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<document id="8FE1D1717D7A62CD08E672525B644CDE" ID-DOI="10.5281/zenodo.179727" ID-GBIF-Dataset="de333d52-b2e2-47cb-b3de-5c313a86ad12" ID-ISSN="1175-5326" ID-Zenodo-Dep="179727" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459987403313" checkinUser="plazi" docAuthor="Zhang, Kuiyan, Yang, Ding &amp; Shih, Dong Ren Chungkun" docDate="2007" docId="03E787C1FFE1FF92FF6BFC2AFBBDF8A1" docLanguage="en" docName="zt01645p017.pdf" docOrigin="Zootaxa 1645" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Calosargus (Calosargus) daohugouensis Zhang, Yang &amp; Ren, sp. nov." docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FFDEFFB9FFE9FF9BFFFCFFA1FFC9FFA5" masterDocTitle="The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1698232378857" updateUser="plazi">
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<mods:title id="87197F66BA12BCED6839139A5EF18242">The oldest Calosargus Mostovski, 1997 from the Middle Jurassic of China (Diptera: Brachycera: Archisargidae)</mods:title>
</mods:titleInfo>
<mods:name id="CCEC303494E9081C121472E5FD4972D9" type="personal">
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<mods:namePart id="468EE850B5481C8C5F7F888D5D4A387F">Zhang, Kuiyan</mods:namePart>
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<mods:namePart id="E553A797CC27A5879559CB5C3A75A25C">Yang, Ding</mods:namePart>
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<mods:namePart id="FDFC29BDA5B99CDBC5B22BA92C2716A4">Shih, Dong Ren Chungkun</mods:namePart>
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<mods:title id="8E89609E5988F33143768C7BD549D550">Zootaxa</mods:title>
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<mods:part id="6B5D5B4F8ADFBB84A64563D38605D31A">
<mods:date id="A8A6B30655941530DCF88AB58A11FC71">2007</mods:date>
<mods:detail id="58B29C7E76FBC2F509C07DE5733EC24C" type="volume">
<mods:number id="053B1841390542FCB90BF430FC0385D0">1645</mods:number>
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<mods:identifier id="DF70E46C68E7BB1981A21420172CF97C" type="DOI">10.5281/zenodo.179727</mods:identifier>
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<mods:identifier id="8F94B0A805DB0D8C507FA91FEA62D3C9" type="Zenodo-Dep">179727</mods:identifier>
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<treatment id="03E787C1FFE1FF92FF6BFC2AFBBDF8A1" ID-DOI="http://doi.org/10.5281/zenodo.6236727" ID-GBIF-Taxon="119379546" ID-Zenodo-Dep="6236727" LSID="urn:lsid:plazi:treatment:03E787C1FFE1FF92FF6BFC2AFBBDF8A1" httpUri="http://treatment.plazi.org/id/03E787C1FFE1FF92FF6BFC2AFBBDF8A1" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C354655CFFE1FF93FF6BFC2AFEFDFC62" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF136D7FFE1FF93FF6BFC2AFC24FC01" blockId="8.[151,1005,906,967]" box="[151,1005,906,933]" pageId="8" pageNumber="9">
<heading id="D0B981BBFFE1FF93FF6BFC2AFC24FC01" bold="true" box="[151,1005,906,933]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B93AEAC5FFE1FF93FF6BFC2AFC24FC01" bold="true" box="[151,1005,906,933]" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D54FFE1FF93FF6BFC2AFC4BFC01" ID-CoL="8BB9T" authority="Zhang, Yang &amp; Ren" authorityName="Zhang, Yang &amp; Ren" box="[151,898,906,933]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov." subGenus="Calosargus">
<emphasis id="B93AEAC5FFE1FF93FF6BFC2AFEE9FC00" bold="true" box="[151,288,907,933]" italics="true" pageId="8" pageNumber="9">Calosargus</emphasis>
(
<emphasis id="B93AEAC5FFE1FF93FECDFC2AFE73FC00" bold="true" box="[305,442,907,933]" italics="true" pageId="8" pageNumber="9">Calosargus</emphasis>
)
<emphasis id="B93AEAC5FFE1FF93FE37FC2AFD48FC00" bold="true" box="[459,641,907,933]" italics="true" pageId="8" pageNumber="9">daohugouensis</emphasis>
Zhang, Yang &amp; Ren
</taxonomicName>
,
<taxonomicNameLabel id="A20957BEFFE1FF93FC6CFC2BFC24FC01" box="[912,1005,906,932]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF136D7FFE1FF93FF6BFC0CFEFDFC62" blockId="8.[151,1005,906,967]" box="[151,308,941,967]" pageId="8" pageNumber="9">
(
<figureCitation id="13752A52FFE1FF93FF63FC0CFEE2FC62" box="[159,299,941,967]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="8.[151,255,1935,1959]" captionStartId-1="9.[151,255,777,801]" captionTargetBox-0="[160,1432,1244,1899]" captionTargetBox-1="[177,1393,200,756]" captionTargetId-0="figure@8.[160,1433,1244,1901]" captionTargetId-1="figure@9.[159,1427,179,768]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 11. Calosargus (Calosargus) daohugouensis sp. nov. Body with wing photograph in lateral view." captionText-1="FIGURE 12. Calosargus (Calosargus) daohugouensis sp. nov. Camera lucida drawing, based on the original photograph, in lateral view." httpUri-0="https://zenodo.org/record/179734/files/figure.png" httpUri-1="https://zenodo.org/record/179735/files/figure.png" pageId="8" pageNumber="9">Figs. 1112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C354655CFFE1FF93FF6BFC5CFB42FBC8" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BF136D7FFE1FF93FF6BFC5CFB42FBC8" blockId="8.[151,1436,1021,1176]" pageId="8" pageNumber="9">
<emphasis id="B93AEAC5FFE1FF93FF6BFC5CFEDEFBB2" bold="true" box="[151,279,1021,1047]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Vei n R s 1 short, nearly as long as vein Rs2; vein R2+3 arising slightly proximal of base of cell d; vein R2+3 ending at vein R1 far before wing margin; crossvein r-m located at basal 1/3 of cell d; mouth of cell r1 narrow, about 1/4 as wide as that of cell r4; mouth of cell m1 narrower than that of cell m2.
</paragraph>
</subSubSection>
<subSubSection id="C354655CFFE1FF93FF3AFBDFFAA9FB3D" box="[198,1376,1150,1176]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BF136D7FFE1FF93FF3AFBDFFAA9FB3D" blockId="8.[151,1436,1021,1176]" box="[198,1376,1150,1176]" pageId="8" pageNumber="9">
<emphasis id="B93AEAC5FFE1FF93FF3AFBDFFE9AFB3D" bold="true" box="[198,339,1150,1176]" pageId="8" pageNumber="9">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="54F58875FFE1FF93FD30FBDFFD37FB3D" box="[716,766,1150,1176]" pageId="8" pageNumber="9">type</typeStatus>
locality: Daohugou village, Inner
<collectingCountry id="F3597647FFE1FF93FB6EFBDFFACEFB3D" box="[1170,1287,1150,1176]" name="Mongolia" pageId="8" pageNumber="9">Mongolia</collectingCountry>
,
<collectingCountry id="F3597647FFE1FF93FAEEFBDFFA95FB3D" box="[1298,1372,1150,1176]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF31665FFFE1FF93FF6BF82EFAC7F803" box="[151,1294,1935,1959]" httpUri="https://zenodo.org/record/179734/files/figure.png" pageId="8" pageNumber="9" targetBox="[160,1432,1244,1899]" targetPageId="8">
<paragraph id="8BF136D7FFE1FF93FF6BF82EFAC7F803" blockId="8.[151,1294,1935,1959]" box="[151,1294,1935,1959]" pageId="8" pageNumber="9">
<emphasis id="B93AEAC5FFE1FF93FF6BF82EFEECF802" bold="true" box="[151,293,1935,1959]" pageId="8" pageNumber="9">FIGURE 11.</emphasis>
<taxonomicName id="4C4E4D54FFE1FF93FED0F82EFD17F803" box="[300,734,1935,1958]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov." subGenus="Calosargus">
<emphasis id="B93AEAC5FFE1FF93FED0F82EFE6FF803" box="[300,422,1935,1958]" italics="true" pageId="8" pageNumber="9">Calosargus</emphasis>
(
<emphasis id="B93AEAC5FFE1FF93FE49F82EFDE6F803" box="[437,559,1935,1958]" italics="true" pageId="8" pageNumber="9">Calosargus</emphasis>
)
<emphasis id="B93AEAC5FFE1FF93FDC2F82EFD17F803" box="[574,734,1935,1958]" italics="true" pageId="8" pageNumber="9">daohugouensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957BEFFE1FF93FD19F82EFCF1F803" box="[741,824,1935,1958]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
Body with wing photograph in lateral view.
</paragraph>
</caption>
<caption id="DF31665FFFE0FF92FF6BFCA8FEB6FCE2" httpUri="https://zenodo.org/record/179735/files/figure.png" pageId="9" pageNumber="10" targetBox="[177,1393,200,756]" targetPageId="9">
<paragraph id="8BF136D7FFE0FF92FF6BFCA8FEB6FCE2" blockId="9.[151,1435,777,839]" pageId="9" pageNumber="10">
<emphasis id="B93AEAC5FFE0FF92FF6BFCA8FEE0FC84" bold="true" box="[151,297,777,801]" pageId="9" pageNumber="10">FIGURE 12.</emphasis>
<taxonomicName id="4C4E4D54FFE0FF92FECFFCABFD25FC84" box="[307,748,777,801]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov." subGenus="Calosargus">
<emphasis id="B93AEAC5FFE0FF92FECFFCABFE64FC84" box="[307,429,778,801]" italics="true" pageId="9" pageNumber="10">Calosargus</emphasis>
(
<emphasis id="B93AEAC5FFE0FF92FE3CFCABFDF3FC84" box="[448,570,778,801]" italics="true" pageId="9" pageNumber="10">Calosargus</emphasis>
)
<emphasis id="B93AEAC5FFE0FF92FDB0FCABFD25FC84" box="[588,748,778,801]" italics="true" pageId="9" pageNumber="10">daohugouensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957BEFFE0FF92FD0AFCA8FC85FC85" box="[758,844,777,800]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
Camera lucida drawing, based on the original photograph, in lateral view.
</paragraph>
</caption>
<subSubSection id="C354655CFFE0FF92FF3AFCDFFC09FC4C" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BF136D7FFE0FF92FF3AFCDFFAB2FC3C" blockId="9.[151,1436,894,1800]" box="[198,1403,894,921]" pageId="9" pageNumber="10">
<emphasis id="B93AEAC5FFE0FF92FF3AFCDFFE89FC3D" bold="true" box="[198,320,894,920]" pageId="9" pageNumber="10">
<typeStatus id="54F58875FFE0FF92FF3AFCDFFEF2FC3D" box="[198,315,894,920]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
.
</emphasis>
CNU-DB-NN2007008, an almost complete adult of unknown sex with wing in lateral view.
</paragraph>
<paragraph id="8BF136D7FFE0FF92FF3AFC07FC09FC4C" blockId="9.[151,1436,894,1800]" pageId="9" pageNumber="10">
<emphasis id="B93AEAC5FFE0FF92FF3AFC07FDD9FC65" bold="true" box="[198,528,934,960]" pageId="9" pageNumber="10">
<typeStatus id="54F58875FFE0FF92FF3AFC07FECAFC65" box="[198,259,934,960]" pageId="9" pageNumber="10">Type</typeStatus>
locality and horizon.
</emphasis>
Daohugou Village, Shantou Township, Ningcheng County, Inner
<collectingCountry id="F3597647FFE0FF92FADFFC06FA51FC64" box="[1315,1432,935,961]" name="Mongolia" pageId="9" pageNumber="10">Mongolia</collectingCountry>
,
<collectingCountry id="F3597647FFE0FF92FF6BFC6EFF28FC4C" box="[151,225,975,1001]" name="China" pageId="9" pageNumber="10">China</collectingCountry>
; Jiulongshan Formation, Middle Jurassic (Aalenian-Bajocian).
</paragraph>
</subSubSection>
<subSubSection id="C354655CFFE0FF92FF3AFC57FB4BFA10" pageId="9" pageNumber="10" type="description">
<paragraph id="8BF136D7FFE0FF92FF3AFC57FB87FBB4" blockId="9.[151,1436,894,1800]" box="[198,1102,1014,1041]" pageId="9" pageNumber="10">
<emphasis id="B93AEAC5FFE0FF92FF3AFC57FE97FBB5" bold="true" box="[198,350,1014,1040]" pageId="9" pageNumber="10">Description.</emphasis>
Body length 8.0 mm; wing length
<quantity id="4CB69B32FFE0FF92FD04FC56FC9CFBB4" box="[760,853,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="10" unit="mm" value="7.8">7.8 mm</quantity>
, wing width
<quantity id="4CB69B32FFE0FF92FC11FC56FB82FBB4" box="[1005,1099,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="10" unit="mm" value="2.5">2.5 mm</quantity>
.
</paragraph>
<paragraph id="8BF136D7FFE0FF92FF3AFBBEFAA3FB9C" blockId="9.[151,1436,894,1800]" box="[198,1386,1055,1081]" pageId="9" pageNumber="10">Head round in lateral view, slightly narrower than thorax. Eyes large, bare; ommatidia visible clearly.</paragraph>
<paragraph id="8BF136D7FFE0FF92FF3AFBE6FDDBFA2F" blockId="9.[151,1436,894,1800]" pageId="9" pageNumber="10">Thorax black. Middle and hind femora long, slightly swollen; hind tibia slender. Wing elongate, hyaline. Vein C ending at wing tip; veins C and R1 markedly thick; vein Sc long, extending far beyond middle of wing. Ve in R 1 long and robust; vein Rs1 short, nearly as long as vein Rs2; vein R2+3 arising slightly proximal of base of cell d; vein R2+3 ending at vein R1 far before wing margin; vein R4 short, convergent to vein R1; vein R5 ending far beyond apex of wing, relative lengths of vein R5 and vein Rs3 about 3: 5. Crossvein r-m located at basal 1/3 of cell d. Vein M3 absent, vein CuA1 arising from cell d. Cell sc wide open, mouth of cell sc about 3 times as wide as that of cell r4, mouth of cell r1 narrow, about 1/4 as wide as that of cell r4. Cell d wide, mouth of cell m1 narrower than that of cell m2.</paragraph>
<paragraph id="8BF136D7FFE0FF92FF3AFA3AFB4BFA10" blockId="9.[151,1436,894,1800]" box="[198,1154,1435,1461]" pageId="9" pageNumber="10">Abdomen slender, cylindrical. Three segments visible. Segment I evidently short.</paragraph>
</subSubSection>
<subSubSection id="C354655CFFE0FF92FF3AFA62FBBDF8A1" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BF136D7FFE0FF92FF3AFA62FBBDF8A1" blockId="9.[151,1436,894,1800]" pageId="9" pageNumber="10">
<emphasis id="B93AEAC5FFE0FF92FF3AFA62FEF6FA78" bold="true" box="[198,319,1475,1501]" pageId="9" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="4C4E4D54FFE0FF92FEB5FA65FD9CFA78" box="[329,597,1475,1501]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="daohugouensis" status="sp. nov." subGenus="Calosargus">
<emphasis id="B93AEAC5FFE0FF92FEB5FA65FEADFA78" box="[329,356,1476,1501]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
(
<emphasis id="B93AEAC5FFE0FF92FE8BFA65FE5BFA78" box="[375,402,1476,1501]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
)
<emphasis id="B93AEAC5FFE0FF92FE59FA65FD9CFA78" box="[421,597,1476,1501]" italics="true" pageId="9" pageNumber="10">daohugouensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957BEFFE0FF92FDA2FA62FD72FA78" box="[606,699,1475,1501]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
could be related to
<taxonomicName id="4C4E4D54FFE0FF92FC50FA65FBA7FA78" box="[940,1134,1475,1501]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="antiquus" status="sp. nov." subGenus="Calosargus">
<emphasis id="B93AEAC5FFE0FF92FC50FA65FC0EFA78" box="[940,967,1476,1501]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
(
<emphasis id="B93AEAC5FFE0FF92FC27FA65FC3CFA78" box="[987,1013,1476,1501]" italics="true" pageId="9" pageNumber="10">C.</emphasis>
)
<emphasis id="B93AEAC5FFE0FF92FBF5FA65FBA7FA78" box="[1033,1134,1476,1501]" italics="true" pageId="9" pageNumber="10">antiquus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957BEFFE0FF92FB8BFA62FB1DFA78" box="[1143,1236,1475,1501]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
in the following aspects: vein R2+3 ending at vein R1 far before wing margin; veins R4 and R5 short; base of cell d sharp. However, we can easily separate it from
<taxonomicName id="4C4E4D54FFE0FF92FDB4F9B6FD64F995" box="[584,685,1559,1584]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="antiquus">
<emphasis id="B93AEAC5FFE0FF92FDB4F9B6FD64F995" box="[584,685,1559,1584]" italics="true" pageId="9" pageNumber="10">antiquus</emphasis>
</taxonomicName>
by vein Rs1 short, nearly as long as vein Rs2; vein R2+3 arising slightly proximal of base of cell d; crossvein r-m located at basal 1/3 of cell d; veins M distinctly long; mouth of cell r1 about 1/4 as wide as that of cell r4; mouth of cell m1 narrower than that of cell m2. In
<taxonomicName id="4C4E4D54FFE0FF92FB09F9C8FA93F927" box="[1269,1370,1641,1666]" class="Insecta" family="Archisargidae" genus="Calosargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="antiquus">
<emphasis id="B93AEAC5FFE0FF92FB09F9C8FA93F927" box="[1269,1370,1641,1666]" italics="true" pageId="9" pageNumber="10">antiquus</emphasis>
</taxonomicName>
, vein Rs1 is long, about twice as long as vein Rs2; vein R2+3 is arising slightly distal of the base of cell d; crossvein rm is nearly located at middle of cell d; the branches of vein M are distinctly short; the mouth of cell r1 is about 1/2 as wide as that of cell r4; the mouth of cell m1 is nearly as wide as that of cell m2.
</paragraph>
</subSubSection>
</treatment>
</document>