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262 lines
35 KiB
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<mods:title id="AC06EE734E2CF3FE051E514700401954">Revision of the montane New Guinean skink genus Lobulia (Squamata: Scincidae), with the description of four new genera and nine new species</mods:title>
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<mods:namePart id="3D829B53E06BD9D0138734B273640ABA">Tamar, Karin</mods:namePart>
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<mods:namePart id="782A22A60C65701F1DA4CDD8B7C5EF1D">Tallowin, Oliver J S</mods:namePart>
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<mods:namePart id="8531B485A7737366E74C799B7CF4CDDF">Kraus, Fred</mods:namePart>
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<mods:namePart id="6FB5367FEAEE8BCF97B2254E7D10B4C0">Allison, Allen</mods:namePart>
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<mods:namePart id="F07C2BD3495674B9CD322DEC684D76CC">Carranza, Salvador</mods:namePart>
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<mods:namePart id="D4F45D47C225592232746B9063E22564">Meiri, Shai</mods:namePart>
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<mods:date id="DF502B0088E1EC56ECDAD8660F3B3F84">2022</mods:date>
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<mods:identifier id="BDCE361806E607B5C3567DD9CBEE70C8" type="DOI">10.1093/zoolinnean/zlab052</mods:identifier>
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<treatment id="03EF4E777C1ACA74FC12FC69242AFA3A" ID-DOI="http://doi.org/10.5281/zenodo.6536308" ID-Zenodo-Dep="6536308" LSID="urn:lsid:plazi:treatment:03EF4E777C1ACA74FC12FC69242AFA3A" httpUri="http://treatment.plazi.org/id/03EF4E777C1ACA74FC12FC69242AFA3A" lastPageId="14" lastPageNumber="234" pageId="13" pageNumber="233">
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<subSubSection id="C35CACEA7C1ACA77FC12FC692571FC52" box="[930,1303,989,1014]" pageId="13" pageNumber="233" type="nomenclature">
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<paragraph id="8BF9FF617C1ACA77FC12FC692571FC52" blockId="13.[930,1303,989,1014]" box="[930,1303,989,1014]" pageId="13" pageNumber="233">
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<heading id="D0B1480D7C1ACA77FC12FC692571FC52" box="[930,1303,989,1014]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="233" reason="2">
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<taxonomicName id="4C4684E27C1ACA77FC12FC6924C7FC52" ID-CoL="CDL4R" ID-ENA="1545017" authority="GREER, 1974" authorityName="Greer" authorityYear="1974" box="[930,1185,989,1014]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
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<emphasis id="B93223737C1ACA77FC12FC692460FC50" box="[930,1030,989,1012]" italics="true" pageId="13" pageNumber="233">LOBULIA</emphasis>
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<bibRefCitation id="EFD782907C1ACA77FBBDFC6924C7FC52" author="Greer AE" box="[1037,1185,989,1014]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">GREER, 1974</bibRefCitation>
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</taxonomicName>
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(CLADE I)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CACEA7C1ACA77FC83FBB724F6FB9F" pageId="13" pageNumber="233" type="description">
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<paragraph id="8BF9FF617C1ACA77FC83FBB724F6FB9F" blockId="13.[819,1414,1027,1083]" pageId="13" pageNumber="233">
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(
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<figureCitation id="137DE3E47C1ACA77FC8BFBB723D7FBB8" box="[827,945,1027,1052]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStart-10="Figure 15" captionStart-2="Figure 7" captionStart-3="Figure 8" captionStart-4="Figure 9" captionStart-5="Figure 10" captionStart-6="Figure 11" captionStart-7="Figure 12" captionStart-8="Figure 13" captionStart-9="Figure 14" captionStartId-0="14.[163,243,1109,1131]" captionStartId-1="15.[145,223,1530,1552]" captionStartId-10="24.[163,242,1447,1469]" captionStartId-2="16.[163,241,1520,1542]" captionStartId-3="17.[145,223,926,948]" captionStartId-4="18.[163,242,1594,1616]" captionStartId-5="19.[146,226,580,602]" captionStartId-6="20.[163,243,1575,1597]" captionStartId-7="21.[147,226,734,756]" captionStartId-8="22.[163,242,1550,1572]" captionStartId-9="23.[145,225,545,567]" captionTargetBox-0="[163,1443,196,1069]" captionTargetBox-1="[145,1425,195,1491]" captionTargetBox-10="[419,1187,195,1407]" captionTargetBox-2="[419,1184,195,1480]" captionTargetBox-3="[154,761,201,871]" captionTargetBox-4="[419,1187,195,1554]" captionTargetBox-5="[146,760,195,540]" captionTargetBox-6="[423,1182,196,1532]" captionTargetBox-7="[145,769,195,695]" captionTargetBox-8="[420,1187,196,1511]" captionTargetBox-9="[145,769,201,506]" captionTargetId-0="figure-329@14.[163,1443,195,1069]" captionTargetId-1="figure-158@15.[145,1425,195,1491]" captionTargetId-10="figure-187@24.[419,1187,195,1407]" captionTargetId-2="figure-26@16.[419,1187,195,1480]" captionTargetId-3="figure-775@17.[145,769,195,886]" captionTargetId-4="figure-119@18.[419,1187,195,1554]" captionTargetId-5="figure-732@19.[145,760,195,541]" captionTargetId-6="figure-87@20.[419,1187,192,1536]" captionTargetId-7="figure-743@21.[145,769,195,695]" captionTargetId-8="figure-169@22.[419,1187,195,1511]" captionTargetId-9="figure-857@23.[145,769,195,506]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-10="24" captionTargetPageId-2="16" captionTargetPageId-3="17" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionTargetPageId-8="22" captionTargetPageId-9="23" captionText-0="Figure 5. Representatives of each of the seven revised genera, not to scale: (A) Lobulia huonensis (BPBM 40322); (B) Prasinohaema flavipes (BPBM 40369); (C) Nubeoscincus glacialis (BPBM 14712); (D) Papuascincus lineage VII (BPBM 44749); (E) Palaia pulchra (BPBM 38789); (F) Alpinoscincus alpinus (BPBM 44218); (G) Ornithuroscincus pterophilus (BPBM 45705). Images by Allen Allison." captionText-1="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images C–E by Allen Allison." captionText-10="Figure 15. Lobulia vogelkopensis holotype (BPBM 6917), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-2="Figure 7. Lobulia lobulus holotype (MCZ R-47067), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Museum of Comparative Zoology, Harvard University." captionText-3="Figure 8. Lobulia lobulus (BPBM 47837) in life, (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-4="Figure 9. Lobulia fortis holotype (BPBM 41149), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-5="Figure 10. Lobulia fortis holotype (BPBM 41149) in life, (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-6="Figure 11. Lobulia huonensis holotype (BPBM 40322), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-7="Figure 12. Lobulia huonensis holotype (BPBM 40322), in life, in (A) dorsal view and (B) ventral view. Images by Allen Allison." captionText-8="Figure 13. Lobulia marmorata holotype (BPBM 34150), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Alex Slavenko." captionText-9="Figure 14. Lobulia marmorata holotype (BPBM 34150), ventral view in life. Image by Fred Kraus." figureDoi-0="http://doi.org/10.5281/zenodo.6530705" figureDoi-1="http://doi.org/10.5281/zenodo.6530711" figureDoi-10="http://doi.org/10.5281/zenodo.6530733" figureDoi-2="http://doi.org/10.5281/zenodo.6530713" figureDoi-3="http://doi.org/10.5281/zenodo.6530715" figureDoi-4="http://doi.org/10.5281/zenodo.6530717" figureDoi-5="http://doi.org/10.5281/zenodo.6530719" figureDoi-6="http://doi.org/10.5281/zenodo.6530721" figureDoi-7="http://doi.org/10.5281/zenodo.6530725" figureDoi-8="http://doi.org/10.5281/zenodo.6530729" figureDoi-9="http://doi.org/10.5281/zenodo.6530731" httpUri-0="https://zenodo.org/record/6530705/files/figure.png" httpUri-1="https://zenodo.org/record/6530711/files/figure.png" httpUri-10="https://zenodo.org/record/6530733/files/figure.png" httpUri-2="https://zenodo.org/record/6530713/files/figure.png" httpUri-3="https://zenodo.org/record/6530715/files/figure.png" httpUri-4="https://zenodo.org/record/6530717/files/figure.png" httpUri-5="https://zenodo.org/record/6530719/files/figure.png" httpUri-6="https://zenodo.org/record/6530721/files/figure.png" httpUri-7="https://zenodo.org/record/6530725/files/figure.png" httpUri-8="https://zenodo.org/record/6530729/files/figure.png" httpUri-9="https://zenodo.org/record/6530731/files/figure.png" pageId="13" pageNumber="233">FIGS 5–15</figureCitation>
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; SUPPORTING INFORMATION,
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<figureCitation id="137DE3E47C1ACA77FAB0FBB72515FBBF" box="[1280,1395,1027,1052]" captionStart="Figure 6" captionStartId="15.[145,223,1530,1552]" captionTargetBox="[145,1425,195,1491]" captionTargetId="figure-158@15.[145,1425,195,1491]" captionTargetPageId="15" captionText="Figure 6. Pictures in life of (A) Lobulia brongersmai (BPBM 34733); (B) Lobulia elegans (BPBM 18692); (C) Lobulia fortis (paratype; BPBM 41130); (D) Lobulia huonensis (holotype; BPBM 40322); (E) Lobulia lobulus (BPBM 47837); (F) Lobulia marmorata (holotype; BPBM 34150). Images A, B, F by Fred Kraus, images C–E by Allen Allison." figureDoi="http://doi.org/10.5281/zenodo.6530711" httpUri="https://zenodo.org/record/6530711/files/figure.png" pageId="13" pageNumber="233">FIGS S6–S</figureCitation>
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<figureCitation id="137DE3E47C1ACA77FAC2FBB725E6FBB8" box="[1394,1408,1027,1052]" captionStart="Figure 7" captionStartId="16.[163,241,1520,1542]" captionTargetBox="[419,1184,195,1480]" captionTargetId="figure-26@16.[419,1187,195,1480]" captionTargetPageId="16" captionText="Figure 7. Lobulia lobulus holotype (MCZ R-47067), in (A) dorsal view, (B) ventral view, (C) lateral view of head, (D) dorsal view of head and (E) ventral view of head. Images by Museum of Comparative Zoology, Harvard University." figureDoi="http://doi.org/10.5281/zenodo.6530713" httpUri="https://zenodo.org/record/6530713/files/figure.png" pageId="13" pageNumber="233">7</figureCitation>
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;
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<tableCitation id="C6C4CADA7C1ACA77FB99FB9624EEFB9F" box="[1065,1160,1058,1083]" captionStart="Table 1" captionStartId="9.[142,206,414,436]" captionTargetPageId="9" captionText="Table 1. Ranges of morphometrics and meristics for specimens of species treated in this study (mean values in parentheses). Only adult specimens were examined for morphometric measurements, but juveniles were included for meristics" httpUri="http://table.plazi.org/id/DF39AFE97C1ECA73FF3EFE2A2460FE75" pageId="13" pageNumber="233" tableUuid="DF39AFE97C1ECA73FF3EFE2A2460FE75">TABLE 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CACEA7C1ACA77FC99FBFE24F9FBDB" pageId="13" pageNumber="233" type="reference_group">
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<paragraph id="8BF9FF617C1ACA77FC99FBFE24F9FBDB" blockId="13.[809,1424,1098,1151]" pageId="13" pageNumber="233">
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<taxonomicName id="4C4684E27C1ACA77FC99FBFE247EFBC4" authority="Greer, 1974" authorityName="Greer" authorityYear="1974" box="[809,1048,1098,1120]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
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<emphasis id="B93223737C1ACA77FC99FBFE23E3FBFB" box="[809,901,1098,1119]" italics="true" pageId="13" pageNumber="233">Lobulia</emphasis>
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<bibRefCitation id="EFD782907C1ACA77FC3EFBFF247EFBC4" author="Greer AE" box="[910,1048,1099,1120]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">Greer, 1974</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="B93223737C1ACA77FB98FBFE2447FBDB" italics="true" pageId="13" pageNumber="233">Australian Journal of Zoology Supplementary Series</emphasis>
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(31): 1–67.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C35CACEA7C1ACA77FC99FB272472FB63" pageId="13" pageNumber="233" type="type_taxon">
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<paragraph id="8BF9FF617C1ACA77FC99FB272472FB63" blockId="13.[809,1425,1170,1223]" pageId="13" pageNumber="233">
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<emphasis id="B93223737C1ACA77FC99FB2723D9FB0C" box="[809,959,1171,1192]" italics="true" pageId="13" pageNumber="233">
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<typeStatus id="54FD41C37C1ACA77FC99FB272306FB0C" box="[809,864,1171,1192]" pageId="13" pageNumber="233">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C4684E27C1ACA77FC7DFB272500FB0C" authority="Boulenger, 1897" authorityName="Boulenger" authorityYear="1897" box="[973,1382,1170,1192]" class="Reptilia" family="Scincidae" genus="Lygosoma" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="species" species="elegans">
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<emphasis id="B93223737C1ACA77FC7DFB2724C6FB03" box="[973,1184,1170,1192]" italics="true" pageId="13" pageNumber="233">Lygosoma elegans</emphasis>
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<bibRefCitation id="EFD782907C1ACA77FB18FB262500FB0C" author="Boulenger GA" box="[1192,1382,1170,1192]" pageId="13" pageNumber="233" pagination="6 - 13" refId="ref43763" refString="Boulenger GA. 1897. Descriptions of new lizards and frogs from Mount Victoria, Owen Stanley Range, New Guinea, collected by Mr. A. S. Anthony. Annals and Magazine of Natural History Series 6 19: 6 - 13." type="journal article" year="1897">Boulenger, 1897</bibRefCitation>
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</taxonomicName>
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, by original designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CACEA7C1ACA77FC99FB4424C9F8CF" pageId="13" pageNumber="233" type="diagnosis">
|
||
<paragraph id="8BF9FF617C1ACA77FC99FB4423B4F99C" blockId="13.[809,1425,1264,1899]" pageId="13" pageNumber="233">
|
||
<emphasis id="B93223737C1ACA77FC99FB4423C2FAA1" box="[809,932,1264,1285]" italics="true" pageId="13" pageNumber="233">Diagnosis:</emphasis>
|
||
Medium-sized (adult SVL
|
||
<quantity id="4CBE52847C1ACA77FB54FB4425EFFAA1" box="[1252,1417,1264,1286]" metricMagnitude="-2" metricUnit="m" metricValue="5.29" metricValueMax="6.39" metricValueMin="4.1899999999999995" pageId="13" pageNumber="233" unit="mm" value="52.9" valueMax="63.9" valueMin="41.9">41.9–63.9 mm</quantity>
|
||
) terrestrial or semi-arboreal skinks with long limbs (forelimbs 32.9–47.6% of SVL, hindlimbs 41.6– 54.8% of SVL); lobules present on anterior edge of ear opening; two pairs of chin shields in medial contact; two supralabials posterior to subocular supralabial; chin shields abutting infralabials; lower eyelid with semi-transparent window; standard three-scale temporal region; nasal scale undivided; frontoparietals either fused or unfused; viviparous; litter size 1–4.
|
||
</paragraph>
|
||
<paragraph id="8BF9FF617C1ACA77FCF1F9F524C9F8CF" blockId="13.[809,1425,1264,1899]" pageId="13" pageNumber="233">
|
||
<taxonomicName id="4C4684E27C1ACA77FCF1F9F523FBF9F2" authorityName="Greer" authorityYear="1974" box="[833,925,1601,1622]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C1ACA77FCF1F9F523FBF9F2" box="[833,925,1601,1622]" italics="true" pageId="13" pageNumber="233">Lobulia</emphasis>
|
||
</taxonomicName>
|
||
differs from all other genera by its much longer limbs (forelimbs 32.9–47.6% vs. 27.7–39.8% of SVL, hindlimbs 41.6–54.8% vs. 29.9–49.6% of SVL). It further differs from
|
||
<taxonomicName id="4C4684E27C1ACA77FBAAF92924A2F916" authorityName="GREER" authorityYear="1974" box="[1050,1220,1693,1714]" class="Reptilia" family="Scincidae" genus="Prasinohaema" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C1ACA77FBAAF92924A2F916" box="[1050,1220,1693,1714]" italics="true" pageId="13" pageNumber="233">Prasinohaema</emphasis>
|
||
</taxonomicName>
|
||
by lacking green blood serum and tissues (
|
||
<bibRefCitation id="EFD782907C1ACA77FBF5F90824A1F975" author="Greer AE" box="[1093,1223,1724,1745]" pageId="13" pageNumber="233" pagination="1 - 67" refId="ref44557" refString="Greer AE. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology Supplementary Series 31: 1 - 67." type="book chapter" year="1974">Greer, 1974</bibRefCitation>
|
||
), a prehensile tail with a glandular tip and basally expanded subdigital lamellae. It differs from
|
||
<taxonomicName id="4C4684E27C1ACA77FBF7F94E2483F8AB" box="[1095,1253,1786,1807]" class="Reptilia" family="Scincidae" genus="Papuascincus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="233" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C1ACA77FBF7F94E2483F8AB" box="[1095,1253,1786,1807]" italics="true" pageId="13" pageNumber="233">Papuascincus</emphasis>
|
||
</taxonomicName>
|
||
by having two pairs of chin shields in medial contact (vs. one), an undivided (vs. divided) nasal scale and a viviparous (vs. oviparous) reproductive mode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF39AFE97C19CA74FF13FBE12221FB67" ID-DOI="http://doi.org/10.5281/zenodo.6530705" ID-Zenodo-Dep="6530705" httpUri="https://zenodo.org/record/6530705/files/figure.png" pageId="14" pageNumber="234" startId="14.[163,243,1109,1131]" targetBox="[163,1443,196,1069]" targetPageId="14">
|
||
<paragraph id="8BF9FF617C19CA74FF13FBE12221FB67" blockId="14.[163,1443,1109,1219]" pageId="14" pageNumber="234">
|
||
<emphasis id="B93223737C19CA74FF13FBE12168FBCE" bold="true" box="[163,270,1109,1131]" pageId="14" pageNumber="234">Figure 5.</emphasis>
|
||
Representatives of each of the seven revised genera, not to scale: (A)
|
||
<taxonomicName id="4C4684E27C19CA74FBB9FBE124A9FBCF" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[1033,1231,1109,1131]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="huonensis">
|
||
<emphasis id="B93223737C19CA74FBB9FBE124A9FBCF" box="[1033,1231,1109,1131]" italics="true" pageId="14" pageNumber="234">Lobulia huonensis</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 40322); (B)
|
||
<taxonomicName id="4C4684E27C19CA74FF13FBC621F1FB2C" baseAuthorityName="Parker" baseAuthorityYear="1936" box="[163,407,1138,1160]" class="Reptilia" family="Scincidae" genus="Prasinohaema" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="flavipes">
|
||
<emphasis id="B93223737C19CA74FF13FBC621F1FB2C" box="[163,407,1138,1160]" italics="true" pageId="14" pageNumber="234">Prasinohaema flavipes</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 40369); (C)
|
||
<taxonomicName id="4C4684E27C19CA74FDC0FBC62302FB2C" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" baseAuthorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" baseAuthorityYear="2005" box="[624,868,1138,1160]" class="Squamata" family="Scincidae" genus="Nubeoscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="glacialis">
|
||
<emphasis id="B93223737C19CA74FDC0FBC62302FB2C" box="[624,868,1138,1160]" italics="true" pageId="14" pageNumber="234">Nubeoscincus glacialis</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 14712); (D)
|
||
<taxonomicName id="4C4684E27C19CA74FB8EFBC624A9FB2C" box="[1086,1231,1138,1160]" class="Reptilia" family="Scincidae" genus="Papuascincus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C19CA74FB8EFBC624A9FB2C" box="[1086,1231,1138,1160]" italics="true" pageId="14" pageNumber="234">Papuascincus</emphasis>
|
||
</taxonomicName>
|
||
lineage VII (BPBM 44749); (E)
|
||
<taxonomicName id="4C4684E27C19CA74FE90FB2421A7FB02" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" baseAuthorityName="Boulenger" baseAuthorityYear="1903" box="[288,449,1168,1190]" class="Squamata" family="Scincidae" genus="Palaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis id="B93223737C19CA74FE90FB2421A7FB02" box="[288,449,1168,1190]" italics="true" pageId="14" pageNumber="234">Palaia pulchra</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 38789); (F)
|
||
<taxonomicName id="4C4684E27C19CA74FD2BFB2423EFFB02" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" baseAuthorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" baseAuthorityYear="2005" box="[667,905,1168,1190]" class="Squamata" family="Scincidae" genus="Alpinoscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="alpinus">
|
||
<emphasis id="B93223737C19CA74FD2BFB2423EFFB02" box="[667,905,1168,1190]" italics="true" pageId="14" pageNumber="234">Alpinoscincus alpinus</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 44218); (G)
|
||
<taxonomicName id="4C4684E27C19CA74FBD6FB2425C5FB02" authorityName="Slavenko & Tamar & Tallowin & Kraus & Allison & Carranza & Meiri" authorityYear="2022" box="[1126,1443,1168,1190]" class="Squamata" family="Scincidae" genus="Ornithuroscincus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Reptilia" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="pterophilus">
|
||
<emphasis id="B93223737C19CA74FBD6FB2425C5FB02" box="[1126,1443,1168,1190]" italics="true" pageId="14" pageNumber="234">Ornithuroscincus pterophilus</emphasis>
|
||
</taxonomicName>
|
||
(BPBM 45705). Images by Allen Allison.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35CACEA7C19CA74FF13FB5B229DFAE6" pageId="14" pageNumber="234" type="discussion">
|
||
<paragraph id="8BF9FF617C19CA74FF13FB5B229DFAE6" blockId="14.[163,780,1263,1346]" pageId="14" pageNumber="234">
|
||
<emphasis id="B93223737C19CA74FF13FB5B210EFAA0" box="[163,360,1263,1284]" italics="true" pageId="14" pageNumber="234">Species included:</emphasis>
|
||
<taxonomicName id="4C4684E27C19CA74FEC6FB5B2363FAA0" authority="Zweifel, 1972" authorityName="Zweifel" authorityYear="1972" box="[374,773,1263,1285]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
|
||
<emphasis id="B93223737C19CA74FEC6FB5B2200FAA0" box="[374,614,1263,1284]" italics="true" pageId="14" pageNumber="234">Lobulia brongersmai</emphasis>
|
||
<bibRefCitation id="EFD782907C19CA74FDDCFB5B2363FAA0" author="Zweifel RG" box="[620,773,1263,1285]" pageId="14" pageNumber="234" pagination="530 - 539" refId="ref46955" refString="Zweifel RG. 1972. A new scincid lizard of the genus Leiolopisma from New Guinea. Zoologische Mededelingen 47: 530 - 539." type="journal article" year="1972">Zweifel, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C4684E27C19CA74FF13FAB92259FA87" authority="(Boulenger, 1897)" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[163,575,1293,1315]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B93223737C19CA74FF13FAB92107FA86" box="[163,353,1293,1314]" italics="true" pageId="14" pageNumber="234">Lobulia elegans</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD782907C19CA74FEC3FAB92252FA86" author="Boulenger GA" box="[371,564,1293,1315]" pageId="14" pageNumber="234" pagination="6 - 13" refId="ref43763" refString="Boulenger GA. 1897. Descriptions of new lizards and frogs from Mount Victoria, Owen Stanley Range, New Guinea, collected by Mr. A. S. Anthony. Annals and Magazine of Natural History Series 6 19: 6 - 13." type="journal article" year="1897">Boulenger, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C4684E27C19CA74FDFDFAB9210BFAE5" authority="(Loveridge, 1945)" baseAuthorityName="Loveridge" baseAuthorityYear="1945" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="lobulus">
|
||
<emphasis id="B93223737C19CA74FDFDFAB9236DFA86" box="[589,779,1293,1314]" italics="true" pageId="14" pageNumber="234">Lobulia lobulus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD782907C19CA74FF1BFA982102FAE6" author="Loveridge A" box="[171,356,1324,1346]" pageId="14" pageNumber="234" pagination="47 - 52" refId="ref45207" refString="Loveridge A. 1945. New scincid lizards of the genera Tropidophorus and Lygosoma from New Guinea. Proceedings of the Biological Society of Washington 58: 47 - 52." type="journal article" year="1945">Loveridge, 1945</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; and new species described below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CACEA7C19CA74FF13FADF21F8F975" pageId="14" pageNumber="234" type="distribution">
|
||
<paragraph id="8BF9FF617C19CA74FF13FADF21F8F975" blockId="14.[163,781,1387,1745]" pageId="14" pageNumber="234">
|
||
<emphasis id="B93223737C19CA74FF13FADF2126FA24" box="[163,320,1387,1408]" italics="true" pageId="14" pageNumber="234">Distribution:</emphasis>
|
||
Members of
|
||
<taxonomicName id="4C4684E27C19CA74FE57FADF2223FA24" authorityName="Greer" authorityYear="1974" box="[487,581,1387,1408]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93223737C19CA74FE57FADF2223FA24" box="[487,581,1387,1408]" italics="true" pageId="14" pageNumber="234">Lobulia</emphasis>
|
||
</taxonomicName>
|
||
are widespread along most of the montane regions of New
|
||
<collectingCountry id="F351BFF17C19CA74FD1DFA3D2363FA3B" box="[685,773,1417,1439]" name="Guinea-Bissau" pageId="14" pageNumber="234">Guinea</collectingCountry>
|
||
, ranging from the central Owen Stanley Mountains in the Papuan Peninsula in the east (
|
||
<taxonomicName id="4C4684E27C19CA74FDD2FA732285FA78" baseAuthorityName="Boulenger" baseAuthorityYear="1897" box="[610,739,1479,1500]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B93223737C19CA74FDD2FA732285FA78" box="[610,739,1479,1500]" italics="true" pageId="14" pageNumber="234">Lo. elegans</emphasis>
|
||
</taxonomicName>
|
||
) to the Arfak Mountains in the Vogelkop Peninsula in the west. Most species are montane, found at elevations up to
|
||
<quantity id="4CBE52847C19CA74FF70F9972170F99C" box="[192,278,1571,1592]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="234" unit="m" value="2700.0">2700 m</quantity>
|
||
a.s.l.; however,
|
||
<taxonomicName id="4C4684E27C19CA74FE7DF99722E1F99C" authorityName="Zweifel" authorityYear="1972" box="[461,647,1571,1592]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
|
||
<emphasis id="B93223737C19CA74FE7DF99722E1F99C" box="[461,647,1571,1592]" italics="true" pageId="14" pageNumber="234">Lo. brongersmai</emphasis>
|
||
</taxonomicName>
|
||
is found in the lowlands and hill regions of the northern versant of New
|
||
<collectingCountry id="F351BFF17C19CA74FF49F9D4212AF9D2" box="[249,332,1632,1654]" name="Guinea-Bissau" pageId="14" pageNumber="234">Guinea</collectingCountry>
|
||
(
|
||
<quantity id="4CBE52847C19CA74FEEBF9D421ADF9D1" box="[347,459,1632,1653]" metricMagnitude="2" metricUnit="m" metricValue="6.7" metricValueMax="13.4" metricValueMin="0.0" pageId="14" pageNumber="234" unit="m" value="670.0" valueMax="1340.0" valueMin="0.0">0–1340 m</quantity>
|
||
a.s.l.).
|
||
<taxonomicName id="4C4684E27C19CA74FDADF9D4236CF9D1" authorityName="Zweifel" authorityYear="1972" box="[541,778,1632,1653]" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="brongersmai">
|
||
<emphasis id="B93223737C19CA74FDADF9D4236CF9D1" box="[541,778,1632,1653]" italics="true" pageId="14" pageNumber="234">Lobulia brongersmai</emphasis>
|
||
</taxonomicName>
|
||
is the only member of the genus to be found below elevations of
|
||
<quantity id="4CBE52847C19CA74FE87F9292118F917" box="[311,382,1693,1715]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="234" unit="m" value="700.0">700 m</quantity>
|
||
, the lowest recorded locality for
|
||
<taxonomicName id="4C4684E27C19CA74FD59F92A21FFF975" authority="(Kraus, 2020)" baseAuthorityName="Kraus" baseAuthorityYear="2020" class="Reptilia" family="Scincidae" genus="Lobulia" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="234" phylum="Chordata" rank="species" species="elegans">
|
||
<emphasis id="B93223737C19CA74FD59F92A2091F975" italics="true" pageId="14" pageNumber="234">Lo. elegans</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD782907C19CA74FEB5F90821E9F975" author="Kraus F" box="[261,399,1724,1745]" pageId="14" pageNumber="234" pagination="201 - 214" refId="ref45028" refString="Kraus F. 2020. A new species of Lobulia (Squamata: Scincidae) from Papua New Guinea. Zootaxa 4779: 201 - 214." type="journal article" year="2020">Kraus, 2020</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CACEA7C19CA74FF13F94E242AFA3A" pageId="14" pageNumber="234" type="discussion">
|
||
<paragraph id="8BF9FF617C19CA74FF13F94E242AFA3A" blockId="14.[163,778,1786,1900]" lastBlockId="14.[827,1443,1262,1438]" pageId="14" pageNumber="234">
|
||
<emphasis id="B93223737C19CA74FF13F94E2177F8AB" box="[163,273,1786,1807]" italics="true" pageId="14" pageNumber="234">Remarks:</emphasis>
|
||
Molecular evidence suggests that at least two other species not already named or described herein occur in the genus: one in Woitape (BPBM 18689–90; WGS 84:
|
||
<geoCoordinate id="EE7299A67C19CA74FE23F8E22189F8C8" box="[403,495,1878,1900]" degrees="8.545" direction="south" orientation="latitude" pageId="14" pageNumber="234" precision="55" value="-8.545">8.545°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C19CA74FE4CF8E22210F8C8" box="[508,630,1878,1900]" degrees="147.251" direction="east" orientation="longitude" pageId="14" pageNumber="234" precision="55" value="147.251">147.251°E</geoCoordinate>
|
||
) and one on Mt Yakapi in the Muller Range (BPBM 34161; WGS 84:
|
||
<geoCoordinate id="EE7299A67C19CA74FCDAFAB923ACFA87" box="[874,970,1293,1315]" degrees="5.666" direction="south" orientation="latitude" pageId="14" pageNumber="234" precision="55" value="-5.666">5.666°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7299A67C19CA74FC6AFAB9243AFA87" box="[986,1116,1293,1315]" degrees="142.643" direction="east" orientation="longitude" pageId="14" pageNumber="234" precision="55" value="142.643">142.643°E</geoCoordinate>
|
||
). However, since both are only known from a few specimens each, and are not extremely morphologically distinct, we refrain from formally describing them until further material can be collected and examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |