161 lines
18 KiB
XML
161 lines
18 KiB
XML
<document id="469A2A1547D98F76326E0AD8055D467D" ID-DOI="10.1093/zoolinnean/zlad078" ID-ISSN="0024-4082" ID-Zenodo-Dep="11240120" ID-ZooBank="5D6C7EBF-E568-4100-9364-2DD357003878" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716361325736" checkinUser="plazi" docAuthor="Marceniuk, Alexandre Pires, Oliveira, Claudio & Ferraris Jr, Carl J." docDate="2024" docId="03F32B77FFC5FF830C79FC97FC4E48A9" docLanguage="en" docName="zlad078.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad078" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Ketengus Bleeker 1846" docType="treatment" docVersion="2" lastPageNumber="460" masterDocId="FFCA530FFFE7FFA10D44FFD6FFAB4A1E" masterDocTitle="A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data" masterLastPageNumber="476" masterPageNumber="426" pageNumber="460" updateTime="1716362088827" updateUser="ExternalLinkService">
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<mods:title id="8716284A914009A2A278102E6E6E2855">A new classification of the family Ariidae (Osteichthyes: Ostariophysi: Siluriformes) based on combined analyses of morphological and molecular data</mods:title>
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<mods:namePart id="E4D9C463A226780A64C51E7D077FD95D">Marceniuk, Alexandre Pires</mods:namePart>
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<mods:affiliation id="730AEA14D10E6CC50ECDB82EC64A7CF2">Programa de Pós-Graduação em Ciências Biológicas, Universidade Federal da Paraíba, conjunto Presidente Castelo Branco III, João Pessoa, PB 58050 - 000, Brazil & Programa de Pós-Graduação em Ecologia e Conservação, Universidade Estadual da Paraíba, Campina Grande, PB 58429 - 500, Brazil & Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
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<mods:nameIdentifier id="8447B9CC0BE69EB11799947B0928BE7A" type="email">a_marceniuk@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="63B98C16047CE3BAA8C2DA284D56E296">Oliveira, Claudio</mods:namePart>
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<mods:affiliation id="3F4932937EB3DC2DFF39BCA558F23D5D">Departamento de Biologia Estrutural e Funcional, Instituto de Biociências, Universidade Estadual Paulista Júlio de Mesquita Filho (UNESP), Botucatu, SP 18618 - 970, Brazil</mods:affiliation>
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<mods:namePart id="E1265567F8E64D1380C12ED2ECF396F3">Ferraris Jr, Carl J.</mods:namePart>
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<mods:affiliation id="67E1B172FBA15279EA38587C67DD00D9">545 NE Floral Pl., Portland, OR 97232, USA</mods:affiliation>
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<mods:title id="CBA77A09B64965ED122F52804A68C24D">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="9DEF7E11EC13C831492C67168BE78AA5">
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<mods:date id="F19692D2A8C55F0A3DEFC497561ACE76">2024</mods:date>
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<mods:number id="868B372AF7207A943D95C09367563E0E">2023-08-21</mods:number>
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<mods:number id="51B3AD383F2ECFCB9EF44809177B79B1">200</mods:number>
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<mods:identifier id="89B4C73F5EC09B8D18296EF920ED0292" type="DOI">10.1093/zoolinnean/zlad078</mods:identifier>
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<mods:identifier id="04FADE7A4AF8392CB6506DEB7928DD47" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F32B77FFC5FF830C79FC97FC4E48A9" LSID="urn:lsid:plazi:treatment:03F32B77FFC5FF830C79FC97FC4E48A9" httpUri="http://treatment.plazi.org/id/03F32B77FFC5FF830C79FC97FC4E48A9" lastPageNumber="460" pageId="34" pageNumber="460">
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<subSubSection id="C340C9EAFFC5FF830C79FC97FD854945" box="[317,558,833,859]" pageId="34" pageNumber="460" type="nomenclature">
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<paragraph id="8BE59A61FFC5FF830C79FC97FD854945" blockId="34.[317,558,833,859]" box="[317,558,833,859]" pageId="34" pageNumber="460">
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<heading id="D0AD2D0DFFC5FF830C79FC97FD854945" bold="true" box="[317,558,833,859]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="460" reason="2">
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<taxonomicName id="4C5AE1E2FFC5FF830C79FC97FD854945" ID-CoL="5847" ID-ENA="337776" authority="Bleeker, 1846" authorityName="Bleeker" authorityYear="1846" box="[317,558,833,859]" family="Ariidae" genus="Ketengus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="460" phylum="Chordata" rank="genus">
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<emphasis id="B92E4673FFC5FF830C79FC97FD854945" bold="true" box="[317,558,833,859]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830C79FC97FE334945" bold="true" box="[317,408,833,859]" italics="true" pageId="34" pageNumber="460">Ketengus</emphasis>
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Bleeker, 1846
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C340C9EAFFC5FF830C2FFCBBFC4E48A9" pageId="34" pageNumber="460" type="description">
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<paragraph id="8BE59A61FFC5FF830C2FFCBBFDAB4999" blockId="34.[363,512,877,904]" box="[363,512,877,904]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830C2FFCBBFDAB4999" bold="true" box="[363,512,877,904]" pageId="34" pageNumber="460">
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(
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<figureCitation id="136186E4FFC5FF830C32FCBBFE644999" box="[374,463,877,903]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[129,194,1805,1829]" captionStartId-1="3.[825,890,1803,1827]" captionStartId-2="4.[113,178,1813,1837]" captionTargetBox-1="[153,1434,148,1768]" captionTargetBox-2="[151,708,146,1775]" captionTargetId-2="figure-643@4.[149,717,144,1776]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. MP analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-1="Figure 2. ML analysis. Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green). Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." captionText-2="Figure 3. BI analysis.Subfamilies (SF) Galeichthyinae (pink), Bagreinae (bright blue), and Ariinae (dark blue). Tribe (T) Ariini (blue), Cathopsini (orange), Genidentini tribe nov. (brown), and Sciadeini tribe nov. (light green).Subtribe (ST) Ariina (light blue) and Doiichthyina (green). Incertae sedis taxa (IS) (red). Numbers over branches correspond to the bootstrap support." figureDoi-1="http://doi.org/10.5281/zenodo.11240122" figureDoi-2="http://doi.org/10.5281/zenodo.11240126" httpUri-1="https://zenodo.org/record/11240122/files/figure.png" httpUri-2="https://zenodo.org/record/11240126/files/figure.png" pageId="34" pageNumber="460">Figs 1–3</figureCitation>
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,
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<figureCitation id="136186E4FFC5FF830C9EFCBBFE5D4999" box="[474,502,877,903]" captionStart="Figure 34" captionStartId="34.[113,178,1954,1978]" captionTargetBox="[114,1458,1562,1926]" captionTargetId="figure-612@34.[114,1458,1562,1926]" captionTargetPageId="34" captionText="Figure 34. Ketengus typus, ANSP 60704, eastern Indian Ocean, type species of the genus." figureDoi="http://doi.org/10.5281/zenodo.11240189" httpUri="https://zenodo.org/record/11240189/files/figure.png" pageId="34" pageNumber="460">34</figureCitation>
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)
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</emphasis>
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830D35FC71FDB149A1" blockId="34.[113,538,935,959]" box="[113,538,935,959]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830D35FC71FF4649A0" box="[113,237,935,958]" italics="true" pageId="34" pageNumber="460">
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<typeStatus id="54E124C3FFC5FF830D35FC71FF0A49A0" box="[113,161,935,958]" pageId="34" pageNumber="460">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C5AE1E2FFC5FF830DB6FC71FDBE49A1" authority="Bleeker, 1846" authorityName="Bleeker" authorityYear="1846" box="[242,533,935,959]" family="Ariidae" genus="Ketengus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="460" phylum="Chordata" rank="species" species="typus">
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<emphasis id="B92E4673FFC5FF830DB6FC71FE2849A0" box="[242,387,935,958]" italics="true" pageId="34" pageNumber="460">
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Ketengus
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<typeStatus id="54E124C3FFC5FF830C0BFC71FE2849A0" box="[335,387,935,958]" pageId="34" pageNumber="460">typus</typeStatus>
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</emphasis>
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Bleeker, 1846
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830CC3FC03FE4F49F0" blockId="34.[391,484,981,1006]" box="[391,484,981,1006]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830CC3FC03FE4F49F0" box="[391,484,981,1006]" italics="true" pageId="34" pageNumber="460">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830D35FC2DFAEF4B3B" blockId="34.[113,763,1019,1513]" lastBlockId="34.[810,1459,144,293]" pageId="34" pageNumber="460">
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Mesethmoid medial notch large and shallow (1, 2> 0); fenestra delimited by mesethmoid and lateral ethmoid present (3, 0> 1); fenestra delimited by mesethmoid and lateral ethmoid large, filled with cartilage (
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<emphasis id="B92E4673FFC5FF830C95FB8EFD984E6E" bold="true" box="[465,563,1112,1137]" pageId="34" pageNumber="460">4,?> 0)</emphasis>
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; mesethmoid posterior horn tubular, narrow and elongate (5, 0> 1); posterior branches of mesethmoid narrow (6, 0> 1); posterior branches of mesethmoid parallel throughout their entire extension (7, 0> 1); lateral ethmoid not contact frontal (
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<emphasis id="B92E4673FFC5FF830F78FB00FD024EF0" bold="true" box="[572,681,1238,1262]" pageId="34" pageNumber="460">14, 3> 0)</emphasis>
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; vomer diamond shaped (49, 1> 0); vomer lateral processes absent (
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<emphasis id="B92E4673FFC5FF830DF6FAC2FE884F32" bold="true" box="[178,291,1300,1325]" pageId="34" pageNumber="460">51, 1> 0)</emphasis>
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; accessory tooth plates absent (58, 1> 0); three anterior branches in lachrymal-antorbital anterior part
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<emphasis id="B92E4673FFC5FF830D35FA85FF424F75" bold="true" box="[113,233,1363,1387]" pageId="34" pageNumber="460">(98, 2> 3)</emphasis>
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; teeth on dentary along entire bone (
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<emphasis id="B92E4673FFC5FF830F3CFA85FD584F75" bold="true" box="[632,755,1363,1387]" pageId="34" pageNumber="460">116, 1> 2)</emphasis>
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; dorsal crest of premaxilla present (125, 0> 1); opercle posterior portion not well developed posteriorly (129, 1> 0); ventral crest of hyomandibula absent (142, 0> 1); dorsal crest of urohyal projected anteriorly (156, 0> 1); one basibranchial series
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<emphasis id="B92E4673FFC5FF830E35FF46FC574AB6" bold="true" box="[881,1020,144,168]" pageId="34" pageNumber="460">(164, 0> 1)</emphasis>
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; first hypobranchial club shaped
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<emphasis id="B92E4673FFC5FF83083CFF46FCC64AD9" bold="true" pageId="34" pageNumber="460">(171, 0> 1)</emphasis>
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; second hypobranchial club shaped (
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<emphasis id="B92E4673FFC5FF8309BAFF79FAD04AD9" bold="true" box="[1278,1403,175,199]" pageId="34" pageNumber="460">175, 0> 1)</emphasis>
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; first epibranchial overlaying second epibranchial (179, 0> 1); third pharyngobranchial funnel shaped (192, 1> 0); seventh vertebra free from ventral superficial ossification (214, 0> 1).
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830962FE93FB1C4B41" blockId="34.[810,1207,325,387]" box="[1062,1207,325,351]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830962FE93FB1C4B41" box="[1062,1207,325,351]" italics="true" pageId="34" pageNumber="460">Included species</emphasis>
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830E6EFEBAFBFA4B9D" blockId="34.[810,1207,325,387]" box="[810,1105,363,387]" pageId="34" pageNumber="460">
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<taxonomicName id="4C5AE1E2FFC5FF830E6EFEBAFBE64B9D" authority="Bleeker, 1846" authorityName="Bleeker" authorityYear="1846" box="[810,1101,363,387]" family="Ariidae" genus="Ketengus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="460" phylum="Chordata" rank="species" species="typus">
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<emphasis id="B92E4673FFC5FF830E6EFEBAFC104B9D" box="[810,955,364,387]" italics="true" pageId="34" pageNumber="460">
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Ketengus
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<typeStatus id="54E124C3FFC5FF830EC3FEBAFC104B9D" box="[903,955,364,387]" pageId="34" pageNumber="460">typus</typeStatus>
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</emphasis>
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Bleeker, 1846
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830E6EFE73FC174BC2" blockId="34.[810,1459,421,476]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830E6EFE73FB8F4BA3" box="[810,1060,421,445]" italics="true" pageId="34" pageNumber="460">Habitat and distribution:</emphasis>
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Brackish waters, South and Southeast Asia (
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<figureCitation id="136186E4FFC5FF830E22FE12FC004BC2" box="[870,939,452,476]" captionStart="Figure 24" captionStartId="27.[129,194,1851,1875]" captionTargetBox="[259,1343,151,1822]" captionTargetId="figure-8@27.[257,1345,145,1823]" captionTargetPageId="27" captionText="Figure 24. Representative skulls, in dorsal view, and distribution of the subtribe Ariina.A, Arius maculatus, AUM 44875. B, Cephalocassis melanochir, CAS 49426. C, Cryptarius truncatus, AUM 50243. D, Hexanematichthys sagor, AUM 50242. E, Kyataphisa nenga, AUM 46280. F, Osteogeneiosus militaris, AUM 44880. G, Pseudosciades sona, AUM 44861. * marine, ** brackish water, *** freshwater." figureDoi="http://doi.org/10.5281/zenodo.11240169" httpUri="https://zenodo.org/record/11240169/files/figure.png" pageId="34" pageNumber="460">Fig. 24</figureCitation>
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).
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830900FE2DFB33480B" blockId="34.[1092,1176,507,533]" box="[1092,1176,507,533]" pageId="34" pageNumber="460">
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<emphasis id="B92E4673FFC5FF830900FE2DFB33480B" box="[1092,1176,507,533]" italics="true" pageId="34" pageNumber="460">Remarks</emphasis>
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</paragraph>
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<paragraph id="8BE59A61FFC5FF830E6EFDF4FC4E48A9" blockId="34.[810,1459,546,695]" pageId="34" pageNumber="460">
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The close relationship of
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<taxonomicName id="4C5AE1E2FFC5FF830916FDF5FB004824" authorityName="Bleeker" authorityYear="1846" box="[1106,1195,547,570]" family="Ariidae" genus="Ketengus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="460" phylum="Chordata" rank="genus">
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<emphasis id="B92E4673FFC5FF830916FDF5FB004824" box="[1106,1195,547,570]" italics="true" pageId="34" pageNumber="460">Ketengus</emphasis>
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</taxonomicName>
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||
plus
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<taxonomicName id="4C5AE1E2FFC5FF8309B0FDF4FA2C4827" authorityName="Bleeker" authorityYear="1846" box="[1268,1415,546,569]" family="Ariidae" genus="Osteogeneiosus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="460" phylum="Chordata" rank="genus">
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||
<emphasis id="B92E4673FFC5FF8309B0FDF4FA2C4827" box="[1268,1415,546,569]" italics="true" pageId="34" pageNumber="460">Osteogeneiosus</emphasis>
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||
</taxonomicName>
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established in previous morphological and molecular studies (
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||
<bibRefCitation id="EFCBE790FFC5FF830E71FDB7FBDB4867" author="Marceniuk AP & Menezes NA" box="[821,1136,609,633]" pageId="34" pageNumber="460" pagination="1 - 126" refId="ref30826" refString="Marceniuk AP, Menezes NA. Systematics of the family Ariidae (Ostariophysi, Siluriformes), with a redefinition of the genera. Zootaxa 2007; 1416: 1 - 126." type="journal article" year="2007">Marceniuk and Menezes 2007</bibRefCitation>
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, Betancur-R. 2009,
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<bibRefCitation id="EFCBE790FFC5FF830806FDB7FC374886" author="Marceniuk AP & Menezes NA & Britto MR" pageId="34" pageNumber="460" pagination="534 - 669" refId="ref30859" refString="Marceniuk AP, Menezes NA, Britto MR. Phylogenetic analysis of the family Ariidae (Ostariophysi: Siluriformes), with a hypothesis on the monophyly and relationships of the genera. Zoological Journal of the Linnean Society 2012; 165: 534 - 669. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00822. x" type="journal article" year="2012">
|
||
Marceniuk
|
||
<emphasis id="B92E4673FFC5FF830E6EFD57FCF54886" box="[810,862,640,664]" italics="true" pageId="34" pageNumber="460">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
) is further supported by the results of the total-evidence analysis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |