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<mods:title id="65F9524CFEA257B57781B80B3791C985">The squat lobster genus Phylladiorhynchus Baba, 1969 in New Zealand and eastern Australia, with description of six new species</mods:title>
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<emphasis id="B939EA817352FFC2FF25B088FE13F979" bold="true" box="[151,468,1695,1722]" italics="true" pageId="10" pageNumber="311">Phylladiorhynchus erebus</emphasis>
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(
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,
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)
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<emphasis id="B939EA817352FFC2FF25B11BFE8EF8E5" bold="true" box="[151,329,1803,1830]" pageId="10" pageNumber="311">Type material.</emphasis>
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Stn.
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.
<materialsCitation id="3B253CCE7352FFC2FE5EB1D7FC34F81A" ID-GBIF-Occurrence="2429072256" box="[492,1011,1983,2010]" collectionCode="NIWA" country="Norfolk Island" location="Norfolk Ridge" pageId="10" pageNumber="311" specimenCode="NIWA 23099" specimenCount="1" specimenCount-male="1">
<specimenCode id="DBEB9EE87352FFC2FE5EB1D7FD45F819" box="[492,642,1984,2010]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="10" pageNumber="311" type="Other institutional type">NIWA 23099</specimenCode>
,
<specimenCount id="9D4BFD1A7352FFC2FD3EB1D7FD12F81A" box="[652,725,1983,2010]" pageId="10" pageNumber="311" type="male">1 male</specimenCount>
(
<quantity id="4CB59B767352FFC2FD51B1A8FCFAF819" box="[739,829,1983,2010]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="311" unit="mm" value="2.6">2.6 mm</quantity>
),
<location id="8E9260487352FFC2FCFDB1A8FC34F81A" LSID="urn:lsid:plazi:treatment:03E487857352FFC6FF25B088FCE9FF14:8E9260487352FFC2FCFDB1A8FC34F81A" box="[847,1011,1983,2009]" country="Norfolk Island" name="Norfolk Ridge" pageId="10" pageNumber="311">Norfolk Ridge</location>
</materialsCitation>
,
<materialsCitation id="3B253CCE7352FFC2FC4FB1D7FEB5F83D" ID-GBIF-Occurrence="2429072320" collectingDate="1975-07-24" collectingDateMax="1994-01" collectingDateMin="1975-07-24" collectionCode="NZOI" collectorName="Stn. I" country="Norfolk Island" elevation="308" latitude="-29.337" longLatPrecision="73" longitude="168.18" pageId="10" pageNumber="311" specimenCount="1">
<collectionCode id="ED5CAE567352FFC2FC4FB1D7FB87F819" box="[1021,1088,1984,2010]" collectionName="NZOI" country="New Zealand" httpUri="http://grbio.org/cool/chf3-57df" name="New Zealand Oceanographic Institute" pageId="10" pageNumber="311">NZOI</collectionCode>
<collectorName id="26B853457352FFC2FBF7B1D7FB46F819" box="[1093,1153,1984,2010]" pageId="10" pageNumber="311">
Stn.
<date id="FFF310537352FFC2FBC5B1D7FB46F819" box="[1143,1153,1984,2010]" pageId="10" pageNumber="311" value="1994-01">I</date>
</collectorName>
94,
<geoCoordinate id="EE7950547352FFC2FB1BB1D7FAD3F819" box="[1193,1300,1984,2010]" degrees="29.337" direction="south" orientation="latitude" pageId="10" pageNumber="311" precision="55" value="-29.337">29.337°S</geoCoordinate>
,
<geoCoordinate id="EE7950547352FFC2FAACB1A8FA5FF81A" box="[1310,1432,1983,2009]" degrees="168.180" direction="east" orientation="longitude" pageId="10" pageNumber="311" precision="55" value="168.18">168.180°E</geoCoordinate>
,
<quantity id="4CB59B767352FFC2FF25B1F3FF27F83D" box="[151,224,2020,2046]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="10" pageNumber="311" unit="m" value="308.0">
<elevation id="0060D1A07352FFC2FF25B1F3FF27F83D" box="[151,224,2020,2046]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="10" pageNumber="311" unit="m" value="308.0">308 m</elevation>
</quantity>
,
<date id="FFF310537352FFC2FF5EB1F3FEB5F83D" box="[236,370,2019,2046]" pageId="10" pageNumber="311" value="1975-07-24">
<collectingDate id="EFB7E9BB7352FFC2FF5EB1F3FEB5F83D" box="[236,370,2019,2046]" pageId="10" pageNumber="311" value="1975-07-24">24 Jul 1975</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35765187353FFC5FF75B68FFA5CFE6C" lastPageId="13" lastPageNumber="314" pageId="11" pageNumber="312" type="diagnosis">
<paragraph id="8BF236937353FFC3FF75B68FFCE6FE6D" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B68FFE85FF70" bold="true" box="[199,322,152,179]" pageId="11" pageNumber="312">Diagnosis.</emphasis>
Carapace with transverse row of 5 epigastric spines (median and 2 on either side of midline); metagastric ridge medially interrupted; anterior branchial margin with 3 spines. Maxilliped 3 merus with 2 prominent spines on flexor margin. Thoracic sternite 3 anterior margin sinuous, with shallow median concavity. Antennular article 1 with 5 or 6 distal spines: distomesial spine distinctly shorter than upper distolateral spine; first lateral spine about as long as upper distolateral spine, followed laterally by 1 or 2 small spines similar to distomesial spine. Antennal article 1 mesial process distally falling well short of second lateral antennular spine; article 2 distolateral and distomesial spines subequal; article 3 with small to distinct mesial spine, laterally unarmed. P24 dactylus extensor margin without upright spines at bases of movable spines.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B7AEFBA9FC09" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B7AEFF2FFE10" bold="true" box="[199,232,441,467]" pageId="11" pageNumber="312">De</emphasis>
s
<emphasis id="B939EA817353FFC3FF41B7AFFE9EFE10" bold="true" box="[243,345,440,467]" pageId="11" pageNumber="312">cription.</emphasis>
<emphasis id="B939EA817353FFC3FEEDB7AEFE0AFE10" box="[351,461,441,467]" italics="true" pageId="11" pageNumber="312">Carapace</emphasis>
: As long as wide; transverse ridges with dense short setae, and few scattered longer setae. Gastric region with 3 interrupted anterior transverse ridges: epigastric ridge indistinct, with 5 spines (1 median and 2 pairs of spines laterally); anterior protogastric ridge with small median gap or continuous, nearly extending laterally to carapace margin; anterior mesogastric ridge minutely interrupted or continuous, broken laterally by cervical groove, laterally continuing uninterrupted to first branchial spine; metagastric ridge medially broken; sometimes with short scales scattered between ridges. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 2 complete, uninterrupted or minutely interrupted ridges, interspersed with 3 short lateral ridges and few short, scattered scales. Lateral margins weakly convex, with 7 or 8 spines: 2 spines (anterolateral, hepatic) in front of, and 5 or 6 spines (3 anterior branchial spines, 2 or 3 posterior branchial spines) behind distinct anterior cervical groove. Anterolateral spine well-developed, reaching or nearly reaching anteriorly to level of lateral orbital spine, subequal in size. Hepatic spine small, set slightly dorsomesially from lateral margin.Anterior branchial spines subequal in size. Posterior branchial spines progressively smaller posteriorly, first spine subequal to anterior branchial spines. Rostrum 0.5 × pcl, length-width ratio 1.51.8, distance between basal rostral spines about 0.3 × carapace width; dorsal surface very slightly concave; lateral margins straight, sub-apical spines obsolescent. Pterygostomian flap without anterior spine; surface unarmed, with series of short striae.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B5C3FB12FBD1" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B5C3FE48FC2D" box="[199,399,980,1006]" italics="true" pageId="11" pageNumber="312">Thoracic sternum</emphasis>
: Sternal plastron about as wide as long, lateral margins of posterior half slightly divergent. Sternite 3 width 23 × length; anterior margin with small median notch; lateral margins rounded.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B20BFBCFFB99" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B20BFEF6FBF5" box="[199,305,1052,1078]" italics="true" pageId="11" pageNumber="312">Abdomen</emphasis>
: Tergite 2 with anterior and posterior transverse ridges; tergites 3 and 4 with anterior transverse ridge, posterior transverse ridge absent; tergites 56 smooth, without elevated ridges.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B272FD39FB61" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B272FF36FBBD" box="[199,241,1125,1150]" italics="true" pageId="11" pageNumber="312">Eye</emphasis>
: Eye length 1.41.6 × width, peduncle distally setose, not distinctly expanded proximally, with few short transverse striae on lateral surfaces; cornea not dilated.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B2BBFB53FB29" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B2BBFEFEFB05" box="[199,313,1196,1222]" italics="true" pageId="11" pageNumber="312">Antennule</emphasis>
: Article 1 with [5]6 well-developed distal spines: distomesial spine small, pair of distolateral spines present; proximal lateral spine small, always present. Short striae covering mesial surfaces.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B2E2FEACFA95" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B2E2FEE3FACC" box="[199,292,1269,1295]" italics="true" pageId="11" pageNumber="312">Antenna</emphasis>
: Article 1 with prominent mesial process, distally falling well short of lateral antennular spine. Article 2 with distinct distal spines laterally and mesially. Article 3 with small to distinct mesial spine, laterally unarmed. Article 4 unarmed.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B377FBD6FA01" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B377FE6BFAB8" box="[199,428,1376,1403]" italics="true" pageId="11" pageNumber="312">Maxilliped 3 (Mxp3)</emphasis>
: Ischium with distinct distal spines on both flexor and extensor margins; crista dentata with regular row of spines along entire margin. Merus 0.70.8 × length of ischium at midline, with small distal spines on extensor margin and 2 much larger spines at flexor margin, otherwise unarmed.
</paragraph>
<paragraph id="8BF236937353FFC3FF75B3DBFE39F8C4" blockId="11.[151,1437,152,2015]" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B3DBFEEBFA25" box="[199,300,1484,1510]" italics="true" pageId="11" pageNumber="312">Cheliped</emphasis>
: Length about 34 × pcl in both sexes; subcylindrical, spiny and with scattered long setae. Ischiomerus 1.21.7 × pcl, slightly exceeding twice carpus length; with rows of spines, mesial spines strongest. Carpus with rows of spines on all surfaces, row of mesial spines most prominent. Palm 1.01.3 × carpus length, length-width ratio 1.52.4, with parallel rows of spines (2 closely running parallel along mesial, 1 at about dorsal surface midline, 1 close to lateral margin). Dactylus and pollex subequal in length to palm; surface with short setiferous striae; distally with row of spines along curved margin; tip excavated. Pollex lateral margin with 1 or 2 spines reaching onto proximal portion; occlusal margin with 2 proximal processes, distinct in massive chela. Dactylus with 1 small to distinct external spine at its base; occlusal margin with basal process, well developed if chela massive. Central portion of gape evenly concave.
</paragraph>
<paragraph id="8BF236937353FFC5FF75B107FC00FE80" blockId="11.[151,1437,152,2015]" lastBlockId="13.[151,1437,153,1043]" lastPageId="13" lastPageNumber="314" pageId="11" pageNumber="312">
<emphasis id="B939EA817353FFC3FF75B107FE6BF8E8" box="[199,428,1808,1835]" italics="true" pageId="11" pageNumber="312">Walking legs (P24)</emphasis>
: Slender, subcylindrical, moderately setose and spinose. Merus successively shorter posteriorly (P2 merus 1.1 × length of P3 merus, P3 merus 1.1 × length of P4 merus). P2 merus 0.8 × pcl, 1.2 × P2 propodus length; length 56 × width. P3 merus 1.2 × P3 propodus length; length about 5 × width. P4 merus as long as P4 propodus; length 56 × width. Extensor margin of P2 and P3 with regular row of spines, proximally diminishing, with prominent distal spine; P4 extensor margin irregular but unarmed other than distal spine; flexor margins of all legs irregular and with distal spine; P4 lateral surface with median row of 3 or 4 small spines, absent on P2 and P3. Carpus with 3 or 4 spines on extensor margin on P23, unarmed on P4; distal spine prominent on P23, minute on P4; row of small spines below extensor margin on lateral surface of P23, unarmed on P4; flexor margin unarmed. P24 propodus length 67 × height; extensor margin irregular, typically unarmed; flexor margin with 24 slender movable spines in addition to distal pair. Dactylus 0.60.7 × length of propodus, ending in incurved, strong, sharp spine; flexor margin with 69 movable spines, otherwise unarmed.
</paragraph>
<caption id="DF32661B7354FFC4FF25B13FFB80F86F" ID-DOI="http://doi.org/10.5281/zenodo.3516753" ID-Zenodo-Dep="3516753" httpUri="https://zenodo.org/record/3516753/files/figure.png" pageId="12" pageNumber="313" startId="12.[151,250,1832,1857]" targetBox="[219,1327,200,1768]" targetPageId="12">
<paragraph id="8BF236937354FFC4FF25B13FFB80F86F" blockId="12.[151,1437,1832,1965]" pageId="12" pageNumber="313">
<emphasis id="B939EA817354FFC4FF25B13FFED3F882" bold="true" box="[151,276,1832,1857]" pageId="12" pageNumber="313">FIGURE 4.</emphasis>
<taxonomicName id="4C4D4D107354FFC4FEA9B13EFDE3F882" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[283,548,1833,1857]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B939EA817354FFC4FEA9B13EFDE3F882" box="[283,548,1833,1857]" italics="true" pageId="12" pageNumber="313">Phylladiorhynchus erebus</emphasis>
</taxonomicName>
<emphasis id="B939EA817354FFC4FD99B13EFDA2F882" bold="true" box="[555,613,1833,1857]" pageId="12" pageNumber="313">
<taxonomicNameLabel id="A20A57FA7354FFC4FD99B13EFDA2F882" box="[555,613,1833,1857]" pageId="12" pageNumber="313" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male, 3.3 mm (NIWA 21269): A, carapace and abdomen, dorsal view. B, sternal plastron. C, telson. D, right antennal peduncle, ventral view. E, left cephalic region, showing antennular and antennal peduncles, ventral view. F, right Mxp3, lateral view. G, H, left and right cheliped, dorsal view. I, J, left P23, lateral view. K, right P4, lateral view. L, distal portion of propodus and dactylus of left P2. Scale bars: 2 mm.
</paragraph>
</caption>
<paragraph id="8BF236937355FFC5FF75B75BFD19FEA4" blockId="13.[151,1437,153,1043]" box="[199,734,332,359]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B75BFF39FEA4" bold="true" box="[199,254,332,359]" pageId="13" pageNumber="314">Size.</emphasis>
Male pcl
<quantity id="4CB59B767355FFC5FEDDB75AFE3EFEA4" box="[367,505,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="3.3" metricValueMin="2.3" pageId="13" pageNumber="314" unit="mm" value="2.8" valueMax="3.3" valueMin="2.3">2.33.3 mm</quantity>
; female pcl
<quantity id="4CB59B767355FFC5FD33B75AFD1DFEA4" box="[641,730,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="314" unit="mm" value="2.1">2.1 mm</quantity>
.
</paragraph>
<paragraph id="8BF236937355FFC5FF75B767FE2EFE48" blockId="13.[151,1437,153,1043]" box="[199,489,368,395]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B767FEAAFE48" bold="true" box="[199,365,368,395]" pageId="13" pageNumber="314">Colour in life.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF236937355FFC5FF75B783FA5CFE6C" blockId="13.[151,1437,153,1043]" box="[199,1435,404,431]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B783FEA1FE6C" bold="true" box="[199,358,404,431]" pageId="13" pageNumber="314">Genetic data.</emphasis>
A COI sequence was generated from a Kermadec Islands sample (AWMM MA73597,
<tableCitation id="C6CF03287355FFC5FA8AB782FA4DFE6C" box="[1336,1418,405,431]" captionStart="TABLE 1" captionStartId="41.[151,244,511,538]" captionTargetBox="[165,2022,621,1679]" captionTargetId="graphics@41.[151,2038,1112,1687]" captionTargetPageId="41" captionText="TABLE 1. Pairwise percentage genetic distances (below diagonal) and genetic similarity (above diagonal) for the partial COI gene between specimens of Phylladi- orhynchus. Cells highlighted in grey indicate intraspecific levels of divergence. The sequence of P. integrirostris from Hawaii was kindly provided as a reference by P. Rodríguez-Flores (National Museum of Natural Sciences, Madrid)." httpUri="http://table.plazi.org/id/DF32661B7371FFE1FF25B7E8FC4FFDA1" pageId="13" pageNumber="314" tableUuid="DF32661B7371FFE1FF25B7E8FC4FFDA1">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35765187355FFC5FF75B7AEFC94FDD8" pageId="13" pageNumber="314" type="etymology">
<paragraph id="8BF236937355FFC5FF75B7AEFBABFE34" blockId="13.[151,1437,153,1043]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B7AEFE89FE10" bold="true" box="[199,334,441,467]" pageId="13" pageNumber="314">Etymology.</emphasis>
From Erebus, of Greek mythology, a primordial deity personifying deep darkness and shadows, alluding to the deep-water occurrence of the new species. Used as a noun in apposition.
</paragraph>
<paragraph id="8BF236937355FFC5FF75B417FC94FDD8" blockId="13.[151,1437,153,1043]" box="[199,851,512,539]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B417FF24FDD8" bold="true" box="[199,227,512,539]" pageId="13" pageNumber="314">Di</emphasis>
s
<emphasis id="B939EA817355FFC5FF5CB417FEA6FDD8" bold="true" box="[238,353,512,539]" pageId="13" pageNumber="314">tribution.</emphasis>
Norfolk and Kermadec Ridges;
<quantity id="4CB59B767355FFC5FD7FB417FC88FDD8" box="[717,847,512,539]" metricMagnitude="2" metricUnit="m" metricValue="3.9799999999999995" metricValueMax="5.06" metricValueMin="2.9" pageId="13" pageNumber="314" unit="m" value="398.0" valueMax="506.0" valueMin="290.0">290506 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35765187355FFC6FF75B432FCE9FF14" lastPageId="14" lastPageNumber="315" pageId="13" pageNumber="314" type="discussion">
<paragraph id="8BF236937355FFC5FF75B432FC4EFD0C" blockId="13.[151,1437,153,1043]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF75B432FEFCFDFC" bold="true" box="[199,315,549,575]" pageId="13" pageNumber="314">Remarks.</emphasis>
<taxonomicName id="4C4D4D107355FFC5FEF3B432FDA3FDFC" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[321,612,549,575]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B939EA817355FFC5FEF3B432FDA3FDFC" box="[321,612,549,575]" italics="true" pageId="13" pageNumber="314">Phylladiorhynchus erebus</emphasis>
</taxonomicName>
<emphasis id="B939EA817355FFC5FDD8B432FD6FFDFC" bold="true" box="[618,680,549,575]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A20A57FA7355FFC5FDD8B432FD6FFDFC" box="[618,680,549,575]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C4D4D107355FFC5FCEBB432FC5AFDFC" authorityName="" baseAuthorityName="Miyake &amp; Baba" baseAuthorityYear="1965" box="[857,925,549,575]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="B939EA817355FFC5FCEBB432FC5AFDFC" box="[857,925,549,575]" italics="true" pageId="13" pageNumber="314">ikedai</emphasis>
</taxonomicName>
/
<emphasis id="B939EA817355FFC5FC17B432FBF4FDFC" box="[933,1075,549,575]" italics="true" pageId="13" pageNumber="314">bengalensis</emphasis>
group that share the five epigastric spines and the maxilliped 3 merus with two prominent spines along the flexor margin. This now also includes
<taxonomicName id="4C4D4D107355FFC5FA36B45DFEFBFD44" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
<emphasis id="B939EA817355FFC5FA36B45DFA52FDA0" box="[1412,1429,586,611]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FF25B47AFEFBFD44" box="[151,316,621,647]" italics="true" pageId="13" pageNumber="314">kermadecensis</emphasis>
</taxonomicName>
<emphasis id="B939EA817355FFC5FEF0B47AFE47FD44" bold="true" box="[322,384,621,647]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A20A57FA7355FFC5FEF0B47AFE47FD44" box="[322,384,621,647]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, which differs in the presence of two instead of three anterior branchial spines along the lateral carapace margin (see comments under that species below). Ongoing studies by Rodríguez-Flores and colleagues indicate at least three further species will be added to this complex.
</paragraph>
<paragraph id="8BF236937355FFC5FF7FB4CEFAF3FC9C" blockId="13.[151,1437,153,1043]" pageId="13" pageNumber="314">
<taxonomicName id="4C4D4D107355FFC5FF7FB4CEFE36FD30" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[205,497,729,755]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B939EA817355FFC5FF7FB4CEFE36FD30" box="[205,497,729,755]" italics="true" pageId="13" pageNumber="314">Phylladiorhynchus erebus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4D4D107355FFC5FD30B4CDFC32FD30" authority="(Miyake &amp; Baba, 1965)" authorityName="" authorityYear="1965" baseAuthorityName="Miyake &amp; Baba" baseAuthorityYear="1965" box="[642,1013,728,755]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="B939EA817355FFC5FD30B4CDFD54FD30" box="[642,659,730,755]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FD13B4CEFD22FD30" box="[673,741,729,755]" italics="true" pageId="13" pageNumber="314">ikedai</emphasis>
(
<bibRefCitation id="EFDC4B627355FFC5FD41B4CEFC2BFD30" author="Miyake, S. &amp; Baba, K." box="[755,1004,728,755]" pageId="13" pageNumber="314" pagination="585 - 593" refId="ref43483" refString="Miyake, S. &amp; Baba, K. (1965) Some galatheids obtained from the Bonin Islands (Crustacea, Anomura). Journal of the Faculty of Agriculture, Kyushu University, 13, 585 - 593." type="journal article" year="1965">Miyake &amp; Baba, 1965</bibRefCitation>
)
</taxonomicName>
in the presence of 56 distal spines of the basal antennular article, the distolateral double spine is always present, and an additional third lateral spine may also be present;
<taxonomicName id="4C4D4D107355FFC5FEFFB535FE76FCF8" authorityName="" baseAuthorityName="Miyake &amp; Baba" baseAuthorityYear="1965" box="[333,433,801,827]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="B939EA817355FFC5FEFFB535FE99FCF8" box="[333,350,802,827]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FEDFB536FE76FCF8" box="[365,433,801,827]" italics="true" pageId="13" pageNumber="314">ikedai</emphasis>
</taxonomicName>
lacks the distolaterally split double spine. All the specimens examined here for
<taxonomicName id="4C4D4D107355FFC5FA81B535FA5AFCF8" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[1331,1437,801,827]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B939EA817355FFC5FA81B535FA83FCF8" box="[1331,1348,802,827]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FAE1B536FA5AFCF8" box="[1363,1437,801,827]" italics="true" pageId="13" pageNumber="314">erebus</emphasis>
</taxonomicName>
also show a medially broken metagastric ridge of the carapace, which appears continuous in
<taxonomicName id="4C4D4D107355FFC5FB17B551FACFFC9C" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[1189,1288,837,863]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" sensu="stricto" species="ikedai">
<emphasis id="B939EA817355FFC5FB17B551FB71FC9C" box="[1189,1206,838,863]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FB76B552FACFFC9C" box="[1220,1288,837,863]" italics="true" pageId="13" pageNumber="314">ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57FA7355FFC5FAA2B552FAF3FC9C" box="[1296,1332,837,863]" pageId="13" pageNumber="314" sensu="stricto">s.s.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF236937355FFC5FF75B57EFED8FBD0" blockId="13.[151,1437,153,1043]" pageId="13" pageNumber="314">
Some variation in the material examined here is observed in the posterior transverse ridge of abdominal tergite 3 (present or absent), the distomesial spine on antennal article 3 being small or obsolescent and the form of the protogastric and mesogastric ridges (continuous to variably interrupted). These characters can be diagnostic in other species of
<taxonomicName id="4C4D4D107355FFC5FEBEB5C2FE18FC2C" authorityName="Baba" authorityYear="1969" box="[268,479,981,1007]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA817355FFC5FEBEB5C2FE18FC2C" box="[268,479,981,1007]" italics="true" pageId="13" pageNumber="314">Phylladiorhynchus</emphasis>
</taxonomicName>
indicating that the
<collectingCountry id="F35A76037355FFC5FD06B5C2FC8DFC2C" box="[692,842,981,1007]" name="New Zealand" pageId="13" pageNumber="314">New Zealand</collectingCountry>
material for
<taxonomicName id="4C4D4D107355FFC5FC6BB5C1FB86FC2C" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[985,1089,981,1007]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="erebus">
<emphasis id="B939EA817355FFC5FC6BB5C1FC2DFC2C" box="[985,1002,982,1007]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FC45B5C2FB86FC2C" box="[1015,1089,981,1007]" italics="true" pageId="13" pageNumber="314">erebus</emphasis>
</taxonomicName>
might still represent more than one species.
</paragraph>
<caption id="DF32661B7355FFC5FF25B1A0FCFCF837" ID-DOI="http://doi.org/10.5281/zenodo.3516755" ID-Zenodo-Dep="3516755" httpUri="https://zenodo.org/record/3516755/files/figure.png" pageId="13" pageNumber="314" startId="13.[151,250,1975,2000]" targetBox="[160,1424,1063,1943]" targetPageId="13">
<paragraph id="8BF236937355FFC5FF25B1A0FCFCF837" blockId="13.[151,1437,1975,2036]" pageId="13" pageNumber="314">
<emphasis id="B939EA817355FFC5FF25B1A0FED4F813" bold="true" box="[151,275,1975,2000]" pageId="13" pageNumber="314">FIGURE 5.</emphasis>
Distribution records for
<taxonomicName id="4C4D4D107355FFC5FDA3B1AFFCDDF813" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[529,794,1976,2000]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B939EA817355FFC5FDA3B1AFFCDDF813" box="[529,794,1976,2000]" italics="true" pageId="13" pageNumber="314">Phylladiorhynchus erebus</emphasis>
</taxonomicName>
<emphasis id="B939EA817355FFC5FC93B1AFFC9DF813" bold="true" box="[801,858,1976,2000]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A20A57FA7355FFC5FC93B1AFFC9DF813" box="[801,858,1976,2000]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(white circles) and
<taxonomicName id="4C4D4D107355FFC5FB97B1AEFB10F813" authority="Schnabel &amp; Ahyong, 2019" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[1061,1239,1976,2000]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="kermadecensis" status="sp. nov.">
<emphasis id="B939EA817355FFC5FB97B1AEFBF3F813" box="[1061,1076,1977,2000]" italics="true" pageId="13" pageNumber="314">P</emphasis>
.
<emphasis id="B939EA817355FFC5FBF0B1AFFB10F813" box="[1090,1239,1976,2000]" italics="true" pageId="13" pageNumber="314">kermadecensis</emphasis>
</taxonomicName>
<emphasis id="B939EA817355FFC5FB6CB1AFFADFF813" bold="true" box="[1246,1304,1976,2000]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A20A57FA7355FFC5FB6CB1AFFADFF813" box="[1246,1304,1976,2000]" pageId="13" pageNumber="314" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(crossed circle). Lines demarcating the EEZ for both Australia and NZ shown.
</paragraph>
</caption>
<paragraph id="8BF236937356FFC6FF75B68EFCE9FF14" blockId="14.[151,1436,153,215]" pageId="14" pageNumber="315">
<bibRefCitation id="EFDC4B627356FFC6FF75B68EFEA2FF70" author="Baba, K." box="[199,357,153,179]" pageId="14" pageNumber="315" pagination="479 - 491" refId="ref40974" refString="Baba, K. (1991) Crustacea Decapoda: Alainius gen. nov., Leiogalathea Baba, 1969, and Phylladiorhynchus Baba, 1969 (Galatheidae) from New Caledonia. In: Crosnier, A. (Ed.), Resultats des Campagnes Musorstom, Memoires du Museum National d'Histoire Naturelle, Paris (A), 152, 479 - 491." type="journal article" year="1991">Babas (1991)</bibRefCitation>
records of
<taxonomicName id="4C4D4D107356FFC6FE56B68DFD80FF70" authorityName="Schnabel &amp; Ahyong" authorityYear="2019" box="[484,583,153,179]" class="Malacostraca" family="Galatheidae" genus="Phylladiorhynchus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="B939EA817356FFC6FE56B68DFE32FF70" box="[484,501,154,179]" italics="true" pageId="14" pageNumber="315">P</emphasis>
.
<emphasis id="B939EA817356FFC6FDB1B68EFD80FF70" box="[515,583,153,179]" italics="true" pageId="14" pageNumber="315">ikedai</emphasis>
</taxonomicName>
from
<collectingCountry id="F35A76037356FFC6FD3EB68EFCFDFF70" box="[652,826,153,179]" name="New Caledonia" pageId="14" pageNumber="315">New Caledonia</collectingCountry>
and the
<collectingRegion id="4989F8717356FFC6FC2BB68EFB81FF70" box="[921,1094,153,179]" country="New Caledonia" name="Province des Iles" pageId="14" pageNumber="315">Loyalty Islands</collectingRegion>
might be referable to either of the new
<collectingCountry id="F35A76037356FFC6FF44B6AAFE4BFF14" box="[246,396,189,215]" name="New Zealand" pageId="14" pageNumber="315">New Zealand</collectingCountry>
species; records should be reviewed.
</paragraph>
</subSubSection>
</treatment>
</document>