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<document id="4AB5A083AC5C4B2FE67E7B99796E2A07" ID-CLB-Dataset="8180" ID-DOI="10.11646/zootaxa.4927.4.9" ID-GBIF-Dataset="dc853953-1449-4ef0-903e-583c3664bafe" ID-ISSN="1175-5326" ID-Zenodo-Dep="4543187" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613475631722" checkinUser="plazi" docAuthor="Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2021" docId="03E787F3FFB9FFE0FF7FFAB8FBFB43D4" docLanguage="en" docName="zootaxa.4927.4.9.pdf" docOrigin="Zootaxa 4927 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aegidium simplex Frolov &amp; Akhmetova 2021, new species" docType="treatment" docVersion="5" lastPageNumber="585" masterDocId="FFDEFF8BFFB9FFE2FFE8FFBDFFFB454B" masterDocTitle="A new species of Aegidium Arrow (Coleoptera: Scarabaeidae: Orphninae) from Colombia" masterLastPageNumber="586" masterPageNumber="583" pageNumber="583" updateTime="1698929769364" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="890E35DD7F6F3FCE8B4E81498A53DADF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F41296DD7217C70429A581452E45CE6A">
<mods:title id="D4696684BFDEE8FE5F9101EEA7F61D39">A new species of Aegidium Arrow (Coleoptera: Scarabaeidae: Orphninae) from Colombia</mods:title>
</mods:titleInfo>
<mods:name id="D2893306AEECBF8962BC851BE04B799F" type="personal">
<mods:role id="EEE37F0FEABA61BAF2CE16C41CC893DD">
<mods:roleTerm id="049AB67E8798248E5D4EE6D5453A99D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E7A5326EAAF147F47AC93AF12ABF586">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="D8CEE57A076290D70CDBD7738C13AE6E" type="personal">
<mods:role id="9A4D7376E42579D7FF31AC878A8C9DA8">
<mods:roleTerm id="6545E63FC221B3048A6767EABBEEB99E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81990D489441D32904EFBA40C02219BB">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="876B13EAAB82BBE5FB0502AD8A9BF735">text</mods:typeOfResource>
<mods:relatedItem id="816024C59FA87AD0E124E0F926B0DC06" type="host">
<mods:titleInfo id="74DE5C61849B469166CF7E835AD4659F">
<mods:title id="EDCA1AD36E26A27015D097F36BE67CF2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="92853A681CD7B33E8559EBD46F09A9D4">
<mods:date id="D43FAF9D702498962BBD54632DD433DE">2021</mods:date>
<mods:detail id="86E29CD9E3372DE2C4927D4D9DB7174D" type="pubDate">
<mods:number id="8B27F2BEEAEAEE97547FD82D50ED707D">2021-02-16</mods:number>
</mods:detail>
<mods:detail id="BC924BFF84E1D9950F182CF2145BE507" type="volume">
<mods:number id="046ED20EA8C317B2C664E58B2325BDAC">4927</mods:number>
</mods:detail>
<mods:detail id="CA069E16960AD9B9170CFE89F5F3D878" type="issue">
<mods:number id="F7AF381229E95A8037565884B91956DB">4</mods:number>
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<mods:start id="391AEB17334D2D971BBBE75E325518EF">583</mods:start>
<mods:end id="3BC176BF40AAB57DD51C6BEAB1B0D2FB">586</mods:end>
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<mods:classification id="D20E9DE9D82C58828E3D0AE2D1A9B7AD">journal article</mods:classification>
<mods:identifier id="BA442FFA8A85805DD8698590F61A18AB" type="CLB-Dataset">8180</mods:identifier>
<mods:identifier id="B061E32F5B8371F3058CCF8E169260A4" type="DOI">10.11646/zootaxa.4927.4.9</mods:identifier>
<mods:identifier id="28D814778EC2B0D038471F7D081AA16D" type="GBIF-Dataset">dc853953-1449-4ef0-903e-583c3664bafe</mods:identifier>
<mods:identifier id="0167F4728645943A3D1790A844BA34A3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E0D44B9761DE5CB2680FDD9EB33098B8" type="Zenodo-Dep">4543187</mods:identifier>
<mods:identifier id="8D9F7DBF4F7A6103F5B0DE34068740AD" type="ZooBank">B6229A63-6691-4D98-B4A6-90345DA7F334</mods:identifier>
</mods:mods>
<treatment id="03E787F3FFB9FFE0FF7FFAB8FBFB43D4" ID-DOI="http://doi.org/10.5281/zenodo.4543189" ID-GBIF-Taxon="177418216" ID-Zenodo-Dep="4543189" LSID="urn:lsid:plazi:treatment:03E787F3FFB9FFE0FF7FFAB8FBFB43D4" httpUri="http://treatment.plazi.org/id/03E787F3FFB9FFE0FF7FFAB8FBFB43D4" lastPageId="2" lastPageNumber="585" pageId="0" pageNumber="583">
<subSubSection id="C354656EFFB9FFE2FF7FFAB8FCDB406B" box="[151,800,1285,1312]" pageId="0" pageNumber="583" type="nomenclature">
<paragraph id="8BF136E5FFB9FFE2FF7FFAB8FCDB406B" blockId="0.[151,1437,1285,2032]" box="[151,800,1285,1312]" pageId="0" pageNumber="583">
<heading id="D0B98189FFB9FFE2FF7FFAB8FCDB406B" bold="true" box="[151,800,1285,1312]" fontSize="11" level="1" pageId="0" pageNumber="583" reason="1">
<emphasis id="B93AEAF7FFB9FFE2FF7FFAB8FCDB406B" bold="true" box="[151,800,1285,1312]" pageId="0" pageNumber="583">
<taxonomicName id="4C4E4D66FFB9FFE2FF7FFAB8FD78406B" authority="Frolov &amp; Akhmetova" authorityName="Frolov &amp; Akhmetova" authorityYear="2021" box="[151,643,1285,1312]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="583" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B93AEAF7FFB9FFE2FF7FFAB8FE8A406B" bold="true" box="[151,369,1285,1312]" italics="true" pageId="0" pageNumber="583">Aegidium simplex</emphasis>
Frolov &amp; Akhmetova
</taxonomicName>
,
<taxonomicNameLabel id="A209578CFFB9FFE2FD78FABBFCDB406B" box="[656,800,1286,1312]" pageId="0" pageNumber="583" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354656EFFB9FFE2FF7FFA91FEFA400F" box="[151,257,1324,1348]" pageId="0" pageNumber="583" type="description">
<paragraph id="8BF136E5FFB9FFE2FF7FFA91FEFA400F" blockId="0.[151,1437,1285,2032]" box="[151,257,1324,1348]" pageId="0" pageNumber="583">
<figureCitation id="13752A60FFB9FFE2FF7FFA91FF06400F" box="[151,253,1324,1348]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–9" captionStartId-0="1.[151,264,1973,1997]" captionStartId-1="2.[151,264,1023,1047]" captionTargetBox-0="[190,1400,429,1941]" captionTargetBox-1="[171,1425,191,993]" captionTargetId-0="figure-128@1.[164,1423,415,1949]" captionTargetId-1="figure-27@2.[151,1435,181,999]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1–4. Aegidium simplex, new species. 1, holotype male, habitus; 2, paratype female, habitus; 3, holotype male, mesotibia in ventral view; 4, holotype male, labels. Scale bar: 1 mm." captionText-1="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4543191" figureDoi-1="http://doi.org/10.5281/zenodo.4543193" httpUri-0="https://zenodo.org/record/4543191/files/figure.png" httpUri-1="https://zenodo.org/record/4543193/files/figure.png" pageId="0" pageNumber="583">Figs. 1–9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354656EFFB9FFE2FF7FFAEDFEEA40C7" pageId="0" pageNumber="583" type="materials_examined">
<paragraph id="8BF136E5FFB9FFE2FF7FFAEDFEEA40C7" blockId="0.[151,1437,1285,2032]" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF7FFAEDFEC5402C" bold="true" box="[151,318,1359,1384]" inLineHeading="true" pageId="0" pageNumber="583" reason="1">Type material.</emphasis>
<materialsCitation id="3B263CB8FFB9FFE2FEAEFAEDFCA44023" ID-GBIF-Occurrence="3037064302" box="[326,863,1360,1384]" collectionCode="ZMHUB" collectorName="Columbia Dr. Will." pageId="0" pageNumber="583" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F58847FFB9FFE2FEAEFAEDFE574023" box="[326,428,1360,1384]" pageId="0" pageNumber="583" type="holotype">Holotype</typeStatus>
, male: “
<collectorName id="26BB5333FFB9FFE2FDECFAEDFD234023" box="[516,728,1360,1384]" pageId="0" pageNumber="583">Columbia Dr. Will.</collectorName>
” (
<collectionCode id="ED5FAE20FFB9FFE2FD19FAEDFCA34023" box="[753,856,1360,1384]" pageId="0" pageNumber="583">ZMHUB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B263CB8FFB9FFE2FC82FAEDFEF740C7" ID-GBIF-Occurrence="3037064301" collectionCode="ZMHUB" collectorName="Columbia Dr. Will." pageId="0" pageNumber="583" specimenCount="1" typeStatus="paratype">
<typeStatus id="54F58847FFB9FFE2FC82FAEDFC314023" box="[874,970,1360,1384]" pageId="0" pageNumber="583" type="paratype">Paratype</typeStatus>
, female with the same data as for holotype (
<collectionCode id="ED5FAE20FFB9FFE2FF77FAC9FEFD40C7" box="[159,262,1396,1420]" pageId="0" pageNumber="583">ZMHUB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354656EFFB9FFE0FF2FFA2AFAE140E9" lastPageId="2" lastPageNumber="585" pageId="0" pageNumber="583" type="description">
<paragraph id="8BF136E5FFB9FFE2FF2FFA2AFBB3409F" blockId="0.[151,1437,1285,2032]" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF2FFA2AFEA940E4" bold="true" box="[199,338,1431,1455]" inLineHeading="true" pageId="0" pageNumber="583" reason="1">Description.</emphasis>
<typeStatus id="54F58847FFB9FFE2FEB1FA25FE4440FB" box="[345,447,1432,1456]" pageId="0" pageNumber="583" type="holotype">Holotype</typeStatus>
, male (
<figureCitation id="13752A60FFB9FFE2FDE4FA25FD9340FB" box="[524,616,1432,1456]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1973,1997]" captionTargetBox="[190,1400,429,1941]" captionTargetId="figure-128@1.[164,1423,415,1949]" captionTargetPageId="1" captionText="FIGURES 1–4. Aegidium simplex, new species. 1, holotype male, habitus; 2, paratype female, habitus; 3, holotype male, mesotibia in ventral view; 4, holotype male, labels. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543191" httpUri="https://zenodo.org/record/4543191/files/figure.png" pageId="0" pageNumber="583">Fig. 1, 3</figureCitation>
,
<figureCitation id="13752A60FFB9FFE2FD9CFA25FD6440FB" box="[628,671,1432,1456]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="0" pageNumber="583">5–9</figureCitation>
). Body length
<quantity id="4CB69B00FFB9FFE2FCD4FA25FC6640E4" box="[828,925,1432,1456]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="0" pageNumber="583" unit="mm" value="10.9">10.9 mm</quantity>
, width of elytra
<quantity id="4CB69B00FFB9FFE2FBA5FA25FB5A40E4" box="[1101,1185,1432,1456]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="0" pageNumber="583" unit="mm" value="5.2">5.2 mm</quantity>
, width of pronotum
<quantity id="4CB69B00FFB9FFE2FA93FA25FF3F4098" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="0" pageNumber="583" unit="mm" value="4.8">4.8 mm</quantity>
. Upper side of body shiny, color uniform brown, head and pronotum slightly darker.
</paragraph>
<paragraph id="8BF136E5FFB9FFE2FF2FFA5DFBFE432F" blockId="0.[151,1437,1285,2032]" pageId="0" pageNumber="583">Clypeus symmetrical with slightly convex anterior margin and rounded anterior angles. Head surface densely punctate with round punctures separated by less than a puncture diameter on clypeus, becoming denser and coarser on frons near eyes. Head without traces of medial horn or tubercle. Mandibles symmetrical, protruding past anterior margin of clypeus. Labrum rounded, small, slightly protruding past clypeus (in dorsal view).</paragraph>
<paragraph id="8BF136E5FFB9FFE2FF2FF9CDFC03439B" blockId="0.[151,1437,1285,2032]" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF2FF9CDFECA43CC" box="[199,305,1648,1671]" italics="true" pageId="0" pageNumber="583">Pronotum</emphasis>
1.5 times wider than long, widest at middle. Anterior margin with a border interrupted medially by a tubercle. Lateral margins crenulate. Base bordered. Disc of pronotum slightly depressed in middle. Surface densely punctate, punctures separated by about a diameter on disc, becoming denser towards sides.
</paragraph>
<paragraph id="8BF136E5FFB9FFE2FF2FF961FC2043BF" blockId="0.[151,1437,1285,2032]" box="[199,987,1756,1780]" pageId="0" pageNumber="583">
<taxonomicName id="4C4E4D66FFB9FFE2FF2FF961FECB43BF" box="[199,304,1756,1780]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="0" pageNumber="583" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAF7FFB9FFE2FF2FF961FECB43BF" box="[199,304,1756,1780]" italics="true" pageId="0" pageNumber="583">Scutellum</emphasis>
</taxonomicName>
elongate, rounded apically, about 1/12 length of elytra, smooth.
</paragraph>
<paragraph id="8BF136E5FFB9FFE2FF2FF8BDFCA842CF" blockId="0.[151,1437,1285,2032]" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF2FF8BDFEF14253" box="[199,266,1792,1816]" italics="true" pageId="0" pageNumber="583">Elytra</emphasis>
moderately convex on disc with maximum width approximately at middle. Humeral and apical humps distinct. Elytra with 1 feebly distinct longitudinal ridge on disc between suture and humeral hump. First (sutural) elytral interval almost not elevated. Striae indistinct. Elytra densely punctate with coarse, irregularly shaped punctures not arranged in distinct longitudinal rows on disc. Base of elytra feebly bordered.
</paragraph>
<paragraph id="8BF136E5FFB9FFE2FF2FF82DFE4642E3" blockId="0.[151,1437,1285,2032]" box="[199,445,1936,1960]" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF2FF82DFEF342EC" box="[199,264,1936,1959]" italics="true" pageId="0" pageNumber="583">Wings</emphasis>
fully developed.
</paragraph>
<paragraph id="8BF136E5FFB9FFE3FF2FF809FB5A4598" blockId="0.[151,1437,1285,2032]" lastBlockId="1.[151,1436,151,392]" lastPageId="1" lastPageNumber="584" pageId="0" pageNumber="583">
<emphasis id="B93AEAF7FFB9FFE2FF2FF809FED04287" box="[199,299,1972,1996]" italics="true" pageId="0" pageNumber="583">Protibiae</emphasis>
with 3 outer teeth and a smaller inner tooth. Lateral margin basad of outer teeth slightly crenulate. Apex and internal margin of tibia with a few slender setae. Protarsi about 1/2 length of protibiae. Claws 1/3 length of tarsomere 5, which is somewhat longer than tarsomeres 3 and 4 combined, and somewhat thicker than other tarsomeres. Tarsomere 1 as long as tarsomeres 2–4 combined. Ventral surface of femora punctate with rounded punctures.
</paragraph>
<paragraph id="8BF136E5FFB8FFE3FF2FFF62FD2E44C3" blockId="1.[151,1436,151,392]" pageId="1" pageNumber="584">
<emphasis id="B93AEAF7FFB8FFE3FF2FFF62FD7F45BC" box="[199,644,223,247]" italics="true" pageId="1" pageNumber="584">Mesothoracic legs and metathoracic legs</emphasis>
similar in shape; metafemora and metatibiae about 1.2 times longer than mesofemora and mesotibiae. Femora sparsely punctate with rounded punctures. Tibiae somewhat triangular, with two apical spurs. Mesotibiae without tuft of setae ventroapically (
<figureCitation id="13752A60FFB8FFE3FCCDFE9AFC9D440B" box="[805,870,295,320]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1973,1997]" captionTargetBox="[190,1400,429,1941]" captionTargetId="figure-128@1.[164,1423,415,1949]" captionTargetPageId="1" captionText="FIGURES 1–4. Aegidium simplex, new species. 1, holotype male, habitus; 2, paratype female, habitus; 3, holotype male, mesotibia in ventral view; 4, holotype male, labels. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543191" httpUri="https://zenodo.org/record/4543191/files/figure.png" pageId="1" pageNumber="584">Fig. 3</figureCitation>
). Upper tibial spur shorter than tarsomeres 1–3 combined; lower spur shorter than tarsomeres 1–2 combined. Claws 1/3 length of tarsomere 5, which is relatively slender, as long as tarsomere 2 and twice as short as tarsomere 1.
</paragraph>
<caption id="DF31666DFFB8FFE3FF7FF808FCDC42BA" ID-DOI="http://doi.org/10.5281/zenodo.4543191" ID-Zenodo-Dep="4543191" httpUri="https://zenodo.org/record/4543191/files/figure.png" pageId="1" pageNumber="584" startId="1.[151,264,1973,1997]" targetBox="[190,1400,429,1941]" targetPageId="1">
<paragraph id="8BF136E5FFB8FFE3FF7FF808FCDC42BA" blockId="1.[151,1436,1973,2033]" pageId="1" pageNumber="584">
<emphasis id="B93AEAF7FFB8FFE3FF7FF808FEC14286" bold="true" box="[151,314,1973,1997]" pageId="1" pageNumber="584">FIGURES 1–4.</emphasis>
<taxonomicName id="4C4E4D66FFB8FFE3FEA9F808FE0E4286" authority="Frolov &amp; Akhmetova, 2021" authorityName="Frolov &amp; Akhmetova" authorityYear="2021" box="[321,501,1973,1997]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="584" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B93AEAF7FFB8FFE3FEA9F808FE0E4286" box="[321,501,1973,1997]" italics="true" pageId="1" pageNumber="584">Aegidium simplex</emphasis>
</taxonomicName>
,
<emphasis id="B93AEAF7FFB8FFE3FDEAF808FD854286" bold="true" box="[514,638,1973,1997]" pageId="1" pageNumber="584">
<taxonomicNameLabel id="A209578CFFB8FFE3FDEAF808FD854286" box="[514,638,1973,1997]" pageId="1" pageNumber="584" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id="B93AEAF7FFB8FFE3FD63F808FD634286" bold="true" box="[651,664,1973,1997]" pageId="1" pageNumber="584">1</emphasis>
, holotype male, habitus;
<emphasis id="B93AEAF7FFB8FFE3FC7FF808FC5F4286" bold="true" box="[919,932,1973,1997]" pageId="1" pageNumber="584">2</emphasis>
, paratype female, habitus;
<emphasis id="B93AEAF7FFB8FFE3FB5DF808FB394286" bold="true" box="[1205,1218,1973,1997]" pageId="1" pageNumber="584">3</emphasis>
, holotype male, mesotibia in ventral view;
<emphasis id="B93AEAF7FFB8FFE3FE95F864FE7142BA" bold="true" box="[381,394,2009,2033]" pageId="1" pageNumber="584">4</emphasis>
, holotype male, labels. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF31666DFFBBFFE0FF7FFC42FC3E4171" ID-DOI="http://doi.org/10.5281/zenodo.4543193" ID-Zenodo-Dep="4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585" startId="2.[151,264,1023,1047]" targetBox="[171,1425,191,993]" targetPageId="2">
<paragraph id="8BF136E5FFBBFFE0FF7FFC42FC3E4171" blockId="2.[151,1436,1023,1083]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF7FFC42FEC7415C" bold="true" box="[151,316,1023,1047]" pageId="2" pageNumber="585">FIGURES 5–9.</emphasis>
<taxonomicName id="4C4E4D66FFBBFFE0FEACFC42FE01415C" authority="Frolov &amp; Akhmetova, 2021" authorityName="Frolov &amp; Akhmetova" authorityYear="2021" box="[324,506,1023,1047]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B93AEAF7FFBBFFE0FEACFC42FE01415C" box="[324,506,1023,1047]" italics="true" pageId="2" pageNumber="585">Aegidium simplex</emphasis>
</taxonomicName>
,
<emphasis id="B93AEAF7FFBBFFE0FDE0FC42FD7C415C" bold="true" box="[520,647,1023,1047]" pageId="2" pageNumber="585">
<taxonomicNameLabel id="A209578CFFBBFFE0FDE0FC42FD7C415C" box="[520,647,1023,1047]" pageId="2" pageNumber="585" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype, male.
<emphasis id="B93AEAF7FFBBFFE0FCD3FC42FCB3415C" bold="true" box="[827,840,1023,1047]" pageId="2" pageNumber="585">5</emphasis>
, aedeagus in lateral view;
<emphasis id="B93AEAF7FFBBFFE0FBB5FC42FB91415C" bold="true" box="[1117,1130,1023,1047]" pageId="2" pageNumber="585">6</emphasis>
, parameres in dorsal view;
<emphasis id="B93AEAF7FFBBFFE0FA61FC42FA6D415C" bold="true" box="[1417,1430,1023,1047]" pageId="2" pageNumber="585">7</emphasis>
, aedeagus in ventral view;
<emphasis id="B93AEAF7FFBBFFE0FE48FB9EFE564170" bold="true" box="[416,429,1059,1083]" pageId="2" pageNumber="585">8</emphasis>
, spiculum gastrale;
<emphasis id="B93AEAF7FFBBFFE0FD90FB9EFD7E4170" bold="true" box="[632,645,1059,1083]" pageId="2" pageNumber="585">9</emphasis>
, endophallus. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BF136E5FFBBFFE0FF2FFBD6FEDD41EC" blockId="2.[151,1437,1131,1695]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF2FFBD6FE6641C8" box="[199,413,1131,1155]" italics="true" pageId="2" pageNumber="585">Abdominal sternites</emphasis>
punctate with coarse irregularly shaped punctures. Sternite 8 medially about 2 times wider than sternites 4–7.
</paragraph>
<paragraph id="8BF136E5FFBBFFE0FF2FFB0EFAEF4181" blockId="2.[151,1437,1131,1695]" box="[199,1300,1203,1227]" pageId="2" pageNumber="585">
<taxonomicName id="4C4E4D66FFBBFFE0FF2FFB0EFED64180" box="[199,301,1203,1227]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="585" phylum="Chordata" rank="genus">
<emphasis id="B93AEAF7FFBBFFE0FF2FFB0EFED64180" box="[199,301,1203,1227]" italics="true" pageId="2" pageNumber="585">Pygidium</emphasis>
</taxonomicName>
triangular, convex, partly hidden under elytra, irregularly punctate with transverse punctures.
</paragraph>
<paragraph id="8BF136E5FFBBFFE0FF2FFB6AFEDC4010" blockId="2.[151,1437,1131,1695]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF2FFB6AFECD41A4" box="[199,310,1239,1263]" italics="true" pageId="2" pageNumber="585">Aedeagus.</emphasis>
Phallobase 2.1 times longer than parameres, tube shaped (
<figureCitation id="13752A60FFBBFFE0FC4AFB6AFC1941A4" box="[930,994,1239,1263]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585">Fig. 5</figureCitation>
). Parameres reasonably wider than apical part of phallobase in dorsal view (
<figureCitation id="13752A60FFBBFFE0FDE9FB46FDB84058" box="[513,579,1275,1299]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585">Fig. 6</figureCitation>
). Ventrobasal transverse plate of parameres absent (
<figureCitation id="13752A60FFBBFFE0FB82FB46FB574058" box="[1130,1196,1275,1299]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585">Fig. 7</figureCitation>
). Endophallus without symmetrical strongly sclerotized sclerites (
<figureCitation id="13752A60FFBBFFE0FDB6FAA2FD5B407C" box="[606,672,1311,1335]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585">Fig. 9</figureCitation>
). Cranial part of spiculum gastrale narrow, tapering and rounded at very apex (
<figureCitation id="13752A60FFBBFFE0FF30FAFEFEE24010" box="[216,281,1347,1371]" captionStart="FIGURES 5–9" captionStartId="2.[151,264,1023,1047]" captionTargetBox="[171,1425,191,993]" captionTargetId="figure-27@2.[151,1435,181,999]" captionTargetPageId="2" captionText="FIGURES 5–9. Aegidium simplex, new species, holotype, male. 5, aedeagus in lateral view; 6, parameres in dorsal view; 7, aedeagus in ventral view; 8, spiculum gastrale; 9, endophallus. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543193" httpUri="https://zenodo.org/record/4543193/files/figure.png" pageId="2" pageNumber="585">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BF136E5FFBBFFE0FF2FFADAFAE140E9" blockId="2.[151,1437,1131,1695]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF2FFADAFEDA4034" bold="true" box="[199,289,1383,1407]" inLineHeading="true" pageId="2" pageNumber="585" reason="1">Female.</emphasis>
Female (
<figureCitation id="13752A60FFBBFFE0FE6BFADAFE394034" box="[387,450,1383,1407]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1973,1997]" captionTargetBox="[190,1400,429,1941]" captionTargetId="figure-128@1.[164,1423,415,1949]" captionTargetPageId="1" captionText="FIGURES 1–4. Aegidium simplex, new species. 1, holotype male, habitus; 2, paratype female, habitus; 3, holotype male, mesotibia in ventral view; 4, holotype male, labels. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4543191" httpUri="https://zenodo.org/record/4543191/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
) differs from male in having a distinct protibial spur and relatively smaller pronotum without depression and anterior tubercle. Body length 12.0 mm, width of elytra
<quantity id="4CB69B00FFBBFFE0FC7CFA36FC1340E9" box="[916,1000,1419,1443]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="585" unit="mm" value="5.5">5.5 mm</quantity>
, width of pronotum
<quantity id="4CB69B00FFBBFFE0FB29FA36FAED40E9" box="[1217,1302,1419,1443]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="585" unit="mm" value="4.7">4.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C354656EFFBBFFE0FF2FFA12FCCC4330" pageId="2" pageNumber="585" type="diagnosis">
<paragraph id="8BF136E5FFBBFFE0FF2FFA12FCCC4330" blockId="2.[151,1437,1131,1695]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF2FFA12FE39408C" bold="true" box="[199,450,1455,1479]" inLineHeading="true" pageId="2" pageNumber="585" reason="1">Differential diagnosis.</emphasis>
In the key to the South American mainland
<taxonomicName id="4C4E4D66FFBBFFE0FC73FA12FBFA408C" box="[923,1025,1455,1479]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAF7FFBBFFE0FC73FA12FBFA408C" box="[923,1025,1455,1479]" italics="true" pageId="2" pageNumber="585">Aegidium</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFDF4B14FFBBFFE0FB8BFA12FACD408C" author="Frolov, A. V. &amp; Akhmetova, L. A. &amp; Vaz-de-Mello, F. Z." box="[1123,1334,1455,1479]" pageId="2" pageNumber="585" pagination="1035 - 1090" refId="ref1713" refString="Frolov, A. V., Akhmetova, L. A. &amp; Vaz-de-Mello, F. Z. (2017 b) Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae). Journal of Natural History, 51, 1035 - 1090. https: // doi. org / 10.1080 / 00222933.2017.1319519" type="journal article" year="2017">
Frolov
<emphasis id="B93AEAF7FFBBFFE0FB5BFA0DFB19408C" box="[1203,1250,1455,1479]" italics="true" pageId="2" pageNumber="585">et al</emphasis>
. 2017b
</bibRefCitation>
), the new species comes out in the couplet with
<taxonomicName id="4C4E4D66FFBBFFE0FDA8FA6EFC5740A1" authority="Preudhomme de Borre" authorityName="Preudhomme de Borre" authorityYear="1886" box="[576,940,1491,1515]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="reichei">
<emphasis id="B93AEAF7FFBBFFE0FDA8FA6EFD5240A0" box="[576,681,1491,1515]" italics="true" pageId="2" pageNumber="585">A. reichei</emphasis>
Preudhomme de Borre
</taxonomicName>
,
<taxonomicName id="4C4E4D66FFBBFFE0FC52FA6EFAB340A0" authority="Frolov &amp; Akhmetova" authorityName="Frolov &amp; Akhmetova" authorityYear="2020" box="[954,1352,1491,1515]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="bolivianum">
<emphasis id="B93AEAF7FFBBFFE0FC52FA6EFBA940A0" box="[954,1106,1491,1515]" italics="true" pageId="2" pageNumber="585">A. bolivianum</emphasis>
Frolov &amp; Akhmetova
</taxonomicName>
, and
<taxonomicName id="4C4E4D66FFBBFFE0FA6DFA6EFE414344" authority="Frolov, Akhmetova" authorityName="Frolov, Akhmetova" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B93AEAF7FFBBFFE0FA6DFA6EFF1C4344" italics="true" pageId="2" pageNumber="585">A. varians</emphasis>
Frolov, Akhmetova
</taxonomicName>
&amp; Vaz-de-Mello. All these species are similar in having the mesotibiae without a tuft of setae ventroapically and in the lack of ventrobasal transverse plate of parameres.
<taxonomicName id="4C4E4D66FFBBFFE0FC5CF9A6FB8A4378" authorityName="Frolov &amp; Akhmetova" authorityYear="2021" box="[948,1137,1563,1587]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B93AEAF7FFBBFFE0FC5CF9A6FB8A4378" box="[948,1137,1563,1587]" italics="true" pageId="2" pageNumber="585">Aegidium simplex</emphasis>
</taxonomicName>
can be separated from these species by the parameres being wider in lateral view and the length/width ratio is approximately 2.0 (
<emphasis id="B93AEAF7FFBBFFE0FB0BF9FDFADC431C" box="[1251,1319,1600,1623]" italics="true" pageId="2" pageNumber="585">versus</emphasis>
being narrower in lateral view and the length/width ratio being 3.5–4.0).
</paragraph>
</subSubSection>
<subSubSection id="C354656EFFBBFFE0FF2FF93AFBFB43D4" box="[199,1024,1671,1695]" pageId="2" pageNumber="585" type="etymology">
<paragraph id="8BF136E5FFBBFFE0FF2FF93AFBFB43D4" blockId="2.[151,1437,1131,1695]" box="[199,1024,1671,1695]" pageId="2" pageNumber="585">
<emphasis id="B93AEAF7FFBBFFE0FF2FF93AFEB243D5" box="[199,329,1671,1695]" inLineHeading="true" pageId="2" pageNumber="585" reason="1">
<emphasis id="B93AEAF7FFBBFFE0FF2FF93AFEB943D4" bold="true" box="[199,322,1671,1695]" pageId="2" pageNumber="585">Etymology</emphasis>
.
</emphasis>
From Latin, “
<taxonomicName id="4C4E4D66FFBBFFE0FE09F93AFDC943D4" authorityName="Frolov &amp; Akhmetova" authorityYear="2021" box="[481,562,1671,1695]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="585" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B93AEAF7FFBBFFE0FE09F93AFDC943D4" box="[481,562,1671,1695]" italics="true" pageId="2" pageNumber="585">simplex</emphasis>
</taxonomicName>
” for an ordinary appearance of the beetles.
</paragraph>
</subSubSection>
</treatment>
</document>
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