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<mods:title id="277E3C5072D24E30CDC0E2650E4B325A">The genus Clubiona Latreille, 1904 (Araneae: Clubionidae) in the Maghreb, with notes on the genevensis group and new records from the Mediterranean Region</mods:title>
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<paragraph id="8BF924430907FF9B1DD1FA541016FA4A" blockId="1.[151,528,1412,1438]" box="[151,528,1412,1438]" pageId="1" pageNumber="2">
<heading id="D0B1932F0907FF9B1DD1FA541016FA4A" bold="true" box="[151,528,1412,1438]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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Genus
<taxonomicName id="4C465FC00907FF9B1DA8FA541016FA4A" ID-CoL="3R86" ID-ENA="286075" authority="Wagner, 1887" authorityName="Wagner" authorityYear="1887" box="[238,528,1412,1438]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B932F8510907FF9B1DA8FA541359FA4A" bold="true" box="[238,351,1412,1438]" italics="true" pageId="1" pageNumber="2">Clubiona</emphasis>
Wagner, 1887
</taxonomicName>
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<subSubSection id="C35C77C80907FF9B1DD1FA151081FA0E" box="[151,647,1476,1498]" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BF924430907FF9B1DD1FA151081FA0E" blockId="1.[151,647,1476,1498]" box="[151,647,1476,1498]" pageId="1" pageNumber="2">
Type species:
<taxonomicName id="4C465FC00907FF9B1C60FA141085FA0E" authority="(Clerck, 1757)" baseAuthorityName="Clerck" baseAuthorityYear="1757" box="[294,643,1476,1498]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pallidula">
<emphasis id="B932F8510907FF9B1C60FA1413E0FA0E" box="[294,486,1476,1498]" italics="true" pageId="1" pageNumber="2">Clubiona pallidula</emphasis>
(Clerck, 1757)
</taxonomicName>
.
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<emphasis id="B932F8510907FF9B1DD1FA2F13A1F9CC" bold="true" box="[151,423,1535,1560]" pageId="1" pageNumber="2">
The
<taxonomicName id="4C465FC00907FF9B1D97FA2F134BF9CC" ID-CoL="92Y9Z" authorityName="L. Koch" authorityYear="1866" box="[209,333,1535,1560]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1D97FA2F134BF9CC" bold="true" box="[209,333,1535,1560]" italics="true" pageId="1" pageNumber="2">genevensis</emphasis>
</taxonomicName>
group.
</emphasis>
<bibRefCitation id="EFD759B20907FF9B1CF4FA2F1071F9CC" author="Mikhailov" box="[434,631,1535,1560]" pageId="1" pageNumber="2" refString="Mikhailov, K. G. (1992) The spider genus Clubiona Latreille, 1804 (Arachnida Aranei Clubionidae) in the USSR fauna: a critical review with taxonomical remarks. Arthropoda Selecta, 1 (3), 3 - 34." type="journal article" year="1992">Mikhailov (1992</bibRefCitation>
;
<bibRefCitation id="EFD759B20907FF9B1FC1FA2F10C7F9CC" author="Mikhailov" box="[647,705,1535,1560]" pageId="1" pageNumber="2" refString="Mikhailov, K. G. (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta, 4 (2), 33 - 48." type="journal article" year="1995">1995</bibRefCitation>
) accommodated some
<taxonomicName id="4C465FC00907FF9B1E89F9D0163EF9CC" ID-CoL="3R86" authorityName="Wagner" authorityYear="1887" box="[975,1080,1536,1560]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B932F8510907FF9B1E89F9D0163EF9CC" box="[975,1080,1536,1560]" italics="true" pageId="1" pageNumber="2">Clubiona</emphasis>
</taxonomicName>
species into the
<taxonomicName id="4C465FC00907FF9B1842F9D1174FF9CC" ID-CoL="WC6N" authorityName="C. L. Koch" authorityYear="1839" box="[1284,1353,1537,1560]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="comta">
<emphasis id="B932F8510907FF9B1842F9D1174FF9CC" box="[1284,1353,1537,1560]" italics="true" pageId="1" pageNumber="2">comta</emphasis>
</taxonomicName>
group, which also comprised the
<taxonomicName id="4C465FC00907FF9B1CF9F9F5103EF9E8" ID-CoL="92Y9Z" authorityName="L. Koch" authorityYear="1866" box="[447,568,1573,1596]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1CF9F9F5103EF9E8" box="[447,568,1573,1596]" italics="true" pageId="1" pageNumber="2">genevensis</emphasis>
</taxonomicName>
subgroup. According to him, these species differed from the others by some characters linked to the embolus in the males and to the copulatory apparatus of the females. All species but
<taxonomicName id="4C465FC00907FF9B1DD1F9BC1349F950" ID-CoL="WC6N" authorityName="C. L. Koch" authorityYear="1839" box="[151,335,1644,1668]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="comta">
<emphasis id="B932F8510907FF9B1DD1F9BC1349F950" box="[151,335,1644,1668]" italics="true" pageId="1" pageNumber="2">Clubiona comta</emphasis>
</taxonomicName>
were part of the
<taxonomicName id="4C465FC00907FF9B1F67F9BD109CF950" ID-CoL="92Y9Z" authorityName="L. Koch" authorityYear="1866" box="[545,666,1645,1668]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1F67F9BD109CF950" box="[545,666,1645,1668]" italics="true" pageId="1" pageNumber="2">genevensis</emphasis>
</taxonomicName>
subgroup. Actually, the representatives of the last group are well distinguished by conspicuous characters, and are treated here as belonging to a species group of their own.
<taxonomicName id="4C465FC00907FF9B1DD1F964134BF918" ID-CoL="WC6N" authorityName="C. L. Koch" authorityYear="1839" box="[151,333,1716,1740]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="comta">
<emphasis id="B932F8510907FF9B1DD1F964134BF918" box="[151,333,1716,1740]" italics="true" pageId="1" pageNumber="2">Clubiona comta</emphasis>
</taxonomicName>
remains therefore the only representative of the
<taxonomicName id="4C465FC00907FF9B1E39F96511C5F918" ID-CoL="WC6N" authorityName="C. L. Koch" authorityYear="1839" box="[895,963,1717,1740]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="comta">
<emphasis id="B932F8510907FF9B1E39F96511C5F918" box="[895,963,1717,1740]" italics="true" pageId="1" pageNumber="2">comta</emphasis>
</taxonomicName>
group, and the newly erected
<taxonomicName id="4C465FC00907FF9B1865F965179AF918" ID-CoL="92Y9Z" authorityName="L. Koch" authorityYear="1866" box="[1315,1436,1717,1740]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1865F965179AF918" box="[1315,1436,1717,1740]" italics="true" pageId="1" pageNumber="2">genevensis</emphasis>
</taxonomicName>
group includes therefore eight species:
<taxonomicName id="4C465FC00907FF9B1F17F90811D7F924" ID-CoL="92Y9X" authority="Blackwall, 1859" authorityName="Blackwall" authorityYear="1859" box="[593,977,1751,1776]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510907FF9B1F17F9081109F924" box="[593,783,1752,1776]" italics="true" pageId="1" pageNumber="2">Clubiona decora</emphasis>
<bibRefCitation id="EFD759B20907FF9B1E50F90711D7F924" author="Blackwall" box="[790,977,1751,1776]" pageId="1" pageNumber="2" refString="Blackwall, J. (1859) Descriptions of newly discovered spiders captured by James Yate Johnson Esq., in the island of Madeira. Annals and Magazine of Natural History, 3 (4), 255 - 267." type="journal article" year="1859">Blackwall, 1859</bibRefCitation>
</taxonomicName>
, C.
<taxonomicName id="4C465FC00907FF9B1EB9F90816F0F924" ID-CoL="92Y9Y" authority="Simon 1878" authorityName="Simon" authorityYear="1878" box="[1023,1270,1751,1776]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="diniensis">
<emphasis id="B932F8510907FF9B1EB9F9081665F924" box="[1023,1123,1752,1776]" italics="true" pageId="1" pageNumber="2">diniensis</emphasis>
<bibRefCitation id="EFD759B20907FF9B192CF90716F0F924" author="Simon" box="[1130,1270,1751,1776]" pageId="1" pageNumber="2" refString="Simon, E. (1878) Les arachnides de France. Tome 4. Librairie Encyclopedique de Roret, Paris, 334 pp." type="book" year="1878">Simon 1878</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B1847F9081342F8C1" ID-CoL="92Y9Z" authority="L. Koch, 1866" authorityName="L. Koch" authorityYear="1866" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1847F908179AF93B" box="[1281,1436,1752,1775]" italics="true" pageId="1" pageNumber="2">C. genevensis</emphasis>
<bibRefCitation id="EFD759B20907FF9B1DD1F92D1342F8C1" author="Koch" box="[151,324,1788,1813]" pageId="1" pageNumber="2" refString="Koch, L. (1866) Die Arachniden-Familie der Drassiden. Hefte 1 - 6. Lotzbeck, Nurnberg, 304 pp." type="book chapter" year="1866">L. Koch, 1866</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B1C15F92C1086F8C0" ID-CoL="92YB4" authority="Simon, 1932" authorityName="Simon" authorityYear="1932" box="[339,640,1788,1813]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="leucaspis">
<emphasis id="B932F8510907FF9B1C15F92C13D9F8C0" box="[339,479,1788,1812]" italics="true" pageId="1" pageNumber="2">C. leucaspis</emphasis>
<bibRefCitation id="EFD759B20907FF9B1CADF92C1086F8C0" author="Simon" box="[491,640,1788,1813]" pageId="1" pageNumber="2" refString="Simon, E. (1932) Les arachnides de France. Tome VI. Encyclopedie Roret, Paris, 205 pp." type="book" year="1932">Simon, 1932</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B1FD6F92C11D7F8C0" ID-CoL="92YB6" authority="Wunderlich, 1987" authorityName="Wunderlich" authorityYear="1987" box="[656,977,1788,1813]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B932F8510907FF9B1FD6F92C10F1F8C0" box="[656,759,1788,1812]" italics="true" pageId="1" pageNumber="2">C. minor</emphasis>
<bibRefCitation id="EFD759B20907FF9B1E44F92C11D7F8C0" author="Wunderlich" box="[770,977,1788,1813]" pageId="1" pageNumber="2" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Triops, Langen, 435 pp." type="book" year="1987">Wunderlich, 1987</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B1EA7F92C1774F8C0" ID-CoL="92YB8" authority="Wunderlich, 1987" authorityName="Wunderlich" authorityYear="1987" box="[993,1394,1788,1813]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pseudominor">
<emphasis id="B932F8510907FF9B1EA7F92C169EF8C0" box="[993,1176,1788,1812]" italics="true" pageId="1" pageNumber="2">C. pseudominor</emphasis>
<bibRefCitation id="EFD759B20907FF9B19E5F92C1774F8C0" author="Wunderlich" box="[1187,1394,1788,1813]" pageId="1" pageNumber="2" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Triops, Langen, 435 pp." type="book" year="1987">Wunderlich, 1987</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B18C4F92C1379F8EC" ID-CoL="92YBB" authority=", Simon, 1918" authorityName=", Simon" authorityYear="1918" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="vegeta">
<emphasis id="B932F8510907FF9B18C4F92C12E6F8EC" italics="true" pageId="1" pageNumber="2">C. vegeta</emphasis>
, Simon, 1918
</taxonomicName>
,
<taxonomicName id="4C465FC00907FF9B1CCFF8F010F7F8EC" ID-CoL="92YBD" authority="Mikhailov, 1992" authorityName="Mikhailov" authorityYear="1992" box="[393,753,1823,1848]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="wunderlichi">
<emphasis id="B932F8510907FF9B1CCFF8F01029F8EC" box="[393,559,1824,1848]" italics="true" pageId="1" pageNumber="2">C. wunderlichi</emphasis>
<bibRefCitation id="EFD759B20907FF9B1F73F8CF10F7F8EC" author="Mikhailov" box="[565,753,1823,1848]" pageId="1" pageNumber="2" refString="Mikhailov, K. G. (1992) The spider genus Clubiona Latreille, 1804 (Arachnida Aranei Clubionidae) in the USSR fauna: a critical review with taxonomical remarks. Arthropoda Selecta, 1 (3), 3 - 34." type="journal article" year="1992">Mikhailov, 1992</bibRefCitation>
</taxonomicName>
. Some of these species are not always easy to distinguish and often have broadly overlapping distributions (
<bibRefCitation id="EFD759B20907FF9B1FFCF8941183F888" author="Helsdingen" box="[698,901,1860,1885]" pageId="1" pageNumber="2" refString="Helsdingen, P. J. van (1979) Remarks concerning Clubionidae. Bulletin of the British Arachnological Society, 4, 298 - 302." type="journal article" year="1979">Helsdingen 1979</bibRefCitation>
). Two species described from
<collectingCountry id="F35164D30907FF9B19BCF8941746F889" box="[1274,1344,1860,1885]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
,
<emphasis id="B932F8510907FF9B1814F8951773F888" box="[1362,1397,1861,1884]" italics="true" pageId="1" pageNumber="2">i.e.</emphasis>
<taxonomicName id="4C465FC00907FF9B18C4F89413B3F854" ID-CoL="5ZCTT" authority="Hu &amp; Li, 1987" authorityName="Hu &amp; Li" authorityYear="1987" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="parallela">
<emphasis id="B932F8510907FF9B18C4F89412F8F854" italics="true" pageId="1" pageNumber="2">C. parallela</emphasis>
Hu &amp; Li, 1987
</taxonomicName>
and
<taxonomicName id="4C465FC00907FF9B1CB6F8B8116DF854" ID-CoL="5ZD58" authority="Hu &amp; Li, 1987" authorityName="Hu &amp; Li" authorityYear="1987" box="[496,875,1895,1920]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="zhangmuensis">
<emphasis id="B932F8510907FF9B1CB6F8B810B7F854" box="[496,689,1896,1920]" italics="true" pageId="1" pageNumber="2">C. zhangmuensis</emphasis>
Hu &amp; Li, 1987
</taxonomicName>
, and originally included in the
<taxonomicName id="4C465FC00907FF9B1991F8B81756F8AB" ID-CoL="92Y9Z" authorityName="L. Koch" authorityYear="1866" box="[1239,1360,1896,1919]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510907FF9B1991F8B81756F8AB" box="[1239,1360,1896,1919]" italics="true" pageId="1" pageNumber="2">genevensis</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EFD759B20907FF9B1DD8F85C135EF871" author="Mikhailov" box="[158,344,1932,1957]" pageId="1" pageNumber="2" refString="Mikhailov, K. G. (1995) Erection of infrageneric groupings within the spider genus Clubiona Latreille, 1804 (Aranei Clubionidae): a typological approach. Arthropoda Selecta, 4 (2), 33 - 48." type="journal article" year="1995">Mikhailov 1995</bibRefCitation>
), do not fit the diagnosis below. The strongly enlarged and protruding tegulum of the male palp and characteristics of the female epigyne and vulva of those two species fit perfectly with the diagnosis of the “
<taxonomicName id="4C465FC00907FF9B1DE5F804130BF838" ID-CoL="WC6Z" baseAuthorityName="Walckenaer" baseAuthorityYear="1802" box="[163,269,2004,2028]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="corticalis">
<emphasis id="B932F8510907FF9B1DE5F804130BF838" box="[163,269,2004,2028]" italics="true" pageId="1" pageNumber="2">corticalis</emphasis>
</taxonomicName>
group” (subgenus
<emphasis id="B932F8510907FF9B1CA7F804107DF838" box="[481,635,2004,2028]" italics="true" pageId="1" pageNumber="2">Paraclubiona</emphasis>
), to which they must be transferred.
</paragraph>
</subSubSection>
<subSubSection id="C35C77C80904FF981D81FF4716BDFEC8" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BF924430904FF981D81FF4716BDFEC8" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="3">
<emphasis id="B932F8510904FF981D81FF471347FF64" bold="true" box="[199,321,151,176]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Representatives of the
<taxonomicName id="4C465FC00904FF981F0AFF4810C3FF7B" authorityName="L. Koch" authorityYear="1866" box="[588,709,152,175]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981F0AFF4810C3FF7B" box="[588,709,152,175]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
group are characterized in the males by the palp with an almost rounded or ovoid bulbus (
<figureCitation id="137D38C60904FF981CFCFF6C1002FF00" box="[442,516,188,213]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 4</figureCitation>
,
<figureCitation id="137D38C60904FF981F56FF6C1028FF00" box="[528,558,188,212]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">12</figureCitation>
,
<figureCitation id="137D38C60904FF981F7DFF6C105FFF00" box="[571,601,188,212]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">17</figureCitation>
,
<figureCitation id="137D38C60904FF981F23FF6C1082FF01" box="[613,644,188,213]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">25</figureCitation>
,
<figureCitation id="137D38C60904FF981FD6FF6C10A8FF00" box="[656,686,188,212]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">38</figureCitation>
,
<figureCitation id="137D38C60904FF981FFCFF6C10DEFF01" box="[698,728,188,213]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">46</figureCitation>
) and bearing a group of modified dark setae on the retrolateral face of the cymbium (
<figureCitation id="137D38C60904FF981CCCFF0F13E9FF2C" box="[394,495,223,248]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 35</figureCitation>
,
<figureCitation id="137D38C60904FF981CBCFF301011FF2C" box="[506,535,224,248]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">11</figureCitation>
,
<figureCitation id="137D38C60904FF981F64FF0F1046FF2C" box="[546,576,223,248]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">16</figureCitation>
,
<figureCitation id="137D38C60904FF981F0AFF30106DFF2C" box="[588,619,224,248]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">24</figureCitation>
,
<figureCitation id="137D38C60904FF981F33FF301095FF2C" box="[629,659,224,248]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">37</figureCitation>
,
<figureCitation id="137D38C60904FF981FD9FF0F10BBFF2C" box="[671,701,223,248]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">45</figureCitation>
,
<figureCitation id="137D38C60904FF981F8EFF0F1115FF2C" box="[712,787,223,248]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">6879</figureCitation>
). The females have an epigyne characterized by wide atria (or copulatory openings) and a deep notch in its ventral margin (
<figureCitation id="137D38C60904FF981E21FED411B6FEC9" box="[871,944,260,285]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 6</figureCitation>
,
<figureCitation id="137D38C60904FF981EFAFED411DCFEC8" box="[956,986,260,284]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">13</figureCitation>
,
<figureCitation id="137D38C60904FF981EA0FED41602FEC8" box="[998,1028,260,284]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">18</figureCitation>
,
<figureCitation id="137D38C60904FF981956FED4165EFEC9" box="[1040,1112,260,285]" captionStart="FIGURES 2633" captionStartId="9.[151,264,1387,1409]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@9.[151,1436,193,1365]" captionTargetPageId="9" captionText="FIGURES 2633. Clubiona genevensis L. Koch, 1866, female genitalia: 26 epigyne (from France), ventral view; 27, 28 same (from Greece); 29 vulva (from France), ventral view, reflected light; 30 same, dorsal view; 31 same, ventral view; 32 vulva (from France), dorsal view; 33 same, ventro-dorsal view, transmitted light. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064426/files/figure.png" pageId="2" pageNumber="3">2628</figureCitation>
,
<figureCitation id="137D38C60904FF981922FED41684FEC8" box="[1124,1154,260,284]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">39</figureCitation>
,
<figureCitation id="137D38C60904FF9819C8FED416A8FEC8" box="[1166,1198,260,284]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C77C80904FF981D88FEF7177DFA81" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BF924430904FF981D88FEF713CBFCEC" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="3">
<emphasis id="B932F8510904FF981D88FEF71344FE94" bold="true" box="[206,322,295,320]" pageId="2" pageNumber="3">Remarks.</emphasis>
The examined species of the
<taxonomicName id="4C465FC00904FF981FD6FEF8110FFEEB" authorityName="L. Koch" authorityYear="1866" box="[656,777,296,319]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981FD6FEF8110FFEEB" box="[656,777,296,319]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
group,
<emphasis id="B932F8510904FF981E26FEF8165BFE94" box="[864,1117,296,320]" italics="true" pageId="2" pageNumber="3">
i.e.
<taxonomicName id="4C465FC00904FF981ECFFEF8165BFE94" authorityName="Simon" authorityYear="1878" box="[905,1117,296,320]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="diniensis">Clubiona diniensis</taxonomicName>
</emphasis>
,
<taxonomicName id="4C465FC00904FF98192CFEF81702FEEB" authorityName="L. Koch" authorityYear="1866" box="[1130,1284,296,319]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF98192CFEF81702FEEB" box="[1130,1284,296,319]" italics="true" pageId="2" pageNumber="3">C. genevensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C465FC00904FF981855FEF8179AFE94" authorityName="Simon" authorityYear="1932" box="[1299,1436,296,320]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="leucaspis">
<emphasis id="B932F8510904FF981855FEF8179AFE94" box="[1299,1436,296,320]" italics="true" pageId="2" pageNumber="3">C. leucaspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C465FC00904FF981D81FE9C1334FEB0" authorityName=", Simon" authorityYear="1918" box="[199,306,332,356]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vegeta">
<emphasis id="B932F8510904FF981D81FE9C1334FEB0" box="[199,306,332,356]" italics="true" pageId="2" pageNumber="3">C. vegeta</emphasis>
</taxonomicName>
, are provided with modified setae situated on the retrolateral face of the cymbium (
<figureCitation id="137D38C60904FF9819ACFE9C1747FEB0" box="[1258,1345,332,357]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">Figs 11</figureCitation>
,
<figureCitation id="137D38C60904FF981808FE9C176AFEB1" box="[1358,1388,332,357]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">16</figureCitation>
,
<figureCitation id="137D38C60904FF98183FFE9C1791FEB0" box="[1401,1431,332,356]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">24</figureCitation>
,
<figureCitation id="137D38C60904FF981DD1FEA012B0FE5C" box="[151,182,368,392]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">37</figureCitation>
,
<figureCitation id="137D38C60904FF981D87FEBF12D9FE5C" box="[193,223,367,392]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">45</figureCitation>
,
<figureCitation id="137D38C60904FF981DADFEBF1001FE5C" box="[235,519,367,392]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">52, 53, 56, 57, 60, 61, 64</figureCitation>
,
<figureCitation id="137D38C60904FF981F55FEA0105BFE5C" box="[531,605,368,392]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">71, 77</figureCitation>
). They also occur in
<taxonomicName id="4C465FC00904FF981E0EFEA011B1FE5C" authorityName="Blackwall" authorityYear="1859" box="[840,951,368,392]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510904FF981E0EFEA011B1FE5C" box="[840,951,368,392]" italics="true" pageId="2" pageNumber="3">C. decora</emphasis>
</taxonomicName>
(see
<figureCitation id="137D38C60904FF981EB4FEBF1652FE5C" box="[1010,1108,367,392]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 35</figureCitation>
) but were not examined here with SEM. The group also includes
<taxonomicName id="4C465FC00904FF981F74FE441091FE78" authorityName="Wunderlich" authorityYear="1987" box="[562,663,404,428]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B932F8510904FF981F74FE441091FE78" box="[562,663,404,428]" italics="true" pageId="2" pageNumber="3">C. minor</emphasis>
</taxonomicName>
from Tenerife and La Gomera (the
<collectingRegion id="4982EAA10904FF981969FE441693FE79" box="[1071,1173,404,429]" country="Spain" name="Canarias" pageId="2" pageNumber="3">Canaries</collectingRegion>
),
<taxonomicName id="4C465FC00904FF9819EFFE44175BFE78" authorityName="Wunderlich" authorityYear="1987" box="[1193,1373,404,428]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pseudominor">
<emphasis id="B932F8510904FF9819EFFE44175BFE78" box="[1193,1373,404,428]" italics="true" pageId="2" pageNumber="3">C. pseudominor</emphasis>
</taxonomicName>
from La Palma (the
<collectingRegion id="4982EAA10904FF981C7CFE671398FE04" box="[314,414,439,464]" country="Spain" name="Canarias" pageId="2" pageNumber="3">Canaries</collectingRegion>
) (
<bibRefCitation id="EFD759B20904FF981CF2FE671084FE04" author="Wunderlich" box="[436,642,439,464]" pageId="2" pageNumber="3" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Triops, Langen, 435 pp." type="book" year="1987">Wunderlich, 1987</bibRefCitation>
, 1991;
<bibRefCitation id="EFD759B20904FF981F95FE671601FE04" author="World" box="[723,1031,439,464]" pageId="2" pageNumber="3" refString="World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 1 March 2017)" type="journal article" year="2017">World Spider Catalog 2017</bibRefCitation>
) and
<taxonomicName id="4C465FC00904FF981902FE6816EDFE04" authorityName="Mikhailov" authorityYear="1992" box="[1092,1259,440,464]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="wunderlichi">
<emphasis id="B932F8510904FF981902FE6816EDFE04" box="[1092,1259,440,464]" italics="true" pageId="2" pageNumber="3">C. wunderlichi</emphasis>
</taxonomicName>
from
<collectingCountry id="F35164D30904FF981869FE67179AFE04" box="[1327,1436,439,464]" name="Mongolia" pageId="2" pageNumber="3">Mongolia</collectingCountry>
(
<bibRefCitation id="EFD759B20904FF981DD8FE0C1351FE20" author="Mikhailov" box="[158,343,476,501]" pageId="2" pageNumber="3" refString="Mikhailov, K. G. (1992) The spider genus Clubiona Latreille, 1804 (Arachnida Aranei Clubionidae) in the USSR fauna: a critical review with taxonomical remarks. Arthropoda Selecta, 1 (3), 3 - 34." type="journal article" year="1992">Mikhailov 1992</bibRefCitation>
;
<bibRefCitation id="EFD759B20904FF981C25FE0C1091FE20" author="World" box="[355,663,476,501]" pageId="2" pageNumber="3" refString="World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 1 March 2017)" type="journal article" year="2017">World Spider Catalog 2017</bibRefCitation>
) but these were not examined. The special setae are inserted near the base of the cymbium and extend up to half its length (
<figureCitation id="137D38C60904FF981E47FE2F116FFDCC" box="[769,873,511,536]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 3, 4</figureCitation>
,
<figureCitation id="137D38C60904FF981E30FDD01195FDCC" box="[886,915,512,536]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">11</figureCitation>
,
<figureCitation id="137D38C60904FF981EE7FE2F11B8FDCC" box="[929,958,511,536]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">52</figureCitation>
) or less (
<figureCitation id="137D38C60904FF98196FFE2F1687FDCC" box="[1065,1153,511,536]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">Figs 24</figureCitation>
,
<figureCitation id="137D38C60904FF9819C8FDD016ABFDCC" box="[1166,1197,512,536]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">37</figureCitation>
,
<figureCitation id="137D38C60904FF9819FFFE2F16DEFDCC" box="[1209,1240,511,536]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">45</figureCitation>
,
<figureCitation id="137D38C60904FF9819A3FE2F175AFDCC" box="[1253,1372,511,536]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">56, 60, 64</figureCitation>
). The basal part is slightly barbed and is provided with a sub-basal enlargement that can be spherical (
<figureCitation id="137D38C60904FF9819A8FDF41775FDE9" box="[1262,1395,548,573]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">Figs 70, 76</figureCitation>
) or plate-shaped (
<figureCitation id="137D38C60904FF981C75FD9713B3FDB4" box="[307,437,583,608]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">Figs 73, 79</figureCitation>
). The extremity is needle like and barbed (
<figureCitation id="137D38C60904FF981ED4FD971614FDB4" box="[914,1042,583,608]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Figs 57, 61</figureCitation>
,
<figureCitation id="137D38C60904FF981958FD981663FDB4" box="[1054,1125,584,608]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">71, 77</figureCitation>
) or flattened and lanceolate (
<figureCitation id="137D38C60904FF981DD9FDBC12F3FD51" box="[159,245,620,645]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">Figs 16</figureCitation>
,
<figureCitation id="137D38C60904FF981C46FDBC1318FD51" box="[256,286,620,645]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">53</figureCitation>
,
<figureCitation id="137D38C60904FF981C6FFDBC134FFD51" box="[297,329,620,645]" captionStart="FIGURES6879" captionStartId="15.[151,264,1495,1517]" captionTargetBox="[151,1436,193,1474]" captionTargetId="figure@15.[151,1436,193,1474]" captionTargetPageId="15" captionText="FIGURES6879. Clubiona spp., male palp, details on the modifiedcymbialsetae, retrolateral view:6870 Clubiona diniensis Simon,1878(from Algeria); 7173 C.genevensis L.Koch, 1866(from Greece); 7476 C. leucaspis Simon, 1932(from Spain); 7779 C. vegeta Simon, 1918 (from Tunisia). 68 detail of setal apices; 69, 72, 74, 75, 78 detail of setal bases; 70, 73, 76, 79 detail of the swellings; 71, 77 global view. Scale bars: 68, 71, 74, 77 = 50 µm; 69, 70, 72, 73, 75, 76, 78, 79 = 10 µm." httpUri="https://zenodo.org/record/1064438/files/figure.png" pageId="2" pageNumber="3">68</figureCitation>
). The retrolateral tibial apophysis (RTA) is sometimes provided with a thin, translucent, needle-like extension at its apex. It was observed in one specimen of
<taxonomicName id="4C465FC00904FF981E61FD401191FD7C" authorityName="Blackwall" authorityYear="1859" box="[807,919,656,680]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510904FF981E61FD401191FD7C" box="[807,919,656,680]" italics="true" pageId="2" pageNumber="3">C. decora</emphasis>
</taxonomicName>
(
<figureCitation id="137D38C60904FF981EEEFD5F11EAFD7C" box="[936,1004,655,680]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
), one specimen of
<taxonomicName id="4C465FC00904FF981983FD401736FD7C" authorityName=", Simon" authorityYear="1918" box="[1221,1328,656,680]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vegeta">
<emphasis id="B932F8510904FF981983FD401736FD7C" box="[1221,1328,656,680]" italics="true" pageId="2" pageNumber="3">C. vegeta</emphasis>
</taxonomicName>
(
<figureCitation id="137D38C60904FF981807FD5F1795FD7C" box="[1345,1427,655,680]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Fig. 65</figureCitation>
) and on one specimen of
<taxonomicName id="4C465FC00904FF981CE2FD64103EFD18" authorityName="L. Koch" authorityYear="1866" box="[420,568,692,716]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981CE2FD64103EFD18" box="[420,568,692,716]" italics="true" pageId="2" pageNumber="3">C genevensis</emphasis>
</taxonomicName>
(P. Oger, pers. comm.; see also on http://arachno.piwigo.com/). This structure appears to be fragile (
<emphasis id="B932F8510904FF981CCAFD0913B2FD24" box="[396,436,729,752]" italics="true" pageId="2" pageNumber="3">e.g.</emphasis>
it could break during copulation) and it was lacking on another
<taxonomicName id="4C465FC00904FF9819C1FD0816F7FD24" authorityName=", Simon" authorityYear="1918" box="[1159,1265,728,752]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vegeta">
<emphasis id="B932F8510904FF9819C1FD0816F7FD24" box="[1159,1265,728,752]" italics="true" pageId="2" pageNumber="3">C. vegeta</emphasis>
</taxonomicName>
male observed with SEM (
<figureCitation id="137D38C60904FF981C58FD2C1373FCC1" box="[286,373,764,789]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Fig. 66</figureCitation>
). This character may be informative but was not used in the key since its presence in other species may be overlooked.
</paragraph>
<paragraph id="8BF924430904FF981D81FC94177DFA81" blockId="2.[151,1437,151,1365]" pageId="2" pageNumber="3">
Inside the
<taxonomicName id="4C465FC00904FF981C7AFC9513B3FC88" authorityName="L. Koch" authorityYear="1866" box="[316,437,837,860]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981C7AFC9513B3FC88" box="[316,437,837,860]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
group, two well-defined subgroups (here called the
<taxonomicName id="4C465FC00904FF981942FC95167BFC88" authorityName="L. Koch" authorityYear="1866" box="[1028,1149,837,860]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981942FC95167BFC88" box="[1028,1149,837,860]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
subgroup and the
<taxonomicName id="4C465FC00904FF981808FC94179AFC88" authorityName="Blackwall" authorityYear="1859" box="[1358,1436,836,860]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510904FF981808FC94179AFC88" box="[1358,1436,836,860]" italics="true" pageId="2" pageNumber="3">decora</emphasis>
</taxonomicName>
subgroup) can be recognized. The
<taxonomicName id="4C465FC00904FF981F5AFCB81093FCAB" authorityName="L. Koch" authorityYear="1866" box="[540,661,872,895]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981F5AFCB81093FCAB" box="[540,661,872,895]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
subgroup includes
<taxonomicName id="4C465FC00904FF981E34FCB8165DFCAB" authorityName="L. Koch" authorityYear="1866" box="[882,1115,872,896]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981E34FCB8165DFCAB" box="[882,1115,872,896]" italics="true" pageId="2" pageNumber="3">Clubiona genevensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C465FC00904FF98192CFCB816F5FC54" authorityName="Simon" authorityYear="1932" box="[1130,1267,872,896]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="leucaspis">
<emphasis id="B932F8510904FF98192CFCB816F5FC54" box="[1130,1267,872,896]" italics="true" pageId="2" pageNumber="3">C. leucaspis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C465FC00904FF98186DFCB81793FC54" authorityName=", Simon" authorityYear="1918" box="[1323,1429,872,896]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vegeta">
<emphasis id="B932F8510904FF98186DFCB81793FC54" box="[1323,1429,872,896]" italics="true" pageId="2" pageNumber="3">C. vegeta</emphasis>
</taxonomicName>
. These species are characterized in the males by the palp with the embolus originating basally and directed laterally (
<figureCitation id="137D38C60904FF981DD9FC7F12F0FC1C" box="[159,246,943,968]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">Figs 25</figureCitation>
,
<figureCitation id="137D38C60904FF981C47FC601319FC1C" box="[257,287,944,968]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">38</figureCitation>
,
<figureCitation id="137D38C60904FF981C6DFC7F134CFC1C" box="[299,330,943,968]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">46</figureCitation>
), by the basolateral extension of the bulbus and cymbium much longer than wide (
<figureCitation id="137D38C60904FF9819A2FC7F1789FC1C" box="[1252,1423,943,968]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Figs 59, 63, 67</figureCitation>
), and by the modified short cymbial setae (length less than half length of cymbium) (
<figureCitation id="137D38C60904FF981970FC04168AFC38" box="[1078,1164,980,1005]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">Figs 24</figureCitation>
,
<figureCitation id="137D38C60904FF9819DEFC0416B0FC38" box="[1176,1206,980,1004]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">37</figureCitation>
,
<figureCitation id="137D38C60904FF981987FC0416D9FC39" box="[1217,1247,980,1005]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">45</figureCitation>
,
<figureCitation id="137D38C60904FF9819ADFC041758FC39" box="[1259,1374,980,1005]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">56, 60, 64</figureCitation>
). The females have an epigyne with large, tightened and strongly convoluted copulatory ducts connected inferiorly to the atrio-spermathecal part (
<figureCitation id="137D38C60904FF981CEFFBCC102DFBE0" box="[425,555,1052,1077]" captionStart="FIGURES 2633" captionStartId="9.[151,264,1387,1409]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@9.[151,1436,193,1365]" captionTargetPageId="9" captionText="FIGURES 2633. Clubiona genevensis L. Koch, 1866, female genitalia: 26 epigyne (from France), ventral view; 27, 28 same (from Greece); 29 vulva (from France), ventral view, reflected light; 30 same, dorsal view; 31 same, ventral view; 32 vulva (from France), dorsal view; 33 same, ventro-dorsal view, transmitted light. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064426/files/figure.png" pageId="2" pageNumber="3">Figs 2932</figureCitation>
,
<figureCitation id="137D38C60904FF981F71FBCC1079FBE0" box="[567,639,1052,1076]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">40, 41</figureCitation>
,
<figureCitation id="137D38C60904FF981FCDFBCC10D0FBE1" box="[651,726,1052,1077]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">4851</figureCitation>
). The
<taxonomicName id="4C465FC00904FF981E5AFBCC116CFBE0" authorityName="Blackwall" authorityYear="1859" box="[796,874,1052,1076]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510904FF981E5AFBCC116CFBE0" box="[796,874,1052,1076]" italics="true" pageId="2" pageNumber="3">decora</emphasis>
</taxonomicName>
subgroup includes
<taxonomicName id="4C465FC00904FF981903FBCC1704FBE0" authorityName="Blackwall" authorityYear="1859" box="[1093,1282,1052,1076]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="decora">
<emphasis id="B932F8510904FF981903FBCC1704FBE0" box="[1093,1282,1052,1076]" italics="true" pageId="2" pageNumber="3">Clubiona decora</emphasis>
</taxonomicName>
,
<taxonomicName id="4C465FC00904FF981856FBCC1793FBE0" authorityName="Simon" authorityYear="1878" box="[1296,1429,1052,1076]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="diniensis">
<emphasis id="B932F8510904FF981856FBCC1793FBE0" box="[1296,1429,1052,1076]" italics="true" pageId="2" pageNumber="3">C. diniensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C465FC00904FF981DD1FB9012FBFB83" authorityName="Wunderlich" authorityYear="1987" box="[151,253,1088,1111]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B932F8510904FF981DD1FB9012FBFB83" box="[151,253,1088,1111]" italics="true" pageId="2" pageNumber="3">C. minor</emphasis>
</taxonomicName>
,
<taxonomicName id="4C465FC00904FF981C48FB9013C3FB8C" authorityName="Wunderlich" authorityYear="1987" box="[270,453,1088,1112]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pseudominor">
<emphasis id="B932F8510904FF981C48FB9013C3FB8C" box="[270,453,1088,1112]" italics="true" pageId="2" pageNumber="3">C. pseudominor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C465FC00904FF981F44FB9010AAFB8C" authorityName="Mikhailov" authorityYear="1992" box="[514,684,1088,1112]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="wunderlichi">
<emphasis id="B932F8510904FF981F44FB9010AAFB8C" box="[514,684,1088,1112]" italics="true" pageId="2" pageNumber="3">C. wunderlichi</emphasis>
</taxonomicName>
. Males of this subgroup are well characterized by the palp with embolus originating in distal or median part of bulbus and directed anteriorly (
<figureCitation id="137D38C60904FF98195EFBB41664FBA8" box="[1048,1122,1124,1149]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 4</figureCitation>
,
<figureCitation id="137D38C60904FF981929FBB4168BFBA8" box="[1135,1165,1124,1148]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">12</figureCitation>
,
<figureCitation id="137D38C60904FF9819DDFBB416BEFBA8" box="[1179,1208,1124,1148]" captionStart="FIGURES 1619" captionStartId="6.[151,264,1640,1662]" captionTargetBox="[169,1411,204,1570]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURES 1619. Clubiona diniensis Simon, 1878: 16 male (from Algeria) palp, retrolateral view; 17 same, ventral view; 18 female (from Algeria) epigyne, ventral view; 19 female (from France) vulva, dorsal view. Abbreviatures: A = atrium, CD = longitudinalcopulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 16, 17 = 0.2 mm; 18,19 = 0.1 mm." httpUri="https://zenodo.org/record/1064422/files/figure.png" pageId="2" pageNumber="3">17</figureCitation>
) (origin of embolus basal and directed laterally in the other species of the
<taxonomicName id="4C465FC00904FF981E6AFB5811A3FB4B" authorityName="L. Koch" authorityYear="1866" box="[812,933,1160,1183]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981E6AFB5811A3FB4B" box="[812,933,1160,1183]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
group:
<figureCitation id="137D38C60904FF98194EFB571665FB74" box="[1032,1123,1159,1184]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">Figs 25</figureCitation>
,
<figureCitation id="137D38C60904FF981932FB581694FB74" box="[1140,1170,1160,1184]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">38</figureCitation>
,
<figureCitation id="137D38C60904FF9819E2FB5716C5FB74" box="[1188,1219,1159,1184]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">46</figureCitation>
), short basolateral extension of bulbus and cymbium (
<figureCitation id="137D38C60904FF981F70FB7C10B8FB11" box="[566,702,1196,1221]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Figs 54, 55</figureCitation>
) (much longer in the other species:
<figureCitation id="137D38C60904FF981928FB7C1725FB11" box="[1134,1315,1196,1221]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">Figs 59, 63, 67</figureCitation>
) and long modified cymbial setae (
<figureCitation id="137D38C60904FF981CFAFB1F102AFB3C" box="[444,556,1231,1256]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 3, 4</figureCitation>
,
<figureCitation id="137D38C60904FF981F78FB00105DFB3C" box="[574,603,1232,1256]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">11</figureCitation>
,
<figureCitation id="137D38C60904FF981F2AFB1F108FFB3C" box="[620,649,1231,1256]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">52</figureCitation>
) (shorter in the others:
<figureCitation id="137D38C60904FF981EEAFB1F1601FB3C" box="[940,1031,1231,1256]" captionStart="FIGURES 2025" captionStartId="8.[151,264,1943,1965]" captionTargetBox="[365,1233,975,1928]" captionTargetId="figure@8.[360,1238,964,1935]" captionTargetPageId="8" captionText="FIGURES 2025. Clubiona genevensis L. Koch, 1866: 20 male (from France) habitus, dorsal view; 21 female (from France) habitus, dorsal view; 22, 23 males (from Greece) habitus, dorsal view; 24 male (from Greece) palp, retrolateral view; 25 same, ventral view. Abbreviature:CS = cymbial modified setae.Scale bars: 2023 = 1 mm; 24, 25 = 0.2 mm." httpUri="https://zenodo.org/record/1064424/files/figure.png" pageId="2" pageNumber="3">Figs 24</figureCitation>
,
<figureCitation id="137D38C60904FF98195EFB001630FB3C" box="[1048,1078,1232,1256]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">37</figureCitation>
,
<figureCitation id="137D38C60904FF98190EFB1F1660FB3C" box="[1096,1126,1231,1256]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">45</figureCitation>
,
<figureCitation id="137D38C60904FF98193EFB1F16F1FB3C" box="[1144,1271,1231,1256]" captionStart="FIGURES 5267" captionStartId="14.[151,264,1845,1867]" captionTargetBox="[182,1406,193,1827]" captionTargetId="figure@14.[182,1406,193,1827]" captionTargetPageId="14" captionText="FIGURES 5267. Clubiona spp., male palp: 5255 Clubiona diniensis Simon, 1878 (from Algeria); 5659 C. genevensis L. Koch, 1866 (from Greece); 6063 C. leucaspis Simon, 1932 (from Spain); 6467 C. vegeta Simon, 1918 (from Tunisia). 52, 56, 60, 64 leftpalp, retrolateral view; 53, 57, 58, 61, 62, 65, 66 detail of RTA and base of cymbium, retrolateral view (65, white arrow, RTA needle-like extension present; 66, black arrow, RTA needle-like extension absent, probably broken); 54, 59, 63, 67 detail of bulbus and basolateral extension of cymbium (stars), retrolateral view; 55 same, ventral view. Scale bars: 52, 56, 60, 64 = 0.1 mm; 53, 57, 59, 61, 63, 67 = 50 µm; 54, 55, 58, 62, 65, 66 = 20 µm." httpUri="https://zenodo.org/record/1064434/files/figure.png" pageId="2" pageNumber="3">56, 60, 64</figureCitation>
). Females are characterized by the epigyne with copulatory ducts loose, slightly coiled and connected laterally to the atriospermathecal part (
<figureCitation id="137D38C60904FF981C33FAC713D8FAE4" box="[373,478,1303,1328]" captionStart="FIGURES 18" captionStartId="4.[151,264,1422,1444]" captionTargetBox="[164,1436,303,1384]" captionTargetId="figure@4.[151,1436,303,1397]" captionTargetPageId="4" captionText="FIGURES 18. Clubiona decora Blackwall, 1859: male and female (from Madeira); 1 male habitus, dorsal view; 2 female habitus, dorsal view; 3 male palp, retrolateral view; 4 same, ventral view. 5 detail of the retrolateral tibial apophysis (RTA) of the male palp, dorsal view (arrow on needle-like extension of RTA); 6 epigyne, ventral view; 7 vulva, ventral view; 8 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars:1, 2 = 1 mm; 3, 4, 68) = 0.2 mm; 5 = 0.1 mm." httpUri="https://zenodo.org/record/1064418/files/figure.png" pageId="2" pageNumber="3">Figs 68</figureCitation>
,
<figureCitation id="137D38C60904FF981CA8FAC71033FAE4" box="[494,565,1303,1328]" captionStart="FIGURES 915" captionStartId="5.[151,264,1836,1858]" captionTargetBox="[173,1436,501,1777]" captionTargetId="figure@5.[151,1436,501,1793]" captionTargetPageId="5" captionText="FIGURES 915. Clubiona diniensis Simon, 1878: 9 male (from Spain) habitus, dorsal view; 10 female (from Spain) habitus, dorsal view; 11 male (from Algeria) palp, retrolateral view; 12 same, ventral view; 13 Female (from Algeria) epigyne, ventral view (arrow on deep ventral notch of the epigynal margin); 14 female (from France) vulva, ventral view; 15 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae. Scale bars: 9, 10 = 1 mm; 1113 = 0.2 mm; 14, 15 = 0.1 mm." httpUri="https://zenodo.org/record/1064420/files/figure.png" pageId="2" pageNumber="3">1315</figureCitation>
) (in the other species of the
<taxonomicName id="4C465FC00904FF981ED5FAC8160AFAFB" authorityName="L. Koch" authorityYear="1866" box="[915,1036,1304,1327]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="genevensis">
<emphasis id="B932F8510904FF981ED5FAC8160AFAFB" box="[915,1036,1304,1327]" italics="true" pageId="2" pageNumber="3">genevensis</emphasis>
</taxonomicName>
group, the tightened and strongly convoluted copulatory ducts are connected inferiorly to the atrio-spermathecal part:
<figureCitation id="137D38C60904FF981903FAEC16C0FA80" box="[1093,1222,1340,1365]" captionStart="FIGURES 2633" captionStartId="9.[151,264,1387,1409]" captionTargetBox="[151,1436,193,1354]" captionTargetId="figure@9.[151,1436,193,1365]" captionTargetPageId="9" captionText="FIGURES 2633. Clubiona genevensis L. Koch, 1866, female genitalia: 26 epigyne (from France), ventral view; 27, 28 same (from Greece); 29 vulva (from France), ventral view, reflected light; 30 same, dorsal view; 31 same, ventral view; 32 vulva (from France), dorsal view; 33 same, ventro-dorsal view, transmitted light. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064426/files/figure.png" pageId="2" pageNumber="3">Figs 2932</figureCitation>
,
<figureCitation id="137D38C60904FF981997FAEC171FFA80" box="[1233,1305,1340,1364]" captionStart="FIGURES3441" captionStartId="10.[151,264,1897,1919]" captionTargetBox="[168,1427,446,1876]" captionTargetId="figure@10.[163,1427,435,1887]" captionTargetPageId="10" captionText="FIGURES3441. Clubiona leucaspis Simon, 1932:34 male (from Spain) habitus, dorsal view; 35, 36 same, females; 37 male (from Spain) palp, retrolateral view; 38 same, ventral view; 39 female (from Spain) epigyne, ventral view; 40 female (from France) vulva, ventral view; 41 same, dorsal view. Abbreviatures: A = atrium, CD = longitudinal copulatory duct, CS = cymbial modified setae, S = spermathecae.Scale bars: 3436 = 1 mm; 3741 = 0.2 mm." httpUri="https://zenodo.org/record/1064428/files/figure.png" pageId="2" pageNumber="3">40, 41</figureCitation>
,
<figureCitation id="137D38C60904FF981863FAEC1769FA81" box="[1317,1391,1340,1365]" captionStart="FIGURES 4551" captionStartId="13.[151,264,1902,1924]" captionTargetBox="[166,1436,237,1832]" captionTargetId="figure@13.[151,1436,220,1850]" captionTargetPageId="13" captionText="FIGURES 4551. Clubiona vegeta Simon, 1918, male and female (from Tunisia): 45 male palp, retrolateral view; 46 same, ventral view; 47 epigyne, ventral view; 48 vulva, ventral view, reflected light; 49 same, dorsal view; 50 vulva, ventral view, transmitted light; 51 same, dorsal view.Abbreviatures:A = atrium, CD = longitudinal copulatory duct,CS = cymbial modified setae, S = spermathecae.Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/1064432/files/figure.png" pageId="2" pageNumber="3">4851</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>