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<document id="F200E1DBEECF63D6638E31622B46D7A5" ID-CLB-Dataset="32316" ID-DOI="10.11646/zootaxa.4311.3.2" ID-GBIF-Dataset="38da213d-7ad2-4afe-bc41-a9087c9e6555" ID-ISSN="1175-5326" ID-Zenodo-Dep="847854" ID-ZooBank="F6Bf844D-60Fe-4D1F-8Fca-Dfbd533F0770" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503550462087" checkinUser="plazi" docAuthor="Achterberg, Cornelis Van" docDate="2017" docId="03F3372EFF98FFB1C387C87FFAA2389B" docLanguage="en" docName="zootaxa.4311.3.2.pdf" docOrigin="Zootaxa 4311 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yezoceryx pacoensis Achterberg 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="359" masterDocId="FFCA4F56FF94FFBFC310CC2FFF9B3A1F" masterDocTitle="First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species" masterLastPageNumber="372" masterPageNumber="345" pageNumber="357" updateTime="1698467142797" updateUser="ExternalLinkService">
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<mods:title id="73779425FD7B81CCF1A70C390BDD45B3">First record of the genus Yezoceryx Uchida (Ichneumonidae: Acaenitinae) from Vietnam, with descriptions of nine new species</mods:title>
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<mods:namePart id="E5DEBCB76BC09266F1B42F901E51DA2F">Achterberg, Cornelis Van</mods:namePart>
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<mods:title id="C8A35EC337514E53AB17DC95623817E0">Zootaxa</mods:title>
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<mods:date id="A334533DB2612A75E45A44C4B19695C4">2017</mods:date>
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<treatment id="03F3372EFF98FFB1C387C87FFAA2389B" ID-DOI="http://doi.org/10.5281/zenodo.6051723" ID-GBIF-Taxon="132923412" ID-Zenodo-Dep="6051723" LSID="urn:lsid:plazi:treatment:03F3372EFF98FFB1C387C87FFAA2389B" httpUri="http://treatment.plazi.org/id/03F3372EFF98FFB1C387C87FFAA2389B" lastPageId="14" lastPageNumber="359" pageId="12" pageNumber="357">
<subSubSection id="C340D5B3FF98FFB3C387C87FFE933E93" pageId="12" pageNumber="357" type="nomenclature">
<paragraph id="8BE58638FF98FFB3C387C87FFE7B3E77" blockId="12.[151,480,1103,1165]" box="[151,480,1103,1128]" pageId="12" pageNumber="357">
<heading id="D0AD3154FF98FFB3C387C87FFE7B3E77" bold="true" box="[151,480,1103,1128]" fontSize="10" level="2" pageId="12" pageNumber="357" reason="2">
<emphasis id="B92E5A2AFF98FFB3C387C87FFE7B3E77" bold="true" box="[151,480,1103,1128]" pageId="12" pageNumber="357">
<taxonomicName id="4C5AFDBBFF98FFB3C387C87FFEE43E77" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[151,383,1104,1128]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="357" phylum="Arthropoda" rank="species" species="pacoensis" status="sp. nov.">
<emphasis id="B92E5A2AFF98FFB3C387C87FFEE43E77" bold="true" box="[151,383,1104,1128]" italics="true" pageId="12" pageNumber="357">Yezoceryx pacoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21DE751FF98FFB3C296C860FE7B3E77" box="[390,480,1103,1128]" pageId="12" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE58638FF98FFB3C387C85BFE933E93" blockId="12.[151,480,1103,1165]" box="[151,264,1140,1165]" pageId="12" pageNumber="357">
(
<figureCitation id="13619ABDFF98FFB3C38EC85BFF653E93" box="[158,254,1140,1165]" captionStart="FIGURE 9" captionStartId="13.[151,250,1885,1907]" captionTargetBox="[160,1420,423,1847]" captionTargetId="figure@13.[160,1420,423,1848]" captionTargetPageId="13" captionText="FIGURE 9. Yezoceryx pacoensis sp. nov.: a. lateral view; b. face; c. lateral view of mesosoma; d. dorsal view of head; e. lateral view of first metasomal tergite; f. dorsal view of propodeum." httpUri="https://zenodo.org/record/847872/files/figure.png" pageId="12" pageNumber="357">Figure 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF98FFB3C387C893FC383F02" pageId="12" pageNumber="357" type="diagnosis">
<paragraph id="8BE58638FF98FFB3C387C893FC383F02" blockId="12.[151,1437,1212,2029]" pageId="12" pageNumber="357">
<emphasis id="B92E5A2AFF98FFB3C387C893FE8A3ECA" bold="true" box="[151,273,1212,1237]" pageId="12" pageNumber="357">Diagnosis.</emphasis>
Body black with head extensively yellow on inner and outer orbits; occipital carina deeply concave dorsally; antenna with median whitish flagellomeres; first tergite 2.4× as long as posterior width; first sternite with sharp anterior-facing hook; ovipositor about 3.3× length of hind tibia.
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF98FFB3C3D7C908FF5A3F7A" pageId="12" pageNumber="357" type="materials_examined">
<paragraph id="8BE58638FF98FFB3C3D7C908FF5A3F7A" blockId="12.[151,1437,1212,2029]" pageId="12" pageNumber="357">
<materialsCitation id="3B328C65FF98FFB3C3D7C908FF5A3F7A" ID-GBIF-Occurrence="1586985645" collectingDate="2002-04-23" collectionCode="IEBR" collectorName="T. V. Hoang" elevation="1100" location="Pa Co" municipality="Mai Chau" pageId="12" pageNumber="357" specimenCount="1" specimenCount-female="1" stateProvince="Hoa Binh" typeStatus="holotype">
<emphasis id="B92E5A2AFF98FFB3C3D7C908FE2A3F5F" bold="true" box="[199,433,1319,1344]" pageId="12" pageNumber="357">Material examined.</emphasis>
<typeStatus id="54E1389AFF98FFB3C2A8C908FDBE3F5F" box="[440,549,1319,1344]" pageId="12" pageNumber="357" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D5C4DB1FF98FFB3C120C907FDDE3F20" box="[560,581,1320,1343]" pageId="12" pageNumber="357" type="female"></specimenCount>
(
<collectionCode id="ED4B1EFDFF98FFB3C144C907FD023F5F" box="[596,665,1320,1344]" pageId="12" pageNumber="357">IEBR</collectionCode>
),
<collectingRegion id="499E48DAFF98FFB3C1BDC907FC873F5F" box="[685,796,1319,1344]" country="Vietnam" name="Hoa Binh" pageId="12" pageNumber="357">Hoa Binh</collectingRegion>
,
<collectingMunicipality id="6B811C42FF98FFB3C038C908FC003F5F" box="[808,923,1319,1344]" pageId="12" pageNumber="357">Mai Chau</collectingMunicipality>
,
<location id="8E85D0E3FF98FFB3C0B7C907FC763F5F" LSID="urn:lsid:plazi:treatment:03F3372EFF98FFB1C387C87FFAA2389B:8E85D0E3FF98FFB3C0B7C907FC763F5F" box="[935,1005,1320,1344]" municipality="Mai Chau" name="Pa Co" pageId="12" pageNumber="357" stateProvince="Hoa Binh">Pa Co</location>
,
<date id="FFE4A0F8FF98FFB3C0E9C908FBE93F5F" box="[1017,1138,1319,1344]" pageId="12" pageNumber="357" value="2002-04-23">
<collectingDate id="EFA05910FF98FFB3C0E9C908FBE93F5F" box="[1017,1138,1319,1344]" pageId="12" pageNumber="357" value="2002-04-23">23.iv.2002</collectingDate>
</date>
,
<elevation id="0077610BFF98FFB3C76EC907FA943F5F" box="[1150,1295,1319,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="357" unit="m" value="1100.0">
<quantity id="4CA22BDDFF98FFB3C76EC907FB493F20" box="[1150,1234,1320,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="357" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
,
<collectorName id="26AFE3EEFF98FFB3C60CC907FA063F5F" box="[1308,1437,1320,1344]" pageId="12" pageNumber="357">T.V. Hoang</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF98FFB1C3D7C940FB2F3822" lastPageId="14" lastPageNumber="359" pageId="12" pageNumber="357" type="description">
<paragraph id="8BE58638FF98FFB3C3D7C940FAAD3C7F" blockId="12.[151,1437,1212,2029]" pageId="12" pageNumber="357">
<emphasis id="B92E5A2AFF98FFB3C3D7C940FDD13F97" bold="true" box="[199,586,1391,1416]" pageId="12" pageNumber="357">
Description of female
<typeStatus id="54E1389AFF98FFB3C2CCC940FDDE3F97" box="[476,581,1391,1416]" pageId="12" pageNumber="357" type="holotype">holotype</typeStatus>
.
</emphasis>
Head. Antenna with 36 flagellomeres, first flagellomere 1.1× length of second; frons concave and polished medially, swollen and punctate laterally; face 0.65× as high as wide, with large punctures laterally, medially swollen and puncto-striate; clypeus about 0.45× as high as wide, with shallow punctures, with median and lateral tubercles; malar space equal to basal width of mandible; mandible with irregular striations, lower tooth of mandible equal to upper tooth, inner margins of eyes divergent ventrally; diameter of lateral ocellus 0.5× ocellar-ocular distance; vertex matt, with moderately dense punctures; occipital carina deeply concave dorsally, ventrally meeting hypostomal carina 1.0× basal mandible width from base of mandible.
</paragraph>
<paragraph id="8BE58638FF98FFB2C3D7CA42FC253ACB" blockId="12.[151,1437,1212,2029]" lastBlockId="13.[151,1437,151,357]" lastPageId="13" lastPageNumber="358" pageId="12" pageNumber="357">
Mesosoma. Epomia 0.9× mandible basal width; pronotum with transverse striations across median groove, upper posterior corner with large, dense punctures; mesoscutum 0.9× as long as wide at anterior level of tegulae, matt, with dense, coarse punctures; convergence area of notauli with irregular sinuous wrinkles; scuto-scutellar groove with two strong plus some weak carinae; scutellum smoother than mesoscutum with shallow punctures, pubescent, lateral carina extending nearly to apex; postscutellum polished, impunctate; mesopleuron with dense, small punctures dorsally and anteriorly, pubescent, ventral half with dense striae mixed with large punctures, epicnemial carina complete, posterior transverse carina of mesosternum present medially; metapleuron with dense, largely punctures, pubescent; propodeum with irregular wrinkles, area basalis round posteriorly, area externa and area dentipara defined, area petiolaris polished, impunctate, pleural parts of anterior and posterior transverse carinae present; fore basitarsus 2.5× second tarsomere; mid basitarsus 2.4× second tarsomere; hind leg with femur 3.0× as long as wide, 0.8× length of tibia, tarsus 0.9× tibia length, basitarsus 1.9× second tarsomere, fifth tarsomere 2.1× longer than third. Fore wing length
<quantity id="4CA22BDDFF99FFB2C17CCCB7FD263AB0" box="[620,701,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="358" unit="mm" value="11.0">11 mm</quantity>
, vein
<emphasis id="B92E5A2AFF99FFB2C014CCB7FCD73AB0" box="[772,844,152,175]" italics="true" pageId="13" pageNumber="358">Rs&amp;M</emphasis>
opposite
<emphasis id="B92E5A2AFF99FFB2C0ADCCB6FC743AAF" box="[957,1007,153,176]" italics="true" pageId="13" pageNumber="358">cu-a</emphasis>
, vein 2
<emphasis id="B92E5A2AFF99FFB2C755CCB6FBE23AAF" box="[1093,1145,153,176]" italics="true" pageId="13" pageNumber="358">rs-m</emphasis>
basad of 2
<emphasis id="B92E5A2AFF99FFB2C7E8CCB6FAAB3AAF" box="[1272,1328,153,176]" italics="true" pageId="13" pageNumber="358">m-cu</emphasis>
by about 0.3× length of 2
<emphasis id="B92E5A2AFF99FFB2C25ACC92FEE63ACB" box="[330,381,189,212]" italics="true" pageId="13" pageNumber="358">rs-m</emphasis>
, first abscissa of hind wing vein
<emphasis id="B92E5A2AFF99FFB2C1FFCC93FC8B3ACC" box="[751,784,188,211]" italics="true" pageId="13" pageNumber="358">Cu</emphasis>
1 equal to
<emphasis id="B92E5A2AFF99FFB2C095CC92FC2C3ACB" box="[901,951,189,212]" italics="true" pageId="13" pageNumber="358">cu-a</emphasis>
.
</paragraph>
<paragraph id="8BE58638FF99FFB2C3D7CCCFFD243B7A" blockId="13.[151,1437,151,357]" pageId="13" pageNumber="358">Metasoma. Tergites with moderately dense, small punctures, pubescent; first tergite 2.4× as long as posterior width, lateral longitudinal carina present anteriorly, not reaching to spiracle; second tergite 0.5× as long as first tergite, and equal to third; first sternite with sharp, anteriorly projecting hook; ovipositor about 3.3× as long as hind tibia; hypopygium 1.8× as long as second tergite.</paragraph>
<caption id="DF25D6B0FF99FFB2C387CB72FC9A3D8D" httpUri="https://zenodo.org/record/847872/files/figure.png" pageId="13" pageNumber="358" targetBox="[160,1420,423,1847]" targetPageId="13">
<paragraph id="8BE58638FF99FFB2C387CB72FC9A3D8D" blockId="13.[151,1435,1885,1938]" pageId="13" pageNumber="358">
<emphasis id="B92E5A2AFF99FFB2C387CB72FE893D6D" bold="true" box="[151,274,1885,1907]" pageId="13" pageNumber="358">FIGURE 9.</emphasis>
<taxonomicName id="4C5AFDBBFF99FFB2C208CB71FE7D3D6C" authority="Achterberg, 2017" authorityName="Achterberg" authorityYear="2017" box="[280,486,1886,1907]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="358" phylum="Arthropoda" rank="species" species="pacoensis" status="sp. nov.">
<emphasis id="B92E5A2AFF99FFB2C208CB71FE7D3D6C" box="[280,486,1886,1907]" italics="true" pageId="13" pageNumber="358">Yezoceryx pacoensis</emphasis>
</taxonomicName>
<emphasis id="B92E5A2AFF99FFB2C2FCCB72FDA73D6D" bold="true" box="[492,572,1885,1906]" pageId="13" pageNumber="358">
<taxonomicNameLabel id="A21DE751FF99FFB2C2FCCB72FDA73D6D" box="[492,572,1885,1906]" pageId="13" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: a. lateral view; b. face; c. lateral view of mesosoma; d. dorsal view of head; e. lateral view of first metasomal tergite; f. dorsal view of propodeum.
</paragraph>
</caption>
<paragraph id="8BE58638FF9AFFB1C3D7CCB8FD333B97" blockId="14.[151,1436,151,645]" pageId="14" pageNumber="359">Colour. Black. Antenna black, except flagellomeres 10-21 whitish or partly whitish, face yellow with median black longitudinal stripe; frons yellow laterally; clypeus black with lateral yellowish mark; ventral two thirds of outer orbit and malar space yellow, tegula and fore and mid legs yellow, except coxae; hind leg with yellow trochanter and trochantellus, femur brown except extreme proximally and distal fifth yellow, tibia and tarsus yellow; metasomal tergites with narrow yellowish brown posterior margins, ovipositor yellow, ovipositor sheath brown, hypopygium black with dorsal margin and posterior narrowed section yellow; fore wing with distal band and area below pterostigma slightly infuscated.</paragraph>
<paragraph id="8BE58638FF9AFFB1C3D7CDBBFE1E3BB2" blockId="14.[151,1436,151,645]" box="[199,389,404,429]" pageId="14" pageNumber="359">Male. Unknown.</paragraph>
<paragraph id="8BE58638FF9AFFB1C3D7CD98FB2F3822" blockId="14.[151,1436,151,645]" pageId="14" pageNumber="359">
Comparison. According to
<bibRefCitation id="EFCBFBC9FF9AFFB1C2EDCD97FD2C3BCF" author="Cameron" box="[509,695,440,464]" pageId="14" pageNumber="359" refString="Cameron, P. (1899) Hymenoptera Orientalia, or Contributions to a knowledge of the Hymenoptera of the Oriental Zoological Region. Part VIII. The Hymenoptera of the Khasia Hills. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 43, 1 - 220." type="journal article" year="1899">Cameron (1899)</bibRefCitation>
and
<bibRefCitation id="EFCBFBC9FF9AFFB1C1E1CD98FCE63BCF" author="Chiu" box="[753,893,439,464]" pageId="14" pageNumber="359" refString="Chiu, S. - C. (1971) The Taiwan Acaenitinae (Hymenoptera: Ichneumonidae). Bulletin of the Taiwan Agricultural Research Institute, 29, 1 - 26." type="journal article" year="1971">Chiu (1971)</bibRefCitation>
, the new species is most similar to
<taxonomicName id="4C5AFDBBFF9AFFB1C605CD96FE8B3BEB" authority="(Cameron)" authorityName="Cameron" baseAuthorityName="Cameron" baseAuthorityYear="1899" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="B92E5A2AFF9AFFB1C605CD96FA003BD0" box="[1301,1435,440,464]" italics="true" pageId="14" pageNumber="359">Y. nigricans</emphasis>
(Cameron)
</taxonomicName>
in the shape of the occipital carina and in general colour. It can be separated from
<taxonomicName id="4C5AFDBBFF9AFFB1C7D6CDF2FAD63BEB" baseAuthorityName="Cameron" baseAuthorityYear="1899" box="[1222,1357,477,500]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="B92E5A2AFF9AFFB1C7D6CDF2FAD63BEB" box="[1222,1357,477,500]" italics="true" pageId="14" pageNumber="359">Y. nigricans</emphasis>
</taxonomicName>
by the median white band on the antennal flagellum and the largely yellow face and legs, whereas
<taxonomicName id="4C5AFDBBFF9AFFB1C7B4CE2EFAB23808" baseAuthorityName="Cameron" baseAuthorityYear="1899" box="[1188,1321,512,536]" class="Insecta" family="Ichneumonidae" genus="Yezoceryx" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="359" phylum="Arthropoda" rank="species" species="nigricans">
<emphasis id="B92E5A2AFF9AFFB1C7B4CE2EFAB23808" box="[1188,1321,512,536]" italics="true" pageId="14" pageNumber="359">Y. nigricans</emphasis>
</taxonomicName>
(
<typeStatus id="54E1389AFF9AFFB1C628CDD0FA073807" box="[1336,1436,511,536]" pageId="14" pageNumber="359" type="holotype">holotype</typeStatus>
female in BMNH examined) is almost entirely black, except for white inner orbits of the face.
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF9AFFB1C3D7CE68FD3B387F" box="[199,672,583,608]" pageId="14" pageNumber="359" type="etymology">
<paragraph id="8BE58638FF9AFFB1C3D7CE68FD3B387F" blockId="14.[151,1436,151,645]" box="[199,672,583,608]" pageId="14" pageNumber="359">
<emphasis id="B92E5A2AFF9AFFB1C3D7CE68FED5387F" bold="true" box="[199,334,583,608]" pageId="14" pageNumber="359">Etymology.</emphasis>
Named after the
<typeStatus id="54E1389AFF9AFFB1C11FCE67FDA4387F" box="[527,575,584,608]" pageId="14" pageNumber="359">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C340D5B3FF9AFFB1C3D7CE43FAA2389B" box="[199,1337,620,645]" pageId="14" pageNumber="359" type="distribution">
<paragraph id="8BE58638FF9AFFB1C3D7CE43FAA2389B" blockId="14.[151,1436,151,645]" box="[199,1337,620,645]" pageId="14" pageNumber="359">
<emphasis id="B92E5A2AFF9AFFB1C3D7CE43FEFB389A" bold="true" box="[199,352,620,645]" pageId="14" pageNumber="359">Distribution.</emphasis>
Currently known only from Mai Chau,
<collectingRegion id="499E48DAFF9AFFB1C030CE42FC61389A" box="[800,1018,620,645]" country="Vietnam" name="Hoa Binh" pageId="14" pageNumber="359">Hoa Binh Province</collectingRegion>
, North
<collectingCountry id="F34DC6A8FF9AFFB1C75DCE43FB37389A" box="[1101,1196,620,645]" name="Vietnam" pageId="14" pageNumber="359">Vietnam</collectingCountry>
(
<figureCitation id="13619ABDFF9AFFB1C7AACE43FAB0389B" box="[1210,1323,620,645]" captionStart="FIGURE 19" captionStartId="25.[151,250,1940,1962]" captionTargetBox="[241,1347,195,1919]" captionTargetId="figure@25.[241,1350,195,1923]" captionTargetPageId="25" captionText="FIGURE 19. Map of sampling sites of Yezoceryx spp in Vietnam: 1. Cao Bang Province; 2. Bac Kan Province; 3. Vinh Phuc Province, 4. Son La Province; 5. Hoa Binh Province; 6. Ninh Binh Province; 7. Nghe An Province; 8. Thua Thien-Hue Province; 9. Kon Tum Province; 10. Dak Lak Province; 11. Dong Nai Province." httpUri="https://zenodo.org/record/847892/files/figure.png" pageId="14" pageNumber="359">Figure 19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>