treatments-xml/data/03/F3/87/03F387C9405B2D17FF34F9DEFD8F4AC1.xml

<document id="629440161CDDE794D3D1FAC255864E55" ID-DOI="10.11646/phytotaxa.663.4.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="14515710" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734535385876" checkinUser="felipe" docAuthor="Vlasenko, Anastasia V., Enkhtuya, Ochirbat, Degidmaa, Turmunkh, Enkhjargal, Enkhtaivan, Chelobanov, Boris P. &amp; Vlasenko, Vyacheslav A." docDate="2024" docId="03F387C9405B2D17FF34F9DEFD8F4AC1" docLanguage="en" docName="phytotaxa.663.4.205-220.pdf" docOrigin="Phytotaxa 663 (4)" docSource="https://doi.org/10.11646/phytotaxa.663.4.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Tubifera khangaiensis A. Vlasenko 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="218" masterDocId="FFCAFFB140512D1AFFBCFFBCFFE3484C" masterDocTitle="Three new species of bright-spored myxomycetes from Asia and the second world record of Perichaena maculosa" masterLastPageNumber="220" masterPageNumber="205" pageNumber="215" updateTime="1734535807056" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C6801DC001D21D57C86A98493F89EF03">Three new species of bright-spored myxomycetes from Asia and the second world record of Perichaena maculosa</mods:title>
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<mods:namePart id="7C955ED981B00CEB762F0482EC0B10DD">Vlasenko, Anastasia V.</mods:namePart>
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<mods:namePart id="C0076BA5B4A606AE7115D1DE6E3B12DD">Enkhtuya, Ochirbat</mods:namePart>
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<mods:affiliation id="E7C7632C61D8E46EFE425F202B80D460">Botanical Garden and Research Institute of the Mongolian Academy of Sciences, 13330, Ulaanbaatar, Mongolia</mods:affiliation>
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<mods:namePart id="2AD406F812008CD48405CCF1D18D9FB8">Enkhjargal, Enkhtaivan</mods:namePart>
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<mods:namePart id="354D0D04D4F4509D1FA55328BF5D685B">Chelobanov, Boris P.</mods:namePart>
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<mods:affiliation id="78B44F3AFB62395E711D6EE196469085">Institute of Chemical Biology and Fundamental Medicine, Siberian Branch, Russian Academy of Sciences, 630090, Novosibirsk, Russia &amp; Novosibirsk State University, 630090, Novosibirsk, Russia</mods:affiliation>
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<mods:namePart id="317469F1FA76D821C37072D8E40DFAF7">Vlasenko, Vyacheslav A.</mods:namePart>
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<mods:affiliation id="E7A8AA75761B745F7A4EBB8F09FFE3D0">Central Siberian Botanical Garden, Siberian Branch, Russian Academy of Sciences, 630090 Novosibirsk, Russia</mods:affiliation>
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<mods:title id="A24A7F75DB57A2FE214B3E9F7A0C78C3">Phytotaxa</mods:title>
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<mods:date id="649AE0E1D531E7B55728A6A2D0988191">2024</mods:date>
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<taxonomicName id="4C5A4D5C405B2D10FF34F9DEFDFD4E30" authority="A. Vlasenko" authorityName="A. Vlasenko" authorityYear="2024" box="[136,542,1634,1661]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="10" pageNumber="215" phylum="Mycetozoa" rank="species" species="khangaiensis" status="sp. nov.">
<emphasis id="B92EEACD405B2D10FF34F9DEFE684E31" bold="true" box="[136,395,1634,1661]" italics="true" pageId="10" pageNumber="215">Tubifera khangaiensis</emphasis>
A. Vlasenko
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. 
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.
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(
<figureCitation id="13612A5A405B2D10FD37F9DEFD3A4E30" box="[651,729,1634,1660]" captionStart="FIGURE 8" captionStartId="11.[136,229,1839,1861]" captionTargetBox="[151,1436,190,1815]" captionTargetId="figure-22@11.[151,1436,190,1815]" captionTargetPageId="11" captionText="FIGURE 8. Tubifera khangaiensis (Holotype). A–B. Pseudoaethalia (RL). C. Part of pseudoaethalia (RL). D. Peridium and capillitium (RL). E–F. Spores on threads of capillitium (TL). G. Capillitium, free rounded node of capillitium and spores (TL). Scale bars:A–B = 2.5 mm, C = 2 mm, D = 1 mm, E = 100 µm, F = 15 µm, G = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.14515730" httpUri="https://zenodo.org/record/14515730/files/figure.png" pageId="10" pageNumber="215">Figs. 8</figureCitation>
, 
<figureCitation id="13612A5A405B2D10FD5AF9DEFD154E30" box="[742,758,1634,1660]" captionStart="FIGURE 9" captionStartId="12.[136,229,1772,1794]" captionTargetBox="[151,1436,281,1744]" captionTargetId="figure-75@12.[151,1436,281,1747]" captionTargetPageId="12" captionText="FIGURE 9. Tubifera khangaiensis (holotype). A. Spores on threads of capillitium (SEM). B–C. Capillitium and spores (SEM). D. Ornamentation of inner surface of the peridium (SEM). E–F. Spores (SEM). Scale bars: A = 100 µm, B = 10 µm, C = 20 µm, D = 10 µm, E–F = 1 µm." figureDoi="http://doi.org/10.5281/zenodo.14515735" httpUri="https://zenodo.org/record/14515735/files/figure.png" pageId="10" pageNumber="215">9</figureCitation>
).
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<paragraph id="8BE536DF405B2D10FF34F93AFE694EED" blockId="10.[136,394,1670,1697]" box="[136,394,1670,1697]" pageId="10" pageNumber="215">MycoBank no: 855058</paragraph>
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<emphasis id="B92EEACD405B2D10FF34F972FECD4EA4" bold="true" box="[136,302,1742,1768]" pageId="10" pageNumber="215">Etymology:—</emphasis>
Named after the Khangai Mountains in Mongolia.
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Pseudoaethalia sessile, umber, with a bluish tint, 
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diam., up to 
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tall, dehiscence irregularly. Individual sporangia in the form of irregular cylinders, curved, merging only in the lower part, middle and upper parts remain free, often expanding towards the upper part. Individual sporangia 
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tall, 0.8–1.1 diam. Hypothallus inconspicuous. Capillitium well developed, threads sinuous, branching, colorless in transmitted light, spores attached to the threads, individual threads up to 5–6 μm in diam., many sporangia free, irregularly rounded nodules are formed, smooth, reddish-brown in transmitted light, 10–40 μm diam. Spores in mass light umber, light brown, with reddish tint in transmitted light, spherical, 6–7 µm diam., ornamented with a mesh of 8–9 cells on the visible part of the spore. Plasmodium unknown.
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<emphasis id="B92EEACD405A2D11FF34F893FF1D4F08" bold="true" box="[136,254,1839,1861]" pageId="11" pageNumber="216">FIGURE 8.</emphasis>
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<emphasis id="B92EEACD405A2D11FEB9F893FE384F08" box="[261,475,1839,1860]" italics="true" pageId="11" pageNumber="216">Tubifera khangaiensis</emphasis>
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(Holotype). A–B. Pseudoaethalia (RL). C. Part of pseudoaethalia (RL). D. Peridium and capillitium (RL). E–F. Spores on threads of capillitium (TL). G. Capillitium, free rounded node of capillitium and spores (TL). Scale bars: A–B = 2.5 mm, C = 2 mm, D = 1 mm, E = 100 µm, F = 15 µm, G = 25 µm.
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:—
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: 
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, near 
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, larch forest, on fallen trunk of 
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<emphasis id="B92EEACD405D2D16FF1AFF7FFF004890" box="[166,227,195,220]" italics="true" pageId="12" pageNumber="217">Larix</emphasis>
sp.
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, 
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, 
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a.s.l.
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, field sample coll. 
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, 
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, ident. 
<collectionCode id="ED4BAE1A405D2D16FAC4FF7EFA6D4890" box="[1400,1422,194,220]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="217" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED4BAE1A405D2D16FA2EFF7EFA4B4890" box="[1426,1448,194,220]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="217" type="Museum">V</collectionCode>
. 
<collectorName id="26AF5309405D2D16FF34FF5AFF16494C" box="[136,245,230,256]" pageId="12" pageNumber="217">Vlasenko</collectorName>
, 
<specimenCode id="DBFC9EA4405D2D16FEBCFF5AFE47494C" box="[256,420,230,256]" collectionCode="NSK" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15284" lsid="urn:lsid:biocol.org:col:15284" name="Central Siberian Botanical Garden, Siberian Branch of Russian Academy of Sciensis" pageId="12" pageNumber="217" type="Botanical Garden (with or without associated herbarium)">NSK 1013200</specimenCode>
, 
<collectorName id="26AF5309405D2D16FE13FF5AFDAE494D" box="[431,589,230,257]" pageId="12" pageNumber="217">GenBank No.</collectorName>
: PQ032479.
</materialsCitation>
</paragraph>
<caption id="DF256657405D2D16FF34F950FEE04F06" ID-DOI="http://doi.org/10.5281/zenodo.14515735" ID-Zenodo-Dep="14515735" httpUri="https://zenodo.org/record/14515735/files/figure.png" pageId="12" pageNumber="217" startId="12.[136,229,1772,1794]" targetBox="[151,1436,281,1744]" targetPageId="12" targetType="figure">
<paragraph id="8BE536DF405D2D16FF34F950FEE04F06" blockId="12.[136,1452,1772,1866]" pageId="12" pageNumber="217">
<emphasis id="B92EEACD405D2D16FF34F950FEE24F4D" bold="true" box="[136,257,1772,1794]" pageId="12" pageNumber="217">FIGURE 9.</emphasis>
<taxonomicName id="4C5A4D5C405D2D16FEB7F950FE004F4D" authorityName="A. Vlasenko" authorityYear="2024" box="[267,483,1772,1793]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="12" pageNumber="217" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405D2D16FEB7F950FE004F4D" box="[267,483,1772,1793]" italics="true" pageId="12" pageNumber="217">Tubifera khangaiensis</emphasis>
</taxonomicName>
(holotype). A. Spores on threads of capillitium (SEM). B–C. Capillitium and spores (SEM). D. Ornamentation of inner surface of the peridium (SEM). E–F. Spores (SEM). Scale bars: A = 100 µm, B = 10 µm, C = 20 µm, D = 10 µm, E–F = 1 µm.
</paragraph>
</caption>
<paragraph id="8BE536DF405D2D16FF01F8C4FCA64FB3" blockId="12.[136,1451,1912,2047]" pageId="12" pageNumber="217">
<emphasis id="B92EEACD405D2D16FF01F8C4FDFB4FDE" bold="true" box="[189,536,1912,1938]" pageId="12" pageNumber="217">Other specimen examined:—</emphasis>
<collectingCountry id="F34D764F405D2D16FDA4F8C5FD574FDF" box="[536,692,1913,1939]" name="Mongolia" pageId="12" pageNumber="217">MONGOLIA</collectingCountry>
. 
<collectingRegion id="499EF83D405D2D16FD01F8C4FCDB4FDE" box="[701,824,1912,1938]" country="Mongolia" name="Arhangay" pageId="12" pageNumber="217">Arkhangai</collectingRegion>
: Khangai Mountains, 
<quantity id="4CA29B3A405D2D16FB90F8C5FB924FDE" box="[1068,1137,1912,1939]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="217" unit="km" value="11.0">11 km</quantity>
southwest of Khunt village, larch forest, on fallen trunk of 
<taxonomicName id="4C5A4D5C405D2D16FE5FF821FDA94FFB" box="[483,586,1949,1975]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="12" pageNumber="217" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B92EEACD405D2D16FE5FF821FDC34FFA" box="[483,544,1949,1974]" italics="true" pageId="12" pageNumber="217">Larix</emphasis>
sp.
</taxonomicName>
, 
<geoCoordinate id="EE6E5018405D2D16FDEBF820FD084FFB" box="[599,747,1948,1975]" degrees="47" direction="north" minutes="46" orientation="latitude" pageId="12" pageNumber="217" precision="15" seconds="32" value="47.775555">47°46’32” N</geoCoordinate>
, 
<geoCoordinate id="EE6E5018405D2D16FD4BF820FC674FFB" box="[759,900,1948,1975]" degrees="99" direction="east" minutes="16" orientation="longitude" pageId="12" pageNumber="217" precision="15" seconds="45" value="99.27917">99°16’45” E</geoCoordinate>
, 
<quantity id="4CA29B3A405D2D16FC2FF821FC0A4FFB" box="[915,1001,1949,1975]" metricMagnitude="3" metricUnit="m" metricValue="2.2009999999999996" pageId="12" pageNumber="217" unit="m" value="2201.0">2201 m</quantity>
a.s.l., field sample coll. 
<date id="FFE4101F405D2D16FB43F821FA444FFB" box="[1279,1447,1949,1975]" pageId="12" pageNumber="217" value="2023-08-08">8 August 2023</date>
, 
<collectionCode id="ED4BAE1A405D2D16FF34F87DFF7D4F97" box="[136,158,1985,2011]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="217" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED4BAE1A405D2D16FF1EF87DFF5B4F97" box="[162,184,1985,2011]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="217" type="Museum">V</collectionCode>
. Vlasenko, ident. 
<collectionCode id="ED4BAE1A405D2D16FEC2F87DFE774F97" box="[382,404,1985,2011]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="12" pageNumber="217" type="Herbarium">A</collectionCode>
. 
<collectionCode id="ED4BAE1A405D2D16FE2BF87DFE4E4F97" box="[407,429,1985,2011]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="217" type="Museum">V</collectionCode>
. Vlasenko, 
<collectionCode id="ED4BAE1A405D2D16FD93F87DFD844F97" box="[559,615,1985,2011]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15284" lsid="urn:lsid:biocol.org:col:15284" name="Central Siberian Botanical Garden, Siberian Branch of Russian Academy of Sciensis" pageId="12" pageNumber="217" type="Botanical Garden (with or without associated herbarium)">NSK</collectionCode>
1013201, GenBank No.: PQ032480. 
<emphasis id="B92EEACD405D2D16FF01F859FEA14FB3" bold="true" box="[189,322,2021,2047]" pageId="12" pageNumber="217">Eсology:—</emphasis>
Grows as a xylobiont on fallen trunk of 
<taxonomicName id="4C5A4D5C405D2D16FCBEF859FCDC4FB2" box="[770,831,2021,2046]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="12" pageNumber="217" phylum="Tracheophyta" rank="genus">
<emphasis id="B92EEACD405D2D16FCBEF859FCDC4FB2" box="[770,831,2021,2046]" italics="true" pageId="12" pageNumber="217">Larix</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3406554405C2D17FF01FF22FD8F4AC1" pageId="13" pageNumber="218" type="discussion">
<paragraph id="8BE536DF405C2D17FF01FF22FE1E4A6D" blockId="13.[136,1452,158,653]" pageId="13" pageNumber="218">
<emphasis id="B92EEACD405C2D17FF01FF22FE8748F4" bold="true" box="[189,356,158,184]" pageId="13" pageNumber="218">Comments:—</emphasis>
The colour of the pseudoaethalia of 
<taxonomicName id="4C5A4D5C405C2D17FCB7FF22FBE548F4" authorityName="A. Vlasenko" authorityYear="2024" box="[779,1030,158,184]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FCB7FF22FBE548F4" box="[779,1030,158,184]" italics="true" pageId="13" pageNumber="218">Tubifera khangaiensis</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C5A4D5C405C2D17FB36FF23FA4748F5" authority="Nann." authorityName="Nann." box="[1162,1444,158,185]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="dimorphotheca">
<emphasis id="B92EEACD405C2D17FB36FF23FB7948F4" box="[1162,1178,159,184]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FB17FF22FAB648F4" box="[1195,1365,158,184]" italics="true" pageId="13" pageNumber="218">dimorphotheca</emphasis>
Nann.
</taxonomicName>
- Bremek. &amp; Loer., 
<taxonomicName id="4C5A4D5C405C2D17FEE9FF7FFC7C4890" authority="(Berk. &amp; M. A. Curtis) G. W. Martin" authorityName="G. W. Martin" authorityYear="1947" baseAuthorityName="Berk. &amp; M. A. Curtis" box="[341,927,194,221]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="microsperma">
<emphasis id="B92EEACD405C2D17FEE9FF7FFE864890" box="[341,357,195,220]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FECFFF7FFDE54890" box="[371,518,195,220]" italics="true" pageId="13" pageNumber="218">microsperma</emphasis>
(Berk. &amp; M.A. Curtis) G.W. Martin
</taxonomicName>
, 
<taxonomicName id="4C5A4D5C405C2D17FC17FF7FFF13494D" authority="G. W. Martin, K. S. Thind &amp; Sohi. However" authorityName="G. W. Martin, K. S. Thind &amp; Sohi. However" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="papillata">
<emphasis id="B92EEACD405C2D17FC17FF7FFC584890" box="[939,955,195,220]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FC75FF7EFBCD4890" box="[969,1070,194,220]" italics="true" pageId="13" pageNumber="218">papillata</emphasis>
G.W. Martin, K.S. Thind &amp; Sohi. However
</taxonomicName>
, the pseudoaethalia of these species have a clearly visible stalk-like structure from the hypothallus (false stalk), whereas the pseudoaethalia of 
<taxonomicName id="4C5A4D5C405C2D17FD9AFEB7FD304968" authorityName="A. Vlasenko" authorityYear="2024" box="[550,723,266,292]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FD9AFEB7FD304968" box="[550,723,266,292]" italics="true" pageId="13" pageNumber="218">T. khangaiensis</emphasis>
</taxonomicName>
are sessile, the hypothallus is inconspicuous, and the false stalk is absent. Sporangia of 
<taxonomicName id="4C5A4D5C405C2D17FEC8FE93FDC14904" authorityName="A. Vlasenko" authorityYear="2024" box="[372,546,302,328]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FEC8FE93FDC14904" box="[372,546,302,328]" italics="true" pageId="13" pageNumber="218">T. khangaiensis</emphasis>
</taxonomicName>
are connected to neighboring sporangia only in the lower part, in the middle and upper parts they remain free, often individual sporangia expand in the upper part, which distinguishes the new species from all previously described species in the genus 
<taxonomicName id="4C5A4D5C405C2D17FD0BFECAFCF549DC" box="[695,790,374,400]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="genus">
<emphasis id="B92EEACD405C2D17FD0BFECAFCF549DC" box="[695,790,374,400]" italics="true" pageId="13" pageNumber="218">Tubifera</emphasis>
</taxonomicName>
. The presence of free, irregularly shaped nodules inside the sporangia in 
<taxonomicName id="4C5A4D5C405C2D17FEA7FE27FE2A49F8" authorityName="A. Vlasenko" authorityYear="2024" box="[283,457,410,436]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FEA7FE27FE2A49F8" box="[283,457,410,436]" italics="true" pageId="13" pageNumber="218">T. khangaiensis</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C5A4D5C405C2D17FDE3FE27FCB649F8" authority="Nann." authorityName="Nann." box="[607,853,410,437]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="dictyoderma">
<emphasis id="B92EEACD405C2D17FDE3FE27FD8C49F8" box="[607,623,411,436]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FDC2FE26FCE949F8" box="[638,778,410,436]" italics="true" pageId="13" pageNumber="218">dictyoderma</emphasis>
Nann.
</taxonomicName>
-Bremek. &amp; Loer and 
<taxonomicName id="4C5A4D5C405C2D17FBEDFE27FE744995" authority="G. W. Martin, K. S. Thind &amp; Sohi. However" authorityName="G. W. Martin, K. S. Thind &amp; Sohi. However" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="papillata">
<emphasis id="B92EEACD405C2D17FBEDFE27FB8249F8" box="[1105,1121,411,436]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FBD3FE26FB3749F8" box="[1135,1236,410,436]" italics="true" pageId="13" pageNumber="218">papillata</emphasis>
G.W. Martin, K.S. Thind &amp; Sohi. However
</taxonomicName>
, the spores of 
<taxonomicName id="4C5A4D5C405C2D17FD8AFE03FD3E4994" box="[566,733,446,472]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="dictyoderma">
<emphasis id="B92EEACD405C2D17FD8AFE03FD3E4994" box="[566,733,446,472]" italics="true" pageId="13" pageNumber="218">T. dictyoderma</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5A4D5C405C2D17FCAFFE03FC764994" box="[787,917,446,473]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="papillata">
<emphasis id="B92EEACD405C2D17FCAFFE03FCC04994" box="[787,803,447,472]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FC8CFE02FC764994" box="[816,917,446,472]" italics="true" pageId="13" pageNumber="218">papillata</emphasis>
</taxonomicName>
are smaller and ornamented with a coarse mesh with 3–4 cells per visible part of the spore, whereas the spores of 
<taxonomicName id="4C5A4D5C405C2D17FCD4FE5FFBFB49B0" authorityName="A. Vlasenko" authorityYear="2024" box="[872,1048,482,509]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FCD4FE5FFC9B49B0" box="[872,888,483,508]" italics="true" pageId="13" pageNumber="218">T</emphasis>
. 
<emphasis id="B92EEACD405C2D17FC3AFE5EFBFB49B0" box="[902,1048,482,508]" italics="true" pageId="13" pageNumber="218">khangaiensis</emphasis>
</taxonomicName>
are ornamented with a mesh of 8–9 cells per visible part of the spore.
</paragraph>
<paragraph id="8BE536DF405C2D17FF78FD96FD8F4AC1" blockId="13.[136,1452,158,653]" pageId="13" pageNumber="218">
The ML analysis based on the 18S nrDNA region (
<figureCitation id="13612A5A405C2D17FD47FD96FCAE4A08" box="[763,845,554,580]" captionStart="FIGURE 10" captionStartId="13.[136,229,1474,1496]" captionTargetBox="[151,1428,685,1441]" captionTargetId="figure-274@13.[151,1436,677,1450]" captionTargetPageId="13" captionText="FIGURE 10. Maximum likelihood tree based on 18S nrDNA sequences shows the phylogenetic relationships between Tubifera khangaiensis and closely related species of the genus Tubifera. BS values with high support and Bayesian posterior probabilities are shown in the branches. GenBank accession numbers are given after the species name." figureDoi="http://doi.org/10.5281/zenodo.14515737" httpUri="https://zenodo.org/record/14515737/files/figure.png" pageId="13" pageNumber="218">Fig. 10</figureCitation>
) showed that 
<taxonomicName id="4C5A4D5C405C2D17FC55FD96FB024A08" authorityName="A. Vlasenko" authorityYear="2024" box="[1001,1249,554,580]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FC55FD96FB024A08" box="[1001,1249,554,580]" italics="true" pageId="13" pageNumber="218">Tubifera khangaiensis</emphasis>
</taxonomicName>
is an independent species, closest to 
<taxonomicName id="4C5A4D5C405C2D17FEE6FDF3FD3F4A24" authority="Leontyev &amp; Fefelov." authorityName="Leontyev &amp; Fefelov." box="[346,732,590,617]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="applanata">
<emphasis id="B92EEACD405C2D17FEE6FDF3FE0B4A24" box="[346,488,590,616]" italics="true" pageId="13" pageNumber="218">T. applanata</emphasis>
Leontyev &amp; Fefelov.
</taxonomicName>
The genetic distance of the 
<taxonomicName id="4C5A4D5C405C2D17FBA1FDF3FB2F4A24" authorityName="A. Vlasenko" authorityYear="2024" box="[1053,1228,590,616]" class="Dictyosteliomycetes" family="Raperosteliaceae" genus="Tubifera" kingdom="Protozoa" order="Dictyosteliales" pageId="13" pageNumber="218" phylum="Mycetozoa" rank="species" species="khangaiensis">
<emphasis id="B92EEACD405C2D17FBA1FDF3FB2F4A24" box="[1053,1228,590,616]" italics="true" pageId="13" pageNumber="218">T. khangaiensis</emphasis>
</taxonomicName>
branch on the SSU tree is 0.138, with 100% bootstrap support.
</paragraph>
</subSubSection>
</treatment>
</document>