treatments-xml/data/03/EC/5C/03EC5C3E1C480C63FF7BFF38FA18FCCB.xml
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<document id="EE41B94230C6A1459B386ACEC0E0E0FD" ID-DOI="10.11646/zootaxa.5361.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10144864" ID-ZooBank="AFEFD8C9-ABC1-4793-8B83-49FB7B526389" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1700163474722" checkinUser="plazi" docAuthor="Koreeda, Reo, Maeda, Ken &amp; Motomura, Hiroyuki" docDate="2023" docId="03EC5C3E1C480C63FF7BFF38FA18FCCB" docLanguage="en" docName="zootaxa.5361.3.5.pdf" docOrigin="Zootaxa 5361 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.3.5/52197" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luciogobius undefined-7" docType="treatment" docVersion="4" lastPageNumber="406" masterDocId="FFD524461C470C73FFECFFAFFF84FFCD" masterDocTitle="A new subtropical species of goby of the genus Luciogobius (Gobiidae) from southwestern Japan" masterLastPageNumber="408" masterPageNumber="390" pageNumber="405" updateTime="1700497287600" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E5DB8C2E139889929415C3C43E59DBA3">A new subtropical species of goby of the genus Luciogobius (Gobiidae) from southwestern Japan</mods:title>
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<mods:namePart id="CA0DA369393CC71C4101F8B8F7FB2313">Koreeda, Reo</mods:namePart>
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<mods:affiliation id="2D53E117502FABF94B51B17FEFD16980">The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 - 0065, Japan</mods:affiliation>
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<mods:namePart id="FECDF591248F605FBE0F79C5BCBBCF8A">Maeda, Ken</mods:namePart>
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<mods:affiliation id="BD4351C1685DC47DF8E734E7A555BD3A">Marine Eco-Evo-Devo Unit, Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 - 0495, Japan</mods:affiliation>
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<mods:namePart id="BF28B1FC62D37201CF9B7EC9798F5097">Motomura, Hiroyuki</mods:namePart>
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<mods:affiliation id="E776CBB85F97324265C9CBFF7B4ADFFC">The Kagoshima University Museum, 1 - 21 - 30 Korimoto, Kagoshima 890 - 0065, Japan</mods:affiliation>
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<paragraph id="8BFAED281C480C7CFF7BFF38FD60FF7C" blockId="15.[151,771,151,249]" box="[151,740,151,178]" pageId="15" pageNumber="405">
<heading id="D0B25A441C480C7CFF7BFF38FD60FF7C" bold="true" box="[151,740,151,178]" fontSize="11" level="1" pageId="15" pageNumber="405" reason="1">
<emphasis id="B931313A1C480C7CFF7BFF38FD60FF7C" bold="true" box="[151,740,151,178]" pageId="15" pageNumber="405">
<taxonomicName id="4C4596AB1C480C7CFF7BFF38FEEAFF7C" box="[151,366,151,178]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C480C7CFF7BFF38FEA9FF7F" bold="true" box="[151,301,151,178]" italics="true" pageId="15" pageNumber="405">Luciogobius</emphasis>
sp. 7
</taxonomicName>
<emphasis id="B931313A1C480C7CFE99FF38FE3DFF7C" bold="true" box="[373,441,151,177]" italics="true" pageId="15" pageNumber="405">sensu</emphasis>
<bibRefCitation id="EFD490D91C480C7CFE2DFF38FD60FF7C" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[449,740,151,178]" pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFDB8FF38FD0BFF7F" bold="true" box="[596,655,151,178]" italics="true" pageId="15" pageNumber="405">et al.</emphasis>
(2019)
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFAED281C480C7CFF7BFF14FC87FF18" blockId="15.[151,771,151,249]" box="[151,771,187,214]" pageId="15" pageNumber="405">[Standard Japanese name: Hauchiwa-yari-mimizuhaze]</paragraph>
</subSubSection>
<subSubSection id="C35FBEA31C480C7CFF7BFF70FEE0FF34" box="[151,356,223,249]" pageId="15" pageNumber="405" type="description">
<paragraph id="8BFAED281C480C7CFF7BFF70FEE0FF34" blockId="15.[151,771,151,249]" box="[151,356,223,249]" pageId="15" pageNumber="405">
<figureCitation id="137EF1AD1C480C7CFF7BFF70FE82FF34" box="[151,262,223,249]" captionStart="FIGURE 10" captionStartId="15.[152,255,1792,1816]" captionTargetBox="[151,1436,952,1768]" captionTargetId="figure-354@15.[151,1436,952,1768]" captionTargetPageId="15" captionText="FIGURE 10. Fresh specimens of Luciogobius sp. 7 sensu Shibukawa et al. (2019) from Kochi Prefecture, Shikoku, Japan. A: KAUMI. 159033, male, 51.3 mm SL; B: KAUMI. 159029, female, 33.2 mm SL; C: KAUMI. 159028, female, 32.1 mm SL. Photographs taken immediately after fixation." figureDoi="http://doi.org/10.5281/zenodo.10144884" httpUri="https://zenodo.org/record/10144884/files/figure.png" pageId="15" pageNumber="405">Figure 10</figureCitation>
;
<tableCitation id="C6C7D8931C480C7CFEFEFF70FEE0FF34" box="[274,356,223,249]" captionStart="TABLE 2" captionStartId="11.[151,244,737,763]" captionTargetBox="[162,2003,778,1447]" captionTargetPageId="11" captionText="TABLE 2. Frequency distribution of meristic counts of the Luciogobius platycephalus complex (this study only)." httpUri="http://table.plazi.org/id/DF3ABDA01C4C0C78FF7BFD4EFAD2FD36" pageId="15" pageNumber="405" tableUuid="DF3ABDA01C4C0C78FF7BFD4EFAD2FD36">Table 2</tableCitation>
</paragraph>
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<treatmentCitationGroup id="AB55CA061C480C7CFF7BFE8BFBD4FEF1" box="[151,1104,291,316]" pageId="15" pageNumber="405">
<taxonomicName id="4C4596AB1C480C7CFF7BFE8BFECAFEF6" box="[151,334,292,316]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C480C7CFF7BFE8BFE90FEF1" box="[151,276,292,316]" italics="true" pageId="15" pageNumber="405">Luciogobius</emphasis>
sp. 7
</taxonomicName>
:
<treatmentCitation id="0AE4CB391C480C7CFEB4FE8BFDE7FEF6" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[344,611,291,316]" page="67" pageId="15" pageNumber="405" year="2019">
<bibRefCitation id="EFD490D91C480C7CFEB4FE8BFDE7FEF6" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[344,611,291,316]" pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFE23FE8BFE78FEF1" box="[463,508,292,316]" italics="true" pageId="15" pageNumber="405">et al</emphasis>
, 2019: 67
</bibRefCitation>
</treatmentCitation>
, figs. 19B, 20EG, 21C (
<collectingRegion id="498123CA1C480C7CFC8FFE8BFC79FEF1" box="[867,1021,292,316]" country="Japan" name="Shizuoka" pageId="15" pageNumber="405">Shizuoka Pref.</collectingRegion>
,
<collectingCountry id="F352ADB81C480C7CFBEBFE8BFBC1FEF1" box="[1031,1093,292,316]" name="Japan" pageId="15" pageNumber="405">Japan</collectingCountry>
).
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</paragraph>
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<emphasis id="B931313A1C480C7CFF7BFEC6FE1EFE4E" bold="true" box="[151,410,361,387]" pageId="15" pageNumber="405">Examined specimens.</emphasis>
<materialsCitation id="3B2DE7751C480C7CFE4FFEC5FA99FE49" box="[419,1309,361,389]" collectionCode="KAUM" location="Japan." pageId="15" pageNumber="405" specimenCount="10" stateProvince="Shizuoka">
<specimenCount id="9D4326A11C480C7CFE4FFEC5FDB9FE49" box="[419,573,362,388]" count="10" pageId="15" pageNumber="405" type="generic">10 specimens</specimenCount>
(
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SL) from
<location id="8E9ABBF31C480C7CFC82FEC5FC32FE49" LSID="urn:lsid:plazi:treatment:03EC5C3E1C480C63FF7BFF38FA18FCCB:8E9ABBF31C480C7CFC82FEC5FC32FE49" box="[878,950,362,388]" name="Japan." pageId="15" pageNumber="405" stateProvince="Shizuoka">Japan.</location>
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<collectingRegion id="498123CA1C480C7CFC52FEC6FB32FE4E" box="[958,1206,361,387]" country="Japan" name="Shizuoka" pageId="15" pageNumber="405">Shizuoka Prefecture</collectingRegion>
:
</emphasis>
<collectionCode id="ED5475ED1C480C7CFB28FEC5FA99FE49" box="[1220,1309,362,388]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
</materialsCitation>
I. 139315,
<quantity id="4CBD40CD1C480C7CFF7BFE21FF7AFE65" box="[151,254,398,424]" metricMagnitude="-2" metricUnit="m" metricValue="4.31" pageId="15" pageNumber="405" unit="mm" value="43.1">43.1 mm</quantity>
SL, collected by
<collectionCode id="ED5475ED1C480C7CFE21FE21FE67FE65" box="[461,483,398,424]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="405">R</collectionCode>
.
<materialsCitation id="3B2DE7751C480C7CFE1CFE21FD4AFE65" box="[496,718,398,424]" collectionCode="KAUM" collectorName="Yamashita." pageId="15" pageNumber="405" specimenCount="1" stateProvince="Shizuoka">
<collectorName id="26B088FE1C480C7CFE1CFE21FDE8FE65" box="[496,620,398,424]" pageId="15" pageNumber="405">Yamashita.</collectorName>
<collectionCode id="ED5475ED1C480C7CFD99FE21FD4AFE65" box="[629,718,398,424]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
</materialsCitation>
I. 157744157747,
<materialsCitation id="3B2DE7751C480C7CFC56FE21FDB1FE01" collectingDate="2021-06-10" collectionCode="KAUM" collectorName="H. Saito" pageId="15" pageNumber="405" specimenCount="4" stateProvince="Shizuoka">
<specimenCount id="9D4326A11C480C7CFC56FE21FBCDFE65" box="[954,1097,398,424]" count="4" pageId="15" pageNumber="405" type="generic">4 specimens</specimenCount>
,
<quantity id="4CBD40CD1C480C7CFBBBFE21FB78FE65" box="[1111,1276,398,425]" metricMagnitude="-2" metricUnit="m" metricValue="4.99" metricValueMax="6.59" metricValueMin="3.3899999999999997" pageId="15" pageNumber="405" unit="mm" value="49.9" valueMax="65.9" valueMin="33.9">33.965.9 mm</quantity>
SL, collected by
<collectorName id="26B088FE1C480C7CFF52FE1DFE9BFE01" box="[190,287,434,460]" pageId="15" pageNumber="405">H. Saito</collectorName>
,
<date id="FFFBCBE81C480C7CFEC2FE1DFE4AFE01" box="[302,462,434,460]" pageId="15" pageNumber="405" value="2021-06-10">
<collectingDate id="EFBF32001C480C7CFEC2FE1DFE4AFE01" box="[302,462,434,460]" pageId="15" pageNumber="405" value="2021-06-10">10 June 2021</collectingDate>
</date>
,
<collectionCode id="ED5475ED1C480C7CFE30FE1DFDB1FE01" box="[476,565,434,460]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
</materialsCitation>
I. 165340,
<quantity id="4CBD40CD1C480C7CFD2CFE1DFCA3FE01" box="[704,807,434,460]" metricMagnitude="-2" metricUnit="m" metricValue="4.93" pageId="15" pageNumber="405" unit="mm" value="49.3">49.3 mm</quantity>
SL,
<collectionCode id="ED5475ED1C480C7CFC88FE1DFC39FE01" box="[868,957,434,460]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
<materialsCitation id="3B2DE7751C480C7CFC27FE1DFCB2FE3D" collectingDate="2022-01-02" collectionCode="KAUM" collectorName="K. Mochizuki" location="Yaizu." pageId="15" pageNumber="405" specimenCount="1" stateProvince="Kochi">
I. 165341,
<quantity id="4CBD40CD1C480C7CFBABFE1DFB2BFE01" box="[1095,1199,434,460]" metricMagnitude="-2" metricUnit="m" metricValue="4.18" pageId="15" pageNumber="405" unit="mm" value="41.8">41.8 mm</quantity>
SL, collected by
<collectorName id="26B088FE1C480C7CFA6DFE1DFE97FE3D" pageId="15" pageNumber="405">K. Mochizuki</collectorName>
,
<date id="FFFBCBE81C480C7CFECCFE79FE21FE3D" box="[288,421,470,496]" pageId="15" pageNumber="405" value="2022-01-02">
<collectingDate id="EFBF32001C480C7CFECCFE79FE21FE3D" box="[288,421,470,496]" pageId="15" pageNumber="405" value="2022-01-02">2 Jan. 2022</collectingDate>
</date>
,
<location id="8E9ABBF31C480C7CFE5CFE79FE7CFE3D" LSID="urn:lsid:plazi:treatment:03EC5C3E1C480C63FF7BFF38FA18FCCB:8E9ABBF31C480C7CFE5CFE79FE7CFE3D" box="[432,504,470,496]" name="Yaizu." pageId="15" pageNumber="405" stateProvince="Kochi">Yaizu.</location>
<emphasis id="B931313A1C480C7CFE13FE7AFD51FE22" bold="true" box="[511,725,469,495]" pageId="15" pageNumber="405">
<collectingRegion id="498123CA1C480C7CFE13FE7AFD54FE22" box="[511,720,469,495]" country="Japan" name="Kochi" pageId="15" pageNumber="405">Kochi Prefecture</collectingRegion>
:
</emphasis>
<collectionCode id="ED5475ED1C480C7CFD31FE79FCB2FE3D" box="[733,822,470,496]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
</materialsCitation>
I. 159028,
<quantity id="4CBD40CD1C480C7CFC50FE79FBA5FE3D" box="[956,1057,470,496]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" pageId="15" pageNumber="405" unit="mm" value="32.1">32.1 mm</quantity>
SL,
<collectionCode id="ED5475ED1C480C7CFBB5FE79FB36FE3D" box="[1113,1202,470,496]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
<materialsCitation id="3B2DE7751C480C7CFB2CFE79FB93FDD9" collectingDate="2021-07-13" collectionCode="KAUM" collectorName="H. Saito" location="Haruno." municipality="Niyodo River" pageId="15" pageNumber="405" specimenCount="1" stateProvince="Kochi">
I. 159029,
<quantity id="4CBD40CD1C480C7CFAD4FE79FA19FE3D" box="[1336,1437,470,496]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="15" pageNumber="405" unit="mm" value="33.2">33.2 mm</quantity>
SL, collected by
<collectorName id="26B088FE1C480C7CFEBBFE55FE32FDD9" box="[343,438,506,532]" pageId="15" pageNumber="405">H. Saito</collectorName>
,
<date id="FFFBCBE81C480C7CFE2EFE55FDD3FDD9" box="[450,599,506,532]" pageId="15" pageNumber="405" value="2021-07-13">
<collectingDate id="EFBF32001C480C7CFE2EFE55FDD3FDD9" box="[450,599,506,532]" pageId="15" pageNumber="405" value="2021-07-13">13 July 2021</collectingDate>
</date>
,
<collectingMunicipality id="6B9E77521C480C7CFD8FFE55FD79FDD9" box="[611,765,506,532]" pageId="15" pageNumber="405">Niyodo River</collectingMunicipality>
mouth,
<location id="8E9ABBF31C480C7CFCB7FE55FC33FDD9" LSID="urn:lsid:plazi:treatment:03EC5C3E1C480C63FF7BFF38FA18FCCB:8E9ABBF31C480C7CFCB7FE55FC33FDD9" box="[859,951,506,532]" municipality="Niyodo River" name="Haruno." pageId="15" pageNumber="405" stateProvince="Kochi">Haruno.</location>
<collectionCode id="ED5475ED1C480C7CFC52FE55FB93FDD9" box="[958,1047,506,532]" country="Japan" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="15" pageNumber="405">KAUM</collectionCode>
</materialsCitation>
<materialsCitation id="3B2DE7751C480C7CFBC9FE55FD7AFDF5" collectingDate="2021-07-22" collectorName="H. Saito" location="Suzaki" municipality="Shinjo River" pageId="15" pageNumber="405" specimenCount="1" stateProvince="Kochi">
I. 159033,
<quantity id="4CBD40CD1C480C7CFB70FE55FA85FDD9" box="[1180,1281,506,533]" metricMagnitude="-2" metricUnit="m" metricValue="5.13" pageId="15" pageNumber="405" unit="mm" value="51.3">51.3 mm</quantity>
SL, collected by
<collectorName id="26B088FE1C480C7CFF56FDB1FE9CFDF5" box="[186,280,542,568]" pageId="15" pageNumber="405">H. Saito</collectorName>
,
<date id="FFFBCBE81C480C7CFEC8FDB1FE3CFDF5" box="[292,440,542,568]" pageId="15" pageNumber="405" value="2021-07-22">
<collectingDate id="EFBF32001C480C7CFEC8FDB1FE3CFDF5" box="[292,440,542,568]" pageId="15" pageNumber="405" value="2021-07-22">22 July 2021</collectingDate>
</date>
,
<collectingMunicipality id="6B9E77521C480C7CFE2FFDB1FDD5FDF5" box="[451,593,542,568]" pageId="15" pageNumber="405">Shinjo River</collectingMunicipality>
mouth,
<location id="8E9ABBF31C480C7CFD42FDB1FD7AFDF5" LSID="urn:lsid:plazi:treatment:03EC5C3E1C480C63FF7BFF38FA18FCCB:8E9ABBF31C480C7CFD42FDB1FD7AFDF5" box="[686,766,542,568]" municipality="Shinjo River" name="Suzaki" pageId="15" pageNumber="405" stateProvince="Kochi">Suzaki</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFAED281C480C7CFF2BFDEEFBCCFD21" blockId="15.[151,1437,361,928]" pageId="15" pageNumber="405">
<emphasis id="B931313A1C480C7CFF2BFDEEFCA1FD96" bold="true" box="[199,805,577,604]" pageId="15" pageNumber="405">Morphological characters of examined specimens.</emphasis>
Total second dorsal-fin rays 1012 (modally 12); total anal-fin rays 1315 (usually 13 or 14); pectoral-fin rays 1316; vertebrae 17 + 2425 = 41 or 42; uppermost 24 rays free on pectoral fin; fewer than 5 pectoral-fin rays branched (central rays branched in specimens&gt;
<quantity id="4CBD40CD1C480C7CFAFAFD25FAE1FD69" box="[1302,1381,650,677]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="405" unit="mm" value="50.0">50 mm</quantity>
SL);
<collectionCode id="ED5475ED1C480C7CFF7BFD01FF56FD05" box="[151,210,686,712]" pageId="15" pageNumber="405">AAA</collectionCode>
longer than half body depth at anus to anal-fin origin; dorsal-fin origin just above or behind anterior 1/3 of anal-fin base; pectoral-fin membrane strongly concave anteriorly; pelvic fins united.
</paragraph>
</subSubSection>
<subSubSection id="C35FBEA31C480C7CFF2BFD5AFDF8FC95" pageId="15" pageNumber="405" type="distribution">
<paragraph id="8BFAED281C480C7CFF2BFD5AFDF8FC95" blockId="15.[151,1437,361,928]" pageId="15" pageNumber="405">
<emphasis id="B931313A1C480C7CFF2BFD5AFEE5FCC2" bold="true" box="[199,353,757,783]" pageId="15" pageNumber="405">Distribution.</emphasis>
Currently known from
<collectingCountry id="F352ADB81C480C7CFD99FD59FD31FCDD" box="[629,693,758,784]" name="Japan" pageId="15" pageNumber="405">Japan</collectingCountry>
[
<collectingRegion id="498123CA1C480C7CFD29FD59FCB7FCDD" box="[709,819,758,784]" country="Japan" name="Shizuoka" pageId="15" pageNumber="405">Shizuoka</collectingRegion>
,
<collectingRegion id="498123CA1C480C7CFCACFD59FC38FCDD" box="[832,956,758,784]" country="Japan" name="Wakayama" pageId="15" pageNumber="405">Wakayama</collectingRegion>
and
<collectingRegion id="498123CA1C480C7CFC1BFD59FBBFFCDD" box="[1015,1083,758,784]" country="Japan" name="Kochi" pageId="15" pageNumber="405">Kochi</collectingRegion>
prefectures (
<bibRefCitation id="EFD490D91C480C7CFB3FFD59FF56FCF9" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFAB7FD58FA0AFCDD" box="[1371,1422,758,784]" italics="true" pageId="15" pageNumber="405">et al</emphasis>
., 2019
</bibRefCitation>
;
<bibRefCitation id="EFD490D91C480C7CFF0CFCB5FD97FCF9" author="Maeda, T. &amp; Hirashima, K." box="[224,531,794,820]" pageId="15" pageNumber="405" pagination="35 - 38" refId="ref11523" refString="Maeda, T. &amp; Hirashima, K. (2021) Records of twelve Luciogobius species from interstitial coast of Haneta, Minabe, Wakayama Prefecture, Japan. Nanki-seibutsu, 63 (1), 35 - 38. [In Japanese]" type="journal article" year="2021">Maeda &amp; Hirashima, 2021</bibRefCitation>
; Okumura
<emphasis id="B931313A1C480C7CFD7FFCB4FD42FCF9" box="[659,710,794,820]" italics="true" pageId="15" pageNumber="405">et al</emphasis>
., 2023)]. The species prefers gravel deposits on beaches, rather than rocky coasts (
<bibRefCitation id="EFD490D91C480C7CFE84FC91FDF4FC95" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[360,624,830,856]" pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFE01FC90FD9BFC95" box="[493,543,830,856]" italics="true" pageId="15" pageNumber="405">et al</emphasis>
., 2019
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35FBEA31C480C63FF2BFCCDFCC5FF18" lastPageId="16" lastPageNumber="406" pageId="15" pageNumber="405" type="discussion">
<paragraph id="8BFAED281C480C7CFF2BFCCDFB34FC6D" blockId="15.[151,1437,361,928]" pageId="15" pageNumber="405">
<emphasis id="B931313A1C480C7CFF2BFCCDFEB0FCB1" bold="true" box="[199,308,866,892]" pageId="15" pageNumber="405">Remarks</emphasis>
. The pre-dorsal fin length + P
<subScript id="17C1EF6D1C480C7CFD69FCDBFD09FC4F" attach="both" box="[645,653,884,898]" fontSize="6" pageId="15" pageNumber="405">2</subScript>
A length of 98.4102.1% of SL in
<taxonomicName id="4C4596AB1C480C7CFBE2FCCDFB50FCB1" box="[1038,1236,866,892]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C480C7CFBE2FCCDFB1CFCB1" box="[1038,1176,866,892]" italics="true" pageId="15" pageNumber="405">Luciogobius</emphasis>
sp. 7
</taxonomicName>
<emphasis id="B931313A1C480C7CFB37FCCCFA9DFCB1" box="[1243,1305,867,892]" italics="true" pageId="15" pageNumber="405">sensu</emphasis>
<bibRefCitation id="EFD490D91C480C7CFACCFCCDFEA5FC6D" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFF7BFC28FF4BFC6D" box="[151,207,902,928]" italics="true" pageId="15" pageNumber="405">et al.</emphasis>
(2019)
</bibRefCitation>
is similar to that of
<taxonomicName id="4C4596AB1C480C7CFDEEFC28FD38FC6D" box="[514,700,902,928]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C480C7CFDEEFC28FD38FC6D" box="[514,700,902,928]" italics="true" pageId="15" pageNumber="405">L. platycephalus</emphasis>
</taxonomicName>
, but shorter than in
<taxonomicName id="4C4596AB1C480C7CFC71FC28FB88FC6D" authority="Koreeda &amp; Maeda &amp; Motomura, 2023" authorityName="Koreeda &amp; Maeda &amp; Motomura" authorityYear="2023" box="[925,1036,903,928]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="griseus" status="n. sp.">
<emphasis id="B931313A1C480C7CFC71FC28FB88FC6D" box="[925,1036,903,928]" italics="true" pageId="15" pageNumber="405">L. griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2028C411C480C7CFBFEFC29FBCAFC6D" box="[1042,1102,902,928]" pageId="15" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="137EF1AD1C480C7CFBB2FC29FB26FC6D" box="[1118,1186,902,928]" captionStart="FIGURE 7" captionStartId="10.[152,255,869,893]" captionTargetBox="[203,1436,370,832]" captionTargetId="figure-89@10.[151,1436,345,846]" captionTargetPageId="10" captionText="FIGURE 7. Relationship of (A) pelvic-fin length (P2F length) as % of pre-dorsal-fin length (PDF) and (B) PDF + P2A length as % of standard length (SL) in the Luciogobius platycephalus complex. Closed star: holotype of L. griseus; open stars: paratypes of L. griseus; circles: non-type specimens of L. platycephalus; triangles: Luciogobius sp. 7 sensu Shibukawa et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10144878" httpUri="https://zenodo.org/record/10144878/files/figure.png" pageId="15" pageNumber="405">Fig. 7</figureCitation>
).
</paragraph>
<caption id="DF3ABDA01C480C7CFF74F8AFFDEDF8AD" ID-DOI="http://doi.org/10.5281/zenodo.10144884" ID-Zenodo-Dep="10144884" httpUri="https://zenodo.org/record/10144884/files/figure.png" pageId="15" pageNumber="405" startId="15.[152,255,1792,1816]" targetBox="[151,1436,952,1768]" targetPageId="15" targetType="figure">
<paragraph id="8BFAED281C480C7CFF74F8AFFDEDF8AD" blockId="15.[151,1437,1792,1888]" pageId="15" pageNumber="405">
<emphasis id="B931313A1C480C7CFF74F8AFFEA1F8D5" bold="true" box="[152,293,1792,1816]" pageId="15" pageNumber="405">FIGURE 10.</emphasis>
Fresh specimens of
<taxonomicName id="4C4596AB1C480C7CFE1BF8AFFD2FF8DA" box="[503,683,1792,1816]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="405" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C480C7CFE1BF8AFFDF0F8D5" box="[503,628,1792,1816]" italics="true" pageId="15" pageNumber="405">Luciogobius</emphasis>
sp. 7
</taxonomicName>
<emphasis id="B931313A1C480C7CFD5DF8AEFD6DF8D5" box="[689,745,1793,1816]" italics="true" pageId="15" pageNumber="405">sensu</emphasis>
<bibRefCitation id="EFD490D91C480C7CFD1CF8AFFC60F8D5" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[752,996,1792,1816]" pageId="15" pageNumber="405" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C480C7CFC8BF8AEFC10F8D5" box="[871,916,1792,1816]" italics="true" pageId="15" pageNumber="405">et al</emphasis>
. (2019)
</bibRefCitation>
from Kochi Prefecture, Shikoku, Japan. A: KAUMI. 159033, male, 51.3 mm SL; B: KAUMI. 159029, female, 33.2 mm SL; C: KAUMI. 159028, female, 32.1 mm SL. Photographs taken immediately after fixation.
</paragraph>
</caption>
<paragraph id="8BFAED281C480C63FF2BF81DFCC5FF18" blockId="15.[151,1437,1970,2033]" lastBlockId="16.[151,1436,151,213]" lastPageId="16" lastPageNumber="406" pageId="15" pageNumber="405">
Three individuals from
<collectingRegion id="498123CA1C480C7CFE22F81DFD08F801" box="[462,652,1970,1996]" country="Japan" name="Kochi" pageId="15" pageNumber="405">Kochi Prefecture</collectingRegion>
had relatively shorter pelvic-fin length (3.03.2 % of SL vs. 3.14.0% of SL) than in
<collectingRegion id="498123CA1C480C7CFEAEF879FE2EF83D" box="[322,426,2006,2032]" country="Japan" name="Shizuoka" pageId="15" pageNumber="405">Shizuoka</collectingRegion>
specimens (
<figureCitation id="137EF1AD1C480C7CFDD5F879FD14F83D" box="[569,656,2006,2032]" captionStart="FIGURE 8" captionStartId="12.[152,255,1453,1477]" captionTargetBox="[203,1396,218,1415]" captionTargetId="figure-18@12.[151,1436,181,1429]" captionTargetPageId="12" captionText="FIGURE 8. Relationships of (A) head length, (B) P2A length, (C) pre-dorsal-fin length (PDF), (D) pre-anal-fin length, (E) prepelvic-fin length, and (F) pelvic-fin length (P2F length) as % of standard length (SL) in the Luciogobius platycephalus complex. Closed star: holotype of L. griseus; open stars: paratypes of L. griseus; closed circle: holotype of L. platycephalus; open circles: non-type specimens of L. platycephalus; triangles: Luciogobius sp. 7 sensu Shibukawa et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10144880" httpUri="https://zenodo.org/record/10144880/files/figure.png" pageId="15" pageNumber="405">Fig. 8F</figureCitation>
), although specimens from both areas shared fewer than 8 branched pectoral-fin rays, a diagnostic character of the species according to
<bibRefCitation id="EFD490D91C570C63FC6FFF38FB15FF7C" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[899,1169,151,177]" pageId="16" pageNumber="406" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C570C63FBE9FF38FBB2FF7C" box="[1029,1078,151,177]" italics="true" pageId="16" pageNumber="406">et al</emphasis>
. (2019)
</bibRefCitation>
. The latter reported also that branched rays were absent in individuals &lt;38.0 mm SL.
</paragraph>
</subSubSection>
<subSubSection id="C35FBEA31C570C63FF7BFE88FA18FCCB" pageId="16" pageNumber="406" type="key">
<paragraph id="8BFAED281C570C63FF7BFE88FCA2FEA8" blockId="16.[151,1436,294,357]" pageId="16" pageNumber="406">
<emphasis id="B931313A1C570C63FF7BFE88FCC7FE8C" bold="true" box="[151,835,294,321]" pageId="16" pageNumber="406">
Key to species of the
<taxonomicName id="4C4596AB1C570C63FE72FE89FD51FE8C" ID-CoL="3WCMP" box="[414,725,294,321]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="406" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C570C63FE72FE89FD51FE8C" bold="true" box="[414,725,294,321]" italics="true" pageId="16" pageNumber="406">Luciogobius platycephalus</emphasis>
</taxonomicName>
complex
</emphasis>
(except larvae and juveniles): each body proportion needs to be aware of changing with growth (see
<figureCitation id="137EF1AD1C570C63FD59FEE4FC87FEA8" box="[693,771,331,357]" captionStart="FIGURE 7" captionStartId="10.[152,255,869,893]" captionTargetBox="[203,1436,370,832]" captionTargetId="figure-89@10.[151,1436,345,846]" captionTargetPageId="10" captionText="FIGURE 7. Relationship of (A) pelvic-fin length (P2F length) as % of pre-dorsal-fin length (PDF) and (B) PDF + P2A length as % of standard length (SL) in the Luciogobius platycephalus complex. Closed star: holotype of L. griseus; open stars: paratypes of L. griseus; circles: non-type specimens of L. platycephalus; triangles: Luciogobius sp. 7 sensu Shibukawa et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10144878" httpUri="https://zenodo.org/record/10144878/files/figure.png" pageId="16" pageNumber="406">Figs. 7</figureCitation>
,
<figureCitation id="137EF1AD1C570C63FCE3FEE4FC99FEA8" box="[783,797,331,357]" captionStart="FIGURE 8" captionStartId="12.[152,255,1453,1477]" captionTargetBox="[203,1396,218,1415]" captionTargetId="figure-18@12.[151,1436,181,1429]" captionTargetPageId="12" captionText="FIGURE 8. Relationships of (A) head length, (B) P2A length, (C) pre-dorsal-fin length (PDF), (D) pre-anal-fin length, (E) prepelvic-fin length, and (F) pelvic-fin length (P2F length) as % of standard length (SL) in the Luciogobius platycephalus complex. Closed star: holotype of L. griseus; open stars: paratypes of L. griseus; closed circle: holotype of L. platycephalus; open circles: non-type specimens of L. platycephalus; triangles: Luciogobius sp. 7 sensu Shibukawa et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.10144880" httpUri="https://zenodo.org/record/10144880/files/figure.png" pageId="16" pageNumber="406">8</figureCitation>
)
</paragraph>
<key id="FFD4BA191C570C63FF7BFE3FFA18FCCB" pageId="16" pageNumber="406">
<keyStep id="30B1F58D1C570C63FF7BFE3FFA18FDD6" pageId="16" pageNumber="406">
<paragraph id="8BFAED281C570C63FF7BFE3FFA18FE0F" blockId="16.[151,1437,400,774]" pageId="16" pageNumber="406">
<keyLead id="30B44E1D1C570C63FF7BFE3FFA18FE0F" pageId="16" pageNumber="406">1a. Pectoral fin with 713 branched rays (usually more than 10); posterior margin of pectoral-fin membrane not strongly concave (except for free rays).................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8BFAED281C570C63FF7BFE65FA18FDD6" blockId="16.[151,1437,400,774]" pageId="16" pageNumber="406">
<keyLead id="30B44E1D1C570C63FF7BFE65FA18FDD6" pageId="16" pageNumber="406">
1b. Pectoral fin with 08 branched rays (usually 6; 0 in specimens &lt;
<quantity id="4CBD40CD1C570C63FC88FE65FC2CFE2D" box="[868,936,458,480]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="406" unit="mm" value="38.0">38 mm</quantity>
in SL); posterior margin of pectoral-fin membrane strongly concave; occurs in accreting beach.........................................................................................
<taxonomicName id="4C4596AB1C570C63FDE0FDAAFD32FDD7" box="[524,694,517,538]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="406" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C570C63FDE0FDAAFD07FDD7" box="[524,643,517,538]" italics="true" pageId="16" pageNumber="406">Luciogobius</emphasis>
sp. 7
</taxonomicName>
<emphasis id="B931313A1C570C63FD50FDA9FD75FDD6" box="[700,753,518,539]" italics="true" pageId="16" pageNumber="406">sensu</emphasis>
<bibRefCitation id="EFD490D91C570C63FD14FDAAFC5BFDD7" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[760,991,517,539]" pageId="16" pageNumber="406" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C570C63FC84FDA9FC17FDD7" box="[872,915,517,539]" italics="true" pageId="16" pageNumber="406">et al</emphasis>
. (2019)
</bibRefCitation>
(Japanese name: Hauchiwa-yari-mimizuhaze)
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</paragraph>
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2a. Pre-dorsal-fin length &lt;70% of SL (except in a specimen
<quantity id="4CBD40CD1C570C63FCFBFD8DFCE9FDF5" box="[791,877,546,568]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="16" pageNumber="406" unit="mm" value="26.1">26.1 mm</quantity>
SL); pelvic-fin length&gt; 5.3% of pre-dorsal-fin length in specimens
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SL or larger (&gt; 5.9% in smaller specimens); pre-dorsal-fin length + P
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A length 95.4101.6% of SL; bright yellow to green body in live and fresh specimens (except individuals&gt;
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SL); usually occurs along temperate rocky coasts in
<collectingCountry id="F352ADB81C570C63FF17FDD4FEB6FD5D" box="[251,306,635,656]" name="Japan" pageId="16" pageNumber="406">Japan</collectingCountry>
(
<collectingRegion id="498123CA1C570C63FED2FDD5FE3BFD5D" box="[318,447,634,656]" country="Japan" name="Aomori" pageId="16" pageNumber="406">Aomori Pref.</collectingRegion>
to Tanega-shima island) and
<collectingCountry id="F352ADB81C570C63FD34FDD4FC97FD5D" box="[728,787,635,656]" name="South Korea" pageId="16" pageNumber="406">Korea</collectingCountry>
…..............
<taxonomicName id="4C4596AB1C570C63FC33FDD5FB64FD42" ID-CoL="3WCMP" box="[991,1248,634,655]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="406" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C570C63FC33FDD5FB64FD42" box="[991,1248,634,655]" italics="true" pageId="16" pageNumber="406">Luciogobius platycephalus</emphasis>
</taxonomicName>
(Yari-mimizuhaze)
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</paragraph>
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2b. Pre-dorsal-fin length&gt; 69.5% of SL; pelvic-fin length &lt;5.3% of pre-dorsal-fin length in specimens
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SL or larger (&lt;6.0% in smaller specimens); pre-dorsal-fin length + P
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A length 103.0107.3% of SL; gray in life (sometimes bluish or purplish), greenish dark brown when fresh; usually occurs near subtropical river mouths or coastal freshwater springs in
<collectingCountry id="F352ADB81C570C63FAF1FD7CFAD0FD25" box="[1309,1364,723,744]" name="Japan" pageId="16" pageNumber="406">Japan</collectingCountry>
(Yaku-shima island to Okinawa-jima island).........................
<taxonomicName id="4C4596AB1C570C63FC96FD5FFBBFFCC8" authority="Koreeda &amp; Maeda &amp; Motomura, 2023" authorityName="Koreeda &amp; Maeda &amp; Motomura" authorityYear="2023" box="[890,1083,752,773]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="406" phylum="Chordata" rank="species" species="griseus" status="sp. nov.">
<emphasis id="B931313A1C570C63FC96FD5FFBBFFCC8" box="[890,1083,752,773]" italics="true" pageId="16" pageNumber="406">Luciogobius griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2028C411C570C63FBAEFD5FFBF2FCC8" box="[1090,1142,752,773]" pageId="16" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
(Sumizome-yari-mimizuhaze)
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</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>