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<document id="6F60FF9ACAC2053C130AEA86B3071468" ID-CLB-Dataset="4264" ID-DOI="10.18590/euscorpius.2010.vol2010.iss105.1" ID-GBIF-Dataset="51dee632-1d92-4ca3-b406-45e90789c50c" ID-ISSN="1536-9307" ID-Zenodo-Dep="4649083" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617137238613" checkinUser="carolina" docAuthor="Teruel, Rolando &amp; Roncallo, César A." docDate="2010" docId="03F387EAFFC1FA33FC35FBCDFE816F8C" docLanguage="en" docName="Euscorpius105.1-15.pdf" docOrigin="Euscorpius 105 (105)" docSource="https://mds.marshall.edu/euscorpius/vol2010/iss105/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Tityus tayrona Lourenco 1991" docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFCAFF92FFC4FA39FF82FFF9FFF96A03" masterDocTitle="Rare or poorly known scorpions from Colombia. IV. Additions, synonymies and new records (Scorpiones: Buthidae, Scorpionidae)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="4" updateTime="1698935563308" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="21C197336932494DB0058963BC9D77F8">Rare or poorly known scorpions from Colombia. IV. Additions, synonymies and new records (Scorpiones: Buthidae, Scorpionidae)</mods:title>
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<mods:namePart id="7E69F89801E3F4BB74BC6495C062DD84">Teruel, Rolando</mods:namePart>
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<treatment id="03F387EAFFC1FA33FC35FBCDFE816F8C" ID-DOI="http://doi.org/10.5281/zenodo.5510372" ID-GBIF-Taxon="187975280" ID-Zenodo-Dep="5510372" LSID="urn:lsid:plazi:treatment:03F387EAFFC1FA33FC35FBCDFE816F8C" httpUri="http://treatment.plazi.org/id/03F387EAFFC1FA33FC35FBCDFE816F8C" lastPageId="10" lastPageNumber="8" pageId="5" pageNumber="4">
<subSubSection id="C3406577FFC1FA3CFC35FBCDFAD16E4D" box="[951,1320,1076,1102]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BE536FCFFC1FA3CFC35FBCDFAD16E4D" blockId="5.[951,1320,1076,1102]" box="[951,1320,1076,1102]" pageId="5" pageNumber="4">
<heading id="D0AD8190FFC1FA3CFC35FBCDFAD16E4D" box="[951,1320,1076,1102]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="4" reason="3">
<taxonomicName id="4C5A4D7FFFC1FA3CFC35FBCDFAD16E4D" ID-CoL="579DY" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" box="[951,1320,1076,1102]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC1FA3CFC35FBCDFB9F6E4D" bold="true" box="[951,1126,1076,1102]" italics="true" pageId="5" pageNumber="4">Tityus tayrona</emphasis>
Lourenço, 1991
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3406577FFC1FA3CFBABFBACFC176F06" pageId="5" pageNumber="4" type="description">
<paragraph id="8BE536FCFFC1FA3CFBABFBACFB4C6E6F" blockId="5.[1065,1205,1109,1132]" box="[1065,1205,1109,1132]" pageId="5" pageNumber="4">
<figureCitation id="13612A79FFC1FA3CFBABFBACFB746E6F" box="[1065,1165,1109,1132]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStartId-0="5.[192,268,912,935]" captionStartId-1="6.[192,268,1836,1859]" captionStartId-2="7.[192,268,872,895]" captionStartId-3="8.[192,268,1838,1861]" captionStartId-4="9.[192,268,1492,1515]" captionTargetBox-0="[191,1440,191,882]" captionTargetBox-1="[191,1441,215,1801]" captionTargetBox-2="[191,1440,191,842]" captionTargetBox-3="[191,1441,215,1801]" captionTargetBox-4="[191,1440,191,1462]" captionTargetId-0="figure-6@5.[192,1439,192,881]" captionTargetId-1="figure-10@6.[192,1440,216,1800]" captionTargetId-2="figure-6@7.[192,1439,192,841]" captionTargetId-3="figure-10@8.[192,1440,216,1800]" captionTargetId-4="figure-6@9.[192,1439,192,1461]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Figure 3: Adult male of Tityus tayrona from Turbaco: a) entire dorsal view; b) entire ventral view." captionText-1="Figure 4: Adult male of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." captionText-2="Figure 5: Adult female of Tityus tayrona from Turbaco: a) entire dorsal view; b) entire ventral view." captionText-3="Figure 6: Adult female of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." captionText-4="Figure 7: Juvenile male of Tityus tayrona from Turbaco: a) entire dorsal view; b) entire ventral view; c) sternite V." figureDoi-0="http://doi.org/10.5281/zenodo.4649089" figureDoi-1="http://doi.org/10.5281/zenodo.4649091" figureDoi-2="http://doi.org/10.5281/zenodo.4649093" figureDoi-3="http://doi.org/10.5281/zenodo.4649095" figureDoi-4="http://doi.org/10.5281/zenodo.4649097" httpUri-0="https://zenodo.org/record/4649089/files/figure.png" httpUri-1="https://zenodo.org/record/4649091/files/figure.png" httpUri-2="https://zenodo.org/record/4649093/files/figure.png" httpUri-3="https://zenodo.org/record/4649095/files/figure.png" httpUri-4="https://zenodo.org/record/4649097/files/figure.png" pageId="5" pageNumber="4">Figs. 37</figureCitation>
,
<figureCitation id="13612A79FFC1FA3CFB18FBACFB4C6E6F" box="[1178,1205,1109,1132]" captionStart="Figures 1213" captionStartId="13.[192,278,1795,1818]" captionTargetBox="[191,1439,215,1759]" captionTargetId="figure-6@13.[192,1438,216,1759]" captionTargetPageId="13" captionText="Figures 1213: 12 (top). Geographical distribution of Tityus tayrona: previous records (black squares), new record (white square), records previously assigned to T. erikae (white squares with black dot). Abbreviations for Departments: Atl. = Atlántico, Magd. = Magdalena. 13 (bottom). Geographical distribution of Tarsoporosus macuira: previous records (black squares), new record (white square)." figureDoi="http://doi.org/10.5281/zenodo.4649105" httpUri="https://zenodo.org/record/4649105/files/figure.png" pageId="5" pageNumber="4">12</figureCitation>
</paragraph>
<paragraph id="8BE536FCFFC1FA3CFCBCFB6BFC176F06" blockId="5.[830,1442,1170,1347]" pageId="5" pageNumber="4">
<taxonomicName id="4C5A4D7FFFC1FA3CFCBCFB6BFB2F6EAA" authority="Lourenco, 1991 a: 275" authorityName="Lourenco" authorityPageNumber="275" authorityYear="1991" box="[830,1238,1170,1193]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC1FA3CFCBCFB6BFC226EAA" box="[830,987,1170,1193]" italics="true" pageId="5" pageNumber="4">Tityus tayrona</emphasis>
Lourenço, 1991a: 275
</taxonomicName>
, 279281, figs. 3 4, tabs. 12; Rojas-Runjaic &amp; Armas, 2002: 63; Botero-Trujillo &amp; Fagua, 2007: 129130, 132133, figs. 1115.
</paragraph>
</subSubSection>
<subSubSection id="C3406577FFC1FA3CFCBCFAF4FC116F40" pageId="5" pageNumber="4" type="reference_group">
<paragraph id="8BE536FCFFC1FA3CFCBCFAF4FC116F40" blockId="5.[830,1442,1170,1347]" pageId="5" pageNumber="4">
<taxonomicName id="4C5A4D7FFFC1FA3CFCBCFAF4FB4B6F27" authority="Lourenco, 1999: 1 - 6" authorityName="Lourenco" authorityPageNumber="1 - 6" authorityYear="1999" box="[830,1202,1293,1316]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC1FA3CFCBCFAF4FC336F27" box="[830,970,1293,1316]" italics="true" pageId="5" pageNumber="4">Tityus erikae</emphasis>
Lourenço, 1999: 16
</taxonomicName>
, figs. 17, tab. I.
<emphasis id="B92EEAEEFFC1FA3CFAECFAF4FC116F40" bold="true" pageId="5" pageNumber="4">New synonym.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3406577FFC1FA3EFCECFA9DFEFC6EFE" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BE536FCFFC1FA3EFCECFA9DFEC06E46" blockId="5.[830,1441,1380,1894]" lastBlockId="7.[192,803,948,1890]" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
<emphasis id="B92EEAEEFFC1FA3CFCECFA9DFC256F78" bold="true" box="[878,988,1380,1403]" pageId="5" pageNumber="4">Diagnosis</emphasis>
(emended): species of small size (males
<quantity id="4CA29B19FFC1FA3CFCBCFA7AFC4E6F99" box="[830,951,1411,1434]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.1" metricValueMin="3.3" pageId="5" pageNumber="4" unit="mm" value="37.0" valueMax="41.0" valueMin="33.0">3341 mm</quantity>
, females
<quantity id="4CA29B19FFC1FA3CFBA9FA7AFB5D6F99" box="[1067,1188,1411,1434]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="5" pageNumber="4" unit="mm" value="36.5" valueMax="38.0" valueMin="35.0">3538 mm</quantity>
) for the genus. Body light yellowish brown, very densely spotted with light to medium brown, tergites with three very irregular dark stripes (better defined in females and juveniles); metasomal segments IV similarly colored (i.e., V not conspicuously darkened), telson dark reddish brown; pedipalp fingers densely spotted, in adult males with basal portion subtly infuscate. Pedipalp chela and metasomal segment V inflate and moderately globular in larger males. Sternite V with a smooth whitish patch in both sexes: always small and flat in the adults (smaller in females), but expanded and bulky in juvenile males. Dorsolateral carinae of metasomal segments IIIV with distal tooth not conspicuously enlarged. Telson vesicle feebly granulose; subaculear tubercle large and rhomboidal, with two large dorsal granules. Pedipalp fixed finger with 1314 principal rows of granules, movable finger with 1415; basal lobe/notch combination vestigial in males, absent in females. Pectinal tooth count
<quantity id="4CA29B19FFC3FA3EFF42FBF6FED96E25" box="[192,288,1039,1062]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="4.826" metricValueMin="4.064" pageId="7" pageNumber="6" unit="in" value="17.5" valueMax="19.0" valueMin="16.0">1619 in</quantity>
males,
<quantity id="4CA29B19FFC3FA3EFEF4FBF6FE2F6E25" box="[374,470,1039,1062]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.572" metricValueMin="3.556" pageId="7" pageNumber="6" unit="in" value="16.0" valueMax="18.0" valueMin="14.0">1418 in</quantity>
females; basal middle lamella not dilated.
</paragraph>
<caption id="DF256674FFC2FA3FFF42F8D5FB876D5F" ID-DOI="http://doi.org/10.5281/zenodo.4649091" ID-Zenodo-Dep="4649091" httpUri="https://zenodo.org/record/4649091/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,268,1836,1859]" targetBox="[191,1441,215,1801]" targetPageId="6">
<paragraph id="8BE536FCFFC2FA3FFF42F8D5FB876D5F" blockId="6.[192,1440,1836,1884]" pageId="6" pageNumber="5">
<emphasis id="B92EEAEEFFC2FA3FFF42F8D5FED16D40" bold="true" box="[192,296,1836,1859]" pageId="6" pageNumber="5">Figure 4:</emphasis>
Adult male of
<taxonomicName id="4C5A4D7FFFC2FA3FFE38F8D7FDBF6D40" authorityName="Lourenco" authorityYear="1991" box="[442,582,1838,1859]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC2FA3FFE38F8D7FDBF6D40" box="[442,582,1838,1859]" italics="true" pageId="6" pageNumber="5">Tityus tayrona</emphasis>
</taxonomicName>
from Turbaco:
<emphasis id="B92EEAEEFFC2FA3FFD5FF8D7FD086D40" bold="true" box="[733,753,1838,1859]" pageId="6" pageNumber="5">a)</emphasis>
carapace;
<emphasis id="B92EEAEEFFC2FA3FFCDAF8D7FC946D40" bold="true" box="[856,877,1838,1859]" pageId="6" pageNumber="5">b)</emphasis>
pedipalp, dorsal view;
<emphasis id="B92EEAEEFFC2FA3FFBCCF8D7FB986D40" bold="true" box="[1102,1121,1838,1859]" pageId="6" pageNumber="5">c)</emphasis>
sternopectinal region;
<emphasis id="B92EEAEEFFC2FA3FFABCF8D7FAAA6D40" bold="true" box="[1342,1363,1838,1859]" pageId="6" pageNumber="5">d)</emphasis>
sternite V;
<emphasis id="B92EEAEEFFC2FA3FFF5CF8BEFF096D5F" bold="true" box="[222,240,1863,1884]" pageId="6" pageNumber="5">e)</emphasis>
metasomal segments IIII, lateral view;
<emphasis id="B92EEAEEFFC2FA3FFDFBF8BEFD736D5F" bold="true" box="[633,650,1863,1884]" pageId="6" pageNumber="5">f)</emphasis>
metasomal segments IVV and telson, lateral view.
</paragraph>
</caption>
<caption id="DF256674FFC3FA3EFF42FC91FB6E697C" ID-DOI="http://doi.org/10.5281/zenodo.4649093" ID-Zenodo-Dep="4649093" box="[192,1175,872,895]" httpUri="https://zenodo.org/record/4649093/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,268,872,895]" targetBox="[191,1440,191,842]" targetPageId="7">
<paragraph id="8BE536FCFFC3FA3EFF42FC91FB6E697C" blockId="7.[192,1175,872,895]" box="[192,1175,872,895]" pageId="7" pageNumber="6">
<emphasis id="B92EEAEEFFC3FA3EFF42FC91FED1697C" bold="true" box="[192,296,872,895]" pageId="7" pageNumber="6">Figure 5:</emphasis>
Adult female of
<taxonomicName id="4C5A4D7FFFC3FA3EFE4FFC93FDA1697C" authorityName="Lourenco" authorityYear="1991" box="[461,600,874,895]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFE4FFC93FDA1697C" box="[461,600,874,895]" italics="true" pageId="7" pageNumber="6">Tityus tayrona</emphasis>
</taxonomicName>
from Turbaco:
<emphasis id="B92EEAEEFFC3FA3EFD6DFC93FCFA697C" bold="true" box="[751,771,874,895]" pageId="7" pageNumber="6">a)</emphasis>
entire dorsal view;
<emphasis id="B92EEAEEFFC3FA3EFC40FC93FC2E697C" bold="true" box="[962,983,874,895]" pageId="7" pageNumber="6">b)</emphasis>
entire ventral view.
</paragraph>
</caption>
<paragraph id="8BE536FCFFC3FA3EFF72FBB4FEFC6EFE" blockId="7.[192,803,948,1890]" pageId="7" pageNumber="6">
<emphasis id="B92EEAEEFFC3FA3EFF72FBB4FE6D6E67" bold="true" box="[240,404,1101,1124]" pageId="7" pageNumber="6">New Records:</emphasis>
<materialsCitation id="3B323CA1FFC3FA3EFE1DFBB4FEF96EFE" ID-GBIF-Occurrence="3352143302" collectingDate="2008-10-07" collectingDateMax="2008-12-31" collectingDateMin="2008-10-07" collectionCode="R" collectorName="C. A. Roncallo &amp; J. A. Vargas" country="Colombia" county="Turbaco" location="Roncallo" municipality="Urbanizacion La Granja" pageId="7" pageNumber="6" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Bolivar">
<collectingCountry id="F34D766CFFC3FA3EFE1DFBB4FDC86E67" box="[415,561,1101,1124]" name="Colombia" pageId="7" pageNumber="6">COLOMBIA</collectingCountry>
:
<emphasis id="B92EEAEEFFC3FA3EFDC2FBB4FCE26E67" box="[576,795,1101,1124]" italics="true" pageId="7" pageNumber="6">
<collectingRegion id="499EF81EFFC3FA3EFDC2FBB4FCE26E67" box="[576,795,1101,1124]" country="Colombia" name="Bolivar" pageId="7" pageNumber="6">Bolívar Department</collectingRegion>
</emphasis>
,
<collectingCounty id="62844E70FFC3FA3EFF42FB92FEE56E81" box="[192,284,1131,1154]" pageId="7" pageNumber="6">Turbaco</collectingCounty>
,
<collectingMunicipality id="6B81AC86FFC3FA3EFEA9FB92FDC06E81" box="[299,569,1131,1154]" pageId="7" pageNumber="6">Urbanización La Granja</collectingMunicipality>
,
<date id="FFE4103CFFC3FA3EFDC5FB92FCE46E81" box="[583,797,1131,1154]" pageId="7" pageNumber="6" value="2008-12-31">
<collectingDate id="EFA0E9D4FFC3FA3EFDC5FB92FCE46E81" box="[583,797,1131,1154]" pageId="7" pageNumber="6" value="2008-12-31">31 December 2008</collectingDate>
</date>
, leg.
<collectorName id="26AF532AFFC3FA3EFF6CFB73FE766EA2" box="[238,399,1162,1185]" pageId="7" pageNumber="6">C. A. Roncallo</collectorName>
,
<collectionCode id="ED4BAE39FFC3FA3EFE1FFB73FE4B6EA2" box="[413,434,1162,1185]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
.
<location id="8E856027FFC3FA3EFE3EFB73FDE56EA2" LSID="urn:lsid:plazi:treatment:03F387EAFFC1FA33FC35FBCDFE816F8C:8E856027FFC3FA3EFE3EFB73FDE56EA2" box="[444,540,1162,1185]" country="Colombia" county="Turbaco" municipality="Urbanizacion La Granja" name="Roncallo" pageId="7" pageNumber="6" stateProvince="Bolivar">Roncallo</location>
&amp;
<collectorName id="26AF532AFFC3FA3EFDBDFB73FD3A6EA2" box="[575,707,1162,1185]" pageId="7" pageNumber="6">J. A. Vargas</collectorName>
,
<specimenCount id="9D5CFD75FFC3FA3EFD50FB73FD0A6EA2" box="[722,755,1162,1185]" pageId="7" pageNumber="6" type="male">1♂</specimenCount>
,
<specimenCount id="9D5CFD75FFC3FA3EFC82FB73FCDB6EA2" box="[768,802,1162,1185]" pageId="7" pageNumber="6" type="female">1♀</specimenCount>
(Sco-0413); Turbaco Botanical Garden,
<date id="FFE4103CFFC3FA3EFDF6FB50FCE26EC3" box="[628,795,1193,1216]" pageId="7" pageNumber="6" value="2008-10-07">
<collectingDate id="EFA0E9D4FFC3FA3EFDF6FB50FCE26EC3" box="[628,795,1193,1216]" pageId="7" pageNumber="6" value="2008-10-07">7 October 2008</collectingDate>
</date>
, leg.
<collectorName id="26AF532AFFC3FA3EFF76FB3EFE586EDD" box="[244,417,1223,1246]" pageId="7" pageNumber="6">C. A. Roncallo</collectorName>
&amp;
<collectionCode id="ED4BAE39FFC3FA3EFE53FB3EFE1F6EDD" box="[465,486,1223,1246]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="6">R</collectionCode>
. Roncallo,
<specimenCount id="9D5CFD75FFC3FA3EFDE8FB3EFD226EDD" box="[618,731,1223,1246]" pageId="7" pageNumber="6" type="juvenile">1 juvenile</specimenCount>
(Sco- 0414)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3406577FFC3FA33FF72FAFCFE816F8C" lastPageId="10" lastPageNumber="9" pageId="7" pageNumber="6" type="discussion">
<paragraph id="8BE536FCFFC3FA3EFF72FAFCFCF26F95" blockId="7.[192,803,948,1890]" pageId="7" pageNumber="6">
<emphasis id="B92EEAEEFFC3FA3EFF72FAFCFE986F1F" bold="true" box="[240,353,1285,1308]" pageId="7" pageNumber="6">Remarks:</emphasis>
the adult specimens were collected among rocks and rotten logs during a night search with UV light (02:30 hrs), in an area which had been cleared for building purposes at an altitude of
<quantity id="4CA29B19FFC3FA3EFDEFFA98FD4D6F7B" box="[621,692,1377,1400]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="6" unit="m" value="100.0">100 m</quantity>
a.s.l. The juvenile was found among bracts of
<taxonomicName id="4C5A4D7FFFC3FA3EFDC6FA86FD546F95" box="[580,685,1407,1430]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="7" pageNumber="6" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEAEEFFC3FA3EFDC6FA86FD546F95" box="[580,685,1407,1430]" italics="true" pageId="7" pageNumber="6">Heliconia</emphasis>
</taxonomicName>
flowers.
</paragraph>
<paragraph id="8BE536FCFFC3FA3EFF72FA67FB3A6E25" blockId="7.[192,803,948,1890]" lastBlockId="7.[830,1442,948,1890]" pageId="7" pageNumber="6">
In a previous paper, Teruel &amp; García (2008b: 10 12) commented on the deep taxonomic problems of the Colombian members of the “
<taxonomicName id="4C5A4D7FFFC3FA3EFD82FA22FD4E6FF1" box="[512,695,1499,1522]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="B92EEAEEFFC3FA3EFD82FA22FD4E6FF1" box="[512,695,1499,1522]" italics="true" pageId="7" pageNumber="6">Tityus clathratus</emphasis>
</taxonomicName>
” speciesgroup, and made a special reference to inadequate or unreliable diagnoses of most species, including
<taxonomicName id="4C5A4D7FFFC3FA3EFD5AF9E0FF0B6C4D" authorityName="Lourenco" authorityYear="1991" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFD5AF9E0FF0B6C4D" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D7FFFC3FA3EFEABF9CEFE736C4D" authorityName="Lourenco" authorityYear="1999" box="[297,394,1591,1614]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC3FA3EFEABF9CEFE736C4D" box="[297,394,1591,1614]" italics="true" pageId="7" pageNumber="6">T. erikae</emphasis>
</taxonomicName>
. Additional specimens studied herein allow updating and correcting the diagnosis of the former according to the current taxonomy of this group. It is worth to note here that the presence of a smooth patch in sternite V is now confirmed for both sexes of
<taxonomicName id="4C5A4D7FFFC3FA3EFC8FF94BFEEA6CEB" authorityName="Lourenco" authorityYear="1991" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFC8FF94BFEEA6CEB" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
, as previously suggested by Teruel &amp; García (2008: 12). This patch is small and flat in the adult male (
<figureCitation id="13612A79FFC3FA3EFF4AF8F7FEE26D26" box="[200,283,1806,1829]" captionStart="Figure 4" captionStartId="6.[192,268,1836,1859]" captionTargetBox="[191,1441,215,1801]" captionTargetId="figure-10@6.[192,1440,216,1800]" captionTargetPageId="6" captionText="Figure 4: Adult male of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4649091" httpUri="https://zenodo.org/record/4649091/files/figure.png" pageId="7" pageNumber="6">Fig. 4d</figureCitation>
), and even less conspicuous in adult female (
<figureCitation id="13612A79FFC3FA3EFF45F8D4FEE36D47" box="[199,282,1837,1860]" captionStart="Figure 6" captionStartId="8.[192,268,1838,1861]" captionTargetBox="[191,1441,215,1801]" captionTargetId="figure-10@8.[192,1440,216,1800]" captionTargetPageId="8" captionText="Figure 6: Adult female of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4649095" httpUri="https://zenodo.org/record/4649095/files/figure.png" pageId="7" pageNumber="6">Fig. 6d</figureCitation>
); thus, it is obvious that it was simply overlooked by these authors in their specimens; this is even demonstrated in the paper by Botero-Trujillo &amp; Fagua (2007), where this patch is explicitly stated to be absent in text page 133, but is in turn clearly shown in the figures 13 and 15 of the same paper.
</paragraph>
<paragraph id="8BE536FCFFC3FA30FCECFBD7FE1D6D0E" blockId="7.[830,1442,948,1890]" lastBlockId="9.[192,803,1568,1897]" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
On the other hand,
<taxonomicName id="4C5A4D7FFFC3FA3EFBCBFBD7FB216E46" authorityName="Lourenco" authorityYear="1999" box="[1097,1240,1070,1093]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC3FA3EFBCBFBD7FB216E46" box="[1097,1240,1070,1093]" italics="true" pageId="7" pageNumber="6">Tityus erikae</emphasis>
</taxonomicName>
was described by Lourenço (1999) on the basis of a single juvenile male, and it was originally separated from
<taxonomicName id="4C5A4D7FFFC3FA3EFB5FFB92FAA96E81" authorityName="Lourenco" authorityYear="1991" box="[1245,1360,1131,1154]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFB5FFB92FAA96E81" box="[1245,1360,1131,1154]" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
on the basis of only three supposedly diagnostic characters: “...
<emphasis id="B92EEAEEFFC3FA3EFCBCFB50FAF86EDD" italics="true" pageId="7" pageNumber="6">a more intense and darker pigmentation generally, a greater distance between median eyes,</emphasis>
[...]
<emphasis id="B92EEAEEFFC3FA3EFAC5FB3EFB696F1F" italics="true" pageId="7" pageNumber="6">and the presence of a smooth and shining expanded zone on the posterior region of sternite V</emphasis>
...” (Lourenço, 1999: 2; italics herein added). Juvenile
<taxonomicName id="4C5A4D7FFFC3FA3EFB08FADAFB336F39" authorityName="C. L. Koch" authorityYear="1836" box="[1162,1226,1315,1338]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEEFFC3FA3EFB08FADAFB336F39" box="[1162,1226,1315,1338]" italics="true" pageId="7" pageNumber="6">Tityus</emphasis>
</taxonomicName>
have no taxonomic value at all, because they lack secondary sexual dimorphism which is critical to diagnose closely related species, and this fact alone renders both Lourenços diagnosis and selection of the
<typeStatus id="54E1885EFFC3FA3EFB2BFA67FAFF6FB6" box="[1193,1286,1438,1461]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5A4D7FFFC3FA3EFABBFA67FA586FB6" authorityName="Lourenco" authorityYear="1999" box="[1337,1441,1438,1461]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC3FA3EFABBFA67FA586FB6" box="[1337,1441,1438,1461]" italics="true" pageId="7" pageNumber="6">T. erikae</emphasis>
</taxonomicName>
useless. Apart from this, the first two characters used by Lourenço are largely known to vary from juvenile through adult instars in all species of
<taxonomicName id="4C5A4D7FFFC3FA3EFB76FA03FACC6C12" authorityName="C. L. Koch" authorityYear="1836" box="[1268,1333,1530,1553]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAEEFFC3FA3EFB76FA03FACC6C12" box="[1268,1333,1530,1553]" italics="true" pageId="7" pageNumber="6">Tityus</emphasis>
</taxonomicName>
and thus cannot be used as diagnostic, so we are only left with character 3 to separate
<taxonomicName id="4C5A4D7FFFC3FA3EFBC7F9CEFB516C4D" authorityName="Lourenco" authorityYear="1999" box="[1093,1192,1591,1614]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC3FA3EFBC7F9CEFB516C4D" box="[1093,1192,1591,1614]" italics="true" pageId="7" pageNumber="6">T. erikae</emphasis>
</taxonomicName>
from
<taxonomicName id="4C5A4D7FFFC3FA3EFB70F9CEFA9C6C4D" authorityName="Lourenco" authorityYear="1991" box="[1266,1381,1591,1614]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFB70F9CEFA9C6C4D" box="[1266,1381,1591,1614]" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
. The examination of the additional specimens of
<taxonomicName id="4C5A4D7FFFC3FA3EFAAEF9AFFA596C6E" authorityName="Lourenco" authorityYear="1991" box="[1324,1440,1622,1645]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFAAEF9AFFA596C6E" box="[1324,1440,1622,1645]" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
herein recorded (which are all clearly conspecific) confirmed that the smooth patch of sternite V is also present in
<taxonomicName id="4C5A4D7FFFC3FA3EFC42F94BFBC16CCA" authorityName="Lourenco" authorityYear="1991" box="[960,1080,1714,1737]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFC3FA3EFC42F94BFBC16CCA" box="[960,1080,1714,1737]" italics="true" pageId="7" pageNumber="6">T. tayrona</emphasis>
</taxonomicName>
, and that it is ontogenetically variable: in males it is expanded and bulky in the juvenile instars (see our
<figureCitation id="13612A79FFC3FA3EFBC0F916FB6F6D05" box="[1090,1174,1775,1798]" captionStart="Figure 7" captionStartId="9.[192,268,1492,1515]" captionTargetBox="[191,1440,191,1462]" captionTargetId="figure-6@9.[192,1439,192,1461]" captionTargetPageId="9" captionText="Figure 7: Juvenile male of Tityus tayrona from Turbaco: a) entire dorsal view; b) entire ventral view; c) sternite V." figureDoi="http://doi.org/10.5281/zenodo.4649097" httpUri="https://zenodo.org/record/4649097/files/figure.png" pageId="7" pageNumber="6">Fig. 7c</figureCitation>
, which shows a patch identical to that depicted in Lourenço [1999: fig. 6] for the
<typeStatus id="54E1885EFFC3FA3EFCE5F8D4FC3C6D47" box="[871,965,1837,1860]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5A4D7FFFC3FA3EFC6FF8D4FBB26D47" authorityName="Lourenco" authorityYear="1999" box="[1005,1099,1837,1860]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFC3FA3EFC6FF8D4FBB26D47" box="[1005,1099,1837,1860]" italics="true" pageId="7" pageNumber="6">T. erikae</emphasis>
</taxonomicName>
), and it becomes much smaller and flat in the adult instar (see our
<figureCitation id="13612A79FFC3FA3EFB6AF8B2FAC46D61" box="[1256,1341,1867,1890]" captionStart="Figure 4" captionStartId="6.[192,268,1836,1859]" captionTargetBox="[191,1441,215,1801]" captionTargetId="figure-10@6.[192,1440,216,1800]" captionTargetPageId="6" captionText="Figure 4: Adult male of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4649091" httpUri="https://zenodo.org/record/4649091/files/figure.png" pageId="7" pageNumber="6">Figs. 4d</figureCitation>
and
<figureCitation id="13612A79FFC3FA3EFAF0F8B2FA696D61" box="[1394,1424,1867,1890]" captionStart="Figure 6" captionStartId="8.[192,268,1838,1861]" captionTargetBox="[191,1441,215,1801]" captionTargetId="figure-10@8.[192,1440,216,1800]" captionTargetPageId="8" captionText="Figure 6: Adult female of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4649095" httpUri="https://zenodo.org/record/4649095/files/figure.png" pageId="7" pageNumber="6">6d</figureCitation>
). Apart from this, the juvenile male
<taxonomicName id="4C5A4D7FFFCDFA30FDD1F9D9FD316C34" authorityName="Lourenco" authorityYear="1991" box="[595,712,1568,1591]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCDFA30FDD1F9D9FD316C34" box="[595,712,1568,1591]" italics="true" pageId="9" pageNumber="8">T. tayrona</emphasis>
</taxonomicName>
studied herein matches every point of the original description and figures of
<taxonomicName id="4C5A4D7FFFCDFA30FEE4F9A4FE306C77" authorityName="Lourenco" authorityYear="1999" box="[358,457,1629,1652]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFCDFA30FEE4F9A4FE306C77" box="[358,457,1629,1652]" italics="true" pageId="9" pageNumber="8">T. erikae</emphasis>
</taxonomicName>
exactly and the distribution of both taxa is also coincident, demonstrating that the
<typeStatus id="54E1885EFFCDFA30FF42F963FEE76CB2" box="[192,286,1690,1713]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
of the latter is merely a juvenile
<taxonomicName id="4C5A4D7FFFCDFA30FD25F963FCE26CB2" authorityName="Lourenco" authorityYear="1991" box="[679,795,1690,1713]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCDFA30FD25F963FCE26CB2" box="[679,795,1690,1713]" italics="true" pageId="9" pageNumber="8">T. tayrona</emphasis>
</taxonomicName>
. Thus, the following synonymy is herein established:
<taxonomicName id="4C5A4D7FFFCDFA30FF42F92EFDE46CED" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" box="[192,541,1751,1774]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCDFA30FF42F92EFE986CED" box="[192,353,1751,1774]" italics="true" pageId="9" pageNumber="8">Tityus tayrona</emphasis>
Lourenço, 1991
</taxonomicName>
=
<taxonomicName id="4C5A4D7FFFCDFA30FDC5F92EFEC16D0E" authority="Lourenco, 1999" authorityName="Lourenco" authorityYear="1999" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="B92EEAEEFFCDFA30FDC5F92EFD216CED" box="[583,728,1751,1774]" italics="true" pageId="9" pageNumber="8">Tityus erikae</emphasis>
Lourenço, 1999
</taxonomicName>
,
<emphasis id="B92EEAEEFFCDFA30FEC0F90FFE246D0E" bold="true" box="[322,477,1782,1805]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="A21D5795FFCDFA30FEC0F90FFE246D0E" box="[322,477,1782,1805]" pageId="9" pageNumber="8">new synonym</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<caption id="DF256674FFCCFA31FF42F8D7FB356D62" ID-DOI="http://doi.org/10.5281/zenodo.4649095" ID-Zenodo-Dep="4649095" httpUri="https://zenodo.org/record/4649095/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,268,1838,1861]" targetBox="[191,1441,215,1801]" targetPageId="8">
<paragraph id="8BE536FCFFCCFA31FF42F8D7FB356D62" blockId="8.[192,1440,1838,1889]" pageId="8" pageNumber="7">
<emphasis id="B92EEAEEFFCCFA31FF42F8D7FED56D46" bold="true" box="[192,300,1838,1861]" pageId="8" pageNumber="7">Figure 6:</emphasis>
Adult female of
<taxonomicName id="4C5A4D7FFFCCFA31FE5FF8C9FD956D46" authorityName="Lourenco" authorityYear="1991" box="[477,620,1840,1861]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCCFA31FE5FF8C9FD956D46" box="[477,620,1840,1861]" italics="true" pageId="8" pageNumber="7">Tityus tayrona</emphasis>
</taxonomicName>
from Turbaco:
<emphasis id="B92EEAEEFFCCFA31FC8FF8C9FCD86D46" bold="true" box="[781,801,1840,1861]" pageId="8" pageNumber="7">a)</emphasis>
carapace;
<emphasis id="B92EEAEEFFCCFA31FC0CF8C9FC5A6D46" bold="true" box="[910,931,1840,1861]" pageId="8" pageNumber="7">b)</emphasis>
pedipalp, dorsal view;
<emphasis id="B92EEAEEFFCCFA31FB13F8C9FB5D6D46" bold="true" box="[1169,1188,1840,1861]" pageId="8" pageNumber="7">c)</emphasis>
sternopectinal region;
<emphasis id="B92EEAEEFFCCFA31FA09F8C9FA596D46" bold="true" box="[1419,1440,1840,1861]" pageId="8" pageNumber="7">d)</emphasis>
sternite V;
<emphasis id="B92EEAEEFFCCFA31FEA9F8B5FEC46D62" bold="true" box="[299,317,1868,1889]" pageId="8" pageNumber="7">e)</emphasis>
metasomal segments IIII, lateral view;
<emphasis id="B92EEAEEFFCCFA31FD44F8B5FD2F6D62" bold="true" box="[710,726,1868,1889]" pageId="8" pageNumber="7">f)</emphasis>
metasomal segments IVV and telson, lateral view.
</paragraph>
</caption>
<caption id="DF256674FFCDFA30FF42FA2DFAD86FE8" ID-DOI="http://doi.org/10.5281/zenodo.4649097" ID-Zenodo-Dep="4649097" box="[192,1313,1492,1515]" httpUri="https://zenodo.org/record/4649097/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,268,1492,1515]" targetBox="[191,1440,191,1462]" targetPageId="9">
<paragraph id="8BE536FCFFCDFA30FF42FA2DFAD86FE8" blockId="9.[192,1313,1492,1515]" box="[192,1313,1492,1515]" pageId="9" pageNumber="8">
<emphasis id="B92EEAEEFFCDFA30FF42FA2DFED06FE8" bold="true" box="[192,297,1492,1515]" pageId="9" pageNumber="8">Figure 7:</emphasis>
Juvenile male of
<taxonomicName id="4C5A4D7FFFCDFA30FE51FA2FFDA66FE8" authorityName="Lourenco" authorityYear="1991" box="[467,607,1494,1515]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCDFA30FE51FA2FFDA66FE8" box="[467,607,1494,1515]" italics="true" pageId="9" pageNumber="8">Tityus tayrona</emphasis>
</taxonomicName>
from Turbaco:
<emphasis id="B92EEAEEFFCDFA30FD74FA2FFCF36FE8" bold="true" box="[758,778,1494,1515]" pageId="9" pageNumber="8">a)</emphasis>
entire dorsal view;
<emphasis id="B92EEAEEFFCDFA30FC4AFA2FFC246FE8" bold="true" box="[968,989,1494,1515]" pageId="9" pageNumber="8">b)</emphasis>
entire ventral view;
<emphasis id="B92EEAEEFFCDFA30FB26FA2FFB4E6FE8" bold="true" box="[1188,1207,1494,1515]" pageId="9" pageNumber="8">c)</emphasis>
sternite V.
</paragraph>
</caption>
<paragraph id="8BE536FCFFCDFA30FF73F8ECFAE96CED" blockId="9.[192,803,1568,1897]" lastBlockId="9.[830,1441,1568,1897]" pageId="9" pageNumber="8">
On the other hand, some wrong information that has been published on
<taxonomicName id="4C5A4D7FFFCDFA30FE1DF8CAFDEC6D49" authorityName="Lourenco" authorityYear="1991" box="[415,533,1843,1866]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B92EEAEEFFCDFA30FE1DF8CAFDEC6D49" box="[415,533,1843,1866]" italics="true" pageId="9" pageNumber="8">T. tayrona</emphasis>
</taxonomicName>
needs to be corrected. First, Rojas-Runjaic &amp; Armas (2002: 63) stated that “
<emphasis id="B92EEAEEFFCDFA30FC8CF8ABFC8B6C34" box="[782,882,1568,1897]" italics="true" pageId="9" pageNumber="8">... male</emphasis>
[...]
<emphasis id="B92EEAEEFFCDFA30FC3AF9D9FA926C34" box="[952,1387,1568,1591]" italics="true" pageId="9" pageNumber="8">has metasoma with the fifth segment</emphasis>
[...]
<emphasis id="B92EEAEEFFCDFA30FCBCF9C7FC526C56" box="[830,939,1598,1621]" italics="true" pageId="9" pageNumber="8">blackish...</emphasis>
” (original text in Spanish, translation and italics added), but this is incorrect according to the original description (Lourenço, 1991a: 280281, figs. 3 4), additional illustrated papers (Lourenço, 1997: 66, figs. 34; Botero-Trujillo &amp; Fagua, 2007: 132133, figs. 1215), and our specimens (
<figureCitation id="13612A79FFCDFA30FBEEF92EFB246CED" box="[1132,1245,1751,1774]" captionStart="Figure 3" captionStartId="5.[192,268,912,935]" captionTargetBox="[191,1440,191,882]" captionTargetId="figure-6@5.[192,1439,192,881]" captionTargetPageId="5" captionText="Figure 3: Adult male of Tityus tayrona from Turbaco: a) entire dorsal view; b) entire ventral view." figureDoi="http://doi.org/10.5281/zenodo.4649089" httpUri="https://zenodo.org/record/4649089/files/figure.png" pageId="9" pageNumber="8">Figs. 3ab</figureCitation>
,
<figureCitation id="13612A79FFCDFA30FB68F92EFAF96CED" box="[1258,1280,1751,1774]" captionStart="Figure 4" captionStartId="6.[192,268,1836,1859]" captionTargetBox="[191,1441,215,1801]" captionTargetId="figure-10@6.[192,1440,216,1800]" captionTargetPageId="6" captionText="Figure 4: Adult male of Tityus tayrona from Turbaco: a) carapace; b) pedipalp, dorsal view; c) sternopectinal region; d) sternite V; e) metasomal segments IIII, lateral view; f) metasomal segments IVV and telson, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4649091" httpUri="https://zenodo.org/record/4649091/files/figure.png" pageId="9" pageNumber="8">4f</figureCitation>
).
</paragraph>
<paragraph id="8BE536FCFFCDFA33FCF2F90FFE816F8C" blockId="9.[830,1441,1568,1897]" lastBlockId="10.[192,802,1308,1423]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
Second, Botero-Trujillo &amp; Fagua (2007: 133) stated that “
<emphasis id="B92EEAEEFFCDFA30FCFEF8ECFBC96D49" italics="true" pageId="9" pageNumber="8">... this species is characterized by the presence of 1417 pectinal teeth...</emphasis>
”, but this only applies to females as given in the original description by Lourenço (1991a: 281; tab. 2), which also gives a range of 1619 for males; our specimens confirm Lourenço's original data, and also allow to know that females may present up to 18 pectinal teeth.
</paragraph>
</subSubSection>
</treatment>
</document>