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<document id="4CDBA168F6026FB053F0D08F8F7FBEE8" ID-CLB-Dataset="288482" ID-DOI="10.5252/zoosystema2024v46a3" ID-GBIF-Dataset="923830b3-43f8-4494-a94a-46be6cb195c1" ID-ISSN="1638-9387" ID-Zenodo-Dep="10672362" ID-ZooBank="1A21F175-68B4-4AB9-99F9-3277D428ED15" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1707899511480" checkinUser="plazi" docAuthor="Hwang, Chung-Chi &amp; Wu, Shu-Ping" docDate="2024" docId="03F4B022E34B5B47FC0C624C862359D7" docLanguage="en" docName="zoosystema.2024.46.3.pdf" docOrigin="Zoosystema 46 (3)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a3.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Satsuma nivnu Hwang &amp; Wu 2024, n. sp." docType="treatment" docUuid="EFF743A3-16B4-4648-BC40-D405E43840B5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="70" masterDocId="FFCDC85AE34C5B4CFFF7603A84715E3C" masterDocTitle="Three new species of Satsuma Adams, 1868 (Gastropoda, Camaenidae) from Taiwan" masterLastPageNumber="76" masterPageNumber="61" pageNumber="66" updateTime="1708115666168" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E5BE4D64EDC15C29AE4F319A5F04CC90" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="28CA8F82C5736B418A268747E9CE5806">
<mods:title id="D5BE899B4C4E48D8D8FDFC0D30B9014B">Three new species of Satsuma Adams, 1868 (Gastropoda, Camaenidae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="FD3228FCDA2DA7606626B7B9F7F2554B" type="personal">
<mods:role id="C9E6AB77992548DF2DECE0CAEF8B2425">
<mods:roleTerm id="20692CB734446526BD970D150220ADDC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81286B92F2DE9DF0777E61DAA510E7D6">Hwang, Chung-Chi</mods:namePart>
<mods:affiliation id="493E823A12034F010A9BA2F534054EB1">Department of Life Sciences, National University of Kaohsiung, No. 700, Kaohsiung University Road, Nan-Tzu District, Kaohsiung 81148 (Taiwan)</mods:affiliation>
<mods:nameIdentifier id="E694FF6FFFFD0D58294417DB17DBD162" type="email">cchwang@nuk.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:name id="D41DAA34F808960C8F157C80B7166399" type="personal">
<mods:role id="3B4B3C915E924283B2212B60F47EAEE5">
<mods:roleTerm id="E8038D6D28B654F80FEF4D434E8C2B12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D4AEF48B8BC71594E2F2228088BD9FB7">Wu, Shu-Ping</mods:namePart>
<mods:affiliation id="F42D2F2E2C17C58E316468C02603B1F6">Department of Earth and Life Sciences, University of Taipei, No. 1, Ai-Guo West Road, Taipei 10048 (Taiwan)</mods:affiliation>
<mods:nameIdentifier id="07CFAEFD0453E30B92C177A0E1DAAAE0" type="email">spwu@utaipei.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3F8F081B491D15BFBD51EDA9C3BD13FD">text</mods:typeOfResource>
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<mods:titleInfo id="83110E851143E9545C8A81DA4F821E10">
<mods:title id="57BA2DDB60C6A288F9F6DC05090945F2">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="C78AB12B5F9AD51F8C2F33FD9F346F73">
<mods:date id="9200E13D50262B78950FD7633FE1012C">2024</mods:date>
<mods:detail id="008549716C3460D5D80757E9B9F376C0" type="pubDate">
<mods:number id="C4C14AB800E7537F7D8286EB8FC08E9E">2024-02-13</mods:number>
</mods:detail>
<mods:detail id="1CC7CA10D4F65C6BA2E572B2C2F7E7BD" type="volume">
<mods:number id="95B7DCACE11C53A766E0003ABDFD703D">46</mods:number>
</mods:detail>
<mods:detail id="859AAF2DBCCE960F06048B6B7D0186F9" type="issue">
<mods:number id="DD9C689783AEB5F367C76F255F461B60">3</mods:number>
</mods:detail>
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<mods:start id="8C7E76C06B0B109DC8B0F8E987E51F7D">61</mods:start>
<mods:end id="51CA199BCE93833EEF85EA853620FD1C">76</mods:end>
</mods:extent>
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<mods:location id="835523D814B23A5B583710906736C345">
<mods:url id="A8A66FE07DD63C18C93385D0648D8F89">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a3.pdf</mods:url>
</mods:location>
<mods:classification id="3BD8E99046A664BDADF2F190DF375509">journal article</mods:classification>
<mods:identifier id="490546F622039D9A519EFB5266150759" type="CLB-Dataset">288482</mods:identifier>
<mods:identifier id="579EF722DFC8C9A104C435956941AB81" type="DOI">10.5252/zoosystema2024v46a3</mods:identifier>
<mods:identifier id="30F1ACE9E0F358D7647D1A40938615C0" type="GBIF-Dataset">923830b3-43f8-4494-a94a-46be6cb195c1</mods:identifier>
<mods:identifier id="6F7AFECFAC0ED8B695B7BA85D8E693D6" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="E2CBD077AEFFA50F2B9F732CB7712503" type="Zenodo-Dep">10672362</mods:identifier>
<mods:identifier id="BC229D4FFDA712189C584DB6CDEF2A0E" type="ZooBank">1A21F175-68B4-4AB9-99F9-3277D428ED15</mods:identifier>
</mods:mods>
<treatment id="03F4B022E34B5B47FC0C624C862359D7" ID-DOI="http://doi.org/10.5281/zenodo.10672374" ID-Zenodo-Dep="10672374" LSID="urn:lsid:zoobank.org:act:EFF743A3-16B4-4648-BC40-D405E43840B5" httpUri="http://treatment.plazi.org/id/03F4B022E34B5B47FC0C624C862359D7" lastPageId="11" lastPageNumber="70" pageId="7" pageNumber="66">
<heading id="D0AAB658E34B5B4BFC0C624C80C55C8C" centered="true" fontSize="11" level="2" pageId="7" pageNumber="66" reason="3">
<subSubSection id="C34752BFE34B5B4BFC0C624C80905CAC" box="[1019,1249,630,656]" pageId="7" pageNumber="66" type="nomenclature">
<paragraph id="8BE20134E34B5B4BFC0C624C80905CAC" blockId="7.[1019,1249,630,720]" box="[1019,1249,630,656]" pageId="7" pageNumber="66">
<taxonomicName id="4C5D7AB7E34B5B4BFC0C624C80EE5CAC" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[1019,1183,630,656]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="66" phylum="Mollusca" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B929DD26E34B5B4BFC0C624C80EE5CAC" bold="true" box="[1019,1183,630,656]" italics="true" pageId="7" pageNumber="66">Satsuma nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE34B5B4BFB51624C80905CAC" box="[1190,1249,630,656]" pageId="7" pageNumber="66" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B4BFBDF62A880C55C8C" box="[1064,1204,658,688]" pageId="7" pageNumber="66" type="multiple">
<paragraph id="8BE20134E34B5B4BFBDF62A880C55C8C" blockId="7.[1019,1249,630,720]" box="[1064,1204,658,688]" pageId="7" pageNumber="66">
<emphasis id="B929DD26E34B5B4BFBDF62A880C55C8C" bold="true" box="[1064,1204,658,688]" pageId="7" pageNumber="66">AE芙º栗ª</emphasis>
</paragraph>
</subSubSection>
</heading>
<subSubSection id="C34752BFE34B5B4BFBB1628F81E35D38" pageId="7" pageNumber="66" type="description">
<paragraph id="8BE20134E34B5B4BFBB1628F80E75CEC" blockId="7.[1019,1249,630,720]" box="[1094,1174,693,720]" pageId="7" pageNumber="66">
(
<figureCitation id="13661DB1E34B5B4BFBB9628F80FC5CEC" box="[1102,1165,693,720]" captionStart="FIG" captionStartId="10.[133,144,1986,2004]" captionTargetBox="[132,1455,216,1944]" captionTargetId="figure-78@10.[698,1267,764,1315]" captionTargetPageId="10" captionText="FIG. 5. — Shell and reproductive system of Satsuma nivnu n. sp.: A, shell of holotype (NMNS-8676-005); B, C, genitalia (paratype NMNS-8676-006):whole genitalia (B), opened genitalia (C); D, living animal. Scale bars: 10 mm. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10672376" httpUri="https://zenodo.org/record/10672376/files/figure.png" pageId="7" pageNumber="66">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="8BE20134E34B5B4BFCBE62C881E35D38" blockId="7.[841,1426,754,772]" box="[841,1426,754,772]" pageId="7" pageNumber="66">
<uri id="FFCC0D36E34B5B4BFCBE62C881E35D38" box="[841,1426,754,772]" pageId="7" pageNumber="66">
urn:lsid:zoobank.org:act:
<uuid id="FFFB3BE1E34B5B4BFBE662C881E35D38" box="[1041,1426,754,772]" pageId="7" pageNumber="66">EFF743A3-16B4-4648-BC40-D405E43840B5</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B4BFCDA631F87D85A79" pageId="7" pageNumber="66" type="materials_examined">
<paragraph id="8BE20134E34B5B4BFCDA631F80C45DB0" blockId="7.[813,1456,805,1094]" pageId="7" pageNumber="66">
<materialsCitation id="3B350B69E34B5B4BFCDA631F80C45DB0" collectingDate="2020-08-14" collectionCode="NMNS" collectorName="S. P. Wu" country="Taiwan" county="Jiayi County" elevation="1970" latitude="23.497778" location="Shitz Village" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" pageId="7" pageNumber="66" specimenCode="NMNS-8676-005" specimenCount="1" typeStatus="holotype">
<smallCapsWord id="8D0497E8E34B5B4BFCDA631F872E5D07" baselines="822,823" box="[813,863,805,828]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="66">TYPE</smallCapsWord>
<smallCapsWord id="8D0497E8E34B5B4BFC9F631087B85D07" baselines="823" box="[872,969,810,827]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="7" pageNumber="66">MATERIAL</smallCapsWord>
. —
<emphasis id="B929DD26E34B5B4BFC0C631F802C5D01" bold="true" box="[1019,1117,805,829]" pageId="7" pageNumber="66">
<typeStatus id="54E6BF96E34B5B4BFC0C631F802C5D01" box="[1019,1117,805,829]" pageId="7" pageNumber="66" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B929DD26E34B5B4BFB9B631C80C95D02" bold="true" box="[1132,1208,806,830]" pageId="7" pageNumber="66">
<collectingCountry id="F34A41A4E34B5B4BFB9B631C80C95D02" box="[1132,1208,806,830]" name="Taiwan" pageId="7" pageNumber="66">Taiwan</collectingCountry>
</emphasis>
•
<collectingCounty id="628379B8E34B5B4BFB24631F81275D01" box="[1235,1366,805,829]" pageId="7" pageNumber="66">Jiayi County</collectingCounty>
,
<collectingMunicipality id="6B869B4EE34B5B4BFA94631F87E55D6B" pageId="7" pageNumber="66">Alishan Township</collectingMunicipality>
,
<location id="8E8257EFE34B5B4BFC57630580555D6B" LSID="urn:lsid:plazi:treatment:03F4B022E34B5B47FC0C624C862359D7:8E8257EFE34B5B4BFC57630580555D6B" box="[928,1060,831,855]" country="Taiwan" county="Jiayi County" latitude="23.497778" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" name="Shitz Village" pageId="7" pageNumber="66" stateProvince="Chiayi">Shitz Village</location>
,
<location id="8E8257EFE34B5B4BFBC7630580815D6B" LSID="urn:lsid:plazi:treatment:03F4B022E34B5B47FC0C624C862359D7:8E8257EFE34B5B4BFBC7630580815D6B" box="[1072,1264,831,855]" country="Taiwan" county="Jiayi County" latitude="23.497778" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" name="Dadongshan forest" pageId="7" pageNumber="66" stateProvince="Chiayi">Dadongshan forest</location>
trail;
<geoCoordinate id="EE6967F3E34B5B4BFADA630581DA5D6B" box="[1325,1451,831,855]" degrees="23" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="66" precision="15" seconds="52" value="23.497778">23°29’52”N</geoCoordinate>
,
<geoCoordinate id="EE6967F3E34B5B4BFCDA636087D95D4E" box="[813,936,858,882]" degrees="120" direction="east" minutes="43" orientation="longitude" pageId="7" pageNumber="66" precision="15" seconds="11" value="120.71972">120°43’11”E</geoCoordinate>
; alt.
<elevation id="0070E607E34B5B4BFC226360803F5D4E" box="[981,1102,858,882]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="7" pageNumber="66" unit="m" value="1970.0">
<quantity id="4CA5ACD1E34B5B4BFC226360806E5D4D" box="[981,1055,858,882]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="7" pageNumber="66" unit="m" value="1970.0">1970 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE327F4E34B5B4BFBA0636080A85D4E" box="[1111,1241,858,882]" pageId="7" pageNumber="66" value="2020-08-14">
<collectingDate id="EFA7DE1CE34B5B4BFBA0636080A85D4E" box="[1111,1241,858,882]" pageId="7" pageNumber="66" value="2020-08-14">14.VIII.2020</collectingDate>
</date>
;
<collectorName id="26A864E2E34B5B4BFB166360815F5D4D" box="[1249,1326,858,882]" pageId="7" pageNumber="66">S. P. Wu</collectorName>
leg.;
<specimenCode id="DBFBA94FE34B5B4BFAAA636087E15DB0" pageId="7" pageNumber="66">NMNS-8676-005</specimenCode>
; SH
<quantity id="4CA5ACD1E34B5B4BFC36634E806F5DB0" box="[961,1054,884,908]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="7" pageNumber="66" unit="mm" value="31.8">31.8 mm</quantity>
, SW
<quantity id="4CA5ACD1E34B5B4BFBA4634E80C05DB0" box="[1107,1201,884,908]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="7" pageNumber="66" unit="mm" value="39.4">39.4 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE20134E34B5B4BFCDA63B587D85A79" blockId="7.[813,1456,805,1094]" pageId="7" pageNumber="66">
<materialsCitation id="3B350B69E34B5B4BFCDA63B587FF5DFD" collectionCode="NMNS" country="Taiwan" location="Taiwan" pageId="7" pageNumber="66" specimenCode="NMNS-8676-006" specimenCount="6" typeStatus="paratype">
<emphasis id="B929DD26E34B5B4BFCDA63B587FC5D9B" bold="true" box="[813,909,911,935]" pageId="7" pageNumber="66">
<typeStatus id="54E6BF96E34B5B4BFCDA63B587FC5D9B" box="[813,909,911,935]" pageId="7" pageNumber="66" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<emphasis id="B929DD26E34B5B4BFC6E63AA87935D94" bold="true" box="[921,994,912,936]" pageId="7" pageNumber="66">
<collectingCountry id="F34A41A4E34B5B4BFC6E63AA87935D94" box="[921,994,912,936]" name="Taiwan" pageId="7" pageNumber="66">Taiwan</collectingCountry>
</emphasis>
•
<specimenCount id="9D5BCABDE34B5B4BFC0F63B580005D9B" box="[1016,1137,911,935]" count="6" pageId="7" pageNumber="66" type="generic">6 specimens</specimenCount>
; same data as holotype;
<specimenCode id="DBFBA94FE34B5B4BFAAB63B587FF5DFD" pageId="7" pageNumber="66">NMNS-8676-006</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B350B69E34B5B4BFC526393816B5DE0" collectingDate="2014-08" collectionCode="NMNS" collectorName="C. F. Lin" country="Taiwan" county="Jiayi County" elevation="1970" latitude="23.497778" location="Shitz Village" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" pageId="7" pageNumber="66" specimenCode="NMNS-8676-007, MNHN-IM-2022-13918" specimenCount="2">
<specimenCount id="9D5BCABDE34B5B4BFC526393806E5DFD" box="[933,1055,937,961]" count="2" pageId="7" pageNumber="66" type="generic">2 specimens</specimenCount>
; from type locality;
<date id="FFE327F4E34B5B4BFB126393813D5DFD" box="[1253,1356,937,961]" pageId="7" pageNumber="66" value="2014-08">
<collectingDate id="EFA7DE1CE34B5B4BFB126393813D5DFD" box="[1253,1356,937,961]" pageId="7" pageNumber="66" value="2014-08">VIII.2014</collectingDate>
</date>
;
<collectorName id="26A864E2E34B5B4BFAA0639381DE5DFD" box="[1367,1455,937,961]" pageId="7" pageNumber="66">C. F. Lin</collectorName>
leg.;
<specimenCode id="DBFBA94FE34B5B4BFCAB63F980625DE7" box="[860,1043,963,987]" pageId="7" pageNumber="66">NMNS-8676-007</specimenCode>
,
<specimenCode id="DBFBA94FE34B5B4BFBEA63FE816B5DE0" box="[1053,1306,964,988]" pageId="7" pageNumber="66">MNHN-IM-2022-13918</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B350B69E34B5B4BFAC563FE87225A2D" collectingDate="2022-08-09" collectionCode="NMNS" collectorName="S. P. Wu" country="Taiwan" county="Jiayi County" elevation="1970" latitude="23.497778" location="Shitz Village" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" pageId="7" pageNumber="66" specimenCode="NMNS-8676-008- 009" specimenCount="2">
<specimenCount id="9D5BCABDE34B5B4BFAC563FE81DD5DE0" box="[1330,1452,964,988]" count="2" pageId="7" pageNumber="66" type="generic">2 specimens</specimenCount>
; from type locality;
<date id="FFE327F4E34B5B4BFC1063E480115DCA" box="[999,1120,990,1014]" pageId="7" pageNumber="66" value="2022-08-09">
<collectingDate id="EFA7DE1CE34B5B4BFC1063E480115DCA" box="[999,1120,990,1014]" pageId="7" pageNumber="66" value="2022-08-09">9.VIII.2022</collectingDate>
</date>
;
<collectorName id="26A864E2E34B5B4BFB9C63E480CC5DCA" box="[1131,1213,990,1014]" pageId="7" pageNumber="66">S. P. Wu</collectorName>
leg.;
<specimenCode id="DBFBA94FE34B5B4BFB0463E487225A2D" pageId="7" pageNumber="66">NMNS-8676-008- 009</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B350B69E34B5B4BFC9E63C387B85A17" collectingDate="2020-08-26" collectionCode="UTM" collectorName="S. P. Wu" country="Taiwan" county="Jiayi County" elevation="1970" latitude="23.497778" location="Shitz Village" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" pageId="7" pageNumber="66" specimenCode="UTM-2023002" specimenCount="1">
<specimenCount id="9D5BCABDE34B5B4BFC9E63C387AB5A2C" box="[873,986,1016,1041]" count="1" pageId="7" pageNumber="66" type="generic">1 specimen</specimenCount>
; from type locality;
<date id="FFE327F4E34B5B4BFB6963C281555A2C" box="[1182,1316,1016,1040]" pageId="7" pageNumber="66" value="2020-08-26">
<collectingDate id="EFA7DE1CE34B5B4BFB6963C281555A2C" box="[1182,1316,1016,1040]" pageId="7" pageNumber="66" value="2020-08-26">26.VIII.2020</collectingDate>
</date>
;
<collectorName id="26A864E2E34B5B4BFAD963C381F15A2C" box="[1326,1408,1017,1041]" pageId="7" pageNumber="66">S. P. Wu</collectorName>
leg.;
<specimenCode id="DBFBA94FE34B5B4BFCDA642987B85A17" box="[813,969,1043,1067]" pageId="7" pageNumber="66">UTM-2023002</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B350B69E34B5B4BFC13642987D45A79" collectingDate="2020-08-26" collectorName="S. P. Wu" country="Taiwan" county="Jiayi County" elevation="1970" latitude="23.497778" location="Shitz Village" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" pageId="7" pageNumber="66" specimenCount="1">
<specimenCount id="9D5BCABDE34B5B4BFC13642980295A17" box="[996,1112,1043,1067]" count="1" pageId="7" pageNumber="66" type="generic">1 specimen</specimenCount>
; from type locality;
<date id="FFE327F4E34B5B4BFAD3642981DA5A17" box="[1316,1451,1043,1067]" pageId="7" pageNumber="66" value="2020-08-26">
<collectingDate id="EFA7DE1CE34B5B4BFAD3642981DA5A17" box="[1316,1451,1043,1067]" pageId="7" pageNumber="66" value="2020-08-26">26.VIII.2020</collectingDate>
</date>
;
<collectorName id="26A864E2E34B5B4BFCD9641487F15A79" box="[814,896,1070,1094]" pageId="7" pageNumber="66">S. P. Wu</collectorName>
leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B4BFCDA646480705A95" pageId="7" pageNumber="66" type="etymology">
<paragraph id="8BE20134E34B5B4BFCDA646480705A95" blockId="7.[813,1457,1117,1194]" pageId="7" pageNumber="66">
<smallCapsWord id="8D0497E8E34B5B4BFCDA646487D95A48" baselines="1135,1136" box="[813,936,1118,1141]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="7" pageNumber="66">ETYMOLOGY</smallCapsWord>
. — Nivnu, treated as a simple noun in apposition, is the goddess of creation and agriculture in the mythology of the indigenous Tsou race.
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B4BFCDA64F8811B5B65" pageId="7" pageNumber="66" type="distribution">
<paragraph id="8BE20134E34B5B4BFCDA64F887A05B32" blockId="7.[812,1455,1218,1295]" pageId="7" pageNumber="66">
<smallCapsWord id="8D0497E8E34B5B4BFCDA64F8872C5AEB" baselines="1235,1235" box="[813,861,1218,1241]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="7" pageNumber="66">TYPE</smallCapsWord>
<smallCapsWord id="8D0497E8E34B5B4BFC9364FD87CE5AE4" baselines="1236" box="[868,959,1223,1240]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="7" pageNumber="66">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F34A41A4E34B5B4BFC1D64F880455AE6" box="[1002,1076,1218,1242]" name="Taiwan" pageId="7" pageNumber="66">Taiwan</collectingCountry>
,
<collectingCounty id="628379B8E34B5B4BFBCA64F880CB5AE6" box="[1085,1210,1218,1242]" pageId="7" pageNumber="66">Jiayi County</collectingCounty>
,
<collectingMunicipality id="6B869B4EE34B5B4BFB3464F881045AE6" box="[1219,1397,1218,1242]" pageId="7" pageNumber="66">Alishan Township</collectingMunicipality>
,
<location id="8E8257EFE34B5B4BFA8964F887095AC8" LSID="urn:lsid:plazi:treatment:03F4B022E34B5B47FC0C624C862359D7:8E8257EFE34B5B4BFA8964F887095AC8" country="Taiwan" county="Jiayi County" latitude="23.497778" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" name="Shitz Village" pageId="7" pageNumber="66">Shitz Village</location>
,
<location id="8E8257EFE34B5B4BFC7264E680F25AC8" LSID="urn:lsid:plazi:treatment:03F4B022E34B5B47FC0C624C862359D7:8E8257EFE34B5B4BFC7264E680F25AC8" box="[901,1155,1244,1268]" country="Taiwan" county="Jiayi County" latitude="23.497778" longLatPrecision="21" longitude="120.71972" municipality="Alishan Township" name="Dadongshan forest trail" pageId="7" pageNumber="66">Dadongshan forest trail</location>
;
<geoCoordinate id="EE6967F3E34B5B4BFB6764E681635AC8" box="[1168,1298,1244,1268]" degrees="23" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="66" precision="15" seconds="52" value="23.497778">23°29’52”N</geoCoordinate>
,
<geoCoordinate id="EE6967F3E34B5B4BFAD764E681D95AC8" box="[1312,1448,1244,1268]" degrees="120" direction="east" minutes="43" orientation="longitude" pageId="7" pageNumber="66" precision="15" seconds="11" value="120.71972">120°43’11”E</geoCoordinate>
; alt.
<quantity id="4CA5ACD1E34B5B4BFCA564CD87EE5B32" box="[850,927,1271,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="7" pageNumber="66" unit="m" value="1970.0">
<elevation id="0070E607E34B5B4BFCA564CD87EE5B32" box="[850,927,1271,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="7" pageNumber="66" unit="m" value="1970.0">1970 m</elevation>
</quantity>
a.s.l.
</paragraph>
<paragraph id="8BE20134E34B5B4BFCDA651D811B5B65" blockId="7.[813,1455,1318,1369]" pageId="7" pageNumber="66">
<smallCapsWord id="8D0497E8E34B5B4BFCDA651D87CB5B01" baselines="1336,1337" box="[813,954,1319,1342]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="7" pageNumber="66">DISTRIBUTION</smallCapsWord>
. — This species was found in
<typeStatus id="54E6BF96E34B5B4BFB13651D817F5B03" box="[1252,1294,1319,1343]" pageId="7" pageNumber="66">type</typeStatus>
locality, Alishan National Forest Recreation Area, and Tefuya Ancient Trail.
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B4BFCDA654B81C15830" pageId="7" pageNumber="66" type="diagnosis">
<paragraph id="8BE20134E34B5B4BFCDA654B81C15830" blockId="7.[811,1457,1392,1548]" pageId="7" pageNumber="66">
<smallCapsWord id="8D0497E8E34B5B4BFCDA654B87EF5BBB" baselines="1410,1411" box="[813,926,1393,1416]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="7" pageNumber="66">DIAGNOSIS</smallCapsWord>
. — Shell dextral, conical, medium to large sized (SW&gt;
<quantity id="4CA5ACD1E34B5B4BFC8465B087CC5B9F" box="[883,957,1418,1443]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="66" unit="mm" value="35.0">35 mm</quantity>
); spiral striation weakly malleated above and below periphery;; whorl not descending behind aperture; upper lip smoothly curved; columellar lip oblique, almost straight; basal lip curved; umbilicus partly open; penis with a muscular and bulging penial pouch; penial caecum conical and short; epiphallus swollen basally.
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE34B5B47FCDA660985935B31" lastPageId="11" lastPageNumber="70" pageId="7" pageNumber="66" type="description">
<paragraph id="8BE20134E34B5B4BFCDA660987CC5877" blockId="7.[811,1458,1587,2028]" box="[813,957,1587,1613]" pageId="7" pageNumber="66">
<smallCapsWord id="8D0497E8E34B5B4BFCDA660987CC5877" baselines="1606,1606" box="[813,957,1587,1613]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="7" pageNumber="66">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BE20134E34B5B47FCDA666886025F2E" blockId="7.[811,1458,1587,2028]" lastBlockId="11.[130,775,215,274]" lastPageId="11" lastPageNumber="70" pageId="7" pageNumber="66">
Shell dextral, with conical spire, brown to dark brown with red-brown peripheral band and umbilicus spot; thin and pale brown area visible immediately below suture and peripheral band. Apex acute.Whorls regularly increasing, slightly convex, not descending behind aperture. Periphery almost round, very weakly angulated. Base of shell convex. Surface covered with small, dense, elongate, axial granules on apical 3.5 whorls, becoming loose, coarse, continuous, oblique axial ridges that are weak on base (
<figureCitation id="13661DB1E34B5B4BFC1D676B804A5950" box="[1002,1083,1873,1900]" captionStart="FIG" captionStartId="13.[131,142,1986,2004]" captionTargetBox="[139,1455,216,1945]" captionTargetId="figure-103@13.[702,1268,760,1313]" captionTargetPageId="13" captionText="FIG. 7. — Shell sculptures on the 3rd whorl (A, B, C); surface of the last whorl (D, E, F); shell base (G, H, I): A, D, G, Satsuma hsuehshan n. sp.; B, E, H, Satsuma nivnu n. sp.; C, F, I, Satsuma sunlinksea n. sp. Scale bars: A-C, 0.5 mm; D-I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10672382" httpUri="https://zenodo.org/record/10672382/files/figure.png" pageId="7" pageNumber="66">Fig. 7B</figureCitation>
); spiral striation wavy and loose on base of shell, becoming weakly malleated above and below periphery. Aperture roundly lunate, light purplish inside. Peristome expanded; outer lip smoothly curved; columellar lip oblique, almost straight, joining curved basal lip in an angle. Parietal callus absent. Umbilicus partly open,
<emphasis id="B929DD26E3475B47FD9360E286045ECD" box="[612,629,216,241]" italics="true" pageId="11" pageNumber="70">c.</emphasis>
1/5 covered by reflected columellar lip, with 2-3 clear creases.
</paragraph>
<caption id="DF2251BCE3445B44FF7367FE86B159D0" ID-DOI="http://doi.org/10.5281/zenodo.10672370" ID-Zenodo-Dep="10672370" httpUri="https://zenodo.org/record/10672370/files/figure.png" pageId="8" pageNumber="67" startId="8.[132,143,1988,2006]" targetBox="[132,1455,215,1945]" targetPageId="8" targetType="figure">
<paragraph id="8BE20134E3445B44FF7367FE86B159D0" blockId="8.[132,1455,1987,2028]" pageId="8" pageNumber="67">
<smallCapsWord id="8D0497E8E3445B44FF7367FE84EC59E8" baselines="2001,2001" box="[132,157,1988,2006]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="8" pageNumber="67">FIG</smallCapsWord>
. 3. — Shell and reproductive system of
<taxonomicName id="4C5D7AB7E3445B44FE1A67F9868F59EA" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[493,766,1987,2006]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="67" phylum="Mollusca" rank="species" species="sericata">
<emphasis id="B929DD26E3445B44FE1A67F9860D59EA" box="[493,636,1987,2006]" italics="true" pageId="8" pageNumber="67">Satsuma sericata</emphasis>
(
<bibRefCitation id="EFCC7CC5E3445B44FD7167FE868859EA" author="KURODA T." box="[646,761,1988,2006]" pageId="8" pageNumber="67" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
:
<emphasis id="B929DD26E3445B44FCF167FE876259EA" bold="true" box="[774,787,1988,2006]" pageId="8" pageNumber="67">A</emphasis>
, shell (HCC-15562a);
<emphasis id="B929DD26E3445B44FC3C67FE87A959EA" bold="true" box="[971,984,1988,2006]" pageId="8" pageNumber="67">B</emphasis>
, whole genitalia (HCC-6921);
<emphasis id="B929DD26E3445B44FB2567F9809159E9" bold="true" box="[1234,1248,1987,2005]" pageId="8" pageNumber="67">C</emphasis>
, opened genitalia (HCC-6921). Scale bars: 10 mm. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF2251BCE3455B45FF736797860059D0" ID-DOI="http://doi.org/10.5281/zenodo.10672372" ID-Zenodo-Dep="10672372" httpUri="https://zenodo.org/record/10672372/files/figure.png" pageId="9" pageNumber="68" startId="9.[132,143,1965,1983]" targetBox="[132,1455,214,1922]" targetPageId="9" targetType="figure">
<paragraph id="8BE20134E3455B45FF736797860059D0" blockId="9.[132,1455,1962,2028]" pageId="9" pageNumber="68">
<smallCapsWord id="8D0497E8E3455B45FF73679784EC5982" baselines="1978,1979" box="[132,157,1965,1983]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="9" pageNumber="68">FIG</smallCapsWord>
. 4. — Shell and reproductive system of
<taxonomicName id="4C5D7AB7E3455B45FE186797862D5983" baseAuthorityName="Mollendorff" baseAuthorityYear="1888" box="[495,604,1965,1983]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="68" phylum="Mollusca" rank="species" species="nux">
<emphasis id="B929DD26E3455B45FE186797862D5983" box="[495,604,1965,1983]" italics="true" pageId="9" pageNumber="68">Satsuma nux</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC7CC5E3455B45FD926790868D5983" author="MOLLENDORFF O. F. &amp; VON" box="[613,764,1962,1983]" pageId="9" pageNumber="68" pagination="38 - 44" refId="ref9200" refString="MOLLENDORFF O. F. VON 1888. - Mittheilungen aus dem Gebiete der Malakozoologie. Diagnoses specierum novarum sinensium. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 20 (3 / 4): 38 - 44." type="journal article" year="1888">MÖllendorff, 1888</bibRefCitation>
):
<emphasis id="B929DD26E3455B45FCFE679787675983" bold="true" box="[777,790,1965,1983]" pageId="9" pageNumber="68">A</emphasis>
, shell (HCC-1925);
<emphasis id="B929DD26E3455B45FC4D679787B65983" bold="true" box="[954,967,1965,1983]" pageId="9" pageNumber="68">B</emphasis>
, whole genitalia (HCC-14121a);
<emphasis id="B929DD26E3455B45FB20679780945983" bold="true" box="[1239,1253,1965,1983]" pageId="9" pageNumber="68">C</emphasis>
, male genitalia showing penial caecum (HCC-1925);
<emphasis id="B929DD26E3455B45FE8667FE850E59EA" bold="true" box="[369,383,1988,2006]" pageId="9" pageNumber="68">D</emphasis>
, opened genitalia (HCC-14121a);
<emphasis id="B929DD26E3455B45FD6B67FE86D959EA" bold="true" box="[668,680,1988,2006]" pageId="9" pageNumber="68">E</emphasis>
, opened male genitalia showing verge (
<emphasis id="B929DD26E3455B45FC0467FE802F59EA" bold="true" box="[1011,1118,1988,2006]" pageId="9" pageNumber="68">red outlined</emphasis>
, HCC-1925). Scale bars: 10 mm, unless otherwise noted. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF2251BCE3465B46FF7267F8807B59D7" ID-DOI="http://doi.org/10.5281/zenodo.10672376" ID-Zenodo-Dep="10672376" httpUri="https://zenodo.org/record/10672376/files/figure.png" pageId="10" pageNumber="69" startId="10.[133,144,1986,2004]" targetBox="[132,1455,216,1944]" targetPageId="10" targetType="figure">
<paragraph id="8BE20134E3465B46FF7267F8807B59D7" blockId="10.[133,1456,1877,2027]" pageId="10" pageNumber="69">
<smallCapsWord id="8D0497E8E3465B46FF7267F884EC59EF" baselines="1999,2000" box="[133,157,1986,2004]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="69">FIG</smallCapsWord>
. 5. — Shell and reproductive system of
<taxonomicName id="4C5D7AB7E3465B46FE1F67F8861359E8" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[488,610,1986,2004]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="69" phylum="Mollusca" rank="species" species="nivnu" status="sp. nov.">
<emphasis id="B929DD26E3465B46FE1F67F8861359E8" box="[488,610,1986,2004]" italics="true" pageId="10" pageNumber="69">Satsuma nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3465B46FD9067F886E859E8" box="[615,665,1986,2004]" pageId="10" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="B929DD26E3465B46FD6867F886DD59E8" bold="true" box="[671,684,1986,2004]" pageId="10" pageNumber="69">A</emphasis>
, shell of holotype (NMNS-8676-005);
<emphasis id="B929DD26E3465B46FC1267F8878359E8" bold="true" box="[997,1010,1986,2004]" pageId="10" pageNumber="69">B</emphasis>
,
<emphasis id="B929DD26E3465B46FC0B67F8807B59E8" bold="true" box="[1020,1034,1986,2004]" pageId="10" pageNumber="69">C</emphasis>
, genitalia (paratype NMNS-8676-006):whole genitalia (
<emphasis id="B929DD26E3465B46FF4667E384CF59D7" bold="true" box="[177,190,2009,2027]" pageId="10" pageNumber="69">B</emphasis>
), opened genitalia (
<emphasis id="B929DD26E3465B46FE9667E3851E59D7" bold="true" box="[353,367,2009,2027]" pageId="10" pageNumber="69">C</emphasis>
);
<emphasis id="B929DD26E3465B46FE8967E385FD59D7" bold="true" box="[382,396,2009,2027]" pageId="10" pageNumber="69">D</emphasis>
, living animal. Scale bars: 10 mm. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BE20134E3475B47FF73610285645F6D" blockId="11.[132,775,312,433]" box="[132,277,312,337]" pageId="11" pageNumber="70">
<emphasis id="B929DD26E3475B47FF73610285645F6D" box="[132,277,312,337]" italics="true" pageId="11" pageNumber="70">Measurements</emphasis>
</paragraph>
<paragraph id="8BE20134E3475B47FF73616D85BE5F8D" blockId="11.[132,775,312,433]" pageId="11" pageNumber="70">
SH
<quantity id="4CA5ACD1E3475B47FF58616D853E5F4D" box="[175,335,343,369]" metricMagnitude="-2" metricUnit="m" metricValue="3.09" metricValueMax="3.29" metricValueMin="2.8899999999999997" pageId="11" pageNumber="70" unit="mm" value="30.9" valueMax="32.9" valueMin="28.9">28.9-32.9 mm</quantity>
, SW
<quantity id="4CA5ACD1E3475B47FE7E616D86585F4D" box="[393,553,343,369]" metricMagnitude="-2" metricUnit="m" metricValue="3.945" metricValueMax="4.26" metricValueMin="3.63" pageId="11" pageNumber="70" unit="mm" value="39.45" valueMax="42.6" valueMin="36.3">36.3-42.6 mm</quantity>
, AH
<quantity id="4CA5ACD1E3475B47FD95616D87725F4D" box="[610,771,343,370]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" metricValueMax="2.03" metricValueMin="1.75" pageId="11" pageNumber="70" unit="mm" value="18.9" valueMax="20.3" valueMin="17.5">17.5-20.3 mm</quantity>
, AW 23.0-
<quantity id="4CA5ACD1E3475B47FF18614D85275FAD" box="[239,342,375,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="11" pageNumber="70" unit="mm" value="26.8">26.8 mm</quantity>
, W# 6.25-6.75, protoconch 2.5 whorls, SH/SW
<geoCoordinate id="EE6967F3E3475B47FF1661AC85505F8C" box="[225,289,406,432]" degrees="0.726" direction="north" orientation="latitude" pageId="11" pageNumber="70" precision="55" value="0.726">0.726</geoCoordinate>
<geoCoordinate id="EE6967F3E3475B47FED661AC85195F8C" box="[289,360,406,432]" degrees="0.822" direction="west" orientation="longitude" pageId="11" pageNumber="70" precision="55" value="-0.822">-0.822</geoCoordinate>
(n = 12).
</paragraph>
<paragraph id="8BE20134E3475B47FF7361EC85B45FCD" blockId="11.[132,776,470,593]" box="[132,453,470,497]" pageId="11" pageNumber="70">
<emphasis id="B929DD26E3475B47FF7361EC85B45FCD" box="[132,453,470,497]" italics="true" pageId="11" pageNumber="70">
External morphology (
<figureCitation id="13661DB1E3475B47FE9D61ED85CC5FCD" box="[362,445,471,497]" captionStart="FIG" captionStartId="10.[133,144,1986,2004]" captionTargetBox="[132,1455,216,1944]" captionTargetId="figure-78@10.[698,1267,764,1315]" captionTargetPageId="10" captionText="FIG. 5. — Shell and reproductive system of Satsuma nivnu n. sp.: A, shell of holotype (NMNS-8676-005); B, C, genitalia (paratype NMNS-8676-006):whole genitalia (B), opened genitalia (C); D, living animal. Scale bars: 10 mm. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10672376" httpUri="https://zenodo.org/record/10672376/files/figure.png" pageId="11" pageNumber="70">Fig. 5D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BE20134E3475B47FF7361CD85295C6D" blockId="11.[132,776,470,593]" pageId="11" pageNumber="70">Brown with irregular, small, dark brown spots and a distinct yellowish line running from head between tentacles to collar. Tentacles dark grey.</paragraph>
<paragraph id="8BE20134E3475B47FF73624C853E5CAC" blockId="11.[130,777,630,1294]" box="[132,335,630,656]" pageId="11" pageNumber="70">
<emphasis id="B929DD26E3475B47FF73624C853E5CAC" box="[132,335,630,656]" italics="true" pageId="11" pageNumber="70">Reproductive system</emphasis>
</paragraph>
<paragraph id="8BE20134E3475B47FF7362AC85935B31" blockId="11.[130,777,630,1294]" pageId="11" pageNumber="70">
Bursa copulatrix oval. Pedunculus of bursa copulatrix as long as spermoviduct,
<quantity id="4CA5ACD1E3475B47FECD628F85A85CEC" box="[314,473,693,720]" metricMagnitude="-2" metricUnit="m" metricValue="3.86" metricValueMax="4.35" metricValueMin="3.37" pageId="11" pageNumber="70" unit="mm" value="38.6" valueMax="43.5" valueMin="33.7">33.7-43.5 mm</quantity>
, thickened basally, gradually slender toward bursa copulatrix. Free oviduct short,
<quantity id="4CA5ACD1E3475B47FD2F62EC84A75D2C" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.1" metricValueMin="0.97" pageId="11" pageNumber="70" unit="mm" value="10.35" valueMax="11.0" valueMin="9.7">9.7- 11 mm</quantity>
. Vagina slender,
<quantity id="4CA5ACD1E3475B47FE7062CF867E5D2C" box="[391,527,757,784]" metricMagnitude="-2" metricUnit="m" metricValue="2.355" metricValueMax="2.5" metricValueMin="2.21" pageId="11" pageNumber="70" unit="mm" value="23.55" valueMax="25.0" valueMin="22.1">22.1-25 mm</quantity>
in length, as thickened as penis, smooth externally, with 13-15 thin internal pilasters. Atrium short, coarsely wrinkled inside. Penis elongate,
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in length; slender shortly near atrium with smooth interior, becoming bulging penial pouch (
<figureCitation id="13661DB1E3475B47FD59634F87705DAC" box="[686,769,885,912]" captionStart="FIG" captionStartId="10.[133,144,1986,2004]" captionTargetBox="[132,1455,216,1944]" captionTargetId="figure-78@10.[698,1267,764,1315]" captionTargetPageId="10" captionText="FIG. 5. — Shell and reproductive system of Satsuma nivnu n. sp.: A, shell of holotype (NMNS-8676-005); B, C, genitalia (paratype NMNS-8676-006):whole genitalia (B), opened genitalia (C); D, living animal. Scale bars: 10 mm. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10672376" httpUri="https://zenodo.org/record/10672376/files/figure.png" pageId="11" pageNumber="70">Fig. 5B</figureCitation>
) with 6-11 smooth and muscular internal pilasters; apparently thickened near epiphallus, twice the thickness of distal portion, furrowed and striped externally corresponding to 16-18 strong, straight, corrugated pilasters internally. Penial caecum conical, short,
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. Verge extending along penial caecum, fusiform, with smooth surface. Epiphallus slender,
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in length, internally with 2 major and 2-3 minor pilasters. Penis retractor muscle attached at basal 1/10-1/5 of epiphallus. Flagellum
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; swollen basally, internally with thick swelling extending from epiphallic pilasters, opening into vas deferense at base; tapering toward tip, internally with 3-5 pilasters.
</paragraph>
</subSubSection>
<subSubSection id="C34752BFE3475B47FF73650E862359D7" pageId="11" pageNumber="70" type="discussion">
<paragraph id="8BE20134E3475B47FF73650E84995B70" blockId="11.[130,777,1332,2028]" box="[132,232,1332,1358]" pageId="11" pageNumber="70">
<smallCapsWord id="8D0497E8E3475B47FF73650E84995B70" baselines="1351,1351" box="[132,232,1332,1358]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="11" pageNumber="70">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BE20134E3475B47FF736569862359D7" blockId="11.[130,777,1332,2028]" pageId="11" pageNumber="70">
The new species is similar to
<taxonomicName id="4C5D7AB7E3475B47FE5B656E86CA5B51" authority="(Kuroda, 1941)" authorityName="Shell" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[428,699,1363,1390]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="arisana">
<emphasis id="B929DD26E3475B47FE5B656E86615B51" box="[428,528,1364,1390]" italics="true" pageId="11" pageNumber="70">S. arisana</emphasis>
(
<bibRefCitation id="EFCC7CC5E3475B47FDEB656986C25B51" author="KURODA T." box="[540,691,1363,1389]" pageId="11" pageNumber="70" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C5D7AB7E3475B47FD33656E85415BB1" authority="(Kuroda, 1941)" baseAuthorityName="Kuroda" baseAuthorityYear="1941" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="tani">
<emphasis id="B929DD26E3475B47FD33656E87765B51" box="[708,775,1364,1390]" italics="true" pageId="11" pageNumber="70">S. tani</emphasis>
(
<bibRefCitation id="EFCC7CC5E3475B47FF7B654985585BB1" author="KURODA T." box="[140,297,1395,1421]" pageId="11" pageNumber="70" pagination="65 - 216" refId="ref9107" refString="KURODA T. 1941. - A catalogue of molluscan shells from Taiwan (Formosa), with description of new species. Memoirs of the Faculty of Science and Agriculture Taihoku Imperial University 12: 65 - 216." type="journal article" year="1941">Kuroda, 1941</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C5D7AB7E3475B47FE98654984CD5B91" authority="Wu, Lin &amp; Hwang, 2007" authorityName="Wu, Lin &amp; Hwang" authorityYear="2007" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="longkiauwensis">
<emphasis id="B929DD26E3475B47FE98654986545BB1" box="[367,549,1395,1421]" italics="true" pageId="11" pageNumber="70">S. longkiauwensis</emphasis>
<bibRefCitation id="EFCC7CC5E3475B47FDDA654E84CD5B91" author="WU S. - P. &amp; LIN Y. - S. &amp; HWANG C. - C." pageId="11" pageNumber="70" pagination="59 - 68" refId="ref9690" refString="WU S. - P., LIN Y. - S. &amp; HWANG C. - C. 2007. - A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan. Zootaxa 1608 (1): 59 - 68. https: // doi. org / 10.11646 / zootaxa. 1608.1.8" type="journal article" year="2007">
Wu, Lin &amp; Hwang,
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</bibRefCitation>
</taxonomicName>
in
<collectingCountry id="F34A41A4E3475B47FF2D65A9855A5B91" box="[218,299,1427,1453]" name="Taiwan" pageId="11" pageNumber="70">Taiwan</collectingCountry>
,
<taxonomicName id="4C5D7AB7E3475B47FEC365A986065B92" authority="(Martens, 1865)" baseAuthorityName="Martens" baseAuthorityYear="1865" box="[308,631,1427,1454]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="myomphala">
<emphasis id="B929DD26E3475B47FEC365A985B55B91" box="[308,452,1427,1453]" italics="true" pageId="11" pageNumber="70">S. myomphala</emphasis>
(Martens, 1865)
</taxonomicName>
in
<collectingCountry id="F34A41A4E3475B47FD6065AE86A75B92" box="[663,726,1428,1454]" name="Japan" pageId="11" pageNumber="70">Japan</collectingCountry>
, and
<taxonomicName id="4C5D7AB7E3475B47FF756589857D5BF1" baseAuthorityName="Pfeiffer" baseAuthorityYear="1845" box="[130,268,1459,1485]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="mercatoria">
<emphasis id="B929DD26E3475B47FF756589857D5BF1" box="[130,268,1459,1485]" italics="true" pageId="11" pageNumber="70">S. mercatoria</emphasis>
</taxonomicName>
in
<collectingRegion id="4999CFD6E3475B47FEC5658985E55BF1" box="[306,404,1459,1485]" country="Japan" name="Okinawa" pageId="11" pageNumber="70">Okinawa</collectingRegion>
Island in having a large shell that is wider than
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.
<taxonomicName id="4C5D7AB7E3475B47FE8965E9865A5BD1" authorityName="Shell" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[382,555,1491,1517]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="arisana">
<emphasis id="B929DD26E3475B47FE8965E9865A5BD1" box="[382,555,1491,1517]" italics="true" pageId="11" pageNumber="70">Satsuma arisana</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5D7AB7E3475B47FDCD65E986BE5BD1" baseAuthorityName="Martens" baseAuthorityYear="1865" box="[570,719,1491,1517]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="myomphala">
<emphasis id="B929DD26E3475B47FDCD65E986BE5BD1" box="[570,719,1491,1517]" italics="true" pageId="11" pageNumber="70">S. myomphala</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5D7AB7E3475B47FF7165C984BA5830" baseAuthorityName="Kuroda" baseAuthorityYear="1941" box="[134,203,1523,1549]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="tani">
<emphasis id="B929DD26E3475B47FF7165C984BA5830" box="[134,203,1523,1549]" italics="true" pageId="11" pageNumber="70">S. tani</emphasis>
</taxonomicName>
have the umbilicus completely covered by a reflected columellar lip. The former two species have been dissected for their genitalia (
<bibRefCitation id="EFCC7CC5E3475B47FEA6660985AD5871" author="AZUMA M." box="[337,476,1586,1613]" pageId="11" pageNumber="70" refId="ref8367" refString="AZUMA M. 1995. - Colored Illustrations of the Land Snails of Japan. Osaka, Hoikusha, xvi + 343 p." type="book" year="1995">Azuma 1995</bibRefCitation>
;
<bibRefCitation id="EFCC7CC5E3475B47FE1E660886005871" author="CHANG K. - M." box="[489,625,1586,1613]" pageId="11" pageNumber="70" pagination="41 - 48" refId="ref8391" refString="CHANG K. - M. 1997. - Anatomy of Coniglobus arisanus (Kuroda) from Taiwan (Pulmonata: Camaenidae). Bulletin of Malacology 21: 41 - 48." type="journal article" year="1997">Chang 1997</bibRefCitation>
) which lack a penial pouch near the atrium and a basally swollen flagellum.
<taxonomicName id="4C5D7AB7E3475B47FF746649850B58B0" authorityName="Wu, Lin &amp; Hwang" authorityYear="2007" box="[131,378,1650,1677]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="longkiauwensis">
<emphasis id="B929DD26E3475B47FF746649850B58B0" box="[131,378,1650,1677]" italics="true" pageId="11" pageNumber="70">Satsuma longkiauwensis</emphasis>
</taxonomicName>
has a lower shell (SH/SW = 0.718 vs
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<geoCoordinate id="EE6967F3E3475B47FF7366A884B55890" box="[132,196,1682,1708]" degrees="0.726" direction="north" orientation="latitude" pageId="11" pageNumber="70" precision="55" value="0.726">0.726</geoCoordinate>
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in
</quantity>
<taxonomicName id="4C5D7AB7E3475B47FEC766A885FB5890" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[304,394,1682,1708]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="nivnu" status="n. sp.">
<emphasis id="B929DD26E3475B47FEC766A885FB5890" box="[304,394,1682,1708]" italics="true" pageId="11" pageNumber="70">S. nivnu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3475B47FE6566A985BE5890" box="[402,463,1682,1709]" pageId="11" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
), lighter shell colour, a more covered umbilicus, an unapparent penial caecum, and a short, truncated oval penial verge that is not extended along penial caecum (
<bibRefCitation id="EFCC7CC5E3475B47FF1D66C885E55930" author="WU S. - P. &amp; LIN Y. - S. &amp; HWANG C. - C." box="[234,404,1778,1804]" pageId="11" pageNumber="70" pagination="59 - 68" refId="ref9690" refString="WU S. - P., LIN Y. - S. &amp; HWANG C. - C. 2007. - A new Satsuma species (Pulmonata: Camaenidae) endemic to Taiwan. Zootaxa 1608 (1): 59 - 68. https: // doi. org / 10.11646 / zootaxa. 1608.1.8" type="journal article" year="2007">
Wu
<emphasis id="B929DD26E3475B47FEED66C985205930" box="[282,337,1778,1804]" italics="true" pageId="11" pageNumber="70">et al.</emphasis>
2007
</bibRefCitation>
).
<taxonomicName id="4C5D7AB7E3475B47FE5A66C886F15930" baseAuthorityName="Pfeiffer" baseAuthorityYear="1845" box="[429,640,1778,1804]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="mercatoria">
<emphasis id="B929DD26E3475B47FE5A66C886F15930" box="[429,640,1778,1804]" italics="true" pageId="11" pageNumber="70">Satsuma mercatoria</emphasis>
</taxonomicName>
has a lower shell, lighter shell colour, descending behind aperture, weaker spiral striations on the base of the shell, and the absence of a penial pouch (
<bibRefCitation id="EFCC7CC5E3475B47FEC4676885CE5950" author="AZUMA M." box="[307,447,1873,1900]" pageId="11" pageNumber="70" refId="ref8367" refString="AZUMA M. 1995. - Colored Illustrations of the Land Snails of Japan. Osaka, Hoikusha, xvi + 343 p." type="book" year="1995">Azuma 1995</bibRefCitation>
). The new species is similar to
<taxonomicName id="4C5D7AB7E3475B47FF736748857759B7" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[132,262,1905,1932]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="hsuehshan" status="n. sp.">
<emphasis id="B929DD26E3475B47FF736748857759B7" box="[132,262,1905,1932]" italics="true" pageId="11" pageNumber="70">S. hsuehshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3475B47FEFC6748853559B0" box="[267,324,1906,1932]" pageId="11" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
in having conical spire and a penial pouch.
<taxonomicName id="4C5D7AB7E3475B47FF7567A885395997" authority="Hwang &amp; Wu, 2024" authorityName="Hwang &amp; Wu" authorityYear="2024" box="[130,328,1937,1964]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="70" phylum="Mollusca" rank="species" species="hsuehshan" status="sp. nov.">
<emphasis id="B929DD26E3475B47FF7567A885395997" box="[130,328,1937,1964]" italics="true" pageId="11" pageNumber="70">Satsuma hsuehshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A605DE3475B47FEBA67A885F75990" box="[333,390,1938,1964]" pageId="11" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
has a smaller shell (maximum SW = 27.0 mm), denser spiral striation on base of shell, a stronger penial pouch than the present new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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