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<document id="5823214E308B33689B36717EA7D18D05" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A3FFFACFC39020E3696B192" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria saurahana Fikáček &amp; Liu 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="247" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="246" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="BDA2148761C3BBDDB6F6E8A9AEFE8E6D">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
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<mods:namePart id="2CA6E79611803729CA79D474CB681442">Fikáček, Martin</mods:namePart>
<mods:affiliation id="33E8A1F7FF64A46847238A1955140DDC">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="B0D5B57B277FC7DB8BCA6BE4671394BB">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="108D4A06F461A43DC7F72EC670956D43">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
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<mods:title id="14E674F8BB0FA76D4FD945A5E79D7718">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="81C44DBF59C8959260D546BEB63BDB0A">2019</mods:date>
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<treatment id="03F387C57A3FFFACFC39020E3696B192" ID-DOI="http://doi.org/10.5281/zenodo.4548841" ID-GBIF-Taxon="177423410" ID-Zenodo-Dep="4548841" LSID="urn:lsid:plazi:treatment:03F387C57A3FFFACFC39020E3696B192" httpUri="http://treatment.plazi.org/id/03F387C57A3FFFACFC39020E3696B192" lastPageId="18" lastPageNumber="247" pageId="17" pageNumber="246">
<subSubSection id="C34065587A3FFFAFFC39020E3119B434" box="[944,1320,806,829]" pageId="17" pageNumber="246" type="nomenclature">
<paragraph id="8BE536D37A3FFFAFFC39020E3119B434" blockId="17.[944,1320,806,829]" box="[944,1320,806,829]" pageId="17" pageNumber="246">
<heading id="D0AD81BF7A3FFFAFFC39020E3119B434" box="[944,1320,806,829]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="246" reason="3">
<emphasis id="B92EEAC17A3FFFAFFC39020E3119B434" bold="true" box="[944,1320,806,829]" pageId="17" pageNumber="246">
<taxonomicName id="4C5A4D507A3FFFAFFC39020E30FDB434" ID-CoL="8YBVW" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[944,1228,806,829]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="saurahana" status="sp. nov.">
<emphasis id="B92EEAC17A3FFFAFFC39020E30FDB434" bold="true" box="[944,1228,806,829]" italics="true" pageId="17" pageNumber="246">Thysanarthria saurahana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3FFFAFFB5A020E3119B434" box="[1235,1320,806,829]" pageId="17" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFBAE026C3080B45E" box="[1063,1201,836,855]" pageId="17" pageNumber="246" type="description">
<paragraph id="8BE536D37A3FFFAFFBAE026C3080B45E" blockId="17.[828,1437,836,1003]" box="[1063,1201,836,855]" pageId="17" pageNumber="246">
(
<figureCitation id="13612A567A3FFFAFFBA4026C3053B45E" box="[1069,1122,836,855]" captionStart="Fig" captionStartId="18.[151,182,1467,1486]" captionTargetBox="[178,1432,202,1437]" captionTargetId="figure-13@18.[151,1436,186,1450]" captionTargetPageId="18" captionText="Fig. 9. Male genitalia of Thysanarthria species, holotypes. AE T. saurahana sp. nov. (AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. siamensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488942" httpUri="https://zenodo.org/record/4488942/files/figure.png" pageId="17" pageNumber="246">Figs 9</figureCitation>
AE,
<figureCitation id="13612A567A3FFFAFFB1C026C309BB45E" box="[1173,1194,836,855]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (AB) and Near East (C), and examples of habitats of Thysanarthria (DF). Localities: D India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; FG Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, FG by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="17" pageNumber="246">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFCB5024630F4B4E2" pageId="17" pageNumber="246" type="materials_examined">
<paragraph id="8BE536D37A3FFFAFFCB5024630F4B4E2" blockId="17.[828,1437,836,1003]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFCB5024637F6B488" bold="true" box="[828,967,878,897]" pageId="17" pageNumber="246">Type material.</emphasis>
<materialsCitation id="3B323C8E7A3FFFAFFC46024630F4B4E2" ID-GBIF-Occurrence="3756360354" collectingDate="2000-04-18" collectingMethod="light trap" collectionCode="SMNS" collectorName="A. Weigel" country="Nepal" determinerName="F. Hebauer'" latitude="84.49695" location="Rapti River bank" longLatPrecision="1" longitude="27.56667" municipality="Sauraha" pageId="17" pageNumber="246" specimenCount="1" specimenCount-male="1" stateProvince="Narayani" typeStatus="holotype">
<typeStatus id="54E188717A3FFFAFFC4602463014B476" box="[975,1061,878,897]" pageId="17" pageNumber="246" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5CFD5A7A3FFFAFFBBD024F3075B489" box="[1076,1092,871,896]" pageId="17" pageNumber="246" type="male"></specimenCount>
(
<collectionCode id="ED4BAE167A3FFFAFFBDB024630A2B488" box="[1106,1171,878,897]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="17" pageNumber="246">SMNS</collectionCode>
):
<emphasis id="B92EEAC17A3FFFAFFB2D024630C5B488" bold="true" box="[1188,1268,878,897]" pageId="17" pageNumber="246">
<collectingCountry id="F34D76437A3FFFAFFB2D024630C1B488" box="[1188,1264,878,897]" name="Nepal" pageId="17" pageNumber="246">NEPAL</collectingCountry>
:
</emphasis>
ʻNepal,
<collectingRegion id="499EF8317A3FFFAFFACF024631A9B488" box="[1350,1432,878,897]" country="Nepal" name="Narayani" pageId="17" pageNumber="246">Narayani</collectingRegion>
,
<collectingMunicipality id="6B81ACA97A3FFFAFFCB502A037B2B492" box="[828,899,904,923]" pageId="17" pageNumber="246">Sauraha</collectingMunicipality>
,
<location id="8E8560087A3FFFAFFC0502A03012B492" LSID="urn:lsid:plazi:treatment:03F387C57A3FFFACFC39020E3696B192:8E8560087A3FFFAFFC0502A03012B492" box="[908,1059,904,923]" country="Nepal" latitude="84.49695" longLatPrecision="1" longitude="27.56667" municipality="Sauraha" name="Rapti River bank" pageId="17" pageNumber="246" stateProvince="Narayani">Rapti River bank</location>
,
<collectingMethod id="521B4EC47A3FFFAFFBA502A0304CB492" box="[1068,1149,904,923]" pageId="17" pageNumber="246">light trap</collectingMethod>
, 180,
<geoCoordinate id="EE6E50147A3FFFAFFB3A02A03134B492" box="[1203,1285,904,923]" degrees="84.49695" direction="north" orientation="latitude" pageId="17" pageNumber="246" precision="1" value="84.49695">84.49695</geoCoordinate>
,
<geoCoordinate id="EE6E50147A3FFFAFFA8602A03150B492" box="[1295,1377,904,923]" degrees="27.56667" direction="east" orientation="longitude" pageId="17" pageNumber="246" precision="1" value="27.56667">27.56667</geoCoordinate>
,
<date id="FFE410137A3FFFAFFAE302A0375EB4BF" pageId="17" pageNumber="246" value="2000-04-18">
<collectingDate id="EFA0E9FB7A3FFFAFFAE302A0375EB4BF" pageId="17" pageNumber="246" value="2000-04-18">2000- 04-18</collectingDate>
</date>
,
<collectorName id="26AF53057A3FFFAFFCFF028B37FCB4BF" box="[886,973,931,950]" pageId="17" pageNumber="246">A. Weigel</collectorName>
, NEPAL, Prov. Narayani / Sauraha, Rapti River / Ufer, 180mNN, 27°34´80´´N, 84°29´49´´E / LF,
<date id="FFE410137A3FFFAFFB3D02963122B4D8" box="[1204,1299,958,977]" pageId="17" pageNumber="246" value="2000-04-18">18.IV.2000</date>
/ leg. Weigel //
<taxonomicName id="4C5A4D507A3FFFAFFCB502F03783B4E2" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[828,946,984,1003]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="252" phylum="Arthropoda" rank="genus" status="gen. nov.">Thysanarthria</taxonomicName>
/ madurensis / det.
<determinerName id="7E39DA177A3FFFAFFBD002F030F0B4E2" box="[1113,1217,984,1003]" pageId="17" pageNumber="246">F. Hebauer</determinerName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFCB5052C3107B1B2" pageId="17" pageNumber="246" type="description">
<paragraph id="8BE536D37A3FFFAFFCB5052C3107B1B2" blockId="17.[828,1437,1028,2075]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFCB5052C3035B312" bold="true" box="[828,1028,1028,1051]" pageId="17" pageNumber="246">
Description.
<emphasis id="B92EEAC17A3FFFAFFC45052C3035B312" bold="true" box="[972,1028,1028,1051]" italics="true" pageId="17" pageNumber="246">Body</emphasis>
</emphasis>
length
<quantity id="4CA29B367A3FFFAFFBDD052C3096B312" box="[1108,1191,1028,1051]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="246" unit="mm" value="1.7">1.7 mm</quantity>
, maximum body width 1.0 mm. Head and labrum black; uniformly yellowish; elytra uniformly yellowish; legs yellowish.
<emphasis id="B92EEAC17A3FFFAFFAAF056C3153B352" bold="true" box="[1318,1378,1092,1115]" italics="true" pageId="17" pageNumber="246">Head</emphasis>
with strong mesh-like microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 2.7× the width of one eye in dorsal view.
<emphasis id="B92EEAC17A3FFFAFFAA4058C31ADB3B2" bold="true" box="[1325,1436,1188,1211]" italics="true" pageId="17" pageNumber="246">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices with strong mesh-like microsculpture.
<emphasis id="B92EEAC17A3FFFAFFAD105CC31ADB3F2" bold="true" box="[1368,1436,1252,1275]" italics="true" pageId="17" pageNumber="246">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (widely around scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
<emphasis id="B92EEAC17A3FFFAFFB7104AC3153B292" bold="true" box="[1272,1378,1412,1435]" italics="true" pageId="17" pageNumber="246">Aedeagus</emphasis>
(
<figureCitation id="13612A567A3FFFAFFAE404AC3779B2B2" captionStart="Fig" captionStartId="18.[151,182,1467,1486]" captionTargetBox="[178,1432,202,1437]" captionTargetId="figure-13@18.[151,1436,186,1450]" captionTargetPageId="18" captionText="Fig. 9. Male genitalia of Thysanarthria species, holotypes. AE T. saurahana sp. nov. (AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ T. siamensis Hebauer, 2001 (FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488942" httpUri="https://zenodo.org/record/4488942/files/figure.png" pageId="17" pageNumber="246">Figs 9</figureCitation>
AE)
<quantity id="4CA29B367A3FFFAFFC03048C37EAB2B2" box="[906,987,1444,1467]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="246" unit="mm" value="0.5">0.5 mm</quantity>
long. Phallobase slightly widened at base of parameres, c. as wide as parameres combined, slightly narrowed towards basal fifth and abruptly widened at base; arcuate in lateral view. Paramere narrowly elongate, subequal in width throughout, apex abruptly narrowing into a short mesoapical ʻtooth. Median lobe much shorter than parameres, with strongly sclerotized shorter part arcuate apically, and membranous apical part rounded apically; gonopore large, triangular, situated subapically.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFAFFCB507EC3066B0D2" pageId="17" pageNumber="246" type="diagnosis">
<paragraph id="8BE536D37A3FFFAFFCB507EC3066B0D2" blockId="17.[828,1437,1028,2075]" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFCB507EC3008B1D2" bold="true" box="[828,1081,1732,1755]" pageId="17" pageNumber="246">Differential diagnosis.</emphasis>
<taxonomicName id="4C5A4D507A3FFFAFFBCB07EC3167B1D2" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1090,1366,1732,1755]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="saurahana">
<emphasis id="B92EEAC17A3FFFAFFBCB07EC3167B1D2" box="[1090,1366,1732,1755]" italics="true" pageId="17" pageNumber="246">Thysanarthria saurahana</emphasis>
</taxonomicName>
is the only species in Himalaya with strong mesh-like microsculpture on the pronotum (in contrast to pronotum without microsculpture in
<taxonomicName id="4C5A4D507A3FFFAFFC72060C30B5B032" authorityName="Knisch" authorityYear="1924" box="[1019,1156,1828,1851]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B92EEAC17A3FFFAFFC72060C30B5B032" box="[1019,1156,1828,1851]" italics="true" pageId="17" pageNumber="246">T. championi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D507A3FFFAFFB3C060C3107B032" authorityName="Hebauer" authorityYear="2001" box="[1205,1334,1828,1851]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B92EEAC17A3FFFAFFB3C060C3107B032" box="[1205,1334,1828,1851]" italics="true" pageId="17" pageNumber="246">T. siamensis</emphasis>
</taxonomicName>
, and with weak granulate microsculpture in
<taxonomicName id="4C5A4D507A3FFFAFFB17066C311EB052" authorityName="Hebauer" authorityYear="2001" box="[1182,1327,1860,1883]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="species" species="madurensis">
<emphasis id="B92EEAC17A3FFFAFFB17066C311EB052" box="[1182,1327,1860,1883]" italics="true" pageId="17" pageNumber="246">T. madurensis</emphasis>
</taxonomicName>
). It can be distinguished from all these species as well as from all other
<taxonomicName id="4C5A4D507A3FFFAFFCB506AC37E4B092" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[828,981,1924,1947]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="246" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3FFFAFFCB506AC37E4B092" box="[828,981,1924,1947]" italics="true" pageId="17" pageNumber="246">Thysanarthria</emphasis>
</taxonomicName>
by the shape of parameres and the unique morphology of the median lobe which is not similar to any other species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3FFFACFCB506CC3647B152" lastPageId="18" lastPageNumber="247" pageId="17" pageNumber="246" type="etymology">
<paragraph id="8BE536D37A3FFFACFCB506CC3647B152" blockId="17.[828,1437,1028,2075]" lastBlockId="18.[151,679,1604,1691]" lastPageId="18" lastPageNumber="247" pageId="17" pageNumber="246">
<emphasis id="B92EEAC17A3FFFAFFCB506CC378DB0F2" bold="true" box="[828,956,2020,2043]" pageId="17" pageNumber="246">Etymology.</emphasis>
The species name refers to the village of Sauraha at the border of the Chitwan National Park in
<collectingCountry id="F34D76437A3FFFAFFAD4092C31ACBF12" box="[1373,1437,2052,2075]" name="Nepal" pageId="17" pageNumber="246">Nepal</collectingCountry>
where the
<typeStatus id="54E188717A3CFFACFE8F076C3552B152" box="[262,355,1604,1627]" pageId="18" pageNumber="247" type="holotype">holotype</typeStatus>
was collected. Adjective.
</paragraph>
</subSubSection>
<caption id="DF25665B7A3CFFACFF1E0493375AB10D" ID-DOI="http://doi.org/10.5281/zenodo.4488942" ID-Zenodo-Dep="4488942" httpUri="https://zenodo.org/record/4488942/files/figure.png" pageId="18" pageNumber="247" startId="18.[151,182,1467,1486]" targetBox="[178,1432,202,1437]" targetPageId="18">
<paragraph id="8BE536D37A3CFFACFF1E0493375AB10D" blockId="18.[151,1437,1467,1540]" pageId="18" pageNumber="247">
Fig. 9. Male genitalia of
<taxonomicName id="4C5A4D507A3CFFACFEF8049335DAB2C7" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[369,491,1467,1486]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="247" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3CFFACFEF8049335DAB2C7" box="[369,491,1467,1486]" italics="true" pageId="18" pageNumber="247">Thysanarthria</emphasis>
</taxonomicName>
species, holotypes. AE
<taxonomicName id="4C5A4D507A3CFFACFD500493377BB2C7" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[729,842,1467,1486]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="247" phylum="Arthropoda" rank="species" species="saurahana" status="sp. nov.">
<emphasis id="B92EEAC17A3CFFACFD500493377BB2C7" box="[729,842,1467,1486]" italics="true" pageId="18" pageNumber="247">T. saurahana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3CFFACFCD8049337A4B2C7" box="[849,917,1467,1486]" pageId="18" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
(AC whole aedeagus in dorsal, lateral and ventral view; DE detail of median lobe and parameres in dorsal and ventral view). FJ
<taxonomicName id="4C5A4D507A3CFFACFCA704FE3026B2E0" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[814,1047,1494,1513]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="247" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B92EEAC17A3CFFACFCA704FE37A4B2E0" box="[814,917,1494,1513]" italics="true" pageId="18" pageNumber="247">T. siamensis</emphasis>
<bibRefCitation id="EFCB4B227A3CFFACFC1304FE3026B2E0" author="Hebauer" box="[922,1047,1494,1513]" firstAuthor="Hebauer" pageId="18" pageNumber="247" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</taxonomicName>
(FH whole aedeagus in dorsal, lateral and ventral view; IJ details of parameres and median lobe in dorsal and ventral view).
</paragraph>
</caption>
<subSubSection id="C34065587A3CFFACFF1E074C355BB172" box="[151,362,1636,1659]" pageId="18" pageNumber="247" type="biology_ecology">
<paragraph id="8BE536D37A3CFFACFF1E074C355BB172" blockId="18.[151,679,1604,1691]" box="[151,362,1636,1659]" pageId="18" pageNumber="247">
<emphasis id="B92EEAC17A3CFFACFF1E074C34C2B172" bold="true" box="[151,243,1636,1659]" pageId="18" pageNumber="247">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3CFFACFF1E07AC3696B192" box="[151,679,1668,1691]" pageId="18" pageNumber="247" type="distribution">
<paragraph id="8BE536D37A3CFFACFF1E07AC3696B192" blockId="18.[151,679,1604,1691]" box="[151,679,1668,1691]" pageId="18" pageNumber="247">
<emphasis id="B92EEAC17A3CFFACFF1E07AC351BB192" bold="true" box="[151,298,1668,1691]" pageId="18" pageNumber="247">Distribution.</emphasis>
Only known from the
<typeStatus id="54E188717A3CFFACFD9407AC367AB192" box="[541,587,1668,1691]" pageId="18" pageNumber="247">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>