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<document id="637D13BA94666110FF7E7C7FE99C54E7" ID-DOI="10.5281/zenodo.195431" ID-GBIF-Dataset="2df8bd5a-e125-42aa-aa6f-ae3e78b83929" ID-ISSN="1175-5326" ID-Zenodo-Dep="195431" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460226955823" checkinUser="plazi" docAuthor="Molineri, Carlos" docDate="2010" docId="03F39207FFB1FFD47EFCFCF8F4E1DFCB" docLanguage="en" docName="zt02481p036.pdf" docOrigin="Zootaxa 2481" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tortopus Needham &amp; Murphy" docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="FFCAEA7FFFB9FFDE7E6BFF82F50BD901" masterDocTitle="A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="9" updateTime="1698238640945" updateUser="plazi">
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<mods:title id="8F34C71814BAC8A095510AC5A73D57D9">A cladistic revision of Tor top us Needham &amp; Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae)</mods:title>
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<mods:namePart id="487A41C717AA8E775CA43B6D607FD8CA">Molineri, Carlos</mods:namePart>
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<mods:date id="808688D0E3B34E8FBD5323E956775939">2010</mods:date>
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<treatment id="03F39207FFB1FFD47EFCFCF8F4E1DFCB" ID-DOI="http://doi.org/10.5281/zenodo.5690563" ID-GBIF-Taxon="119386655" ID-Zenodo-Dep="5690563" LSID="urn:lsid:plazi:treatment:03F39207FFB1FFD47EFCFCF8F4E1DFCB" httpUri="http://treatment.plazi.org/id/03F39207FFB1FFD47EFCFCF8F4E1DFCB" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="C340709AFFB1FFD67EFCFCF8F6B4DAF8" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BE52311FFB1FFD67EFCFCF8F705DA95" blockId="8.[151,526,890,916]" box="[151,526,890,916]" pageId="8" pageNumber="9">
<heading id="D0AD947DFFB1FFD67EFCFCF8F705DA95" bold="true" box="[151,526,890,916]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<taxonomicName id="4C5A5892FFB1FFD67EFCFCF8F705DA95" ID-CoL="7XX3" ID-ENA="309600" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" box="[151,526,890,916]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB1FFD67EFCFCF8F705DA95" bold="true" box="[151,526,890,916]" pageId="8" pageNumber="9">
<emphasis id="B92EFF03FFB1FFD67EFCFCF8F5F4DA95" bold="true" box="[151,255,890,916]" italics="true" pageId="8" pageNumber="9">Tortopus</emphasis>
Needham &amp; Murphy
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE52311FFB1FFD67EFCFC40F6B4DAF8" blockId="8.[151,1276,962,1049]" pageId="8" pageNumber="9">
<taxonomicName id="4C5A5892FFB1FFD67EFCFC40F4D1DAD8" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" box="[151,474,962,985]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB1FFD67EFCFC40F5FEDAD8" box="[151,245,962,985]" italics="true" pageId="8" pageNumber="9">Tortopus</emphasis>
Needham &amp; Murphy
</taxonomicName>
(in part), 1924: 23; Ulmer (in part), 1933: 197; Traver (in part), 1950: 596. Type-species:
<taxonomicName id="4C5A5892FFB1FFD67F5BFC60F7D2DAF8" authority="Needham &amp; Murphy" authorityName="Needham &amp; Murphy" box="[304,729,994,1017]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB1FFD67F5BFC60F4F8DAF8" box="[304,499,994,1017]" italics="true" pageId="8" pageNumber="9">Tortopus igaranus</emphasis>
Needham &amp; Murphy
</taxonomicName>
, original designation.
</paragraph>
</subSubSection>
<subSubSection id="C340709AFFB1FFD47EFCFB80F4E1DFCB" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
<paragraph id="8BE52311FFB1FFD67EFCFB80F1BFDD18" blockId="8.[151,1276,962,1049]" box="[151,1204,1026,1049]" pageId="8" pageNumber="9">
Species composition:
<taxonomicName id="4C5A5892FFB1FFD67FEAFB80F4F1DD18" box="[385,506,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB1FFD67FEAFB80F4F1DD18" box="[385,506,1026,1049]" italics="true" pageId="8" pageNumber="9">T. igaranus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A5892FFB1FFD67C6CFB80F795DD18" box="[519,670,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="circumfluus">
<emphasis id="B92EFF03FFB1FFD67C6CFB80F795DD18" box="[519,670,1026,1049]" italics="true" pageId="8" pageNumber="9">T. circumfluus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A5892FFB1FFD67CC0FB80F605DD18" box="[683,782,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFB1FFD67CC0FB80F605DD18" box="[683,782,1026,1049]" italics="true" pageId="8" pageNumber="9">T. harrisi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A5892FFB1FFD67D77FB80F67ADD18" box="[796,881,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zottai">
<emphasis id="B92EFF03FFB1FFD67D77FB80F67ADD18" box="[796,881,1026,1049]" italics="true" pageId="8" pageNumber="9">T. zottai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A5892FFB1FFD67D15FB80F6D3DD18" box="[894,984,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFB1FFD67D15FB80F6D3DD18" box="[894,984,1026,1049]" italics="true" pageId="8" pageNumber="9">T. bellus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A5892FFB1FFD67D8EFB80F150DD18" box="[997,1115,1026,1049]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="arenales" status="sp. nov.">
<emphasis id="B92EFF03FFB1FFD67D8EFB80F150DD18" box="[997,1115,1026,1049]" italics="true" pageId="8" pageNumber="9">T. arenales</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4278FFB1FFD67A0AFB80F1BFDD18" box="[1121,1204,1026,1049]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BE52311FFB1FFD67EFCFBCAF00EDF25" blockId="8.[151,1437,1096,1908]" pageId="8" pageNumber="9">
Adult: fore legs of male normal, remaining legs of male and all legs of female distorted and non-functional. Mesosternum: furcasternal protuberances approximating each other anteriorly (Fig. 4) so furcasternal longitudinal impression narrow basally and progressively wider posteriorly. Female pronotum with thin anterior portion (or ring), about 1/6 of the total length of pronotum in dorsal view. Wings sexually dimorphic, all veins of female thickened; posterior margin of hind wings of female with a reticulated pattern of veinlets. Female fore wing with veins R3 and short intercalary before it present (Fig. 8). Abdomen. Female abdominal sternum VIII with relatively small and deep sockets (forceps receptors), with its openings directed anteriorly, and with well marked longitudinal furrows on sternum associated to them (
<figureCitation id="13613F94FFB1FFD67A5DFAD2F1C5DC6B" box="[1078,1230,1360,1386]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="8" pageNumber="9">Figs. 70–71</figureCitation>
). Male genitalia: abdominal sternum IX divided by a mediolongitudinal membranous furrow (Fig. 2); parastyli short, straight and pointed, less than 3 times the length of their bases; penes bladelike with their basal 1/5–1/4 fused (Fig. 2); lateral margin of penes narrowly sclerotized, apex not forming a hooked spine. Penial arms formed by a single transverse bar which is articulated to the posterior corners of tergum IX. Forceps two-segmented, basal segment short and small, with a ventral knob (Figs. 2); distal segment long, slender and clublike.
</paragraph>
<paragraph id="8BE52311FFB1FFD67EADF9B2F4F7DE05" blockId="8.[151,1437,1096,1908]" pageId="8" pageNumber="9">
Nymph (only the nymph of
<taxonomicName id="4C5A5892FFB1FFD67C7BF9B2F776DF48" box="[528,637,1584,1609]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="harrisi">
<emphasis id="B92EFF03FFB1FFD67C7BF9B2F776DF48" box="[528,637,1584,1609]" italics="true" pageId="8" pageNumber="9">T. harrisi</emphasis>
</taxonomicName>
is known, for a complete characterization see
<bibRefCitation id="EFCB5EE0FFB1FFD67ACBF9B2F09ADF4B" author="Molineri" box="[1184,1425,1584,1610]" pageId="8" pageNumber="35" refString="Molineri, C., Siegloch A. &amp; Righi-Cavallaro, K. O. (2010) The nymph of Tortopus harrisi Traver (Ephemeroptera: Polymitarcyidae). Zootaxa, 2436, 65 - 68." type="journal article" year="2010">Molineri et al. 2010</bibRefCitation>
): head with fronto-clypeal region expanded, straight and not surpassing ventrally the mandibular tusks (
<figureCitation id="13613F94FFB1FFD67B00F9D7F5B3DF95" captionStart="FIGURES 93 – 96" captionStartId="30.[151,272,1872,1896]" captionTargetBox="[188,1398,197,1845]" captionTargetId="figure@30.[188,1398,197,1846]" captionTargetPageId="30" captionText="FIGURES 93 – 96. Polymitarcyidae, nymphal head: 93, Asthenopus sp.; 94, Campsurus near violaceus; 95, Tortopus harrisi; 96, Tortopsis obscuripennis." httpUri="https://zenodo.org/record/195446/files/figure.png" pageId="8" pageNumber="9">Fig. 95</figureCitation>
); mandibular tusks long with two large tubercles (submedian and subapical) on inner margin; maxilla with a basal finger like gill. Large filtering setae present on mouthparts and fore legs. Apex of tibia-tarsus in fore legs not strongly expanded (dorsal projection 2/5 of total length of claw). Abdomen with vestigial unilamellate gill on segment I.
</paragraph>
<paragraph id="8BE52311FFB1FFD67EADF892F7C0DE75" blockId="8.[151,1437,1096,1908]" pageId="8" pageNumber="9">
Eggs. Hemispherical, bowl-shaped (
<figureCitation id="13613F94FFB1FFD67C06F892F623DE2B" box="[621,808,1808,1834]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="8" pageNumber="9">Figs. 67–69, 72</figureCitation>
). Length, 325–400 µm; width, 260–345 µm. No polar caps or attachment structures. Chorion with microsculptures consisting of slightly elevated pentagonal and hexagonal cells, or shallow subcircular grooves.
</paragraph>
<paragraph id="8BE52311FFB0FFD77EFCF8A1F4C6DEDF" blockId="9.[151,1436,1827,2014]" pageId="9" pageNumber="10">
<emphasis id="B92EFF03FFB0FFD77EFCF8A1F448DE3A" bold="true" box="[151,323,1827,1851]" pageId="9" pageNumber="10">FIGURES 2–9.</emphasis>
<taxonomicName id="4C5A5892FFB0FFD77F21F8A6F706DE3A" box="[330,525,1828,1851]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="igaranus">
<emphasis id="B92EFF03FFB0FFD77F21F8A6F706DE3A" box="[330,525,1828,1851]" italics="true" pageId="9" pageNumber="10">Tortopus igaranus</emphasis>
</taxonomicName>
. 2, male genitalia, ventral view (f1 and f2 = forceps segment 1 and 2, k = knob, mf = median furrow, pe = penes, ps = parastilus, sIX = ninth abdominal sternum); 3, same, lateral view; 4, mesofurcasternal plates (female
<typeStatus id="54E19DB3FFB0FFD77F5FF8E6F498DE7A" box="[308,403,1892,1915]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
, fsi = furcasternal impression, fsp = furcasternal plates); 5, head (female
<typeStatus id="54E19DB3FFB0FFD77AF4F8E6F00BDE7A" box="[1183,1280,1892,1915]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
); 6, sockets on abdominal sternum VIII (female
<typeStatus id="54E19DB3FFB0FFD77F9EF806F75FDE9A" box="[501,596,1924,1947]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
, da = depressed area, s = socket, so = sockets´openning); 7, same (
<typeStatus id="54E19DB3FFB0FFD77B78F806F07CDE9A" box="[1299,1399,1924,1947]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
); 8, fore wing (female
<typeStatus id="54E19DB3FFB0FFD77F37F826F4B1DEBA" box="[348,442,1956,1979]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
, R2, R3 and R4+5 = main veins of radial sector, IR = radial intercalary, iv = intercalary vein); 9, hind wing (female
<typeStatus id="54E19DB3FFB0FFD77F0AF845F4C9DEDF" box="[353,450,1991,2014]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
).
</paragraph>
<caption id="DF257399FFB3FFD47EFCFBDDF76EDD96" httpUri="https://zenodo.org/record/195433/files/figure.png" pageId="10" pageNumber="11" targetBox="[169,1402,195,1092]" targetPageId="10">
<paragraph id="8BE52311FFB3FFD47EFCFBDDF76EDD96" blockId="10.[151,1436,1119,1175]" pageId="10" pageNumber="11">
<emphasis id="B92EFF03FFB3FFD47EFCFBDDF454DD76" bold="true" box="[151,351,1119,1143]" pageId="10" pageNumber="11">FIGURES 10–13.</emphasis>
<taxonomicName id="4C5A5892FFB3FFD47F0DFBE2F707DD76" box="[358,524,1120,1143]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="B92EFF03FFB3FFD47F0DFBE2F707DD76" box="[358,524,1120,1143]" italics="true" pageId="10" pageNumber="11">Tortopus bellus</emphasis>
</taxonomicName>
: 10, male genitalia, v.v.; 11, details of apex of penes; 12, detail of pedestal, v.v.; 13, sockets on female abdominal sternum VIII.
</paragraph>
</caption>
<paragraph id="8BE52311FFB3FFD47EADFB47F4E1DFCB" blockId="10.[151,1437,1221,1738]" pageId="10" pageNumber="11">
Discussion and diagnosis. The adults of
<taxonomicName id="4C5A5892FFB3FFD47CF4FB47F60DDDDF" box="[671,774,1221,1246]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47CF4FB47F60DDDDF" box="[671,774,1221,1246]" italics="true" pageId="10" pageNumber="11">Tortopus</emphasis>
</taxonomicName>
can be separated from those of its sister genus
<taxonomicName id="4C5A5892FFB3FFD47B5BFB47F092DDDF" box="[1328,1433,1221,1246]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47B5BFB47F092DDDF" box="[1328,1433,1221,1246]" italics="true" pageId="10" pageNumber="11">Tortopsis</emphasis>
</taxonomicName>
by: 1) mesofurcasternal plates approximating each other only basally, with inner margins diverging posteriorly from their basal 1/3 or less (Fig. 4); 2) male abdominal sternum IX longitudinally divided by a median furrow (Fig. 2); 3) parastyli short, less than 3 times the length of their bases (pedestals); 4) penes bladelike and basally fused, at least on basal 1/5; 5) female fore wing with veins R3 and short intercalary present (Fig. 8) (veins IRSa and RSa
<quantity id="4CA28EF4FFB3FFD47C2FFA11F761DC9D" box="[580,618,1410,1441]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="11" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="EFCB5EE0FFB3FFD47C1AFA00F60BDC9D" author="Kluge" box="[625,768,1410,1436]" pageId="10" pageNumber="34" refString="Kluge, N. Yu. (2004) The phylogenetic system of Ephemeroptera. Kluwer, Dordrecht, 442 pp." type="book" year="2004">Kluge 2004</bibRefCitation>
); 6) sockets on female abdominal sternum VIII relatively small and submedian in position (
<figureCitation id="13613F94FFB3FFD47C48FA29F770DCC4" box="[547,635,1451,1477]" captionStart="FIGURES 67 – 72" captionStartId="25.[151,269,1458,1482]" captionTargetBox="[187,1399,197,1432]" captionTargetId="figure@25.[187,1399,197,1433]" captionTargetPageId="25" captionText="FIGURES 67 – 72. SEM photographs. Tortopus igaranus: 67, egg, general view; 68, egg, detail of convex side; 69, egg, detail of concave side. Tortopus arenales: 70, female abdominal sternum VIII; 71, same, detail of sockets; 72, eggs." httpUri="https://zenodo.org/record/195442/files/figure.png" pageId="10" pageNumber="11">Fig. 70</figureCitation>
), openings towards fore margin of sternum. The characters useful to separate
<taxonomicName id="4C5A5892FFB3FFD47F68FA53F465DCEB" box="[259,366,1489,1514]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47F68FA53F465DCEB" box="[259,366,1489,1514]" italics="true" pageId="10" pageNumber="11">Tortopus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A5892FFB3FFD47FC1FA53F711DCEB" box="[426,538,1489,1514]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47FC1FA53F711DCEB" box="[426,538,1489,1514]" italics="true" pageId="10" pageNumber="11">Tortopsis</emphasis>
</taxonomicName>
from the other genera of the family are listed in the phylogenetic section (apomorphies of node c). The coupling apparatus of
<taxonomicName id="4C5A5892FFB3FFD47D48FA74F684DF0E" box="[803,911,1526,1551]" class="Insecta" family="Polymitarcyidae" genus="Tortopus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B92EFF03FFB3FFD47D48FA74F684DF0E" box="[803,911,1526,1551]" italics="true" pageId="10" pageNumber="11">Tortopus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D4278FFB3FFD47DF3FA74F6CBDF11" box="[920,960,1526,1552]" pageId="10" pageNumber="11" sensu="stricto">s.s.</taxonomicNameLabel>
apparently involves male forceps (not parastyli, poorly developed) and female sockets. Male forceps are not large enough to embrace female abdomen, but they seem to fit in the submedian sockets. Parastyli are so poorly developed that they could not reach female sockets during copula. In contrast, the coupling apparatus of
<taxonomicName id="4C5A5892FFB3FFD47A6BF9E4F167DF7E" box="[1024,1132,1638,1663]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47A6BF9E4F167DF7E" box="[1024,1132,1638,1663]" italics="true" pageId="10" pageNumber="11">Tortopsis</emphasis>
</taxonomicName>
(described in
<bibRefCitation id="EFCB5EE0FFB3FFD47B7FF9E4F48EDFA4" author="McCafferty" pageId="10" pageNumber="34" refString="McCafferty, W. P. &amp; Bloodgood, D. W. (1989) The female and male coupling apparatus in Tortopus mayflies. Aquatic Insects, 11, 141 - 146." type="journal article" year="1989">McCafferty &amp; Bloodgood, 1989</bibRefCitation>
, for a species now moved to
<taxonomicName id="4C5A5892FFB3FFD47CB4F909F640DFA5" box="[735,843,1675,1700]" class="Insecta" family="Polymitarcyidae" genus="Tortopsis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92EFF03FFB3FFD47CB4F909F640DFA5" box="[735,843,1675,1700]" italics="true" pageId="10" pageNumber="11">Tortopsis</emphasis>
</taxonomicName>
, see below) involves the strongly developed male parastyli and female sockets.
</paragraph>
</subSubSection>
</treatment>
</document>
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