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<mods:title id="16714021EE0C38F447E431FA0B761592">Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): a sixth species of Pseudholophylla Blackburn, 1911, and notes on the other known species</mods:title>
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<mods:namePart id="0DA4DF15A157132937519C05A6A878C1">Hutchinson, Paul M.</mods:namePart>
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<paragraph id="8BF9A318FFBBBB53098DBC77FD420159" blockId="5.[151,730,1877,1940]" box="[151,730,1877,1904]" pageId="5" pageNumber="35">
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<taxonomicName id="4C46D89BFFBBBB53098DBC77FD420159" authority="(Burmeister, 1855)" baseAuthorityName="Burmeister" baseAuthorityYear="1855" box="[151,730,1877,1904]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFBBBB53098DBC77FD420159" bold="true" box="[151,730,1877,1904]" pageId="5" pageNumber="35">
<emphasis id="B9327F0AFFBBBB53098DBC77FE730146" bold="true" box="[151,491,1877,1904]" italics="true" pageId="5" pageNumber="35">Pseudholophylla furfuracea</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBBBB5308E0BC74FD4A0159" author="Burmeister, H. C. C." box="[506,722,1878,1904]" pageId="5" pageNumber="35" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855</bibRefCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C35CF093FFBBBB53098DBC58FF6201BD" box="[151,250,1914,1940]" pageId="5" pageNumber="35" type="description">
<paragraph id="8BF9A318FFBBBB53098DBC58FF6201BD" blockId="5.[151,730,1877,1940]" box="[151,250,1914,1940]" pageId="5" pageNumber="35">
(
<figureCitation id="137DBF9DFFBBBB5309BABC58FF6901BD" box="[160,241,1914,1940]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="5" pageNumber="35">Fig. 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBBBB50098DBC9CFE3706FC" lastPageId="6" lastPageNumber="36" pageId="5" pageNumber="35" type="reference_group">
<paragraph id="8BF9A318FFBBBB53098DBC9CFDF201FF" blockId="5.[151,1373,1982,2037]" box="[151,618,1982,2006]" pageId="5" pageNumber="35">
<treatmentCitationGroup id="AB568436FFBBBB53098DBC9CFDF201FF" box="[151,618,1982,2006]" pageId="5" pageNumber="35">
<treatmentCitation id="0AE78509FFBBBB53098DBC9CFDFE01FF" author="Burmeister, H. C. C." box="[151,614,1982,2006]" page="426" pageId="5" pageNumber="35" year="1855">
<taxonomicName id="4C46D89BFFBBBB53098DBC9CFDFE01FF" authority="Burmeister, 1855: 426" authorityName="Burmeister" authorityPageNumber="426" authorityYear="1855" box="[151,614,1982,2006]" class="Insecta" family="Melolonthidae" genus="Holophylla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFBBBB53098DBC9CFEE301FF" box="[151,379,1982,2006]" italics="true" pageId="5" pageNumber="35">Holophylla furfuracea</emphasis>
<bibRefCitation id="EFD7DEE9FFBBBB53089BBC9CFDFE01FF" author="Burmeister, H. C. C." box="[385,614,1982,2006]" pageId="5" pageNumber="35" refId="ref5821" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Coleoptera Lamellicornia Phyllophaga Chaenochela. Vol. 4. Part 2. Reimer, Berlin. [unknown pagination]" type="book" year="1855">Burmeister, 1855: 426</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BF9A318FFBBBB53098DBCFFFAC501DC" blockId="5.[151,1373,1982,2037]" box="[151,1373,2013,2037]" pageId="5" pageNumber="35">
<treatmentCitation id="0AE78509FFBBBB53098DBCFFFD0001DC" author="Blackburn, T." box="[151,664,2013,2037]" page="196" pageId="5" pageNumber="35" year="1911">
<taxonomicName id="4C46D89BFFBBBB53098DBCFFFD0001DC" authority=": Blackburn 1911: 196" authorityName="Blackburn" authorityPageNumber="196" authorityYear="1911" box="[151,664,2013,2037]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="furfuracea">
<emphasis id="B9327F0AFFBBBB53098DBCFFFE2B01DC" box="[151,435,2013,2037]" italics="true" pageId="5" pageNumber="35">Pseudholophylla furfuracea</emphasis>
:
<bibRefCitation id="EFD7DEE9FFBBBB5308DABCFFFD0001DC" author="Blackburn, T." box="[448,664,2013,2037]" pageId="5" pageNumber="35" pagination="173 - 203" refId="ref5686" refString="Blackburn, T. (1911) Further notes on Australian Coleoptera, with descriptions of new genera and species. No. XLI. Transactions of the Royal Society of South Australia, 35, 173 - 203." type="journal article" year="1911">Blackburn 1911: 196</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AE78509FFBBBB530BB8BCFFFCD401DC" author="Britton, E. B." box="[674,844,2013,2037]" page="35" pageId="5" pageNumber="35" year="1978">
<bibRefCitation id="EFD7DEE9FFBBBB530BB8BCFFFCD401DC" author="Britton, E. B." box="[674,844,2013,2037]" pageId="5" pageNumber="35" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton 1978: 35</bibRefCitation>
</treatmentCitation>
, Figs. 103105, 275A;
<treatmentCitation id="0AE78509FFBBBB530D23BCFFFA9C01DC" ID-DOI="http://doi.org/10.5281/zenodo.4340877" ID-Zenodo-Dep="4340877" author="Allsopp, P. G." box="[1081,1284,2013,2037]" httpUri="http://treatment.plazi.org/id/C264702F452135052BE799FEFD9DFDFD" page="478" pageId="5" pageNumber="35" year="2020">
<bibRefCitation id="EFD7DEE9FFBBBB530D23BCFFFA9C01DC" author="Allsopp, P. G." box="[1081,1284,2013,2037]" pageId="5" pageNumber="35" pagination="451 - 486" refId="ref5561" refString="Allsopp, P. G. (2020 b) Clarification of the status of the types of Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae) described before 1950. Zootaxa, 4885 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4885.4.1" type="journal article" year="2020">Allsopp 2020b: 478</bibRefCitation>
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, Fig. 37.
</paragraph>
<paragraph id="8BF9A318FFB8BB50098DBBB5FE3706FC" blockId="6.[151,1437,150,573]" pageId="6" pageNumber="36">
<emphasis id="B9327F0AFFB8BB50098DBBB5FEBD0699" bold="true" box="[151,293,150,177]" pageId="6" pageNumber="36">Type series.</emphasis>
<typeStatus id="54FD1DBAFFB8BB500835BBB5FE070698" box="[303,415,151,177]" pageId="6" pageNumber="36" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D406891FFB8BB5008B2BBB5FE270698" box="[424,447,151,177]" pageId="6" pageNumber="36" type="male"></specimenCount>
, designated by
<bibRefCitation id="EFD7DEE9FFB8BB500B69BBB5FCAA069B" author="Allsopp, P. G." box="[627,818,151,178]" pageId="6" pageNumber="36" pagination="451 - 486" refId="ref5561" refString="Allsopp, P. G. (2020 b) Clarification of the status of the types of Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae) described before 1950. Zootaxa, 4885 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4885.4.1" type="journal article" year="2020">Allsopp (2020b)</bibRefCitation>
.
<collectingCountry id="F351E388FFB8BB500A27BBB5FC470698" box="[829,991,151,177]" name="Australia" pageId="6" pageNumber="36">AUSTRALIA</collectingCountry>
. Holl. N. [New Holland, Australia]; in ZMH [image examined].
</paragraph>
</treatmentCitationGroup>
</subSubSection>
<subSubSection id="C35CF093FFB8BB5009DDBBFCFC4E0784" pageId="6" pageNumber="36" type="diagnosis">
<paragraph id="8BF9A318FFB8BB5009DDBBFCFC4E0784" blockId="6.[151,1437,150,573]" pageId="6" pageNumber="36">
<emphasis id="B9327F0AFFB8BB5009DDBBFCFE3106D0" bold="true" box="[199,425,222,249]" pageId="6" pageNumber="36">Diagnosis of males.</emphasis>
Body
<quantity id="4CBE0EFDFFB8BB5008EBBBFDFDA706D0" box="[497,575,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="36" unit="mm" value="16.0">16 mm</quantity>
long. Head, pronotum, elytra, pygidium, and legs dark brown, ventral surface brown. Lamella of antennomere 4 as long as lamella of antennomere 5, antennomere 3 broadly triangular. Maxillary palp with apical palpomere broad, with an elliptical concavity on the upper surface. Pronotum with greatest width 1.75x mid length; anterior margin and posterior margins well defined, posterior margin with a sparse fringe of erect setae. Parameres (
<figureCitation id="137DBF9DFFB8BB500879BA4DFE2D07A0" box="[355,437,367,393]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="6" pageNumber="36">Fig. 10</figureCitation>
) without a transverse apical process, dorsal margin sinuate convexly and ventral margin concave arcuate in lateral view, apical third evenly curved in ventral view.
</paragraph>
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<subSubSection id="C35CF093FFB8BB5009DDBA95FF780430" pageId="6" pageNumber="36" type="discussion">
<paragraph id="8BF9A318FFB8BB5009DDBA95FF780430" blockId="6.[151,1437,150,573]" pageId="6" pageNumber="36">
<emphasis id="B9327F0AFFB8BB5009DDBA95FEA307F8" bold="true" box="[199,315,439,465]" pageId="6" pageNumber="36">Remarks.</emphasis>
This name was used by sugarcane entomologists in the early to mid 1900s (
<emphasis id="B9327F0AFFB8BB500D92BA95FB2807F9" box="[1160,1200,439,464]" italics="true" pageId="6" pageNumber="36">e.g.</emphasis>
,
<bibRefCitation id="EFD7DEE9FFB8BB500DA4BA95FF4807DC" author="Illingworth, J. F. &amp; Dodd, A. P." pageId="6" pageNumber="36" pagination="1 - 104" refId="ref5957" refString="Illingworth, J. F. &amp; Dodd, A. P. (1921) Australian sugar-cane beetles and their allies. Bulletin, Division of Entomology, Bureau of Sugar Experiment Stations, Queensland, 16, 1 - 104." type="journal article" year="1921">Illingworth &amp; Dodd 1921</bibRefCitation>
) for the species later described as
<taxonomicName id="4C46D89BFFB8BB500B47BAF9FC4507DC" authority="Britton, 1978" authorityName="Britton" authorityYear="1978" box="[605,989,475,501]" class="Insecta" family="Melolonthidae" genus="Antitrogus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B9327F0AFFB8BB500B47BAF9FCA507DC" box="[605,829,475,501]" italics="true" pageId="6" pageNumber="36">Antitrogus parvulus</emphasis>
<bibRefCitation id="EFD7DEE9FFB8BB500A5CBAF9FC4507DC" author="Britton, E. B." box="[838,989,475,501]" pageId="6" pageNumber="36" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton, 1978</bibRefCitation>
</taxonomicName>
(Childers canegrub) (
<bibRefCitation id="EFD7DEE9FFB8BB500DCCBAF9FF4C0430" author="Miller, L. J. &amp; Allsopp, P. G." pageId="6" pageNumber="36" pagination="377 - 409" refId="ref6172" refString="Miller, L. J. &amp; Allsopp, P. G. (2000) Identification of Australian canegrubs (Coleoptera: Scarabaeidae: Melolonthini). Invertebrate Taxonomy, 14, 377 - 409. https: // doi. org / 10.1071 / IT 98019" type="journal article" year="2000">Miller &amp; Allsopp 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="8BF9A318FFB8BB5009DDB900FA960414" blockId="6.[151,1437,150,573]" box="[199,1294,546,573]" pageId="6" pageNumber="36">
<emphasis id="B9327F0AFFB8BB5009DDB900FEF90415" bold="true" box="[199,353,546,572]" pageId="6" pageNumber="36">Distribution.</emphasis>
Known only from the
<typeStatus id="54FD1DBAFFB8BB500B7AB901FD5D0414" box="[608,709,547,573]" pageId="6" pageNumber="36" type="holotype">holotype</typeStatus>
, without a precise locality (
<bibRefCitation id="EFD7DEE9FFB8BB500AE3B901FB3B0414" author="Allsopp, P. G." box="[1017,1187,547,573]" pageId="6" pageNumber="36" pagination="451 - 486" refId="ref5561" refString="Allsopp, P. G. (2020 b) Clarification of the status of the types of Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae) described before 1950. Zootaxa, 4885 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4885.4.1" type="journal article" year="2020">Allsopp 2020b</bibRefCitation>
, Fig. 37).
</paragraph>
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