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<document id="B7895D5142EC0ECC589FEE3DF070392D" ID-CLB-Dataset="20932" ID-DOI="10.11646/zootaxa.5092.5.5" ID-GBIF-Dataset="7167ab3c-aa29-4166-aafd-5619777f1b9c" ID-ISSN="1175-5326" ID-Zenodo-Dep="5896455" ID-ZooBank="19DB83B4-ED9B-4310-AECA-9C1323A50941" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643010864285" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Koo, Jun-Mo" docDate="2022" docId="03EF4C71950EFFBAFF0CFF1FFAFDF9A8" docLanguage="en" docName="zootaxa.5092.5.5.pdf" docOrigin="Zootaxa 5092 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora manalis Park &amp; Koo 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="584" masterDocId="FFD634099506FFB2FF9BFF88FFD2FFA2" masterDocTitle="Description of Spiniola hanaro Park, gen. and sp. nov., and three new species of Torodora Meyrick, 1894 (Lepidoptera, Lecithoceridae, Torodorinae) from the Afrotropical Region" masterLastPageNumber="586" masterPageNumber="576" pageNumber="584" updateTime="1699303981264" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="50CADF417462A128519106971CE2B122" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7093BE5C920FCBFD8BBC4B4A70D97C0C">
<mods:title id="3E7984E0675938439558FBA8F8800C7A">Description of Spiniola hanaro Park, gen. and sp. nov., and three new species of Torodora Meyrick, 1894 (Lepidoptera, Lecithoceridae, Torodorinae) from the Afrotropical Region</mods:title>
</mods:titleInfo>
<mods:name id="1384B4043038FB0EE254BFFBA234CC1E" type="personal">
<mods:role id="75AA6220A4F5842B81D39420B3C830B4">
<mods:roleTerm id="A77386089E9797E5B22EFD48D10990F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D1104BBAC24FFA526E297861AAD3604">Park, Kyu-Tek</mods:namePart>
<mods:nameIdentifier id="CE6E5FAE20F7A8511E08252AE118A499" type="ORCID">0000-0001-9933-4497</mods:nameIdentifier>
<mods:affiliation id="CBE893641CC91891B53CCF21A2BC411E">Bioresource and Environmental Center, Incheon National University, Incheon, 22012 Korea.</mods:affiliation>
<mods:nameIdentifier id="31C74F1ACE40846C480F32D791F54065" type="email">ktpark02@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B4AB2FABB90646E104742F55AA9C4E73" type="personal">
<mods:role id="C777110218CE356B4BB5208B73B973E5">
<mods:roleTerm id="E970DAB892CD148EE8E65D19E61A89AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81AA65B1477D6FEB9FEE57E994A6FA30">Koo, Jun-Mo</mods:namePart>
<mods:affiliation id="9AE439D8FEB83B711A85BC831FB6A63E">Insect Systematic Bioinformatics Laboratory, Department of Plant Medicine, Chungbuk National University, Cheongju, 28644 Korea. ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
</mods:name>
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<mods:titleInfo id="E59AC1BC070B32A24FA6CB77ED02C4BE">
<mods:title id="2BF2C0464A6A20A65E191B0390B9FCFA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6E9C5F3AEC90EDE0E03BD3656512EB9B">
<mods:date id="45C5916927CE26612FA6E3CA3DD25803">2022</mods:date>
<mods:detail id="B0BE7711F7039D625AC4F74218013C3F" type="pubDate">
<mods:number id="092F622EED15B99321679E3EBE255FE5">2022-01-24</mods:number>
</mods:detail>
<mods:detail id="7D55CB0F2CB017C38D06A38AF984FD7B" type="volume">
<mods:number id="4B2218CFC4D0E217F891CFDA0C0A4930">5092</mods:number>
</mods:detail>
<mods:detail id="AC253A87F6EA153135EACC8532DBFEFE" type="issue">
<mods:number id="A0930F5AF3CAEFF7617AAB153DB11573">5</mods:number>
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<mods:start id="F3F0230BECAF3EFCE8AEC97C194CC053">576</mods:start>
<mods:end id="51097C0F54C64BFB232B25B6B33C06A7">586</mods:end>
</mods:extent>
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<mods:classification id="AD3D8B7BF10738288CD3D6D746BC15D0">journal article</mods:classification>
<mods:identifier id="2955A8D89CE1EDEB4ECC491825740A17" type="CLB-Dataset">20932</mods:identifier>
<mods:identifier id="5DDC1699186DF636EA859A6787C5E6BE" type="DOI">10.11646/zootaxa.5092.5.5</mods:identifier>
<mods:identifier id="4C0F95D0372E963967768D3F20786E94" type="GBIF-Dataset">7167ab3c-aa29-4166-aafd-5619777f1b9c</mods:identifier>
<mods:identifier id="20EDA688D70A31FA43080B3A9FC09074" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="10F846209EE555CC0F3784408821BE57" type="Zenodo-Dep">5896455</mods:identifier>
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</mods:mods>
<treatment id="03EF4C71950EFFBAFF0CFF1FFAFDF9A8" ID-DOI="http://doi.org/10.5281/zenodo.5913879" ID-GBIF-Taxon="192696644" ID-Zenodo-Dep="5913879" LSID="urn:lsid:plazi:treatment:03EF4C71950EFFBAFF0CFF1FFAFDF9A8" httpUri="http://treatment.plazi.org/id/03EF4C71950EFFBAFF0CFF1FFAFDF9A8" lastPageNumber="584" pageId="8" pageNumber="584">
<subSubSection id="C35CAEEC950EFFBAFF0CFF1FFBD5FF74" pageId="8" pageNumber="584" type="nomenclature">
<paragraph id="8BF9FD67950EFFBAFF0CFF1FFDA9FF13" blockId="8.[151,1031,151,250]" box="[151,635,151,178]" pageId="8" pageNumber="584">
<heading id="D0B14A0B950EFFBAFF0CFF1FFDA9FF13" bold="true" box="[151,635,151,178]" fontSize="11" level="1" pageId="8" pageNumber="584" reason="1">
<emphasis id="B9322175950EFFBAFF0CFF1FFDA9FF13" bold="true" box="[151,635,151,178]" pageId="8" pageNumber="584">
<taxonomicName id="4C4686E4950EFFBAFF0CFF1FFDC2FF13" ID-CoL="9FPXS" authority="Park &amp; Koo" authorityName="Park &amp; Koo" authorityYear="2022" box="[151,528,151,178]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="manalis" status="sp. nov.">
<emphasis id="B9322175950EFFBAFF0CFF1FFEBCFF10" bold="true" box="[151,366,151,178]" italics="true" pageId="8" pageNumber="584">Torodora manalis</emphasis>
Park &amp; Koo
</taxonomicName>
,
<taxonomicNameLabel id="A2019C0E950EFFBAFD86FF1FFDA9FF13" box="[541,635,151,177]" pageId="8" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9FD67950EFFBAFF0CFF33FBD5FF74" blockId="8.[151,1031,151,250]" box="[151,1031,187,214]" pageId="8" pageNumber="584">
LSID:
<uri id="FFD7F165950EFFBAFF7FFF33FBD5FF74" box="[228,1031,187,214]" pageId="8" pageNumber="584">
urn:lsid:zoobank.org:act:
<uuid id="FFE0C7B2950EFFBAFE6CFF33FBD5FF74" box="[503,1031,187,214]" pageId="8" pageNumber="584">D29247A4-3C17-4717-BF1D-95EB36859980</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF0CFF57FEFFFF58" box="[151,301,223,250]" pageId="8" pageNumber="584" type="description">
<paragraph id="8BF9FD67950EFFBAFF0CFF57FEFFFF58" blockId="8.[151,1031,151,250]" box="[151,301,223,250]" pageId="8" pageNumber="584">
(
<figureCitation id="137DE1E2950EFFBAFF04FF57FEF7FF58" box="[159,293,223,250]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Figs. 5A–H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF0CFEAFFABAFEC7" pageId="8" pageNumber="584" type="materials_examined">
<paragraph id="8BF9FD67950EFFBAFF0CFEAFFABAFEC7" blockId="8.[151,1437,294,1546]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF0CFEAFFE96FEE3" bold="true" box="[151,324,295,321]" pageId="8" pageNumber="584">Type material.</emphasis>
<materialsCitation id="3B2EF73A950EFFBAFED1FEAFFABAFEC7" ID-GBIF-Occurrence="3458591306" collectingDate="2014-11-19" collectingDateMax="2014-11-24" collectingDateMin="2014-11-19" collectionCode="MfN" collectorName="W. Mey &amp; Wings" country="Uganda" location="National Park" municipality="Biol. Field Station" pageId="8" pageNumber="584" specimenCode="CBNU063" specimenCount="1" stateProvince="Kibale" typeStatus="holotype">
<typeStatus id="54FD43C5950EFFBAFED1FEAFFE64FEE3" box="[330,438,295,321]" pageId="8" pageNumber="584" type="holotype">Holotype</typeStatus>
. [
<collectingCountry id="F351BDF7950EFFBAFE5DFEAEFDF1FEE3" box="[454,547,294,321]" name="Uganda" pageId="8" pageNumber="584">Uganda</collectingCountry>
] male,
<collectingMunicipality id="6B9D671D950EFFBAFDEFFEAFFC91FEE3" box="[628,835,295,321]" pageId="8" pageNumber="584">Biol. Field Station</collectingMunicipality>
,
<collectingRegion id="49823385950EFFBAFCD5FEAFFC4BFEE3" box="[846,921,295,321]" country="Uganda" name="Kibale" pageId="8" pageNumber="584">Kibale</collectingRegion>
<location id="8E99ABBC950EFFBAFC05FEAFFBE9FEE3" LSID="urn:lsid:plazi:treatment:03EF4C71950EFFBAFF0CFF1FFAFDF9A8:8E99ABBC950EFFBAFC05FEAFFBE9FEE3" box="[926,1083,295,321]" country="Uganda" municipality="Biol. Field Station" name="National Park" pageId="8" pageNumber="584" stateProvince="Kibale">National Park</location>
,
<collectingCountry id="F351BDF7950EFFBAFBDDFEAFFB4FFEE3" box="[1094,1181,295,321]" name="Uganda" pageId="8" pageNumber="584">Uganda</collectingCountry>
,
<date id="FFF8DBA7950EFFBAFB31FEAFFAB5FEE3" box="[1194,1383,295,322]" pageId="8" pageNumber="584" value="2014-11-19" valueMax="2014-11-24" valueMin="2014-11-19">
<collectingDate id="EFBC224F950EFFBAFB31FEAFFAB5FEE3" box="[1194,1383,295,322]" pageId="8" pageNumber="584" value="2014-11-19" valueMax="2014-11-24" valueMin="2014-11-19">19.–24. XI. 2014</collectingDate>
</date>
(LF, leg.
<collectorName id="26B398B1950EFFBAFF5CFEC4FEF6FEC4" box="[199,292,332,358]" pageId="8" pageNumber="584">W. Mey</collectorName>
), gen. slide no. CIS-7039,
<collectorName id="26B398B1950EFFBAFDD5FEC3FD44FEC7" box="[590,662,331,357]" pageId="8" pageNumber="584">Wings</collectorName>
slide no. CIS-7053,
<emphasis id="B9322175950EFFBAFCE5FEC4FC7CFEC4" box="[894,942,332,358]" italics="true" pageId="8" pageNumber="584">COI</emphasis>
barcode
<specimenCode id="DBE0551C950EFFBAFB8EFEC3FB5DFEC7" box="[1045,1167,331,357]" collectionCode="CBNU" pageId="8" pageNumber="584">CBNU063</specimenCode>
, deposited in
<collectionCode id="ED5765A2950EFFBAFAB1FEC3FAB6FEC7" box="[1322,1380,331,357]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="8" pageNumber="584" type="Museum">MfN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF5CFEE7FCC6FD24" pageId="8" pageNumber="584" type="diagnosis">
<paragraph id="8BF9FD67950EFFBAFF5CFEE7FCC6FD24" blockId="8.[151,1437,294,1546]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF5CFEE7FE90FE2B" bold="true" box="[199,322,367,393]" pageId="8" pageNumber="584">Diagnosis.</emphasis>
This new species is superficially similar to
<taxonomicName id="4C4686E4950EFFBAFCC2FEE7FAC1FE2B" authority="(Meyrick, 1933)" baseAuthorityName="Meyrick" baseAuthorityYear="1933" box="[857,1299,367,393]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="semnodora">
<emphasis id="B9322175950EFFBAFCC2FEE7FB9BFE2B" box="[857,1097,367,393]" italics="true" pageId="8" pageNumber="584">Torodora semnodora</emphasis>
(
<bibRefCitation id="EFD78096950EFFBAFBC5FEE7FAD9FE2B" author="Meyrick, E." box="[1118,1291,367,393]" pageId="8" pageNumber="584" pagination="353 - 448" refId="ref5425" refString="Meyrick, E. (1933) Exotic Microlepidoptera, Vol. IV. Thornhanger, Marlborough, Wilts, pp. 353 - 448." type="book chapter" year="1933">Meyrick, 1933</bibRefCitation>
)
</taxonomicName>
which was described from
<collectingCountry id="F351BDF7950EFFBAFED2FE1CFE17FE0F" box="[329,453,403,430]" name="Democratic Republic of the Congo" pageId="8" pageNumber="584">DR Congo</collectingCountry>
, but it can be distinguished by the wing pattern: forewing lacking a costal patch in the new species, but with an orange-yellow costal patch beyond 2/
<quantity id="4CBE5082950EFFBAFCCDFE3FFC52FE73" box="[854,896,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="584" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C4686E4950EFFBAFC1DFE30FBCCFE73" baseAuthorityName="Meyrick" baseAuthorityYear="1933" box="[902,1054,439,465]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="semnodora">
<emphasis id="B9322175950EFFBAFC1DFE30FBCCFE73" box="[902,1054,439,465]" italics="true" pageId="8" pageNumber="584">T. semnodora</emphasis>
</taxonomicName>
. The male genitalia also resemble each other, but they differ in the following characteristics: 1) cucullus narrowed with sharply pointed apex in the new species, but cucullus rounded apically in
<taxonomicName id="4C4686E4950EFFBAFD3AFD88FCE8FDBB" baseAuthorityName="Meyrick" baseAuthorityYear="1933" box="[673,826,511,537]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="semnodora">
<emphasis id="B9322175950EFFBAFD3AFD88FCE8FDBB" box="[673,826,511,537]" italics="true" pageId="8" pageNumber="584">T. semnodora</emphasis>
</taxonomicName>
; 2) juxta narrowed in basal half, with slightly convex caudal margin medially in the new species, but juxta with almost constant width, and a nearly straight caudal margin in
<taxonomicName id="4C4686E4950EFFBAFF28FDC0FE98FDC3" baseAuthorityName="Meyrick" baseAuthorityYear="1933" box="[179,330,583,609]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="semnodora">
<emphasis id="B9322175950EFFBAFF28FDC0FE98FDC3" box="[179,330,583,609]" italics="true" pageId="8" pageNumber="584">T. semnodora</emphasis>
</taxonomicName>
; 3) aedeagus roundly produced dorsally and triangularly expanded ventrally in distal 1/
<quantity id="4CBE5082950EFFBAFA8EFDCFFAEDFDC3" box="[1301,1343,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="584" unit="in" value="3.0">3 in</quantity>
the new species, but aedeagus bifurcate apically in
<taxonomicName id="4C4686E4950EFFBAFDEEFDE4FCDFFD27" baseAuthorityName="Meyrick" baseAuthorityYear="1933" box="[629,781,619,645]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="semnodora">
<emphasis id="B9322175950EFFBAFDEEFDE4FCDFFD27" box="[629,781,619,645]" italics="true" pageId="8" pageNumber="584">T. semnodora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF5CFD07FE43FA63" pageId="8" pageNumber="584" type="description">
<paragraph id="8BF9FD67950EFFBAFF5CFD07FB30FBFB" blockId="8.[151,1437,294,1546]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF5CFD07FE8BFD0B" bold="true" box="[199,345,655,681]" pageId="8" pageNumber="584">Description.</emphasis>
Adult (
<figureCitation id="137DE1E2950EFFBAFE2AFD07FD8FFD08" box="[433,605,655,682]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Figs. 5A, C, D</figureCitation>
). Wingspan 20.0 mm.
<emphasis id="B9322175950EFFBAFCC5FD07FC49FD0B" box="[862,923,655,681]" italics="true" pageId="8" pageNumber="584">Head</emphasis>
: Vertex yellowish brown with erect scales of same color laterally. Antenna 0.7 times shorter than forewing; scape elongated, yellowish white with dark brown scales on outer surface; flagellum filiform, ciliate, yellowish white without annulations. 2
<superScript id="7C33502F950EFFBAFBECFD5FFB55FD47" attach="left" box="[1143,1159,727,741]" fontSize="6" pageId="8" pageNumber="584">nd</superScript>
segment of labial palpus yellowish brown with yellowish-white scales on outer surface, yellowish white on inner surface; 3
<superScript id="7C33502F950EFFBAFB45FD73FB3EFCAB" attach="left" box="[1246,1260,763,777]" fontSize="6" pageId="8" pageNumber="584">rd</superScript>
segment nearly same length as 2
<superScript id="7C33502F950EFFBAFECCFC97FEB5FC8F" attach="left" box="[343,359,799,813]" fontSize="6" pageId="8" pageNumber="584">nd</superScript>
segment, yellowish white with dark brown scales dorsally (
<figureCitation id="137DE1E2950EFFBAFB81FC97FB76FC98" box="[1050,1188,799,826]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Figs. 5C, D</figureCitation>
).
<emphasis id="B9322175950EFFBAFB21FC97FAC6FC9B" box="[1210,1300,799,825]" italics="true" pageId="8" pageNumber="584">Thorax:</emphasis>
Notum and tegula yellowish brown. Forewing ground color yellowish brown, without markings; apex obtuse; termen slightly concave; fringe concolorous with ground color; venation (
<figureCitation id="137DE1E2950EFFBAFCABFCEFFC54FC20" box="[816,902,871,898]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Fig. 5E</figureCitation>
) with R
<subScript id="17C2FF22950EFFBAFC7EFCF2FC3FFC2A" attach="left" box="[997,1005,890,904]" fontSize="6" pageId="8" pageNumber="584">1</subScript>
arising from basal half of discal cell; distance between origins of R
<subScript id="17C2FF22950EFFBAFE7AFC16FE3BFC0E" attach="left" box="[481,489,926,940]" fontSize="6" pageId="8" pageNumber="584">1</subScript>
and R
<subScript id="17C2FF22950EFFBAFDAAFC16FDEBFC0E" attach="left" box="[561,569,926,940]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
3.1 times longer than that of R
<subScript id="17C2FF22950EFFBAFC0AFC16FC4BFC0E" attach="left" box="[913,921,926,940]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
and R
<subScript id="17C2FF22950EFFBAFC7AFC15FBDEFC09" attach="left" box="[993,1036,925,939]" fontSize="6" pageId="8" pageNumber="584">3+4+5</subScript>
; R
<subScript id="17C2FF22950EFFBAFBB6FC16FBE7FC0E" attach="left" box="[1069,1077,926,940]" fontSize="6" pageId="8" pageNumber="584">3</subScript>
stalked with R
<subScript id="17C2FF22950EFFBAFB47FC15FB24FC09" attach="left" box="[1244,1270,925,939]" fontSize="6" pageId="8" pageNumber="584">4+5</subScript>
for basal 3/10; R
<subScript id="17C2FF22950EFFBAFF31FC4AFF60FC72" attach="left" box="[170,178,962,976]" fontSize="6" pageId="8" pageNumber="584">4</subScript>
stalked with R
<subScript id="17C2FF22950EFFBAFEC7FC49FEB6FC6D" attach="left" box="[348,356,961,975]" fontSize="6" pageId="8" pageNumber="584">5</subScript>
for basal 3/5; M
<subScript id="17C2FF22950EFFBAFDBBFC4AFDFAFC72" attach="left" box="[544,552,962,976]" fontSize="6" pageId="8" pageNumber="584">1</subScript>
remote from R
<subScript id="17C2FF22950EFFBAFD4FFC49FCD2FC6D" attach="left" box="[724,768,961,975]" fontSize="6" pageId="8" pageNumber="584">3+4+5</subScript>
at base; M
<subScript id="17C2FF22950EFFBAFCE7FC4AFC56FC72" attach="left" box="[892,900,962,976]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
free from M
<subScript id="17C2FF22950EFFBAFB8EFC4AFBCFFC72" attach="left" box="[1045,1053,962,976]" fontSize="6" pageId="8" pageNumber="584">3</subScript>
; CuA
<subScript id="17C2FF22950EFFBAFBFAFC4AFBBBFC72" attach="left" box="[1121,1129,962,976]" fontSize="6" pageId="8" pageNumber="584">1</subScript>
and CuA
<subScript id="17C2FF22950EFFBAFB4EFC4AFB0FFC72" attach="left" box="[1237,1245,962,976]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
stalked for basal 1/4; 1A+2A forked basally; discal cell closed. Hindwing ground color similar to that of forewing; apex obtuse; termen oblique; fringe concolorous with ground color; venation with M
<subScript id="17C2FF22950EFFBAFC5DFB82FC1CFBBA" attach="left" box="[966,974,1034,1048]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
present; M
<subScript id="17C2FF22950EFFBAFBCBFB82FB8AFBBA" attach="left" box="[1104,1112,1034,1048]" fontSize="6" pageId="8" pageNumber="584">2</subScript>
free from M
<subScript id="17C2FF22950EFFBAFB77FB82FB26FBBA" attach="left" box="[1260,1268,1034,1048]" fontSize="6" pageId="8" pageNumber="584">3</subScript>
; M
<subScript id="17C2FF22950EFFBAFA85FB82FAF4FBBA" attach="left" box="[1310,1318,1034,1048]" fontSize="6" pageId="8" pageNumber="584">3</subScript>
and CuA
<subScript id="17C2FF22950EFFBAFA0FFB82FA4EFBBA" attach="left" box="[1428,1436,1034,1048]" fontSize="6" pageId="8" pageNumber="584">1</subScript>
short-stalked at base; discal cell weakly closed.
<emphasis id="B9322175950EFFBAFD23FB93FCF0FB97" box="[696,802,1051,1077]" italics="true" pageId="8" pageNumber="584">Abdomen</emphasis>
(
<figureCitation id="137DE1E2950EFFBAFCA9FB93FC5FFB94" box="[818,909,1051,1078]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Fig. 5H</figureCitation>
): Spinose zones developed widely in posterior margin of each II–VII tergite. Sternites weakly sclerotized; VIII sternite not specifically modified.
</paragraph>
<paragraph id="8BF9FD67950EFFBAFF5CFBEBFD66FA3F" blockId="8.[151,1437,294,1546]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF5CFBEBFEBCFBDF" box="[199,366,1123,1149]" italics="true" pageId="8" pageNumber="584">Male genitalia</emphasis>
(
<figureCitation id="137DE1E2950EFFBAFEE4FBEBFDD1FBDF" box="[383,515,1123,1150]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[186,1403,184,1915]" captionTargetId="figure-34@9.[184,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Torodora manalis Park &amp; Koo, sp. nov., holotype: A, adult; B, label; C, outer surface of labial palpus; D, inner surface of labial palpus; E, venation, wings slide no. CIS-7053; F, male genitalia, gen. slide no. CIS-7039, with close-up of uncus+gnathos; G, aedeagus; H, abdomen." figureDoi="http://doi.org/10.5281/zenodo.5896467" httpUri="https://zenodo.org/record/5896467/files/figure.png" pageId="8" pageNumber="584">Figs. 5F, G</figureCitation>
): Uncus elongated, with pointed apex, exceeding apex of basal plate of gnathos. Median process of gnathos relatively large, strongly bent beyond middle, narrowly produced apically. Tegumen deeply concave on anterior margin. Valva broad basally, strongly upturned beyond middle, with narrow concavity below costa in basal 1/5; costa slightly extended in basal 1/3; oblique crescent line extended from below end of concavity below costa to lower corner of cucullus; cucullus densely setose, narrowed apically with inwardly acute apex. Vinculum narrow, heavily sclerotized. Juxta large, shield-shaped, broadened in distal half; caudal margin slightly convex medially. Aedeagus nearly same length as valva, slender and weakly sclerotized in basal 2/3; distal 1/3 of aedeagus roundly produced dorsally with numerous pegs, triangularly expanded ventrally with relatively smaller numerous ventral pegs; cornutus absent.
</paragraph>
<paragraph id="8BF9FD67950EFFBAFF5CFA2FFE43FA63" blockId="8.[151,1437,294,1546]" box="[199,401,1447,1473]" pageId="8" pageNumber="584">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF5CFA43FD49FA44" box="[199,667,1483,1510]" pageId="8" pageNumber="584" type="distribution">
<paragraph id="8BF9FD67950EFFBAFF5CFA43FD49FA44" blockId="8.[151,1437,294,1546]" box="[199,667,1483,1510]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF5CFA43FEB3FA47" bold="true" box="[199,353,1483,1509]" pageId="8" pageNumber="584">Distribution.</emphasis>
<collectingCountry id="F351BDF7950EFFBAFEFCFA43FE6CFA47" box="[359,446,1483,1509]" name="Uganda" pageId="8" pageNumber="584">Uganda</collectingCountry>
(
<collectingRegion id="49823385950EFFBAFE56FA43FDCBFA47" box="[461,537,1483,1509]" country="Uganda" name="Kibale" pageId="8" pageNumber="584">Kibale</collectingRegion>
Nat. Park).
</paragraph>
</subSubSection>
<subSubSection id="C35CAEEC950EFFBAFF5CFA67FAFDF9A8" box="[199,1327,1519,1546]" pageId="8" pageNumber="584" type="etymology">
<paragraph id="8BF9FD67950EFFBAFF5CFA67FAFDF9A8" blockId="8.[151,1437,294,1546]" box="[199,1327,1519,1546]" pageId="8" pageNumber="584">
<emphasis id="B9322175950EFFBAFF5CFA67FE9CF9AB" bold="true" box="[199,334,1519,1545]" pageId="8" pageNumber="584">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="B9322175950EFFBAFCDCFA78FC56F9AB" box="[839,900,1520,1545]" italics="true" pageId="8" pageNumber="584">mane</emphasis>
(= morning) with a Latin suffix, -
<emphasis id="B9322175950EFFBAFA9BFA67FAFAF9AB" box="[1280,1320,1519,1545]" italics="true" pageId="8" pageNumber="584">alis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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