383 lines
47 KiB
XML
383 lines
47 KiB
XML
<document id="FA23C96D4FC0FA96E172543F093847DA" ID-DOI="10.11646/zootaxa.4802.1.14" ID-ISSN="1175-5326" ID-Zenodo-Dep="3904872" ID-ZooBank="15115C18-B0E3-4F58-BD93-1E95EEE9D90F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592922866963" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2020" docId="03E287A2FFFCFFB074F596F8AFA3AE31" docLanguage="en" docName="zootaxa.4802.1.14.pdf" docOrigin="Zootaxa 4802 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nipponophloeostiba thayeri Shavrin 2020, sp.n." docType="treatment" docVersion="3" lastPageNumber="199" masterDocId="FFDBFFDAFFFDFFB374629619AD69AB4A" masterDocTitle="Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini)" masterLastPageNumber="200" masterPageNumber="196" pageNumber="197" updateTime="1700276889372" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="9DF6164085674EFB2880B2E4B802E5F1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8041FB8C9E07BB12FE608AAA93D83791">
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<mods:title id="32AE33CDD61D4670D2393ABE3BC41AC5">Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini)</mods:title>
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</mods:titleInfo>
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<mods:name id="B0BBE237BA115A31AAD951D7F3ED551D" type="personal">
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<mods:role id="6807F8F17CEF2702BB087ADC745A692F">
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<mods:roleTerm id="668C48031D734AD61BB95EFB66CB5D9E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="918F40FF4220B726B19D0752F00471A8">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="9750399C369760BAC6A7AA80F65E8768">text</mods:typeOfResource>
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<mods:titleInfo id="07FD501EBB75D3FF88078BEF4C9C240F">
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<mods:title id="2EA54D3E79523838FD5228F41D715049">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E04F86195C218A055ED5E6C2901317F0">
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<mods:date id="EA2189EE3BE04548538F3BBB3240F20D">2020</mods:date>
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<mods:detail id="31E9882B722D7C8223806912B6839045" type="pubDate">
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<mods:number id="0D25280C13E9CE9E10B7E3B520BA3B7B">2020-06-22</mods:number>
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</mods:detail>
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<mods:detail id="C829B24B89DB57DEDEB21D35FA94A0BB" type="volume">
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<mods:number id="E094EF4F45BB193BB5F675598920ACDE">4802</mods:number>
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</mods:detail>
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<mods:detail id="D5B12AB3E228A9D4A50ACF316196B18B" type="issue">
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<mods:number id="788863137582262B3B3AFED488D7F29F">1</mods:number>
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</mods:detail>
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<mods:extent id="54A1CB6D40A639571CD207237E61E717" unit="page">
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<mods:start id="21CF69962BF1CC6DFBBBE93C30B0ECFE">196</mods:start>
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<mods:end id="2E89CB83B63060BA129F997BDE41EC0A">200</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="59928363123A599F1C01882553BC66D9">journal article</mods:classification>
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<mods:identifier id="810E58DB316F831EC1F20E2ACCF3C704" type="DOI">10.11646/zootaxa.4802.1.14</mods:identifier>
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<mods:identifier id="4434E3D3625DF26CAE51F897FBB94474" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="944193A9FC30F61FB6BB5FA55CC3A4BA" type="Zenodo-Dep">3904872</mods:identifier>
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<mods:identifier id="D863FBBB48CB2BBC1A97F07110153342" type="ZooBank">15115C18-B0E3-4F58-BD93-1E95EEE9D90F</mods:identifier>
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</mods:mods>
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<treatment id="03E287A2FFFCFFB074F596F8AFA3AE31" LSID="urn:lsid:plazi:treatment:03E287A2FFFCFFB074F596F8AFA3AE31" httpUri="http://treatment.plazi.org/id/03E287A2FFFCFFB074F596F8AFA3AE31" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197">
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<subSubSection id="C351653FFFFCFFB274F596F8AFF2ABB6" box="[151,667,225,252]" pageId="1" pageNumber="197" type="nomenclature">
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<paragraph id="8BF436B4FFFCFFB274F596F8AFF2ABB6" blockId="1.[151,667,225,287]" box="[151,667,225,252]" pageId="1" pageNumber="197">
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<heading id="D0BC81D8FFFCFFB274F596F8AFF2ABB6" bold="true" box="[151,667,225,252]" fontSize="11" level="1" pageId="1" pageNumber="197" reason="1">
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<emphasis id="B93FEAA6FFFCFFB274F596F8AFF2ABB6" bold="true" box="[151,667,225,252]" pageId="1" pageNumber="197">
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<taxonomicName id="4C4B4D37FFFCFFB274F596F8AF3BABB1" ID-CoL="8Y6P9" authority="Shavrin" authorityName="Shavrin" authorityYear="2020" box="[151,594,225,252]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="thayeri" status="sp. nov.">
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<emphasis id="B93FEAA6FFFCFFB274F596F8AC8FABB6" bold="true" box="[151,486,225,252]" italics="true" pageId="1" pageNumber="197">Nipponophloeostiba thayeri</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20C57DDFFFCFFB2760296FBAFF2ABB6" box="[608,667,226,252]" pageId="1" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C351653FFFFCFFB274F5971CAC7BAA55" box="[151,274,261,287]" pageId="1" pageNumber="197" type="description">
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<paragraph id="8BF436B4FFFCFFB274F5971CAC7BAA55" blockId="1.[151,667,225,287]" box="[151,274,261,287]" pageId="1" pageNumber="197">
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(
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<figureCitation id="13702A31FFFCFFB274FD971CAC60AA55" box="[159,265,261,287]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="1" pageNumber="197">Figs. 1–9</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C351653FFFFCFFB274F59757AD89A881" pageId="1" pageNumber="197" type="materials_examined">
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<paragraph id="8BF436B4FFFCFFB274F59757A973AA99" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="197">
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<emphasis id="B93FEAA6FFFCFFB274F59757ACAAAA22" bold="true" box="[151,451,333,360]" pageId="1" pageNumber="197">Type material examined:</emphasis>
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<materialsCitation id="3B233CE9FFFCFFB275A99754AEBFAAE5" collectingDate="1987-05-17" collectionCode="FIELD, MUSEUM" collectorName="Kinabalu Nat. Pk. & Liwagu Rv. & A. Smetana' & D. E. Bright & Collector' & H. G. Nelson'" elevation="1500" location="SABAH" pageId="1" pageNumber="197" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="54F08816FFFCFFB275A99754AF5AAA2D" box="[459,563,333,359]" pageId="1" pageNumber="197" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D4DFD3DFFFCFFB276599757AF3AAA22" box="[571,595,334,360]" pageId="1" pageNumber="197" type="male">♂</specimenCount>
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: ‘BORNEO
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<collectingRegion id="498FF856FFFCFFB276879754AE2DAA2D" box="[741,836,333,359]" country="Malaysia" name="Sabah" pageId="1" pageNumber="197">SABAH</collectingRegion>
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Mt. |
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<collectorName id="26BE5362FFFCFFB277EB9754A93DAA2D" box="[905,1108,333,360]" pageId="1" pageNumber="197">Kinabalu Nat. Pk.</collectorName>
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| HQ at
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<collectorName id="26BE5362FFFCFFB270D79754A850AA22" box="[1205,1337,333,360]" pageId="1" pageNumber="197">Liwagu Rv.</collectorName>
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|
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<quantity id="4CB39B51FFFCFFB2712C9754A8F4AA22" box="[1358,1437,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="197" unit="m" value="1500.0">
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||
<elevation id="0066D187FFFCFFB2712C9754A8F4AA22" box="[1358,1437,333,360]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="197" unit="m" value="1500.0">1500m</elevation>
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</quantity>
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<date id="FFF51074FFFCFFB274F59768AC7CAAC1" box="[151,277,369,396]" pageId="1" pageNumber="197" value="1987-05-17">
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<collectingDate id="EFB1E99CFFFCFFB274F59768AC7CAAC1" box="[151,277,369,396]" pageId="1" pageNumber="197" value="1987-05-17">17. V. 1987</collectingDate>
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</date>
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|
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<collectorName id="26BE5362FFFCFFB275469768ACC1AAC1" box="[292,424,369,395]" pageId="1" pageNumber="197">A.Smetana’</collectorName>
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<rectangular label, printed>, ‘
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<collectorName id="26BE5362FFFCFFB276979768AE13AAC1" box="[757,890,369,395]" pageId="1" pageNumber="197">D.E. Bright</collectorName>
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|
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<collectorName id="26BE5362FFFCFFB277E99768AE92AAC1" box="[907,1019,369,395]" pageId="1" pageNumber="197">Collector’</collectorName>
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<rectangular label, printed>, ‘
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<collectionCode id="ED5AAE71FFFCFFB2712A9768AC7BAAE5" pageId="1" pageNumber="197">FIELD MUSEUM</collectionCode>
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| ex collection of |
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<collectorName id="26BE5362FFFCFFB2759B978FAFF3AAE5" box="[505,666,405,432]" pageId="1" pageNumber="197">H. G. Nelson’</collectorName>
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<rectangular label, printed
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</materialsCitation>
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>, ‘
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<materialsCitation id="3B233CE9FFFCFFB27795978CA97FAA99" collectionCode="V, FMNH" collectorName="Shavrin A." pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus id="54F08816FFFCFFB27795978CA9F9AAE5" box="[1015,1168,405,431]" pageId="1" pageNumber="197" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C4B4D37FFFCFFB270C8978CA8E4AAE5" authorityName="Watanabe" authorityYear="1962" box="[1194,1421,405,431]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">Nipponophloeostiba</taxonomicName>
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| thayeri
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<taxonomicNameLabel id="A20C57DDFFFCFFB2748E97A3AC48AA9E" box="[236,289,442,468]" pageId="1" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
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||
|
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||
<collectorName id="26BE5362FFFCFFB2755697A0ACC2AA9E" box="[308,427,441,468]" pageId="1" pageNumber="197">Shavrin A.</collectorName>
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<collectionCode id="ED5AAE71FFFCFFB275C997A3ACA8AA9E" box="[427,449,442,468]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="197" type="Museum">V</collectionCode>
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. des. 2020’ <red rectangular label, printed> (
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<collectionCode id="ED5AAE71FFFCFFB277D497A0A964AA99" box="[950,1037,441,467]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
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)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BF436B4FFFCFFB274A597C7AD89A881" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="197">
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||
<emphasis id="B93FEAA6FFFCFFB274A597C7AC56AAB2" bold="true" box="[199,319,478,504]" pageId="1" pageNumber="197">
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||
<typeStatus id="54F08816FFFCFFB274A597C7AC56AAB2" box="[199,319,478,504]" pageId="1" pageNumber="197" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
(
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<specimenCount id="9D4DFD3DFFFCFFB2752D97C4AC87AAB2" box="[335,494,477,504]" pageId="1" pageNumber="197" type="generic">23 specimens</specimenCount>
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||
):
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<materialsCitation id="3B233CE9FFFCFFB2766297C4A844AABD" box="[512,1325,477,504]" collectionCode="FMNH" collectorName="Shavrin A." pageId="1" pageNumber="197" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
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<specimenCount id="9D4DFD3DFFFCFFB2766297C4AF42AAB2" box="[512,555,477,504]" pageId="1" pageNumber="197" type="male">3 ♂</specimenCount>
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||
,
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||
<specimenCount id="9D4DFD3DFFFCFFB2762397C4AF02AAB2" box="[577,619,477,504]" pageId="1" pageNumber="197" type="female">2 ♀</specimenCount>
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(one specimen dissected): same data as the holotype (
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<collectionCode id="ED5AAE71FFFCFFB270AF97C4A84DAABD" box="[1229,1316,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
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||
)
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</materialsCitation>
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||
;
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<specimenCount id="9D4DFD3DFFFCFFB2715A97C4A80AAAB2" box="[1336,1379,477,504]" pageId="1" pageNumber="197" type="male">5 ♂</specimenCount>
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(one specimen dissected),
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<materialsCitation id="3B233CE9FFFCFFB275E69418A807A975" collectionCode="FIELD, MUSEUM" collectorName="Kinabalu Nat. Pk. & D. E. Bright & Collector'" elevation="1560" location="Mt." pageId="1" pageNumber="197" specimenCount="5" specimenCount-female="5" stateProvince="Sabah" typeStatus="paratype">
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||
<specimenCount id="9D4DFD3DFFFCFFB275E69418ACC4A956" box="[388,429,513,540]" pageId="1" pageNumber="197" type="female">5 ♀</specimenCount>
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(one specimen dissected): ‘BORNEO
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<collectingRegion id="498FF856FFFCFFB277389418AED0A951" box="[858,953,513,539]" country="Malaysia" name="Sabah" pageId="1" pageNumber="197">SABAH</collectingRegion>
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||
<location id="8E94606FFFFCFFB277DD941BAE8EA956" LSID="urn:lsid:plazi:treatment:03E287A2FFFCFFB074F596F8AFA3AE31:8E94606FFFFCFFB277DD941BAE8EA956" box="[959,999,514,540]" name="Mt." pageId="1" pageNumber="197" stateProvince="Sabah">Mt.</location>
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||
|
|
||
<collectorName id="26BE5362FFFCFFB2779A9418A9D6A951" box="[1016,1215,513,540]" pageId="1" pageNumber="197">Kinabalu Nat. Pk.</collectorName>
|
||
| HQ
|
||
<quantity id="4CB39B51FFFCFFB2709D9418A824A956" box="[1279,1357,513,540]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="197" unit="m" value="1560.0">
|
||
<elevation id="0066D187FFFCFFB2709D9418A824A956" box="[1279,1357,513,540]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="1" pageNumber="197" unit="m" value="1560.0">1560m</elevation>
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||
</quantity>
|
||
24. IV. [19]87’ <rectangular label, printed>, ‘
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||
<collectorName id="26BE5362FFFCFFB2765E943CAFABA975" box="[572,706,549,575]" pageId="1" pageNumber="197">D.E. Bright</collectorName>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB276B4943CAE2FA975" box="[726,838,549,575]" pageId="1" pageNumber="197">Collector’</collectorName>
|
||
<rectangular label, printed>, ‘
|
||
<collectionCode id="ED5AAE71FFFCFFB270FA943CA807A975" box="[1176,1390,549,575]" pageId="1" pageNumber="197">FIELD MUSEUM</collectionCode>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B233CE9FFFCFFB271E0943FA9C5A92E" collectionCode="FMNH" location="Nelson'" pageId="1" pageNumber="197" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Sabah" typeStatus="paratype">
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||
ex collection of | H. G.
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||
<location id="8E94606FFFFCFFB2751D9450ACBFA929" LSID="urn:lsid:plazi:treatment:03E287A2FFFCFFB074F596F8AFA3AE31:8E94606FFFFCFFB2751D9450ACBFA929" box="[383,470,585,611]" name="Nelson'" pageId="1" pageNumber="197" stateProvince="Sabah">Nelson’</location>
|
||
<rectangular label, printed> (
|
||
<specimenCount id="9D4DFD3DFFFCFFB2774B9450AE3DA92E" box="[809,852,585,612]" pageId="1" pageNumber="197" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4DFD3DFFFCFFB277039450AEE4A92E" box="[865,909,585,612]" pageId="1" pageNumber="197" type="female">1 ♀</specimenCount>
|
||
: CSh;
|
||
<specimenCount id="9D4DFD3DFFFCFFB277B99450A96EA92E" box="[987,1031,585,612]" pageId="1" pageNumber="197" type="male">4 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4DFD3DFFFCFFB270769450A929A92E" box="[1044,1088,585,612]" pageId="1" pageNumber="197" type="female">4 ♀</specimenCount>
|
||
:
|
||
<collectionCode id="ED5AAE71FFFCFFB2702C9453A9CDA92E" box="[1102,1188,586,612]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D4DFD3DFFFCFFB270D59450A98BA92E" box="[1207,1250,585,612]" pageId="1" pageNumber="197" type="male">1 ♂</specimenCount>
|
||
(dissected),
|
||
<materialsCitation id="3B233CE9FFFCFFB271109450A9C0A9E1" collectionCode="FIELD, MUSEUM" collectorName="Mt. Kinabalu N. P. & D. E. Bright & Collector'" elevation="1558" location="Headquarters" pageId="1" pageNumber="197" specimenCount="3" specimenCount-female="3" stateProvince="Sabah" typeStatus="paratype">
|
||
<specimenCount id="9D4DFD3DFFFCFFB271109450A8F5A92E" box="[1394,1436,585,612]" pageId="1" pageNumber="197" type="female">3 ♀</specimenCount>
|
||
(one specimen dissected): ‘BORNEO:
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||
<collectingRegion id="498FF856FFFCFFB2762B9474AFE4A9CD" box="[585,653,621,647]" country="Malaysia" name="Sabah" pageId="1" pageNumber="197">Sabah</collectingRegion>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB276C29477AE1BA9C2" box="[672,882,621,648]" pageId="1" pageNumber="197">Mt. Kinabalu N. P.</collectorName>
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|
|
||
<location id="8E94606FFFFCFFB277E69474A970A9CD" LSID="urn:lsid:plazi:treatment:03E287A2FFFCFFB074F596F8AFA3AE31:8E94606FFFFCFFB277E69474A970A9CD" box="[900,1049,621,647]" name="Headquarters" pageId="1" pageNumber="197" stateProvince="Sabah">Headquarters</location>
|
||
|
|
||
<quantity id="4CB39B51FFFCFFB2704E9474A913A9C2" box="[1068,1146,621,648]" metricMagnitude="3" metricUnit="m" metricValue="1.558" pageId="1" pageNumber="197" unit="m" value="1558.0">
|
||
<elevation id="0066D187FFFCFFB2704E9474A913A9C2" box="[1068,1146,621,648]" metricMagnitude="3" metricUnit="m" metricValue="1.558" pageId="1" pageNumber="197" unit="m" value="1558.0">1558m</elevation>
|
||
</quantity>
|
||
IV-29-[19]87’ <rectangu- lar label, printed>, ‘
|
||
<collectorName id="26BE5362FFFCFFB275169488AC92A9E1" box="[372,507,657,683]" pageId="1" pageNumber="197">D.E. Bright</collectorName>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB2766D9488AF16A9E1" box="[527,639,657,683]" pageId="1" pageNumber="197">Collector’</collectorName>
|
||
<rectangular label, printed>, ‘
|
||
<collectionCode id="ED5AAE71FFFCFFB277B09488A9C0A9E1" box="[978,1193,657,683]" pageId="1" pageNumber="197">FIELD MUSEUM</collectionCode>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B233CE9FFFCFFB270DC948BAFD5A985" collectionCode="FMNH" collectorName="Nelson'" pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
|
||
ex collection of | H. G.
|
||
<collectorName id="26BE5362FFFCFFB274DB94ACAC79A985" box="[185,272,693,719]" pageId="1" pageNumber="197">Nelson’</collectorName>
|
||
<rectangular label, printed> (
|
||
<collectionCode id="ED5AAE71FFFCFFB2763F94ACAFDDA985" box="[605,692,693,719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D4DFD3DFFFCFFB276A594ACAF98A99A" box="[711,753,693,720]" pageId="1" pageNumber="197" type="male">1 ♂</specimenCount>
|
||
(right antennomeres 2–11 missing),
|
||
<materialsCitation id="3B233CE9FFFCFFB270E994ACA9D4A9BE" collectionCode="V" collectorName="Mt. & Kinabalu Nat. Pk. & Liwagu Rv." elevation="1500" location="SABAH" pageId="1" pageNumber="197" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
|
||
<specimenCount id="9D4DFD3DFFFCFFB270E994ACA9DCA99A" box="[1163,1205,693,720]" pageId="1" pageNumber="197" type="female">1 ♀</specimenCount>
|
||
(right antennomeres 4–11 missing): ‘BORNEO
|
||
<collectingRegion id="498FF856FFFCFFB275AB94C0AF41A9B9" box="[457,552,729,755]" country="Malaysia" name="Sabah" pageId="1" pageNumber="197">SABAH</collectingRegion>
|
||
<collectorName id="26BE5362FFFCFFB2764D94C3AF3EA9BE" box="[559,599,730,756]" pageId="1" pageNumber="197">Mt.</collectorName>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB2760994C0AE5DA9B9" box="[619,820,729,756]" pageId="1" pageNumber="197">Kinabalu Nat. Pk.</collectorName>
|
||
| HQ at
|
||
<collectorName id="26BE5362FFFCFFB277F194C0A97EA9BE" box="[915,1047,729,756]" pageId="1" pageNumber="197">Liwagu Rv.</collectorName>
|
||
|
|
||
<quantity id="4CB39B51FFFCFFB2704894C0A911A9BE" box="[1066,1144,729,756]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="197" unit="m" value="1500.0">
|
||
<elevation id="0066D187FFFCFFB2704894C0A911A9BE" box="[1066,1144,729,756]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="197" unit="m" value="1500.0">1500m</elevation>
|
||
</quantity>
|
||
<date id="FFF51074FFFCFFB2701D94C0A9D4A9BE" box="[1151,1213,729,756]" pageId="1" pageNumber="197" value="1987-05-21">
|
||
21.
|
||
<collectionCode id="ED5AAE71FFFCFFB270CB94C3A9D4A9BE" box="[1193,1213,730,756]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="197" type="Museum">V</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B233CE9FFFCFFB270DF94C3A9EFA85D" collectionCode="FIELD, MUSEUM" collectorName="D. E. Bright & Collector'" pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
|
||
. 1987’ <rectangular label, printed>, ‘
|
||
<collectorName id="26BE5362FFFCFFB2752D94E4ACBFA85D" box="[335,470,765,791]" pageId="1" pageNumber="197">D.E. Bright</collectorName>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB2758894E4AF33A85D" box="[490,602,765,791]" pageId="1" pageNumber="197">Collector’</collectorName>
|
||
<rectangular label, printed>, ‘
|
||
<collectionCode id="ED5AAE71FFFCFFB277CD94E4A9EFA85D" box="[943,1158,765,791]" pageId="1" pageNumber="197">FIELD MUSEUM</collectionCode>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B233CE9FFFCFFB270F894E7AFC8A871" collectionCode="FMNH" collectorName="Nelson'" pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
|
||
ex collection of | H. G.
|
||
<collectorName id="26BE5362FFFCFFB274F59538AD87A871" box="[151,238,801,827]" pageId="1" pageNumber="197">Nelson’</collectorName>
|
||
<rectangular label, printed> (
|
||
<collectionCode id="ED5AAE71FFFCFFB276209538AFF0A871" box="[578,665,801,827]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B233CE9FFFCFFB276CC9538AF23A8C9" collectionCode="FIELD, MUSEUM" collectorName="Mt. Kinabalu N. P. & Large & D. E. Bright & Collector'" elevation="1558" location="Headquarters" pageId="1" pageNumber="197" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
|
||
<specimenCount id="9D4DFD3DFFFCFFB276CC9538AFB2A876" box="[686,731,801,828]" pageId="1" pageNumber="197" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4DFD3DFFFCFFB2768B9538AE7FA876" box="[745,790,801,828]" pageId="1" pageNumber="197" type="female">1 ♀</specimenCount>
|
||
: ‘BORNEO:
|
||
<collectingRegion id="498FF856FFFCFFB277D39538AE9CA871" box="[945,1013,801,827]" country="Malaysia" name="Sabah" pageId="1" pageNumber="197">Sabah</collectingRegion>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB2706E953BA98AA876" box="[1036,1251,801,828]" pageId="1" pageNumber="197">Mt. Kinabalu N. P.</collectorName>
|
||
|
|
||
<location id="8E94606FFFFCFFB2709B9538A8E7A871" LSID="urn:lsid:plazi:treatment:03E287A2FFFCFFB074F596F8AFA3AE31:8E94606FFFFCFFB2709B9538A8E7A871" box="[1273,1422,801,827]" name="Headquarters" pageId="1" pageNumber="197" stateProvince="Sabah">Headquarters</location>
|
||
|
|
||
<quantity id="4CB39B51FFFCFFB274F5955CAD8CA82A" box="[151,229,837,864]" metricMagnitude="3" metricUnit="m" metricValue="1.558" pageId="1" pageNumber="197" unit="m" value="1558.0">
|
||
<elevation id="0066D187FFFCFFB274F5955CAD8CA82A" box="[151,229,837,864]" metricMagnitude="3" metricUnit="m" metricValue="1.558" pageId="1" pageNumber="197" unit="m" value="1558.0">1558m</elevation>
|
||
</quantity>
|
||
IV-27-[19]87’ <rectangular label, printed>, ‘
|
||
<collectorName id="26BE5362FFFCFFB276BA955CAE77A815" box="[728,798,837,863]" pageId="1" pageNumber="197">Large</collectorName>
|
||
cut | tree-Quercus | sp.
|
||
<collectorName id="26BE5362FFFCFFB27079955CA9F1A815" box="[1051,1176,837,863]" pageId="1" pageNumber="197">D.E.Bright</collectorName>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB270C9955CA872A815" box="[1195,1307,837,863]" pageId="1" pageNumber="197">Collector’</collectorName>
|
||
<rectangu- lar label, printed>, ‘
|
||
<collectionCode id="ED5AAE71FFFCFFB275119570AF23A8C9" box="[371,586,873,899]" pageId="1" pageNumber="197">FIELD MUSEUM</collectionCode>
|
||
</materialsCitation>
|
||
|
|
||
<materialsCitation id="3B233CE9FFFCFFB2763D9573AFE0A8ED" collectionCode="FMNH" collectorName="H. G. Nelson' & All" pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
|
||
ex collection of |
|
||
<collectorName id="26BE5362FFFCFFB277419573AEA9A8C9" box="[803,960,873,900]" pageId="1" pageNumber="197">H. G. Nelson’</collectorName>
|
||
<rectangular label, printed> (
|
||
<collectionCode id="ED5AAE71FFFCFFB271729570A80EA8C9" box="[1296,1383,873,899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="1" pageNumber="197" type="Museum">FMNH</collectionCode>
|
||
).
|
||
<collectorName id="26BE5362FFFCFFB2711A9570A8F5A8C9" box="[1400,1436,873,899]" pageId="1" pageNumber="197">All</collectorName>
|
||
<typeStatus id="54F08816FFFCFFB274F59597AC6BA8E2" box="[151,258,910,936]" pageId="1" pageNumber="197" type="paratype">paratypes</typeStatus>
|
||
with red rectangular printed label
|
||
</materialsCitation>
|
||
: ‘
|
||
<materialsCitation id="3B233CE9FFFCFFB276FA9594ADBEA881" collectionCode="V" collectorName="Shavrin A." pageId="1" pageNumber="197" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
|
||
<typeStatus id="54F08816FFFCFFB276FA9594AE41A8ED" box="[664,808,909,935]" pageId="1" pageNumber="197" type="paratype">PARATYPE</typeStatus>
|
||
|
|
||
<taxonomicName id="4C4B4D37FFFCFFB2775C9594A948A8ED" authorityName="Watanabe" authorityYear="1962" box="[830,1057,909,935]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="199" phylum="Arthropoda" rank="genus">Nipponophloeostiba</taxonomicName>
|
||
| thayeri
|
||
<taxonomicNameLabel id="A20C57DDFFFCFFB270EF9597A9ABA8E2" box="[1165,1218,910,936]" pageId="1" pageNumber="197" rank="species">sp.n.</taxonomicNameLabel>
|
||
|
|
||
<collectorName id="26BE5362FFFCFFB270B59594A826A8E2" box="[1239,1359,909,936]" pageId="1" pageNumber="197">Shavrin A.</collectorName>
|
||
<collectionCode id="ED5AAE71FFFCFFB2712D9597A80CA8E2" box="[1359,1381,910,936]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="197" type="Museum">V</collectionCode>
|
||
. des. 2020
|
||
</materialsCitation>
|
||
’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351653FFFFCFFB074A595CCAC10AFE9" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197" type="description">
|
||
<paragraph id="8BF436B4FFFCFFB274A595CCADB0AE45" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="197">
|
||
<emphasis id="B93FEAA6FFFCFFB274A595CCAC38A8BA" bold="true" box="[199,337,981,1008]" pageId="1" pageNumber="197">Description</emphasis>
|
||
. Measurements (n=24, in mm): maximum width of head including eyes: 0.36–0.41; length of head (from base of labrum to neck contriction along head midline): 0.24–0.27; length of antenna (
|
||
<typeStatus id="54F08816FFFCFFB270CC95E0A87EAF5E" box="[1198,1303,1017,1044]" pageId="1" pageNumber="197" type="holotype">holotype</typeStatus>
|
||
): 0.44; ocular length (longitudinal): 0.10–0.11; length of temple: 0.05–0.06; length of pronotum: 0.27–0.32; maximum width of pronotum: 0.38–0.42; minimum width of pronotum: 0.27–0.33; sutural length of elytra (length of elytra from the apex of scutellum to the posterior margin of sutural angle): 0.44–0.52; maximum width of elytra: 0.44–0.49; maximum width of abdomen: 0.46–0.50; length of metatibia (
|
||
<typeStatus id="54F08816FFFCFFB277259290AED9AFEE" box="[839,944,1161,1188]" pageId="1" pageNumber="197" type="holotype">holotype</typeStatus>
|
||
): 0.22; length of metatarsus (
|
||
<typeStatus id="54F08816FFFCFFB270939290A833AFEE" box="[1265,1370,1161,1188]" pageId="1" pageNumber="197" type="holotype">holotype</typeStatus>
|
||
): 0.11 (metatarsomeres 1–4: 0.03; metatarsomere 5: 0.08); length of aedeagus (from base of the median lobe to apex of parameres): 0.30–0.32; total length (from anterior margin of clypeus to apex of abdomen): 1.75–2.22 (
|
||
<typeStatus id="54F08816FFFCFFB2715392C8A8F0AFA1" box="[1329,1433,1233,1259]" pageId="1" pageNumber="197" type="holotype">holotype</typeStatus>
|
||
: 1.97).
|
||
</paragraph>
|
||
<paragraph id="8BF436B4FFFCFFB274959300AE43ADA9" blockId="1.[151,1437,333,2016]" pageId="1" pageNumber="197">
|
||
Body somewhat parallel-sided, significantly flattened. Head and abdomen brown; pronotum reddishbrown; antennomeres 6–11, elytra (scutellar and apical portions slightly darker) and sometimes apical margins and intersegmental membranes of abdominal tergites 3–6 yellow-brown; mouthparts, antennomeres 1–5 and legs yellow. Middle portion of head with several sparse, irregular, fine punctures; pronotum with regular, fine punctation, sparser in medioapical portion and along midline; scutellum without punctures; elytra with punctation similar to that in pronotum, but slightly deeper and denser, smooth in apical and lateral portions; abdomen without or with very indistinct and sparse fine punctures. Head with dense, irregular and relatively coarse microsculpture: transverse in frontal, diagonal in middle and longitudinal in lateral portions; neck without microsculpture in middle, with isodiametric meshes in lateral portions; pronotum with dense, longitudinal microsculpture as that in head; scutellum without or with indistinct isodiametric microreticulation; elytra with longitudinal and diagonal microsculpture, denser and coarser than that in pronotum; abdomen with irregular, dense and coarse, transverse and isodiametric sculpture, markedly coarser on abdominal tergite VII. Pubescence of forebody very sparse and short; paratergites and tergite VIII with moderately long erect setae. Habitus as in
|
||
<figureCitation id="13702A31FFFCFFB2768090D0AE4DADA9" box="[738,804,1737,1763]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="1" pageNumber="197">Fig. 1</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436B4FFFCFFB074A590F4AFBAAA2D" blockId="1.[151,1437,333,2016]" lastBlockId="3.[151,1437,153,1403]" lastPageId="3" lastPageNumber="199" pageId="1" pageNumber="197">Head wide, subtrapezoidal, 1.5 times as broad as long, with long and moderately deep straight anteocellar foveae extending level of middle or apical third of eyes; basal margin of head straight; posterior portions between margin of ocelli and temples with distinct narrow and straight carina, almost reaching posterior margin of eye; postantennal depressions indistinct; eyes large, convex; ocelli moderately large, located very close to posterior constriction, distance between ocelli about as long as distance between ocellus and posterior margin of eye, or slightly longer; postocular parts moderately long, slightly convex, not expanded laterad, about twice as long as longitudinal length of eyes. Neck separated from head by distinct, narrow and deep constriction. Maxillary palpi moderately long; preapical palpomere swollen, slightly longer than wide; apical palpomere narrow, realtively parallel-sided, with conical apex, about 1.3 times as long as preceding palpomere. Antennae short, reaching apical third of pronotum when reclined; basal antennomere suboval, about twice as long as 2, antennomere 2 small, indistinctly longer than wide, 3 narrower, about as long as 2, 4 very small, transverse, 5 markedly wider than 4, twice wider than long, antennomeres 5–10 progressively widened apicad, distinctly transverse and covered by dense pubescence, apical antennomere wide, oval, about twice as long as 10.</paragraph>
|
||
<caption id="DF34663CFFFFFFB174F5917CADA4ACA0" ID-DOI="http://doi.org/10.5281/zenodo.3904874" ID-Zenodo-Dep="3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="2" pageNumber="198" startId="2.[151,264,1893,1918]" targetBox="[157,1425,192,1862]" targetPageId="2">
|
||
<paragraph id="8BF436B4FFFFFFB174F5917CADA4ACA0" blockId="2.[151,1437,1893,2026]" pageId="2" pageNumber="198">
|
||
<emphasis id="B93FEAA6FFFFFFB174F5917CAC53AC34" bold="true" box="[151,314,1893,1918]" pageId="2" pageNumber="198">FIGURES 1–9.</emphasis>
|
||
<taxonomicName id="4C4B4D37FFFFFFB17522917FAF35AC34" authorityName="Shavrin" authorityYear="2020" box="[320,604,1894,1918]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="thayeri">
|
||
<emphasis id="B93FEAA6FFFFFFB17522917FAF35AC34" box="[320,604,1894,1918]" italics="true" pageId="2" pageNumber="198">Nipponophloeostiba thayeri</emphasis>
|
||
</taxonomicName>
|
||
: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF436B4FFFEFFB074A59768AFBAA951" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">Pronotum subtrapezoidal, slightly wider than long and about as wide as head, wider in apical portion, from middle strongly narrowed towards widely rounded posterior angles; apical angles widely rounded; apical margin straight or slightly rounded, with two small, convex tubercles at level of lateral margins of neck; basal margin rounded; median part with longitudinal narrow impunctated band; lateral portions bordered at margins and narrowly flattened, laterobasal portions distinctly impressed.</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A5943CACF4A9CD" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">Elytra about as wide as long, 1.6 times as long as pronotum, somewhat parallel-sided, indistinctly widened in apical third towards widely rounded apical angles; apical margin straight or slightly rounded, truncated at suture. Wings fully developed.</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A59488AD8BA985" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">Metatarsus twice as long as metatibia; apical metatarsomere significantly longer than previous four tarsomeres.</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A594C0A9EBA85D" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">Abdomen parallel-sided, about same width as elytra, with two small, round wing-folding patches (tomentose spots) on abdominal tergite IV and narrow palisade fringe on apical margin of tergite VII.</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A59538A915A881" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
Male. Apical margin of abdominal tergite VIII rounded (
|
||
<figureCitation id="13702A31FFFEFFB0773C9538AECEA871" box="[862,935,801,827]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 4</figureCitation>
|
||
). Apical margin of abdominal sternite VIII slightly sinuate (
|
||
<figureCitation id="13702A31FFFEFFB07537955CACF5A815" box="[341,412,836,863]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 5</figureCitation>
|
||
). Genital segment as in
|
||
<figureCitation id="13702A31FFFEFFB076D7955CAF95A815" box="[693,764,837,863]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 6</figureCitation>
|
||
. Aedeagus (
|
||
<figureCitation id="13702A31FFFEFFB077EA955CAEA4A815" box="[904,973,837,863]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 2</figureCitation>
|
||
) long, with wide basal portion and very narrow median lobe; parameres distinctly widened apically, with three long apical, three preapical, and two lateral setae, and with three long additional setae along entire length of inner edge; internal sac without large sclerotized structures, with very small round spines in apical portion. Aedeagus laterally as in
|
||
<figureCitation id="13702A31FFFEFFB0705695A8A91EA881" box="[1076,1143,945,971]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074AC95CCAE09AF59" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
Female. Abdominal tergite VIII (
|
||
<figureCitation id="13702A31FFFEFFB0765F95CCAFE9A8A5" box="[573,640,981,1007]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 7</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="13702A31FFFEFFB0773795CCAEFEA8A5" box="[853,919,981,1007]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 8</figureCitation>
|
||
) rounded. Genital segment as in
|
||
<figureCitation id="13702A31FFFEFFB0716595CCA823A8A5" box="[1287,1354,981,1007]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1893,1918]" captionTargetBox="[157,1425,192,1862]" captionTargetId="figure@2.[151,1436,179,1866]" captionTargetPageId="2" captionText="FIGURES 1–9. Nipponophloeostiba thayeri: 1—habitus, 2—aedeagus (parameral view), 3—aedeagus (lateral view), 4—male tergite VIII (dorsal view), 5—male sternite VIII (ventral view), 6—male genital segment (dorsal view), 7—female tergite VIII (dorsal view), 8—female sternite VIII (ventral view), 9—female genital segment. Scale bar: 1.00 mm (Fig. 1), 0.10 mm (Figs. 2–9)." figureDoi="http://doi.org/10.5281/zenodo.3904874" httpUri="https://zenodo.org/record/3904874/files/figure.png" pageId="3" pageNumber="199">Fig. 9</figureCitation>
|
||
; acces- sory sclerite moderately wide, suboval. Spermatheca not found.
|
||
</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A59205AC10AFE9" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
<emphasis id="B93FEAA6FFFEFFB074A59205ACDDAF7D" bold="true" box="[199,436,1052,1079]" pageId="3" pageNumber="199">Comparative notes.</emphasis>
|
||
<taxonomicName id="4C4B4D37FFFEFFB075DC9204AF92AF7D" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[446,763,1053,1079]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="thayeri" status="sp. nov.">
|
||
<emphasis id="B93FEAA6FFFEFFB075DC9204AF92AF7D" box="[446,763,1053,1079]" italics="true" pageId="3" pageNumber="199">Nipponophloeostiba thayeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93FEAA6FFFEFFB077649204AE54AF7D" bold="true" box="[774,829,1053,1079]" pageId="3" pageNumber="199">
|
||
<taxonomicNameLabel id="A20C57DDFFFEFFB077649204AE54AF7D" box="[774,829,1053,1079]" pageId="3" pageNumber="199" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from Japanese
|
||
<taxonomicName id="4C4B4D37FFFEFFB070289207A985AF7D" authorityName="Watanabe" authorityYear="1962" box="[1098,1260,1053,1079]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="species" species="verrucifera">
|
||
<emphasis id="B93FEAA6FFFEFFB070289207A934AF7D" box="[1098,1117,1054,1079]" italics="true" pageId="3" pageNumber="199">N</emphasis>
|
||
.
|
||
<emphasis id="B93FEAA6FFFEFFB0700C9204A985AF7D" box="[1134,1260,1053,1079]" italics="true" pageId="3" pageNumber="199">verrucifera</emphasis>
|
||
</taxonomicName>
|
||
by the smaller body, darker abdomen, more prominent eyes, somewhat narrower head, with different shape of the postocular parts, not expanded laterad, slightly shorter elytra, narrower median lobe and wider apical portions of the parameres, with more setae on them.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351653FFFFEFFB074A592B5AC0DAE79" pageId="3" pageNumber="199" type="distribution">
|
||
<paragraph id="8BF436B4FFFEFFB074A592B5AF6DAFA1" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
<emphasis id="B93FEAA6FFFEFFB074A592B5AC08AF8D" bold="true" box="[199,353,1196,1223]" pageId="3" pageNumber="199">Distribution.</emphasis>
|
||
The species is known from several locations close to each other in Kinabalu National Park in
|
||
<collectingRegion id="498FF856FFFEFFB074F592C8ADB7AFA1" box="[151,222,1233,1259]" country="Malaysia" name="Sabah" pageId="3" pageNumber="199">Sabah</collectingRegion>
|
||
, Borneo Island,
|
||
<collectingCountry id="F35C7624FFFEFFB075F792C8AC96AFA1" box="[405,511,1233,1259]" name="Malaysia" pageId="3" pageNumber="199">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF436B4FFFEFFB074A592EDAC0DAE79" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
<emphasis id="B93FEAA6FFFEFFB074A592EDAC22AE45" bold="true" box="[199,331,1268,1295]" pageId="3" pageNumber="199">Bionomics.</emphasis>
|
||
Specimens were collected at elevations
|
||
<quantity id="4CB39B51FFFEFFB0777B92EDAEA6AE45" box="[793,975,1268,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="199" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">
|
||
<date id="FFF51074FFFEFFB0777B92EDAED9AE45" box="[793,944,1268,1295]" pageId="3" pageNumber="199" value="1500" valueMax="1600">1500 to 1600</date>
|
||
m
|
||
</quantity>
|
||
a.s.l. Some specimens were collected in large cut
|
||
<taxonomicName id="4C4B4D37FFFEFFB0749D9300AC34AE79" box="[255,349,1305,1331]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="3" pageNumber="199" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B93FEAA6FFFEFFB0749D9300AC34AE79" box="[255,349,1305,1331]" italics="true" pageId="3" pageNumber="199">Quercus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C351653FFFFEFFB074A59324AFA3AE31" pageId="3" pageNumber="199" type="etymology">
|
||
<paragraph id="8BF436B4FFFEFFB074A59324AFA3AE31" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="199">
|
||
<emphasis id="B93FEAA6FFFEFFB074A59324AC27AE1D" bold="true" box="[199,334,1341,1367]" pageId="3" pageNumber="199">Etymology.</emphasis>
|
||
Eponymic, the species is named to honour my colleague Margaret K. Thayer (Chicago,
|
||
<collectingCountry id="F35C7624FFFEFFB071519324A804AE1D" box="[1331,1389,1341,1367]" name="United States of America" pageId="3" pageNumber="199">USA</collectingCountry>
|
||
) for help in the study of
|
||
<taxonomicName id="4C4B4D37FFFEFFB075149378AC82AE31" authorityName="MacLeay" authorityYear="1825" box="[374,491,1377,1403]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="199" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
|
||
during many years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |