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<document id="2639D380749C21717B711C737CCAF51C" ID-CLB-Dataset="307877" ID-DOI="10.11646/zootaxa.5569.3.10" ID-GBIF-Dataset="5c5b991e-e18e-4399-8957-689bf1581cb3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14734270" ID-ZooBank="829FE323-1C2E-4896-AA71-C20B7EA0BFF3" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1737735007098" checkinUser="plazi" docAuthor="Williams, Jason L., Williams, John F., Lapolla, John S. &amp; Lucky, Andrea" docDate="2025" docId="03EC28231547326AFF692CA2FD45FD95" docLanguage="en" docName="zootaxa.5569.3.10.pdf" docOrigin="Zootaxa 5569 (3)" docSource="https://doi.org/10.11646/zootaxa.5569.3.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nylanderia usul Williams, Williams, Lapolla &amp; Lucky, 2025, sp. nov." docType="treatment" docVersion="6" lastPageNumber="587" masterDocId="FFD5505B1564324DFFFE285AFF8AFFF4" masterDocTitle="Nylanderia of the World, Part IV: Taxonomic contributions to the American Clade I of New World Nylanderia (Hymenoptera: Formicidae)" masterLastPageNumber="593" masterPageNumber="549" pageNumber="584" updateTime="1738069513367" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="55F52F12C8A40ECAF71B09B9A7BCE469" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="75E92FA56D40E43671DFFA0264ABC25B">
<mods:title id="6FCBC70DA872B8653D3B5808EE46B5AF">Nylanderia of the World, Part IV: Taxonomic contributions to the American Clade I of New World Nylanderia (Hymenoptera: Formicidae)</mods:title>
</mods:titleInfo>
<mods:name id="A30DC90A9102DFB2AEF92AA618022AAA" type="personal">
<mods:role id="E78A17C48907CB6611DAD1912B1D6374">
<mods:roleTerm id="8C83D3236054FE922C8D3D5F9FEEC4BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86FC75C0C527FC2B71284E2E1DF603F4">Williams, Jason L.</mods:namePart>
<mods:affiliation id="92D4B53D53B90F713B9D636C3B8BEDC8">Entomology &amp; Nematology Department, University of Florida, Gainesville, Florida, U. S. A.</mods:affiliation>
</mods:name>
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<mods:role id="169427D38BD536C33C5589C9A9BC189F">
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</mods:role>
<mods:namePart id="9490965BDEDEFC5A8C090A5FC2A25D9D">Williams, John F.</mods:namePart>
<mods:affiliation id="8D803FE877F199AB9C0CFC16871497E3">Entomology &amp; Nematology Department, University of Florida, Gainesville, Florida, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="58B0CA82646F0182323E49E891AC2872" type="personal">
<mods:role id="D05E9EA1DD617929C1C393417724726D">
<mods:roleTerm id="00AC25E3E0C3ACA515B0D786615820B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7BD1D0236D401B49D71D6ACEC3495C4">Lapolla, John S.</mods:namePart>
<mods:affiliation id="477BEF2DB0525792C258E3146989AA5F">Department of Biological Sciences, Towson University, Towson, Maryland, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="5F135C634F1FC640DB7E9700B1397FD0" type="personal">
<mods:role id="D28A77BE14D1CD4C09831F2111919AF1">
<mods:roleTerm id="1EE1FDD7452DA7483FBCB30CE9C3D70C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F9B91EB20497247704E263A5F6D100A3">Lucky, Andrea</mods:namePart>
<mods:affiliation id="036F19D15CF08460015C9EBB6156116E">Entomology &amp; Nematology Department, University of Florida, Gainesville, Florida, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C3167667DD68434C45A2A66F29081841">text</mods:typeOfResource>
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<mods:titleInfo id="8B6311749B9E9A3238881E394D3BCEE8">
<mods:title id="AA20D5B154A37B8326C626199B62F04A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="145ABC1476C1E1DA080FA57D98506C9A">
<mods:date id="BE1523B17AD8CD6DE2AD224C02AB846B">2025</mods:date>
<mods:detail id="F1B727070474E9FB858DB1FBD23844A0" type="pubDate">
<mods:number id="82C62F6BED753BD230B5036F9546430B">2025-01-15</mods:number>
</mods:detail>
<mods:detail id="8E0C48E242D3C59DE25C75EDD2D688C4" type="volume">
<mods:number id="88BBCD834B4BCB0C5160DCF5F68517F9">5569</mods:number>
</mods:detail>
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<mods:number id="154526F83AB1F89BCE21FC43E1AFDE98">3</mods:number>
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<mods:start id="8016647EC2BA56470DF6B20DC9577F0D">549</mods:start>
<mods:end id="DF2095E53A5EEFFD36F38D538A555032">593</mods:end>
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<mods:location id="681919C6974E684E717566DE33D0EBB5">
<mods:url id="F87285D12DF6E50143F06D1E14A69BEB">https://doi.org/10.11646/zootaxa.5569.3.10</mods:url>
</mods:location>
<mods:classification id="189A55B7D923B3ACDF6D2FDF65CEBC01">journal article</mods:classification>
<mods:identifier id="31AEBD4AD555093F669DD2AFFE8682FF" type="CLB-Dataset">307877</mods:identifier>
<mods:identifier id="B7A37CA50A9A2975EAFF58CE2C226196" type="DOI">10.11646/zootaxa.5569.3.10</mods:identifier>
<mods:identifier id="A420B125D693043DAF7E8A28F721A647" type="GBIF-Dataset">5c5b991e-e18e-4399-8957-689bf1581cb3</mods:identifier>
<mods:identifier id="16AC51753A865C41FC429825D5A234CE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2EFEDE072BA2F713D3A0F85E09711EDB" type="Zenodo-Dep">14734270</mods:identifier>
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</mods:mods>
<treatment id="03EC28231547326AFF692CA2FD45FD95" ID-DOI="http://doi.org/10.5281/zenodo.14754813" ID-Zenodo-Dep="14754813" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95" httpUri="http://treatment.plazi.org/id/03EC28231547326AFF692CA2FD45FD95" lastPageId="39" lastPageNumber="587" pageId="35" pageNumber="584">
<subSubSection id="C35FCABE1547326EFF692CA2FE4CFAE7" box="[151,454,1272,1299]" pageId="35" pageNumber="584" type="nomenclature">
<paragraph id="8BFA99351547326EFF692CA2FE4CFAE7" blockId="35.[151,855,1272,1336]" box="[151,454,1272,1299]" pageId="35" pageNumber="584">
<heading id="D0B22E591547326EFF692CA2FE4CFAE7" bold="true" box="[151,454,1272,1299]" fontSize="11" level="1" pageId="35" pageNumber="584" reason="1">
<emphasis id="B93145271547326EFF692CA2FE4CFAE7" bold="true" box="[151,454,1272,1299]" pageId="35" pageNumber="584">
<taxonomicName id="4C45E2B61547326EFF692CA2FED0FAE7" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[151,346,1272,1299]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="584" phylum="Arthropoda" rank="species" species="usul" status="sp. nov.">
<emphasis id="B93145271547326EFF692CA2FED0FAE7" bold="true" box="[151,346,1272,1299]" italics="true" pageId="35" pageNumber="584">Nylanderia usul</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A202F85C1547326EFE972CA3FE4CFAE7" box="[361,454,1273,1299]" pageId="35" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE1547326EFF692D47FCDDFAC3" box="[151,855,1309,1336]" pageId="35" pageNumber="584" type="description">
<paragraph id="8BFA99351547326EFF692D47FCDDFAC3" blockId="35.[151,855,1272,1336]" box="[151,855,1309,1336]" pageId="35" pageNumber="584">
<figureCitation id="137E85B01547326EFF692D47FE96FAC3" box="[151,284,1309,1335]" captionStart="FIGURES 62–64" captionStartId="36.[152,270,1495,1519]" captionTargetBox="[151,1435,181,1471]" captionTargetId="figure-26@36.[151,1435,181,1471]" captionTargetPageId="36" captionText="FIGURES 62–64. Nylanderia usul holotype worker (USNMENT01131798). Lateral, full-face, and dorsal view of the body." figureDoi="http://doi.org/10.5281/zenodo.14734326" httpUri="https://zenodo.org/record/14734326/files/figure.png" pageId="35" pageNumber="584">Figs. 62–64</figureCitation>
(worker);
<figureCitation id="137E85B01547326EFE6D2D47FD9DFACC" box="[403,535,1309,1336]" captionStart="FIGURES 65–66" captionStartId="37.[152,270,715,739]" captionTargetBox="[151,1414,181,691]" captionTargetId="figure-17@37.[151,1435,181,691]" captionTargetPageId="37" captionText="FIGURES 65–66. Nylanderia usul paratype queen (USNMENT01131819). Full-face and lateral view of the body." figureDoi="http://doi.org/10.5281/zenodo.14734328" httpUri="https://zenodo.org/record/14734328/files/figure.png" pageId="35" pageNumber="584">Figs. 65–66</figureCitation>
(queen);
<figureCitation id="137E85B01547326EFD7D2D47FC8DFAC3" box="[643,775,1309,1335]" captionStart="FIGURES 67–70" captionStartId="38.[152,270,1581,1605]" captionTargetBox="[151,1434,274,1558]" captionTargetId="figure-61@38.[151,1435,273,1558]" captionTargetPageId="38" captionText="FIGURES 67–70. Nylanderia usul paratype male (USNMENT01131831). Full-face and lateral view of the body; ventral and dorsal view of the genital capsule." figureDoi="http://doi.org/10.5281/zenodo.14734332" httpUri="https://zenodo.org/record/14734332/files/figure.png" pageId="35" pageNumber="584">Figs. 67–70</figureCitation>
(male)
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE1547326EFF692D3BFB5BFA22" pageId="35" pageNumber="584" type="materials_examined">
<paragraph id="8BFA99351547326EFF692D3BFB5BFA22" blockId="35.[151,1437,1377,1494]" pageId="35" pageNumber="584">
<materialsCitation id="3B2D93681547326EFF692D3BFC03FA6C" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="1950" latitude="9.55301" location="San Jose" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131798" specimenCount="1" specimenCount-male="1" specimenCount-queen="1" specimenCount-worker="5" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="54FE27971547326EFF692D3BFF7FFA8D" box="[151,245,1377,1401]" pageId="35" pageNumber="584" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4352BC1547326EFF062D3BFECBFA8D" box="[248,321,1377,1401]" pageId="35" pageNumber="584" type="worker">worker</specimenCount>
,
<emphasis id="B93145271547326EFEB92D3BFE6CFA8D" bold="true" box="[327,486,1377,1401]" pageId="35" pageNumber="584">
<collectingCountry id="F352D9A51547326EFEB92D3BFE6BFA8D" box="[327,481,1377,1401]" name="Costa Rica" pageId="35" pageNumber="584">COSTA RICA</collectingCountry>
:
</emphasis>
<collectingRegion id="498157D71547326EFE162D38FDB4FA8E" box="[488,574,1378,1402]" country="Costa Rica" name="San Jose" pageId="35" pageNumber="584">San José</collectingRegion>
,
<locationDeviation id="9A4880171547326EFDBA2D3BFCACFA8E" box="[580,806,1377,1402]" pageId="35" pageNumber="584">11km SSE Santa María</locationDeviation>
,
<geoCoordinate id="EE71FFF21547326EFCD22D3BFCF4FA8D" box="[812,894,1377,1401]" degrees="9.55301" direction="north" orientation="latitude" pageId="35" pageNumber="584" precision="10" value="9.55301">9.55301</geoCoordinate>
<geoCoordinate id="EE71FFF21547326EFC7E2D3BFC62FA8D" box="[896,1000,1377,1401]" degrees="83.94523" direction="west" orientation="longitude" pageId="35" pageNumber="584" precision="10" value="-83.94523">-83.94523</geoCoordinate>
±10m,elev.1950m,
<date id="FFFBBFF51547326EFB532D3BFA96FA8D" box="[1197,1308,1377,1401]" pageId="35" pageNumber="584" value="2015-06-24">
<collectingDate id="EFBF461D1547326EFB532D3BFA96FA8D" box="[1197,1308,1377,1401]" pageId="35" pageNumber="584" value="2015-06-24">24.vi.2015</collectingDate>
</date>
,
<collectedFrom id="3859DAC81547326EFADC2D3BFA12FA8D" box="[1314,1432,1377,1401]" pageId="35" pageNumber="584">cloud forest</collectedFrom>
,
<collectingMethod id="5204E1221547326EFF392DDAFEBFFA6C" box="[199,309,1408,1432]" pageId="35" pageNumber="584">sifted litter</collectingMethod>
, ADMAC#Wa-E-04-1-49 (
<collectionCode id="ED5401F01547326EFDBB2DDAFD19FA6C" box="[581,659,1408,1432]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNM</collectionCode>
:
<specimenCode id="DBE3314E1547326EFD652DDAFCF5FA6C" box="[667,895,1408,1432]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131798</specimenCode>
).
</materialsCitation>
<materialsCitation id="3B2D93681547326EFC732DDAFC4BFA43" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131109" specimenCount="7" specimenCount-male="1" specimenCount-worker="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCount id="9D4352BC1547326EFC732DDAFBC4FA6C" box="[909,1102,1408,1433]" count="5" pageId="35" pageNumber="584" type="worker" typeStatus="paratype">5 paratype workers</specimenCount>
,
<specimenCount id="9D4352BC1547326EFBA82DDAFA88FA6C" box="[1110,1282,1408,1433]" count="1" pageId="35" pageNumber="584" type="queen" typeStatus="paratype">1 paratype queen</specimenCount>
, and
<specimenCount id="9D4352BC1547326EFACA2DDAFF72FA43" count="1" pageId="35" pageNumber="584" type="male" typeStatus="paratype">1 paratype male</specimenCount>
with same locality data as holotype (
<collectionCode id="ED5401F01547326EFD7C2DC5FD5AFA43" box="[642,720,1439,1463]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNM</collectionCode>
:
<specimenCode id="DBE3314E1547326EFD222DC5FC37FA43" box="[732,957,1439,1463]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131109</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2D93681547326EFC372DC5FB25FA43" box="[969,1199,1439,1463]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131799" specimenCount="1" specimenCount-worker="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCode id="DBE3314E1547326EFC372DC5FB21FA43" box="[969,1195,1439,1463]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131799</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2D93681547326EFB492DC5FA17FA43" box="[1207,1437,1439,1463]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131800" specimenCount="1" specimenCount-worker="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCode id="DBE3314E1547326EFB492DC5FA12FA43" box="[1207,1432,1439,1463]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131800</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2D93681547326EFF392DE4FE27FA22" box="[199,429,1470,1494]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131819" specimenCount="1" specimenCount-worker="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCode id="DBE3314E1547326EFF392DE4FE23FA22" box="[199,425,1470,1494]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131819</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2D93681547326EFE4D2DE4FD13FA22" box="[435,665,1470,1494]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131831" specimenCount="1" specimenCount-worker="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCode id="DBE3314E1547326EFE4D2DE4FD1FFA22" box="[435,661,1470,1494]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="584" type="Museum">USNMENT01131831</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B2D93681547326EFD612DE4FC56FA22" box="[671,988,1470,1494]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="MNCR" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131802" specimenCount="1" specimenCount-queen="1" stateProvince="San Jose" typeStatus="paratype">
<collectionCode id="ED5401F01547326EFD612DE4FD64FA22" box="[671,750,1470,1494]" collectionName="Costa Rica, San Jose, Museo Nacional de Costa Rica" pageId="35" pageNumber="584">MNCR</collectionCode>
:
<specimenCode id="DBE3314E1547326EFD092DE4FC52FA22" box="[759,984,1470,1494]" pageId="35" pageNumber="584">USNMENT01131802</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B2D93681547326EFC1C2DE4FB5BFA22" box="[994,1233,1470,1494]" collectedFrom="cloud forest" collectingDate="2015-06-24" collectingMethod="sifted litter" collectionCode="USNM" country="Costa Rica" elevation="10" latitude="9.55301" location="Santa Maria" longLatPrecision="14" longitude="-83.94523" pageId="35" pageNumber="584" specimenCode="USNMENT01131804" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<specimenCode id="DBE3314E1547326EFC1C2DE4FB4CFA22" box="[994,1222,1470,1494]" pageId="35" pageNumber="584">USNMENT01131804</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE1547326EFF692E5EFD0CF925" pageId="35" pageNumber="584" type="diagnosis">
<paragraph id="8BFA99351547326EFF692E5EFDD8F95A" blockId="35.[151,1437,1539,1926]" pageId="35" pageNumber="584">
<emphasis id="B93145271547326EFF692E5EFEF9F9E9" bold="true" box="[151,371,1539,1566]" pageId="35" pageNumber="584">Worker diagnosis:</emphasis>
Antennae relatively short, with scapes surpassing posterior margin by about a quarter to a third of their length; color variable from yellow to dark brown, with meso- and metacoxae at most somewhat lighter than mesosoma; fine cuticular microsculpture resulting in moderate to strong blue iridescence on head, mesosoma, and gaster; moderate to dense cephalic pubescence; propodeum high and evenly rounded, with posterior face steep and at most slightly longer than dorsal face.
</paragraph>
<paragraph id="8BFA99351547326EFF392EEDFD0CF925" blockId="35.[151,1437,1539,1926]" box="[199,646,1719,1746]" pageId="35" pageNumber="584">
<emphasis id="B93145271547326EFF392EEDFEF6F925" bold="true" box="[199,380,1719,1745]" pageId="35" pageNumber="584">Compare with:</emphasis>
<emphasis id="B93145271547326EFE7D2EE2FD0CF925" box="[387,646,1720,1746]" italics="true" pageId="35" pageNumber="584">
<taxonomicName id="4C45E2B61547326EFE7D2EE2FE74F926" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[387,510,1720,1746]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="584" phylum="Arthropoda" rank="species" species="mendax">N. mendax</taxonomicName>
,
<taxonomicName id="4C45E2B61547326EFDF72EE2FD0CF925" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[521,646,1720,1745]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="584" phylum="Arthropoda" rank="species" species="mosaica">N. mosaica</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE15473268FF392E86FEE1FA64" lastPageId="37" lastPageNumber="586" pageId="35" pageNumber="584" type="description">
<paragraph id="8BFA99351547326EFF392E86FEA2F872" blockId="35.[151,1437,1539,1926]" pageId="35" pageNumber="584">
WORKER.
<emphasis id="B93145271547326EFEB02E86FDCFF902" box="[334,581,1756,1782]" italics="true" pageId="35" pageNumber="584">Measurements (n=11)</emphasis>
: TL: 2.00–2.35; EL: 0.11–0.13; EW: 0.08–0.10; IOD: 0.32–0.39; HE: 0.01– 0.02; HL: 0.58–0.65; HW: 0.47–0.53; HLA: 0.18–0.23; HLP: 0.27–0.30; SL: 0.65–0.74; PW: 0.36–0.40; MW: 0.18–0.21; PrW: 0.26–0.32; PDH: 0.18–0.27; PTW: 0.13–0.15; LHT: 0.58–0.69; WL: 0.69–0.80; GL: 0.70–0.94; SMC: 9–12; PMC: 2–3; MMC: 2–3.
<emphasis id="B93145271547326EFDC52F12FD06F896" box="[571,652,1864,1890]" italics="true" pageId="35" pageNumber="584">Indices</emphasis>
: BLI: 144–154; CI: 81–84; EPI: 67–80; HTI: 113–131; REL: 18–21; SI: 128–140.
</paragraph>
<caption id="DF3AC9BD15403269FF662D8DFA03FA1B" ID-DOI="http://doi.org/10.5281/zenodo.14734326" ID-Zenodo-Dep="14734326" box="[152,1417,1495,1519]" httpUri="https://zenodo.org/record/14734326/files/figure.png" pageId="36" pageNumber="585" startId="36.[152,270,1495,1519]" targetBox="[151,1435,181,1471]" targetPageId="36" targetType="figure">
<paragraph id="8BFA993515403269FF662D8DFA03FA1B" blockId="36.[152,1417,1495,1519]" box="[152,1417,1495,1519]" pageId="36" pageNumber="585">
<emphasis id="B931452715403269FF662D8DFED0FA1B" bold="true" box="[152,346,1495,1519]" pageId="36" pageNumber="585">FIGURES 62–64.</emphasis>
<taxonomicName id="4C45E2B615403269FE9E2D8DFD89FA1B" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[352,515,1495,1519]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="585" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B931452715403269FE9E2D8DFD89FA1B" box="[352,515,1495,1519]" italics="true" pageId="36" pageNumber="585">Nylanderia usul</emphasis>
</taxonomicName>
holotype worker (USNMENT01131798). Lateral, full-face, and dorsal view of the body.
</paragraph>
</caption>
<paragraph id="8BFA993515403269FF392E1BFB8FF837" blockId="36.[151,1437,1601,1988]" pageId="36" pageNumber="585">
<emphasis id="B931452715403269FF392E1BFE82F9AF" box="[199,264,1601,1627]" italics="true" pageId="36" pageNumber="585">Color</emphasis>
: variable, from yellow to dark brown; darker specimens often with yellow to yellow-brown mandibles and leg joints; in darker specimens, meso-and metacoxae somewhat lighter than mesosoma; overall cuticle moderately to finely microsculptured, resulting in moderate to strong blue iridescence on head, mesosoma, and gaster.
<emphasis id="B931452715403269FAC12ED3FA1FF957" box="[1343,1429,1673,1699]" italics="true" pageId="36" pageNumber="585">Pilosity</emphasis>
: moderate to dense cephalic pubescence; mesosoma pubescence sparse to absent, limited at most to scattered hairs on pronotum and propodeum, and a fringe of hairs concentrated just posterior to the metanotal groove; gaster pubescence sparse to absent; pair of small, erect setae on apex of petiole.
<emphasis id="B931452715403269FC3C2EAFFC75F8FB" box="[962,1023,1781,1807]" italics="true" pageId="36" pageNumber="585">Head</emphasis>
: longer than broad and quadrate with distinct posterolateral corners and flattened posterior margin, with moderate medial emargination; scapes surpass posterior margin of head by about a quarter to a third of their length; median ocellus small, but often apparent; lateral ocelli absent or not apparent.
<emphasis id="B931452715403269FE162F38FDEAF88F" box="[488,608,1890,1915]" italics="true" pageId="36" pageNumber="585">Mesosoma</emphasis>
: promesonotal area evenly convex in profile view, with posterior margin of pronotum continuous or nearly continuous with anterior margin of mesonotum; propodeum high and evenly rounded, with posterior face steep and at most slightly longer than dorsal face.
</paragraph>
<caption id="DF3AC9BD15413268FF662A91FAA3FD17" ID-DOI="http://doi.org/10.5281/zenodo.14734328" ID-Zenodo-Dep="14734328" box="[152,1321,715,740]" httpUri="https://zenodo.org/record/14734328/files/figure.png" pageId="37" pageNumber="586" startId="37.[152,270,715,739]" targetBox="[151,1414,181,691]" targetPageId="37" targetType="figure">
<paragraph id="8BFA993515413268FF662A91FAA3FD17" blockId="37.[152,1321,715,740]" box="[152,1321,715,740]" pageId="37" pageNumber="586">
<emphasis id="B931452715413268FF662A91FED0FD17" bold="true" box="[152,346,715,739]" pageId="37" pageNumber="586">FIGURES 65–66.</emphasis>
<taxonomicName id="4C45E2B615413268FE9E2A91FD89FD17" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[352,515,715,739]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="586" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B931452715413268FE9E2A91FD89FD17" box="[352,515,715,739]" italics="true" pageId="37" pageNumber="586">Nylanderia usul</emphasis>
</taxonomicName>
paratype queen (USNMENT01131819). Full-face and lateral view of the body.
</paragraph>
</caption>
<paragraph id="8BFA993515413268FF392B6CFDA3FC80" blockId="37.[151,1437,821,2036]" pageId="37" pageNumber="586">
QUEEN.
<emphasis id="B931452715413268FECD2B6CFD97FCA4" box="[307,541,822,848]" italics="true" pageId="37" pageNumber="586">Measurements (n=1)</emphasis>
: TL: 3.73; EL: 0.19; HL: 0.72; HW: 0.69; SL: 0.86; WL: 1.44; GL: 1.58.
<emphasis id="B931452715413268FABA2B6CFA1FFCA4" box="[1348,1429,822,848]" italics="true" pageId="37" pageNumber="586">Indices</emphasis>
: BLI: 200; CI: 97; REL: 26; SI: 125.
</paragraph>
<paragraph id="8BFA993515413268FF392B24FD73FBF0" blockId="37.[151,1437,821,2036]" pageId="37" pageNumber="586">
Relatively large.
<emphasis id="B931452715413268FE702B24FE45FC6C" box="[398,463,894,920]" italics="true" pageId="37" pageNumber="586">Color</emphasis>
: medium to dark brown, with meso- and metacoxae at most somewhat lighter than mesosoma.
<emphasis id="B931452715413268FEE22BF8FEF8FC48" box="[284,370,930,956]" italics="true" pageId="37" pageNumber="586">Pilosity</emphasis>
: most of body covered in moderate to dense pubescence; pubescence sparse to absent laterally on pronotum, sparse on most of anepisternum, absent on dorsal half of katepisternum. and absent on posterior face of propodeum; 4-5 small, erect setae on anepisternum.
</paragraph>
<paragraph id="8BFA993515413268FF392C54FDA3FBB8" blockId="37.[151,1437,821,2036]" pageId="37" pageNumber="586">
MALE.
<emphasis id="B931452715413268FEDA2C54FD85FBDC" box="[292,527,1038,1064]" italics="true" pageId="37" pageNumber="586">Measurements (n=1)</emphasis>
: TL: 2.20; EL: 0.18; HL: 0.53; HW: 0.46; SL: 0.70; WL: 0.82; GL: 0.85.
<emphasis id="B931452715413268FABA2C54FA1FFBDC" box="[1348,1429,1038,1064]" italics="true" pageId="37" pageNumber="586">Indices</emphasis>
: BLI: 180; CI: 86; REL: 34; SI: 154.
</paragraph>
<paragraph id="8BFA993515413268FF392C0CFEE1FA64" blockId="37.[151,1437,821,2036]" pageId="37" pageNumber="586">
<emphasis id="B931452715413268FF392C0CFE82FB84" box="[199,264,1110,1136]" italics="true" pageId="37" pageNumber="586">Color</emphasis>
: medium brown with yellow mandibles; meso- and metacoxae lighter than mesosoma.
<emphasis id="B931452715413268FB082C0CFAC6FB84" box="[1270,1356,1110,1136]" italics="true" pageId="37" pageNumber="586">Pilosity</emphasis>
: dense pubescence on head, mesoscutum, and mesoscutellum; pubescence sparse to absent on pronotum, anepisternum, katepisternum, propodeum, and gaster; 1-2 small, erect setae on anepisternum.
<emphasis id="B931452715413268FBEA2CC4FBDBFB4C" box="[1044,1105,1182,1208]" italics="true" pageId="37" pageNumber="586">Head</emphasis>
: as broad as long; compound eyes surpass lateral margins of head in full-face view; scapes curve sinuously and surpass posterior margin of head by about a third of their length.
<emphasis id="B931452715413268FDFA2CBCFDF6FB0B" box="[516,636,1254,1279]" italics="true" pageId="37" pageNumber="586">Mesosoma</emphasis>
: In profile view, propodeum gently curved with dorsal face somewhat longer than posterior face.
<emphasis id="B931452715413268FE3B2D50FDA5FAD0" box="[453,559,1290,1316]" italics="true" pageId="37" pageNumber="586">Genitalia</emphasis>
: gonopod long, narrow, and evenly triangular to weakly digitiform, with apex tapered and coming to a point (
<figureCitation id="137E85B015413268FE0E2D74FDC8FABC" box="[496,578,1326,1352]" captionStart="FIGURE 71" captionStartId="39.[152,255,1385,1409]" captionTargetBox="[156,1433,638,1357]" captionTargetId="figure-253@39.[151,1435,633,1362]" captionTargetPageId="39" captionText="FIGURE 71. Line drawings of male gonopods of N. austroccidua, N. breviscapa, N. contraria, N. cf. docilis, N. lazulina, and N. usul." figureDoi="http://doi.org/10.5281/zenodo.14734334" httpUri="https://zenodo.org/record/14734334/files/figure.png" pageId="37" pageNumber="586">Fig. 71</figureCitation>
); in ventral view, ventromedial edge of basivolsella unevenly bracket-shaped and crenate, with edge at base of cuspis forming a right angle (
<figureCitation id="137E85B015413268FCB02D08FC2BFA9F" box="[846,929,1361,1388]" captionStart="FIGURES 67–70" captionStartId="38.[152,270,1581,1605]" captionTargetBox="[151,1434,274,1558]" captionTargetId="figure-61@38.[151,1435,273,1558]" captionTargetPageId="38" captionText="FIGURES 67–70. Nylanderia usul paratype male (USNMENT01131831). Full-face and lateral view of the body; ventral and dorsal view of the genital capsule." figureDoi="http://doi.org/10.5281/zenodo.14734332" httpUri="https://zenodo.org/record/14734332/files/figure.png" pageId="37" pageNumber="586">Fig. 69</figureCitation>
); gonossiculus straight and nearly three times longer than cuspis.
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE1541326BFF392DC3FD4CFF0D" lastPageId="38" lastPageNumber="587" pageId="37" pageNumber="586" type="materials_examined">
<paragraph id="8BFA99351541326BFF392DC3FD4CFF0D" blockId="37.[151,1437,821,2036]" lastBlockId="38.[151,1437,150,249]" lastPageId="38" lastPageNumber="587" pageId="37" pageNumber="586">
<emphasis id="B931452715413268FF392DC3FD8EFA47" bold="true" box="[199,516,1433,1459]" pageId="37" pageNumber="586">Other material examined:</emphasis>
<emphasis id="B931452715413268FDF02DC3FD4CFA47" bold="true" box="[526,710,1433,1459]" pageId="37" pageNumber="586">
<collectingCountry id="F352D9A515413268FDF02DC3FD4BFA47" box="[526,705,1433,1459]" name="Costa Rica" pageId="37" pageNumber="586">COSTA RICA</collectingCountry>
:
</emphasis>
<collectingRegion id="498157D715413268FD2E2DC0FCA6FA40" box="[720,812,1434,1460]" country="Costa Rica" name="Heredia" pageId="37" pageNumber="586">Heredia</collectingRegion>
,
<location id="8E9ACFEE15413268FCC42DC3FBBFFA40" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FCC42DC3FBBFFA40" box="[826,1077,1433,1460]" country="Costa Rica" latitude="10.26667" longLatPrecision="1" longitude="-84.08333" name="16 km SSE La Virgen" pageId="37" pageNumber="586" stateProvince="Heredia">
<quantity id="4CBD34D015413268FCC42DC3FCF0FA47" box="[826,890,1433,1459]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="586" unit="km" value="16.0">16km</quantity>
SSE La Virgen
</location>
,
<geoCoordinate id="EE71FFF215413268FBBC2DC3FB21FA47" box="[1090,1195,1433,1459]" degrees="10.26667" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="1" value="10.26667">10.26667</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FB482DC0FAA2FA40" box="[1206,1320,1434,1460]" degrees="84.08333" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="1" value="-84.08333">-84.08333</geoCoordinate>
±m, elev.
<quantity id="4CBD34D015413268FF692DE4FF6DFA2C" box="[151,231,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="586" unit="m" value="1100.0">
<elevation id="00687E0615413268FF692DE4FF6DFA2C" box="[151,231,1470,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="586" unit="m" value="1100.0">1100m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FF0E2DE4FEDFFA2C" box="[240,341,1470,1496]" pageId="37" pageNumber="586" value="2001-02-08">
<collectingDate id="EFBF461D15413268FF0E2DE4FEDFFA2C" box="[240,341,1470,1496]" pageId="37" pageNumber="586" value="2001-02-08">8.ii.2001</collectingDate>
</date>
, montane wet forest, leaf litter,ALAS#11/B/all;
<collectingRegion id="498157D715413268FC982DE4FC48FA2C" box="[870,962,1470,1496]" country="Costa Rica" name="Heredia" pageId="37" pageNumber="586">Heredia</collectingRegion>
,
<location id="8E9ACFEE15413268FC342DE7FB3EFA2C" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FC342DE7FB3EFA2C" box="[970,1204,1469,1496]" country="Costa Rica" latitude="10.26863" longLatPrecision="281" longitude="-84.08287" name="16 km SSE La Virgen" pageId="37" pageNumber="586" stateProvince="Heredia">
<quantity id="4CBD34D015413268FC342DE7FB80FA23" box="[970,1034,1469,1495]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="586" unit="km" value="16.0">16km</quantity>
SSE La Virgen
</location>
,
<geoCoordinate id="EE71FFF215413268FB422DE7FAAFFA23" box="[1212,1317,1469,1495]" degrees="10.26863" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="200" value="10.26863">10.26863</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FAD42DE4FA16FA2C" box="[1322,1436,1470,1496]" degrees="84.08287" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="200" value="-84.08287">-84.08287</geoCoordinate>
±
<quantity id="4CBD34D015413268FF582DB8FF63FA08" box="[166,233,1506,1532]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="200.0">
<elevation id="00687E0615413268FF582DB8FF63FA08" box="[166,233,1506,1532]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="200.0">200m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FECE2DB8FEF5FA08" box="[304,383,1506,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="37" pageNumber="586" unit="m" value="1110.0">
<elevation id="00687E0615413268FECE2DB8FEF5FA08" box="[304,383,1506,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="37" pageNumber="586" unit="m" value="1110.0">1110m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FE742DB8FD8CFA08" box="[394,518,1506,1532]" pageId="37" pageNumber="586" value="2001-03-14">
<collectingDate id="EFBF461D15413268FE742DB8FD8CFA08" box="[394,518,1506,1532]" pageId="37" pageNumber="586" value="2001-03-14">14.iii.2001</collectingDate>
</date>
, montane wet forest, leaf litter, ALAS#11/WF/02;
<collectingRegion id="498157D715413268FBBE2DB8FB16FA08" box="[1088,1180,1506,1532]" country="Costa Rica" name="Heredia" pageId="37" pageNumber="586">Heredia</collectingRegion>
,
<location id="8E9ACFEE15413268FB592DBBFA13FA08" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FB592DBBFA13FA08" box="[1191,1433,1505,1532]" country="Costa Rica" latitude="10.26863" longLatPrecision="281" longitude="-84.08287" name="16 km SSE La Virgen" pageId="37" pageNumber="586" stateProvince="Heredia">
<quantity id="4CBD34D015413268FB592DBBFB6CFA0F" box="[1191,1254,1505,1531]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="37" pageNumber="586" unit="km" value="16.0">16km</quantity>
SSE La Virgen
</location>
,
<geoCoordinate id="EE71FFF215413268FF692E5FFE8AF9EB" box="[151,256,1541,1567]" degrees="10.26863" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="200" value="10.26863">10.26863</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FEF82E5CFEF2F9D4" box="[262,376,1542,1568]" degrees="84.08287" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="200" value="-84.08287">-84.08287</geoCoordinate>
±
<quantity id="4CBD34D015413268FE702E5CFE5AF9D4" box="[398,464,1542,1568]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="200.0">
<elevation id="00687E0615413268FE702E5CFE5AF9D4" box="[398,464,1542,1568]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="200.0">200m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FDEA2E5CFDE9F9D4" box="[532,611,1542,1568]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="37" pageNumber="586" unit="m" value="1110.0">
<elevation id="00687E0615413268FDEA2E5CFDE9F9D4" box="[532,611,1542,1568]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="37" pageNumber="586" unit="m" value="1110.0">1110m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FD932E5CFD63F9D4" box="[621,745,1542,1568]" pageId="37" pageNumber="586" value="2001-03-14">
<collectingDate id="EFBF461D15413268FD932E5CFD63F9D4" box="[621,745,1542,1568]" pageId="37" pageNumber="586" value="2001-03-14">14.iii.2001</collectingDate>
</date>
, montane wet forest, leaf litter, ALAS#15/WF/02;
<collectingRegion id="498157D715413268FAE42E5CFA12F9D4" box="[1306,1432,1542,1568]" country="Costa Rica" name="Puntarenas" pageId="37" pageNumber="586">Puntarenas</collectingRegion>
,
<location id="8E9ACFEE15413268FF692E70FEABF9B0" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FF692E70FEABF9B0" box="[151,289,1578,1604]" country="Costa Rica" latitude="10.30327" longLatPrecision="70" longitude="-84.81099" name="Monteverde" pageId="37" pageNumber="586" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="EE71FFF215413268FED22E70FE1FF9B0" box="[300,405,1578,1604]" degrees="10.30327" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="10.30327">10.30327</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FE622E70FD85F9B0" box="[412,527,1578,1604]" degrees="84.81099" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-84.81099">-84.81099</geoCoordinate>
±
<quantity id="4CBD34D015413268FDDB2E73FDD3F9B0" box="[549,601,1577,1604]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FDDB2E73FDD3F9B0" box="[549,601,1577,1604]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FD5E2E70FD7AF9B0" box="[672,752,1578,1604]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="586" unit="m" value="1400.0">
<elevation id="00687E0615413268FD5E2E70FD7AF9B0" box="[672,752,1578,1604]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="586" unit="m" value="1400.0">1400m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FD052E73FCD0F9B7" box="[763,858,1577,1603]" pageId="37" pageNumber="586" value="2020-01-06">
<collectingDate id="EFBF461D15413268FD052E73FCD0F9B7" box="[763,858,1577,1603]" pageId="37" pageNumber="586" value="2020-01-06">6.i.2020</collectingDate>
</date>
, moist forest, leaf litter,
<collectorName id="26B0FCE315413268FB942E70FB72F9B0" box="[1130,1272,1578,1604]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL10993-s;
<collectingRegion id="498157D715413268FF692E14FE9FF99C" box="[151,277,1614,1640]" country="Costa Rica" name="Puntarenas" pageId="37" pageNumber="586">Puntarenas</collectingRegion>
,
<location id="8E9ACFEE15413268FEDF2E14FE21F99C" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FEDF2E14FE21F99C" box="[289,427,1614,1640]" country="Costa Rica" latitude="10.30259" longLatPrecision="70" longitude="-84.79269" name="Monteverde" pageId="37" pageNumber="586" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="EE71FFF215413268FE492E17FDAAF99C" box="[439,544,1613,1640]" degrees="10.30259" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="10.30259">10.30259</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FDD92E17FD13F993" box="[551,665,1613,1639]" degrees="84.79269" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-84.79269">-84.79269</geoCoordinate>
±
<quantity id="4CBD34D015413268FD4F2E17FD6FF99C" box="[689,741,1613,1640]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FD4F2E17FD6FF99C" box="[689,741,1613,1640]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FCD22E17FCF7F99C" box="[812,893,1613,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="37" pageNumber="586" unit="m" value="1590.0">
<elevation id="00687E0615413268FCD22E17FCF7F99C" box="[812,893,1613,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.59" pageId="37" pageNumber="586" unit="m" value="1590.0">1590m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FC772E14FC70F99C" box="[905,1018,1614,1640]" pageId="37" pageNumber="586" value="2014-05-13">
<collectingDate id="EFBF461D15413268FC772E14FC70F99C" box="[905,1018,1614,1640]" pageId="37" pageNumber="586" value="2014-05-13">13.v.2014</collectingDate>
</date>
, cloud forest, leaf litter,
<collectorName id="26B0FCE315413268FAF02E14FA16F99C" box="[1294,1436,1614,1640]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL8625-s;
<collectingRegion id="498157D715413268FED02E28FE26F978" box="[302,428,1650,1676]" country="Costa Rica" name="Puntarenas" pageId="37" pageNumber="586">Puntarenas</collectingRegion>
,
<location id="8E9ACFEE15413268FE492E28FDCBF978" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FE492E28FDCBF978" box="[439,577,1650,1676]" country="Costa Rica" latitude="10.28987" longLatPrecision="70" longitude="-84.79322" name="Monteverde" pageId="37" pageNumber="586" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="EE71FFF215413268FDB32E28FD3CF978" box="[589,694,1650,1676]" degrees="10.28987" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="10.28987">10.28987</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FD402E28FCBAF978" box="[702,816,1650,1676]" degrees="84.79322" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-84.79322">-84.79322</geoCoordinate>
±
<quantity id="4CBD34D015413268FCB92E2BFCF1F978" box="[839,891,1649,1676]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FCB92E2BFCF1F978" box="[839,891,1649,1676]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FC3D2E28FB99F978" box="[963,1043,1650,1676]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="37" pageNumber="586" unit="m" value="1310.0">
<elevation id="00687E0615413268FC3D2E28FB99F978" box="[963,1043,1650,1676]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="37" pageNumber="586" unit="m" value="1310.0">1310m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FBE12E28FB1AF978" box="[1055,1168,1650,1676]" pageId="37" pageNumber="586" value="2014-05-13">
<collectingDate id="EFBF461D15413268FBE12E28FB1AF978" box="[1055,1168,1650,1676]" pageId="37" pageNumber="586" value="2014-05-13">13.v.2014</collectingDate>
</date>
, moist forest, leaf litter,
<collectorName id="26B0FCE315413268FF692ECCFEA0F944" box="[151,298,1686,1712]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL8631-s;
<collectingRegion id="498157D715413268FE2E2ECCFDC4F944" box="[464,590,1686,1712]" country="Costa Rica" name="Puntarenas" pageId="37" pageNumber="586">Puntarenas</collectingRegion>
,
<location id="8E9ACFEE15413268FDA02ECCFD62F944" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FDA02ECCFD62F944" box="[606,744,1686,1712]" country="Costa Rica" latitude="10.28744" longLatPrecision="70" longitude="-84.79266" name="Monteverde" pageId="37" pageNumber="586" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="EE71FFF215413268FD062ECCFCEBF944" box="[760,865,1686,1712]" degrees="10.28744" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="10.28744">10.28744</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FC922ECFFC55F95B" box="[876,991,1685,1711]" degrees="84.79266" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-84.79266">-84.79266</geoCoordinate>
±
<quantity id="4CBD34D015413268FC042ECFFBA4F944" box="[1018,1070,1685,1712]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FC042ECFFBA4F944" box="[1018,1070,1685,1712]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FB802ECCFB47F944" box="[1150,1229,1686,1712]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="37" pageNumber="586" unit="m" value="1180.0">
<elevation id="00687E0615413268FB802ECCFB47F944" box="[1150,1229,1686,1712]" metricMagnitude="3" metricUnit="m" metricValue="1.18" pageId="37" pageNumber="586" unit="m" value="1180.0">1180m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FB232ECCFAC4F944" box="[1245,1358,1686,1712]" pageId="37" pageNumber="586" value="2014-05-13">
<collectingDate id="EFBF461D15413268FB232ECCFAC4F944" box="[1245,1358,1686,1712]" pageId="37" pageNumber="586" value="2014-05-13">13.v.2014</collectingDate>
</date>
, moist forest, leaf litter,
<collectorName id="26B0FCE315413268FEA62EE0FE6CF920" box="[344,486,1722,1748]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL8632-s;
<collectingRegion id="498157D715413268FD7F2EE0FD69F920" box="[641,739,1722,1748]" country="Costa Rica" name="Alajuela" pageId="37" pageNumber="586">Alajuela</collectingRegion>
,
<location id="8E9ACFEE15413268FD102EE3FC43F920" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FD102EE3FC43F920" box="[750,969,1721,1748]" country="Costa Rica" latitude="10.29824" longLatPrecision="70" longitude="-84.78163" name="5 km E Monteverde" pageId="37" pageNumber="586" stateProvince="Alajuela">
<quantity id="4CBD34D015413268FD102EE3FCAAF920" box="[750,800,1721,1748]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="km" value="5.0">5km</quantity>
E Monteverde
</location>
,
<geoCoordinate id="EE71FFF215413268FC2A2EE0FBB7F920" box="[980,1085,1722,1748]" degrees="10.29824" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="10.29824">10.29824</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FBBA2EE3FB3CF927" box="[1092,1206,1721,1747]" degrees="84.78163" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-84.78163">-84.78163</geoCoordinate>
±
<quantity id="4CBD34D015413268FB332EE3FA8BF920" box="[1229,1281,1721,1748]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FB332EE3FA8BF920" box="[1229,1281,1721,1748]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FAB92EE3FA12F920" box="[1351,1432,1721,1748]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="37" pageNumber="586" unit="m" value="1495.0">
<elevation id="00687E0615413268FAB92EE3FA12F920" box="[1351,1432,1721,1748]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="37" pageNumber="586" unit="m" value="1495.0">1495m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FF692E84FE82F90C" box="[151,264,1758,1784]" pageId="37" pageNumber="586" value="2014-05-18">
<collectingDate id="EFBF461D15413268FF692E84FE82F90C" box="[151,264,1758,1784]" pageId="37" pageNumber="586" value="2014-05-18">18.v.2014</collectingDate>
</date>
, cloud forest, leaf litter,
<collectorName id="26B0FCE315413268FDE42E84FD22F90C" box="[538,680,1758,1784]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL8698-s;
<collectingRegion id="498157D715413268FCB82E84FC20F90C" box="[838,938,1758,1784]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FC4B2E87FAAFF90C" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FC4B2E87FAAFF90C" box="[949,1317,1757,1784]" country="Costa Rica" latitude="9.58689" longLatPrecision="70" longitude="-83.94758" name="7.5 km SSE Santa Maria de Dota" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FC4B2E87FC76F90C" box="[949,1020,1757,1784]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="37" pageNumber="586" unit="km" value="7.5">7.5km</quantity>
SSE Santa Maria de Dota
</location>
,
<geoCoordinate id="EE71FFF215413268FACF2E87FA06F90C" box="[1329,1420,1757,1784]" degrees="9.58689" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="50" value="9.58689">9.58689</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FA6D2E84FE8AF8E8" degrees="83.94758" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="50" value="-83.94758">- 83.94758</geoCoordinate>
±
<quantity id="4CBD34D015413268FEEB2F5BFEC3F8E8" box="[277,329,1793,1820]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">
<elevation id="00687E0615413268FEEB2F5BFEC3F8E8" box="[277,329,1793,1820]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="586" unit="m" value="50.0">50m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FE732F5BFE57F8E8" box="[397,477,1793,1820]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="37" pageNumber="586" unit="m" value="1520.0">
<elevation id="00687E0615413268FE732F5BFE57F8E8" box="[397,477,1793,1820]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="37" pageNumber="586" unit="m" value="1520.0">1520m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FE192F58FDD2F8E8" box="[487,600,1794,1820]" pageId="37" pageNumber="586" value="2014-05-27">
<collectingDate id="EFBF461D15413268FE192F58FDD2F8E8" box="[487,600,1794,1820]" pageId="37" pageNumber="586" value="2014-05-27">27.v.2014</collectingDate>
</date>
, 2nd growth cloud forest, leaf litter,
<collectorName id="26B0FCE315413268FC182F58FBE7F8E8" box="[998,1133,1794,1820]" pageId="37" pageNumber="586">T. Sumnicht</collectorName>
&amp;
<collectorName id="26B0FCE315413268FB702F58FA90F8E8" box="[1166,1306,1794,1820]" pageId="37" pageNumber="586">J.T. Longino</collectorName>
#JTL8704- s;
<collectingRegion id="498157D715413268FF4D2F7CFE93F8B4" box="[179,281,1830,1856]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FED92F7CFDBCF8B4" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FED92F7CFDBCF8B4" box="[295,566,1830,1856]" country="Costa Rica" latitude="9.55387" longLatPrecision="14" longitude="-83.94547" name="11 km SSE Santa Maria" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FED92F7CFEECF8B4" box="[295,358,1830,1856]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="586" unit="km" value="11.0">11km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF215413268FDBD2F7FFD14F8B4" box="[579,670,1829,1856]" degrees="9.55387" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="10" value="9.55387">9.55387</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FD592F7FFC93F8B4" box="[679,793,1829,1856]" degrees="83.94547" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="10" value="-83.94547">-83.94547</geoCoordinate>
±
<quantity id="4CBD34D015413268FCCC2F7CFCECF8B4" box="[818,870,1830,1856]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">
<elevation id="00687E0615413268FCCC2F7CFCECF8B4" box="[818,870,1830,1856]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">10m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FC4F2F7CFB8BF8B4" box="[945,1025,1830,1856]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">
<elevation id="00687E0615413268FC4F2F7CFB8BF8B4" box="[945,1025,1830,1856]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">1940m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FBF12F7FFB03F8B4" box="[1039,1161,1829,1856]" pageId="37" pageNumber="586" value="2015-06-24">
<collectingDate id="EFBF461D15413268FBF12F7FFB03F8B4" box="[1039,1161,1829,1856]" pageId="37" pageNumber="586" value="2015-06-24">24.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wa-E-04-1-21;
<collectingRegion id="498157D715413268FE3E2F10FDACF890" box="[448,550,1866,1892]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FDCA2F10FCC9F890" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FDCA2F10FCC9F890" box="[564,835,1866,1892]" country="Costa Rica" latitude="9.55378" longLatPrecision="14" longitude="-83.94551" name="11 km SSE Santa Maria" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FDCA2F10FDF9F890" box="[564,627,1866,1892]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="586" unit="km" value="11.0">11km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF215413268FCAE2F13FC21F890" box="[848,939,1865,1892]" degrees="9.55378" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="10" value="9.55378">9.55378</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FC4B2F13FBADF890" box="[949,1063,1865,1892]" degrees="83.94551" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="10" value="-83.94551">-83.94551</geoCoordinate>
±
<quantity id="4CBD34D015413268FBBF2F10FBFEF890" box="[1089,1140,1866,1892]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">
<elevation id="00687E0615413268FBBF2F10FBFEF890" box="[1089,1140,1866,1892]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">10m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FB412F10FA9AF890" box="[1215,1296,1866,1892]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">
<elevation id="00687E0615413268FB412F10FA9AF890" box="[1215,1296,1866,1892]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">1940m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FAE32F13FA12F890" box="[1309,1432,1865,1892]" pageId="37" pageNumber="586" value="2015-06-24">
<collectingDate id="EFBF461D15413268FAE32F13FA12F890" box="[1309,1432,1865,1892]" pageId="37" pageNumber="586" value="2015-06-24">24.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wa-E-04-1-24;
<collectingRegion id="498157D715413268FD412F34FCA9F87C" box="[703,803,1902,1928]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FCD02F34FBBFF87C" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FCD02F34FBBFF87C" box="[814,1077,1902,1928]" country="Costa Rica" latitude="9.55296" longLatPrecision="14" longitude="-83.94551" name="11 km SSE Santa Maria" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FCD02F34FCE7F87C" box="[814,877,1902,1928]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="37" pageNumber="586" unit="km" value="11.0">11km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF215413268FBC12F37FB10F87C" box="[1087,1178,1901,1928]" degrees="9.55296" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="10" value="9.55296">9.55296</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FB5F2F37FA99F87C" box="[1185,1299,1901,1928]" degrees="83.94551" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="10" value="-83.94551">-83.94551</geoCoordinate>
±
<quantity id="4CBD34D015413268FAD42F34FAD7F87C" box="[1322,1373,1902,1928]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">
<elevation id="00687E0615413268FAD42F34FAD7F87C" box="[1322,1373,1902,1928]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="586" unit="m" value="10.0">10m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FF692FC8FF6DF858" box="[151,231,1938,1964]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">
<elevation id="00687E0615413268FF692FC8FF6DF858" box="[151,231,1938,1964]" metricMagnitude="3" metricUnit="m" metricValue="1.94" pageId="37" pageNumber="586" unit="m" value="1940.0">1940m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FF082FCBFEFBF858" box="[246,369,1937,1964]" pageId="37" pageNumber="586" value="2015-06-24">
<collectingDate id="EFBF461D15413268FF082FCBFEFBF858" box="[246,369,1937,1964]" pageId="37" pageNumber="586" value="2015-06-24">24.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wa-E-04-1-42;
<collectingRegion id="498157D715413268FC472FC8FBAAF858" box="[953,1056,1938,1964]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FBD12FC8FAB9F858" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FBD12FC8FAB9F858" box="[1071,1331,1938,1964]" country="Costa Rica" latitude="9.58425" longLatPrecision="28" longitude="-83.94826" name="8 km SSE Santa Maria" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FBD12FC8FBEAF858" box="[1071,1120,1938,1964]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="586" unit="km" value="8.0">8km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF215413268FABF2FCBFA16F858" box="[1345,1436,1937,1964]" degrees="9.58425" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="20" value="9.58425">9.58425</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FF692FEFFE83F83B" box="[151,265,1973,1999]" degrees="83.94826" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="20" value="-83.94826">-83.94826</geoCoordinate>
±
<quantity id="4CBD34D015413268FEDF2FECFEDFF824" box="[289,341,1974,2000]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="20.0">
<elevation id="00687E0615413268FEDF2FECFEDFF824" box="[289,341,1974,2000]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="20.0">20m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FE612FEFFE65F824" box="[415,495,1973,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="37" pageNumber="586" unit="m" value="1580.0">
<elevation id="00687E0615413268FE612FEFFE65F824" box="[415,495,1973,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="37" pageNumber="586" unit="m" value="1580.0">1580m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FE022FEFFDFCF824" box="[508,630,1973,2000]" pageId="37" pageNumber="586" value="2015-06-23">
<collectingDate id="EFBF461D15413268FE022FEFFDFCF824" box="[508,630,1973,2000]" pageId="37" pageNumber="586" value="2015-06-23">23.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wm-E-04-1-02;
<collectingRegion id="498157D715413268FB3E2FECFAAFF824" box="[1216,1317,1974,2000]" country="Costa Rica" name="San Jose" pageId="37" pageNumber="586">San José</collectingRegion>
,
<location id="8E9ACFEE15413268FACC2FECFEAFF800" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE15413268FACC2FECFEAFF800" country="Costa Rica" latitude="9.57877" longLatPrecision="28" longitude="-83.95085" name="8 km SSE Santa Maria" pageId="37" pageNumber="586" stateProvince="San Jose">
<quantity id="4CBD34D015413268FACC2FECFAEEF824" box="[1330,1380,1974,2000]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="586" unit="km" value="8.0">8km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF215413268FECC2F83FE07F800" box="[306,397,2009,2036]" degrees="9.57877" direction="north" orientation="latitude" pageId="37" pageNumber="586" precision="20" value="9.57877">9.57877</geoCoordinate>
<geoCoordinate id="EE71FFF215413268FE692F83FD83F800" box="[407,521,2009,2036]" degrees="83.95085" direction="west" orientation="longitude" pageId="37" pageNumber="586" precision="20" value="-83.95085">-83.95085</geoCoordinate>
±
<quantity id="4CBD34D015413268FDDC2F80FDDCF800" box="[546,598,2010,2036]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="20.0">
<elevation id="00687E0615413268FDDC2F80FDDCF800" box="[546,598,2010,2036]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="586" unit="m" value="20.0">20m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D015413268FD5F2F83FD7BF807" box="[673,753,2009,2035]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="37" pageNumber="586" unit="m" value="1620.0">
<elevation id="00687E0615413268FD5F2F83FD7BF807" box="[673,753,2009,2035]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="37" pageNumber="586" unit="m" value="1620.0">1620m</elevation>
</quantity>
,
<date id="FFFBBFF515413268FD012F83FCF3F800" box="[767,889,2009,2036]" pageId="37" pageNumber="586" value="2015-06-25">
<collectingDate id="EFBF461D15413268FD012F83FCF3F800" box="[767,889,2009,2036]" pageId="37" pageNumber="586" value="2015-06-25">25.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wm-E-04-1- 06;
<collectingRegion id="498157D71542326BFF3A28CDFEA0FF45" box="[196,298,151,177]" country="Costa Rica" name="San Jose" pageId="38" pageNumber="587">San José</collectingRegion>
,
<location id="8E9ACFEE1542326BFEC928CDFDB2FF45" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE1542326BFEC928CDFDB2FF45" box="[311,568,151,177]" country="Costa Rica" latitude="9.56191" longLatPrecision="28" longitude="-83.94807" name="9 km SSE Santa Maria" pageId="38" pageNumber="587" stateProvince="San Jose">
<quantity id="4CBD34D01542326BFEC928CDFEE3FF45" box="[311,361,151,177]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="587" unit="km" value="9.0">9km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF21542326BFDBB28CCFD2AFF45" box="[581,672,150,177]" degrees="9.56191" direction="north" orientation="latitude" pageId="38" pageNumber="587" precision="20" value="9.56191">9.56191</geoCoordinate>
<geoCoordinate id="EE71FFF21542326BFD5728CDFC91FF45" box="[681,795,151,177]" degrees="83.94807" direction="west" orientation="longitude" pageId="38" pageNumber="587" precision="20" value="-83.94807">-83.94807</geoCoordinate>
±
<quantity id="4CBD34D01542326BFCCA28CDFCE2FF45" box="[820,872,151,177]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="587" unit="m" value="20.0">
<elevation id="00687E061542326BFCCA28CDFCE2FF45" box="[820,872,151,177]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="587" unit="m" value="20.0">20m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D01542326BFC4C28CDFB88FF45" box="[946,1026,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="38" pageNumber="587" unit="m" value="1830.0">
<elevation id="00687E061542326BFC4C28CDFB88FF45" box="[946,1026,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="38" pageNumber="587" unit="m" value="1830.0">1830m</elevation>
</quantity>
,
<date id="FFFBBFF51542326BFBEE28CCFB00FF45" box="[1040,1162,150,177]" pageId="38" pageNumber="587" value="2015-06-25">
<collectingDate id="EFBF461D1542326BFBEE28CCFB00FF45" box="[1040,1162,150,177]" pageId="38" pageNumber="587" value="2015-06-25">25.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wm-E-04-1-09;
<collectingRegion id="498157D71542326BFE3428E1FDA5FF21" box="[458,559,187,213]" country="Costa Rica" name="San Jose" pageId="38" pageNumber="587">San José</collectingRegion>
,
<location id="8E9ACFEE1542326BFDC228E1FCC3FF21" LSID="urn:lsid:plazi:treatment:03EC28231547326AFF692CA2FD45FD95:8E9ACFEE1542326BFDC228E1FCC3FF21" box="[572,841,187,213]" country="Costa Rica" latitude="9.55908" longLatPrecision="28" longitude="-83.94646" name="10 km SSE Santa Maria" pageId="38" pageNumber="587" stateProvince="San Jose">
<quantity id="4CBD34D01542326BFDC228E1FDF6FF21" box="[572,636,187,213]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="587" unit="km" value="10.0">10km</quantity>
SSE Santa María
</location>
,
<geoCoordinate id="EE71FFF21542326BFCA828E0FC3BFF21" box="[854,945,186,213]" degrees="9.55908" direction="north" orientation="latitude" pageId="38" pageNumber="587" precision="20" value="9.55908">9.55908</geoCoordinate>
<geoCoordinate id="EE71FFF21542326BFC4728E1FBA1FF21" box="[953,1067,187,213]" degrees="83.94646" direction="west" orientation="longitude" pageId="38" pageNumber="587" precision="20" value="-83.94646">-83.94646</geoCoordinate>
±
<quantity id="4CBD34D01542326BFBBD28E1FBFDFF21" box="[1091,1143,187,213]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="587" unit="m" value="20.0">
<elevation id="00687E061542326BFBBD28E1FBFDFF21" box="[1091,1143,187,213]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="587" unit="m" value="20.0">20m</elevation>
</quantity>
, elev.
<quantity id="4CBD34D01542326BFB3E28E1FA9BFF21" box="[1216,1297,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="38" pageNumber="587" unit="m" value="1870.0">
<elevation id="00687E061542326BFB3E28E1FA9BFF21" box="[1216,1297,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="38" pageNumber="587" unit="m" value="1870.0">1870m</elevation>
</quantity>
,
<date id="FFFBBFF51542326BFAE328E0FA1DFF21" box="[1309,1431,186,213]" pageId="38" pageNumber="587" value="2015-06-25">
<collectingDate id="EFBF461D1542326BFAE328E0FA1DFF21" box="[1309,1431,186,213]" pageId="38" pageNumber="587" value="2015-06-25">25.vi.2015</collectingDate>
</date>
, cloud forest, leaf litter, ADMAC#Wm-E-04-1-10.
</paragraph>
</subSubSection>
<caption id="DF3AC9BD1542326BFF662E77FE78F99E" ID-DOI="http://doi.org/10.5281/zenodo.14734332" ID-Zenodo-Dep="14734332" httpUri="https://zenodo.org/record/14734332/files/figure.png" pageId="38" pageNumber="587" startId="38.[152,270,1581,1605]" targetBox="[151,1434,274,1558]" targetPageId="38" targetType="figure">
<paragraph id="8BFA99351542326BFF662E77FE78F99E" blockId="38.[151,1437,1581,1642]" pageId="38" pageNumber="587">
<emphasis id="B93145271542326BFF662E77FED1F9B1" bold="true" box="[152,347,1581,1605]" pageId="38" pageNumber="587">FIGURES 67–70.</emphasis>
<taxonomicName id="4C45E2B61542326BFE9F2E74FD8EF9B2" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[353,516,1582,1606]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B93145271542326BFE9F2E74FD8EF9B2" box="[353,516,1582,1606]" italics="true" pageId="38" pageNumber="587">Nylanderia usul</emphasis>
</taxonomicName>
paratype male (USNMENT01131831). Full-face and lateral view of the body; ventral and dorsal view of the genital capsule.
</paragraph>
</caption>
<subSubSection id="C35FCABE1542326BFF392EE6FE7CF87E" pageId="38" pageNumber="587" type="etymology">
<paragraph id="8BFA99351542326BFF392EE6FE7CF87E" blockId="38.[151,1437,1724,2038]" pageId="38" pageNumber="587">
<emphasis id="B93145271542326BFF392EE6FED8F922" bold="true" box="[199,338,1724,1750]" pageId="38" pageNumber="587">Etymology:</emphasis>
The species epithet is derived from Frank Herbert’s science fiction novel Dune. In the novel, “Usul” is the secret name given to the protagonist, Paul Atreides, by the Fremen, the desert dwellers of the planet Arrakis. This name references the species’ moderate to intense blue iridescence, alluding to the “blue-within-blue” eyes that Paul acquires after joining the Fremen and consuming the spice melange. The term “Usul” translates to “the strength of the base of the pillar” and is derived from the Arabic word
<taxonomicName id="4C45E2B61542326BFC282F16FB8FF892" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[982,1029,1868,1894]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B93145271542326BFC282F16FB8FF892" box="[982,1029,1868,1894]" italics="true" pageId="38" pageNumber="587">usul</emphasis>
</taxonomicName>
, the plural form of
<emphasis id="B93145271542326BFB222F16FB77F892" box="[1244,1277,1868,1894]" italics="true" pageId="38" pageNumber="587">asl</emphasis>
, which means “origin,” “root,” or “principle.”
</paragraph>
</subSubSection>
<subSubSection id="C35FCABE1542326AFF392FCEFD45FD95" lastPageId="39" lastPageNumber="588" pageId="38" pageNumber="587" type="discussion">
<paragraph id="8BFA99351542326AFF392FCEFD45FD95" blockId="38.[151,1437,1724,2038]" lastBlockId="39.[151,1437,151,609]" lastPageId="39" lastPageNumber="588" pageId="38" pageNumber="587">
<emphasis id="B93145271542326BFF392FCEFE99F85A" bold="true" box="[199,275,1940,1966]" pageId="38" pageNumber="587">Notes:</emphasis>
<taxonomicName id="4C45E2B61542326BFEE22FCEFE58F85A" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[284,466,1940,1966]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B93145271542326BFEE22FCEFE58F85A" box="[284,466,1940,1966]" italics="true" pageId="38" pageNumber="587">Nylanderia usul</emphasis>
</taxonomicName>
workers most strongly resemble those of
<taxonomicName id="4C45E2B61542326BFC4D2FCFFBA6F85A" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[947,1068,1940,1966]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B93145271542326BFC4D2FCFFBA6F85A" box="[947,1068,1940,1966]" italics="true" pageId="38" pageNumber="587">N. mendax</emphasis>
</taxonomicName>
and
<taxonomicName id="4C45E2B61542326BFB982FCFFB6CF85A" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[1126,1254,1941,1966]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="mosaica">
<emphasis id="B93145271542326BFB982FCFFB6CF85A" box="[1126,1254,1941,1966]" italics="true" pageId="38" pageNumber="587">N. mosaica</emphasis>
</taxonomicName>
. This species is most readily separable from
<taxonomicName id="4C45E2B61542326BFE2B2FE3FDC6F826" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[469,588,1976,2002]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B93145271542326BFE2B2FE3FDC6F826" box="[469,588,1976,2002]" italics="true" pageId="38" pageNumber="587">N. mendax</emphasis>
</taxonomicName>
by its relatively short scapes, which only surpass the posterior margin of the head by a quarter to a third of their length, whereas in
<taxonomicName id="4C45E2B61542326BFCFF2F87FCF3F802" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[769,889,2012,2038]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="587" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B93145271542326BFCFF2F87FCF3F802" box="[769,889,2012,2038]" italics="true" pageId="38" pageNumber="587">N. mendax</emphasis>
</taxonomicName>
they surpass the posterior margin by nearly half their length. There are two distinct color morphs of this species. The dark form is dark brown with yellow to yellow-brown mandibles and tarsi, meso- and metacoxae somewhat lighter than the mesosoma, and has moderate cephalic pubescence, concentrated mostly as patches around the compound eyes (
<figureCitation id="137E85B01543326AFC182885FBC8FF0D" box="[998,1090,223,249]" captionStart="FIGURE 4" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[378,1207,1557,1916]" captionTargetId="figure-1745@7.[372,1215,1550,1919]" captionTargetPageId="7" captionText="FIGURE 4. Line drawings depicting examples of cephalic pubescence patterns in full-face view: (A) N. austroccidua with uniformly dense cephalic pubescence; (B) N. lazulina with moderate pubescence, concentrated primarily in areas anterior and posterior to the compound eyes; and (C) N. cf. docilis with sparse cephalic pubescence." figureDoi="http://doi.org/10.5281/zenodo.14734278" httpUri="https://zenodo.org/record/14734278/files/figure.png" pageId="39" pageNumber="588">Fig. 4B</figureCitation>
). The light form is uniformly yellow, with dense cephalic pubescence (
<figureCitation id="137E85B01543326AFD942959FD4CFEE9" box="[618,710,259,285]" captionStart="FIGURE 4" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[378,1207,1557,1916]" captionTargetId="figure-1745@7.[372,1215,1550,1919]" captionTargetPageId="7" captionText="FIGURE 4. Line drawings depicting examples of cephalic pubescence patterns in full-face view: (A) N. austroccidua with uniformly dense cephalic pubescence; (B) N. lazulina with moderate pubescence, concentrated primarily in areas anterior and posterior to the compound eyes; and (C) N. cf. docilis with sparse cephalic pubescence." figureDoi="http://doi.org/10.5281/zenodo.14734278" httpUri="https://zenodo.org/record/14734278/files/figure.png" pageId="39" pageNumber="588">Fig. 4A</figureCitation>
). The light form is more likely to be confused with
<taxonomicName id="4C45E2B61543326AFAE82959FA1FFEE8" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[1302,1429,259,284]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="588" phylum="Arthropoda" rank="species" species="mosaica">
<emphasis id="B93145271543326AFAE82959FA1FFEE8" box="[1302,1429,259,284]" italics="true" pageId="39" pageNumber="588">N. mosaica</emphasis>
</taxonomicName>
, but the latter is smaller in overall size and bicolored with a yellow to yellow-brown head and mesosoma and medium to dark brown gaster. Interestingly, the light form, represented by NT002, NT003, NT020, and NT024 in the phylogeny (
<figureCitation id="137E85B01543326AFEB92935FE06FE7D" box="[327,396,367,393]" captionStart="FIGURE 3" captionStartId="6.[152,255,1895,1919]" captionTargetBox="[166,1423,181,1871]" captionTargetId="figure-26@6.[164,1423,181,1871]" captionTargetPageId="6" captionText="FIGURE 3. Maximum likelihood phylogeny of clade AC1 reconstructed using the 90% complete SWSC partitioned matrix. Nodal support (SH-aLRT/UFBoot) is displayed only for nodes with &lt;95% support for either measure. The map inset in the bottom-right indicates the extent of geographical overlap between the Nearctic and Neotropical subclades around the Isthmus of Tehuantepec." figureDoi="http://doi.org/10.5281/zenodo.14734276" httpUri="https://zenodo.org/record/14734276/files/figure.png" pageId="39" pageNumber="588">Fig. 3</figureCitation>
), resolved as paraphyletic with respect to the dark form. This pattern may indicate a case of incipient speciation, in which one is beginning to diverge from the other but has not yet achieved full reproductive isolation. Consequently, the two forms may be interpretable as two distinct species based on their morphology. However, for now we conservatively treat them all as a single species. If someday these two forms are split into separate species, the dark form will keep the name
<taxonomicName id="4C45E2B61543326AFD3029A5FC97FDED" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[718,797,511,537]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="588" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B93145271543326AFD3029A5FC97FDED" box="[718,797,511,537]" italics="true" pageId="39" pageNumber="588">N. usul</emphasis>
</taxonomicName>
and the light form will take a new name.
<taxonomicName id="4C45E2B61543326AFB1729A5FA17FDED" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" box="[1257,1437,511,537]" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="588" phylum="Arthropoda" rank="species" species="usul">
<emphasis id="B93145271543326AFB1729A5FA17FDED" box="[1257,1437,511,537]" italics="true" pageId="39" pageNumber="588">Nylanderia usul</emphasis>
</taxonomicName>
is only known from moderate to high elevation forests in
<collectingCountry id="F352D9A51543326AFCDD2A79FC17FDC9" box="[803,925,547,573]" name="Costa Rica" pageId="39" pageNumber="588">Costa Rica</collectingCountry>
(
<figureCitation id="137E85B01543326AFC502A79FB8CFDC9" box="[942,1030,547,573]" captionStart="FIGURE 2" captionStartId="5.[152,255,1791,1815]" captionTargetBox="[158,1434,188,1760]" captionTargetId="figure-17@5.[151,1436,181,1767]" captionTargetPageId="5" captionText="FIGURE 2. Distributions of species: (A) N. austroccidua in the United States and northern Mexico, (B) species found across southern Mexico and Mesoamerica, and (C) N. docilis in South America. Maps were created using GPS coordinates listed in Table S1 and from the Global Ant Biodiversity Informatics (GABI; Guénard et al. 2017) database." figureDoi="http://doi.org/10.5281/zenodo.14734274" httpUri="https://zenodo.org/record/14734274/files/figure.png" pageId="39" pageNumber="588">Fig. 2B</figureCitation>
), where it is sympatric with only
<taxonomicName id="4C45E2B61543326AFA7D2A79FF7EFD94" authorityName="Williams &amp; Williams &amp; Lapolla &amp; Lucky" authorityYear="2025" class="Insecta" family="Formicidae" genus="Nylanderia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="588" phylum="Arthropoda" rank="species" species="mosaica">
<emphasis id="B93145271543326AFA7D2A79FF7EFD94" italics="true" pageId="39" pageNumber="588">N. mosaica</emphasis>
</taxonomicName>
and no other species in American Clade I.
</paragraph>
</subSubSection>
</treatment>
</document>
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