368 lines
58 KiB
XML
368 lines
58 KiB
XML
<document id="AA1E89243E37E54150E492FF2B2CB61E" ID-CLB-Dataset="86005" ID-DOI="10.5252/geodiversitas2022v44a21" ID-GBIF-Dataset="b760c638-30c3-40d1-9c3f-d3cc43476b53" ID-ISSN="1638-9395" ID-Zenodo-Dep="6787056" ID-ZooBank="urn:lsid:zoobank.org:pub:1928AA5E-1D54-434A-829F-68DB923EBF1B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656341311969" checkinUser="felipe" docAuthor="Béthoux, Olivier & Grauvogel-Stamm, Léa" docDate="2022" docId="03EF0D12816DFFCBFF62FA405A6BB8F2" docLanguage="en" docName="Geodiversitas.44.21.603-608.pdf" docOrigin="Geodiversitas 44 (21)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Parabintoniella papierae Béthoux & Grauvogel-Stamm 2022, n. gen., n. sp." docType="treatment" docUuid="1AAFA563-F408-41C0-A519-B1E25EC3ABCA" docUuidSource="ZooBank" docVersion="8" lastPageNumber="607" masterDocId="FFD6756A8169FFCDFF90FFF55F51BD5F" masterDocTitle="Parabintoniella papierae n. gen., n. sp., a new Triassic representative of the Bintoniellidae, a poorly known extinct family of Orthoptera (Insecta)" masterLastPageNumber="608" masterPageNumber="603" pageNumber="605" updateTime="1699352364461" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="31D09836764379EE6FB814083BCDEF2B">Parabintoniella papierae n. gen., n. sp., a new Triassic representative of the Bintoniellidae, a poorly known extinct family of Orthoptera (Insecta)</mods:title>
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<mods:namePart id="2CAFBA9F4CC4B6782B5C953575800160">Béthoux, Olivier</mods:namePart>
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<mods:roleTerm id="381236F691720ADE60A782B0A65CEFFF">Author</mods:roleTerm>
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<mods:namePart id="C6619E74BB7DBD4D61F2954A1F96F0B4">Grauvogel-Stamm, Léa</mods:namePart>
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<mods:typeOfResource id="4BD1E323544CDB4D52585F9C41F908B9">text</mods:typeOfResource>
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<mods:title id="9B36288D3DB292A228B7DC09AC5F5D30">Geodiversitas</mods:title>
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<mods:part id="5DAC469310A69CBCE8990A6DB0705853">
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<mods:date id="EC3A51FB7C497ABCB1AE1A666426BA86">2022</mods:date>
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<mods:number id="B8F367BBD8E1B1CE1FAD08B1B1A5BE28">2022-06-23</mods:number>
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<mods:number id="2BD2026661C370B6A382D256AF4263FD">44</mods:number>
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<treatment id="03EF0D12816DFFCBFF62FA405A6BB8F2" ID-DOI="http://doi.org/10.5281/zenodo.6787054" ID-GBIF-Taxon="196373164" ID-Zenodo-Dep="6787054" LSID="urn:lsid:zoobank.org:act:1AAFA563-F408-41C0-A519-B1E25EC3ABCA" httpUri="http://treatment.plazi.org/id/03EF0D12816DFFCBFF62FA405A6BB8F2" lastPageId="6" lastPageNumber="607" pageId="4" pageNumber="605">
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<subSubSection id="C35CEF8F816DFFC9FF62FA405DCBB88F" box="[242,666,1461,1488]" pageId="4" pageNumber="605" type="nomenclature">
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<paragraph id="8BF9BC04816DFFC9FF62FA405DCBB88F" blockId="4.[242,666,1461,1520]" box="[242,666,1461,1488]" pageId="4" pageNumber="605">
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<heading id="D0B10B68816DFFC9FF62FA405DCBB88F" box="[242,666,1461,1488]" level="2" pageId="4" pageNumber="605">
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<taxonomicName id="4C46C787816DFFC9FF62FA405D51B890" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[242,512,1461,1488]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="605" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
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<emphasis id="B9326016816DFFC9FF62FA405D51B890" bold="true" box="[242,512,1461,1488]" italics="true" pageId="4" pageNumber="605">Parabintoniella papierae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A201DD6D816DFFC9FD97FA435DCBB88F" box="[519,666,1462,1488]" pageId="4" pageNumber="605" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CEF8F816DFFC9FE1FFA205C56BB76" pageId="4" pageNumber="605" type="description">
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<paragraph id="8BF9BC04816DFFC9FE1FFA205EADB8B0" blockId="4.[242,666,1461,1520]" box="[399,508,1493,1520]" pageId="4" pageNumber="605">
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(
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<figureCitation id="137DA081816DFFC9FE08FA205E86B8B0" box="[408,471,1493,1519]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Figs 1</figureCitation>
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;
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<figureCitation id="137DA081816DFFC9FE75FA205EA2B8B0" box="[485,499,1493,1519]" captionStart="FIG" captionStartId="6.[132,143,600,617]" captionTargetBox="[820,1456,206,1010]" captionTargetId="figure-495@6.[132,776,214,644]" captionTargetPageId="6" captionText="FIG. 2. — Parabintoniella papierae n. gen., n. sp., specimen 9122b (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’), photograph of detail of medio-cubital area (location as shown with a white frame in Fig.1C); the sign * indicates the area where the free basal portion of CuPaα, present in the Orthoptera groundplan but absent in the Bintoniellidae, could be expected to occur. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787065" httpUri="https://zenodo.org/record/6787065/files/figure.png" pageId="4" pageNumber="605">2</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF9BC04816DFFC9FF14F9ED5C56BB76" blockId="4.[132,775,1560,1577]" box="[132,775,1560,1577]" pageId="4" pageNumber="605">
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<uri id="FFD7B006816DFFC9FF14F9ED5C56BB76" box="[132,775,1560,1577]" pageId="4" pageNumber="605">
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urn:lsid:zoobank.org:act:
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CEF8F816DFFC9FF14F9A25EDBBA5A" pageId="4" pageNumber="605" type="materials_examined">
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<paragraph id="8BF9BC04816DFFC9FF14F9A25E38BBFB" blockId="4.[132,776,1622,1700]" pageId="4" pageNumber="605">
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<materialsCitation id="3B2EB659816DFFC9FF14F9A25E38BBFB" ID-GBIF-Occurrence="3821853302" country="France" location="Gottenhouse" municipality="Gottenhouse" pageId="4" pageNumber="605" specimenCode="5589/5590" specimenCount="1" stateProvince="Bas-Rhin" typeStatus="holotype">
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<typeStatus id="54FD02A6816DFFC9FF14F9A25FBEBB31" box="[132,239,1623,1646]" pageId="4" pageNumber="605" type="holotype">HOLOTYPE</typeStatus>
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. —
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<specimenCount id="9D40778D816DFFC9FE8CF9A25E2ABB30" box="[284,379,1623,1647]" pageId="4" pageNumber="605" type="generic">Specimen</specimenCount>
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<specimenCode id="DBE0147F816DFFC9FE12F9A35EA1BB31" box="[386,496,1622,1646]" pageId="4" pageNumber="605">5589/5590</specimenCode>
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(collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’, Stuttgart, Germany) (
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<figureCitation id="137DA081816DFFC9FF67F97E5E0CBBFB" box="[247,349,1675,1700]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Fig. 1A, B</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF9BC04816DFFC9FF14F94C5EDBBA5A" blockId="4.[132,777,1720,1797]" pageId="4" pageNumber="605">
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<materialsCitation id="3B2EB659816DFFC9FF14F94C5EDBBA5A" ID-GBIF-Occurrence="3821853301" country="France" location="Bust" municipality="Bust" pageId="4" pageNumber="605" specimenCode="9122a, b" specimenCount="1" stateProvince="Bas-Rhin" typeStatus="paratype">
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<typeStatus id="54FD02A6816DFFC9FF14F94C5FB2BB8F" box="[132,227,1721,1744]" pageId="4" pageNumber="605" type="paratype">PARATYPE</typeStatus>
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. —
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<specimenCount id="9D40778D816DFFC9FE83F94D5E23BB8F" box="[275,370,1720,1744]" pageId="4" pageNumber="605" type="generic">Specimen</specimenCount>
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<specimenCode id="DBE0147F816DFFC9FEEAF94D5E83BB8F" box="[378,466,1720,1744]" pageId="4" pageNumber="605">9122a, b</specimenCode>
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(collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’, Stuttgart, Germany) (
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<figureCitation id="137DA081816DFFC9FF67F9185E34BA5A" box="[247,357,1773,1797]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Figs 1C, D</figureCitation>
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;
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<figureCitation id="137DA081816DFFC9FEFFF9185E2DBA5A" box="[367,380,1773,1797]" captionStart="FIG" captionStartId="6.[132,143,600,617]" captionTargetBox="[820,1456,206,1010]" captionTargetId="figure-495@6.[132,776,214,644]" captionTargetPageId="6" captionText="FIG. 2. — Parabintoniella papierae n. gen., n. sp., specimen 9122b (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’), photograph of detail of medio-cubital area (location as shown with a white frame in Fig.1C); the sign * indicates the area where the free basal portion of CuPaα, present in the Orthoptera groundplan but absent in the Bintoniellidae, could be expected to occur. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787065" httpUri="https://zenodo.org/record/6787065/files/figure.png" pageId="4" pageNumber="605">2</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CEF8F816DFFC9FF14F8EE5E39BAB4" pageId="4" pageNumber="605" type="diagnosis">
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<paragraph id="8BF9BC04816DFFC9FF14F8EE5E39BAB4" blockId="4.[130,776,1818,2027]" pageId="4" pageNumber="605">
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DIAGNOSIS. — Forewing (and see
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<figureCitation id="137DA081816DFFC9FE59F8EF5EAEBA6D" box="[457,511,1818,1842]" captionStart="FIG" captionStartId="6.[813,824,1035,1052]" captionTargetBox="[820,1456,206,1010]" captionTargetPageId="6" captionText="FIG. 3. — Summary of the assumed phylogenetic relationships of the taxa and species considered in this contribution (character states invariant under different hypotheses in dark grey): A, favoured hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Eubintoniella ferganica Gorochov, 1987 and Bintoniella brodiei Handlirsch, 1939, owing to the occurrence of a simple MA in the forewing of these species; B, alternative hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Oshiellinae, Eubintoniella ferganica and Bintoniella brodiei, owing to the occurrence of cross-veins between ScP veinlets in the forewing of Parabintoniella papierae n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.6787067" httpUri="https://zenodo.org/record/6787067/files/figure.png" pageId="4" pageNumber="605">Fig. 3</figureCitation>
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): no distinct base of CuPaα diverging from CuPa (and fusing with CuA or M+CuA) (derived state shared with all
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<taxonomicName id="4C46C787816DFFC9FED6F8BA5E9EBA38" authorityName="Handlirsch" authorityYear="1938" box="[326,463,1871,1895]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="605" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
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as delimited herein); MA simple (derived state shared with
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<taxonomicName id="4C46C787816DFFC9FE12F89C5D61BADE" authorityName="Handlirsch" authorityYear="1938" box="[386,560,1897,1921]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="605" phylum="Arthropoda" rank="species" species="brodiei">
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<emphasis id="B9326016816DFFC9FE12F89C5D61BADE" box="[386,560,1897,1921]" italics="true" pageId="4" pageNumber="605">Bintoniella brodiei</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C46C787816DFFC9FDF1F89C5F91BAC3" authorityName="Gorochov" authorityYear="1987" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="605" phylum="Arthropoda" rank="species" species="ferganica">
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<emphasis id="B9326016816DFFC9FDF1F89C5F91BAC3" italics="true" pageId="4" pageNumber="605">Eubintoniella ferganica</emphasis>
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</taxonomicName>
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); CuA+CuPaα with more than two branches (plesiomorphy shared with
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<taxonomicName id="4C46C787816DFFC9FF63F86B5E3ABAE9" authorityName="Gorochov" authorityYear="1989" box="[243,363,1950,1974]" higherTaxonomySource="Manual Input" pageId="4" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
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and Oshiellinae); cross-veins forming loose, convex intercalaries between ScP branches (shared with
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<taxonomicName id="4C46C787816DFFC9FD3FF84C5FE4BAB4" authorityName="Gorochov" authorityYear="1989" higherTaxonomySource="Manual Input" pageId="4" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
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, polarity unclear).
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</paragraph>
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</subSubSection>
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<subSubSection id="C35CEF8F816DFFC9FCBDFF2D5B5ABC7B" pageId="4" pageNumber="605" type="etymology">
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<paragraph id="8BF9BC04816DFFC9FCBDFF2D5B5ABC7B" blockId="4.[813,1455,215,293]" pageId="4" pageNumber="605">ETYMOLOGY. — The species epithet honours the late Francine Papier, who significantly contributed to the knowledge of the ‘Grès à Voltzia’ fossil insects.</paragraph>
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</subSubSection>
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<subSubSection id="C35CEF8F816DFFC9FCBDFEC85AFCBCE2" pageId="4" pageNumber="605" type="materials_examined">
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<paragraph id="8BF9BC04816DFFC9FCBDFEC85AFCBCE2" blockId="4.[813,1457,316,446]" pageId="4" pageNumber="605">
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<collectionCode id="ED5724C1816DFFC9FCBDFEC85C0EBC0B" box="[813,863,317,340]" pageId="4" pageNumber="605">TYPE</collectionCode>
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LOCALITY AND STRATIGRAPHY. — The
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<typeStatus id="54FD02A6816DFFC9FB60FEC95A19BC0B" box="[1264,1352,316,340]" pageId="4" pageNumber="605" type="holotype">holotype</typeStatus>
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specimen 5589/5590 was collected at
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<collectingMunicipality id="6B9D267E816DFFC9FBD5FEA35B95BC31" box="[1093,1220,342,366]" pageId="4" pageNumber="605">Gottenhouse</collectingMunicipality>
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(
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<collectingRegion id="498272E6816DFFC9FB42FEA35A60BC31" box="[1234,1329,342,366]" pageId="4" pageNumber="605">Bas-Rhin</collectingRegion>
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||
,
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<collectingCountry id="F351FC94816DFFC9FAACFEA25A2EBC30" box="[1340,1407,343,367]" name="France" pageId="4" pageNumber="605">France</collectingCountry>
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||
) and the
|
||
<typeStatus id="54FD02A6816DFFC9FCC4FE845CF6BCD6" box="[852,935,369,393]" pageId="4" pageNumber="605" type="paratype">paratype</typeStatus>
|
||
specimen 9122a, b was collected at
|
||
<collectingMunicipality id="6B9D267E816DFFC9FA84FE845A11BCD7" box="[1300,1344,369,392]" pageId="4" pageNumber="605">Bust</collectingMunicipality>
|
||
(
|
||
<collectingRegion id="498272E6816DFFC9FADEFE845AFCBCD6" box="[1358,1453,369,393]" pageId="4" pageNumber="605">Bas-Rhin</collectingRegion>
|
||
,
|
||
<collectingCountry id="F351FC94816DFFC9FCBEFE795C23BCFB" box="[814,882,396,420]" name="France" pageId="4" pageNumber="605">France</collectingCountry>
|
||
). Both specimens come from the ‘Grès à Voltzia’ (Upper Buntsandstein; early Middle Triassic; Gall & Grauvogel-Stamm 2005).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CEF8F816DFFC9FCBDFE035B1BBBB3" pageId="4" pageNumber="605" type="description">
|
||
<paragraph id="8BF9BC04816DFFC9FCBDFE035B75BF50" blockId="4.[811,1462,502,1262]" box="[813,1060,502,528]" pageId="4" pageNumber="605">GENERAL DESCRIPTION</paragraph>
|
||
<paragraph id="8BF9BC04816DFFC9FCBDFDE35B3FB9B1" blockId="4.[811,1462,502,1262]" pageId="4" pageNumber="605">
|
||
Forewing: length about 22.4-28.0 mm, width 6.4-8.0 mm; ScA conspicuous; in its median part, area between anterior wing margin and ScP broad, with cross-veins forming loose, convex intercalaries between ScP branches (concave); ScP reaching the anterior wing margin from two-thirds to threequarters of wing length; RA/RP split slightly distal to wing mid-length; RA with anterior veinlets and cross-veins between them; RP without well-organized branching pattern, with five to eight terminal branches/veinlets covering the apex; M+CuA(+CuPaα) splitting into M and CuA+CuPaα at about the first third of wing length; free basal portion of M (between the M/CuA(+CuPaα) split and the MA/MP split) relatively long; both MA and MP simple (known in
|
||
<typeStatus id="54FD02A6816DFFC9FA9AFC605A38BEF0" box="[1290,1385,917,943]" pageId="4" pageNumber="605" type="holotype">holotype</typeStatus>
|
||
only); CuA+CuPaα posteriorly pectinate, with four branches; no distinct base of CuPaα diverging from CuPa (and fusing with CuA or M+CuA; better visible in specimen 9122); CuPaβ and CuPb simple; CuPb reaching posterior wing margin at wing mid-length; AA with at least four, simple veins; cross-veins reticulated (i.e. forming 2-3 cells between two main veins) between ScP veinlets, in the apical half, elsewhere mostly scalariform; along the posterior wing margin, areas between CuPb and AA branches with numerous, sigmoidal cross-veins; membrane uniformly hyaline.
|
||
</paragraph>
|
||
<paragraph id="8BF9BC04816DFFC9FCBDFAE15B64B873" blockId="4.[810,1457,1300,1772]" box="[813,1077,1300,1326]" pageId="4" pageNumber="605">SPECIMEN DESCRIPTIONS</paragraph>
|
||
<paragraph id="8BF9BC04816DFFC9FCBDFAC65B1BBBB3" blockId="4.[810,1457,1300,1772]" pageId="4" pageNumber="605">
|
||
Specimen 5589/5590 (
|
||
<figureCitation id="137DA081816DFFC9FBB3FAC65BC0B812" box="[1059,1169,1331,1358]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Fig. 1A, B</figureCitation>
|
||
): adpression of an isolated, nearly complete left forewing, in both dorsal (5590) and ventral (5589) aspects, presumably of a female insect (see Remarks section); preserved length
|
||
<quantity id="4CBE11E1816DFFC9FB56FA665A61B8F1" box="[1222,1328,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="4" pageNumber="605" unit="mm" value="26.6">26.6 mm</quantity>
|
||
(estimated complete length about 28.0 mm), width 8.0 mm (at its widest); RP with either eight terminal branches or seven with an intercalary vein between its most posterior branch and MA. Specimen 9122a,b (
|
||
<figureCitation id="137DA081816DFFC9FB83F9E75BD6BB72" box="[1043,1159,1554,1581]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Figs 1C, D</figureCitation>
|
||
;
|
||
<figureCitation id="137DA081816DFFC9FB1EF9E65BCDBB72" box="[1166,1180,1555,1581]" captionStart="FIG" captionStartId="6.[132,143,600,617]" captionTargetBox="[820,1456,206,1010]" captionTargetId="figure-495@6.[132,776,214,644]" captionTargetPageId="6" captionText="FIG. 2. — Parabintoniella papierae n. gen., n. sp., specimen 9122b (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’), photograph of detail of medio-cubital area (location as shown with a white frame in Fig.1C); the sign * indicates the area where the free basal portion of CuPaα, present in the Orthoptera groundplan but absent in the Bintoniellidae, could be expected to occur. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787065" httpUri="https://zenodo.org/record/6787065/files/figure.png" pageId="4" pageNumber="605">2</figureCitation>
|
||
): adpression of an isolated left forewing, in both dorsal (9122a) and ventral (9122b) aspects, with most of the postero-distal area and area basal to the claval furrow (itself located along CuPb; i.e. the remigulum) missing, presumably of a male insect (see Remarks section); preserved length
|
||
<quantity id="4CBE11E1816DFFC9FC72F9475B15BB92" box="[994,1092,1714,1741]" metricMagnitude="-2" metricUnit="m" metricValue="2.06" pageId="4" pageNumber="605" unit="mm" value="20.6">20.6 mm</quantity>
|
||
(estimated complete length about
|
||
<quantity id="4CBE11E1816DFFC9FCBBF9275CC2BBB3" box="[811,915,1746,1772]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="4" pageNumber="605" unit="mm" value="22.4">22.4 mm</quantity>
|
||
), width
|
||
<quantity id="4CBE11E1816DFFC9FC7CF9275B17BBB3" box="[1004,1094,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="4" pageNumber="605" unit="mm" value="6.4">6.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35CEF8F816DFFCBFCBDF8E75A6BB8F2" lastPageId="6" lastPageNumber="607" pageId="4" pageNumber="605" type="discussion">
|
||
<paragraph id="8BF9BC04816DFFC9FCBDF8E75CC0BA73" blockId="4.[811,1455,1810,2027]" box="[813,913,1810,1836]" pageId="4" pageNumber="605">REMARKS</paragraph>
|
||
<paragraph id="8BF9BC04816DFFC8FCBDF8C75A58BAB4" blockId="4.[811,1455,1810,2027]" lastBlockId="5.[811,1455,1905,2028]" lastPageId="5" lastPageNumber="606" pageId="4" pageNumber="605">
|
||
From the difference in width of the two forewings at their widest, which is the single and most accurate comparable dimension between them, the size of the smallest one (
|
||
<figureCitation id="137DA081816DFFC9FACBF8845C12BAF4" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Fig. 1C, D</figureCitation>
|
||
) is 80% that of the largest (
|
||
<figureCitation id="137DA081816DFFC9FBE0F8645BB1BAF4" box="[1136,1248,1937,1963]" captionStart="FIG" captionStartId="5.[132,143,1751,1768]" captionTargetBox="[132,1410,185,1731]" captionTargetId="figure-104@5.[745,1316,754,1296]" captionTargetPageId="5" captionText="FIG. 1. — Parabintoniella papierae n. gen., n. sp.: A, B, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); A, photograph, ventral aspect (5589), light-mirrored; B, drawing of venation (reconstructed parts in grey); C, D, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’); C, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2); D, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm." figureDoi="http://doi.org/10.5281/zenodo.6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="4" pageNumber="605">Fig. 1A, B</figureCitation>
|
||
). This difference in size falls within the range of intra-specific variability and more particularly within that of sexual size dimorphism (SSD), commonly female-biased in extant
|
||
<taxonomicName id="4C46C787816CFFC8FE62F8845E7BBAF4" authority="(Hochkirch & Groning 2008)" baseAuthorityName="Hochkirch & Groning" baseAuthorityYear="2008" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="order">
|
||
Orthoptera (
|
||
<bibRefCitation id="EFD7C1F5816CFFC8FDE8F8845E4EBAF4" author="HOCHKIRCH A. & GRONING J." pageId="5" pageNumber="606" pagination="189 - 196" refId="ref5185" refString="HOCHKIRCH A. & GRONING J. 2008. - Sexual size dimorphism in Orthoptera (sens. str.) - a review. Journal of Orthoptera Research 17 (2): 189 - 196. https: // doi. org / 10.1665 / 1082 - 6467 - 17.2.189" type="journal article" year="2008">Hochkirch & Gröning 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Moreover, such phenomenon is also known to occur in early stem-Orthoptera (
|
||
<bibRefCitation id="EFD7C1F5816CFFC8FD95F8475DF2BA94" author="DU S. - L. & BETHOUX O. & GU J. - J. & REN D." box="[517,675,1969,1996]" pageId="5" pageNumber="606" pagination="193 - 204" refId="ref4691" refString="DU S. - L., BETHOUX O., GU J. - J. & REN D. 2017. - Protomiamia yangi gen. et sp. nov. (Early Pennsylvanian; Xiaheyan, China), a sexually dimorphic Palaeozoic stem-Orthoptera. Journal of Systematic Palaeontology 15 (3): 193 - 204. https: // doi. org / 10.10 80 / 14772019.2016.1154899" type="journal article" year="2017">
|
||
Du
|
||
<emphasis id="B9326016816CFFC8FDA0F8475D35BA94" box="[560,612,1969,1995]" italics="true" pageId="5" pageNumber="606">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
) and has already been assumed for the related species
|
||
<taxonomicName id="4C46C787816CFFC8FDD4F8245DAFBAB4" authorityName="Handlirsch" authorityYear="1938" box="[580,766,2001,2027]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816CFFC8FDD4F8245DAFBAB4" box="[580,766,2001,2027]" italics="true" pageId="5" pageNumber="606">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
, with a similar ratio (
|
||
<bibRefCitation id="EFD7C1F5816CFFC8FB96F8845BCFBAD4" author="WHALLEY P. E. S." box="[1030,1182,1905,1931]" pageId="5" pageNumber="606" pagination="143 - 149" refId="ref5581" refString="WHALLEY P. E. S. 1982. - Bintoniella brodiei Handlirsch (Orthoptera) from the Lower Lias of the English Channel, with a review of Britisch bintoniellid fossils. Bulletin of the British Museum of Natural History (Geology) 36 (2): 143 - 149. https: // www. biodiversitylibrary. org / page / 2364267" type="journal article" year="1982">Whalley 1982</bibRefCitation>
|
||
). As the other differences are minimal, it is doubtless that the
|
||
<specimenCount id="9D40778D816CFFC8FB38F8675A15BAF4" box="[1192,1348,1937,1964]" pageId="5" pageNumber="606" type="generic">two specimens</specimenCount>
|
||
belong to the same species, the specimen 5589/5590 presumably being a female, and the specimen 9122a, b a male.
|
||
</paragraph>
|
||
<caption id="DF39EC8C816CFFC8FF14F9225B01BA73" ID-DOI="http://doi.org/10.5281/zenodo.6787063" ID-Zenodo-Dep="6787063" httpUri="https://zenodo.org/record/6787063/files/figure.png" pageId="5" pageNumber="606" startId="5.[132,143,1751,1768]" targetBox="[132,1410,185,1731]" targetPageId="5">
|
||
<paragraph id="8BF9BC04816CFFC8FF14F9225B01BA73" blockId="5.[132,1456,1751,1836]" pageId="5" pageNumber="606">
|
||
FIG. 1. —
|
||
<taxonomicName id="4C46C787816CFFC8FF44F9225ECABBB7" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[212,411,1751,1768]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="606" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
|
||
<emphasis id="B9326016816CFFC8FF44F9225ECABBB7" box="[212,411,1751,1768]" italics="true" pageId="5" pageNumber="606">Parabintoniella papierae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A201DD6D816CFFC8FE30F9225D44BBB7" box="[416,533,1751,1768]" pageId="5" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B9326016816CFFC8FD8CF9225D78BBB7" bold="true" box="[540,553,1751,1768]" pageId="5" pageNumber="606">A</emphasis>
|
||
,
|
||
<emphasis id="B9326016816CFFC8FDA3F9225D11BBB7" bold="true" box="[563,576,1751,1768]" pageId="5" pageNumber="606">B</emphasis>
|
||
, holotype, specimen 5589/5590, left forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’);
|
||
<emphasis id="B9326016816CFFC8FED8F91B5E04BBA0" bold="true" box="[328,341,1774,1791]" pageId="5" pageNumber="606">A</emphasis>
|
||
, photograph, ventral aspect (5589), light-mirrored;
|
||
<emphasis id="B9326016816CFFC8FD6DF91B5C5BBBA0" bold="true" box="[765,778,1774,1791]" pageId="5" pageNumber="606">B</emphasis>
|
||
, drawing of venation (reconstructed parts in grey);
|
||
<emphasis id="B9326016816CFFC8FB21F91B5BEEBBA0" bold="true" box="[1201,1215,1774,1791]" pageId="5" pageNumber="606">C</emphasis>
|
||
,
|
||
<emphasis id="B9326016816CFFC8FB59F91B5B86BBA0" bold="true" box="[1225,1239,1774,1791]" pageId="5" pageNumber="606">D</emphasis>
|
||
, specimen 9122a, b, right forewing (collection Louis Grauvogel at the ‘Staatlisches Museum für Naturkunde Stuttgart’);
|
||
<emphasis id="B9326016816CFFC8FC08F8F05CF7BA49" bold="true" box="[920,934,1797,1814]" pageId="5" pageNumber="606">C</emphasis>
|
||
, photograph, ventral aspect (9122b), light-mirrored (the white frame indicates the section enlarged in Fig. 2);
|
||
<emphasis id="B9326016816CFFC8FD9AF8EE5D49BA73" bold="true" box="[522,536,1819,1836]" pageId="5" pageNumber="606">D</emphasis>
|
||
, drawing of venation (reconstructed parts in grey). Scale bar: 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF39EC8C816FFFCBFF14FDAD5E81BF84" ID-DOI="http://doi.org/10.5281/zenodo.6787065" ID-Zenodo-Dep="6787065" httpUri="https://zenodo.org/record/6787065/files/figure.png" pageId="6" pageNumber="607" startId="6.[132,143,600,617]" targetBox="[820,1456,206,1010]" targetPageId="6">
|
||
<paragraph id="8BF9BC04816FFFCBFF14FDAD5E81BF84" blockId="6.[132,775,600,731]" pageId="6" pageNumber="607">
|
||
FIG. 2. —
|
||
<taxonomicName id="4C46C787816FFFCBFF45FDAD5ECCBF36" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[213,413,600,617]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
|
||
<emphasis id="B9326016816FFFCBFF45FDAD5ECCBF36" box="[213,413,600,617]" italics="true" pageId="6" pageNumber="607">Parabintoniella papierae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A201DD6D816FFFCBFE33FDAD5D4BBF36" box="[419,538,600,617]" pageId="6" pageNumber="607" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
, specimen 9122b (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’), photograph of detail of medio-cubital area (location as shown with a white frame in Fig.1C); the sign
|
||
<emphasis id="B9326016816FFFCBFEB6FD695E7FBFF2" bold="true" box="[294,302,668,685]" pageId="6" pageNumber="607">*</emphasis>
|
||
indicates the area where the free basal portion of CuPaα, present in the
|
||
<taxonomicName id="4C46C787816FFFCBFF6FFD465E0BBF9B" baseAuthorityName="Hochkirch & Groning" baseAuthorityYear="2008" box="[255,346,691,708]" class="Insecta" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
|
||
groundplan but absent in the
|
||
<taxonomicName id="4C46C787816FFFCBFDCDFD465D81BF9B" authorityName="Handlirsch" authorityYear="1938" box="[605,720,691,708]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
|
||
, could be expected to occur. Scale bar: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9BC04816FFFCBFF0BFCE05D2BB971" blockId="6.[131,777,789,2028]" pageId="6" pageNumber="607">
|
||
With the aid of the systematic conventions posited above, the relationships between the species to which the new specimens belong can be discussed (
|
||
<figureCitation id="137DA081816FFFCBFE77FCA05D76BE30" box="[487,551,853,879]" captionStart="FIG" captionStartId="6.[813,824,1035,1052]" captionTargetBox="[820,1456,206,1010]" captionTargetPageId="6" captionText="FIG. 3. — Summary of the assumed phylogenetic relationships of the taxa and species considered in this contribution (character states invariant under different hypotheses in dark grey): A, favoured hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Eubintoniella ferganica Gorochov, 1987 and Bintoniella brodiei Handlirsch, 1939, owing to the occurrence of a simple MA in the forewing of these species; B, alternative hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Oshiellinae, Eubintoniella ferganica and Bintoniella brodiei, owing to the occurrence of cross-veins between ScP veinlets in the forewing of Parabintoniella papierae n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.6787067" httpUri="https://zenodo.org/record/6787067/files/figure.png" pageId="6" pageNumber="607">Fig. 3</figureCitation>
|
||
). The occurrence of the character state ‘in forewing, no distinct base of CuPaα diverging from CuPa (and fusing with CuA or M+CuA)’ (to be understood as ‘beyond the wing base’) qualifies an assignment to the
|
||
<taxonomicName id="4C46C787816FFFCBFEDEFC215EA1BEB1" authorityName="Handlirsch" authorityYear="1938" box="[334,496,980,1006]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
|
||
. In other words, affinities with the
|
||
<taxonomicName id="4C46C787816FFFCBFF7DFC015E2CB951" authorityName="Gorochov" authorityYear="1989" box="[237,381,1012,1038]" higherTaxonomySource="Manual Input" pageId="6" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
|
||
can be excluded. This condition is best observed in the specimen 9122 (
|
||
<figureCitation id="137DA081816FFFCBFDB8FBE15D38B971" box="[552,617,1044,1070]" captionStart="FIG" captionStartId="6.[132,143,600,617]" captionTargetBox="[820,1456,206,1010]" captionTargetId="figure-495@6.[132,776,214,644]" captionTargetPageId="6" captionText="FIG. 2. — Parabintoniella papierae n. gen., n. sp., specimen 9122b (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’), photograph of detail of medio-cubital area (location as shown with a white frame in Fig.1C); the sign * indicates the area where the free basal portion of CuPaα, present in the Orthoptera groundplan but absent in the Bintoniellidae, could be expected to occur. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6787065" httpUri="https://zenodo.org/record/6787065/files/figure.png" pageId="6" pageNumber="607">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF39EC8C816FFFCBFCBDFBFE5BD7B9B7" ID-DOI="http://doi.org/10.5281/zenodo.6787067" ID-Zenodo-Dep="6787067" httpUri="https://zenodo.org/record/6787067/files/figure.png" pageId="6" pageNumber="607" startId="6.[813,824,1035,1052]" targetBox="[820,1456,206,1010]" targetPageId="6">
|
||
<paragraph id="8BF9BC04816FFFCBFCBDFBFE5BD7B9B7" blockId="6.[813,1456,1035,1256]" pageId="6" pageNumber="607">
|
||
FIG. 3. — Summary of the assumed phylogenetic relationships of the taxa and species considered in this contribution (character states invariant under different hypotheses in dark grey):
|
||
<emphasis id="B9326016816FFFCBFBCFFBCD5B3DB916" bold="true" box="[1119,1132,1080,1097]" pageId="6" pageNumber="607">A</emphasis>
|
||
, favoured hypothesis, assuming that
|
||
<taxonomicName id="4C46C787816FFFCBFCBDFBBA5CADB93F" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[813,1020,1103,1120]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
|
||
<emphasis id="B9326016816FFFCBFCBDFBBA5CADB93F" box="[813,1020,1103,1120]" italics="true" pageId="6" pageNumber="607">Parabintoniella papierae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A201DD6D816FFFCBFB93FBBA5B2FB93F" box="[1027,1150,1103,1120]" pageId="6" pageNumber="607" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
is the sister-group of
|
||
<taxonomicName id="4C46C787816FFFCBFAD1FBBA5B56B929" authority="Gorochov, 1987" authorityName="Gorochov" authorityYear="1987" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFAD1FBBA5C2BB929" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFCEFFB905B56B929" author="GOROCHOV A. V." box="[895,1031,1125,1142]" pageId="6" pageNumber="607" pagination="18 - 23" refId="ref4819" refString="GOROCHOV A. V. 1987. - New fossil Orthoptera of the families Bintoniellidae, Mesoedischiidae fam. n. and Pseudelcanidae fam. n. (Orthoptera, Ensifera) from Permian and Triassic deposits of the USSR territory. Vestnik Zoologii 1: 18 - 23 [in Russian with English summary]." type="journal article" year="1987">Gorochov, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46C787816FFFCBFBA1FB905A0EB929" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[1073,1375,1125,1142]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFBA1FB905B99B929" box="[1073,1224,1125,1142]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
Handlirsch, 1939
|
||
</taxonomicName>
|
||
, owing to the occurrence of a simple MA in the forewing of these species;
|
||
<emphasis id="B9326016816FFFCBFAC6FB895A32B9D2" bold="true" box="[1366,1379,1148,1165]" pageId="6" pageNumber="607">B</emphasis>
|
||
, alternative hypothesis, assuming that
|
||
<taxonomicName id="4C46C787816FFFCBFBA5FB665A51B9FB" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[1077,1280,1171,1188]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
|
||
<emphasis id="B9326016816FFFCBFBA5FB665A51B9FB" box="[1077,1280,1171,1188]" italics="true" pageId="6" pageNumber="607">Parabintoniella papierae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A201DD6D816FFFCBFA96FB665A2DB9FB" box="[1286,1404,1171,1188]" pageId="6" pageNumber="607" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
is the sister-group of Oshiellinae,
|
||
<taxonomicName id="4C46C787816FFFCBFB8EFB5C5BB0B9E5" authorityName="Gorochov" authorityYear="1987" box="[1054,1249,1193,1210]" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFB8EFB5C5BB0B9E5" box="[1054,1249,1193,1210]" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46C787816FFFCBFA9FFB5C5AFBB9E5" authorityName="Handlirsch" authorityYear="1939" box="[1295,1450,1193,1210]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFA9FFB5C5AFBB9E5" box="[1295,1450,1193,1210]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
, owing to the occurrence of cross-veins between ScP veinlets in the forewing of
|
||
<taxonomicName id="4C46C787816FFFCBFCD2FB225B5DB9B7" authority="Béthoux & Grauvogel-Stamm, 2022" authorityName="Béthoux & Grauvogel-Stamm" authorityYear="2022" box="[834,1036,1239,1256]" class="Insecta" family="Bintoniellidae" genus="Parabintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="papierae" status="n. gen., n. sp.">
|
||
<emphasis id="B9326016816FFFCBFCD2FB225B5DB9B7" box="[834,1036,1239,1256]" italics="true" pageId="6" pageNumber="607">Parabintoniella papierae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A201DD6D816FFFCBFB81FB225BD7B9B7" box="[1041,1158,1239,1256]" pageId="6" pageNumber="607" rank="species">n. gen., n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9BC04816FFFCBFF0BFBC05D79BB12" blockId="6.[131,777,789,2028]" pageId="6" pageNumber="607">
|
||
In contrast to all the
|
||
<taxonomicName id="4C46C787816FFFCBFE1BFBC15D4AB911" authorityName="Gorochov" authorityYear="1989" box="[395,539,1076,1102]" higherTaxonomySource="Manual Input" pageId="6" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
|
||
and the Oshiellinae, in which MA commonly has 3-4 branches in the forewing, the species to which the new specimens belong displays a simple MA. This feature, which is shared with
|
||
<taxonomicName id="4C46C787816FFFCBFD1FFB615F81B991" authorityName="Handlirsch" authorityYear="1939" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFD1FFB615F81B991" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFE99FB415EF9B991" author="WHALLEY P. E. S." box="[265,424,1204,1230]" pageId="6" pageNumber="607" pagination="143 - 149" refId="ref5581" refString="WHALLEY P. E. S. 1982. - Bintoniella brodiei Handlirsch (Orthoptera) from the Lower Lias of the English Channel, with a review of Britisch bintoniellid fossils. Bulletin of the British Museum of Natural History (Geology) 36 (2): 143 - 149. https: // www. biodiversitylibrary. org / page / 2364267" type="journal article" year="1982">Whalley 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFE26FB415D07B991" author="BETHOUX O." box="[438,598,1204,1230]" pageId="6" pageNumber="607" pagination="179 - 225" refId="ref4280" refString="BETHOUX O. 2012. - King crickets, raspy crickets & weta, their wings, their fossil relatives. Journal of Orthoptera Research 21 (2): 179 - 225. https: // doi. org / 10.1665 / 034.021.0206" type="journal article" year="2012">Béthoux 2012</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C46C787816FFFCBFD09FB415E4FB9B1" authorityName="Gorochov" authorityYear="1987" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFD09FB415E4FB9B1" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
</taxonomicName>
|
||
(see the original description), is definitely derived and represents the diagnostic trait of a yet unnamed taxon. Furthermore, in contrast to
|
||
<taxonomicName id="4C46C787816FFFCBFD9BFAE65D87B872" authorityName="Handlirsch" authorityYear="1939" box="[523,726,1299,1325]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFD9BFAE65D87B872" box="[523,726,1299,1325]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46C787816FFFCBFF14FAC65EC1B812" authorityName="Gorochov" authorityYear="1987" box="[132,400,1331,1357]" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFF14FAC65EC1B812" box="[132,400,1331,1357]" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
</taxonomicName>
|
||
, in which CuA+CuPaα has two branches (rare exceptions are known in
|
||
<taxonomicName id="4C46C787816FFFCBFDA6FAA65DAFB832" authorityName="Handlirsch" authorityYear="1939" box="[566,766,1363,1389]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFDA6FAA65DAFB832" box="[566,766,1363,1389]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFF3AFA865E17B8D2" author="BETHOUX O." box="[170,326,1395,1421]" pageId="6" pageNumber="607" pagination="179 - 225" refId="ref4280" refString="BETHOUX O. 2012. - King crickets, raspy crickets & weta, their wings, their fossil relatives. Journal of Orthoptera Research 21 (2): 179 - 225. https: // doi. org / 10.1665 / 034.021.0206" type="journal article" year="2012">Béthoux 2012</bibRefCitation>
|
||
), the species to which the new specimens belong displays a CuA+CuPaα with four branches. This trait must be regarded as a plesiomorphy, as
|
||
<taxonomicName id="4C46C787816FFFCBFDE8FA465C56B892" authorityName="Gorochov" authorityYear="1989" box="[632,775,1459,1485]" higherTaxonomySource="Manual Input" pageId="6" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
|
||
and Oshiellinae commonly possess a 4- to 5-branched CuA+CuPaα. It follows that the species to which the new specimens belong is the sister-group of
|
||
<taxonomicName id="4C46C787816FFFCBFDACF9E75C56BB73" authorityName="Handlirsch" authorityYear="1939" box="[572,775,1554,1580]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFDACF9E75C56BB73" box="[572,775,1554,1580]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46C787816FFFCBFF26F9C75EE2BB12" authorityName="Gorochov" authorityYear="1987" box="[182,435,1586,1613]" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFF26F9C75EE2BB12" box="[182,435,1586,1613]" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137DA081816FFFCBFE54F9C75D4DBB12" box="[452,540,1586,1613]" captionStart="FIG" captionStartId="6.[813,824,1035,1052]" captionTargetBox="[820,1456,206,1010]" captionTargetPageId="6" captionText="FIG. 3. — Summary of the assumed phylogenetic relationships of the taxa and species considered in this contribution (character states invariant under different hypotheses in dark grey): A, favoured hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Eubintoniella ferganica Gorochov, 1987 and Bintoniella brodiei Handlirsch, 1939, owing to the occurrence of a simple MA in the forewing of these species; B, alternative hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Oshiellinae, Eubintoniella ferganica and Bintoniella brodiei, owing to the occurrence of cross-veins between ScP veinlets in the forewing of Parabintoniella papierae n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.6787067" httpUri="https://zenodo.org/record/6787067/files/figure.png" pageId="6" pageNumber="607">Fig. 3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF9BC04816FFFCBFF0BF9A75A6BB8F2" blockId="6.[131,777,789,2028]" lastBlockId="6.[812,1456,1363,1454]" pageId="6" pageNumber="607">
|
||
Unlike
|
||
<taxonomicName id="4C46C787816FFFCBFF63F9A75ECBBB33" authorityName="Handlirsch" authorityYear="1938" box="[243,410,1618,1644]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
|
||
, in which no cross-veins occur between ScP branches, the species to which the new specimens belong displays cross-veins forming loose, convex intercalaries between ScP branches. This state occurring in
|
||
<taxonomicName id="4C46C787816FFFCBFF33F9275E67BBB3" authorityName="Gorochov" authorityYear="1989" box="[163,310,1746,1772]" higherTaxonomySource="Manual Input" pageId="6" pageNumber="605" rank="subFamily" subFamily="Proshiellinae">Proshiellinae</taxonomicName>
|
||
, it can then be regarded as a plesiomorphy within
|
||
<taxonomicName id="4C46C787816FFFCBFF40F9075E3FBA53" authorityName="Handlirsch" authorityYear="1938" box="[208,366,1778,1804]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
|
||
. Accordingly, the species to which the new specimens belong would be the sister-group of all other
|
||
<taxonomicName id="4C46C787816FFFCBFF13F8C45E4FBA14" authorityName="Handlirsch" authorityYear="1938" box="[131,286,1841,1867]" class="Insecta" family="Bintoniellidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="family">Bintoniellidae</taxonomicName>
|
||
(namely, Oshiellinae,
|
||
<taxonomicName id="4C46C787816FFFCBFD81F8C45D86BA14" authorityName="Handlirsch" authorityYear="1939" box="[529,727,1841,1867]" class="Insecta" family="Bintoniellidae" genus="Bintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="brodiei">
|
||
<emphasis id="B9326016816FFFCBFD81F8C45D86BA14" box="[529,727,1841,1867]" italics="true" pageId="6" pageNumber="607">Bintoniella brodiei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C46C787816FFFCBFF13F8A45ED0BA33" authorityName="Gorochov" authorityYear="1987" box="[131,385,1873,1900]" class="Insecta" family="Bintoniellidae" genus="Eubintoniella" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="species" species="ferganica">
|
||
<emphasis id="B9326016816FFFCBFF13F8A45ED0BA33" box="[131,385,1873,1900]" italics="true" pageId="6" pageNumber="607">Eubintoniella ferganica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="137DA081816FFFCBFE1FF8A45EB5BA33" box="[399,484,1873,1900]" captionStart="FIG" captionStartId="6.[813,824,1035,1052]" captionTargetBox="[820,1456,206,1010]" captionTargetPageId="6" captionText="FIG. 3. — Summary of the assumed phylogenetic relationships of the taxa and species considered in this contribution (character states invariant under different hypotheses in dark grey): A, favoured hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Eubintoniella ferganica Gorochov, 1987 and Bintoniella brodiei Handlirsch, 1939, owing to the occurrence of a simple MA in the forewing of these species; B, alternative hypothesis, assuming that Parabintoniella papierae n. gen., n. sp. is the sister-group of Oshiellinae, Eubintoniella ferganica and Bintoniella brodiei, owing to the occurrence of cross-veins between ScP veinlets in the forewing of Parabintoniella papierae n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.6787067" httpUri="https://zenodo.org/record/6787067/files/figure.png" pageId="6" pageNumber="607">Fig. 3B</figureCitation>
|
||
). Considering (i) that this scenario implies that the number of branches of MA would have been subjected to homoplasic evolution, and, on the other hand, (ii) that the occurrence of cross-veins between ScP veinlets, and their lack thereof, can both occur within a single genus (such as in the Permian stem-Orthoptera genus
|
||
<taxonomicName id="4C46C787816FFFCBFCE5FA865B9CB8D2" authority="Zalessky, 1929" authorityName="Zalessky" authorityYear="1929" box="[885,1229,1395,1421]" class="Insecta" family="Tcholmanvissiidae" genus="Tcholmanvissia" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="607" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9326016816FFFCBFCE5FA865B4DB8D2" box="[885,1052,1395,1421]" italics="true" pageId="6" pageNumber="607">Tcholmanvissia</emphasis>
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFBB4FA865B9CB8D2" author="ZALESSKY M. D." box="[1060,1229,1395,1421]" pageId="6" pageNumber="607" pagination="1 - 30" refId="ref5646" refString="ZALESSKY M. D. 1929. - About new insects from the Permian deposits of the basins of the Kama, Vyatka and Belaya rivers. Trudy obshchestva estestvoispytatelej pri kazanskom universitet 52 (1): 1 - 30 [in Russian]." type="journal article" year="1929">Zalessky, 1929</bibRefCitation>
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="EFD7C1F5816FFFCBFA92FA865C24B8F2" author="BETHOUX O. & NEL A." pageId="6" pageNumber="607" pagination="223 - 235" refId="ref4468" refString="BETHOUX O. & NEL A. 2002 b. - New data on Tcholmanvissiidae (Orthoptera; Permian). Journal of Orthoptera Research 11 (2): 223 - 235. https: // doi. org / ftx 74 v" type="journal article" year="2002">Béthoux & Nel 2002b</bibRefCitation>
|
||
), we consider this hypothesis less likely.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |