1016,1040]" class="Malacostraca" family="Hypsimetopidae" genus="Hyperoedesipus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FF75FC7F707DFBDD" box="[151,331,1016,1040]" italics="true" pageId="5" pageNumber="42">Hyperoedesipus</emphasis>
|
||
</taxonomicName>
|
||
is moved to be sister to the
|
||
<emphasis id="B9286C50FA27FFB3FD40FC7F72CEFBDD" box="[674,1016,1016,1040]" italics="true" pageId="5" pageNumber="42">
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FD40FC7F727CFBDD" box="[674,842,1016,1040]" class="Malacostraca" family="Hypsimetopidae" genus="Phreatoicoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Phreatoicoides</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FCBBFC7E72CEFBDD" box="[857,1016,1017,1040]" class="Malacostraca" family="Hypsimetopidae" genus="Hypsimetopus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Hypsimetopus</taxonomicName>
|
||
</emphasis>
|
||
clade and the Pilbaran-Indian clade (
|
||
<emphasis id="B9286C50FA27FFB3FF42FB9B7390FBF9" box="[160,678,1052,1076]" italics="true" pageId="5" pageNumber="42">
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FF42FB9B7008FBF9" box="[160,318,1052,1076]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Andhracoides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FEA9FB9B70F7FBF9" box="[331,449,1052,1076]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Nichollsia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FE2CFB9B7390FBF9" box="[462,678,1052,1076]" class="Malacostraca" family="Hypsimetopidae" genus="Pilbarophreatoicus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">Pilbarophreatoicus</taxonomicName>
|
||
</emphasis>
|
||
) emerges basally in the hypsimetopid clade. These results support the biogeographic relationship between
|
||
<collectingCountry id="F34BF0D2FA27FFB3FDB8FBC773A5FB95" box="[602,659,1088,1112]" name="India" pageId="5" pageNumber="42">India</collectingCountry>
|
||
and Western
|
||
<collectingCountry id="F34BF0D2FA27FFB3FCCDFBC772AFFB95" box="[815,921,1088,1112]" name="Australia" pageId="5" pageNumber="42">Australia</collectingCountry>
|
||
, if based on tectonic vicariance, suggests that these lineages are older than 135 million years (
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FD4CFBE37297FBB1" author="Wilson" box="[686,929,1124,1148]" pageId="5" pageNumber="42" refString="Wilson, G. D. F. (2008) Gondwanan groundwater: subterranean connections of Australian phreatoicidean isopods (Crustacea) to India and New Zealand. Invertebrate Systematics, 22, 301 - 310." type="journal article" year="2008">Wilson, 2008, table 2</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE3B042FA27FFB3FF27FB0E7269FA0D" blockId="5.[151,1436,151,1473]" pageId="5" pageNumber="42">
|
||
In his introduction,
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FE46FB0F7367FB6D" author="Gupta" box="[420,593,1160,1184]" pageId="5" pageNumber="42" refString="Gupta, L. P. (1989) Monograph on Nichollsia kashiensis [sic] Chopra & Tiwari, 1950 (Crustacea: Isopoda: Phreatoicidea: Nichollsidae). Memoirs of the Zoological Survey of India, 17, 1 - 160." type="journal article" year="1989">Gupta (1989:1)</bibRefCitation>
|
||
writes "In a personal communication to Dr. K. K.Tiwari, Prof. P. J. Sanjeeva Raj of Madras Christian College, Tarnbaram (Madras), informs that he has material of phreatoicid isopods from Andhra Pradesh (locality not revealed) in South
|
||
<collectingCountry id="F34BF0D2FA27FFB3FD0DFB57721FFB25" box="[751,809,1232,1256]" name="India" pageId="5" pageNumber="42">India</collectingCountry>
|
||
obtained during deep drilling operations for boring tube wells." This is almost certainly the first report of
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FD2AFB737252FAC1" box="[712,868,1268,1292]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B9286C50FA27FFB3FD2AFB737252FAC1" box="[712,868,1268,1292]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21BD12BFA27FFB3FC8EFB7372E0FAC1" box="[876,982,1268,1292]" pageId="5" pageNumber="42" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FC8EFB7372E0FAC1" bold="true" box="[876,982,1268,1292]" pageId="5" pageNumber="42">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
but further details have not appeared in the published literature. A conference abstract (
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FD52FA9F726BFAFD" author="Messana" box="[688,861,1304,1328]" pageId="5" pageNumber="42" refString="Messana, G. (2004) A new Nichollsia (Isopoda, Phreatoicidea, Nichollsidae) from Indian caves. XVII International Symposium on Biospeleology, Raipur, India. Abstracts, p. 48." type="book chapter" year="2004">Messana, 2004</bibRefCitation>
|
||
) mentions a species from Andhra Pradesh referred to
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FF57FABB701EFA99" box="[181,296,1340,1364]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FF57FABB701EFA99" box="[181,296,1340,1364]" italics="true" pageId="5" pageNumber="42">Nichollsia</emphasis>
|
||
</taxonomicName>
|
||
; this species belongs to
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FDA0FABB73E8FA99" box="[578,734,1340,1364]" class="Malacostraca" family="Hypsimetopidae" genus="Andhracoides" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FDA0FABB73E8FA99" box="[578,734,1340,1364]" italics="true" pageId="5" pageNumber="42">Andhracoides</emphasis>
|
||
</taxonomicName>
|
||
but is a different species with a heavily setose pleotelson. Its locality is reported to be Belum Cave (Messana, pers. comm.).
|
||
<bibRefCitation id="EFCDCDB3FA27FFB3FC97FAE77538FAB5" author="Gupta" box="[885,1038,1376,1400]" pageId="5" pageNumber="42" refString="Gupta, L. P. (1989) Monograph on Nichollsia kashiensis [sic] Chopra & Tiwari, 1950 (Crustacea: Isopoda: Phreatoicidea: Nichollsidae). Memoirs of the Zoological Survey of India, 17, 1 - 160." type="journal article" year="1989">Gupta (1989)</bibRefCitation>
|
||
mentions that
|
||
<taxonomicName id="4C5CCBC1FA27FFB3FB59FAE77418FAB5" box="[1211,1326,1376,1400]" class="Malacostraca" family="Hypsimetopidae" genus="Nichollsia" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9286C50FA27FFB3FB59FAE77418FAB5" box="[1211,1326,1376,1400]" italics="true" pageId="5" pageNumber="42">Nichollsia</emphasis>
|
||
</taxonomicName>
|
||
has been recorded from a number of isolated localities in Uttar Pradesh and Bihar in the Gangetic Plains. These two genera appear to be isolated by major river systems in these two states.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |