486 lines
55 KiB
XML
486 lines
55 KiB
XML
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<mods:title id="E9DFC139C47F35B03D162D1C101EB8B2">Revision of the montane New Guinean skink genus Lobulia (Squamata: Scincidae), with the description of four new genera and nine new species</mods:title>
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<mods:namePart id="B07CFC8378437BDE860B810E263C81B0">Slavenko, Alex</mods:namePart>
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<mods:namePart id="82E7056ADA02EA52DF77019711B4C48B">Tamar, Karin</mods:namePart>
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<mods:namePart id="D966B8EAB8B8371A926F79168A7DD62D">Tallowin, Oliver J S</mods:namePart>
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<mods:namePart id="1C61620F2F0158716E5F38DF25457632">Kraus, Fred</mods:namePart>
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<mods:namePart id="E4E4FD502C61A8A64FA4F833DFE30773">Allison, Allen</mods:namePart>
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<mods:namePart id="A62230E40F9B47099F35B83CCEC6D592">Carranza, Salvador</mods:namePart>
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<mods:namePart id="3FF17A23D20335D34B432D086B6CE77F">Meiri, Shai</mods:namePart>
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<mods:title id="FA550FDB43CEB511B8DA2B8203D0DF08">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="6A5F39149E4882B96AC0E0A608107DD1">2022</mods:date>
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<mods:number id="69FBF9ABD9645A8D2E849C69CADC655E">2022-05-01</mods:number>
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<mods:number id="3A0F07DC147040B9982D38CC71134810">195</mods:number>
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<mods:identifier id="B440C04798D5703BBD5AEFF600248238" type="DOI">10.1093/zoolinnean/zlab052</mods:identifier>
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<mods:identifier id="7FF2EE984FD1B2B1B2FB595B9495AF59" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03EF4E777C32CA5DFEBEFF7021AFFE37" ID-DOI="http://doi.org/10.5281/zenodo.6536330" ID-Zenodo-Dep="6536330" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37" httpUri="http://treatment.plazi.org/id/03EF4E777C32CA5DFEBEFF7021AFFE37" lastPageId="39" lastPageNumber="258" pageId="37" pageNumber="257">
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<subSubSection id="C35CACEA7C32CA5FFEBEFF702222FEA6" pageId="37" pageNumber="257" type="nomenclature">
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<paragraph id="8BF9FF617C32CA5FFEBEFF70221DFF78" blockId="37.[270,635,196,220]" box="[270,635,196,220]" pageId="37" pageNumber="257">
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<heading id="D0B1480D7C32CA5FFEBEFF70221DFF78" box="[270,635,196,220]" centered="true" fontSize="9" level="2" pageId="37" pageNumber="257" reason="2">
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<taxonomicName id="4C4684E27C32CA5FFEBEFF702277FF7E" ID-CoL="8SHRD" authority="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri, 2022" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[270,529,196,219]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="marmorata" status="sp. nov.">LOBULIA MARMORATA</taxonomicName>
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<taxonomicNameLabel id="A2019E087C32CA5FFDA8FF73221DFF78" box="[536,635,196,220]" pageId="37" pageNumber="257" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BF9FF617C32CA5FFEF5FF5E2222FEA6" blockId="37.[325,580,234,259]" box="[325,580,234,259]" pageId="37" pageNumber="257">MARBLED MOSS SKINK</paragraph>
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</subSubSection>
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<subSubSection id="C35CACEA7C32CA5DFE87FEA521AFFE37" lastPageId="39" lastPageNumber="259" pageId="37" pageNumber="257" type="description">
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<paragraph id="8BF9FF617C32CA5FFE87FEA52234FE8E" blockId="37.[311,594,273,298]" box="[311,594,273,298]" pageId="37" pageNumber="257">
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(
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<figureCitation id="137DE3E47C32CA5FFEF0FEA521ECFE8E" box="[320,394,273,298]" captionStart="Figure 6" captionStartId="15.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1491]" captionTargetId="figure-158@15.[145,1425,195,1491]" captionTargetPageId="15" captionText="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images C–E by Allen Allison." figureDoi="http://doi.org/10.5281/zenodo.6530711" httpUri="https://zenodo.org/record/6530711/files/figure.png" pageId="37" pageNumber="257">FIGS 6</figureCitation>
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,
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<figureCitation id="137DE3E47C32CA5FFE28FEA521B8FE8E" box="[408,478,273,298]" captionStart-0="Figure 13" captionStart-1="Figure 14" captionStartId-0="22.[163,242,1550,1572]" captionStartId-1="23.[145,225,545,567]" captionTargetBox-0="[420,1187,196,1511]" captionTargetBox-1="[145,769,201,506]" captionTargetId-0="figure-169@22.[419,1187,195,1511]" captionTargetId-1="figure-857@23.[145,769,195,506]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Figure 13. Lobulia marmorata holotype (BPBM 34150), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-1="Figure 14. Lobulia marmorata holotype (BPBM 34150), ventral view in life. Image by Fred Kraus." figureDoi-0="http://doi.org/10.5281/zenodo.6530729" figureDoi-1="http://doi.org/10.5281/zenodo.6530731" httpUri-0="https://zenodo.org/record/6530729/files/figure.png" httpUri-1="https://zenodo.org/record/6530731/files/figure.png" pageId="37" pageNumber="257">13–14</figureCitation>
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;
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<tableCitation id="C6C4CADA7C32CA5FFE5AFEA5222CFE8E" box="[490,586,273,298]" captionStart="Table 1" captionStartId="9.[142,206,414,436]" captionTargetPageId="9" captionText="Table 1. Ranges of morphometrics and meristics for specimens of species treated in this study (mean values in parentheses). Only adult specimens were examined for morphometric measurements, but juveniles were included for meristics" httpUri="http://table.plazi.org/id/DF39AFE97C1ECA73FF3EFE2A2460FE75" pageId="37" pageNumber="257" tableUuid="DF39AFE97C1ECA73FF3EFE2A2460FE75">TABLE 1</tableCitation>
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)
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</paragraph>
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<paragraph id="8BF9FF617C32CA5FFF21FE8D229FFECB" blockId="37.[145,761,312,367]" pageId="37" pageNumber="257">
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<emphasis id="B93223737C32CA5FFF21FE8D21B8FEF4" box="[145,478,312,336]" italics="true" pageId="37" pageNumber="257">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n:l s i d: z o o b a n k. org:act:
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</paragraph>
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<paragraph id="8BF9FF617C32CA5FFF21FE2B224AFDB4" blockId="37.[145,761,415,529]" pageId="37" pageNumber="257">
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<materialsCitation id="3B2EF53C7C32CA5FFF21FE2B224AFDB4" collectingDate="2009-03-23" collectorName="F. Kraus" country="Papua New Guinea" elevation="1900" latitude="-5.639" location="Muller Range" longLatPrecision="78" longitude="142.625" pageId="37" pageNumber="257" specimenCount="1" stateProvince="Hela" typeStatus="holotype">
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<typeStatus id="54FD41C37C32CA5FFF21FE2B209DFE10" box="[145,251,415,436]" pageId="37" pageNumber="257" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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BPBM 34150 (field tag FK 12731), adult male, collected by
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at
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,
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(WGS 84),
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a.s.l.
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</elevation>
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,
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,
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,
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<typeStatus id="54FD41C37C32CA5FFF21FDF62098FDF3" box="[145,254,578,599]" pageId="37" pageNumber="257" type="paratype">Paratypes</typeStatus>
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(N = 32):
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</emphasis>
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:
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<collectingRegion id="498231837C32CA5FFDE6FDF62290FDFC" box="[598,758,578,600]" country="Papua New Guinea" name="Hela" pageId="37" pageNumber="257">Hela Province</collectingRegion>
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:
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<location id="8E99A9BA7C32CA5FFF21FDD4214DFDD2" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFF21FDD4214DFDD2" box="[145,299,608,630]" country="Papua New Guinea" name="Muller Range" pageId="37" pageNumber="257" stateProvince="Hela">Muller Range</location>
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: same locality as holotype (BPBM 34151– 55, 34157;
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<specimenCount id="9D4034E87C32CA5FFEA8FDCB21F3FD31" box="[280,405,639,661]" pageId="37" pageNumber="257" type="male">two males</specimenCount>
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,
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,
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<specimenCount id="9D4034E87C32CA5FFDF7FDCB228AFD31" box="[583,748,639,661]" pageId="37" pageNumber="257" type="juvenile">two juveniles</specimenCount>
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)
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;
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,
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<geoCoordinate id="EE7299A67C32CA5FFF4AFD2A2112FD10" box="[250,372,670,692]" degrees="142.634" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.634">142.634°E</geoCoordinate>
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(WGS 84),
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a.s.l. (BPBM 34156, 34162–23;
|
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<specimenCount id="9D4034E87C32CA5FFEECFD0921B6FD76" box="[348,464,700,722]" pageId="37" pageNumber="257" type="male">two males</specimenCount>
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,
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);
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<materialsCitation id="3B2EF53C7C32CA5FFDDAFD082263FCB4" country="Papua New Guinea" elevation="1966" latitude="-5.666" location="Mt Yakapi" longLatPrecision="78" longitude="142.643" pageId="37" pageNumber="257" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hela Province" typeStatus="paratype">
|
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“
|
||
<location id="8E99A9BA7C32CA5FFDC5FD08228DFD76" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFDC5FD08228DFD76" box="[629,747,700,722]" country="Papua New Guinea" latitude="-5.666" longLatPrecision="78" longitude="142.643" name="Mt Yakapi" pageId="37" pageNumber="257" stateProvince="Hela Province">Mt Yakapi</location>
|
||
”,
|
||
<geoCoordinate id="EE7299A67C32CA5FFF21FD6F208AFD55" box="[145,236,731,753]" degrees="5.666" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="55" value="-5.666">5.666°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFF4AFD6F2112FD55" box="[250,372,731,753]" degrees="142.643" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.643">142.643°E</geoCoordinate>
|
||
(WGS 84),
|
||
<elevation id="006B18527C32CA5FFDB1FD6F22FFFD55" box="[513,665,731,753]" metricMagnitude="3" metricUnit="m" metricValue="1.966" pageId="37" pageNumber="257" unit="m" value="1966.0">
|
||
<quantity id="4CBE52847C32CA5FFDB1FD6F223FFD54" box="[513,601,731,753]" metricMagnitude="3" metricUnit="m" metricValue="1.966" pageId="37" pageNumber="257" unit="m" value="1966.0">1966 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(BPBM 34158–60;
|
||
<specimenCount id="9D4034E87C32CA5FFEB8FD4E211CFCB4" box="[264,378,762,784]" pageId="37" pageNumber="257" type="male">two males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFE34FD4E2198FCB4" box="[388,510,762,784]" pageId="37" pageNumber="257" type="female">one female</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2EF53C7C32CA5FFDBFFD4E21C1FCC8" country="Papua New Guinea" elevation="1859" latitude="-5.64" location="Dickson's Village" longLatPrecision="558" longitude="142.628" municipality="Point" pageId="37" pageNumber="257" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Hela Province" typeStatus="paratype">
|
||
“
|
||
<collectingMunicipality id="6B9D651B7C32CA5FFDA9FD4E2235FCB4" box="[537,595,762,784]" pageId="37" pageNumber="257">Point</collectingMunicipality>
|
||
17 =
|
||
<location id="8E99A9BA7C32CA5FFD20FD4E2080FC8A" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFD20FD4E2080FC8A" country="Papua New Guinea" latitude="-5.64" longLatPrecision="558" longitude="142.628" municipality="Point" name="Dickson's Village" pageId="37" pageNumber="257" stateProvince="Hela Province">Dickson’s Village</location>
|
||
”,
|
||
<geoCoordinate id="EE7299A67C32CA5FFF4EFCAC212DFC8A" box="[254,331,792,814]" degrees="5.64" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="555" value="-5.64">5.64°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFEEAFCAC21B5FC8A" box="[346,467,792,814]" degrees="142.628" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.628">142.628°E</geoCoordinate>
|
||
(WGS 84),
|
||
<elevation id="006B18527C32CA5FFDD0FCAC229EFC8A" box="[608,760,792,814]" metricMagnitude="3" metricUnit="m" metricValue="1.859" pageId="37" pageNumber="257" unit="m" value="1859.0">
|
||
<quantity id="4CBE52847C32CA5FFDD0FCAC22DEFC8A" box="[608,696,792,814]" metricMagnitude="3" metricUnit="m" metricValue="1.859" pageId="37" pageNumber="257" unit="m" value="1859.0">1859 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(BPBM 34164–5, 34169–70, PNGNM 25284;
|
||
<specimenCount id="9D4034E87C32CA5FFD33FC832293FCE9" box="[643,757,823,845]" pageId="37" pageNumber="257" type="male">two males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFF21FCE22161FCC8" box="[145,263,854,876]" pageId="37" pageNumber="257" type="female">one female</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFEBEFCE221F9FCC8" box="[270,415,854,876]" pageId="37" pageNumber="257" type="juvenile">two juveniles</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2EF53C7C32CA5FFE1EFCE22096FC0D" country="Papua New Guinea" elevation="1910" latitude="-5.6431" location="Kunida" longLatPrecision="55" longitude="142.634" pageId="37" pageNumber="257" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hela Province" typeStatus="paratype">
|
||
“
|
||
<location id="8E99A9BA7C32CA5FFE08FCE2226CFCC8" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFE08FCE2226CFCC8" box="[440,522,854,876]" country="Papua New Guinea" latitude="-5.6431" longLatPrecision="55" longitude="142.634" name="Kunida" pageId="37" pageNumber="257" stateProvince="Hela Province">Kunida</location>
|
||
”,
|
||
<geoCoordinate id="EE7299A67C32CA5FFDACFCE12219FCCF" box="[540,639,853,875]" degrees="5.6431" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="5" value="-5.6431">5.6431°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFD37FCE2229FFCC8" box="[647,761,854,876]" degrees="142.634" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.634">142.634°E</geoCoordinate>
|
||
(WGS 84),
|
||
<elevation id="006B18527C32CA5FFEB9FCC021F3FC2E" box="[265,405,884,906]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="37" pageNumber="257" unit="m" value="1910.0">
|
||
<quantity id="4CBE52847C32CA5FFEB9FCC0213AFC2E" box="[265,348,884,906]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="37" pageNumber="257" unit="m" value="1910.0">1910 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(BPBM 34166–8;
|
||
<specimenCount id="9D4034E87C32CA5FFDD0FCC122A3FC2E" box="[608,709,884,906]" pageId="37" pageNumber="257" type="male">one male</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFD60FCC1208FFC0D" pageId="37" pageNumber="257" type="female">two females</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B2EF53C7C32CA5FFF4AFC272246FC42" country="Papua New Guinea" elevation="1777" latitude="-5.6454" location="Dickson's House" longLatPrecision="55" longitude="142.639" pageId="37" pageNumber="257" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hela Province" typeStatus="paratype">
|
||
“
|
||
<location id="8E99A9BA7C32CA5FFEB4FC2721DCFC0C" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFEB4FC2721DCFC0C" box="[260,442,915,937]" country="Papua New Guinea" latitude="-5.6454" longLatPrecision="55" longitude="142.639" name="Dickson's House" pageId="37" pageNumber="257" stateProvince="Hela Province">Dickson’s House</location>
|
||
”,
|
||
<geoCoordinate id="EE7299A67C32CA5FFE7FFC272255FC0D" box="[463,563,915,937]" degrees="5.6454" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="5" value="-5.6454">5.6454°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFD8DFC2722D6FC0D" box="[573,688,915,937]" degrees="142.639" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.639">142.639°E</geoCoordinate>
|
||
(WGS 84),
|
||
<elevation id="006B18527C32CA5FFF74FC06213FFC6C" box="[196,345,946,968]" metricMagnitude="3" metricUnit="m" metricValue="1.777" pageId="37" pageNumber="257" unit="m" value="1777.0">
|
||
<quantity id="4CBE52847C32CA5FFF74FC06217DFC63" box="[196,283,946,967]" metricMagnitude="3" metricUnit="m" metricValue="1.777" pageId="37" pageNumber="257" unit="m" value="1777.0">1777 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(BPBM 34171–73, 34177, PNGNM 25281–83;
|
||
<specimenCount id="9D4034E87C32CA5FFEB6FC64211DFC42" box="[262,379,976,998]" pageId="37" pageNumber="257" type="male">four males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFE33FC64227CFC42" box="[387,538,976,998]" pageId="37" pageNumber="257" type="female">three females</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<geoCoordinate id="EE7299A67C32CA5FFD99FC642218FC42" box="[553,638,976,998]" degrees="5.652" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="55" value="-5.652">5.652°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFD37FC64229FFC42" box="[647,761,976,998]" degrees="142.643" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.643">142.643°E</geoCoordinate>
|
||
(WGS 84),
|
||
<quantity id="4CBE52847C32CA5FFEB6FC5B213EFBA0" box="[262,344,1007,1029]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="37" pageNumber="257" unit="m" value="1860.0">1860 m</quantity>
|
||
a.s.l. (BPBM 34174–76;
|
||
<specimenCount id="9D4034E87C32CA5FFDD2FC5B22A1FBA1" box="[610,711,1007,1029]" pageId="37" pageNumber="257" type="male">one male</specimenCount>
|
||
,
|
||
<specimenCount id="9D4034E87C32CA5FFD60FC5B208BFB80" pageId="37" pageNumber="257" type="female">two females</specimenCount>
|
||
);
|
||
<geoCoordinate id="EE7299A67C32CA5FFEB1FBB9213DFB87" box="[257,347,1037,1059]" degrees="5.639" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="55" value="-5.639">5.639°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFED7FBB921B8FB87" box="[359,478,1037,1059]" degrees="142.625" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.625">142.625°E</geoCoordinate>
|
||
(WGS 84),
|
||
<quantity id="4CBE52847C32CA5FFDD5FBBA22DDFB87" box="[613,699,1038,1059]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="37" pageNumber="257" unit="m" value="1900.0">1900 m</quantity>
|
||
a.s.l. (BPBM 34178; male);
|
||
<materialsCitation id="3B2EF53C7C32CA5FFE33FB9822FEFBC5" country="Papua New Guinea" elevation="1910" latitude="-5.6591" location="Top House" longLatPrecision="55" longitude="142.635" pageId="37" pageNumber="257" specimenCount="1" stateProvince="Hela Province" typeStatus="paratype">
|
||
“
|
||
<location id="8E99A9BA7C32CA5FFE3DFB982265FBE5" LSID="urn:lsid:plazi:treatment:03EF4E777C32CA5DFEBEFF7021AFFE37:8E99A9BA7C32CA5FFE3DFB982265FBE5" box="[397,515,1068,1089]" country="Papua New Guinea" latitude="-5.6591" longLatPrecision="55" longitude="142.635" name="Top House" pageId="37" pageNumber="257" stateProvince="Hela Province">Top House</location>
|
||
”,
|
||
<geoCoordinate id="EE7299A67C32CA5FFDA8FB98221DFBE6" box="[536,635,1068,1090]" degrees="5.6591" direction="south" orientation="latitude" pageId="37" pageNumber="257" precision="5" value="-5.6591">5.6591°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C32CA5FFD37FB98229FFBE6" box="[647,761,1068,1090]" degrees="142.635" direction="east" orientation="longitude" pageId="37" pageNumber="257" precision="55" value="142.635">142.635°E</geoCoordinate>
|
||
(WGS 84),
|
||
<elevation id="006B18527C32CA5FFEB7FBFF21F7FBC5" box="[263,401,1099,1121]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="37" pageNumber="257" unit="m" value="1910.0">
|
||
<quantity id="4CBE52847C32CA5FFEB7FBFF213FFBC4" box="[263,345,1099,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.91" pageId="37" pageNumber="257" unit="m" value="1910.0">1910 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(PNGNM 25285; male)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C32CA5FFF21FB21229CF976" blockId="37.[145,762,1172,1746]" pageId="37" pageNumber="257">
|
||
<emphasis id="B93223737C32CA5FFF21FB21216FFB0E" box="[145,265,1173,1194]" italics="true" pageId="37" pageNumber="257">Diagnosis:</emphasis>
|
||
A medium-sized species of
|
||
<taxonomicName id="4C4684E27C32CA5FFDE3FB2022CAFB0D" authorityName="Greer" authorityYear="1974" box="[595,684,1172,1193]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C32CA5FFDE3FB2022CAFB0D" box="[595,684,1172,1193]" italics="true" pageId="37" pageNumber="257">Lobulia</emphasis>
|
||
</taxonomicName>
|
||
(adult SVL
|
||
<quantity id="4CBE52847C32CA5FFF60FB0721E4FB6C" box="[208,386,1203,1225]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" metricValueMax="5.6899999999999995" metricValueMin="4.1899999999999995" pageId="37" pageNumber="257" unit="mm" value="49.4" valueMax="56.9" valueMin="41.9">41.9–56.9 mm</quantity>
|
||
), characterized by the unique combination of frontoparietals fused; supraorbital ridges typically pronounced; nuchals 2–4 pairs; paravertebral scales 46–56; mid-body scale rows 30–34; 4
|
||
<superScript id="7C3352297C32CA5FFF40FA992098FA9D" attach="left" box="[240,254,1325,1337]" fontSize="5" pageId="37" pageNumber="257">th</superScript>
|
||
digit on front foot longer than 3
|
||
<superScript id="7C3352297C32CA5FFDDBFA99221CFA9D" attach="none" box="[619,634,1325,1337]" fontSize="5" pageId="37" pageNumber="257">rd</superScript>
|
||
; subdigital lamellae 20–25 under 4
|
||
<superScript id="7C3352297C32CA5FFE29FAFF21C1FAF3" attach="left" box="[409,423,1355,1367]" fontSize="5" pageId="37" pageNumber="257">th</superScript>
|
||
toe; single supradigital scales 3–4 on 4
|
||
<superScript id="7C3352297C32CA5FFF40FADE2098FAD2" attach="left" box="[240,254,1386,1398]" fontSize="5" pageId="37" pageNumber="257">th</superScript>
|
||
toe; mid-dorsum with two rows of large dark brown spots converging to a single row roughly around midbody; top of tail with two rows of dark brown spots joining ventrally with dark lateral stripes along the tail length; light blue or white dorsolateral stripes absent; flanks dark brown spotted with grey; light blue or white lateral stripes absent; uniform coloration on abdomen and base of tail, lemon yellow in life, light blue in preservative; thighs and precloacal region without brown spotting; tail and chin uniform light blue speckled with brown spots; palmar and plantar surfaces dark yellow in life, light brown in preservative.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C32CA5FFF21F94F2426FED0" blockId="37.[145,762,1787,1901]" lastBlockId="37.[809,1425,197,372]" pageId="37" pageNumber="257">
|
||
<emphasis id="B93223737C32CA5FFF21F94F2250F8B5" box="[145,566,1787,1809]" italics="true" pageId="37" pageNumber="257">
|
||
Comparisons:
|
||
<taxonomicName id="4C4684E27C32CA5FFEFAF94F2250F8B5" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[330,566,1787,1809]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="marmorata">Lobulia marmorata</taxonomicName>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C4684E27C32CA5FFD66F9482083F88B" baseAuthorityName="Kraus" baseAuthorityYear="2020" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B93223737C32CA5FFD66F9482083F88B" italics="true" pageId="37" pageNumber="257">Lo. elegans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4684E27C32CA5FFF41F8AE2108F88B" authorityName=", Kraus" authorityYear="2020" box="[241,366,1818,1839]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="lobulus">
|
||
<emphasis id="B93223737C32CA5FFF41F8AE2108F88B" box="[241,366,1818,1839]" italics="true" pageId="37" pageNumber="257">Lo. lobulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4684E27C32CA5FFE17F8AE2224F88B" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[423,578,1818,1839]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="huonensis">
|
||
<emphasis id="B93223737C32CA5FFE17F8AE2224F88B" box="[423,578,1818,1839]" italics="true" pageId="37" pageNumber="257">Lo. huonensis</emphasis>
|
||
</taxonomicName>
|
||
in having fused vs. unfused frontoparietals.
|
||
<taxonomicName id="4C4684E27C32CA5FFE77F88C22C3F8EA" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[455,677,1848,1870]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="B93223737C32CA5FFE77F88C22C3F8EA" box="[455,677,1848,1870]" italics="true" pageId="37" pageNumber="257">Lobulia marmorata</emphasis>
|
||
</taxonomicName>
|
||
is most similar in scalation and coloration to
|
||
<taxonomicName id="4C4684E27C32CA5FFD8CF8EC2295F8C8" authorityName="Zweifel" authorityYear="1972" box="[572,755,1879,1901]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="257" phylum="Chordata" rank="species" species="brongersmai">
|
||
<emphasis id="B93223737C32CA5FFD8CF8EC2295F8C8" box="[572,755,1879,1901]" italics="true" pageId="37" pageNumber="257">Lo. brongersmai</emphasis>
|
||
</taxonomicName>
|
||
, but differs from it in having a higher average count of midbody scale rows [31.5 (30–34) vs. 29 (27–32)], a dark blotch on the nuchal region, posterior to the parietals, and a generally darker coloration due to larger size of the dark brown dorsal spots relative to the base grey coloration.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C32CA5CFC99FE222283FC5C" blockId="37.[809,1425,405,1899]" lastBlockId="38.[163,779,197,1016]" lastPageId="38" lastPageNumber="258" pageId="37" pageNumber="257">
|
||
<emphasis id="B93223737C32CA5FFC99FE222430FE0E" box="[809,1110,405,427]" italics="true" pageId="37" pageNumber="257">
|
||
Description of the
|
||
<typeStatus id="54FD41C37C32CA5FFC41FE212436FE0E" box="[1009,1104,405,426]" pageId="37" pageNumber="257" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Rostral broad and shallow, wider than deep, projecting slightly onto top of snout; nasals more or less rectangular, separated by rostral and frontonasal contact, projecting anterodorsally onto dorsum of snout; nostril circular, centred within nasal; frontonasal large, with eight sides, extending laterally to slightly above the level of nares, in broad contact with frontal; prefrontals large, separated by frontonasal and frontal contact, bordered ventrolaterally by two loreals; supraoculars four, anterior two in contact with frontal, posterior three in contact with frontoparietal; frontal kite shaped, widest anteriorly; frontoparietal single, anteriorly in contact with frontal, posteriorly with interparietal and parietals; interparietal smaller than fused frontoparietal, kite shaped, widest anteriorly; parietal eye spot absent; parietals in contact behind interparietal, in contact anteriorly with frontoparietal, posteriormost supraocular and two pretemporals; nuchals two pairs, transversely enlarged, wider than long, separated from secondary temporal by a single intercalated scale, with a third nuchal on the right side. Anterior loreal slightly smaller than posterior loreal, roughly as long as high; posterior loreal longer than high; lower preocular roughly square in shape; upper preocular much smaller, longer than high; presubocular single; postsuboculars three, lowest interdigitated between subocular supralabial and penultimate supralabial; lower eyelid scaly, moveable, with a clear palpebral disc roughly the size of ear opening; supraciliaries nine, anteriormost in narrow contact with frontal, posteriormost projecting medially and interdigitated between posteriormost supraocular and upper pretemporal; primary temporal single, ventrally contacting posteriormost supralabial; secondary temporals two, upper larger and overlapping lower; supralabials seven, fifth in contact with small scales of lower eyelid; postsupralabials two; ear opening moderately large, with lobules along anterior margin. Mental single; postmental single, contacting two anteriormost infralabials; infralabials seven; enlarged chin shields four pairs, the first two pairs in medial contact, third pair narrowly separated by a single medial scale, fourth pair separated by three medial scales; posteriormost chin shield in contact with penultimate infralabial. Body scales smooth, in 31 rows at midbody; paravertebral scales 50; medial precloacal scales enlarged, overlapping lateral precloacals. Scales on dorsal surface of 4th toe in two rows proximally, single row distally beginning at third interphalangeal joint, three single scales; subdigital lamellae under 4th toe 20, smooth. In preservative (
|
||
<figureCitation id="137DE3E47C31CA5CFF1BFEB6209CFEBC" box="[171,250,258,280]" captionStart="Figure 13" captionStartId="22.[163,242,1550,1572]" captionTargetBox="[420,1187,196,1511]" captionTargetId="figure-169@22.[419,1187,195,1511]" captionTargetPageId="22" captionText="Figure 13. Lobulia marmorata holotype (BPBM 34150), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." figureDoi="http://doi.org/10.5281/zenodo.6530729" httpUri="https://zenodo.org/record/6530729/files/figure.png" pageId="38" pageNumber="258">Fig. 13</figureCitation>
|
||
), base dorsal coloration grey-brown, with two mid-dorsal parallel rows of large dark brown spots two to four scales long, converging posterior to forelimbs to form a single continuous mid-dorsal dark stripe with transverse lateral projections, extending to base of tail; on dorsal surface of tail, two parallel rows of dark brown blotches extending medially from dark brown lateral stripes; light blue or white dorsolateral stripes absent, but parallel dorsolateral rows of dark brown blotches present, extending laterally to lateral fields; lateral field dark brown, speckled with grey spots; light blue or white lateral stripe absent; head similar in coloration to dorsum; dark brown spotting on head scales, mostly along scale margins, with large dark spot in centre of frontal; large dark blotch in nuchal region formed by dark coloration on the posterior margins of parietals and medial margins of nuchals; ventral surfaces uniform light blue; brown dusting present on chin; scales on palmar and plantar surfaces light brown, contrasting with dark brown digits. In life (
|
||
<figureCitation id="137DE3E47C31CA5CFD6AFCFD20D7FCDA" captionStart="Figure 6" captionStartId="15.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1491]" captionTargetId="figure-158@15.[145,1425,195,1491]" captionTargetPageId="15" captionText="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images C–E by Allen Allison." figureDoi="http://doi.org/10.5281/zenodo.6530711" httpUri="https://zenodo.org/record/6530711/files/figure.png" pageId="38" pageNumber="258">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="137DE3E47C31CA5CFF70FCDC20BAFCD9" box="[192,220,872,893]" captionStart="Figure 14" captionStartId="23.[145,225,545,567]" captionTargetBox="[145,769,201,506]" captionTargetId="figure-857@23.[145,769,195,506]" captionTargetPageId="23" captionText="Figure 14. Lobulia marmorata holotype (BPBM 34150), ventral view in life. Image by Fred Kraus." figureDoi="http://doi.org/10.5281/zenodo.6530731" httpUri="https://zenodo.org/record/6530731/files/figure.png" pageId="38" pageNumber="258">14</figureCitation>
|
||
), dorsum greyish with black mid-dorsal spots; lateral field brown with white spotting; chin white; chest, abdomen, thighs, precloacal region and base of tail lemon yellow, with tail becoming white posteriorly; scales on palmar and plantar surfaces dark yellow.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFF13FBAE24C8FD88" blockId="38.[163,779,1049,1899]" lastBlockId="38.[827,1443,197,893]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFF13FBAE2142FB8B" box="[163,292,1050,1071]" italics="true" pageId="38" pageNumber="258">Variation:</emphasis>
|
||
Adult body size
|
||
<quantity id="4CBE52847C31CA5CFDA4FBAD22ACFB8A" box="[532,714,1049,1071]" metricMagnitude="-2" metricUnit="m" metricValue="4.9399999999999995" metricValueMax="5.6899999999999995" metricValueMin="4.1899999999999995" pageId="38" pageNumber="258" unit="mm" value="49.4" valueMax="56.9" valueMin="41.9">41.9–56.9 mm</quantity>
|
||
SVL (mean = 48.1, SD = 3.8,
|
||
<emphasis id="B93223737C31CA5CFE1DFB8C21B0FBE9" box="[429,470,1080,1101]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
29). Females (mean = 50.5, range: 42–56.9, SD = 4.0,
|
||
<emphasis id="B93223737C31CA5CFE65FBE32267FBC8" box="[469,513,1111,1132]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
12) larger than males (mean = 46.4, range: 41.9–50.5, SD = 2.6,
|
||
<emphasis id="B93223737C31CA5CFD00FBC222BBFB2F" box="[688,733,1142,1163]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
17;
|
||
<emphasis id="B93223737C31CA5CFF13FB2020CAFB0D" box="[163,172,1172,1193]" italics="true" pageId="38" pageNumber="258">t</emphasis>
|
||
= 3.4,
|
||
<emphasis id="B93223737C31CA5CFEBAFB20217CFB0D" box="[266,282,1172,1193]" italics="true" pageId="38" pageNumber="258">P</emphasis>
|
||
<0.01). Forelimbs 36.8–47.0% of SVL (mean = 42.4%, SD = 2.5,
|
||
<emphasis id="B93223737C31CA5CFE6BFB072261FB6C" box="[475,519,1203,1224]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
29). Hindlimbs 44.1– 53.4% of SVL (mean = 47.5%, SD = 2.8,
|
||
<emphasis id="B93223737C31CA5CFDDBFB6622F3FB43" box="[619,661,1234,1255]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
29). Scale rows at midbody 30–34 (mean = 31.5, SD = 0.9,
|
||
<emphasis id="B93223737C31CA5CFD03FB4422BDFAA1" box="[691,731,1264,1285]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
33); paravertebral scales 46–56 (mean = 51.2, SD = 2.2,
|
||
<emphasis id="B93223737C31CA5CFD47FABB20D4FAE7" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
33). Lamellae under 4th toe 20–25 (mean = 22.5, SD = 1.2,
|
||
<emphasis id="B93223737C31CA5CFEAAFAF82120FAC5" box="[282,326,1356,1377]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
32); single supradigital scales on 4th toe 3–4 (mean = 3.2, SD = 0.4,
|
||
<emphasis id="B93223737C31CA5CFDABFADF2221FA24" box="[539,583,1387,1408]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
32). Mostly 2–4 pairs of nuchals, but BPBM 34150–51, 34155–56, 34166, 34169–71, 34173, 34178 and PNGNM 25281 have an asymmetrical number of nuchals, with one more nuchal either on left side (
|
||
<emphasis id="B93223737C31CA5CFD81FA522223FA5F" box="[561,581,1510,1531]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 6) or on right side (
|
||
<emphasis id="B93223737C31CA5CFF51F9B02093F9BD" box="[225,245,1540,1561]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 5). Primary nuchals usually separated from secondary temporals by single smaller intercalated scale (
|
||
<emphasis id="B93223737C31CA5CFF5CF9F62166F9F3" box="[236,256,1602,1623]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 27), rarely by none on left side and one on right (
|
||
<emphasis id="B93223737C31CA5CFF5BF9D42099F9D1" box="[235,255,1632,1653]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 1), one on left side and two on right (
|
||
<emphasis id="B93223737C31CA5CFD0EF9D422B4F9D1" box="[702,722,1632,1653]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 1), two on left side and one on right (
|
||
<emphasis id="B93223737C31CA5CFDAEF9CB2254F930" box="[542,562,1663,1684]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 3) or two on both sides (
|
||
<emphasis id="B93223737C31CA5CFF5CF92A2166F917" box="[236,256,1694,1715]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 1). Supraorbital ridges usually pronounced (
|
||
<emphasis id="B93223737C31CA5CFF1CF90820A6F975" box="[172,192,1724,1745]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 25), but occasionally not (
|
||
<emphasis id="B93223737C31CA5CFDACF9082256F975" box="[540,560,1724,1745]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 8). Frontonasal usually as wide as long (
|
||
<emphasis id="B93223737C31CA5CFE76F96F21BCF954" box="[454,474,1755,1776]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 25), occasionally longer than wide (
|
||
<emphasis id="B93223737C31CA5CFE91F94E2153F8AB" box="[289,309,1786,1807]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 8). Frontonasal fragmented in PNGNM 25285. Interparietal fused with frontoparietal in BPBM 34151 and 34153. Loreals typically two (
|
||
<emphasis id="B93223737C31CA5CFD46F883236CF8E8" box="[758,778,1847,1868]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 29), rarely three (
|
||
<emphasis id="B93223737C31CA5CFE34F8E221FEF8CF" box="[388,408,1878,1899]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 4). Supraciliaries either eight (
|
||
<emphasis id="B93223737C31CA5CFCF3FF722331FF7F" box="[835,855,198,219]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 15) or nine (
|
||
<emphasis id="B93223737C31CA5CFBB8FF72247AFF7F" box="[1032,1052,198,219]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 18). Anteriormost supraciliary usually not in contact with frontal (
|
||
<emphasis id="B93223737C31CA5CFB7BFF5024B9FF5D" box="[1227,1247,228,249]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 22), sometimes in narrow contact (
|
||
<emphasis id="B93223737C31CA5CFBA5FEB7244FFEBC" box="[1045,1065,259,280]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 11). Postsuboculars rarely two (
|
||
<emphasis id="B93223737C31CA5CFCF3FE962331FE93" box="[835,855,290,311]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 2), typically three (
|
||
<emphasis id="B93223737C31CA5CFBE5FE96240FFE93" box="[1109,1129,290,311]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 24), occasionally four (
|
||
<emphasis id="B93223737C31CA5CFA3EFE9625C4FE93" box="[1422,1442,290,311]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 7). Supralabials rarely six (
|
||
<emphasis id="B93223737C31CA5CFB28FEF424CAFEF1" box="[1176,1196,320,341]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 1), typically seven (
|
||
<emphasis id="B93223737C31CA5CFCF4FEEB233EFED0" box="[836,856,351,372]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 20), occasionally eight (
|
||
<emphasis id="B93223737C31CA5CFB13FEEB24D1FED0" box="[1187,1207,351,372]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 12). Infralabials typically seven (
|
||
<emphasis id="B93223737C31CA5CFBB0FEC92472FE36" box="[1024,1044,381,402]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 25), occasionally eight (
|
||
<emphasis id="B93223737C31CA5CFAE1FEC92503FE36" box="[1361,1381,381,402]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 8). Mental split medially in BPBM 34160. Chin shields typically four on both sides (
|
||
<emphasis id="B93223737C31CA5CFB26FE0F24CCFE74" box="[1174,1194,443,464]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 31), rarely five on right side (
|
||
<emphasis id="B93223737C31CA5CFC72FE6D23B0FE4A" box="[962,982,473,494]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 2). Infralabials posterior to contact with chin shields occasionally zero (
|
||
<emphasis id="B93223737C31CA5CFB40FE4C2562FDA9" box="[1264,1284,504,525]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 7), usually one (
|
||
<emphasis id="B93223737C31CA5CFCC3FDA323E1FD88" box="[883,903,535,556]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 24), rarely two (
|
||
<emphasis id="B93223737C31CA5CFBEFFDA32415FD88" box="[1119,1139,535,556]" italics="true" pageId="38" pageNumber="258">N</emphasis>
|
||
= 2).
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFCE3FD8124C9FCD9" blockId="38.[827,1443,197,893]" pageId="38" pageNumber="258">
|
||
Colour pattern of all specimens generally similar to
|
||
<typeStatus id="54FD41C37C31CA5CFC8BFDE723FBFDCD" box="[827,925,595,617]" pageId="38" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
, with a few exceptions. Size of mid-dorsal dark brown spots, and location at which two anterior rows converge to form single posterior row vary between individuals. In some individuals, dark blotches form a continuous thick stripe posterior to convergence; in others form a row of dark blotches. BPBM 34174 has continuous, unfrgamented lateral stripes. Degree of brown speckling on chin varies from almost no brown spots (BPBM 34156) to chin shields being almost uniformly brown (BPBM 34152).
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFC8BFC122454FB37" blockId="38.[827,1443,934,1171]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BFC12239FFC1F" box="[827,1017,934,956]" italics="true" pageId="38" pageNumber="258">Colour in life:</emphasis>
|
||
Notes for the
|
||
<typeStatus id="54FD41C37C31CA5CFB7AFC12255FFC18" box="[1226,1337,934,956]" pageId="38" pageNumber="258" type="holotype">holotype</typeStatus>
|
||
(BPBM 34150) stated “Dorsum tan with irregular black zigzags vertebrally and dorsolaterally, tan scales margined in black. Face black. Chest, belly, and under rear legs deep orange-yellow; chin and throat white with black spots. Mouth lining and tongue blue-black.”
|
||
<typeStatus id="54FD41C37C31CA5CFC8BFBEA23D2FBD7" box="[827,948,1118,1139]" pageId="38" pageNumber="258" type="paratype">Paratypes</typeStatus>
|
||
BPBM 34152 and BPBM 34155 had the venter entirely white.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFC8BFB0F248AFAAB" blockId="38.[827,1443,1211,1295]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BFB0F23AFFB74" box="[827,969,1211,1232]" italics="true" pageId="38" pageNumber="258">Etymology:</emphasis>
|
||
Feminine Latin adjective meaning “marbled”, in reference to the marbled grey and dark brown dorsal coloration of the species.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFC8BFA862490FA67" blockId="38.[827,1442,1330,1475]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BFA8623A8FAE3" box="[827,974,1330,1351]" italics="true" pageId="38" pageNumber="258">Distribution:</emphasis>
|
||
Known only from
|
||
<quantity id="4CBE52847C31CA5CFB1AFA862522FAE3" box="[1194,1348,1330,1352]" metricMagnitude="3" metricUnit="m" metricValue="1.8715" metricValueMax="1.966" metricValueMin="1.777" pageId="38" pageNumber="258" unit="m" value="1871.5" valueMax="1966.0" valueMin="1777.0">1777–1966 m</quantity>
|
||
a.s.l. on the north-eastern slopes of the Muller Range,
|
||
<collectingRegion id="498231837C31CA5CFADAFAE523C5FA21" country="Papua New Guinea" name="Hela" pageId="38" pageNumber="258">Hela Province</collectingRegion>
|
||
, where it occurs in sympatry with another, currently undescribed, species of
|
||
<taxonomicName id="4C4684E27C31CA5CFB51FA3A2526FA07" authorityName="Greer" authorityYear="1974" box="[1249,1344,1422,1443]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="258" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C31CA5CFB51FA3A2526FA07" box="[1249,1344,1422,1443]" italics="true" pageId="38" pageNumber="258">Lobulia</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 34161) having unfused frontoparietals.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFC8BFA522483F99D" blockId="38.[827,1443,1510,1593]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BFA522390FA5F" box="[827,1014,1510,1531]" italics="true" pageId="38" pageNumber="258">Natural history:</emphasis>
|
||
This species was locally common in the village areas and surrounding cleared areas; it was never observed in forested situations.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5CFC8BF9D62453F933" blockId="38.[827,1442,1634,1687]" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BF9D623BDF9D3" box="[827,987,1634,1655]" italics="true" pageId="38" pageNumber="258">Reproduction:</emphasis>
|
||
Viviparous. Litter size varies between 2–3 (mean 2.4,
|
||
<emphasis id="B93223737C31CA5CFC58F9352474F932" box="[1000,1042,1665,1686]" italics="true" pageId="38" pageNumber="258">N =</emphasis>
|
||
9).
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C31CA5DFC8BF90E21AFFE37" blockId="38.[827,1445,1722,1898]" lastBlockId="39.[145,761,197,403]" lastPageId="39" lastPageNumber="259" pageId="38" pageNumber="258">
|
||
<emphasis id="B93223737C31CA5CFC8BF90E244FF974" box="[827,1065,1722,1744]" italics="true" pageId="38" pageNumber="258">Conservation status:</emphasis>
|
||
The species is locally abundant in the areas where collected although the population trend is unknown. Based on the available samples,
|
||
<taxonomicName id="4C4684E27C31CA5CFA32F94C23DAF888" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="258" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="B93223737C31CA5CFA32F94C23DAF888" italics="true" pageId="38" pageNumber="258">Lo. marmorata</emphasis>
|
||
</taxonomicName>
|
||
has an extent of occurrence of
|
||
<quantity id="4CBE52847C31CA5CFAABF8A2250FF888" box="[1307,1385,1814,1836]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="258" unit="km" value="2.5">2.5 km</quantity>
|
||
<superScript id="7C3352297C31CA5CFAD9F8A22517F886" attach="left" box="[1385,1393,1814,1826]" fontSize="5" pageId="38" pageNumber="258">2</superScript>
|
||
and an area of occupancy of
|
||
<quantity id="4CBE52847C31CA5CFB8EF88124E5F8EF" box="[1086,1155,1845,1867]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="258" unit="km" value="16.0">16 km</quantity>
|
||
<superScript id="7C3352297C31CA5CFB33F88024EDF8E4" attach="left" box="[1155,1163,1844,1856]" fontSize="5" pageId="38" pageNumber="258">2</superScript>
|
||
(based on occupation of
|
||
<quantity id="4CBE52847C31CA5CFA25F8812339F8CE" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="38" pageNumber="258" unit="km" value="4.0">4 km</quantity>
|
||
<superScript id="7C3352297C31CA5CFCEFF8E72301F8FB" attach="left" box="[863,871,1875,1887]" fontSize="5" pageId="38" pageNumber="258">2</superScript>
|
||
cells; both calculated using http://geocat.kew.org/). However, it is likely distributed in other localities of suitable habitat in the Muller Range. It does not occur near any protected areas. Further research is needed to discern its true distributional boundaries, potential threats and demographic trends for a proper assessment. We recommend assigning a status of Data Deficient to
|
||
<taxonomicName id="4C4684E27C30CA5DFEAAFECA21A4FE37" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[282,450,382,403]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="259" phylum="Chordata" rank="species" species="marmorata">
|
||
<emphasis id="B93223737C30CA5DFEAAFECA21A4FE37" box="[282,450,382,403]" italics="true" pageId="39" pageNumber="259">Lo. marmorata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |