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<document id="B2F5AE7DF8E6B1F5AEF1D98806A3E112" ID-DOI="10.11646/zootaxa.4093.1.9" ID-GBIF-Dataset="7c162abc-6b9d-4560-aba8-fac6f2524cfe" ID-ISSN="1175-5326" ID-Zenodo-Dep="265878" ID-ZooBank="AE48DA77-FE06-4B2B-88CB-9195D3D855FD" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458561840274" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="03ECAF62FFBA2B26FF4E353BFDE081F8" docLanguage="en" docName="zootaxa.4093.1.9.pdf.imf" docOrigin="Zootaxa 4093 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liliputella Jałoszyński, 2016, gen. nov." docType="treatment" docVersion="8" lastPageNumber="139" masterDocId="FFD5D71AFFBB2B22FFD93159FFEE8558" masterDocTitle="Liliputella, a new genus of microscopic, wingless and blind Scydmaenini of the Bismarck Islands (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="142" masterPageNumber="135" pageNumber="136" updateTime="1698185737167" updateUser="plazi">
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<mods:title id="999834A5BA880ADFAD1DF13A8A4762F7">Liliputella, a new genus of microscopic, wingless and blind Scydmaenini of the Bismarck Islands (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="174524767DBB3F9E35F724C940AD53C8" type="personal">
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<mods:namePart id="E8ACF6D4B294266EFC7CBB228D75EC40">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="B3EDA1A2D342E17F068B42A9ADD423EC">Zootaxa</mods:title>
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<mods:part id="3B7BC25041E5C5DC28754E37574D2FBD">
<mods:date id="702FBE99B7B1564F4ED54C6A2534C674">2016</mods:date>
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<mods:number id="1BB27FD105D4DCB4A0439E90A78B1B9F">4093</mods:number>
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<mods:identifier id="1245AF8537D431005EBB3806402A8C86" type="DOI">10.11646/zootaxa.4093.1.9</mods:identifier>
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<treatment id="03ECAF62FFBA2B26FF4E353BFDE081F8" ID-DOI="http://doi.org/10.5281/zenodo.6082684" ID-GBIF-Taxon="119208903" ID-Zenodo-Dep="6082684" LSID="urn:lsid:plazi:treatment:03ECAF62FFBA2B26FF4E353BFDE081F8" httpUri="http://treatment.plazi.org/id/03ECAF62FFBA2B26FF4E353BFDE081F8" lastPageId="4" lastPageNumber="139" pageId="1" pageNumber="136">
<subSubSection id="C35F4DFFFFBA2B23FF4E353BFCF781E7" pageId="1" pageNumber="136" type="nomenclature">
<paragraph id="8BFA1E74FFBA2B23FF4E353BFE678125" blockId="1.[151,393,1122,1149]" box="[151,393,1122,1149]" pageId="1" pageNumber="136">
<heading id="D0B2A918FFBA2B23FF4E353BFE678125" bold="true" box="[151,393,1122,1149]" fontSize="11" level="1" pageId="1" pageNumber="136" reason="1">
<emphasis id="B931C266FFBA2B23FF4E353BFE678125" bold="true" box="[151,393,1122,1149]" pageId="1" pageNumber="136">
<taxonomicName id="4C4565F7FFBA2B23FF4E353BFEFA8124" ID-CoL="8XMCY" box="[151,276,1122,1148]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931C266FFBA2B23FF4E353BFEFA8124" bold="true" box="[151,276,1122,1148]" italics="true" pageId="1" pageNumber="136">Liliputella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2027F1DFFBA2B23FEC5353AFE678125" box="[284,393,1123,1149]" pageId="1" pageNumber="136" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA1E74FFBA2B23FF4E35F3FCF781E7" blockId="1.[151,793,1193,1215]" box="[151,793,1193,1215]" pageId="1" pageNumber="136">
<typeStatus id="54FEA0D6FFBA2B23FF4E35F3FF2781E7" box="[151,201,1194,1215]" pageId="1" pageNumber="136">Type</typeStatus>
species:
<taxonomicName id="4C4565F7FFBA2B23FEFF35F0FDF381E7" box="[294,541,1193,1215]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="microscopica" status="sp. nov.">
<emphasis id="B931C266FFBA2B23FEFF35F0FDF381E7" box="[294,541,1193,1215]" italics="true" pageId="1" pageNumber="136">Liliputella microscopica</emphasis>
</taxonomicName>
<emphasis id="B931C266FFBA2B23FDFA35F0FDB281E6" bold="true" box="[547,604,1193,1214]" pageId="1" pageNumber="136">
<taxonomicNameLabel id="A2027F1DFFBA2B23FDFA35F0FDB281E6" box="[547,604,1193,1214]" pageId="1" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(here designated).
</paragraph>
</subSubSection>
<subSubSection id="C35F4DFFFFBA2B23FF4E35B5FD9C80E0" pageId="1" pageNumber="136" type="diagnosis">
<paragraph id="8BFA1E74FFBA2B23FF4E35B5FEFC8014" blockId="1.[151,1437,1260,2005]" pageId="1" pageNumber="136">
<emphasis id="B931C266FFBA2B23FF4E35B5FEFF805D" bold="true" box="[151,273,1260,1285]" pageId="1" pageNumber="136">Diagnosis.</emphasis>
A genus of
<taxonomicName id="4C4565F7FFBA2B23FE7635B5FDAF805D" box="[431,577,1260,1285]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
differing from all other taxa of this tribe in large ventral cavity behind mouthparts; pronotum and elytra lacking any traces of pits and impressions; and aedeagus with slender, free parameres.
</paragraph>
<paragraph id="8BFA1E74FFBA2B23FF1E340EFD9C80E0" blockId="1.[151,1437,1260,2005]" pageId="1" pageNumber="136">
<emphasis id="B931C266FFBA2B23FF1E340EFE568028" bold="true" box="[199,440,1367,1392]" pageId="1" pageNumber="136">General body shape</emphasis>
(
<figureCitation id="137E02F1FFBA2B23FE11340EFDEA8028" box="[456,516,1367,1392]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="1" pageNumber="136">Fig 1</figureCitation>
) elongate, body strongly flattened, distinctly but not deeply constricted between head and pronotum and between pronotum and elytra; appendages short and robust; vestiture composed of short setae; pigmentation of cuticle light brown.
</paragraph>
</subSubSection>
<subSubSection id="C35F4DFFFFBA2B26FF1E349DFC8E86B4" lastPageId="4" lastPageNumber="139" pageId="1" pageNumber="136" type="description">
<paragraph id="8BFA1E74FFBA2B23FF1E349DFD61821D" blockId="1.[151,1437,1260,2005]" pageId="1" pageNumber="136">
<emphasis id="B931C266FFBA2B23FF1E349DFE868085" bold="true" box="[199,360,1476,1501]" pageId="1" pageNumber="136">Head capsule</emphasis>
(
<figureCitation id="137E02F1FFBA2B23FEA1349DFE2F8084" box="[376,449,1476,1501]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="1" pageNumber="136">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFBA2B23FE08349DFE318084" box="[465,479,1476,1500]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">3</figureCitation>
) divided by occipital constriction into large anterior and small posterior part ('neck' region), posterior part retracted into pronotum. 'Neck' region distinctly narrower than vertex, short and subcylindrical; narrowest site of occipital constriction (
<figureCitation id="137E02F1FFBA2B23FD273755FCAC837C" box="[766,834,1548,1573]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFBA2B23FC973754FC9B837C" box="[846,885,1549,1572]" italics="true" pageId="1" pageNumber="136">occ</emphasis>
) much wider than half HW. Anterior part of head (
<figureCitation id="137E02F1FFBA2B23FF463776FF0B8310" box="[159,229,1583,1608]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="1" pageNumber="136">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFBA2B23FF2A3769FEEF8310" box="[243,257,1584,1608]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">3</figureCitation>
) subtriangular and rounded, broadest in posterior half. Compound eyes absent; vertex posteriorly slightly impressed medially, anteriorly confluent with subtriangular frons; clypeus and mouthparts shifted to the ventral surface of head (
<figureCitation id="137E02F1FFBA2B23FE82372EFE4F83C8" box="[347,417,1655,1680]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
), which is flattened. Antennal insertions located on ventral surface of strongly declined frons, nearly touching at middle. Gular plate (
<figureCitation id="137E02F1FFBA2B23FD4637C5FD0B83EC" box="[671,741,1692,1717]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFBA2B23FD2B37C4FCE083EC" box="[754,782,1693,1716]" italics="true" pageId="1" pageNumber="136">gp</emphasis>
) subtrapezoidal with rounded sides, gular sutures (
<figureCitation id="137E02F1FFBA2B23FA8837C5FA7883EC" box="[1361,1430,1692,1717]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFBA2B23FF4E3798FF5E8380" box="[151,176,1729,1752]" italics="true" pageId="1" pageNumber="136">gs</emphasis>
) distinct, sharply marked and anteriorly meeting at middle. Posterior tentorial pits not visible. Median area between gular plate and mouthparts occupied by large transverse and deep ventral cavity (
<figureCitation id="137E02F1FFBA2B23FB5B37BDFB2883A4" box="[1154,1222,1764,1789]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFBA2B23FB0A37BCFB0283A4" box="[1235,1260,1765,1788]" italics="true" pageId="1" pageNumber="136">vc</emphasis>
) surrounded by dense setae. Head with indistinct microsculpture: faint transverse cells on gular plate and polygonal reticulation of anterolateral area of ventral surface of 'neck'.
</paragraph>
<paragraph id="8BFA1E74FFBA2B20FF1E3616FA6D843D" blockId="1.[151,1437,1260,2005]" lastBlockId="2.[151,1436,151,752]" lastPageId="2" lastPageNumber="137" pageId="1" pageNumber="136">
<emphasis id="B931C266FFBA2B23FF1E3616FEB88230" bold="true" box="[199,342,1871,1896]" pageId="1" pageNumber="136">Mouthparts</emphasis>
(
<figureCitation id="137E02F1FFBA2B23FEBE3616FE458230" box="[359,427,1871,1896]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
) in the studied specimen only partly exposed, details of clypeus, labrum, mandibles and prementum not visible because of unusual ventral position of mouthparts and tightly closed mandibles. Submentum strongly declining into ventral cavity, not visible in ventral view, but its position marked by pair of setae, which in
<taxonomicName id="4C4565F7FFBA2B23FF4E36E5FED4828D" box="[151,314,1980,2005]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
are typically inserted near middle or in anterior region of submentum. Mentum (
<figureCitation id="137E02F1FFBA2B23FACA36E5FAB0828C" box="[1299,1374,1980,2005]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFBA2B23FAA936E4FA7C828C" box="[1392,1426,1981,2004]" italics="true" pageId="1" pageNumber="136">mn</emphasis>
) subhexagonal and elongate, broadest in front of middle. Prementum largely obscured by maxillae; labial palps seem minute and their insertions broadly separated; cardo (
<figureCitation id="137E02F1FFB92B20FCE831E5FC98858C" box="[817,886,188,213]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FC5A31E5FC70858D" box="[899,926,188,213]" italics="true" pageId="2" pageNumber="137">cd</emphasis>
) transverse; basistipes (
<figureCitation id="137E02F1FFB92B20FB7531E5FB1E858C" box="[1196,1264,188,213]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FB2731E5FAF1858D" box="[1278,1311,188,213]" italics="true" pageId="2" pageNumber="137">bst</emphasis>
) small and triangular, with one seta; mediostipes (
<figureCitation id="137E02F1FFB92B20FD903186FD6085A0" box="[585,654,223,248]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FD4231B8FD2C85A0" box="[667,706,225,248]" italics="true" pageId="2" pageNumber="137">mst</emphasis>
) elongate; lacinia and galea (
<figureCitation id="137E02F1FFB92B20FBD53186FBBF85A0" box="[1036,1105,223,248]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FB873186FB6E85A0" box="[1118,1152,223,248]" italics="true" pageId="2" pageNumber="137">lac</emphasis>
,
<emphasis id="B931C266FFB92B20FB573186FB5C85A0" box="[1166,1202,223,248]" italics="true" pageId="2" pageNumber="137">gal</emphasis>
) elongate, each with dense distal setae; palpifer (
<figureCitation id="137E02F1FFB92B20FE01305DFDF18444" box="[472,543,260,285]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FDF7305DFDBF8444" box="[558,593,260,284]" italics="true" pageId="2" pageNumber="137">ppf</emphasis>
) large, with short seta. Maxillary palps (
<figureCitation id="137E02F1FFB92B20FBF2305DFB9C8444" box="[1067,1138,260,285]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 3</figureCitation>
;
<emphasis id="B931C266FFB92B20FB58305CFB598444" box="[1153,1207,261,284]" italics="true" pageId="2" pageNumber="137">mxpi</emphasis>
) short, palpomere I slightly longer than broad; palpomere II clavate, broad and only about 2.5× as long as broad; palpomere III pedunculate, broad and about twice as long as broad, broadest near middle; palpomere IV minute, dome-shaped.
</paragraph>
<paragraph id="8BFA1E74FFB92B20FF1E3036FAD484F5" blockId="2.[151,1436,151,752]" pageId="2" pageNumber="137">
<emphasis id="B931C266FFB92B20FF1E3036FED484D0" bold="true" box="[199,314,367,392]" pageId="2" pageNumber="137">Antennae</emphasis>
(
<figureCitation id="137E02F1FFB92B20FE943036FE7984D0" box="[333,407,367,392]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="2" pageNumber="137">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFB92B20FE703029FE3684D0" box="[425,472,368,392]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">3, 4</figureCitation>
) moderately slender and moderately long, gradually thickened distally; scape with shallow and broad ventral and dorsal apical emarginations; flagellomeres relatively compactly assembled.
</paragraph>
<paragraph id="8BFA1E74FFB92B20FF1E30EEFDC987F0" blockId="2.[151,1436,151,752]" pageId="2" pageNumber="137">
<emphasis id="B931C266FFB92B20FF1E30EEFEAC8488" bold="true" box="[199,322,439,464]" pageId="2" pageNumber="137">Prothorax</emphasis>
(
<figureCitation id="137E02F1FFB92B20FE8830EEFE778488" box="[337,409,439,464]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="2" pageNumber="137">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFB92B20FE7030EEFE3A8488" box="[425,468,439,464]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">56</figureCitation>
) in dorsal view (
<figureCitation id="137E02F1FFB92B20FD4E30EEFD368488" box="[663,728,439,464]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="2" pageNumber="137">Fig. 1</figureCitation>
) indistinctly bell-shaped, broadest in front of middle; anterior corners indistinct, posterior corners obtuse-angled and blunt. Pronotum lacking any carinae, pits, grooves or impressions. Prosternum (
<figureCitation id="137E02F1FFB92B20FE1B30A6FDE68740" box="[450,520,511,536]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
) nearly half as long as pronotum, with basisternal part (
<figureCitation id="137E02F1FFB92B20FB4230A6FB0D8740" box="[1179,1251,511,536]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
;
<emphasis id="B931C266FFB92B20FB2A30A6FAFA8740" box="[1267,1300,511,536]" italics="true" pageId="2" pageNumber="137">bst</emphasis>
) very long, several times longer than coxal part and distinctly demarcated from procoxal cavities (
<figureCitation id="137E02F1FFB92B20FB4A337DFB328765" box="[1171,1244,548,573]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
;
<emphasis id="B931C266FFB92B20FB34337CFAFA8764" box="[1261,1300,549,572]" italics="true" pageId="2" pageNumber="137">pcc</emphasis>
) by double carina; prosternal intercoxal process (
<figureCitation id="137E02F1FFB92B20FDE2331EFD918738" box="[571,639,583,608]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
;
<emphasis id="B931C266FFB92B20FD553310FD5D8738" box="[652,691,585,608]" italics="true" pageId="2" pageNumber="137">psp</emphasis>
) subtriangular and weakly elevated; procoxal sockets (
<figureCitation id="137E02F1FFB92B20FAC5331EFA8C8738" box="[1308,1378,583,608]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
;
<emphasis id="B931C266FFB92B20FAB43310FA7C8738" box="[1389,1426,585,608]" italics="true" pageId="2" pageNumber="137">pcs</emphasis>
) broadly open. Hypomera (
<figureCitation id="137E02F1FFB92B20FE193335FDE887DD" box="[448,518,620,645]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 6</figureCitation>
;
<emphasis id="B931C266FFB92B20FDCD3335FDC187DD" box="[532,559,620,645]" italics="true" pageId="2" pageNumber="137">hy</emphasis>
) elongate and confluent with sides of pronotum, lacking hypomeral ridges or grooves; notosternal sutures absent.
</paragraph>
<paragraph id="8BFA1E74FFB92B20FF1E33EDFD2E87A8" blockId="2.[151,1436,151,752]" pageId="2" pageNumber="137">
<emphasis id="B931C266FFB92B20FF1E33EDFEB88795" bold="true" box="[199,342,692,717]" pageId="2" pageNumber="137">Mesothorax</emphasis>
(
<figureCitation id="137E02F1FFB92B20FEBC33EDFE248794" box="[357,458,692,717]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Figs 5, 7</figureCitation>
). Mesonotum with remarkably large, triangular and strongly transverse scutellum (
<figureCitation id="137E02F1FFB92B20FAB633EDFF4987A8" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="2" pageNumber="137">Fig. 5</figureCitation>
;
<emphasis id="B931C266FFB92B20FF6D338EFF0F87A8" box="[180,225,727,752]" italics="true" pageId="2" pageNumber="137">scl2</emphasis>
), only its tip visible between elytral bases.
</paragraph>
<caption id="DF3A4EFCFFB92B20FF4E362FFF0C82F3" httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="2" pageNumber="137" targetBox="[163,1436,833,1886]" targetPageId="2">
<paragraph id="8BFA1E74FFB92B20FF4E362FFF0C82F3" blockId="2.[151,1436,1910,1963]" pageId="2" pageNumber="137">
<emphasis id="B931C266FFB92B20FF4E362FFED482D4" bold="true" box="[151,314,1910,1933]" pageId="2" pageNumber="137">FIGURES 12.</emphasis>
<taxonomicName id="4C4565F7FFB92B20FE98362EFDD782D4" box="[321,569,1911,1932]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="microscopica">
<emphasis id="B931C266FFB92B20FE98362EFDD782D4" box="[321,569,1911,1932]" italics="true" pageId="2" pageNumber="137">Liliputella microscopica</emphasis>
</taxonomicName>
<emphasis id="B931C266FFB92B20FDE6362EFD3082D4" bold="true" box="[575,734,1910,1932]" pageId="2" pageNumber="137">gen. &amp; sp. nov.</emphasis>
Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow).
</paragraph>
</caption>
<caption id="DF3A4EFCFFB82B21FF4E365BFD3382B7" httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="3" pageNumber="138" targetBox="[185,1380,194,1749]" targetPageId="3">
<paragraph id="8BFA1E74FFB82B21FF4E365BFD3382B7" blockId="3.[151,1436,1794,2031]" pageId="3" pageNumber="138">
<emphasis id="B931C266FFB82B21FF4E365BFED5824F" bold="true" box="[151,315,1794,1817]" pageId="3" pageNumber="138">FIGURES 39.</emphasis>
<taxonomicName id="4C4565F7FFB82B21FE9B365AFDD58240" box="[322,571,1795,1816]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="microscopica">
<emphasis id="B931C266FFB82B21FE9B365AFDD58240" box="[322,571,1795,1816]" italics="true" pageId="3" pageNumber="138">Liliputella microscopica</emphasis>
</taxonomicName>
gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl2, mesocutellum; st38, sternite III VIII; vc, ventral cavity.
</paragraph>
</caption>
<paragraph id="8BFA1E74FFBF2B26FF1E31CEFC8C843D" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
Mesoventrite (
<figureCitation id="137E02F1FFBF2B26FEB331CEFE4085E8" box="[362,430,151,176]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
) short, much broader than long, lacking demarcated anterior ridge; procoxal rests weakly defined and hidden under procoxae (
<figureCitation id="137E02F1FFBF2B26FD9131E5FD7F858C" box="[584,657,188,213]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
); mesoventral intercoxal process (
<figureCitation id="137E02F1FFBF2B26FBFA31E5FB82858C" box="[1059,1132,188,213]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FBA431E4FB59858C" box="[1149,1207,189,212]" italics="true" pageId="4" pageNumber="139">msvp</emphasis>
) moderately broad, nearly parallel-sided, weakly elevated; mesocoxal sockets (
<figureCitation id="137E02F1FFBF2B26FC943186FC7985AF" box="[845,919,223,248]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FC7131B8FC3085A0" box="[936,990,225,248]" italics="true" pageId="4" pageNumber="139">mscs</emphasis>
) located ventromesally on mesocoxal projections (
<figureCitation id="137E02F1FFBF2B26FEFC305DFE828444" box="[293,364,260,285]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FEA2305CFE478444" box="[379,425,261,284]" italics="true" pageId="4" pageNumber="139">mcp</emphasis>
) and in ventral view partly exposed; mesofurcal foveae (
<figureCitation id="137E02F1FFBF2B26FBE4305DFB6D8444" box="[1085,1155,260,285]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FB4B305DFB2E8444" box="[1170,1216,260,284]" italics="true" pageId="4" pageNumber="139">msff</emphasis>
) small and located near anterior margins of mesocoxal cavities (
<figureCitation id="137E02F1FFBF2B26FD64307EFCE78467" box="[701,777,295,320]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FCC73070FCB88418" box="[798,854,297,320]" italics="true" pageId="4" pageNumber="139">mscc</emphasis>
). Prepectus (
<figureCitation id="137E02F1FFBF2B26FC20307EFBAB8467" box="[1017,1093,295,320]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FB803070FB938418" box="[1113,1149,297,320]" italics="true" pageId="4" pageNumber="139">pre</emphasis>
) large. Median area of mesoventrite in front of mesocoxae with polygonal reticulation.
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E3036FDAF8764" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E3036FEBA84D0" bold="true" box="[199,340,367,392]" pageId="4" pageNumber="139">Metathorax</emphasis>
. Metaventrite (
<figureCitation id="137E02F1FFBF2B26FDC93036FDB584D0" box="[528,603,367,392]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
) shorter than mesoventrite, strongly transverse, anteriorly fused with mesoventrite, lateral margins arcuate, lateral (admetacoxal) parts of posterior margin strongly concave, at middle posterior margin broadly concave and forming broad and short metaventral intercoxal process (
<figureCitation id="137E02F1FFBF2B26FB2230EEFAAA8497" box="[1275,1348,439,464]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FA8C30E0FA658488" box="[1365,1419,441,464]" italics="true" pageId="4" pageNumber="139">mtvp</emphasis>
). Metanepisterna concealed under elytra. Anteriorly and laterally metaventrite covered with polygonal reticulation; pattern of connected and raised edges of cells forms pair of indistinct lateral carinae extending from each mesocoxal projection up to metacoxa.
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E331EFA818738" blockId="4.[151,1437,151,1184]" box="[199,1391,583,608]" pageId="4" pageNumber="139">
Metafurca (metendosternite) (
<figureCitation id="137E02F1FFBF2B26FDCF331EFDB78738" box="[534,601,583,608]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
) with broadly separated insertions of lateral furcal arms (
<figureCitation id="137E02F1FFBF2B26FB05331EFACF8707" box="[1244,1313,583,608]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FAF5331EFAB08738" box="[1324,1374,583,608]" italics="true" pageId="4" pageNumber="139">lmfa</emphasis>
).
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E3335FE4987F0" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E3335FEFA87DD" bold="true" box="[199,276,620,645]" pageId="4" pageNumber="139">Elytra</emphasis>
(
<figureCitation id="137E02F1FFBF2B26FEFE3335FE9D87DC" box="[295,371,620,645]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="4" pageNumber="139">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFBF2B26FE5E3335FE5987DC" box="[391,439,620,645]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">5, 7</figureCitation>
) elongate, lacking basal foveae, impressions or humeral calli, with distinct humeral denticles (
<figureCitation id="137E02F1FFBF2B26FED033D6FE8087FF" box="[265,366,655,680]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Figs 5, 7</figureCitation>
;
<emphasis id="B931C266FFBF2B26FEA233D6FE7987F0" box="[379,407,655,680]" italics="true" pageId="4" pageNumber="139">hd</emphasis>
).
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E33EDFDE78795" blockId="4.[151,1437,151,1184]" box="[199,521,692,717]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E33EDFE5C8795" bold="true" box="[199,434,692,717]" pageId="4" pageNumber="139">Metathoracic wings</emphasis>
absent.
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E338EFC838660" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E338EFED287A8" bold="true" box="[199,316,727,752]" pageId="4" pageNumber="139">Abdomen</emphasis>
(
<figureCitation id="137E02F1FFBF2B26FE92338EFE6087A8" box="[331,398,727,752]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Fig. 7</figureCitation>
) longer than meso- and metaventrite together; sternite III (first visible) nearly as long as IV VII combined, its anterior intermetacoxal region distinctly reticulate; suture between sternites VII and VIII distinct. Pygidium not exposed; propygidium lacking median impression.
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E321DFBD786FC" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E321DFF118605" bold="true" box="[199,255,836,861]" pageId="4" pageNumber="139">Legs</emphasis>
(
<figureCitation id="137E02F1FFBF2B26FECA321DFEB08604" box="[275,350,836,861]" captionStart="FIGURES 1 2" captionStartId="2.[151,264,1910,1933]" captionTargetBox="[163,1436,833,1886]" captionTargetId="figure@2.[151,1436,833,1886]" captionTargetPageId="2" captionText="FIGURES 1 2. Liliputella microscopica gen. &amp; sp. nov. Dorsal habitus of holotype male (1) and distribution (2; indicated by arrow)." httpUri="https://zenodo.org/record/265879/files/figure.png" pageId="4" pageNumber="139">Figs 1</figureCitation>
,
<figureCitation id="137E02F1FFBF2B26FEA8321DFE738605" box="[369,413,836,861]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">67</figureCitation>
) short and robust; pro- and mesocoxae subconical; metacoxae oval and transverse; all trochanters subtriangular and short; all femora gradually and weakly clavate; tibiae flattened, broadened distally; tarsi strikingly short, tarsomeres I and V elongate, IIIV subquadrate or transverse.
</paragraph>
<paragraph id="8BFA1E74FFBF2B26FF1E32F6FC8E86B4" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E32F6FED58690" bold="true" box="[199,315,943,968]" pageId="4" pageNumber="139">Aedeagus</emphasis>
(
<figureCitation id="137E02F1FFBF2B26FE9032F6FE438690" box="[329,429,943,968]" captionStart="FIGURES 3 9" captionStartId="3.[151,264,1794,1817]" captionTargetBox="[185,1380,194,1749]" captionTargetId="figure@3.[177,1391,193,1758]" captionTargetPageId="3" captionText="FIGURES 3 9. Liliputella microscopica gen. &amp; sp. nov. Head in ventral view (3); left antenna in ventral view (4), prothorax and elytral base in dorsal view (5), prothorax in ventral view (6), pterothorax in ventral view (7), aedeagus in ventral (8) and lateral (9) views. Abbreviations: bst, basisternal part of prosternum; bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hd, humeral denticle; lac, lacinia; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mediostipes; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp, maxillary palp; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; ppf, palpifer; pre, prepectus; psp, prosternal process; scl 2, mesocutellum; st 3 8, sternite III VIII; vc, ventral cavity." httpUri="https://zenodo.org/record/265880/files/figure.png" pageId="4" pageNumber="139">Figs 89</figureCitation>
) with symmetrical median lobe, elongate; diaphragm located sub-basally on ventral wall; flagellum looped; parameres free and slender, with apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C35F4DFFFFBF2B26FF1E32AEFD17816D" pageId="4" pageNumber="139" type="etymology">
<paragraph id="8BFA1E74FFBF2B26FF1E32AEFD17816D" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E32AEFEA08148" bold="true" box="[199,334,1015,1040]" pageId="4" pageNumber="139">Etymology.</emphasis>
<taxonomicName id="4C4565F7FFBF2B26FE8E32AEFE228148" box="[343,460,1015,1040]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931C266FFBF2B26FE8E32AEFE228148" box="[343,460,1015,1040]" italics="true" pageId="4" pageNumber="139">Liliputella</emphasis>
</taxonomicName>
is derived from the Liliput nation of
<emphasis id="B931C266FFBF2B26FC5932AEFBA48148" box="[896,1098,1015,1040]" italics="true" pageId="4" pageNumber="139">Gulliver's Travels</emphasis>
by Jonathan Swift; the name refers to the tiny body of this beetle. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C35F4DFFFFBF2B26FF1E3566FDE081F8" pageId="4" pageNumber="139" type="discussion">
<paragraph id="8BFA1E74FFBF2B26FF1E3566FDE081F8" blockId="4.[151,1437,151,1184]" pageId="4" pageNumber="139">
<emphasis id="B931C266FFBF2B26FF1E3566FED58100" bold="true" box="[199,315,1087,1112]" pageId="4" pageNumber="139">Remarks.</emphasis>
The structure of maxillary palpomeres clearly places
<taxonomicName id="4C4565F7FFBF2B26FC1D3566FBD78100" box="[964,1081,1087,1112]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931C266FFBF2B26FC1D3566FBD78100" box="[964,1081,1087,1112]" italics="true" pageId="4" pageNumber="139">Liliputella</emphasis>
</taxonomicName>
in
<taxonomicName id="4C4565F7FFBF2B26FBBE3566FB138100" box="[1127,1277,1087,1112]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
. Indeed, in a preliminary phylogenetic analysis,
<taxonomicName id="4C4565F7FFBF2B26FDED353DFD478125" box="[564,681,1124,1149]" class="Insecta" family="Staphylinidae" genus="Liliputella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B931C266FFBF2B26FDED353DFD478125" box="[564,681,1124,1149]" italics="true" pageId="4" pageNumber="139">Liliputella</emphasis>
</taxonomicName>
was placed as a sister group to
<taxonomicName id="4C4565F7FFBF2B26FB98353DFB088125" box="[1089,1254,1124,1149]" class="Insecta" family="Staphylinidae" genus="Pseudoeudesis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B931C266FFBF2B26FB98353DFB088125" box="[1089,1254,1124,1149]" italics="true" pageId="4" pageNumber="139">Pseudoeudesis</emphasis>
</taxonomicName>
, clearly within
<taxonomicName id="4C4565F7FFBF2B26FF4E35DEFEC781F8" box="[151,297,1159,1184]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="139" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
(results not shown).
</paragraph>
</subSubSection>
</treatment>
</document>