treatments-xml/data/03/E4/87/03E487B6FFD5FFCEFF41886FFC4F791A.xml

<document id="52800A6795CEAC97332F8B4852011F0E" ID-CLB-Dataset="6096" ID-DOI="10.11646/zootaxa.4980.1.8" ID-GBIF-Dataset="465ec546-64e0-4ba3-84bb-b485feab12ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="4883127" ID-ZooBank="DF5C6B16-642A-4238-A4C2-AFDF9B2A1A29" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622463809428" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Cz-, Petr Baňař" docDate="2021" docId="03E487B6FFD5FFCEFF41886FFC4F791A" docLanguage="en" docName="zootaxa.4980.1.8.pdf" docOrigin="Zootaxa 4980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astenus ambohitantelyensis Janak &amp; Banar 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="130" masterDocId="FFDDFFCEFFD4FFCDFFD68B57FFDD7A0F" masterDocTitle="Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="141" masterPageNumber="127" pageNumber="128" updateTime="1698995102808" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BDE847F14A367EF6C9F81156EE38A9DE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F88EB7EB2C997EBB453C8D395737EB40">
<mods:title id="D3CD18811BA4ACAA692764A05C1652E0">Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="116B27B0C2DD5A0512329550DD2290AA" type="personal">
<mods:role id="3BE1B144E30B07F67BE6D39F686A436C">
<mods:roleTerm id="B27607E16B8397DDEC58701B53CEB8A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BC5A9D7C2E970339583CF30C2CAC84F">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="786C68C9174759C1B8F6F7FAD37FBB5A" type="personal">
<mods:role id="BEFA895FEE8BC13D7D73819C63A5586A">
<mods:roleTerm id="415CF69079D9B9AF7167F0CC7B75A5E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B55D23C5F329419687D695E71AEAD50">Cz-, Petr Baňař</mods:namePart>
</mods:name>
<mods:typeOfResource id="90D36321C65D214B3D92720B49438FB6">text</mods:typeOfResource>
<mods:relatedItem id="921F9E627722BAE105D26AC59E5F8773" type="host">
<mods:titleInfo id="933A5132C03700CF6CDD403EB5BD07C4">
<mods:title id="42F13D51AF74D0FA4518D986E69006C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F5695171DD2E5EC1D29CB6DDA475A2E6">
<mods:date id="853D84B4ADC77882FA6E3B25E41FC0EE">2021</mods:date>
<mods:detail id="056AC2D175E2F9644AE3238B72CEF41B" type="pubDate">
<mods:number id="5D93C49F5EBC1E0CC560C34D1DADFDF5">2021-05-31</mods:number>
</mods:detail>
<mods:detail id="D177A701419AF2F76CDBB3C133AA77DD" type="volume">
<mods:number id="038F907EF2525B9DDFD0CB010173C47B">4980</mods:number>
</mods:detail>
<mods:detail id="3A28BC039A5F41AFAF8555D4649E3B7F" type="issue">
<mods:number id="8D87CD94D70753ECB9D6C3BF78259E74">1</mods:number>
</mods:detail>
<mods:extent id="2CD46BE746735C7F85ED2E62A9C74983" unit="page">
<mods:start id="16E4B83210DA9EE3878C67773CEE3F65">127</mods:start>
<mods:end id="7B2663ADC412DD9C37DB83245D2C23F9">141</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7635D085891BA06D25DA2674677A7521">journal article</mods:classification>
<mods:identifier id="04EBE249549313C57BEF3C24FEB5D5E8" type="CLB-Dataset">6096</mods:identifier>
<mods:identifier id="D7EFE00D340E6282FB291420D51D0C16" type="DOI">10.11646/zootaxa.4980.1.8</mods:identifier>
<mods:identifier id="290E10C843D2BB5718345C53D1BB64B1" type="GBIF-Dataset">465ec546-64e0-4ba3-84bb-b485feab12ab</mods:identifier>
<mods:identifier id="D2EB88B429664B72B915D3AF5E059010" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B2CAF4A128F7581B0E2B128A3520AA7B" type="Zenodo-Dep">4883127</mods:identifier>
<mods:identifier id="CB8B0E930C8F4980240C45B203861289" type="ZooBank">DF5C6B16-642A-4238-A4C2-AFDF9B2A1A29</mods:identifier>
</mods:mods>
<treatment id="03E487B6FFD5FFCEFF41886FFC4F791A" ID-DOI="http://doi.org/10.5281/zenodo.4883131" ID-GBIF-Taxon="182741903" ID-Zenodo-Dep="4883131" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A" httpUri="http://treatment.plazi.org/id/03E487B6FFD5FFCEFF41886FFC4F791A" lastPageId="3" lastPageNumber="130" pageId="1" pageNumber="128">
<subSubSection id="C357652BFFD5FFCCFF41886FFCC0795D" box="[151,797,823,850]" pageId="1" pageNumber="128" type="nomenclature">
<paragraph id="8BF236A0FFD5FFCCFF41886FFCC0795D" blockId="1.[151,797,823,886]" box="[151,797,823,850]" pageId="1" pageNumber="128">
<heading id="D0BA81CCFFD5FFCCFF41886FFCC0795D" bold="true" box="[151,797,823,850]" fontSize="11" level="1" pageId="1" pageNumber="128" reason="1">
<emphasis id="B939EAB2FFD5FFCCFF41886FFCC0795D" bold="true" box="[151,797,823,850]" pageId="1" pageNumber="128">
<taxonomicName id="4C4D4D23FFD5FFCCFF41886FFD6E795D" ID-CoL="8XV53" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[151,691,823,850]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">
<emphasis id="B939EAB2FFD5FFCCFF41886FFE35795D" bold="true" box="[151,488,823,850]" italics="true" pageId="1" pageNumber="128">Astenus ambohitantelyensis</emphasis>
Janák &amp; Baňař
</taxonomicName>
, 
<taxonomicNameLabel id="A20A57C9FFD5FFCCFD16886FFCC0795D" box="[704,797,824,850]" pageId="1" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD5FFCCFF41880BFEAE7979" box="[151,371,860,886]" pageId="1" pageNumber="128" type="description">
<paragraph id="8BF236A0FFD5FFCCFF41880BFEAE7979" blockId="1.[151,797,823,886]" box="[151,371,860,886]" pageId="1" pageNumber="128">
(
<figureCitation id="13762A25FFD5FFCCFF49880BFECA7979" box="[159,279,860,886]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Figs. 1–11</figureCitation>
, 
<figureCitation id="13762A25FFD5FFCCFEF2880BFEB67979" box="[292,363,860,886]" captionStart="FIGURES 40–41" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[171,1416,181,1920]" captionTargetId="figure-17@9.[171,1416,181,1920]" captionTargetPageId="9" captionText="FIGURES 40–41. Forest of Ambohitantely Special Reserve. 41, View from administrative buildings close to main entrance; 42, detail of forest undergrowth." figureDoi="http://doi.org/10.5281/zenodo.4883137" httpUri="https://zenodo.org/record/4883137/files/figure.png" pageId="1" pageNumber="128">40–41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD5FFCCFF4188F3FCB87F45" pageId="1" pageNumber="128" type="materials_examined">
<paragraph id="8BF236A0FFD5FFCCFF4188F3FCEC79B0" blockId="1.[151,1437,932,2038]" box="[151,817,932,959]" pageId="1" pageNumber="128">
<materialsCitation id="3B253CFDFFD5FFCCFF4188F3FCEC79B0" ID-GBIF-Occurrence="3131331341" box="[151,817,932,959]" country="Madagascar" location="Ambohitantely Special Reserve" pageId="1" pageNumber="128" specimenCount="1" typeStatus="holotype">
<emphasis id="B939EAB2FFD5FFCCFF4188F3FEEB79B1" bold="true" box="[151,310,932,958]" pageId="1" pageNumber="128">
<typeStatus id="54F68802FFD5FFCCFF4188F3FF0F79B1" box="[151,210,932,958]" pageId="1" pageNumber="128">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35A7630FFD5FFCCFEEA88F3FE1879B1" box="[316,453,932,958]" name="Madagascar" pageId="1" pageNumber="128">Madagascar</collectingCountry>
, 
<location id="8E92607BFFD5FFCCFE1988F3FCEC79B0" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A:8E92607BFFD5FFCCFE1988F3FCEC79B0" box="[463,817,932,959]" country="Madagascar" name="Ambohitantely Special Reserve" pageId="1" pageNumber="128">Ambohitantely Special Reserve</location>
</materialsCitation>
</paragraph>
<paragraph id="8BF236A0FFD5FFCCFF11889FFCB87F45" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
<emphasis id="B939EAB2FFD5FFCCFF11889FFE5C79ED" bold="true" box="[199,385,968,994]" pageId="1" pageNumber="128">Type material.</emphasis>
<materialsCitation id="3B253CFDFFD5FFCCFE45889FFED87E41" ID-GBIF-Occurrence="3131331355" collectingDate="2011-09" collectingDateMax="2011-09-05" collectingDateMin="2011-09" collectionCode="MMBC" collectorName="L. S. Rahanitriniaina" country="Madagascar" determinerName="Banar" elevation="1640" latitude="-18.19536" location="MADAGASCAR" longLatPrecision="1" longitude="47.28958" pageId="1" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B939EAB2FFD5FFCCFE45889FFDDD79ED" bold="true" box="[403,512,968,994]" pageId="1" pageNumber="128">
<typeStatus id="54F68802FFD5FFCCFE45889FFDDD79ED" box="[403,512,968,994]" pageId="1" pageNumber="128" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D4BFD29FFD5FFCCFDC5889EFDF679EC" box="[531,555,969,995]" pageId="1" pageNumber="128" type="male">♂</specimenCount>
: “ABT/ 
<date id="FFF31060FFD5FFCCFD58889FFD2079ED" box="[654,765,968,994]" pageId="1" pageNumber="128" value="2011-09">
<collectingDate id="EFB7E988FFD5FFCCFD58889FFD2079ED" box="[654,765,968,994]" pageId="1" pageNumber="128" value="2011-09">Sept.2011</collectingDate>
</date>
/24 
<collectingCountry id="F35A7630FFD5FFCCFCE3889FFC2279ED" box="[821,1023,968,994]" name="Madagascar" pageId="1" pageNumber="128">MADAGASCAR</collectingCountry>
, AMBOHITANTELY Spec. Res.; 
<date id="FFF31060FFD5FFCCFF4188BBFEDE7E08" box="[151,259,1004,1031]" pageId="1" pageNumber="128" value="2011-09-05">
<collectingDate id="EFB7E988FFD5FFCCFF4188BBFEDE7E08" box="[151,259,1004,1031]" pageId="1" pageNumber="128" value="2011-09-05">5.ix.2011</collectingDate>
</date>
; 
<geoCoordinate id="EE795067FFD5FFCCFEC788BBFE737E09" box="[273,430,1004,1030]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="1" pageNumber="128" precision="1" seconds="43.3" value="-18.19536">S18°11’43.3‘‘</geoCoordinate>
<geoCoordinate id="EE795067FFD5FFCCFE6188BBFD857E08" box="[439,600,1004,1031]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="128" precision="1" seconds="22.5" value="47.28958">E47°17’22.5‘‘</geoCoordinate>
, 
<quantity id="4CB59B45FFD5FFCCFDB188BBFD647E09" box="[615,697,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="1" pageNumber="128" unit="m" value="1640.0">
<elevation id="0060D193FFD5FFCCFDB188BBFD647E09" box="[615,697,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="1" pageNumber="128" unit="m" value="1640.0">1640m</elevation>
</quantity>
; sifting of forest litter under palm tree, Winkler app. Extraction, 
<collectorName id="26B85376FFD5FFCCFF418F47FE5F7E25" box="[151,386,1040,1066]" pageId="1" pageNumber="128">L.S. Rahanitriniaina</collectorName>
lgt. // 
<typeStatus id="54F68802FFD5FFCCFE348F47FD557E25" box="[482,648,1040,1066]" pageId="1" pageNumber="128" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C4D4D23FFD5FFCCFD408F47FC0A7E25" authority="Janák &amp; Cz-, 2021" authorityName="Janák &amp; Cz-" authorityYear="2021" box="[662,983,1040,1066]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">Astenus ambohitantelyensis</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD5FFCCFC308F46FB9A7E24" box="[998,1095,1041,1067]" pageId="1" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp; 
<determinerName id="7E2EDA64FFD5FFCCFB188F47FACC7E25" box="[1230,1297,1040,1066]" pageId="1" pageNumber="128">Baňař</determinerName>
det. 2020” (
<collectionCode id="ED5CAE65FFD5FFCCFF498F63FF217E41" box="[159,252,1076,1102]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="128">MMBC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B253CFDFFD5FFCCFED98F63FC7F7E41" ID-GBIF-Occurrence="3131331304" box="[271,930,1076,1103]" collectionCode="MMBC" country="Madagascar" pageId="1" pageNumber="128" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B939EAB2FFD5FFCCFED98F63FE5A7E41" bold="true" box="[271,391,1076,1102]" pageId="1" pageNumber="128">
<typeStatus id="54F68802FFD5FFCCFED98F63FE5A7E41" box="[271,391,1076,1102]" pageId="1" pageNumber="128" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D4BFD29FFD5FFCCFE408F63FE1E7E40" box="[406,451,1076,1103]" pageId="1" pageNumber="128" type="female">1 ♀</specimenCount>
: first label same as the 
<typeStatus id="54F68802FFD5FFCCFD1D8F63FCF07E41" box="[715,813,1076,1102]" pageId="1" pageNumber="128" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED5CAE65FFD5FFCCFCEA8F63FC447E41" box="[828,921,1076,1102]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="128">MMBC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B253CFDFFD5FFCCFC7B8F63FD347E99" ID-GBIF-Occurrence="3131331373" collectingDate="2011-09" collectingDateMax="2011-09-03" collectingDateMin="2011-09" collectionCode="JJRC" collectorName="L. S. Rahanitriniaina" country="Madagascar" elevation="1544" latitude="-18.182667" location="Spec. Res." longLatPrecision="1" longitude="47.288807" pageId="1" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD5FFCCFC7B8F63FC067E40" box="[941,987,1076,1103]" pageId="1" pageNumber="128" type="male">1 ♂</specimenCount>
: “ABT/ 
<date id="FFF31060FFD5FFCCFBE28F63FB7E7E41" box="[1076,1187,1076,1102]" pageId="1" pageNumber="128" value="2011-09">
<collectingDate id="EFB7E988FFD5FFCCFBE28F63FB7E7E41" box="[1076,1187,1076,1102]" pageId="1" pageNumber="128" value="2011-09">Sept.2011</collectingDate>
</date>
/14 
<collectingCountry id="F35A7630FFD5FFCCFB188F63FA457E41" box="[1230,1432,1076,1102]" name="Madagascar" pageId="1" pageNumber="128">MADAGASCAR</collectingCountry>
, AMBOHITANTELY 
<location id="8E92607BFFD5FFCCFE5D8F0FFDD87E7C" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A:8E92607BFFD5FFCCFE5D8F0FFDD87E7C" box="[395,517,1112,1139]" country="Madagascar" latitude="-18.182667" longLatPrecision="1" longitude="47.288807" name="Spec. Res." pageId="1" pageNumber="128">Spec. Res.</location>
; 
<date id="FFF31060FFD5FFCCFDC78F0FFDA07E7D" box="[529,637,1112,1138]" pageId="1" pageNumber="128" value="2011-09-03">
<collectingDate id="EFB7E988FFD5FFCCFDC78F0FFDA07E7D" box="[529,637,1112,1138]" pageId="1" pageNumber="128" value="2011-09-03">3.ix.2011</collectingDate>
</date>
; 
<geoCoordinate id="EE795067FFD5FFCCFD5F8F0FFCFA7E7C" box="[649,807,1112,1139]" degrees="18" direction="south" minutes="10" orientation="latitude" pageId="1" pageNumber="128" precision="1" seconds="57.6" value="-18.182667">S18°10’57.6‘‘</geoCoordinate>
<geoCoordinate id="EE795067FFD5FFCCFCF88F0FFC0D7E7D" box="[814,976,1112,1138]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="128" precision="1" seconds="19.7" value="47.288807">E47°17’19.7‘‘</geoCoordinate>
, 
<quantity id="4CB59B45FFD5FFCCFC0A8F0FFBF07E7C" box="[988,1069,1112,1139]" metricMagnitude="3" metricUnit="m" metricValue="1.544" pageId="1" pageNumber="128" unit="m" value="1544.0">
<elevation id="0060D193FFD5FFCCFC0A8F0FFBF07E7C" box="[988,1069,1112,1139]" metricMagnitude="3" metricUnit="m" metricValue="1.544" pageId="1" pageNumber="128" unit="m" value="1544.0">1544m</elevation>
</quantity>
; sifting forest litter and dry fern, Winkler app. extr.; 
<collectorName id="26B85376FFD5FFCCFEA58F2BFD8A7E99" box="[371,599,1148,1174]" pageId="1" pageNumber="128">L.S. Rahanitriniaina</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD5FFCCFD768F2BFD3C7E99" box="[672,737,1148,1174]" pageId="1" pageNumber="128">JJRC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B253CFDFFD5FFCCFD208F2BFD047ED0" ID-GBIF-Occurrence="3131331314" collectionCode="MMBC" collectorName="L. S. Rahanitriniaina" country="Madagascar" elevation="1577" latitude="-18.184027" location="Ambohitantely Spec. Res." longLatPrecision="1" longitude="47.286915" pageId="1" pageNumber="128" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD5FFCCFD208F2BFCF97E98" box="[758,804,1148,1175]" pageId="1" pageNumber="128" type="male">1 ♂</specimenCount>
: “ABT/06/2011 
<collectingCountry id="F35A7630FFD5FFCCFC368F2BFB7A7E99" box="[992,1191,1148,1174]" name="Madagascar" pageId="1" pageNumber="128">MADAGASCAR</collectingCountry>
2011, 
<location id="8E92607BFFD5FFCCFB228F2BFECE7EB4" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A:8E92607BFFD5FFCCFB228F2BFECE7EB4" country="Madagascar" latitude="-18.184027" longLatPrecision="1" longitude="47.286915" name="Ambohitantely Spec. Res." pageId="1" pageNumber="128">Ambohitantely Spec. Res.</location>
; 19.iv., 
<geoCoordinate id="EE795067FFD5FFCCFEB98FF7FDD37EB4" box="[367,526,1184,1211]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="1" pageNumber="128" precision="1" seconds="02.5" value="-18.184027">S18°11’02.5‘‘</geoCoordinate>
, 
<geoCoordinate id="EE795067FFD5FFCCFDCA8FF7FD627EB5" box="[540,703,1184,1210]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="128" precision="1" seconds="12.9" value="47.286915">E47°17’12.9‘‘</geoCoordinate>
; 
<quantity id="4CB59B45FFD5FFCCFD1B8FF7FCC37EB4" box="[717,798,1184,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.577" pageId="1" pageNumber="128" unit="m" value="1577.0">
<elevation id="0060D193FFD5FFCCFD1B8FF7FCC37EB4" box="[717,798,1184,1211]" metricMagnitude="3" metricUnit="m" metricValue="1.577" pageId="1" pageNumber="128" unit="m" value="1577.0">1577m</elevation>
</quantity>
, sifting forest litter, Winkler apparatus extraction; 
<collectorName id="26B85376FFD5FFCCFABB8FF7FE997ED1" pageId="1" pageNumber="128">L.S. Rahanitriniaina</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD5FFCCFE408F93FE2C7ED1" box="[406,497,1220,1246]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="128">MMBC</collectionCode>
, aedeagus missing)
</materialsCitation>
, 
<materialsCitation id="3B253CFDFFD5FFCCFD3C8F93FCE07F29" ID-GBIF-Occurrence="3131331323" collectingDate="2011-09" collectingDateMax="2011-09-05" collectingDateMin="2011-09" collectionCode="MMBC" collectorName="L. S. Rahanitriniaina" country="Madagascar" elevation="1635" latitude="-18.189749" location="Ambohitantely Spec. Res." longLatPrecision="1" longitude="47.293915" pageId="1" pageNumber="128" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD5FFCCFD3C8F93FCC17ED0" box="[746,796,1220,1247]" pageId="1" pageNumber="128" type="female">1 ♀</specimenCount>
: “ABT/ 
<date id="FFF31060FFD5FFCCFCAD8F93FC367ED1" box="[891,1003,1220,1246]" pageId="1" pageNumber="128" value="2011-09">
<collectingDate id="EFB7E988FFD5FFCCFCAD8F93FC367ED1" box="[891,1003,1220,1246]" pageId="1" pageNumber="128" value="2011-09">Sept.2011</collectingDate>
</date>
/21 
<collectingCountry id="F35A7630FFD5FFCCFBCD8F93FB387ED1" box="[1051,1253,1220,1246]" name="Madagascar" pageId="1" pageNumber="128">MADAGASCAR</collectingCountry>
, 
<location id="8E92607BFFD5FFCCFB228F93FECF7F0C" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A:8E92607BFFD5FFCCFB228F93FECF7F0C" country="Madagascar" latitude="-18.189749" longLatPrecision="1" longitude="47.293915" name="Ambohitantely Spec. Res." pageId="1" pageNumber="128">Ambohitantely Spec. Res.</location>
; 
<date id="FFF31060FFD5FFCCFEF68FBFFE517F0C" box="[288,396,1256,1283]" pageId="1" pageNumber="128" value="2011-09-05">
<collectingDate id="EFB7E988FFD5FFCCFEF68FBFFE517F0C" box="[288,396,1256,1283]" pageId="1" pageNumber="128" value="2011-09-05">5.ix.2011</collectingDate>
</date>
; 
<geoCoordinate id="EE795067FFD5FFCCFE4C8FBFFDEA7F0D" box="[410,567,1256,1282]" degrees="18" direction="south" minutes="11" orientation="latitude" pageId="1" pageNumber="128" precision="1" seconds="23.1" value="-18.189749">S18°11’23.1‘‘</geoCoordinate>
<geoCoordinate id="EE795067FFD5FFCCFD978FBFFD3F7F0D" box="[577,738,1256,1282]" degrees="47" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="128" precision="1" seconds="38.1" value="47.293915">E47°17’38.1‘‘</geoCoordinate>
, 
<quantity id="4CB59B45FFD5FFCCFD268FBFFC9F7F0C" box="[752,834,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.635" pageId="1" pageNumber="128" unit="m" value="1635.0">
<elevation id="0060D193FFD5FFCCFD268FBFFC9F7F0C" box="[752,834,1256,1283]" metricMagnitude="3" metricUnit="m" metricValue="1.635" pageId="1" pageNumber="128" unit="m" value="1635.0">1635m</elevation>
</quantity>
; sifting of forest litter and dry fern; secodary forest; Winkler app. extraction; 
<collectorName id="26B85376FFD5FFCCFE788E5BFD4C7F29" box="[430,657,1292,1318]" pageId="1" pageNumber="128">L.S. Rahanitriniaina</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD5FFCCFD0E8E5BFCE87F29" box="[728,821,1292,1318]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="1" pageNumber="128">MMBC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B253CFDFFD5FFCCFC908E5BFCBC7F45" ID-GBIF-Occurrence="3131331344" country="Madagascar" determinerName="Janak &amp; Banar" location="All" pageId="1" pageNumber="128" specimenCount="2" typeStatus="paratype">
<location id="8E92607BFFD5FFCCFC908E5BFCB77F29" LSID="urn:lsid:plazi:treatment:03E487B6FFD5FFCEFF41886FFC4F791A:8E92607BFFD5FFCCFC908E5BFCB77F29" box="[838,874,1292,1318]" country="Madagascar" name="All" pageId="1" pageNumber="128">All</location>
<typeStatus id="54F68802FFD5FFCCFCA78E5AFC017F28" box="[881,988,1293,1319]" pageId="1" pageNumber="128" type="paratype">paratypes</typeStatus>
with an additional label “ 
<typeStatus id="54F68802FFD5FFCCFB2D8E5BFA417F29" box="[1275,1436,1292,1318]" pageId="1" pageNumber="128" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C4D4D23FFD5FFCCFF418E67FE0D7F45" authority="Janák &amp; Cz-, 2021" authorityName="Janák &amp; Cz-" authorityYear="2021" box="[151,464,1328,1354]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">Astenus ambohitantelyensis</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD5FFCCFE0E8E66FDEC7F44" box="[472,561,1329,1355]" pageId="1" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
, 
<determinerName id="7E2EDA64FFD5FFCCFDEA8E67FDA17F45" box="[572,636,1328,1354]" pageId="1" pageNumber="128">Janák</determinerName>
&amp; 
<determinerName id="7E2EDA64FFD5FFCCFD498E67FD3F7F45" box="[671,738,1328,1354]" pageId="1" pageNumber="128">Baňař</determinerName>
det. 2020”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD5FFCEFF118E03FD2E7B4E" lastPageId="3" lastPageNumber="130" pageId="1" pageNumber="128" type="description">
<paragraph id="8BF236A0FFD5FFCCFF118E03FE047FD5" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
<emphasis id="B939EAB2FFD5FFCCFF118E03FE8C7F61" bold="true" box="[199,337,1364,1390]" pageId="1" pageNumber="128">Description</emphasis>
: Body length 
<quantity id="4CB59B45FFD5FFCCFE208E03FDA27F60" box="[502,639,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.4" metricValueMin="4.7" pageId="1" pageNumber="128" unit="mm" value="5.55" valueMax="6.4" valueMin="4.7">4.7–6.4 mm</quantity>
(M = 
<quantity id="4CB59B45FFD5FFCCFD1D8E03FCF87F60" box="[715,805,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="1" pageNumber="128" unit="mm" value="5.4">5.4 mm</quantity>
, HT = 
<quantity id="4CB59B45FFD5FFCCFCAE8E03FC097F60" box="[888,980,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="128" unit="mm" value="4.7">4.7 mm</quantity>
), forebody length 
<quantity id="4CB59B45FFD5FFCCFB738E03FAF07F60" box="[1189,1325,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="1" pageNumber="128" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 mm</quantity>
(M = 
<quantity id="4CB59B45FFD5FFCCFAAF8E03FF187F9C" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="128" unit="mm" value="2.6">2.6 mm</quantity>
, HT = 
<quantity id="4CB59B45FFD5FFCCFEC48E2FFEB17F9C" box="[274,364,1400,1427]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="128" unit="mm" value="2.4">2.4 mm</quantity>
). Black to dark brown, anterior part of head and abdomen dark brown, femora and tibiae brown, knees and tarsi light reddish yellow, antennomeres 1–7 dark reddish brown, antennomeres 8–11 and mouthparts light reddish yellow (
<figureCitation id="13762A25FFD5FFCCFE518E97FE157FD5" box="[391,456,1472,1498]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BF236A0FFD5FFCCFF118EB3FC797C81" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
Head (
<figureCitation id="13762A25FFD5FFCCFEC68EB3FE8F7FF1" box="[272,338,1508,1534]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Fig. 2</figureCitation>
) moderately convex, a fifth longer than wide (L/W = 1.17–1.22, M = 1.20, HT = 1.20). Surface with coarse and dense umbilicate punctures. Eyes moderately large, temples on average about 1.2 times as long as eyes (TL/EL 1.16–1.30, M = 1.20, HT = 1.18). Temples moderately narrowed posteriorly. Antennae moderately short, fifth antennomere about 2.5 times longer than wide (L/W = 2.50–2.73, M = 2.63, HT = 2.50), tenth antennomere about twice as long as wide (L/W = 1.93–2.11, M = 2.02, HT = 2.08).
</paragraph>
<paragraph id="8BF236A0FFD5FFCCFF118DCEFEC87D11" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
Pronotum (
<figureCitation id="13762A25FFD5FFCCFE928DCFFE5A7CBD" box="[324,391,1688,1714]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Fig. 2</figureCitation>
) egg-shaped, moderately convex, a quarter longer than wide (L/W = 1.19–1.26, M = 1.24, HT = 1.25) and moderately narrower than head (PW/HW = 0.87–0.89, M = 0.88, HT = 0.87). Sides markedly narrowed posteriorly. Surface, except for very narrow unpunctured midline with coarse and dense umbilicate punctures similar as on head.
</paragraph>
<paragraph id="8BF236A0FFD5FFCCFF118C7FFB037DA1" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
Elytra (
<figureCitation id="13762A25FFD5FFCCFECB8C7FFE547D4D" box="[285,393,1832,1858]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Figs. 2, 3</figureCitation>
) elongate with rounded sides, concave along suture, about 1.3 times longer than wide (R = 1.26–1.34, M = 1.30, HT = 1.30). Surface moderately coarsely and densely punctured. Interstices shining, without microscupture. Shoulders with 2 tubercules bearing long, protruding fragile spines (
<figureCitation id="13762A25FFD5FFCCFBB28C27FB747D85" box="[1124,1193,1904,1930]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Fig. 3</figureCitation>
) [in most specimens including HT broken and missing]. Sides with additional a few shorter and less protruding spines.
</paragraph>
<paragraph id="8BF236A0FFD5FFCCFF118CEFFE9A7DF9" blockId="1.[151,1437,932,2038]" pageId="1" pageNumber="128">
Abdomen (
<figureCitation id="13762A25FFD5FFCCFE908CEFFE557DDD" box="[326,392,1976,2002]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="1" pageNumber="128">Fig. 1</figureCitation>
) slightly widened to segment VI, finely and densely punctured. Interstices shining, without microsculpture.
</paragraph>
<caption id="DF326628FFD6FFCFFF418C22FEE67DF6" ID-DOI="http://doi.org/10.5281/zenodo.4883129" ID-Zenodo-Dep="4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="2" pageNumber="129" startId="2.[151,264,1909,1933]" targetBox="[181,1409,192,1878]" targetPageId="2">
<paragraph id="8BF236A0FFD6FFCFFF418C22FEE67DF6" blockId="2.[151,1437,1909,2041]" pageId="2" pageNumber="129">
<emphasis id="B939EAB2FFD6FFCFFF418C22FE9C7D82" bold="true" box="[151,321,1909,1933]" pageId="2" pageNumber="129">FIGURES 1–11</emphasis>
. 
<taxonomicName id="4C4D4D23FFD6FFCFFE988C22FCDB7D82" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[334,774,1909,1933]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">
<emphasis id="B939EAB2FFD6FFCFFE988C22FDBA7D82" box="[334,615,1909,1933]" italics="true" pageId="2" pageNumber="129">Astenus ambohitantelyensis</emphasis>
Janák &amp; Baňař
</taxonomicName>
<emphasis id="B939EAB2FFD6FFCFFCDB8C22FC827D82" bold="true" box="[781,863,1909,1933]" pageId="2" pageNumber="129">
<taxonomicNameLabel id="A20A57C9FFD6FFCFFCDB8C22FC827D82" box="[781,863,1909,1933]" pageId="2" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9).
</paragraph>
</caption>
<paragraph id="8BF236A0FFD7FFCEFF118BC0FAAC7B12" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="130">
Male. Posterior margin of sternite VII very slightly, almost invisible emarginate and with almost 40 rod shaped moderately long spines (
<figureCitation id="13762A25FFD7FFCEFE728BECFE297ADA" box="[420,500,187,213]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="3" pageNumber="130">Fig. 10</figureCitation>
), sternite VIII long and narrow, with triangular emargination in posterior quarter (
<figureCitation id="13762A25FFD7FFCEFAB98BECFF697AF6" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="3" pageNumber="130">Fig. 11</figureCitation>
). Aedeagus (
<figureCitation id="13762A25FFD7FFCEFE928B88FE737AF6" box="[324,430,223,249]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="3" pageNumber="130">Figs. 4–9</figureCitation>
) 
<quantity id="4CB59B45FFD7FFCEFE6A8B88FDFD7AF6" box="[444,544,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="3" pageNumber="130" unit="mm" value="0.77">0.77 mm</quantity>
long. Median lobe in ventral view moderately wide, sides slightly emarginate in middle part and with small narrow sub-rounded point. Median lobe in lateral view with distinct subapical dent.
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118A70FD2E7B4E" blockId="3.[151,1437,151,789]" box="[199,755,295,321]" pageId="3" pageNumber="130">Female. Sternite VIII largely rounded posteriorly.</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFCEFF118A1DFC81786D" pageId="3" pageNumber="130" type="diagnosis">
<paragraph id="8BF236A0FFD7FFCEFF118A1DFC81786D" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF118A1DFE0D7B6B" bold="true" box="[199,464,330,356]" pageId="3" pageNumber="130">Differential diagnosis.</emphasis>
The new species is to be arranged in the last key of Madagascan 
<taxonomicName id="4C4D4D23FFD7FFCEFB058A1CFEF57B86" authority="(Janak &amp; Lecoq 2007)" baseAuthorityName="Janak &amp; Lecoq" baseAuthorityYear="2007" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAB2FFD7FFCEFB058A1CFAF77B6A" box="[1235,1322,331,357]" italics="true" pageId="3" pageNumber="130">Astenus</emphasis>
(
<bibRefCitation id="EFDC4B51FFD7FFCEFAED8A1CFEFC7B86" author="Janak, J. &amp; Lecoq, J. - C." pageId="3" pageNumber="130" pagination="299 - 330" refId="ref7827" refString="Janak, J. &amp; Lecoq, J. - C. (2007) Treize nouvelles especes et nouvelles recoltes du genre Astenus Dejean, 1833 de Madagascar et des iles voisines (Coleoptera, Staphylinidae, Paederinae). Bulletin de la Societe entomologique de France, 112 (3), 299 - 330." type="journal article" year="2007">Janák &amp; Lecoq 2007</bibRefCitation>
)
</taxonomicName>
in the couplet 12b near 
<taxonomicName id="4C4D4D23FFD7FFCEFD908A38FBD47B86" authority="Janak &amp; Lecoq, 2007" authorityName="Janak &amp; Lecoq" authorityYear="2007" box="[582,1033,367,393]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="ambondrombe">
<emphasis id="B939EAB2FFD7FFCEFD908A38FCD47B86" box="[582,777,367,393]" italics="true" pageId="3" pageNumber="130">A. ambondrombe</emphasis>
<bibRefCitation id="EFDC4B51FFD7FFCEFCC48A38FBD47B86" author="Janak, J. &amp; Lecoq, J. - C." box="[786,1033,367,393]" pageId="3" pageNumber="130" pagination="299 - 330" refId="ref7827" refString="Janak, J. &amp; Lecoq, J. - C. (2007) Treize nouvelles especes et nouvelles recoltes du genre Astenus Dejean, 1833 de Madagascar et des iles voisines (Coleoptera, Staphylinidae, Paederinae). Bulletin de la Societe entomologique de France, 112 (3), 299 - 330." type="journal article" year="2007">Janák &amp; Lecoq, 2007</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C4D4D23FFD7FFCEFB938A38FAB77B86" authority="Janak, 2008" authorityName="Janak" authorityYear="2008" box="[1093,1386,367,393]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="ambrensis">
<emphasis id="B939EAB2FFD7FFCEFB938A38FB057B86" box="[1093,1240,367,393]" italics="true" pageId="3" pageNumber="130">A. ambrensis</emphasis>
<bibRefCitation id="EFDC4B51FFD7FFCEFB348A38FAB77B86" author="Janak, J." box="[1250,1386,367,393]" pageId="3" pageNumber="130" pagination="125 - 134" refId="ref7885" refString="Janak, J. (2008) Deux nouvelles especes du genre Astenus Dejean et de nouvelles recoltes des genres Astenus Dejean et Dibelonetes Sahlberg de Madagascar et des iles Sechelles (Coleoptera: Staphylinidae: Paederinae). Studies and reports of District Museum Prague East, Taxonomical Series, 4 (1 - 2), 125 - 134." type="journal article" year="2008">Janák, 2008</bibRefCitation>
</taxonomicName>
and differs from these species by very slightly, almost invisible emarginated posterior margin of the male sternite VII with moderately long rod shaped spines and different shape of aedeagus (see 
<figureCitation id="13762A25FFD7FFCEFBD78AE0FB587BDE" box="[1025,1157,439,465]" captionStart-0="FIGURES 31–39" captionStart-1="FIGURES 40–41" captionStart-2="FIGURES 42–43" captionStart-3="FIGURES 44–45" captionStartId-0="8.[151,264,1908,1932]" captionStartId-1="9.[151,264,1943,1967]" captionStartId-2="10.[151,264,1943,1967]" captionStartId-3="11.[151,264,1908,1932]" captionTargetBox-0="[169,1418,316,1871]" captionTargetBox-1="[171,1416,181,1920]" captionTargetBox-2="[173,1415,200,1920]" captionTargetBox-3="[154,1434,181,1884]" captionTargetId-0="figure-73@8.[158,1430,309,1885]" captionTargetId-1="figure-17@9.[171,1416,181,1920]" captionTargetId-2="figure-20@10.[173,1415,181,1920]" captionTargetId-3="figure-17@11.[154,1434,181,1884]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURES 31–39. Astenus duflosi Lecoq. 31, 32, 36–39, Vohimana; 33–35, Andasibé; 31, habitus; 32, forebody; 33, 35, aedeagus, lateral view; 34, aedeagus, ventral view; 36, 37, apical part of aedeagus, ventral view; 38, posterior part of male sternite VII; 39, male sternite VIII; 33–35, reflected light; 36, 37, transmitted light. Scale bar: 1.0 mm (Fig. 31), 0.5 mm (Figs. 33–35, 38–39), 0.1 mm (Figs. 36 –37)." captionText-1="FIGURES 40–41. Forest of Ambohitantely Special Reserve. 41, View from administrative buildings close to main entrance; 42, detail of forest undergrowth." captionText-2="FIGURES 42–43. Forest of Anjozorobe-Angavo. 42, View from camping site; 43, forest detail (both photos by M. Trýzna)." captionText-3="FIGURES 44–45. Forest of Anjozorobe-Angavo, microhabitats of Astenus grebennikovi Janák &amp; Baňař sp. nov. 42, detail of forest undergrowth; 43, detail of microhabitat of sift sample nr. 4 under Pandanus plant." figureDoi-0="http://doi.org/10.5281/zenodo.4945834" figureDoi-1="http://doi.org/10.5281/zenodo.4883137" figureDoi-2="http://doi.org/10.5281/zenodo.4883141" figureDoi-3="http://doi.org/10.5281/zenodo.4883145" httpUri-0="https://zenodo.org/record/4945834/files/figure.png" httpUri-1="https://zenodo.org/record/4883137/files/figure.png" httpUri-2="https://zenodo.org/record/4883141/files/figure.png" httpUri-3="https://zenodo.org/record/4883145/files/figure.png" pageId="3" pageNumber="130">
Figs. 
<quantity id="4CB59B45FFD7FFCEFB968AE0FB587BDE" box="[1088,1157,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.1176" metricValueMin="0.9652" pageId="3" pageNumber="130" unit="in" value="41.0" valueMax="44.0" valueMin="38.0">38–44</quantity>
</figureCitation>
in 
<bibRefCitation id="EFDC4B51FFD7FFCEFB7D8AE0FA417BDE" author="Janak, J. &amp; Lecoq, J. - C." box="[1195,1436,439,465]" pageId="3" pageNumber="130" pagination="299 - 330" refId="ref7827" refString="Janak, J. &amp; Lecoq, J. - C. (2007) Treize nouvelles especes et nouvelles recoltes du genre Astenus Dejean, 1833 de Madagascar et des iles voisines (Coleoptera, Staphylinidae, Paederinae). Bulletin de la Societe entomologique de France, 112 (3), 299 - 330." type="journal article" year="2007">Janák &amp; Lecoq, 2007</bibRefCitation>
and 
<figureCitation id="13762A25FFD7FFCEFF1C8A8CFEE87BFA" box="[202,309,475,501]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="3" pageNumber="130">
Figs. 
<quantity id="4CB59B45FFD7FFCEFEDA8A8CFEE87BFA" box="[268,309,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.524" metricValueMin="0.508" pageId="3" pageNumber="130" unit="in" value="4.0" valueMax="6.0" valueMin="2.0">2–6</quantity>
</figureCitation>
in 
<bibRefCitation id="EFDC4B51FFD7FFCEFEB68A8CFE337BFA" author="Janak, J." box="[352,494,475,501]" pageId="3" pageNumber="130" pagination="125 - 134" refId="ref7885" refString="Janak, J. (2008) Deux nouvelles especes du genre Astenus Dejean et de nouvelles recoltes des genres Astenus Dejean et Dibelonetes Sahlberg de Madagascar et des iles Sechelles (Coleoptera: Staphylinidae: Paederinae). Studies and reports of District Museum Prague East, Taxonomical Series, 4 (1 - 2), 125 - 134." type="journal article" year="2008">Janák, 2008</bibRefCitation>
). There are also some external differences among these species – the color of 
<taxonomicName id="4C4D4D23FFD7FFCEFA528A8CFDC07816" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">
<emphasis id="B939EAB2FFD7FFCEFA528A8CFEB27816" italics="true" pageId="3" pageNumber="130">A. ambohitantelyensis</emphasis>
Janák &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD7FFCEFDF28AA8FDA67816" box="[548,635,511,537]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
is much darker and the pronotum and elytra are on average longer than in remaining two species, but these characters are quite variable (see 
<figureCitation id="13762A25FFD7FFCEFC4D8974FC367832" box="[923,1003,547,573]" captionStart="FIGURES 31–39" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[169,1418,316,1871]" captionTargetId="figure-73@8.[158,1430,309,1885]" captionTargetPageId="8" captionText="FIGURES 31–39. Astenus duflosi Lecoq. 31, 32, 36–39, Vohimana; 33–35, Andasibé; 31, habitus; 32, forebody; 33, 35, aedeagus, lateral view; 34, aedeagus, ventral view; 36, 37, apical part of aedeagus, ventral view; 38, posterior part of male sternite VII; 39, male sternite VIII; 33–35, reflected light; 36, 37, transmitted light. Scale bar: 1.0 mm (Fig. 31), 0.5 mm (Figs. 33–35, 38–39), 0.1 mm (Figs. 36 –37)." figureDoi="http://doi.org/10.5281/zenodo.4945834" httpUri="https://zenodo.org/record/4945834/files/figure.png" pageId="3" pageNumber="130">
Fig. 
<quantity id="4CB59B45FFD7FFCEFC198974FC367832" box="[975,1003,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="3" pageNumber="130" unit="in" value="39.0">39</quantity>
</figureCitation>
in 
<bibRefCitation id="EFDC4B51FFD7FFCEFBD88974FAD47832" author="Janak, J. &amp; Lecoq, J. - C." box="[1038,1289,547,573]" pageId="3" pageNumber="130" pagination="299 - 330" refId="ref7827" refString="Janak, J. &amp; Lecoq, J. - C. (2007) Treize nouvelles especes et nouvelles recoltes du genre Astenus Dejean, 1833 de Madagascar et des iles voisines (Coleoptera, Staphylinidae, Paederinae). Bulletin de la Societe entomologique de France, 112 (3), 299 - 330." type="journal article" year="2007">Janák &amp; Lecoq (2007)</bibRefCitation>
and 
<figureCitation id="13762A25FFD7FFCEFAE88974FA5D7832" box="[1342,1408,547,573]" captionStart="FIGURES 1–11" captionStartId="2.[151,264,1909,1933]" captionTargetBox="[181,1409,192,1878]" captionTargetId="figure-20@2.[171,1417,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–11. Astenus ambohitantelyensis Janák &amp; Baňař sp. nov. 1, 2, 4, 5, 8–11, holotype; 3, 6, 7, paratypes; 1, habitus; 2, forebody; 3, elytra; 4, 6, 8, aedeagus, lateral view; 5, 7, 9, aedeagus, ventral view; 10, posterior part of male sternite VII; 11, male sternite VIII; 4, 5, 8, 9, transmitted light; 6, 7, reflected light. Scale bar: 1.0 mm (Fig. 1); 0.5 mm (Figs. 2–7, 10–11), 0.1 mm (Figs. 8–9)." figureDoi="http://doi.org/10.5281/zenodo.4883129" httpUri="https://zenodo.org/record/4883129/files/figure.png" pageId="3" pageNumber="130">
Fig. 
<quantity id="4CB59B45FFD7FFCEFAA48974FA5D7832" box="[1394,1408,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="130" unit="in" value="1.0">1</quantity>
</figureCitation>
in 
<bibRefCitation id="EFDC4B51FFD7FFCEFF418910FEF4786E" author="Janak, J." box="[151,297,583,609]" pageId="3" pageNumber="130" pagination="125 - 134" refId="ref7885" refString="Janak, J. (2008) Deux nouvelles especes du genre Astenus Dejean et de nouvelles recoltes des genres Astenus Dejean et Dibelonetes Sahlberg de Madagascar et des iles Sechelles (Coleoptera: Staphylinidae: Paederinae). Studies and reports of District Museum Prague East, Taxonomical Series, 4 (1 - 2), 125 - 134." type="journal article" year="2008">Janák (2008)</bibRefCitation>
and measurements in the descriptions for details).
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFCEFF11893CFB30788A" box="[199,1261,619,645]" pageId="3" pageNumber="130" type="etymology">
<paragraph id="8BF236A0FFD7FFCEFF11893CFB30788A" blockId="3.[151,1437,151,789]" box="[199,1261,619,645]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF11893CFE93788A" bold="true" box="[199,334,619,645]" pageId="3" pageNumber="130">Etymology.</emphasis>
This species is named the after the 
<typeStatus id="54F68802FFD7FFCEFD0B893CFCD0788A" box="[733,781,619,645]" pageId="3" pageNumber="130">type</typeStatus>
locality – Ambohitantely Special Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFCEFF1189D9FC4F791A" pageId="3" pageNumber="130" type="distribution">
<paragraph id="8BF236A0FFD7FFCEFF1189D9FC0578C2" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF1189D9FEBC78A7" bold="true" box="[199,353,654,680]" pageId="3" pageNumber="130">Distribution.</emphasis>
<taxonomicName id="4C4D4D23FFD7FFCEFEB989D8FCAF78A6" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[367,882,655,681]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="ambohitantelyensis" status="sp. nov.">
<emphasis id="B939EAB2FFD7FFCEFEB989D8FD7078A6" box="[367,685,655,681]" italics="true" pageId="3" pageNumber="130">Astenus ambohitantelyensis</emphasis>
Janák &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD7FFCEFC5789D8FC3D78A6" box="[897,992,655,681]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
is currently recorded only from the Ambohitantely Special Reserve in the northern central part of 
<collectingCountry id="F35A7630FFD7FFCEFC8789E4FC0578C2" box="[849,984,691,717]" name="Madagascar" pageId="3" pageNumber="130">Madagascar</collectingCountry>
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118981FC4F791A" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF118981FE9678FF" bold="true" box="[199,331,726,752]" pageId="3" pageNumber="130">Bionomics.</emphasis>
All specimens were collected in siftings of forest litter, palm trees and dry ferns in an indigenous or secondary forests at an altitude of 
<quantity id="4CB59B45FFD7FFCEFDC189ADFD6F791A" box="[535,690,762,789]" metricMagnitude="3" metricUnit="m" metricValue="1.59" metricValueMax="1.64" metricValueMin="1.54" pageId="3" pageNumber="130" unit="m" value="1590.0" valueMax="1640.0" valueMin="1540.0">1540–1640 m</quantity>
a.s.l. (
<figureCitation id="13762A25FFD7FFCEFD2A89ACFC58791A" box="[764,901,763,789]" captionStart="FIGURES 40–41" captionStartId="9.[151,264,1943,1967]" captionTargetBox="[171,1416,181,1920]" captionTargetId="figure-17@9.[171,1416,181,1920]" captionTargetPageId="9" captionText="FIGURES 40–41. Forest of Ambohitantely Special Reserve. 41, View from administrative buildings close to main entrance; 42, detail of forest undergrowth." figureDoi="http://doi.org/10.5281/zenodo.4883137" httpUri="https://zenodo.org/record/4883137/files/figure.png" pageId="3" pageNumber="130">Figs. 40, 41</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>