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<document id="D3285AB7E0F9D670B165EB760490ADDB" ID-CLB-Dataset="6242" ID-DOI="10.11646/zootaxa.3802.2.4" ID-GBIF-Dataset="1338921a-4e48-4b52-9d48-72ae85d525a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915487" ID-ZooBank="C92CE0BD-2FFE-4B45-8DA7-37C300E8D042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623200198152" checkinUser="felipe" docAuthor="Winterbottom, Richard" docDate="2014" docId="03E2B921FFE5FFD8FF6FEDFC5F20FC9B" docLanguage="en" docName="zt03802p216.pdf" docOrigin="Zootaxa 3802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trimma irinae Winterbottom 2014, n. sp." docType="treatment" docVersion="7" lastPageNumber="215" masterDocId="FFDBC159FFE4FFDEFFF8E9315B32FF8E" masterDocTitle="Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea" masterLastPageNumber="216" masterPageNumber="209" pageNumber="210" updateTime="1699006540437" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2B71CF75D5DD882613A7810F88800B2E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CCBF8AB44B0B026A7EE587490447FBD0">
<mods:title id="427FCDDF9637CCAF9B9F341BAF2D3056">Trimma irinae, (Pisces; Gobioidei) a new species of gobiid fish from Milne Bay Province, Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="48F6F8647D29C8989BE787C8044B1A43" type="personal">
<mods:role id="4CA8A1B9971225A7B85E27C30400B18D">
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<mods:namePart id="0171249F168A7AB432F866831A75822D">Winterbottom, Richard</mods:namePart>
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<mods:title id="33CF4A968FFA11E81E8FD034B2851B36">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AFE72897B67875558456D6F04E6C6F42">
<mods:date id="BEC9066508E5A52CD75CDCB79834F778">2014</mods:date>
<mods:detail id="CABE0D09D8B2916E62A1A453D48DF5C4" type="pubDate">
<mods:number id="D0847B81984C92153BA620A0E756D25D">2014-05-26</mods:number>
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<mods:number id="ECDD156446E6E6198E24B085A994F4FA">3802</mods:number>
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<mods:classification id="798FD7A5510E6D063E6F473D5536510C">journal article</mods:classification>
<mods:identifier id="6766087A06E4E74201019E55EAF81307" type="CLB-Dataset">6242</mods:identifier>
<mods:identifier id="FD9C596B7F7082DCCF5F216069C8BD5E" type="DOI">10.11646/zootaxa.3802.2.4</mods:identifier>
<mods:identifier id="8B0A511EA95085EF54F8A7B5A8C362AC" type="GBIF-Dataset">1338921a-4e48-4b52-9d48-72ae85d525a7</mods:identifier>
<mods:identifier id="2C885E6673797BD7A176DC00D13E3E17" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B234A43AA9C08E184CD991382AA828C2" type="Zenodo-Dep">4915487</mods:identifier>
<mods:identifier id="6D4B0C7BB341381D0E216D1A4310ADE6" type="ZooBank">C92CE0BD-2FFE-4B45-8DA7-37C300E8D042</mods:identifier>
</mods:mods>
<treatment id="03E2B921FFE5FFD8FF6FEDFC5F20FC9B" ID-DOI="http://doi.org/10.5281/zenodo.4915463" ID-GBIF-Taxon="182639311" ID-Zenodo-Dep="4915463" LSID="urn:lsid:plazi:treatment:03E2B921FFE5FFD8FF6FEDFC5F20FC9B" httpUri="http://treatment.plazi.org/id/03E2B921FFE5FFD8FF6FEDFC5F20FC9B" lastPageId="6" lastPageNumber="215" pageId="1" pageNumber="210">
<subSubSection id="C3515BBCFFE5FFDFFF6FEDFC5AB6FB68" box="[151,388,1229,1254]" pageId="1" pageNumber="210" type="nomenclature">
<paragraph id="8BF40837FFE5FFDFFF6FEDFC5AB6FB68" blockId="1.[151,388,1229,1290]" box="[151,388,1229,1254]" pageId="1" pageNumber="210">
<heading id="D0BCBF5BFFE5FFDFFF6FEDFC5AB6FB68" bold="true" box="[151,388,1229,1254]" fontSize="10" level="2" pageId="1" pageNumber="210" reason="2">
<emphasis id="B93FD425FFE5FFDFFF6FEDFC5AB6FB68" bold="true" box="[151,388,1229,1254]" pageId="1" pageNumber="210">
<taxonomicName id="4C4B73B4FFE5FFDFFF6FEDFC5A0CFB68" ID-CoL="58V6B" authority="Winterbottom, 2014" authorityName="Winterbottom" authorityYear="2014" box="[151,318,1229,1254]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="irinae" status="sp. nov.">Trimma irinae</taxonomicName>
<taxonomicNameLabel id="A20C695EFFE5FFDFFEBDEDFC5AB6FB68" box="[325,388,1229,1254]" pageId="1" pageNumber="210" rank="species">
<emphasis id="B93FD425FFE5FFDFFEBDEDFC5A6EFB68" box="[325,348,1229,1254]" inLineHeading="true" pageId="1" pageNumber="210" reason="1">n.</emphasis>
sp.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE5FFDFFF6FEDC05992FADC" pageId="1" pageNumber="210" type="description">
<paragraph id="8BF40837FFE5FFDFFF6FEDC05A1CFA84" blockId="1.[151,388,1229,1290]" box="[151,302,1265,1290]" pageId="1" pageNumber="210">
(
<figureCitation id="137014B2FFE5FFDFFF66EDC05A17FA84" box="[158,293,1265,1290]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1778,1800]" captionStartId-1="3.[151,250,1881,1903]" captionStartId-2="4.[151,250,1077,1099]" captionStartId-3="4.[151,250,1833,1855]" captionStartId-4="5.[151,250,1823,1845]" captionTargetBox-0="[193,1393,844,1755]" captionTargetBox-1="[193,1393,841,1859]" captionTargetBox-2="[193,1393,177,1055]" captionTargetBox-3="[193,1393,1179,1811]" captionTargetBox-4="[193,1393,193,1802]" captionTargetId-0="figure-462@2.[193,1393,844,1755]" captionTargetId-1="figure-370@3.[193,1393,841,1859]" captionTargetId-2="figure-141@4.[193,1393,171,1055]" captionTargetId-3="figure-150@4.[193,1393,1179,1811]" captionTargetId-4="figure-106@5.[193,1393,193,1802]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="FIGURE 1. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Left lateral view of head to show cephalic sensory papillae (image reversed). Specimen stained with cyanine blue. Photo: R. Winterbottom." captionText-1="FIGURE 2. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Dorsal view of head to show cephalic sensory papillae. Specimen stained with cyanine blue. Photo: R. Winterbottom." captionText-2="FIGURE 3. Trimma irinae, 18.6 mm SL paratype, ROM 94506. Ventral view of head to show cephalic sensory papillae. Specimen not stained. Photo: R. Winterbottom." captionText-3="FIGURE 4. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Freshly collected, left lateral view. Photo: G. R. Allen." captionText-4="FIGURE 5. Trimma irinae. A and B: Live specimens. Photos: M. V. Erdmann." figureDoi-0="http://doi.org/10.5281/zenodo.4915489" figureDoi-1="http://doi.org/10.5281/zenodo.4915491" figureDoi-2="http://doi.org/10.5281/zenodo.4915493" figureDoi-3="http://doi.org/10.5281/zenodo.4915495" figureDoi-4="http://doi.org/10.5281/zenodo.4915497" httpUri-0="https://zenodo.org/record/4915489/files/figure.png" httpUri-1="https://zenodo.org/record/4915491/files/figure.png" httpUri-2="https://zenodo.org/record/4915493/files/figure.png" httpUri-3="https://zenodo.org/record/4915495/files/figure.png" httpUri-4="https://zenodo.org/record/4915497/files/figure.png" pageId="1" pageNumber="210">Figures 1–5</figureCitation>
)
</paragraph>
<paragraph id="8BF40837FFE5FFDFFF6FEC085992FADC" blockId="1.[151,1436,1337,2010]" box="[151,672,1337,1362]" pageId="1" pageNumber="210">Suggested common name: Irina’s pygmy goby</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE5FFDFFF3FEC6C5A32F9C0" pageId="1" pageNumber="210" type="materials_examined">
<paragraph id="8BF40837FFE5FFDFFF3FEC6C5E40FA30" blockId="1.[151,1436,1337,2010]" pageId="1" pageNumber="210">
<emphasis id="B93FD425FFE5FFDFFF3FEC6C5A8AFAF8" bold="true" box="[199,440,1373,1398]" pageId="1" pageNumber="210">Material Examined.</emphasis>
<materialsCitation id="3B23026AFFE5FFDFFE47EC6C5E5CFA30" ID-GBIF-Occurrence="3127814302" accessionNumber="T12776" collectionCode="T" country="Papua New Guinea" location="The" pageId="1" pageNumber="210" specimenCount="3" stateProvince="Milne Bay">
One lot,
<specimenCount id="9D4DC3BEFFE5FFDFFDDBEC6C599CFAF8" box="[547,686,1373,1398]" pageId="1" pageNumber="210" type="generic">3 specimens</specimenCount>
(
<quantity id="4CB3A5D2FFE5FFDFFD45EC6C5850FAFB" box="[701,866,1373,1398]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.97" metricValueMin="1.43" pageId="1" pageNumber="210" unit="mm" value="17.0" valueMax="19.7" valueMin="14.3">14.3–19.7 mm</quantity>
SL, all females) from
<collectingRegion id="498FC6D5FFE5FFDFFB9DEC6C5E79FAF8" box="[1125,1355,1373,1398]" country="Papua New Guinea" name="Milne Bay" pageId="1" pageNumber="210">Milne Bay Province</collectingRegion>
,
<collectingCountry id="F35C48A7FFE5FFDFFAA0EC6F5A1FFA14" name="Papua New Guinea" pageId="1" pageNumber="210">Papua New Guinea</collectingCountry>
.
<location id="8E945EECFFE5FFDFFEC5ECB05A5BFA14" LSID="urn:lsid:plazi:treatment:03E2B921FFE5FFD8FF6FEDFC5F20FC9B:8E945EECFFE5FFDFFEC5ECB05A5BFA14" box="[317,361,1409,1434]" country="Papua New Guinea" name="The" pageId="1" pageNumber="210" stateProvince="Milne Bay">The</location>
description is based primarily on the two formalin-fixed specimens, with some additional information from the tissue specimen (
<collectionCode id="ED5A90F2FFE5FFDFFDB0EC945968FA30" box="[584,602,1445,1470]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="210">T</collectionCode>
12776, originally preserved in 95% ethanol, genetic analysis pending)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF40837FFE5FFDFFF3FECF85F20F988" blockId="1.[151,1436,1337,2010]" pageId="1" pageNumber="210">
<materialsCitation id="3B23026AFFE5FFDFFF3FECF85F20F988" ID-GBIF-Occurrence="3127814303" accessionNumber="ROM 94505" collectionCode="ROM, V" county="East Cape" elevation="65" latitude="-10.256583" location="July" longLatPrecision="1" longitude="150.69958" municipality="Lawadi" pageId="1" pageNumber="210" specimenCount="1" stateProvince="Milne Bay" typeStatus="holotype">
<emphasis id="B93FD425FFE5FFDFFF3FECF85A09FA6C" bold="true" box="[199,315,1481,1506]" inLineHeading="true" pageId="1" pageNumber="210" reason="1">
<typeStatus id="54F0B695FFE5FFDFFF3FECF85A04FA6C" box="[199,310,1481,1506]" pageId="1" pageNumber="210" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED5A90F2FFE5FFDFFEBBECF85AB1FA6C" box="[323,387,1481,1506]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="210" type="Museum">ROM</collectionCode>
94505,
<quantity id="4CB3A5D2FFE5FFDFFE18ECF85977FA6F" box="[480,581,1481,1506]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="1" pageNumber="210" unit="mm" value="19.7">19.7 mm</quantity>
SL female,
<collectingRegion id="498FC6D5FFE5FFDFFD28ECF85885FA6C" box="[720,951,1481,1506]" country="Papua New Guinea" name="Milne Bay" pageId="1" pageNumber="210">Milne Bay Province</collectingRegion>
, on northern shore of
<collectingCounty id="629570BBFFE5FFDFFB4AECFB5E16FA6C" box="[1202,1316,1481,1506]" pageId="1" pageNumber="210">East Cape</collectingCounty>
at
<collectingMunicipality id="6B90924DFFE5FFDFFAB0ECF85EAEFA6C" box="[1352,1436,1481,1506]" pageId="1" pageNumber="210">Lawadi</collectingMunicipality>
(
<geoCoordinate id="EE7F6EF0FFE5FFDFFF58ECDC5A0FF988" box="[160,317,1517,1542]" degrees="10" direction="south" minutes="15.395" orientation="latitude" pageId="1" pageNumber="210" precision="1" value="-10.256583">10° 15.395’ S</geoCoordinate>
,
<geoCoordinate id="EE7F6EF0FFE5FFDFFEB1ECDC5AC2F988" box="[329,496,1517,1542]" degrees="150" direction="east" minutes="41.975" orientation="longitude" pageId="1" pageNumber="210" precision="1" value="150.69958">150° 41.975’E</geoCoordinate>
),
<quantity id="4CB3A5D2FFE5FFDFFDFCECDC590DF98B" box="[516,575,1517,1542]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="210" unit="m" value="65.0">
<elevation id="0066EF04FFE5FFDFFDFCECDC590DF98B" box="[516,575,1517,1542]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="210" unit="m" value="65.0">65 m</elevation>
</quantity>
, clove oil. 15
<location id="8E945EECFFE5FFDFFD25ECDC583FF988" LSID="urn:lsid:plazi:treatment:03E2B921FFE5FFD8FF6FEDFC5F20FC9B:8E945EECFFE5FFDFFD25ECDC583FF988" box="[733,781,1517,1542]" county="East Cape" latitude="-10.256583" longLatPrecision="1" longitude="150.69958" municipality="Lawadi" name="July" pageId="1" pageNumber="210" stateProvince="Milne Bay">July</location>
, 2013, M.
<collectionCode id="ED5A90F2FFE5FFDFFC7DECDF58ABF98B" box="[901,921,1518,1541]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="210" type="Museum">V</collectionCode>
. Erdmann.
</materialsCitation>
</paragraph>
<paragraph id="8BF40837FFE5FFDFFF3FEF205A32F9C0" blockId="1.[151,1436,1337,2010]" pageId="1" pageNumber="210">
<materialsCitation id="3B23026AFFE5FFDFFF3FEF205827F9A4" ID-GBIF-Occurrence="3127814301" accessionNumber="ROM 94506" box="[199,789,1553,1578]" collectionCode="ROM" pageId="1" pageNumber="210" specimenCount="1" typeStatus="paratype">
<emphasis id="B93FD425FFE5FFDFFF3FEF205A74F9A4" bold="true" box="[199,326,1553,1578]" inLineHeading="true" pageId="1" pageNumber="210" reason="1">
<typeStatus id="54F0B695FFE5FFDFFF3FEF205A73F9A4" box="[199,321,1553,1578]" pageId="1" pageNumber="210" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED5A90F2FFE5FFDFFEA8EF205AA2F9A4" box="[336,400,1553,1578]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="210" type="Museum">ROM</collectionCode>
94506,
<quantity id="4CB3A5D2FFE5FFDFFE0AEF205968F9A7" box="[498,602,1553,1578]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="1" pageNumber="210" unit="mm" value="18.6">18.6 mm</quantity>
SL female, and
</materialsCitation>
<materialsCitation id="3B23026AFFE5FFDFFCD8EF205BCFF9C0" ID-GBIF-Occurrence="3127814304" accessionNumber="T12776" collectionCode="ROM, T" pageId="1" pageNumber="210" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED5A90F2FFE5FFDFFCD8EF205852F9A4" box="[800,864,1553,1578]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="210" type="Museum">ROM</collectionCode>
<collectionCode id="ED5A90F2FFE5FFDFFC92EF20584EF9A4" box="[874,892,1553,1578]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="210">T</collectionCode>
12776,
<quantity id="4CB3A5D2FFE5FFDFFC2BEF205F09F9A7" box="[979,1083,1553,1578]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="210" unit="mm" value="14.3">14.3 mm</quantity>
SL female, collected with the
<typeStatus id="54F0B695FFE5FFDFFF6FEF045BCFF9C0" box="[151,253,1589,1614]" pageId="1" pageNumber="210" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE5FFDFFF3FEF685915F8A8" pageId="1" pageNumber="210" type="diagnosis">
<paragraph id="8BF40837FFE5FFDFFF3FEF685915F8A8" blockId="1.[151,1436,1337,2010]" pageId="1" pageNumber="210">
<emphasis id="B93FD425FFE5FFDFFF3FEF685A73F9FC" bold="true" box="[199,321,1625,1650]" inLineHeading="true" pageId="1" pageNumber="210" reason="1">Diagnosis.</emphasis>
<taxonomicName id="4C4B73B4FFE5FFDFFEB4EF6B5AC0F9FF" authorityName="Winterbottom" authorityYear="2014" box="[332,498,1626,1649]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="210" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE5FFDFFEB4EF6B5AC0F9FF" box="[332,498,1626,1649]" italics="true" pageId="1" pageNumber="210">Trimma irinae</emphasis>
</taxonomicName>
has 8–9 scales in the predorsal midline, usually cycloid scales along the upper border of the opercle which may be in up to three horizontal rows, no cheek scales, a very elongate second dorsal spine reaching posteriorly to the end of the second dorsal fin or beyond this (in specimens&gt;
<quantity id="4CB3A5D2FFE5FFDFFB61EF905FD5F937" box="[1177,1255,1697,1722]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="210" unit="mm" value="18.0">18 mm</quantity>
SL), the middle rays of the pectoral fin branched, a fifth pelvic fin ray which branches once dichotomously and is 52–64% the length of the fourth ray, and the basal membrane connecting the inner margins of the fifth pelvic fins rays is less than 15% the length of the fifth ray.
</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE5FFD8FF3FEE005FCFFDEE" lastPageId="6" lastPageNumber="215" pageId="1" pageNumber="210" type="description">
<paragraph id="8BF40837FFE5FFDCFF3FEE0058C4FC9B" blockId="1.[151,1436,1337,2010]" lastBlockId="2.[151,1436,151,789]" lastPageId="2" lastPageNumber="211" pageId="1" pageNumber="210">
<emphasis id="B93FD425FFE5FFDFFF3FEE005A6BF8C4" bold="true" box="[199,345,1841,1866]" inLineHeading="true" pageId="1" pageNumber="210" reason="1">Description.</emphasis>
Dorsal fins VI + I 9, second spine elongated, reaching posteriorly to between the base of ray 3 of second dorsal fin (
<quantity id="4CB3A5D2FFE5FFDFFE8AEE645AEEF8E3" box="[370,476,1877,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="210" unit="mm" value="14.3">14.3 mm</quantity>
SL) to beyond the base of the last ray when adpressed, rays all branched except posterior element of last ray, last ray reaching posteriorly
<emphasis id="B93FD425FFE5FFDFFCAFEE485841F81C" bold="true" box="[855,883,1913,1938]" pageId="1" pageNumber="210">49</emphasis>
–65% of distance from its base to first dorsal procurrent ray; anal fin I 8, all but first and posterior element of posteriormost ray branched; posteriormost ray
<emphasis id="B93FD425FFE5FFDFFF6FEEF05B81F854" bold="true" box="[151,179,1985,2010]" pageId="1" pageNumber="210">50</emphasis>
–54% of distance from its base to first ventral procurrent ray; pectoral fin 17, 3 and 3–
<emphasis id="B93FD425FFE5FFDFFB7FEEF05FA7F854" bold="true" box="[1159,1173,1985,2010]" pageId="1" pageNumber="210">5</emphasis>
unbranched dorsal and ventral rays respectively, middle
<emphasis id="B93FD425FFE6FFDCFDF4E9A65928FF3E" bold="true" box="[524,538,151,176]" pageId="2" pageNumber="211">9</emphasis>
–11 rays branched, fin reaching posteriorly to a vertical line between urogenital papilla and anal-fin origin; pelvic fin I 5, no frenum, basal membrane reduced and
<emphasis id="B93FD425FFE6FFDCFBCFE98D5F61FF5B" bold="true" box="[1079,1107,188,213]" pageId="2" pageNumber="211">10</emphasis>
–13% length of fifth ray, first four rays with one sequential branch, fifth ray branched once dichotomously and 52–
<emphasis id="B93FD425FFE6FFDCFB8FE9EE5FA1FF76" bold="true" box="[1143,1171,223,248]" pageId="2" pageNumber="211">55</emphasis>
–64% length of fourth, fourth ray reaching posteriorly to between bases of anal rays
<emphasis id="B93FD425FFE6FFDCFCAEE8355856FE93" bold="true" box="[854,868,260,285]" pageId="2" pageNumber="211">3</emphasis>
–4. Lateral scales 23; anterior transverse scales 9; posterior transverse scales 8; predorsal scales
<emphasis id="B93FD425FFE6FFDCFD5EE8165986FECE" bold="true" box="[678,692,295,320]" pageId="2" pageNumber="211">8</emphasis>
–9; no scales on cheek; opercle apparently with 2–
<emphasis id="B93FD425FFE6FFDCFB03E8165E3BFECE" bold="true" box="[1275,1289,295,320]" pageId="2" pageNumber="211">3</emphasis>
rows of 2–
<emphasis id="B93FD425FFE6FFDCFA76E8165EAEFECE" bold="true" box="[1422,1436,295,320]" pageId="2" pageNumber="211">3</emphasis>
usually cycloid scales (scales mostly missing); pectoral base with 3 vertical rows of scales with 4 cycloid scales in posterior row; 6–
<emphasis id="B93FD425FFE6FFDCFEA2E85E5A5AFE06" bold="true" box="[346,360,367,392]" pageId="2" pageNumber="211">7</emphasis>
cycloid prepelvic scales (in midline anterior to basal membrane); 12 circumpeduncular scales; body scales ctenoid except for cycloid scales on anterior belly midline; and on, beneath and just posterior to pectoral fin base; body scales extend anteriorly to just behind eye. Gill opening extending anteroventrally to a vertical below mid- pupil. Teeth not examined in detail due to the paucity of specimens, but appear to be typical of other
<taxonomicName id="4C4B73B4FFE6FFDCFF20EB315A1DFD99" box="[216,303,512,535]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="211" phylum="Chordata" rank="genus">
<emphasis id="B93FD425FFE6FFDCFF20EB315A1DFD99" box="[216,303,512,535]" italics="true" pageId="2" pageNumber="211">Trimma</emphasis>
</taxonomicName>
, although apparently less robust. Cephalic sensory papillae (
<figureCitation id="137014B2FFE6FFDCFC26E8CE5F7AFD96" box="[990,1096,511,536]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1778,1800]" captionStartId-1="3.[151,250,1881,1903]" captionStartId-2="4.[151,250,1077,1099]" captionTargetBox-0="[193,1393,844,1755]" captionTargetBox-1="[193,1393,841,1859]" captionTargetBox-2="[193,1393,177,1055]" captionTargetId-0="figure-462@2.[193,1393,844,1755]" captionTargetId-1="figure-370@3.[193,1393,841,1859]" captionTargetId-2="figure-141@4.[193,1393,171,1055]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Left lateral view of head to show cephalic sensory papillae (image reversed). Specimen stained with cyanine blue. Photo: R. Winterbottom." captionText-1="FIGURE 2. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Dorsal view of head to show cephalic sensory papillae. Specimen stained with cyanine blue. Photo: R. Winterbottom." captionText-2="FIGURE 3. Trimma irinae, 18.6 mm SL paratype, ROM 94506. Ventral view of head to show cephalic sensory papillae. Specimen not stained. Photo: R. Winterbottom." figureDoi-0="http://doi.org/10.5281/zenodo.4915489" figureDoi-1="http://doi.org/10.5281/zenodo.4915491" figureDoi-2="http://doi.org/10.5281/zenodo.4915493" httpUri-0="https://zenodo.org/record/4915489/files/figure.png" httpUri-1="https://zenodo.org/record/4915491/files/figure.png" httpUri-2="https://zenodo.org/record/4915493/files/figure.png" pageId="2" pageNumber="211">Figs. 1–3</figureCitation>
) row counts given in
<tableCitation id="C6C93D8CFFE6FFDCFAB9E8CE5EA4FD96" box="[1345,1430,511,536]" captionStart="TABLE 1" captionStartId="1.[151,239,152,175]" captionTargetPageId="1" captionText="TABLE 1. Number of cephalic sensory papillae in each row in Trimma irinae. Nomenclature follows that given in Winterbottom (2011)." httpUri="http://table.plazi.org/id/DF3458BFFFE5FFDFFF6FE9A95A4DFF41" pageId="2" pageNumber="211" tableUuid="DF3458BFFFE5FFDFFF6FE9A95A4DFF41">Table 1</tableCitation>
. Tongue truncate and bilobed, with rounded edges. Gill rakers on first arch 4 +
<emphasis id="B93FD425FFE6FFDCFBCFEB155F61FDB3" bold="true" box="[1079,1107,548,573]" pageId="2" pageNumber="211">15</emphasis>
–16 =
<emphasis id="B93FD425FFE6FFDCFB59EB155F8FFDB3" bold="true" box="[1185,1213,548,573]" pageId="2" pageNumber="211">19</emphasis>
–20. Anterior nasal opening a short tube extending out over upper lip, posterior nasal opening a pore with a raised rim, both protruding from slightly raised oval sac confined to anterior half of snout. Bony interorbital
<emphasis id="B93FD425FFE6FFDCFBE6EB5D5F08FD0B" bold="true" box="[1054,1082,620,645]" pageId="2" pageNumber="211">38</emphasis>
–42% pupil width, with a broad moderately developed U-shaped interorbital furrow and no postorbital groove. No ridge of tissue (or dermal crest) extending anterior to first dorsal spine. Epaxialis extending anteriorly to above a vertical with posterior margin of pupil. Vertebral transition (as defined by
<bibRefCitation id="EFDA75C6FFE6FFDCFD9FEBE658B1FD7E" author="Winterbottom, R." box="[615,899,727,752]" pageId="2" pageNumber="211" pagination="127 - 162" refId="ref3388" refString="Winterbottom, R. (2011) Six new species of the genus Trimma (Percomorpha; Gobiidae) from the Raja Ampat Islands, Indonesia. aqua, International Journal of Ichthyology, 17 (3), 127 - 162." type="journal article" year="2011">Winterbottom, 2011: 130</bibRefCitation>
) not examined, but predicted to be
<typeStatus id="54F0B695FFE6FFDCFAEDEBE95E7FFD7E" box="[1301,1357,728,752]" pageId="2" pageNumber="211">Type</typeStatus>
B (see Winterbottom, 2003, Fig. 14, inset, and
<bibRefCitation id="EFDA75C6FFE6FFDCFDAFEBCD58B5FC9B" author="Winterbottom, R. &amp; Zur, M." box="[599,903,764,789]" pageId="2" pageNumber="211" pagination="13 - 24" refId="ref3565" refString="Winterbottom, R. &amp; Zur, M. (2007) Three new species of the Trimma from Palau, Western Pacific (Percomorpha; Gobiidae). aqua, International Journal of Ichthyology, 13 (1), 13 - 24." type="journal article" year="2007">Winterbottom &amp; Zur, 2007</bibRefCitation>
,
<figureCitation id="137014B2FFE6FFDCFC69EBCD58DAFC9B" box="[913,1000,764,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1077,1099]" captionTargetBox="[193,1393,177,1055]" captionTargetId="figure-141@4.[193,1393,171,1055]" captionTargetPageId="4" captionText="FIGURE 3. Trimma irinae, 18.6 mm SL paratype, ROM 94506. Ventral view of head to show cephalic sensory papillae. Specimen not stained. Photo: R. Winterbottom." figureDoi="http://doi.org/10.5281/zenodo.4915493" httpUri="https://zenodo.org/record/4915493/files/figure.png" pageId="2" pageNumber="211">Fig. 3B</figureCitation>
).
</paragraph>
<caption id="DF3458BFFFE6FFDCFF6FEFC358F3F8A9" ID-DOI="http://doi.org/10.5281/zenodo.4915489" ID-Zenodo-Dep="4915489" httpUri="https://zenodo.org/record/4915489/files/figure.png" pageId="2" pageNumber="211" startId="2.[151,250,1778,1800]" targetBox="[193,1393,844,1755]" targetPageId="2">
<paragraph id="8BF40837FFE6FFDCFF6FEFC358F3F8A9" blockId="2.[151,1436,1778,1831]" pageId="2" pageNumber="211">
<emphasis id="B93FD425FFE6FFDCFF6FEFC35A26F889" bold="true" box="[151,276,1778,1800]" pageId="2" pageNumber="211">FIGURE 1.</emphasis>
<taxonomicName id="4C4B73B4FFE6FFDCFEE4EFC25A9CF886" authorityName="Winterbottom" authorityYear="2014" box="[284,430,1779,1800]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="211" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE6FFDCFEE4EFC25A9CF886" box="[284,430,1779,1800]" italics="true" pageId="2" pageNumber="211">Trimma irinae</emphasis>
</taxonomicName>
, 19.7 mm SL holotype, ROM 94505. Left lateral view of head to show cephalic sensory papillae (image reversed). Specimen stained with cyanine blue. Photo: R. Winterbottom.
</paragraph>
</caption>
<paragraph id="8BF40837FFE6FFDDFF3FEE7F5BE7FC9B" blockId="2.[151,1436,1870,2003]" lastBlockId="3.[151,1437,151,789]" lastPageId="3" lastPageNumber="212" pageId="2" pageNumber="211">
Colour pattern (live and freshly collected, from a total of four images of at least
<specimenCount id="9D4DC3BEFFE6FFDCFBB1EE7E5FC3F8E9" box="[1097,1265,1870,1895]" pageId="2" pageNumber="211" type="generic">two specimens</specimenCount>
,
<figureCitation id="137014B2FFE6FFDCFB04EE7F5E78F8E9" box="[1276,1354,1870,1895]" captionStart="FIGURE 4" captionStartId="4.[151,250,1833,1855]" captionTargetBox="[193,1393,1179,1811]" captionTargetId="figure-150@4.[193,1393,1179,1811]" captionTargetPageId="4" captionText="FIGURE 4. Trimma irinae, 19.7 mm SL holotype, ROM 94505. Freshly collected, left lateral view. Photo: G. R. Allen." figureDoi="http://doi.org/10.5281/zenodo.4915495" httpUri="https://zenodo.org/record/4915495/files/figure.png" pageId="2" pageNumber="211">Figs. 4</figureCitation>
,
<figureCitation id="137014B2FFE6FFDCFAAEEE7F5B9EF805" captionStart="FIGURE 5" captionStartId="5.[151,250,1823,1845]" captionTargetBox="[193,1393,193,1802]" captionTargetId="figure-106@5.[193,1393,193,1802]" captionTargetPageId="5" captionText="FIGURE 5. Trimma irinae. A and B: Live specimens. Photos: M. V. Erdmann." figureDoi="http://doi.org/10.5281/zenodo.4915497" httpUri="https://zenodo.org/record/4915497/files/figure.png" pageId="2" pageNumber="211">5 A &amp; B</figureCitation>
). Background colour of head and anterior half of body pinkish-grey, grading to translucent orange at level of origin of second dorsal fin. Snout and cheek immaculate. Numerous orange to red spots, often with diffuse darker centers and pupil-diameter in width, on top of head, nape, opercular region and anterior half of body. Spots separated by narrow interstices which are frequently invested heavily with iridocytes. Up to three such spots on vertical limb of preopercle, which may coalesce to form an irregularly outlined bar. Below this, a dark red halfpupil diameter sized spot on branchiostegal membrane. Interorbital region crossed by up to three thin transverse white lines. Spots on body decrease somewhat in size posteriorly, and grade into irregular rows of diffuse yellow spots which may be dorso-ventrally elongate behind second dorsal fin. First dorsal fin with two dark, basal markings, an oval-shaped one about eye diameter in width between shafts of the first to fourth spines, and second smaller rounded spot centered on shaft of sixth spine. Both are dark red to black, ocellated with lighter red to orange. A similar, but much smaller rounded spot centered on shaft of fourth spine about two-thirds of the way up shaft, and this may be preceded by a thin, diffuse orange stripe. Membrane of fin a mix of melanophores and iridocytes (and may appear to be white –
<figureCitation id="137014B2FFE7FFDDFD9CE8ED59F3FE7B" box="[612,705,476,501]" captionStart="FIGURE 5" captionStartId="5.[151,250,1823,1845]" captionTargetBox="[193,1393,193,1802]" captionTargetId="figure-106@5.[193,1393,193,1802]" captionTargetPageId="5" captionText="FIGURE 5. Trimma irinae. A and B: Live specimens. Photos: M. V. Erdmann." figureDoi="http://doi.org/10.5281/zenodo.4915497" httpUri="https://zenodo.org/record/4915497/files/figure.png" pageId="3" pageNumber="212">Fig. 5 B</figureCitation>
). Tip of first spine and main shaft of second spine bright white or pale blue. Second dorsal fin with two small basal ocellated spots similar to the one on fourth spine situated between rays 1 and 2, and on shaft of ray 5, and, in middle portion of fin, there may be an irregular series of small yellow spots centered on some of fin rays. Distal part of fin with a flush of yellow, whole membrane with scattered iridocytes. Caudal fin rays mostly reddish-orange with scattered small yellow spots. Anal fin rays pale reddishyellow with some diffuse yellow spots, fin membranes with iridocytes and dark pigment (unclear whether these are melanophores or very dark chromatophores). Pectoral and pelvic fin rays pale pinkish red, membranes hyaline. A specimen photographed just after collection is similar to above, but scattered iridocytes are visible over most of body.
</paragraph>
<caption id="DF3458BFFFE7FFDDFF6FEE685838F803" ID-DOI="http://doi.org/10.5281/zenodo.4915491" ID-Zenodo-Dep="4915491" httpUri="https://zenodo.org/record/4915491/files/figure.png" pageId="3" pageNumber="212" startId="3.[151,250,1881,1903]" targetBox="[193,1393,841,1859]" targetPageId="3">
<paragraph id="8BF40837FFE7FFDDFF6FEE685838F803" blockId="3.[151,1437,1881,1933]" pageId="3" pageNumber="212">
<emphasis id="B93FD425FFE7FFDDFF6FEE685A24F8E0" bold="true" box="[151,278,1881,1903]" pageId="3" pageNumber="212">FIGURE 2.</emphasis>
<taxonomicName id="4C4B73B4FFE7FFDDFED8EE6B5A87F8E1" authorityName="Winterbottom" authorityYear="2014" box="[288,437,1882,1903]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="212" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE7FFDDFED8EE6B5A87F8E1" box="[288,437,1882,1903]" italics="true" pageId="3" pageNumber="212">Trimma irinae</emphasis>
</taxonomicName>
, 19.7 mm SL holotype, ROM 94505. Dorsal view of head to show cephalic sensory papillae. Specimen stained with cyanine blue. Photo: R. Winterbottom.
</paragraph>
</caption>
<caption id="DF3458BFFFE0FFDAFF6FED045944FBE4" ID-DOI="http://doi.org/10.5281/zenodo.4915493" ID-Zenodo-Dep="4915493" httpUri="https://zenodo.org/record/4915493/files/figure.png" pageId="4" pageNumber="213" startId="4.[151,250,1077,1099]" targetBox="[193,1393,177,1055]" targetPageId="4">
<paragraph id="8BF40837FFE0FFDAFF6FED045944FBE4" blockId="4.[151,1437,1077,1130]" pageId="4" pageNumber="213">
<emphasis id="B93FD425FFE0FFDAFF6FED045A24FBC4" bold="true" box="[151,278,1077,1099]" pageId="4" pageNumber="213">FIGURE 3.</emphasis>
<taxonomicName id="4C4B73B4FFE0FFDAFED8ED075A86FBC5" authorityName="Winterbottom" authorityYear="2014" box="[288,436,1078,1099]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="213" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE0FFDAFED8ED075A86FBC5" box="[288,436,1078,1099]" italics="true" pageId="4" pageNumber="213">Trimma irinae</emphasis>
</taxonomicName>
, 18.6 mm SL paratype, ROM 94506. Ventral view of head to show cephalic sensory papillae. Specimen not stained. Photo: R. Winterbottom.
</paragraph>
</caption>
<caption id="DF3458BFFFE0FFDAFF6FEE185E63F8B1" ID-DOI="http://doi.org/10.5281/zenodo.4915495" ID-Zenodo-Dep="4915495" box="[151,1361,1833,1855]" httpUri="https://zenodo.org/record/4915495/files/figure.png" pageId="4" pageNumber="213" startId="4.[151,250,1833,1855]" targetBox="[193,1393,1179,1811]" targetPageId="4">
<paragraph id="8BF40837FFE0FFDAFF6FEE185E63F8B1" blockId="4.[151,1361,1833,1855]" box="[151,1361,1833,1855]" pageId="4" pageNumber="213">
<emphasis id="B93FD425FFE0FFDAFF6FEE185A20F8B0" bold="true" box="[151,274,1833,1855]" pageId="4" pageNumber="213">FIGURE 4.</emphasis>
<taxonomicName id="4C4B73B4FFE0FFDAFEE1EE1B5A98F8B1" authorityName="Winterbottom" authorityYear="2014" box="[281,426,1834,1855]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="213" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE0FFDAFEE1EE1B5A98F8B1" box="[281,426,1834,1855]" italics="true" pageId="4" pageNumber="213">Trimma irinae</emphasis>
</taxonomicName>
, 19.7 mm SL holotype, ROM 94505. Freshly collected, left lateral view. Photo: G. R. Allen.
</paragraph>
</caption>
<paragraph id="8BF40837FFE0FFDAFF3FEE5D5E02F87F" blockId="4.[151,1436,1900,2033]" pageId="4" pageNumber="213">
Colour pattern in alcohol: Background colour pale straw-yellow, all traces or red, orange, yellow and pink absent. Centres of head and anterior body spots with numerous dark pigment cells, much fewer at periphery of spots, thin white network separating spots in live fishes now consists of lines of brown chromatophores. Black spots in dorsal fins remain black, dark spot on branchiostegal membrane is also black (
<figureCitation id="137014B2FFE0FFDAFB9BEEE95F95F87F" box="[1123,1191,2008,2033]" captionStart="FIGURE 3" captionStartId="4.[151,250,1077,1099]" captionTargetBox="[193,1393,177,1055]" captionTargetId="figure-141@4.[193,1393,171,1055]" captionTargetPageId="4" captionText="FIGURE 3. Trimma irinae, 18.6 mm SL paratype, ROM 94506. Ventral view of head to show cephalic sensory papillae. Specimen not stained. Photo: R. Winterbottom." figureDoi="http://doi.org/10.5281/zenodo.4915493" httpUri="https://zenodo.org/record/4915493/files/figure.png" pageId="4" pageNumber="213">Fig. 3</figureCitation>
, red arrow).
</paragraph>
<caption id="DF3458BFFFE1FFDBFF6FEE2E588BF8BB" ID-DOI="http://doi.org/10.5281/zenodo.4915497" ID-Zenodo-Dep="4915497" box="[151,953,1823,1845]" httpUri="https://zenodo.org/record/4915497/files/figure.png" pageId="5" pageNumber="214" startId="5.[151,250,1823,1845]" targetBox="[193,1393,193,1802]" targetPageId="5">
<paragraph id="8BF40837FFE1FFDBFF6FEE2E588BF8BB" blockId="5.[151,953,1823,1845]" box="[151,953,1823,1845]" pageId="5" pageNumber="214">
<emphasis id="B93FD425FFE1FFDBFF6FEE2E5A20F8BB" bold="true" box="[151,274,1823,1845]" pageId="5" pageNumber="214">FIGURE 5.</emphasis>
<taxonomicName id="4C4B73B4FFE1FFDBFEE1EE115A98F8BB" authorityName="Winterbottom" authorityYear="2014" box="[281,426,1824,1845]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE1FFDBFEE1EE115A98F8BB" box="[281,426,1824,1845]" italics="true" pageId="5" pageNumber="214">Trimma irinae</emphasis>
</taxonomicName>
. A and B: Live specimens. Photos: M. V. Erdmann.
</paragraph>
</caption>
<paragraph id="8BF40837FFE1FFD8FF3FEE535FCFFDEE" blockId="5.[151,1438,1890,2023]" lastBlockId="6.[151,1438,151,789]" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214">
<emphasis id="B93FD425FFE1FFDBFF3FEE535A5FF8F5" bold="true" box="[199,365,1890,1915]" inLineHeading="true" pageId="5" pageNumber="214" reason="1">Comparisons.</emphasis>
<taxonomicName id="4C4B73B4FFE1FFDBFE8DEE52592AF8F4" authorityName="Winterbottom" authorityYear="2014" box="[373,536,1891,1914]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE1FFDBFE8DEE52592AF8F4" box="[373,536,1891,1914]" italics="true" pageId="5" pageNumber="214">Trimma irinae</emphasis>
</taxonomicName>
belongs to a group of nine described species which all have a scaled predorsal midline, no cheek scales, a scaled opercle, and at least the middle rays of the pectoral fin branched (
<bibRefCitation id="EFDA75C6FFE1FFDBFB0FEEB75BE6F84D" author="Winterbottom, R." pageId="5" pageNumber="214" pagination="55 - 68" refId="ref3349" refString="Winterbottom, R. (2006) Two new species of the gobiid fish Trimma from the coral reefs ofthe western Pacific Ocean (Pisces; Perciformes; Gobioidei). Zootaxa, 1331, 55 - 68." type="journal article" year="2006">Winterbottom, 2006</bibRefCitation>
). Within this group,
<taxonomicName id="4C4B73B4FFE1FFDBFE30EE9A59B9F84C" authorityName="Jordan &amp; Seale" authorityYear="1906" box="[456,651,1963,1986]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="caesiura">
<emphasis id="B93FD425FFE1FFDBFE30EE9A59B9F84C" box="[456,651,1963,1986]" italics="true" pageId="5" pageNumber="214">Trimma caesiura</emphasis>
</taxonomicName>
<bibRefCitation id="EFDA75C6FFE1FFDBFD6DEE9B58A7F84D" author="Jordan, D. S. &amp; Seale, A." box="[661,917,1962,1987]" pageId="5" pageNumber="214" pagination="173 - 455" refId="ref3096" refString="Jordan, D. S. &amp; Seale, A. (1906) The fishes of Samoa. Description of the species found in the archipelago, with a provisional check-list of the fishes of Oceania. Bulletin of the Bureau of Fisheries, 25 (for 1905), 173 - 455 + index 457 - 488, pls. 33 - 53." type="journal article" year="1906">
<collectingCountry id="F35C48A7FFE1FFDBFD6DEE9B59D2F84D" box="[661,736,1962,1987]" name="Jordan" pageId="5" pageNumber="214">Jordan</collectingCountry>
&amp; Seale, 1906
</bibRefCitation>
,
<taxonomicName id="4C4B73B4FFE1FFDBFC5CEE9A5E32F84D" authority="Winterbottom, 1984" authorityName="Winterbottom" authorityYear="1984" box="[932,1280,1962,1987]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="fraena">
<emphasis id="B93FD425FFE1FFDBFC5CEE9A5F3FF84D" box="[932,1037,1962,1987]" italics="true" pageId="5" pageNumber="214">T. fraena</emphasis>
<bibRefCitation id="EFDA75C6FFE1FFDBFBE0EE9B5E32F84D" author="Winterbottom, R." box="[1048,1280,1962,1987]" pageId="5" pageNumber="214" pagination="695 - 715" refId="ref3211" refString="Winterbottom, R. (1984) A review of the gobiid fish genus Trimma from the Chagos Archipelago, Central Indian Ocean, with the description of seven new species. Canadian Journal of Zoology, 62, 695 - 715. http: // dx. doi. org / 10.1139 / z 84 - 101" type="journal article" year="1984">Winterbottom, 1984</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C4B73B4FFE1FFDBFABBEE9A5A4BF869" authority="Winterbottom, 2002" authorityName="Winterbottom" authorityYear="2002" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="milta">
<emphasis id="B93FD425FFE1FFDBFABBEE9A5EAEF84D" box="[1347,1436,1962,1987]" italics="true" pageId="5" pageNumber="214">T. milta</emphasis>
<bibRefCitation id="EFDA75C6FFE1FFDBFF6FEEFF5A4BF869" author="Winterbottom, R." box="[151,377,1998,2023]" pageId="5" pageNumber="214" pagination="45 - 52" refId="ref3269" refString="Winterbottom, R. (2002) Two new species of Trimma (Gobiidae) from the central, western, and south Pacific. aqua, Journal of Ichthyology and Aquatic Biology, 5 (2), 45 - 52." type="journal article" year="2002">Winterbottom, 2002</bibRefCitation>
</taxonomicName>
lack an elongate second spine in the first dorsal fin (vs. present). Of the remaining species,
<taxonomicName id="4C4B73B4FFE1FFD8FA7FEEFE5AE8FF3E" authority="(Goren, 1978)" baseAuthorityName="Goren" baseAuthorityYear="1978" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" lastPageId="6" lastPageNumber="215" order="Perciformes" pageId="5" pageNumber="214" phylum="Chordata" rank="species" species="mendelssohni">
<emphasis id="B93FD425FFE1FFD8FA7FEEFE5A1DFF3E" italics="true" lastPageId="6" lastPageNumber="215" pageId="5" pageNumber="214">T. mendelssohni</emphasis>
(
<bibRefCitation id="EFDA75C6FFE2FFD8FEB8E9A65AE3FF3E" author="Goren, M." box="[320,465,151,176]" pageId="6" pageNumber="215" pagination="191 - 203" refId="ref3022" refString="Goren, M. (1978) A new gobiid genus and seven new species from Sinai coasts. Senckenbergiana biologica, 59 (3 / 4), 191 - 203." type="journal article" year="1978">Goren, 1978</bibRefCitation>
)
</taxonomicName>
has multiple branches in the fifth pelvic fin ray, and
<taxonomicName id="4C4B73B4FFE2FFD8FBBCE9A85BFDFF5B" authority="Suzuki &amp; Senou, 2007" authorityName="Suzuki &amp; Senou" authorityYear="2007" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="nomourai">
<emphasis id="B93FD425FFE2FFD8FBBCE9A85FFDFF21" box="[1092,1231,152,176]" italics="true" pageId="6" pageNumber="215">T. nomourai</emphasis>
<bibRefCitation id="EFDA75C6FFE2FFD8FB21E9A65BFDFF5B" author="Suzuki, T. &amp; Senou, H." pageId="6" pageNumber="215" pagination="175 - 184" refId="ref3162" refString="Suzuki, T. &amp; Senou, H. (2007) Two new species of the gobiid fish genus Trimma (Perciformes: Gobioidei) from southern Japan. Bulletin of the National Museum of Natural Science, Series A, Supplement 1, 175 - 184." type="journal article" year="2007">Suzuki &amp; Senou, 2007</bibRefCitation>
</taxonomicName>
has two branches (for a total of four tips) whereas all the others have a single branch point in this ray.
<taxonomicName id="4C4B73B4FFE2FFD8FABDE98C591FFF76" authority="Winterbottom &amp; Villa, 2003" authorityName="Winterbottom &amp; Villa" authorityYear="2003" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="lantana">
<emphasis id="B93FD425FFE2FFD8FABDE98C5BDFFF76" italics="true" pageId="6" pageNumber="215">Trimma lantana</emphasis>
<bibRefCitation id="EFDA75C6FFE2FFD8FF0BE9EE591FFF76" author="Winterbottom, R. &amp; Villa, C. A." box="[243,557,223,248]" pageId="6" pageNumber="215" pagination="13 - 28" refId="ref3499" refString="Winterbottom, R. &amp; Villa, C. A. (2003) A new species of the Trimma caesiura complex (Gobiidae, Teleostei) from the northeastern margin of the Australian Plate, with a redescription of the other nominal species in the complex. aqua, Journal of Ichthyology and Aquatic Biology, 7 (1), 13 - 28." type="journal article" year="2003">Winterbottom &amp; Villa, 2003</bibRefCitation>
</taxonomicName>
has deep, vertically sided trenches in the interorbital and postorbital regions of the head (vs. moderate trough and none respectively in
<taxonomicName id="4C4B73B4FFE2FFD8FCD7E83458A1FE92" authorityName="Winterbottom" authorityYear="2014" box="[815,915,261,284]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE2FFD8FCD7E83458A1FE92" box="[815,915,261,284]" italics="true" pageId="6" pageNumber="215">T. irinae</emphasis>
</taxonomicName>
).
<taxonomicName id="4C4B73B4FFE2FFD8FC57E8345E70FE93" authority="Winterbottom, 2011" authorityName="Winterbottom" authorityYear="2011" box="[943,1346,260,285]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="cheni">
<emphasis id="B93FD425FFE2FFD8FC57E8345F7DFE93" box="[943,1103,260,285]" italics="true" pageId="6" pageNumber="215">Trimma cheni</emphasis>
<bibRefCitation id="EFDA75C6FFE2FFD8FBA2E8355E70FE93" author="Winterbottom, R." box="[1114,1346,260,285]" pageId="6" pageNumber="215" pagination="127 - 162" refId="ref3388" refString="Winterbottom, R. (2011) Six new species of the genus Trimma (Percomorpha; Gobiidae) from the Raja Ampat Islands, Indonesia. aqua, International Journal of Ichthyology, 17 (3), 127 - 162." type="journal article" year="2011">Winterbottom, 2011</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C4B73B4FFE2FFD8FA7FE8345A86FECE" authority="(Aoyagi, 1949)" baseAuthorityName="Aoyagi" baseAuthorityYear="1949" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="okinawae">
<emphasis id="B93FD425FFE2FFD8FA7FE8345A31FECE" italics="true" pageId="6" pageNumber="215">T. okinawae</emphasis>
(
<bibRefCitation id="EFDA75C6FFE2FFD8FEE9E8165A9EFECE" author="Aoyagi, H." box="[273,428,295,320]" pageId="6" pageNumber="215" pagination="171 - 173" refId="ref2980" refString="Aoyagi, H. (1949) Studies on the coral reef fishes of the Riu-Kiu Islands. V. The Zoological Magazine, The Zoological Society of Japan (Tokyo), 58 (9), 171 - 173." type="journal article" year="1949">Aoyagi, 1949</bibRefCitation>
)
</taxonomicName>
have only moderately elongate second dorsal spines that do not extend beyond about the middle of the second dorsal fin when adpressed (vs to last ray or further back).
<taxonomicName id="4C4B73B4FFE2FFD8FBD2E87C5BFCFE06" authority="Winterbottom, 2011" authorityName="Winterbottom" authorityYear="2011" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="papayum">
<emphasis id="B93FD425FFE2FFD8FBD2E87C5FC2FEEA" box="[1066,1264,333,356]" italics="true" pageId="6" pageNumber="215">Trimma papayum</emphasis>
<bibRefCitation id="EFDA75C6FFE2FFD8FB01E87D5BFCFE06" author="Winterbottom, R." pageId="6" pageNumber="215" pagination="127 - 162" refId="ref3388" refString="Winterbottom, R. (2011) Six new species of the genus Trimma (Percomorpha; Gobiidae) from the Raja Ampat Islands, Indonesia. aqua, International Journal of Ichthyology, 17 (3), 127 - 162." type="journal article" year="2011">Winterbottom, 2011</bibRefCitation>
</taxonomicName>
appears closest morphologically to
<taxonomicName id="4C4B73B4FFE2FFD8FD8DE84059EAFE09" authorityName="Winterbottom" authorityYear="2014" box="[629,728,368,392]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE2FFD8FD8DE84059EAFE09" box="[629,728,368,392]" italics="true" pageId="6" pageNumber="215">T. irinae</emphasis>
</taxonomicName>
in having a poorly developed postorbital trough or none, in possessing a greatly elongate second dorsal spine, and in several details of colour pattern. However,
<taxonomicName id="4C4B73B4FFE2FFD8FAE0E8A45EAEFE22" authorityName="Winterbottom" authorityYear="2011" box="[1304,1436,405,428]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="papayum">
<emphasis id="B93FD425FFE2FFD8FAE0E8A45EAEFE22" box="[1304,1436,405,428]" italics="true" pageId="6" pageNumber="215">T. papayum</emphasis>
</taxonomicName>
has more pectoral fin rays (18–19 vs. 17) and fewer lower gill rakers (13–14 vs. 15–16) than
<taxonomicName id="4C4B73B4FFE2FFD8FB5CE8885E30FE41" authorityName="Winterbottom" authorityYear="2014" box="[1188,1282,440,464]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE2FFD8FB5CE8885E30FE41" box="[1188,1282,440,464]" italics="true" pageId="6" pageNumber="215">T. irinae</emphasis>
</taxonomicName>
, in additional to trenchant differences in live and preserved colouration. These include the presence in
<taxonomicName id="4C4B73B4FFE2FFD8FB72E8EC5FD8FE7A" authorityName="Winterbottom" authorityYear="2014" box="[1162,1258,477,500]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE2FFD8FB72E8EC5FD8FE7A" box="[1162,1258,477,500]" italics="true" pageId="6" pageNumber="215">T. irinae</emphasis>
</taxonomicName>
of a distinctive dark spot on the branchiostegal membrane below the vertical limb of the preopercle (vs. absent - a larger such spot is also evident in
<taxonomicName id="4C4B73B4FFE2FFD8FEA4EB145867FDB3" authority="Hagiwara &amp; Winterbottom, 2007" authorityName="Hagiwara &amp; Winterbottom" authorityYear="2007" box="[348,853,548,573]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="hayashii">
<emphasis id="B93FD425FFE2FFD8FEA4EB145AE5FDB3" box="[348,471,548,573]" italics="true" pageId="6" pageNumber="215">T. hayashii</emphasis>
<bibRefCitation id="EFDA75C6FFE2FFD8FE26EB155867FDB3" author="Hagiwara, K. &amp; Winterbottom, R." box="[478,853,548,573]" pageId="6" pageNumber="215" pagination="163 - 174" refId="ref3055" refString="Hagiwara, K. &amp; Winterbottom, R. (2007) Two new species of Trimma (Gobiidae) from the western Pacific. Bulletin of National Science Museum, Series A, Supplement 1, 163 - 174." type="journal article" year="2007">Hagiwara &amp; Winterbottom, 2007</bibRefCitation>
</taxonomicName>
, which usually lacks scales in the predorsal midline and on the opercle), and two large black spots basally in the first dorsal fin (vs. one small dark spot).
</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE2FFD8FF3FEB5D5AF3FD29" pageId="6" pageNumber="215" type="distribution">
<paragraph id="8BF40837FFE2FFD8FF3FEB5D5AF3FD29" blockId="6.[151,1438,151,789]" pageId="6" pageNumber="215">
<emphasis id="B93FD425FFE2FFD8FF3FEB5D5A52FD0B" bold="true" box="[199,352,620,645]" inLineHeading="true" pageId="6" pageNumber="215" reason="1">Distribution.</emphasis>
<taxonomicName id="4C4B73B4FFE2FFD8FE9EEB5C5935FD0A" authorityName="Winterbottom" authorityYear="2014" box="[358,519,621,644]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="irinae">
<emphasis id="B93FD425FFE2FFD8FE9EEB5C5935FD0A" box="[358,519,621,644]" italics="true" pageId="6" pageNumber="215">Trimma irinae</emphasis>
</taxonomicName>
is currently known only from the
<typeStatus id="54F0B695FFE2FFD8FC7AEB5D5880FD0A" box="[898,946,620,644]" pageId="6" pageNumber="215">type</typeStatus>
locality in
<collectingRegion id="498FC6D5FFE2FFD8FBD7EB5D5E23FD0B" box="[1071,1297,620,645]" country="Papua New Guinea" name="Milne Bay" pageId="6" pageNumber="215">Milne Bay Province</collectingRegion>
,
<collectingCountry id="F35C48A7FFE2FFD8FAE4EB5D5BDAFD26" name="Papua New Guinea" pageId="6" pageNumber="215">Papua New Guinea</collectingCountry>
at a depth of
<specimenCount id="9D4DC3BEFFE2FFD8FE79EBBE5AF3FD29" box="[385,449,655,680]" pageId="6" pageNumber="215" type="generic">
<quantity id="4CB3A5D2FFE2FFD8FE79EBBE5A8FFD29" box="[385,445,655,680]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="215" unit="m" value="65.0">65 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C3515BBCFFE2FFD8FF3FEB855F20FC9B" pageId="6" pageNumber="215" type="etymology">
<paragraph id="8BF40837FFE2FFD8FF3FEB855F20FC9B" blockId="6.[151,1438,151,789]" pageId="6" pageNumber="215">
<emphasis id="B93FD425FFE2FFD8FF3FEB855A7CFD43" bold="true" box="[199,334,692,717]" inLineHeading="true" pageId="6" pageNumber="215" reason="1">Etymology.</emphasis>
The species is named in honour of my wife, Irina, as a small token of my appreciation and gratitude for her patience and forbearance of what she once referred to as my “magnificent obsession” with coral reef fishes. This species has been informally referred to as
<taxonomicName id="4C4B73B4FFE2FFD8FCD1EBCC58E8FC9D" box="[809,986,764,788]" class="Actinopterygii" family="Gobiidae" genus="Trimma" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="215" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B93FD425FFE2FFD8FCD1EBCC58B3FC9A" box="[809,897,765,788]" italics="true" pageId="6" pageNumber="215">Trimma</emphasis>
RW sp.
</taxonomicName>
100.
</paragraph>
</subSubSection>
</treatment>
</document>
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