491 lines
69 KiB
XML
491 lines
69 KiB
XML
<document id="CC14B814CD872C3E2DBDE5B876EC74ED" ID-CLB-Dataset="193695" ID-DOI="10.11646/zootaxa.5209.5.1" ID-GBIF-Dataset="80fa12f4-e62b-487d-91b1-b1ec37da5ba4" ID-ISSN="1175-5326" ID-Zenodo-Dep="7469353" ID-ZooBank="360B047E-5396-48AE-B26F-00B34BCB835A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1669021335121" checkinUser="plazi" docAuthor="Silva, Ruan Felipe Da, Zilberman, Bruno & Carvalho-Filho, Fernando Da Silva" docDate="2022" docId="03E787F8EB3CFFB45DF0F9BAFB843931" docLanguage="en" docName="zootaxa.5209.5.1.pdf" docOrigin="Zootaxa 5209 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Termitozophilus tenebrus Silva & Zilberman & Carvalho-Filho 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="529" masterDocId="FFDEFF80EB29FFA85D67FFC9FFD63B50" masterDocTitle="Hidden Brazilian Amazon diversity: four new species, redescription and notes on natural history of Termitozophilus Silvestri, 1901 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="534" masterPageNumber="501" pageNumber="522" updateTime="1699411831406" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="815DC162777181F0179873364D30FCAF">Hidden Brazilian Amazon diversity: four new species, redescription and notes on natural history of Termitozophilus Silvestri, 1901 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="4925DE16905713911494EBC55FC35DB9" type="personal">
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<mods:roleTerm id="3E97F2D8EBBF355DB85231770BFACCE3">Author</mods:roleTerm>
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<mods:namePart id="71550AB11414B95719675E890D84F5DC">Silva, Ruan Felipe Da</mods:namePart>
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<mods:nameIdentifier id="87E074D4CF777103D9D46421BA5DC669" type="ORCID">0000-0002-7096-226X</mods:nameIdentifier>
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<mods:affiliation id="C7F16E79B0CE0DD5573F3312231446BC">Programa de Pós-graduação em Biodiversidade e Evolução. & Museu Paraense Emílio Goeldi. Coordenação de Zoologia. Av. Perimetral, 1901, 66077 - 830, Belém, Pará, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="77E31683E6B0CDFF04F9DCF7DDCF0BCD" type="email">silvaruanbio@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="3700624E916554063415A1FBD67079B7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="695C63BF4FB27A079A0129A53260D9D3">Zilberman, Bruno</mods:namePart>
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<mods:nameIdentifier id="AAE94FB462669B5CC3AFE41DBE42736B" type="ORCID">0000-0003-2613-4827</mods:nameIdentifier>
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<mods:affiliation id="9E4265836286EDB9EABB6FCD5827A5DE">Museu de Zoologia da Universidade de São Paulo, Caixa Postal 42494; 04218 - 970, São Paulo, Brazil. & Programa de Pós-graduação em Sistemática, Taxonomia Animal e Biodiversidade.</mods:affiliation>
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<mods:nameIdentifier id="2FA72ACAE236BD0EF52E5DAA272C3A93" type="email">brunozilberman@usp.br</mods:nameIdentifier>
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<mods:roleTerm id="A9180760311A6ACD0F9D0CB0189242C3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D004BC77A457029E2706CE6245E9439F">Carvalho-Filho, Fernando Da Silva</mods:namePart>
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<mods:nameIdentifier id="9A79A97EFAF1E57B64CAFBD6B4EBD955" type="ORCID">0000-0002-2480-3874</mods:nameIdentifier>
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<mods:affiliation id="00EEECBCCFCF3531B34CDCF6D7B5BD52">Museu Paraense Emílio Goeldi. Coordenação de Zoologia. Av. Perimetral, 1901, 66077 - 830, Belém, Pará, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="3A7980035D6DFCFA0826B4764F95F065" type="email">fernandofilho@museu-goeldi.br</mods:nameIdentifier>
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<mods:title id="F99D397EB899037D66966077B880E9D1">Zootaxa</mods:title>
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<mods:part id="0795A27BBB388CE47B5773BD997C3ED8">
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<mods:date id="C4AE89E39EDE470F013F916DC1E09698">2022</mods:date>
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<mods:number id="AE24BDD4020EC3045FD0F247AD5B7BB2">2022-11-21</mods:number>
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<treatment id="03E787F8EB3CFFB45DF0F9BAFB843931" ID-DOI="http://doi.org/10.5281/zenodo.7469391" ID-GBIF-Taxon="205134459" ID-Zenodo-Dep="7469391" LSID="urn:lsid:plazi:treatment:03E787F8EB3CFFB45DF0F9BAFB843931" httpUri="http://treatment.plazi.org/id/03E787F8EB3CFFB45DF0F9BAFB843931" lastPageId="28" lastPageNumber="529" pageId="21" pageNumber="522">
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<subSubSection id="C3546565EB3CFFBD5DF0F9BAFDF83DDE" box="[151,558,1651,1678]" pageId="21" pageNumber="522" type="nomenclature">
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<paragraph id="8BF136EEEB3CFFBD5DF0F9BAFDF83DDE" blockId="21.[151,558,1651,1714]" box="[151,558,1651,1678]" pageId="21" pageNumber="522">
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<heading id="D0B98182EB3CFFBD5DF0F9BAFDF83DDE" bold="true" box="[151,558,1651,1678]" fontSize="11" level="1" pageId="21" pageNumber="522" reason="1">
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<emphasis id="B93AEAFCEB3CFFBD5DF0F9BAFDF83DDE" bold="true" box="[151,558,1651,1678]" pageId="21" pageNumber="522">
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<taxonomicName id="4C4E4D6DEB3CFFBD5DF0F9BAFE1F3DDE" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[151,457,1651,1678]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
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<emphasis id="B93AEAFCEB3CFFBD5DF0F9BAFE1F3DDE" bold="true" box="[151,457,1651,1678]" italics="true" pageId="21" pageNumber="522">Termitozophilus tenebrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2095787EB3CFFBD5CB6F9BDFDF83DDE" box="[465,558,1652,1678]" pageId="21" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3546565EB3CFFBD5DF0F951FE453DE2" box="[151,403,1688,1714]" pageId="21" pageNumber="522" type="description">
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<paragraph id="8BF136EEEB3CFFBD5DF0F951FE453DE2" blockId="21.[151,558,1651,1714]" box="[151,403,1688,1714]" pageId="21" pageNumber="522">
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(
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<figureCitation id="13752A6BEB3CFFBD5DF8F951FEF13DE2" box="[159,295,1688,1714]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStart-2="FIGURE 21" captionStart-3="FIGURE 22" captionStart-4="FIGURE 23" captionStart-5="FIGURE 24" captionStartId-0="22.[151,250,1744,1768]" captionStartId-1="23.[151,250,1943,1967]" captionStartId-2="24.[151,250,1914,1938]" captionStartId-3="25.[151,250,1857,1881]" captionStartId-4="26.[151,250,1921,1945]" captionStartId-5="27.[151,250,1613,1637]" captionTargetBox-0="[151,1436,181,1720]" captionTargetBox-1="[203,1385,181,1919]" captionTargetBox-2="[151,1431,420,1881]" captionTargetBox-3="[232,1355,181,1833]" captionTargetBox-4="[151,1436,273,1898]" captionTargetBox-5="[151,1436,181,1589]" captionTargetId-0="figure-22@22.[151,1436,181,1720]" captionTargetId-1="figure-19@23.[203,1385,181,1919]" captionTargetId-2="figure-131@24.[151,1436,417,1891]" captionTargetId-3="figure-19@25.[232,1355,181,1833]" captionTargetId-4="figure-64@26.[151,1436,273,1898]" captionTargetId-5="figure-19@27.[151,1436,181,1589]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionTargetPageId-3="25" captionTargetPageId-4="26" captionTargetPageId-5="27" captionText-0="FIGURE 19. Termitozophilus tenebrus sp. nov., male. A) Habitus, lateral view. B) Habitus, dorsal view." captionText-1="FIGURE 20. Termitozophilus tenebrus sp. nov., female. A) Habitus, dorsal view. B) Abdomen, ventral view. C) Habitus, lateral view." captionText-2="FIGURE 21. Termitozophilus tenebrus sp. nov. A) Head, dorsal view. B) Left and right mandibles, dorsal view. C) Labrum, dorsal view. D) Maxilla, dorsal view. E) Pre-mentum, mentum and sub-mentum, ventral view. Scales: A = 2 mm; D = 1,5 mm; E = 1 mm; B, C = 0,5 mm." captionText-3="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." captionText-4="FIGURE 23. Termitozophilus tenebrus sp. nov. A) Elytron, internal view. B) Detail of the anteromesial border of elytron, internal view. C–D) Mandible, dorsal view (arrow showing pores). Scales: A = 100 µm; B = 20 µm; C = 50 µm; D = 10 µm." captionText-5="FIGURE 24. Termitozophilus tenebrus sp. nov. A) Mentum and sub-mentum (arrow showing suture. B) Maxillary palpus. C) Tergite VII. D) Detail of glandular área of tergite VII. Scales: C = 200 µm; A, D = 20 µm; B = 10 µm; B = 2 µm." figureDoi-0="http://doi.org/10.5281/zenodo.7469419" figureDoi-1="http://doi.org/10.5281/zenodo.7469421" figureDoi-2="http://doi.org/10.5281/zenodo.7469423" figureDoi-3="http://doi.org/10.5281/zenodo.7469427" figureDoi-4="http://doi.org/10.5281/zenodo.7469429" figureDoi-5="http://doi.org/10.5281/zenodo.7469431" httpUri-0="https://zenodo.org/record/7469419/files/figure.png" httpUri-1="https://zenodo.org/record/7469421/files/figure.png" httpUri-2="https://zenodo.org/record/7469423/files/figure.png" httpUri-3="https://zenodo.org/record/7469427/files/figure.png" httpUri-4="https://zenodo.org/record/7469429/files/figure.png" httpUri-5="https://zenodo.org/record/7469431/files/figure.png" pageId="21" pageNumber="522">
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Figs. 19–
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</figureCitation>
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,
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<figureCitation id="13752A6BEB3CFFBD5C54F951FEB53DE2" box="[307,355,1688,1714]" captionStart="FIGURE 27" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[234,1355,189,1913]" captionTargetId="figure-22@30.[221,1367,181,1919]" captionTargetPageId="30" captionText="FIGURE 27. Phylogenetic recapitulation hypothesis based on the condition of abdominal sclerites in Termitozophilus species." figureDoi="http://doi.org/10.5281/zenodo.7469437" httpUri="https://zenodo.org/record/7469437/files/figure.png" pageId="21" pageNumber="522">27E</figureCitation>
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,
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<figureCitation id="13752A6BEB3CFFBD5C09F951FE5D3DE2" box="[366,395,1688,1714]" captionStart="FIGURE 28" captionStartId="31.[151,250,1557,1581]" captionTargetBox="[156,1436,675,1526]" captionTargetId="figure-243@31.[152,1436,669,1533]" captionTargetPageId="31" captionText="FIGURE 28. Geographic distribution of species of Termitozophilus in the Brazilian Amazon region (Para)." figureDoi="http://doi.org/10.5281/zenodo.7469441" httpUri="https://zenodo.org/record/7469441/files/figure.png" pageId="21" pageNumber="522">28</figureCitation>
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)
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</paragraph>
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<subSubSection id="C3546565EB3CFFBD5DF0F929FE073C36" pageId="21" pageNumber="522" type="materials_examined">
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<emphasis id="B93AEAFCEB3CFFBD5DF0F929FE903DAA" bold="true" box="[151,326,1760,1786]" pageId="21" pageNumber="522">Type material.</emphasis>
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<materialsCitation id="3B263CB3EB3CFFBD5C29F929FBBD3C4F" ID-GBIF-Occurrence="3970431306" collectingDate="2021-03-11" collectionCode="MPEG, R" collectorName="de Calafate & F. Silva" country="Brazil" location="Vila de Calafate" municipality="Magalhaes Barata" pageId="21" pageNumber="522" specimenCount="1" specimenCount-male="1" stateProvince="Para" typeStatus="holotype">
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<typeStatus id="54F5884CEB3CFFBD5C29F929FE343DAA" box="[334,482,1760,1786]" pageId="21" pageNumber="522" type="holotype">HOLOTYPE</typeStatus>
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<specimenCount id="9D48FD67EB3CFFBD5C8EF928FE283DAB" box="[489,510,1761,1787]" pageId="21" pageNumber="522" type="male">♂</specimenCount>
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(
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<collectionCode id="ED5FAE2BEB3CFFBD5F69F929FDB53DAA" box="[526,611,1760,1786]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="522">MPEG</collectionCode>
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): Brasil [=
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<collectingCountry id="F359767EEB3CFFBD5F85F929FCFA3DAA" box="[738,812,1760,1786]" name="Brazil" pageId="21" pageNumber="522">Brazil</collectingCountry>
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],
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<collectingRegion id="498AF80CEB3CFFBD5E59F929FCA53DAA" box="[830,883,1760,1786]" country="Brazil" name="Para" pageId="21" pageNumber="522">Pará</collectingRegion>
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,
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<collectingMunicipality id="6B95AC94EB3CFFBD5E18F929FB9F3DAB" box="[895,1097,1760,1787]" pageId="21" pageNumber="522">Magalhães Barata</collectingMunicipality>
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/
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<location id="8E916035EB3CFFBD5907F929FAC03DAA" LSID="urn:lsid:plazi:treatment:03E787F8EB3CFFB45DF0F9BAFB843931:8E916035EB3CFFBD5907F929FAC03DAA" box="[1120,1302,1760,1786]" country="Brazil" municipality="Magalhaes Barata" name="Vila de Calafate" pageId="21" pageNumber="522" stateProvince="Para">
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Vila
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<collectorName id="26BB5338EB3CFFBD59F0F929FAC03DAA" box="[1175,1302,1760,1786]" pageId="21" pageNumber="522">de Calafate</collectorName>
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</location>
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[= Calafate Village],
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<date id="FFF0102EEB3CFFBD5D98F8CDFEAA3C4E" box="[255,380,1796,1822]" pageId="21" pageNumber="522" value="2021-03-11">
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<collectingDate id="EFB4E9C6EB3CFFBD5D98F8CDFEAA3C4E" box="[255,380,1796,1822]" pageId="21" pageNumber="522" value="2021-03-11">11.III.2021</collectingDate>
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</date>
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/
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<collectionCode id="ED5FAE2BEB3CFFBD5CF5F8CCFE7E3C4F" box="[402,424,1797,1823]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="522">R</collectionCode>
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.
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||
<collectorName id="26BB5338EB3CFFBD5CD4F8CCFDDE3C4E" box="[435,520,1796,1823]" pageId="21" pageNumber="522">F. Silva</collectorName>
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||
[collector] / hospedeiro [= host]:
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||
<taxonomicName id="4C4E4D6DEB3CFFBD5EE5F8CDFBB23C4E" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[898,1124,1796,1822]" class="Insecta" family="Termitidae" genus="Cornitermes" kingdom="Animalia" order="Blattodea" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="pilosus">
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||
<emphasis id="B93AEAFCEB3CFFBD5EE5F8CDFBB23C4E" box="[898,1124,1796,1822]" italics="true" pageId="21" pageNumber="522">Cornitermes pilosus</emphasis>
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||
</taxonomicName>
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||
.
|
||
</materialsCitation>
|
||
</paragraph>
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||
<paragraph id="8BF136EEEB3CFFBD5DF0F8E1FE073C36" blockId="21.[151,1436,1760,2039]" pageId="21" pageNumber="522">
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<materialsCitation id="3B263CB3EB3CFFBD5DF0F8E1FE1B3C36" ID-GBIF-Occurrence="3970431302" collectingDate="2021-03-11" collectionCode="MPEG, MZSUP" collectorName="de Calafate & F. Silva" country="Brazil" location="Vila de Calafate" municipality="Magalhaes Barata" pageId="21" pageNumber="522" specimenCode="MZSP 21288" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Para" typeStatus="paratype">
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<typeStatus id="54F5884CEB3CFFBD5DF0F8E1FEE33C12" box="[151,309,1832,1858]" pageId="21" pageNumber="522" type="paratype">PARATYPES</typeStatus>
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||
. Same data as holotype,
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||
<specimenCount id="9D48FD67EB3CFFBD5F20F8E1FD503C13" box="[583,646,1832,1859]" pageId="21" pageNumber="522" type="female">5 ♀♀</specimenCount>
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and
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<specimenCount id="9D48FD67EB3CFFBD5FDBF8E1FD2D3C13" box="[700,763,1832,1859]" pageId="21" pageNumber="522" type="male">5 ♂♂</specimenCount>
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in
|
||
<collectionCode id="ED5FAE2BEB3CFFBD5E79F8E1FCBA3C12" box="[798,876,1832,1858]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="522">MPEG</collectionCode>
|
||
(in alcohol 70%),
|
||
<specimenCount id="9D48FD67EB3CFFBD595CF8E1FBB33C13" box="[1083,1125,1832,1859]" pageId="21" pageNumber="522" type="female">1 ♀</specimenCount>
|
||
and
|
||
<specimenCount id="9D48FD67EB3CFFBD59FCF8E1FB133C13" box="[1179,1221,1832,1859]" pageId="21" pageNumber="522" type="male">1 ♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED5FAE2BEB3CFFBD598FF8E1FA923C12" box="[1256,1348,1832,1858]" pageId="21" pageNumber="522">MZSUP</collectionCode>
|
||
(
|
||
<specimenCode id="DBE89E95EB3CFFBD5835F8E1FF083C36" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="21" pageNumber="522" type="Museum">MZSP 21288</specimenCode>
|
||
)(dissected on slides)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3546565EB3CFFBD5DA0F8B9FAF23CA6" pageId="21" pageNumber="522" type="diagnosis">
|
||
<paragraph id="8BF136EEEB3CFFBD5DA0F8B9FAF23CA6" blockId="21.[151,1436,1760,2039]" pageId="21" pageNumber="522">
|
||
<emphasis id="B93AEAFCEB3CFFBD5DA0F8B9FEEC3CDA" bold="true" box="[199,314,1904,1930]" pageId="21" pageNumber="522">Diagnosis</emphasis>
|
||
. Paratergites fused neither to each other nor to tergites and sternites; sclerotized areas mostly dark brown, with a large white membranous area exposed in dorsal view. Head with vertex bearing two pairs of bristles, one behind the eye and a pair on medial-posterior region. Hind wings present in post-imaginal growth. Elytra trapezoidal, with anteromesial border densely scaled. Posterior margin of sternite VIII with two bristles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF316666EB3FFFBE5DF0F919FB693DB8" ID-DOI="http://doi.org/10.5281/zenodo.7469419" ID-Zenodo-Dep="7469419" box="[151,1215,1744,1768]" httpUri="https://zenodo.org/record/7469419/files/figure.png" pageId="22" pageNumber="523" startId="22.[151,250,1744,1768]" targetBox="[151,1436,181,1720]" targetPageId="22">
|
||
<paragraph id="8BF136EEEB3FFFBE5DF0F919FB693DB8" blockId="22.[151,1215,1744,1768]" box="[151,1215,1744,1768]" pageId="22" pageNumber="523">
|
||
<emphasis id="B93AEAFCEB3FFFBE5DF0F919FEF63DB8" bold="true" box="[151,288,1744,1768]" pageId="22" pageNumber="523">FIGURE 19.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB3FFFBE5C41F919FDF03DB8" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[294,550,1744,1768]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="523" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB3FFFBE5C41F919FDF03DB8" box="[294,550,1744,1768]" italics="true" pageId="22" pageNumber="523">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB3FFFBE5F4BF919FDAB3DB8" bold="true" box="[556,637,1744,1768]" pageId="22" pageNumber="523">
|
||
<taxonomicNameLabel id="A2095787EB3FFFBE5F4BF919FDAB3DB8" box="[556,637,1744,1768]" pageId="22" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis id="B93AEAFCEB3FFFBE5FAFF919FD343DB8" bold="true" box="[712,738,1744,1768]" pageId="22" pageNumber="523">A)</emphasis>
|
||
<emphasis id="B93AEAFCEB3FFFBE5F8EF919FCEF3DB8" box="[745,825,1744,1768]" italics="true" pageId="22" pageNumber="523">Habitus</emphasis>
|
||
, lateral view.
|
||
<emphasis id="B93AEAFCEB3FFFBE5EAFF919FC373DB8" bold="true" box="[968,993,1744,1768]" pageId="22" pageNumber="523">B)</emphasis>
|
||
<emphasis id="B93AEAFCEB3FFFBE5E80F919FBE13DB8" box="[999,1079,1744,1768]" italics="true" pageId="22" pageNumber="523">Habitus</emphasis>
|
||
, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3546565EB3FFFB45DA0F8DFFBE93B85" lastPageId="28" lastPageNumber="529" pageId="22" pageNumber="523" type="description">
|
||
<paragraph id="8BF136EEEB3FFFBE5DA0F8DFFB3F3C60" blockId="22.[151,1437,1814,2021]" box="[199,1257,1814,1841]" pageId="22" pageNumber="523">
|
||
<emphasis id="B93AEAFCEB3FFFBE5DA0F8DFFE8F3C60" bold="true" box="[199,345,1814,1840]" pageId="22" pageNumber="523">Description.</emphasis>
|
||
Male—length ~
|
||
<quantity id="4CB69B0BEB3FFFBE5F76F8DFFDB13C61" box="[529,615,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="523" unit="mm" value="1.6">1.6 mm</quantity>
|
||
(
|
||
<figureCitation id="13752A6BEB3FFFBE5F11F8DFFCDD3C60" box="[630,779,1814,1840]" captionStart="FIGURE 19" captionStartId="22.[151,250,1744,1768]" captionTargetBox="[151,1436,181,1720]" captionTargetId="figure-22@22.[151,1436,181,1720]" captionTargetPageId="22" captionText="FIGURE 19. Termitozophilus tenebrus sp. nov., male. A) Habitus, lateral view. B) Habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7469419" httpUri="https://zenodo.org/record/7469419/files/figure.png" pageId="22" pageNumber="523">Figs. 19A–B</figureCitation>
|
||
); female—length ~
|
||
<quantity id="4CB69B0BEB3FFFBE5E84F8DFFBEF3C61" box="[995,1081,1814,1841]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="523" unit="mm" value="1.8">1.8 mm</quantity>
|
||
(
|
||
<figureCitation id="13752A6BEB3FFFBE592FF8DFFB0B3C60" box="[1096,1245,1814,1840]" captionStart="FIGURE 20" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-19@23.[203,1385,181,1919]" captionTargetPageId="23" captionText="FIGURE 20. Termitozophilus tenebrus sp. nov., female. A) Habitus, dorsal view. B) Abdomen, ventral view. C) Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7469421" httpUri="https://zenodo.org/record/7469421/files/figure.png" pageId="22" pageNumber="523">Figs. 20A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF136EEEB3FFFB05DA0F8F3FE133AD9" blockId="22.[151,1437,1814,2021]" lastBlockId="24.[151,1437,151,393]" lastPageId="24" lastPageNumber="525" pageId="22" pageNumber="523">
|
||
<emphasis id="B93AEAFCEB3FFFBE5DA0F8F3FED13C04" bold="true" box="[199,263,1850,1876]" pageId="22" pageNumber="523">Head</emphasis>
|
||
subquadrate, about as wide as long, widest behind eye, narrowing conspicuously through posterior region; eye about 1/3 of head length; vertex bearing two pairs of bristles: one bristle behind eye and one on medialposterior region (
|
||
<figureCitation id="13752A6BEB3FFFBE5C06F84BFE183CCC" box="[353,462,1922,1948]" captionStart="FIGURE 21" captionStartId="24.[151,250,1914,1938]" captionTargetBox="[151,1431,420,1881]" captionTargetId="figure-131@24.[151,1436,417,1891]" captionTargetPageId="24" captionText="FIGURE 21. Termitozophilus tenebrus sp. nov. A) Head, dorsal view. B) Left and right mandibles, dorsal view. C) Labrum, dorsal view. D) Maxilla, dorsal view. E) Pre-mentum, mentum and sub-mentum, ventral view. Scales: A = 2 mm; D = 1,5 mm; E = 1 mm; B, C = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7469423" httpUri="https://zenodo.org/record/7469423/files/figure.png" pageId="22" pageNumber="523">Fig. 21A</figureCitation>
|
||
).
|
||
<emphasis id="B93AEAFCEB3FFFBE5C83F84BFD823CCC" bold="true" box="[484,596,1922,1948]" pageId="22" pageNumber="523">Antenna:</emphasis>
|
||
scape almost longer than remainder antennomeres, except antennomere XI; antennomere XI about three times longer than wide and more than twice longer than X; antennomeres II–VI elongate, gradually decreasing in length, VII–X moniliform.
|
||
<emphasis id="B93AEAFCEB3FFFBE5E55F803FBE73CB4" bold="true" box="[818,1073,1994,2020]" pageId="22" pageNumber="523">Mouthparts. Labrum</emphasis>
|
||
more than twice wider than long, with almost sinuous anterior region; five main long bristles on each lateral half, distributed in two diagonal rows, each with two bristles and one bristle placed on middle of labrum length (
|
||
<figureCitation id="13752A6BEB31FFB05E9AFF72FBB13B85" box="[1021,1127,187,213]" captionStart="FIGURE 21" captionStartId="24.[151,250,1914,1938]" captionTargetBox="[151,1431,420,1881]" captionTargetId="figure-131@24.[151,1436,417,1891]" captionTargetPageId="24" captionText="FIGURE 21. Termitozophilus tenebrus sp. nov. A) Head, dorsal view. B) Left and right mandibles, dorsal view. C) Labrum, dorsal view. D) Maxilla, dorsal view. E) Pre-mentum, mentum and sub-mentum, ventral view. Scales: A = 2 mm; D = 1,5 mm; E = 1 mm; B, C = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7469423" httpUri="https://zenodo.org/record/7469423/files/figure.png" pageId="24" pageNumber="525">Fig. 21C</figureCitation>
|
||
); epipharynx covered with sensilla and numerous short bristles, two of which longer.
|
||
<emphasis id="B93AEAFCEB31FFB05E45FF17FC433BA8" bold="true" box="[802,917,222,248]" pageId="24" pageNumber="525">Mandible</emphasis>
|
||
apex narrow, bearing a tooth on mesal region; with many punctuations on basal two thirds, and with some pores on outer margin (
|
||
<figureCitation id="13752A6BEB31FFB05933FECAFB113A4D" box="[1108,1223,259,285]" captionStart="FIGURE 21" captionStartId="24.[151,250,1914,1938]" captionTargetBox="[151,1431,420,1881]" captionTargetId="figure-131@24.[151,1436,417,1891]" captionTargetPageId="24" captionText="FIGURE 21. Termitozophilus tenebrus sp. nov. A) Head, dorsal view. B) Left and right mandibles, dorsal view. C) Labrum, dorsal view. D) Maxilla, dorsal view. E) Pre-mentum, mentum and sub-mentum, ventral view. Scales: A = 2 mm; D = 1,5 mm; E = 1 mm; B, C = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7469423" httpUri="https://zenodo.org/record/7469423/files/figure.png" pageId="24" pageNumber="525">Figs. 21B</figureCitation>
|
||
,
|
||
<figureCitation id="13752A6BEB31FFB059B3FECAFAFF3A4D" box="[1236,1321,259,285]" captionStart="FIGURE 23" captionStartId="26.[151,250,1921,1945]" captionTargetBox="[151,1436,273,1898]" captionTargetId="figure-64@26.[151,1436,273,1898]" captionTargetPageId="26" captionText="FIGURE 23. Termitozophilus tenebrus sp. nov. A) Elytron, internal view. B) Detail of the anteromesial border of elytron, internal view. C–D) Mandible, dorsal view (arrow showing pores). Scales: A = 100 µm; B = 20 µm; C = 50 µm; D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7469429" httpUri="https://zenodo.org/record/7469429/files/figure.png" pageId="24" pageNumber="525">23C–D</figureCitation>
|
||
).
|
||
<emphasis id="B93AEAFCEB31FFB0585AFECBFA4A3A4C" bold="true" box="[1341,1436,258,284]" pageId="24" pageNumber="525">Labium</emphasis>
|
||
with ligula narrowed at base and with a median-apical emargination, resulting in two lobes; mentum and submentum distinct, both trapezoidal, with three bristles distributed on each lateral margin, as follows: apical, pre-apical, and proximal (
|
||
<figureCitation id="13752A6BEB31FFB05C6BFEA6FEAC3AD9" box="[268,378,367,393]" captionStart="FIGURE 21" captionStartId="24.[151,250,1914,1938]" captionTargetBox="[151,1431,420,1881]" captionTargetId="figure-131@24.[151,1436,417,1891]" captionTargetPageId="24" captionText="FIGURE 21. Termitozophilus tenebrus sp. nov. A) Head, dorsal view. B) Left and right mandibles, dorsal view. C) Labrum, dorsal view. D) Maxilla, dorsal view. E) Pre-mentum, mentum and sub-mentum, ventral view. Scales: A = 2 mm; D = 1,5 mm; E = 1 mm; B, C = 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7469423" httpUri="https://zenodo.org/record/7469423/files/figure.png" pageId="24" pageNumber="525">Figs. 21E</figureCitation>
|
||
,
|
||
<figureCitation id="13752A6BEB31FFB05CE1FEA6FE6F3AD9" box="[390,441,367,393]" captionStart="FIGURE 24" captionStartId="27.[151,250,1613,1637]" captionTargetBox="[151,1436,181,1589]" captionTargetId="figure-19@27.[151,1436,181,1589]" captionTargetPageId="27" captionText="FIGURE 24. Termitozophilus tenebrus sp. nov. A) Mentum and sub-mentum (arrow showing suture. B) Maxillary palpus. C) Tergite VII. D) Detail of glandular área of tergite VII. Scales: C = 200 µm; A, D = 20 µm; B = 10 µm; B = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.7469431" httpUri="https://zenodo.org/record/7469431/files/figure.png" pageId="24" pageNumber="525">24C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF316666EB3EFFBF5DF0F85EFF183C84" ID-DOI="http://doi.org/10.5281/zenodo.7469421" ID-Zenodo-Dep="7469421" httpUri="https://zenodo.org/record/7469421/files/figure.png" pageId="23" pageNumber="524" startId="23.[151,250,1943,1967]" targetBox="[203,1385,181,1919]" targetPageId="23">
|
||
<paragraph id="8BF136EEEB3EFFBF5DF0F85EFF183C84" blockId="23.[151,1436,1943,2004]" pageId="23" pageNumber="524">
|
||
<emphasis id="B93AEAFCEB3EFFBF5DF0F85EFEC93CFF" bold="true" box="[151,287,1943,1967]" pageId="23" pageNumber="524">FIGURE 20.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB3EFFBF5C43F851FDF43CE0" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[292,546,1944,1968]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="524" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB3EFFBF5C43F851FDF43CE0" box="[292,546,1944,1968]" italics="true" pageId="23" pageNumber="524">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB3EFFBF5F4FF851FDA13CE0" bold="true" box="[552,631,1944,1968]" pageId="23" pageNumber="524">
|
||
<taxonomicNameLabel id="A2095787EB3EFFBF5F4FF851FDA13CE0" box="[552,631,1944,1968]" pageId="23" pageNumber="524" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female.
|
||
<emphasis id="B93AEAFCEB3EFFBF5FB4F85EFD383CFF" bold="true" box="[723,750,1943,1967]" pageId="23" pageNumber="524">A)</emphasis>
|
||
<emphasis id="B93AEAFCEB3EFFBF5F94F851FC953CE0" box="[755,835,1944,1968]" italics="true" pageId="23" pageNumber="524">Habitus</emphasis>
|
||
, dorsal view.
|
||
<emphasis id="B93AEAFCEB3EFFBF5EAAF85EFC313CFF" bold="true" box="[973,999,1943,1967]" pageId="23" pageNumber="524">B)</emphasis>
|
||
Abdomen, ventral view.
|
||
<emphasis id="B93AEAFCEB3EFFBF5986F85EFB2A3CFF" bold="true" box="[1249,1276,1943,1967]" pageId="23" pageNumber="524">C)</emphasis>
|
||
<emphasis id="B93AEAFCEB3EFFBF5866F851FA873CE0" box="[1281,1361,1944,1968]" italics="true" pageId="23" pageNumber="524">Habitus</emphasis>
|
||
, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF316666EB31FFB05DF0F8B3FE7E3C8A" ID-DOI="http://doi.org/10.5281/zenodo.7469423" ID-Zenodo-Dep="7469423" httpUri="https://zenodo.org/record/7469423/files/figure.png" pageId="24" pageNumber="525" startId="24.[151,250,1914,1938]" targetBox="[151,1431,420,1881]" targetPageId="24">
|
||
<paragraph id="8BF136EEEB31FFB05DF0F8B3FE7E3C8A" blockId="24.[151,1437,1914,2010]" pageId="24" pageNumber="525">
|
||
<emphasis id="B93AEAFCEB31FFB05DF0F8B3FEF73CC2" bold="true" box="[151,289,1914,1938]" pageId="24" pageNumber="525">FIGURE 21.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB31FFB05C4EF8B3FDFC3CC2" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[297,554,1914,1938]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="525" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB31FFB05C4EF8B3FDFC3CC2" box="[297,554,1914,1938]" italics="true" pageId="24" pageNumber="525">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB31FFB05F55F8B3FD733CC2" bold="true" box="[562,677,1914,1938]" pageId="24" pageNumber="525">
|
||
<taxonomicNameLabel id="A2095787EB31FFB05F55F8B3FD523CC2" box="[562,644,1914,1938]" pageId="24" pageNumber="525" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A)
|
||
</emphasis>
|
||
Head, dorsal view.
|
||
<emphasis id="B93AEAFCEB31FFB05E14F8B3FC5B3CC2" bold="true" box="[883,909,1914,1938]" pageId="24" pageNumber="525">B)</emphasis>
|
||
Left and right mandibles, dorsal view.
|
||
<emphasis id="B93AEAFCEB31FFB05844F8B3FAE83CC2" bold="true" box="[1315,1342,1914,1938]" pageId="24" pageNumber="525">C)</emphasis>
|
||
Labrum, dorsal view.
|
||
<emphasis id="B93AEAFCEB31FFB05C7FF857FEE43CE6" bold="true" box="[280,306,1950,1974]" pageId="24" pageNumber="525">D)</emphasis>
|
||
Maxilla, dorsal view.
|
||
<emphasis id="B93AEAFCEB31FFB05F72F857FDF83CE6" bold="true" box="[533,558,1950,1974]" pageId="24" pageNumber="525">E)</emphasis>
|
||
Pre-mentum, mentum and sub-mentum, ventral view. Scales: A
|
||
<emphasis id="B93AEAFCEB31FFB059DAF857FB1D3CE6" bold="true" box="[1213,1227,1950,1974]" pageId="24" pageNumber="525">=</emphasis>
|
||
2 mm; D
|
||
<emphasis id="B93AEAFCEB31FFB05854F857FA973CE6" bold="true" box="[1331,1345,1950,1974]" pageId="24" pageNumber="525">=</emphasis>
|
||
1,5 mm; E
|
||
<emphasis id="B93AEAFCEB31FFB05DCAF80BFF6D3C8A" bold="true" box="[173,187,1986,2010]" pageId="24" pageNumber="525">=</emphasis>
|
||
1 mm; B, C
|
||
<emphasis id="B93AEAFCEB31FFB05C59F80BFE9A3C8A" bold="true" box="[318,332,1986,2010]" pageId="24" pageNumber="525">=</emphasis>
|
||
0,5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF316666EB30FFB15DF0F888FED73CB9" ID-DOI="http://doi.org/10.5281/zenodo.7469427" ID-Zenodo-Dep="7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="25" pageNumber="526" startId="25.[151,250,1857,1881]" targetBox="[232,1355,181,1833]" targetPageId="25">
|
||
<paragraph id="8BF136EEEB30FFB15DF0F888FED73CB9" blockId="25.[151,1437,1857,2025]" pageId="25" pageNumber="526">
|
||
<emphasis id="B93AEAFCEB30FFB15DF0F888FEF43C09" bold="true" box="[151,290,1857,1881]" pageId="25" pageNumber="526">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB30FFB15C4DF888FDFA3C09" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[298,556,1857,1881]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="526" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB30FFB15C4DF888FDFA3C09" box="[298,556,1857,1881]" italics="true" pageId="25" pageNumber="526">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB30FFB15F52F888FD7C3C09" bold="true" box="[565,682,1857,1881]" pageId="25" pageNumber="526">
|
||
<taxonomicNameLabel id="A2095787EB30FFB15F52F888FD5E3C09" box="[565,648,1857,1881]" pageId="25" pageNumber="526" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A)
|
||
</emphasis>
|
||
Meso-metanotum and tergite I, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB159E7F888FB4C3C09" bold="true" box="[1152,1178,1857,1881]" pageId="25" pageNumber="526">B)</emphasis>
|
||
Elytron, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB158E6F888FA4A3C09" bold="true" box="[1409,1436,1857,1881]" pageId="25" pageNumber="526">C)</emphasis>
|
||
Tergite VII, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB15CF4F8ACFE7B3C2D" bold="true" box="[403,429,1893,1917]" pageId="25" pageNumber="526">D)</emphasis>
|
||
Male tergites IX and X, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB15E40F8ACFC963C2D" bold="true" box="[807,832,1893,1917]" pageId="25" pageNumber="526">E)</emphasis>
|
||
Male sternite IX, ventral view.
|
||
<emphasis id="B93AEAFCEB30FFB159E6F8ACFB4F3C2D" bold="true" box="[1153,1177,1893,1917]" pageId="25" pageNumber="526">F)</emphasis>
|
||
Tergite VIII, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB15DF0F840FF653CF1" bold="true" box="[151,179,1929,1953]" pageId="25" pageNumber="526">G)</emphasis>
|
||
Sternite VIII, ventral view.
|
||
<emphasis id="B93AEAFCEB30FFB15CABF840FE3E3CF1" bold="true" box="[460,488,1929,1953]" pageId="25" pageNumber="526">H)</emphasis>
|
||
Female tergites IX and X, dorsal view.
|
||
<emphasis id="B93AEAFCEB30FFB15E10F840FC5F3CF1" bold="true" box="[887,905,1929,1953]" pageId="25" pageNumber="526">I)</emphasis>
|
||
Median lobe of aedeagus, lateral view.
|
||
<emphasis id="B93AEAFCEB30FFB15870F840FAFA3CF1" bold="true" box="[1303,1324,1929,1953]" pageId="25" pageNumber="526">J)</emphasis>
|
||
Aedeagus, paramere, lateral view.
|
||
<emphasis id="B93AEAFCEB30FFB15CE2F864FE773C95" bold="true" box="[389,417,1965,1989]" pageId="25" pageNumber="526">K)</emphasis>
|
||
Spermatheca, lateral view. Scales: A, C, F–H
|
||
<emphasis id="B93AEAFCEB30FFB15E1EF864FC513C95" bold="true" box="[889,903,1965,1989]" pageId="25" pageNumber="526">=</emphasis>
|
||
2 mm; B
|
||
<emphasis id="B93AEAFCEB30FFB15E88F864FC2B3C95" bold="true" box="[1007,1021,1965,1989]" pageId="25" pageNumber="526">=</emphasis>
|
||
1,5 mm; D, E
|
||
<emphasis id="B93AEAFCEB30FFB159F2F864FB753C95" bold="true" box="[1173,1187,1965,1989]" pageId="25" pageNumber="526">=</emphasis>
|
||
1 mm; K
|
||
<emphasis id="B93AEAFCEB30FFB1586BF864FACC3C95" bold="true" box="[1292,1306,1965,1989]" pageId="25" pageNumber="526">=</emphasis>
|
||
0,5 mm; I, J
|
||
<emphasis id="B93AEAFCEB30FFB15DF0F818FF733CB9" bold="true" box="[151,165,2001,2025]" pageId="25" pageNumber="526">=</emphasis>
|
||
0,1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF136EEEB33FFB25DA0FF5EFB573BA9" blockId="26.[151,1436,151,249]" pageId="26" pageNumber="527">
|
||
<emphasis id="B93AEAFCEB33FFB25DA0FF5EFEAD3BE1" bold="true" box="[199,379,151,177]" pageId="26" pageNumber="527">Thorax. Elytra</emphasis>
|
||
trapezoidal, almost longer than wide and wider at posterior region, covering mesonotum and metanotum (
|
||
<figureCitation id="13752A6BEB33FFB25C4EFF72FE423B85" box="[297,404,187,213]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="26" pageNumber="527">Fig. 22B</figureCitation>
|
||
); anteromesial border densely scaled in internal view (
|
||
<figureCitation id="13752A6BEB33FFB2597EFF72FB673B85" box="[1049,1201,187,213]" captionStart="FIGURE 23" captionStartId="26.[151,250,1921,1945]" captionTargetBox="[151,1436,273,1898]" captionTargetId="figure-64@26.[151,1436,273,1898]" captionTargetPageId="26" captionText="FIGURE 23. Termitozophilus tenebrus sp. nov. A) Elytron, internal view. B) Detail of the anteromesial border of elytron, internal view. C–D) Mandible, dorsal view (arrow showing pores). Scales: A = 100 µm; B = 20 µm; C = 50 µm; D = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7469429" httpUri="https://zenodo.org/record/7469429/files/figure.png" pageId="26" pageNumber="527">Figs. 23A–B</figureCitation>
|
||
); covered with long bristles.
|
||
<emphasis id="B93AEAFCEB33FFB25D91FF17FEA23BA8" bold="true" box="[246,372,222,248]" pageId="26" pageNumber="527">Hind wing</emphasis>
|
||
present, membrane not shed during post-imaginal growth (
|
||
<figureCitation id="13752A6BEB33FFB2596CFF16FBA23BA9" box="[1035,1140,223,249]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="26" pageNumber="527">Fig. 22A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF316666EB33FFB25DF0F848FA503CEE" ID-DOI="http://doi.org/10.5281/zenodo.7469429" ID-Zenodo-Dep="7469429" httpUri="https://zenodo.org/record/7469429/files/figure.png" pageId="26" pageNumber="527" startId="26.[151,250,1921,1945]" targetBox="[151,1436,273,1898]" targetPageId="26">
|
||
<paragraph id="8BF136EEEB33FFB25DF0F848FA503CEE" blockId="26.[151,1436,1921,1982]" pageId="26" pageNumber="527">
|
||
<emphasis id="B93AEAFCEB33FFB25DF0F848FEF53CCA" bold="true" box="[151,291,1921,1946]" pageId="26" pageNumber="527">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB33FFB25C4BF84BFDF83CCA" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[300,558,1922,1946]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="527" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB33FFB25C4BF84BFDF83CCA" box="[300,558,1922,1946]" italics="true" pageId="26" pageNumber="527">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB33FFB25F5FF84BFD793CC9" bold="true" box="[568,687,1921,1946]" pageId="26" pageNumber="527">
|
||
<taxonomicNameLabel id="A2095787EB33FFB25F5FF84BFD5D3CCA" box="[568,651,1922,1946]" pageId="26" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A)
|
||
</emphasis>
|
||
Elytron, internal view.
|
||
<emphasis id="B93AEAFCEB33FFB25ECDF848FC123CC9" bold="true" box="[938,964,1921,1945]" pageId="26" pageNumber="527">B)</emphasis>
|
||
Detail of the anteromesial border of elytron, internal view.
|
||
<emphasis id="B93AEAFCEB33FFB25C40F86CFEB63CED" bold="true" box="[295,352,1957,1981]" pageId="26" pageNumber="527">C–D)</emphasis>
|
||
Mandible, dorsal view (arrow showing pores). Scales: A
|
||
<emphasis id="B93AEAFCEB33FFB25ECFF86FFC603CEE" bold="true" box="[936,950,1958,1982]" pageId="26" pageNumber="527">=</emphasis>
|
||
100 µm; B
|
||
<emphasis id="B93AEAFCEB33FFB25957F86FFBE83CEE" bold="true" box="[1072,1086,1958,1982]" pageId="26" pageNumber="527">=</emphasis>
|
||
20 µm; C
|
||
<emphasis id="B93AEAFCEB33FFB259CCF86FFB6F3CEE" bold="true" box="[1195,1209,1958,1982]" pageId="26" pageNumber="527">=</emphasis>
|
||
50 µm; D
|
||
<emphasis id="B93AEAFCEB33FFB2584FF86FFAE03CEE" bold="true" box="[1320,1334,1958,1982]" pageId="26" pageNumber="527">=</emphasis>
|
||
10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF316666EB32FFB35DF0F984FAC63DDA" ID-DOI="http://doi.org/10.5281/zenodo.7469431" ID-Zenodo-Dep="7469431" httpUri="https://zenodo.org/record/7469431/files/figure.png" pageId="27" pageNumber="528" startId="27.[151,250,1613,1637]" targetBox="[151,1436,181,1589]" targetPageId="27">
|
||
<paragraph id="8BF136EEEB32FFB35DF0F984FAC63DDA" blockId="27.[151,1436,1613,1674]" pageId="27" pageNumber="528">
|
||
<emphasis id="B93AEAFCEB32FFB35DF0F984FEF63D35" bold="true" box="[151,288,1613,1637]" pageId="27" pageNumber="528">FIGURE 24.</emphasis>
|
||
<taxonomicName id="4C4E4D6DEB32FFB35C40F984FDF13D35" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[295,551,1613,1637]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="528" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB32FFB35C40F984FDF13D35" box="[295,551,1613,1637]" italics="true" pageId="27" pageNumber="528">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB32FFB35F49F984FD773D35" bold="true" box="[558,673,1613,1637]" pageId="27" pageNumber="528">
|
||
<taxonomicNameLabel id="A2095787EB32FFB35F49F984FDA93D35" box="[558,639,1613,1637]" pageId="27" pageNumber="528" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A)
|
||
</emphasis>
|
||
Mentum and sub-mentum (arrow showing suture.
|
||
<emphasis id="B93AEAFCEB32FFB359C0F984FB173D35" bold="true" box="[1191,1217,1613,1637]" pageId="27" pageNumber="528">B)</emphasis>
|
||
Maxillary palpus.
|
||
<emphasis id="B93AEAFCEB32FFB358E6F984FA4A3D35" bold="true" box="[1409,1436,1613,1637]" pageId="27" pageNumber="528">C)</emphasis>
|
||
Tergite VII.
|
||
<emphasis id="B93AEAFCEB32FFB35C73F9B8FEF83DD9" bold="true" box="[276,302,1649,1673]" pageId="27" pageNumber="528">D)</emphasis>
|
||
Detail of glandular área of tergite VII. Scales: C
|
||
<emphasis id="B93AEAFCEB32FFB35E45F9B8FCE63DD9" bold="true" box="[802,816,1649,1673]" pageId="27" pageNumber="528">=</emphasis>
|
||
200 µm; A, D
|
||
<emphasis id="B93AEAFCEB32FFB35EAEF9B8FC013DD9" bold="true" box="[969,983,1649,1673]" pageId="27" pageNumber="528">=</emphasis>
|
||
20 µm; B
|
||
<emphasis id="B93AEAFCEB32FFB35923F9B8FB843DD9" bold="true" box="[1092,1106,1649,1673]" pageId="27" pageNumber="528">=</emphasis>
|
||
10 µm; B
|
||
<emphasis id="B93AEAFCEB32FFB359D8F9B8FB1B3DD9" bold="true" box="[1215,1229,1649,1673]" pageId="27" pageNumber="528">=</emphasis>
|
||
2 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF136EEEB32FFB45DA0F97EFBE93B85" blockId="27.[151,1437,1719,2034]" lastBlockId="28.[151,1437,151,609]" lastPageId="28" lastPageNumber="529" pageId="27" pageNumber="528">
|
||
<emphasis id="B93AEAFCEB32FFB35DA0F97EFEEA3D81" bold="true" box="[199,316,1719,1745]" pageId="27" pageNumber="528">Abdomen</emphasis>
|
||
with short tergites and with secondary sclerotization not reaching inner paratergites; outer paratergites fused to sternites by former secondary sclerotization; inner and outer paratergites fused (
|
||
<figureCitation id="13752A6BEB32FFB359EBF912FB253DA6" box="[1164,1267,1755,1782]" captionStart="FIGURE 27" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[234,1355,189,1913]" captionTargetId="figure-22@30.[221,1367,181,1919]" captionTargetPageId="30" captionText="FIGURE 27. Phylogenetic recapitulation hypothesis based on the condition of abdominal sclerites in Termitozophilus species." figureDoi="http://doi.org/10.5281/zenodo.7469437" httpUri="https://zenodo.org/record/7469437/files/figure.png" pageId="27" pageNumber="528">Fig. 27E</figureCitation>
|
||
); first segment represented by a strong sclerotized tergite attached to metanotum, with medial region strongly projected backwards (
|
||
<figureCitation id="13752A6BEB32FFB35DF8F8EAFEDC3C6E" box="[159,266,1827,1854]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22A</figureCitation>
|
||
); sternites III–V with pores clustered; tergite VII (
|
||
<figureCitation id="13752A6BEB32FFB35E20F8EAFC783C6E" box="[839,942,1827,1854]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22C</figureCitation>
|
||
) almost subquadrate, wider than long, with two rows of bristles, two bristles on medial region and six bristles on posterior half, anterior margin of tergite VII with conspicuous glandular areas (
|
||
<figureCitation id="13752A6BEB32FFB35F4CF8A2FD683CD6" box="[555,702,1899,1926]" captionStart="FIGURE 24" captionStartId="27.[151,250,1613,1637]" captionTargetBox="[151,1436,181,1589]" captionTargetId="figure-19@27.[151,1436,181,1589]" captionTargetPageId="27" captionText="FIGURE 24. Termitozophilus tenebrus sp. nov. A) Mentum and sub-mentum (arrow showing suture. B) Maxillary palpus. C) Tergite VII. D) Detail of glandular área of tergite VII. Scales: C = 200 µm; A, D = 20 µm; B = 10 µm; B = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.7469431" httpUri="https://zenodo.org/record/7469431/files/figure.png" pageId="27" pageNumber="528">Figs. 24E–F</figureCitation>
|
||
); tergite VIII (
|
||
<figureCitation id="13752A6BEB32FFB35E0DF8A2FC193CD6" box="[874,975,1899,1926]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22F</figureCitation>
|
||
) almost subquadrate, with two rows of bristles, six main long bristles on medial and posterior regions; sternite VIII (
|
||
<figureCitation id="13752A6BEB32FFB35E9BF846FBB53CFA" box="[1020,1123,1935,1962]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22G</figureCitation>
|
||
) subquadrate, with one pair of median bristle, one row of six bristle close to posterior margin and eight bristles along posterior margin; male sternite IX (
|
||
<figureCitation id="13752A6BEB32FFB35C7BF81EFEA93CA2" box="[284,383,2007,2034]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22E</figureCitation>
|
||
) elongate and tongue-shaped, with two bristles on posterior half; female sternite IX (
|
||
<figureCitation id="13752A6BEB32FFB35848F81EFA433CA2" box="[1327,1429,2007,2034]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="27" pageNumber="528">Fig. 22H</figureCitation>
|
||
) represented by a pair of hemisternites laterally attached to tergite IX; tergite X (
|
||
<figureCitation id="13752A6BEB35FFB45944FF5EFB5D3BE1" box="[1059,1163,151,177]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="28" pageNumber="529">Fig. 22D</figureCitation>
|
||
) not bilobed, with three long bristles on each lateral half, two on medial region and two on posterior region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3546565EB35FFB45DA0FF16FE513A4D" pageId="28" pageNumber="529" type="etymology">
|
||
<paragraph id="8BF136EEEB35FFB45DA0FF16FE513A4D" blockId="28.[151,1437,151,609]" pageId="28" pageNumber="529">
|
||
<emphasis id="B93AEAFCEB35FFB45DA0FF16FE983BA9" bold="true" box="[199,334,223,249]" pageId="28" pageNumber="529">Etymology.</emphasis>
|
||
The specific epithet is a Latin adjective that means “dark”, alluding to dull coloration of the examined specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3546565EB35FFB45DA0FEEFFD5B3A11" box="[199,653,294,321]" pageId="28" pageNumber="529" type="distribution">
|
||
<paragraph id="8BF136EEEB35FFB45DA0FEEFFD5B3A11" blockId="28.[151,1437,151,609]" box="[199,653,294,321]" pageId="28" pageNumber="529">
|
||
<emphasis id="B93AEAFCEB35FFB45DA0FEEFFE393A10" bold="true" box="[199,495,294,320]" pageId="28" pageNumber="529">Geographic distribution.</emphasis>
|
||
<collectingCountry id="F359767EEB35FFB45C91FEEEFDEC3A11" box="[502,570,295,321]" name="Brazil" pageId="28" pageNumber="529">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="498AF80CEB35FFB45F2EFEEEFD573A11" box="[585,641,295,321]" country="Brazil" name="Para" pageId="28" pageNumber="529">Pará</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3546565EB35FFB45DA0FE83FB843931" pageId="28" pageNumber="529" type="discussion">
|
||
<paragraph id="8BF136EEEB35FFB45DA0FE83FB843931" blockId="28.[151,1437,151,609]" pageId="28" pageNumber="529">
|
||
<emphasis id="B93AEAFCEB35FFB45DA0FE83FE933A34" bold="true" box="[199,325,330,356]" pageId="28" pageNumber="529">Discussion</emphasis>
|
||
.
|
||
<taxonomicName id="4C4E4D6DEB35FFB45C30FE82FDA23A35" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[343,628,331,357]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB35FFB45C30FE82FDA23A35" box="[343,628,331,357]" italics="true" pageId="28" pageNumber="529">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB35FFB45F18FE82FD0A3A35" bold="true" box="[639,732,331,357]" pageId="28" pageNumber="529">
|
||
<taxonomicNameLabel id="A2095787EB35FFB45F18FE82FD0A3A35" box="[639,732,331,357]" pageId="28" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C4E4D6DEB35FFB45E1CFE82FC093A35" authorityName="Silvestri" authorityYear="1901" box="[891,991,331,357]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB45E1CFE82FC5D3A34" box="[891,907,331,356]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45EFBFE82FC093A35" box="[924,991,331,357]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
in having inner and outer paratergites fused, and paratergites fused to sternites (
|
||
<figureCitation id="13752A6BEB35FFB45F07FEA6FD243AD9" box="[608,754,367,393]" captionStart="FIGURE 27" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[234,1355,189,1913]" captionTargetId="figure-22@30.[221,1367,181,1919]" captionTargetPageId="30" captionText="FIGURE 27. Phylogenetic recapitulation hypothesis based on the condition of abdominal sclerites in Termitozophilus species." figureDoi="http://doi.org/10.5281/zenodo.7469437" httpUri="https://zenodo.org/record/7469437/files/figure.png" pageId="28" pageNumber="529">Figs. 27A–E</figureCitation>
|
||
). It differs from
|
||
<taxonomicName id="4C4E4D6DEB35FFB45EC1FEA6FBD33AD9" authorityName="Silvestri" authorityYear="1901" box="[934,1029,367,393]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB45EC1FEA6FC603AD8" box="[934,950,367,392]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45EA5FEA6FBD33AD9" box="[962,1029,367,393]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
in having head, thorax and abdomen with dark brown sclerites (light brown in
|
||
<taxonomicName id="4C4E4D6DEB35FFB45F19FE5AFD343AFD" authorityName="Silvestri" authorityYear="1901" box="[638,738,403,429]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB45F19FE5AFD583AFC" box="[638,654,403,428]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45FF8FE5AFD343AFD" box="[671,738,403,429]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
) and pores on sternites distributed in circular clusters (
|
||
<figureCitation id="13752A6BEB35FFB45803FE5AFF3C3A81" captionStart="FIGURE 19" captionStartId="22.[151,250,1744,1768]" captionTargetBox="[151,1436,181,1720]" captionTargetId="figure-22@22.[151,1436,181,1720]" captionTargetPageId="22" captionText="FIGURE 19. Termitozophilus tenebrus sp. nov., male. A) Habitus, lateral view. B) Habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7469419" httpUri="https://zenodo.org/record/7469419/files/figure.png" pageId="28" pageNumber="529">Figs. 19A–B</figureCitation>
|
||
,
|
||
<figureCitation id="13752A6BEB35FFB45D92FE7EFE913A81" box="[245,327,439,465]" captionStart="FIGURE 20" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-19@23.[203,1385,181,1919]" captionTargetPageId="23" captionText="FIGURE 20. Termitozophilus tenebrus sp. nov., female. A) Habitus, dorsal view. B) Abdomen, ventral view. C) Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7469421" httpUri="https://zenodo.org/record/7469421/files/figure.png" pageId="28" pageNumber="529">20A–C</figureCitation>
|
||
) (scattered in
|
||
<taxonomicName id="4C4E4D6DEB35FFB45C83FE7EFD953A81" authorityName="Silvestri" authorityYear="1901" box="[484,579,439,465]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB45C83FE7EFE223A80" box="[484,500,439,464]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45F67FE7EFD953A81" box="[512,579,439,465]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
). In addition,
|
||
<taxonomicName id="4C4E4D6DEB35FFB45F87FE7EFC8C3A81" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[736,858,439,465]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB35FFB45F87FE7EFC8C3A81" box="[736,858,439,465]" italics="true" pageId="28" pageNumber="529">T. tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB35FFB45E06FE7EFC6F3A81" bold="true" box="[865,953,439,465]" pageId="28" pageNumber="529">
|
||
<taxonomicNameLabel id="A2095787EB35FFB45E06FE7EFC6F3A81" box="[865,953,439,465]" pageId="28" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a trapezoidal elytron (
|
||
<figureCitation id="13752A6BEB35FFB459BAFE7EFA943A81" box="[1245,1346,439,465]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="28" pageNumber="529">Fig. 22B</figureCitation>
|
||
), tergite VII with bristles (
|
||
<figureCitation id="13752A6BEB35FFB45C3DFE12FE6B3AA5" box="[346,445,475,501]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="28" pageNumber="529">Fig. 22C</figureCitation>
|
||
) and posterior margin of tergite VIII without bristles (
|
||
<figureCitation id="13752A6BEB35FFB45977FE12FBA43AA5" box="[1040,1138,475,501]" captionStart="FIGURE 22" captionStartId="25.[151,250,1857,1881]" captionTargetBox="[232,1355,181,1833]" captionTargetId="figure-19@25.[232,1355,181,1833]" captionTargetPageId="25" captionText="FIGURE 22. Termitozophilus tenebrus sp. nov. A) Meso-metanotum and tergite I, dorsal view. B) Elytron, dorsal view. C) Tergite VII, dorsal view. D) Male tergites IX and X, dorsal view. E) Male sternite IX, ventral view. F) Tergite VIII, dorsal view. G) Sternite VIII, ventral view. H) Female tergites IX and X, dorsal view. I) Median lobe of aedeagus, lateral view. J) Aedeagus, paramere, lateral view. K) Spermatheca, lateral view. Scales: A, C, F–H = 2 mm; B = 1,5 mm; D, E = 1 mm; K = 0,5 mm; I, J = 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.7469427" httpUri="https://zenodo.org/record/7469427/files/figure.png" pageId="28" pageNumber="529">Fig. 22F</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C4E4D6DEB35FFB459C6FE12FAD63AA5" authorityName="Silvestri" authorityYear="1901" box="[1185,1280,475,501]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB459C6FE12FB673AA4" box="[1185,1201,475,500]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB459DAFE12FAD63AA5" box="[1213,1280,475,501]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
, the elytron is subquadrate, the bristles on tergite VII are absent and the posterior margin of tergite VIII has four bristles (see figs.
|
||
<quantity id="4CB69B0BEB35FFB45DF0FDEAFF2A396D" box="[151,252,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="28" pageNumber="529" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
|
||
Zilberman (2019)).
|
||
<taxonomicName id="4C4E4D6DEB35FFB45C82FDEAFCD7396D" authority="Silva & Zilberman & Carvalho-Filho, 2022" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[485,769,547,573]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="tenebrus" status="sp. nov.">
|
||
<emphasis id="B93AEAFCEB35FFB45C82FDEAFCD7396D" box="[485,769,547,573]" italics="true" pageId="28" pageNumber="529">Termitozophilus tenebrus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93AEAFCEB35FFB45E6CFDEAFCB0396D" bold="true" box="[779,870,547,573]" pageId="28" pageNumber="529">
|
||
<taxonomicNameLabel id="A2095787EB35FFB45E6CFDEAFCB0396D" box="[779,870,547,573]" pageId="28" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is associated with nests of
|
||
<taxonomicName id="4C4E4D6DEB35FFB459C6FDEAFAC4396D" authorityName="Silva & Zilberman & Carvalho-Filho" authorityYear="2022" box="[1185,1298,547,573]" class="Insecta" family="Termitidae" genus="Cornitermes" kingdom="Animalia" order="Blattodea" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="pilosus">
|
||
<emphasis id="B93AEAFCEB35FFB459C6FDEAFB62396D" box="[1185,1204,547,573]" italics="true" pageId="28" pageNumber="529">C</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB459A4FDEAFAC4396D" box="[1219,1298,547,573]" italics="true" pageId="28" pageNumber="529">pilosus</emphasis>
|
||
</taxonomicName>
|
||
in pastures, while
|
||
<taxonomicName id="4C4E4D6DEB35FFB45DBBFD8EFEEB3931" authorityName="Silvestri" authorityYear="1901" box="[220,317,583,609]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="laetus">
|
||
<emphasis id="B93AEAFCEB35FFB45DBBFD8EFF3A3930" box="[220,236,583,608]" italics="true" pageId="28" pageNumber="529">T</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45D9DFD8EFEEB3931" box="[250,317,583,609]" italics="true" pageId="28" pageNumber="529">laetus</emphasis>
|
||
</taxonomicName>
|
||
is associated with nests of
|
||
<taxonomicName id="4C4E4D6DEB35FFB45F09FD8EFD2F3931" baseAuthorityName="Kollar" baseAuthorityYear="1832" box="[622,761,583,609]" class="Insecta" family="Termitidae" genus="Cornitermes" kingdom="Animalia" order="Blattodea" pageId="28" pageNumber="529" phylum="Arthropoda" rank="species" species="cumulans">
|
||
<emphasis id="B93AEAFCEB35FFB45F09FD8EFD573931" box="[622,641,583,609]" italics="true" pageId="28" pageNumber="529">C</emphasis>
|
||
.
|
||
<emphasis id="B93AEAFCEB35FFB45FE9FD8EFD2F3931" box="[654,761,583,609]" italics="true" pageId="28" pageNumber="529">cumulans</emphasis>
|
||
</taxonomicName>
|
||
in savanna-like environments.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |