359 lines
46 KiB
XML
359 lines
46 KiB
XML
<document id="7C63AEC375CB0284A169046E90A7CD57" ID-CLB-Dataset="58476" ID-DOI="10.11646/zootaxa.5315.2.8" ID-GBIF-Dataset="e09fa98b-98b2-4837-9ad3-d03be006936a" ID-ISSN="1175-5326" ID-Zenodo-Dep="8130447" ID-ZooBank="490C3836-0D20-47A8-8AE9-E67071BED336" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1688980734631" checkinUser="plazi" docAuthor="Salles, Frederico F. & Jabeen, Fatima" docDate="2023" docId="03E2F73EA111FFB6869CFF7A4135D07E" docLanguage="en" docName="zootaxa.5315.2.8.pdf" docOrigin="Zootaxa 5315 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5315.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Askola fortuita Salles & Jabeen 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="199" masterDocId="FFDB8F46A110FFB2860BFFED4221D677" masterDocTitle="A new species of Askola Peters, 1969 (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil" masterLastPageNumber="200" masterPageNumber="195" pageNumber="196" updateTime="1689165467620" updateUser="tatiana" zenodo-license-document="CLOSED">
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<mods:mods id="20D6BBD4CB8915F28C648ED66590EE65" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A777E3710A9BECAE1788BE10D2C87785">
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<mods:title id="B6722D30A53FFE882E447AB09BACEA05">A new species of Askola Peters, 1969 (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil</mods:title>
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</mods:titleInfo>
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<mods:name id="1727CBA18590D3A09F09CEAA5E4E7499" type="personal">
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<mods:role id="F07FE6B1287774CDFEAA92794B3B490A">
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<mods:roleTerm id="F7850D850584CC0B909BA25474227DC1">Author</mods:roleTerm>
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<mods:namePart id="5C856CCACF31F4E3CD785CFE151940B9">Salles, Frederico F.</mods:namePart>
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<mods:affiliation id="F34AD99C2990C2091D92150432947AC8">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="9556DC798F776AACA16CAC64643B4344" type="personal">
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<mods:roleTerm id="A95169563CAC024579BCF719C0798A35">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50D3A3B8843708B289767B5E5EB1B224">Jabeen, Fatima</mods:namePart>
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<mods:nameIdentifier id="CD5EA3000D334F18E4AFFB69A8044CAB" type="ORCID">0000-0003-0407-8713</mods:nameIdentifier>
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<mods:affiliation id="9EB82D6421DB7AA52BF5B722F0395307">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil & Programa de Pós-graduação em Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil & fatima. jabeen @ ufv. br; https: // orcid. org / 0000 - 0003 - 0407 - 8713</mods:affiliation>
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<mods:nameIdentifier id="15BA470C2586967EB43EC2D2E06D3E6A" type="email">fatima.jabeen@ufv.br</mods:nameIdentifier>
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<mods:typeOfResource id="5010BDA8100A8540FC7CBE00207C6A44">text</mods:typeOfResource>
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<mods:title id="45EFD52915F204F0B15FD2AEB3130E3C">Zootaxa</mods:title>
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<mods:part id="7E6A253695CB108285B5BFA598AD9633">
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<mods:date id="B69353D4606F8634472C63C0360D3A41">2023</mods:date>
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<mods:number id="0703B11AA35E94CB5D64BDB353027D60">2023-07-07</mods:number>
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<mods:detail id="3437E0B2DB9A9BEA5269647E0A250789" type="volume">
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<mods:number id="5B886F089246BA913E5CE74EB5FFCF90">5315</mods:number>
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<mods:start id="F8605D194F4FAB4C31624604002693FD">195</mods:start>
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<mods:url id="781338B1DC8B08021F2EED3409A44F28">http://dx.doi.org/10.11646/zootaxa.5315.2.8</mods:url>
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<mods:identifier id="F8BF9FAAC60A0FD1C392ADA687175366" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E2F73EA111FFB6869CFF7A4135D07E" ID-DOI="http://doi.org/10.5281/zenodo.8130545" ID-GBIF-Taxon="210892583" ID-Zenodo-Dep="8130545" LSID="urn:lsid:plazi:treatment:03E2F73EA111FFB6869CFF7A4135D07E" httpUri="http://treatment.plazi.org/id/03E2F73EA111FFB6869CFF7A4135D07E" lastPageId="4" lastPageNumber="199" pageId="1" pageNumber="196">
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<subSubSection id="C35115A3A111FFB3869CFF7A40A8D6C6" box="[151,649,151,178]" pageId="1" pageNumber="196" type="nomenclature">
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<paragraph id="8BF44628A111FFB3869CFF7A40A8D6C6" blockId="1.[151,649,151,213]" box="[151,649,151,178]" pageId="1" pageNumber="196">
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<heading id="D0BCF144A111FFB3869CFF7A40A8D6C6" bold="true" box="[151,649,151,178]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
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<emphasis id="B93F9A3AA111FFB3869CFF7A40A8D6C6" bold="true" box="[151,649,151,178]" pageId="1" pageNumber="196">
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<taxonomicName id="4C4B3DABA111FFB3869CFF7A403CD6C6" authority="Salles & Jabeen" authorityName="Salles & Jabeen" authorityYear="2023" box="[151,541,151,178]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
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||
<emphasis id="B93F9A3AA111FFB3869CFF7A436CD6C6" bold="true" box="[151,333,151,178]" italics="true" pageId="1" pageNumber="196">Askola fortuita</emphasis>
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Salles & Jabeen
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A20C2741A111FFB38421FF7A40A8D6C6" box="[554,649,151,177]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35115A3A111FFB3869CFF564333D6A2" box="[151,274,187,213]" pageId="1" pageNumber="196" type="description">
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||
<paragraph id="8BF44628A111FFB3869CFF564333D6A2" blockId="1.[151,649,151,213]" box="[151,274,187,213]" pageId="1" pageNumber="196">
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||
(
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<figureCitation id="13705AADA111FFB38694FF564328D6A2" box="[159,265,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1484,1508]" captionStartId-1="2.[151,250,1931,1955]" captionStartId-2="3.[151,250,1929,1953]" captionTargetBox-0="[173,1414,567,1453]" captionTargetBox-1="[209,1378,181,1896]" captionTargetBox-2="[206,1382,196,1905]" captionTargetId-0="figure-124@1.[166,1421,561,1460]" captionTargetId-1="figure-24@2.[209,1378,181,1907]" captionTargetId-2="figure-19@3.[206,1382,181,1905]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Map of South America and known distribution of Askola fortuita sp. nov. in Caparaó mountain range." captionText-1="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." captionText-2="FIGURE 3. Askola fortuita sp. nov. male imago: A, genitalia, ventral view; B, genitalia, dorsal view; C, genitalia, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.8130449" figureDoi-1="http://doi.org/10.5281/zenodo.8130451" figureDoi-2="http://doi.org/10.5281/zenodo.8130453" httpUri-0="https://zenodo.org/record/8130449/files/figure.png" httpUri-1="https://zenodo.org/record/8130451/files/figure.png" httpUri-2="https://zenodo.org/record/8130453/files/figure.png" pageId="1" pageNumber="196">Figs. 1–3</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35115A3A111FFB3869CFEEE41CBD46E" pageId="1" pageNumber="196" type="diagnosis">
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<paragraph id="8BF44628A111FFB3869CFEEE4331D76A" blockId="1.[151,272,259,285]" box="[151,272,259,285]" pageId="1" pageNumber="196">
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||
<heading id="D0BCF144A111FFB3869CFEEE4331D76A" bold="true" box="[151,272,259,285]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
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||
<emphasis id="B93F9A3AA111FFB3869CFEEE4331D76A" bold="true" box="[151,272,259,285]" pageId="1" pageNumber="196">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF44628A111FFB3869CFEA641CBD46E" blockId="1.[151,1436,331,538]" pageId="1" pageNumber="196">
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The male imago of
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<taxonomicName id="4C4B3DABA111FFB3878AFEA643D6D712" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[385,503,331,357]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
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<emphasis id="B93F9A3AA111FFB3878AFEA643D6D712" box="[385,503,331,357]" italics="true" pageId="1" pageNumber="196">A. fortuita</emphasis>
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||
</taxonomicName>
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<emphasis id="B93F9A3AA111FFB38409FEA64041D712" bold="true" box="[514,608,331,357]" pageId="1" pageNumber="196">
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<taxonomicNameLabel id="A20C2741A111FFB38409FEA64041D712" box="[514,608,331,357]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can be distinguished from other species of the genus by the following combination of characteristics: dorsal portion of compound eyes widely meeting on meson of head (
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<figureCitation id="13705AADA111FFB38339FE8247AFD7FE" box="[1330,1422,367,393]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Fig. 2B</figureCitation>
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); forewings hyaline (
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<figureCitation id="13705AADA111FFB38771FE7E43F7D7DA" box="[378,470,403,429]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Fig. 2E</figureCitation>
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); femora white with a submedial mark and an apical band, both orangish brown; abdominal coloration mostly translucent white (
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<figureCitation id="13705AADA111FFB384BEFE5A4131D7A6" box="[693,784,439,465]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Fig. 2D</figureCitation>
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); genitalia with styliger plate washed with brown and red laterally, forceps segment I reddish-brown on basal ¼, remainder translucent white, penes yellowish brown; and penes not extending beyond posterior margin of styliger plate (
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<figureCitation id="13705AADA111FFB3855CFE1241FFD46E" box="[855,990,511,537]" captionStart="FIGURE 3" captionStartId="3.[151,250,1929,1953]" captionTargetBox="[206,1382,196,1905]" captionTargetId="figure-19@3.[206,1382,181,1905]" captionTargetPageId="3" captionText="FIGURE 3. Askola fortuita sp. nov. male imago: A, genitalia, ventral view; B, genitalia, dorsal view; C, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8130453" httpUri="https://zenodo.org/record/8130453/files/figure.png" pageId="1" pageNumber="196">Figs. 3A–C</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF3416A0A111FFB3869CFA214707D392" ID-DOI="http://doi.org/10.5281/zenodo.8130449" ID-Zenodo-Dep="8130449" box="[151,1318,1484,1509]" httpUri="https://zenodo.org/record/8130449/files/figure.png" pageId="1" pageNumber="196" startId="1.[151,250,1484,1508]" targetBox="[173,1414,567,1453]" targetPageId="1" targetType="figure">
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<paragraph id="8BF44628A111FFB3869CFA214707D392" blockId="1.[151,1318,1484,1509]" box="[151,1318,1484,1509]" pageId="1" pageNumber="196">
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<emphasis id="B93F9A3AA111FFB3869CFA214332D393" bold="true" box="[151,275,1484,1508]" pageId="1" pageNumber="196">FIGURE 1.</emphasis>
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Map of South America and known distribution of
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<taxonomicName id="4C4B3DABA111FFB3851EFA21418CD393" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[789,941,1484,1508]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
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<emphasis id="B93F9A3AA111FFB3851EFA21418CD393" box="[789,941,1484,1508]" italics="true" pageId="1" pageNumber="196">Askola fortuita</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B93F9A3AA111FFB385B8FA214627D393" bold="true" box="[947,1030,1484,1508]" pageId="1" pageNumber="196">
|
||
<taxonomicNameLabel id="A20C2741A111FFB385B8FA214627D393" box="[947,1030,1484,1508]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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in Caparaó mountain range.
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</paragraph>
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</caption>
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||
<subSubSection id="C35115A3A111FFB686CCF9DB40A3D4DE" lastPageId="4" lastPageNumber="199" pageId="1" pageNumber="196" type="description">
|
||
<paragraph id="8BF44628A111FFB386CCF9DB4052D027" blockId="1.[151,1437,1590,1941]" box="[199,627,1590,1616]" pageId="1" pageNumber="196">
|
||
<emphasis id="B93F9A3AA111FFB386CCF9DB4345D027" bold="true" box="[199,356,1590,1616]" pageId="1" pageNumber="196">Descriptions.</emphasis>
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Male imago (
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||
<figureCitation id="13705AADA111FFB3840BF9DB4071D027" box="[512,592,1590,1616]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Figs. 2</figureCitation>
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,
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<figureCitation id="13705AADA111FFB38457F9DB404BD027" box="[604,618,1590,1616]" captionStart="FIGURE 3" captionStartId="3.[151,250,1929,1953]" captionTargetBox="[206,1382,196,1905]" captionTargetId="figure-19@3.[206,1382,181,1905]" captionTargetPageId="3" captionText="FIGURE 3. Askola fortuita sp. nov. male imago: A, genitalia, ventral view; B, genitalia, dorsal view; C, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8130453" httpUri="https://zenodo.org/record/8130453/files/figure.png" pageId="1" pageNumber="196">3</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF44628A111FFB386CCF9B74738D002" blockId="1.[151,1437,1590,1941]" box="[199,1305,1626,1653]" pageId="1" pageNumber="196">
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<emphasis id="B93F9A3AA111FFB386CCF9B74360D003" bold="true" box="[199,321,1626,1652]" pageId="1" pageNumber="196">Measures.</emphasis>
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Body:
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<quantity id="4CB3EBCDA111FFB3879FF9B7403FD002" box="[404,542,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.8" metricValueMin="5.2" pageId="1" pageNumber="196" unit="mm" value="5.5" valueMax="5.8" valueMin="5.2">5.2–5.8 mm</quantity>
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; forewing: 6.0–
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<quantity id="4CB3EBCDA111FFB384C4F9B74106D002" box="[719,807,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="196" unit="mm" value="6.7">6.7 mm</quantity>
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. Foreleg:
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<quantity id="4CB3EBCDA111FFB3859DF9B741CED002" box="[918,1007,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="1" pageNumber="196" unit="mm" value="5.9">5.9 mm</quantity>
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. Caudal filament:
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<quantity id="4CB3EBCDA111FFB382B7F9B74734D002" box="[1212,1301,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="196" unit="mm" value="9.9">9.9 mm</quantity>
|
||
.
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</paragraph>
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<paragraph id="8BF44628A111FFB386CCF9934644D172" blockId="1.[151,1437,1590,1941]" pageId="1" pageNumber="196">
|
||
<emphasis id="B93F9A3AA111FFB386CCF9934332D0EF" bold="true" box="[199,275,1662,1688]" pageId="1" pageNumber="196">Ratios</emphasis>
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.
|
||
<emphasis id="B93F9A3AA111FFB3872AF9924346D0EF" box="[289,359,1663,1688]" italics="true" pageId="1" pageNumber="196">Wings</emphasis>
|
||
: forewing width 0.35× forewing length; fork on MA
|
||
<subScript id="17CF446DA111FFB385BCF97C419ED0E8" attach="left" box="[951,959,1681,1695]" fontSize="6" pageId="1" pageNumber="196">2</subScript>
|
||
of forewing
|
||
<quantity id="4CB3EBCDA111FFB3825AF993468ED0EF" box="[1105,1199,1662,1688]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="1" pageNumber="196" unit="mm" value="0.47">0.47mm</quantity>
|
||
× total length of MA.
|
||
<emphasis id="B93F9A3AA111FFB3869CF94F4320D0CB" box="[151,257,1698,1724]" italics="true" pageId="1" pageNumber="196">Genitalia</emphasis>
|
||
: medial length of styliger plate 0.8× maximum width; lateral length of styliger plate 1.2× medial length of styliger plate. Forceps segment III 1.0× forceps segment II; forceps segment III 0.16× forceps segment I; forceps segment II 1.6× forceps segment I. Penis length approximately 1.0× forceps segment I.
|
||
</paragraph>
|
||
<paragraph id="8BF44628A111FFB386CCF8E34720D15F" blockId="1.[151,1437,1590,1941]" box="[199,1281,1806,1832]" pageId="1" pageNumber="196">
|
||
<emphasis id="B93F9A3AA111FFB386CCF8E3403AD15F" bold="true" box="[199,539,1806,1832]" pageId="1" pageNumber="196">Coloration and morphology.</emphasis>
|
||
General coloration (
|
||
<figureCitation id="13705AADA111FFB38508F8E3417FD15F" box="[771,862,1806,1832]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Fig. 2A</figureCitation>
|
||
): brown anteriorly, white posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BF44628A111FFB386CCF8DF4619D1E3" blockId="1.[151,1437,1590,1941]" pageId="1" pageNumber="196">
|
||
Head: head brown, ocelli surrounded by black, frons orange. Compound eyes widely meet dorsally on median portion of head (
|
||
<figureCitation id="13705AADA111FFB38745F8BB4384D107" box="[334,421,1878,1904]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Fig. 2B</figureCitation>
|
||
); dorsal surface oblong, wider than long; dorsal portion gray, lower portion black (
|
||
<figureCitation id="13705AADA111FFB3833DF8BB428CD1E3" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="1" pageNumber="196">Figs. 2B, C</figureCitation>
|
||
). Antenna translucent white, except for scape with outer margin orangish brown.
|
||
</paragraph>
|
||
<caption id="DF3416A0A112FFB0869CF8664087D1B0" ID-DOI="http://doi.org/10.5281/zenodo.8130451" ID-Zenodo-Dep="8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="2" pageNumber="197" startId="2.[151,250,1931,1955]" targetBox="[209,1378,181,1896]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BF44628A112FFB0869CF8664087D1B0" blockId="2.[151,1437,1931,1991]" pageId="2" pageNumber="197">
|
||
<emphasis id="B93F9A3AA112FFB0869CF866432CD1D4" bold="true" box="[151,269,1931,1955]" pageId="2" pageNumber="197">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B3DABA112FFB08712F8664391D1D4" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[281,432,1931,1955]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
|
||
<emphasis id="B93F9A3AA112FFB08712F8664391D1D4" box="[281,432,1931,1955]" italics="true" pageId="2" pageNumber="197">Askola fortuita</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9A3AA112FFB087BDF8664029D1D4" bold="true" box="[438,520,1931,1955]" pageId="2" pageNumber="197">
|
||
<taxonomicNameLabel id="A20C2741A112FFB087BDF8664029D1D4" box="[438,520,1931,1955]" pageId="2" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3416A0A113FFB1869CF86442F1D1B2" ID-DOI="http://doi.org/10.5281/zenodo.8130453" ID-Zenodo-Dep="8130453" httpUri="https://zenodo.org/record/8130453/files/figure.png" pageId="3" pageNumber="198" startId="3.[151,250,1929,1953]" targetBox="[206,1382,196,1905]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BF44628A113FFB1869CF86442F1D1B2" blockId="3.[151,1436,1929,1989]" pageId="3" pageNumber="198">
|
||
<emphasis id="B93F9A3AA113FFB1869CF8644331D1D6" bold="true" box="[151,272,1929,1953]" pageId="3" pageNumber="198">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B3DABA113FFB18714F864439BD1D6" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[287,442,1929,1953]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
|
||
<emphasis id="B93F9A3AA113FFB18714F864439BD1D6" box="[287,442,1929,1953]" italics="true" pageId="3" pageNumber="198">Askola fortuita</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9A3AA113FFB187C8F8644038D1D6" bold="true" box="[451,537,1929,1953]" pageId="3" pageNumber="198">
|
||
<taxonomicNameLabel id="A20C2741A113FFB187C8F8644038D1D6" box="[451,537,1929,1953]" pageId="3" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
male imago: A, genitalia, ventral view; B, genitalia, dorsal view; C, genitalia, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF44628A114FFB686CCFF7A46A9D7DA" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="199">
|
||
Thorax (
|
||
<figureCitation id="13705AADA114FFB68723FF7A438CD6C6" box="[296,429,151,177]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="4" pageNumber="199">Figs. 2B, C</figureCitation>
|
||
): Pronotum brown with white mark on posterior margin. Mesonotum brown with median longitudinal suture yellowish brown, anteronotal impression and medioparapsidal suture dark brown, posterior scutal protuberance and scutellum yellowish brown. Metanotum yellowish brown. Pleurae yellowish brown, with some areas dark brown; sterna dark brown. Fore femur white with submedial mark and apical band, both orangish brown; fore tibia white with weak orangish brown mark on apex; tarsi and claw white. Mid and hind legs similar to fore leg, except for tarsi, darker toward apex, and for mark on apex of femora, brown. Wings (
|
||
<figureCitation id="13705AADA114FFB682B3FEA64731D712" box="[1208,1296,331,357]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="4" pageNumber="199">Fig. 2E</figureCitation>
|
||
). Membrane hyaline. Longitudinal veins light yellow, cross veins white; base of wing and base of veins C, Sc and RA orange brown; costal arc dark brown; 3 cross veins basal to bullae, 14 cross veins distal to bullae.
|
||
</paragraph>
|
||
<paragraph id="8BF44628A114FFB686CCFE5A42F4D4F2" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="199">
|
||
Abdomen (
|
||
<figureCitation id="13705AADA114FFB6874CFE5A4384D7A6" box="[327,421,439,465]" captionStart="FIGURE 2" captionStartId="2.[151,250,1931,1955]" captionTargetBox="[209,1378,181,1896]" captionTargetId="figure-24@2.[209,1378,181,1907]" captionTargetPageId="2" captionText="FIGURE 2. Askola fortuita sp. nov. male imago: A, habitus, lateral view; B, head and thorax, dorsal view; C, head and thorax, lateral view; D, abdomen, dorsal view; E, forewing." figureDoi="http://doi.org/10.5281/zenodo.8130451" httpUri="https://zenodo.org/record/8130451/files/figure.png" pageId="4" pageNumber="199">Fig. 2D</figureCitation>
|
||
): translucent white. Tergum I with triangular brown mark anteromedially; terga I to VII translucent white, darker posterolaterally; posterior half of tergum VII and terga VIII to X heavily washed with brown and slightly tinged with orange. Sterna white; sterna I to VII with small posterolateral gray mark. Genitalia (
|
||
<figureCitation id="13705AADA114FFB68694FDCE430ED44A" box="[159,303,547,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1929,1953]" captionTargetBox="[206,1382,196,1905]" captionTargetId="figure-19@3.[206,1382,181,1905]" captionTargetPageId="3" captionText="FIGURE 3. Askola fortuita sp. nov. male imago: A, genitalia, ventral view; B, genitalia, dorsal view; C, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8130453" httpUri="https://zenodo.org/record/8130453/files/figure.png" pageId="4" pageNumber="199">Figs. 3 A–C</figureCitation>
|
||
). Styliger plate washed with brown and red laterally; forceps segment I washed with brown and red on basal 1/4, remainder of forceps white; penes yellowish brown, not extending beyond posterior margin of styliger plate.
|
||
</paragraph>
|
||
<paragraph id="8BF44628A114FFB686CCFD6240A3D4DE" blockId="4.[151,1437,151,1545]" box="[199,642,654,681]" pageId="4" pageNumber="199">
|
||
<emphasis id="B93F9A3AA114FFB686CCFD624024D4DE" bold="true" box="[199,517,654,681]" pageId="4" pageNumber="199">Female imago and nymph.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35115A3A114FFB686CCFD5E4194D4BA" box="[199,949,691,717]" pageId="4" pageNumber="199" type="etymology">
|
||
<paragraph id="8BF44628A114FFB686CCFD5E4194D4BA" blockId="4.[151,1437,151,1545]" box="[199,949,691,717]" pageId="4" pageNumber="199">
|
||
<emphasis id="B93F9A3AA114FFB686CCFD5E436ED4BA" bold="true" box="[199,335,691,717]" pageId="4" pageNumber="199">Etymology.</emphasis>
|
||
From the latin adjective
|
||
<emphasis id="B93F9A3AA114FFB68463FD5E40E6D4BA" box="[616,711,691,717]" italics="true" pageId="4" pageNumber="199">fortuitus</emphasis>
|
||
, meaning at random.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35115A3A114FFB686CCFD3B469BD486" box="[199,1210,726,753]" pageId="4" pageNumber="199" type="distribution">
|
||
<paragraph id="8BF44628A114FFB686CCFD3B469BD486" blockId="4.[151,1437,151,1545]" box="[199,1210,726,753]" pageId="4" pageNumber="199">
|
||
<emphasis id="B93F9A3AA114FFB686CCFD3B4340D487" bold="true" box="[199,353,726,752]" pageId="4" pageNumber="199">Distribution.</emphasis>
|
||
<collectingCountry id="F35C06B8A114FFB6876CFD3A4391D486" box="[359,432,727,753]" name="Brazil" pageId="4" pageNumber="199">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="498F88CAA114FFB687B0FD3A40B8D486" box="[443,665,727,753]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="199">Espírito Santo State</collectingRegion>
|
||
(close to the border with
|
||
<collectingRegion id="498F88CAA114FFB685B2FD3A4670D486" box="[953,1105,727,753]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="199">Minas Gerais</collectingRegion>
|
||
) (
|
||
<figureCitation id="13705AADA114FFB68263FD3A4688D486" box="[1128,1193,727,753]" captionStart="FIGURE 1" captionStartId="1.[151,250,1484,1508]" captionTargetBox="[173,1414,567,1453]" captionTargetId="figure-124@1.[166,1421,561,1460]" captionTargetPageId="1" captionText="FIGURE 1. Map of South America and known distribution of Askola fortuita sp. nov. in Caparaó mountain range." figureDoi="http://doi.org/10.5281/zenodo.8130449" httpUri="https://zenodo.org/record/8130449/files/figure.png" pageId="4" pageNumber="199">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35115A3A114FFB686CCFD164644D52A" pageId="4" pageNumber="199" type="materials_examined">
|
||
<paragraph id="8BF44628A114FFB686CCFD164644D52A" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="199">
|
||
<emphasis id="B93F9A3AA114FFB686CCFD1643CDD562" bold="true" box="[199,492,762,789]" pageId="4" pageNumber="199">
|
||
Type material.
|
||
<materialsCitation id="3B234C75A114FFB68774FD16430DD52A" collectingDate="2022-12-29" collectionCode="UFVB" collectorName="Salles, F. F." country="Brazil" elevation="1340" latitude="-20.5325" location="Pedra Menina" longLatPrecision="21" longitude="-41.790276" municipality="Dores do Rio Preto" pageId="4" pageNumber="199" specimenCount="1" specimenCount-male="1" stateProvince="Espirito Santo State" typeStatus="holotype">
|
||
<typeStatus id="54F0F88AA114FFB68774FD1643CDD562" box="[383,492,763,789]" pageId="4" pageNumber="199">Holotype</typeStatus>
|
||
:
|
||
<specimenCount id="9D4D8DA1A114FFB687F7FD114030D560" box="[508,529,764,791]" pageId="4" pageNumber="199" type="male">♂</specimenCount>
|
||
imago in alcohol,
|
||
<collectingCountry id="F35C06B8A114FFB684EEFD16410DD562" box="[741,812,763,789]" name="Brazil" pageId="4" pageNumber="199">Brazil</collectingCountry>
|
||
,
|
||
<collectingRegion id="498F88CAA114FFB68533FD16463AD562" box="[824,1051,763,789]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="199">Espírito Santo State</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B90DC52A114FFB6822DFD164721D562" box="[1062,1280,763,789]" pageId="4" pageNumber="199">Dores do Rio Preto</collectingMunicipality>
|
||
municipality,
|
||
<location id="8E9410F3A114FFB6869CFCF24314D54E" LSID="urn:lsid:plazi:treatment:03E2F73EA111FFB6869CFF7A4135D07E:8E9410F3A114FFB6869CFCF24314D54E" box="[151,309,799,825]" country="Brazil" latitude="-20.5325" longLatPrecision="21" longitude="-41.790276" municipality="Dores do Rio Preto" name="Pedra Menina" pageId="4" pageNumber="199" stateProvince="Espirito Santo State">Pedra Menina</location>
|
||
,
|
||
<location id="8E9410F3A114FFB68734FCF243E2D54E" LSID="urn:lsid:plazi:treatment:03E2F73EA111FFB6869CFF7A4135D07E:8E9410F3A114FFB68734FCF243E2D54E" box="[319,451,799,825]" country="Brazil" latitude="-20.5325" longLatPrecision="21" longitude="-41.790276" municipality="Dores do Rio Preto" name="Vo Nininho" pageId="4" pageNumber="199" stateProvince="Espirito Santo State">Vô Nininho</location>
|
||
property, S„o Bento stream,
|
||
<geoCoordinate id="EE7F20EFA114FFB6850EFCF341AAD54E" box="[773,907,798,825]" degrees="20" direction="south" minutes="31" orientation="latitude" pageId="4" pageNumber="199" precision="15" seconds="57" value="-20.5325">S20°31’57”</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7F20EFA114FFB6859DFCF34609D54E" box="[918,1064,798,825]" degrees="41" direction="west" minutes="47" orientation="longitude" pageId="4" pageNumber="199" precision="15" seconds="25" value="-41.790276">W41°47’25”</geoCoordinate>
|
||
,
|
||
<quantity id="4CB3EBCDA114FFB68239FCF246A9D54E" box="[1074,1160,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="4" pageNumber="199" unit="m" value="1340.0">
|
||
<elevation id="0066A11BA114FFB68239FCF246A9D54E" box="[1074,1160,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.34" pageId="4" pageNumber="199" unit="m" value="1340.0">1340 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFF560E8A114FFB68298FCF24734D54E" box="[1171,1301,799,825]" pageId="4" pageNumber="199" value="2022-12-29">
|
||
<collectingDate id="EFB19900A114FFB68298FCF24734D54E" box="[1171,1301,799,825]" pageId="4" pageNumber="199" value="2022-12-29">29.xii.2022</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BE23FEA114FFB68314FCF247BDD54E" box="[1311,1436,799,825]" pageId="4" pageNumber="199">Salles, F.F.</collectorName>
|
||
col. (
|
||
<collectionCode id="ED5ADEEDA114FFB686C4FCAE4301D52A" box="[207,288,835,861]" pageId="4" pageNumber="199">UFVB</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</emphasis>
|
||
<materialsCitation id="3B234C75A114FFB68738FCAE4644D52A" box="[307,1125,835,863]" collectingDate="2022-12-29" collectionCode="UFVB, MNRJ" collectorName="Salles, F. F." country="Brazil" elevation="1340" latitude="-20.5325" location="Pedra Menina" longLatPrecision="21" longitude="-41.790276" municipality="Dores do Rio Preto" pageId="4" pageNumber="199" specimenCount="4" specimenCount-male="4" stateProvince="Espirito Santo State" typeStatus="paratype">
|
||
<emphasis id="B93F9A3AA114FFB68738FCAE438AD52A" bold="true" box="[307,427,835,861]" pageId="4" pageNumber="199">
|
||
<typeStatus id="54F0F88AA114FFB68738FCAE438AD52A" box="[307,427,835,861]" pageId="4" pageNumber="199">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D4D8DA1A114FFB687B2FCAE43C2D528" box="[441,483,835,863]" pageId="4" pageNumber="199" type="male">4 ♂</specimenCount>
|
||
imagos, same data as holotype (2 at
|
||
<collectionCode id="ED5ADEEDA114FFB6858BFCAE41ECD52A" box="[896,973,835,861]" collectionName="Brazil, Minas Gerais, Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="4" pageNumber="199">UFVB</collectionCode>
|
||
, 2 at
|
||
<collectionCode id="ED5ADEEDA114FFB68202FCAE4678D52A" box="[1033,1113,835,861]" collectionName="Brazil, Rio de Janeiro, Sao Cristovao, Universidade do Rio Janeiro, Museu Nacional" pageId="4" pageNumber="199">MNRJ</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35115A3A114FFB686CCFC8B4135D07E" pageId="4" pageNumber="199" type="discussion">
|
||
<paragraph id="8BF44628A114FFB686CCFC8B4135D07E" blockId="4.[151,1437,151,1545]" pageId="4" pageNumber="199">
|
||
<emphasis id="B93F9A3AA114FFB686CCFC8B4369D5F7" bold="true" box="[199,328,870,896]" pageId="4" pageNumber="199">Comments</emphasis>
|
||
. According to
|
||
<bibRefCitation id="EFDA3BD9A114FFB687F2FC8A40B0D5F6" author="Peters, W. L." box="[505,657,871,897]" pageId="4" pageNumber="199" pagination="253 - 258" refId="ref2422" refString="Peters, W. L. (1969) Askola froehlichi a new genus and species from southern Brazil (Leptophlebiidae: Ephemeroptera). Florida Entomologist, 52, 253 - 258. https: // doi. org / 10.2307 / 3493877" type="journal article" year="1969">Peters (1969)</bibRefCitation>
|
||
in the original description of
|
||
<taxonomicName id="4C4B3DABA114FFB685E4FC8A465FD5F6" authorityName="Peters" authorityYear="1969" box="[1007,1150,871,897]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB685E4FC8A465FD5F6" box="[1007,1150,871,897]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
, there is variation in the abdominal color pattern of the species: some male subimagos present more extensive dark-brown marks on terga II to IX (an abdominal color pattern approaching that of the female), while in others male subimagos markings on the sterna are faded to light-brown. These slight discrepancies are expected due to the fading of the specimens after a period in ethanol, or even as a result of temporal and/or geographic variation.
|
||
<bibRefCitation id="EFDA3BD9A114FFB68211FC1A472BD266" author="Campos, R. & Mariano, R. & Calor, A. R." box="[1050,1290,1015,1041]" pageId="4" pageNumber="199" pagination="69 - 92" refId="ref2147" refString="Campos, R., Mariano, R. & Calor, A. R. (2019) Askola Peters 1969 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae): an updated review under cladistics approach. Zoologischer Anzeiger, 283, 69 - 92. https: // doi. org / 10.1016 / j. jcz. 2019.08.006" type="journal article" year="2019">
|
||
Campos
|
||
<emphasis id="B93F9A3AA114FFB68276FC1A468FD266" box="[1149,1198,1015,1041]" italics="true" pageId="4" pageNumber="199">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
, on the other hand, interpreted a remarkable variation in the abdominal coloration pattern of
|
||
<taxonomicName id="4C4B3DABA114FFB6820DFBF646B0D242" authorityName="Peters" authorityYear="1969" box="[1030,1169,1051,1077]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB6820DFBF646B0D242" box="[1030,1169,1051,1077]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
, as illustrated in figures 3A–F of their paper. It’s our opinion, however, that some of these morphotypes might represent distinct and new species. In the case of
|
||
<taxonomicName id="4C4B3DABA114FFB68798FB8E4027D20A" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[403,518,1123,1149]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
|
||
<emphasis id="B93F9A3AA114FFB68798FB8E4027D20A" box="[403,518,1123,1149]" italics="true" pageId="4" pageNumber="199">A. fortuita</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9A3AA114FFB68406FB8E4049D20A" bold="true" box="[525,616,1123,1149]" pageId="4" pageNumber="199">
|
||
<taxonomicNameLabel id="A20C2741A114FFB68406FB8E4049D20A" box="[525,616,1123,1149]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which is similar to
|
||
<bibRefCitation id="EFDA3BD9A114FFB68547FB8E4619D20A" author="Campos, R. & Mariano, R. & Calor, A. R." box="[844,1080,1123,1149]" pageId="4" pageNumber="199" pagination="69 - 92" refId="ref2147" refString="Campos, R., Mariano, R. & Calor, A. R. (2019) Askola Peters 1969 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae): an updated review under cladistics approach. Zoologischer Anzeiger, 283, 69 - 92. https: // doi. org / 10.1016 / j. jcz. 2019.08.006" type="journal article" year="2019">
|
||
Campos
|
||
<emphasis id="B93F9A3AA114FFB685A4FB8E41C1D20A" box="[943,992,1123,1149]" italics="true" pageId="4" pageNumber="199">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
morphotype C (also distributed in Caparaó—the closest point to the letter C on figure 3 of
|
||
<bibRefCitation id="EFDA3BD9A114FFB68527FB6A462BD2D6" author="Campos, R. & Mariano, R. & Calor, A. R." box="[812,1034,1159,1185]" pageId="4" pageNumber="199" pagination="69 - 92" refId="ref2147" refString="Campos, R., Mariano, R. & Calor, A. R. (2019) Askola Peters 1969 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae): an updated review under cladistics approach. Zoologischer Anzeiger, 283, 69 - 92. https: // doi. org / 10.1016 / j. jcz. 2019.08.006" type="journal article" year="2019">
|
||
Campos
|
||
<emphasis id="B93F9A3AA114FFB68584FB6A41E1D2D6" box="[911,960,1159,1185]" italics="true" pageId="4" pageNumber="199">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
), important differences are found in relation to the original description of
|
||
<taxonomicName id="4C4B3DABA114FFB6844AFB4640EFD2B2" authorityName="Peters" authorityYear="1969" box="[577,718,1195,1221]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB6844AFB4640EFD2B2" box="[577,718,1195,1221]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
, even considering the variation noted by
|
||
<bibRefCitation id="EFDA3BD9A114FFB682ADFB46471ED2B2" author="Peters, W. L." box="[1190,1343,1195,1221]" pageId="4" pageNumber="199" pagination="253 - 258" refId="ref2422" refString="Peters, W. L. (1969) Askola froehlichi a new genus and species from southern Brazil (Leptophlebiidae: Ephemeroptera). Florida Entomologist, 52, 253 - 258. https: // doi. org / 10.2307 / 3493877" type="journal article" year="1969">Peters (1969)</bibRefCitation>
|
||
, as well as in relation to material of this species examined by us from other areas in
|
||
<collectingRegion id="498F88CAA114FFB685FCFB2246AFD29E" box="[1015,1166,1231,1257]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="199">Minas Gerais</collectingRegion>
|
||
(Brigadeiro State Park) and especially from
|
||
<collectingRegion id="498F88CAA114FFB6878AFB1E4002D37A" box="[385,547,1267,1293]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="199">Espírito Santo</collectingRegion>
|
||
(Augusto Ruchi Biological Reserve). In that case more than
|
||
<specimenCount id="9D4D8DA1A114FFB682E9FB1E475DD37A" box="[1250,1404,1267,1293]" pageId="4" pageNumber="199" type="generic">40 specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C4B3DABA114FFB6869CFAFA4304D346" authorityName="Peters" authorityYear="1969" box="[151,293,1303,1329]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB6869CFAFA4304D346" box="[151,293,1303,1329]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
were examined, collected monthly for more than one year (
|
||
<date id="FFF560E8A114FFB685D9FAFA465FD346" box="[978,1150,1303,1329]" pageId="4" pageNumber="199" value="2017-07-26">26th July 2017</date>
|
||
to
|
||
<date id="FFF560E8A114FFB682AEFAFA47B4D346" box="[1189,1429,1303,1329]" pageId="4" pageNumber="199" value="2018-11-21">21st November 2018</date>
|
||
) in three streams. The differences are: a) the coloration of the femora, especially of the posterior pair, which in
|
||
<taxonomicName id="4C4B3DABA114FFB6838FFAD64325D30E" authorityName="Peters" authorityYear="1969" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB6838FFAD64325D30E" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
is light brown and always represented by a medial and an apical brown band, while in the new species is white with a submedial mark and an apical band, both orangish brown; b) the abdominal coloration, which is mostly translucent white in
|
||
<taxonomicName id="4C4B3DABA114FFB68789FA4A43D6D3B6" authority="Salles & Jabeen, 2023" authorityName="Salles & Jabeen" authorityYear="2023" box="[386,503,1447,1473]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="fortuita" status="sp. nov.">
|
||
<emphasis id="B93F9A3AA114FFB68789FA4A43D6D3B6" box="[386,503,1447,1473]" italics="true" pageId="4" pageNumber="199">A. fortuita</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93F9A3AA114FFB6840BFA4A407CD3B6" bold="true" box="[512,605,1447,1473]" pageId="4" pageNumber="199">
|
||
<taxonomicNameLabel id="A20C2741A114FFB6840BFA4A407CD3B6" box="[512,605,1447,1473]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but light-brown with dark brown marks in
|
||
<taxonomicName id="4C4B3DABA114FFB68250FA4A46CBD3B6" authorityName="Peters" authorityYear="1969" box="[1115,1258,1447,1473]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB68250FA4A46CBD3B6" box="[1115,1258,1447,1473]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
(see comments above); and c) the forceps segment I, basal ¼ of
|
||
<taxonomicName id="4C4B3DABA114FFB684BAFA26411CD392" authorityName="Peters" authorityYear="1969" box="[689,829,1483,1509]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="froehlichi">
|
||
<emphasis id="B93F9A3AA114FFB684BAFA26411CD392" box="[689,829,1483,1509]" italics="true" pageId="4" pageNumber="199">A. froehlichi</emphasis>
|
||
</taxonomicName>
|
||
is brown, remainder is paler, while basal ¼ of the new species is reddish-brown, remainder is translucent white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |