475 lines
92 KiB
XML
475 lines
92 KiB
XML
<document id="C9ED499A87286E3B789556522B00916D" ID-DOI="10.5281/zenodo.281468" ID-GBIF-Dataset="c8c7cfd9-2eb5-4be9-9d30-f3741b06c619" ID-ISSN="1175-5326" ID-Zenodo-Dep="281468" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460489221128" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Anker, Arthur" docDate="2012" docId="03E2878C8C7F7602FF2EA1D0E159EEB9" docLanguage="en" docName="zt03344p068.pdf" docOrigin="Zootaxa 3344" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Salmoneus pinguis Komai & Anker, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="FFDBFFF48C7E7605FFB9A10DE03CEB5D" masterDocTitle="A distinctive new species of the alpheid genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the northwestern Pacific" masterLastPageNumber="68" masterPageNumber="60" pageNumber="61" updateTime="1698309771148" updateUser="plazi">
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<mods:titleInfo id="6C52723ED60FCB700DDD7E912F3D95A9">
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<mods:title id="120C81BAD42A05A535369CA25E031692">A distinctive new species of the alpheid genus Salmoneus Holthuis, 1955 (Crustacea: Decapoda: Caridea) from the northwestern Pacific</mods:title>
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<mods:name id="7194BB5DCB952811A28853C7F4765606" type="personal">
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<mods:namePart id="24998AA34EE5784E6EA33D74307BC571">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="5D78F7D5EE6056D580D7A0BEEF79AE38">Anker, Arthur</mods:namePart>
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<mods:typeOfResource id="2BE90076D9909F3FB9449FDCD9FBC7DC">text</mods:typeOfResource>
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<mods:title id="50DDC05637A218BD590A59A630824988">Zootaxa</mods:title>
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<mods:part id="AA700795093C7EF30173AD6A03687FE8">
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<mods:date id="6FD8443CDE36528378554DCAA01FB17A">2012</mods:date>
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<mods:detail id="30C5D07F5EBD8CC73F8F472BA45F3E85" type="volume">
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<mods:number id="B34425774DA926E0C741CFDB243A0E90">3344</mods:number>
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<mods:classification id="41D07D2AC2530B11524A1D084BECBACB">journal article</mods:classification>
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<mods:identifier id="E8FA9A2D597C7EE92E1E01513D727492" type="DOI">10.5281/zenodo.281468</mods:identifier>
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<mods:identifier id="1CDF4E6FBD0ED23EFDBB37A89B6C1A58" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DF9E8D4D244518B990B68B40BC11A2C5" type="Zenodo-Dep">281468</mods:identifier>
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<treatment id="03E2878C8C7F7602FF2EA1D0E159EEB9" ID-DOI="http://doi.org/10.5281/zenodo.6167410" ID-GBIF-Taxon="119519043" ID-Zenodo-Dep="6167410" LSID="urn:lsid:plazi:treatment:03E2878C8C7F7602FF2EA1D0E159EEB9" httpUri="http://treatment.plazi.org/id/03E2878C8C7F7602FF2EA1D0E159EEB9" lastPageId="7" lastPageNumber="67" pageId="1" pageNumber="61">
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<subSubSection id="C35165118C7F7604FF2EA1D0E0C3EA4A" pageId="1" pageNumber="61" type="nomenclature">
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<paragraph id="8BF4369A8C7F7604FF2EA1D0E1DEEBAA" blockId="1.[151,482,221,279]" box="[151,482,221,247]" pageId="1" pageNumber="61">
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<heading id="D0BC81F68C7F7604FF2EA1D0E1DEEBAA" bold="true" box="[151,482,221,247]" fontSize="11" level="1" pageId="1" pageNumber="61" reason="1">
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<emphasis id="B93FEA888C7F7604FF2EA1D0E1DEEBAA" bold="true" box="[151,482,221,247]" pageId="1" pageNumber="61">
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<taxonomicName id="4C4B4D198C7F7604FF2EA1D0E142EBAA" ID-CoL="85Y2M" box="[151,382,221,247]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="61" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
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<emphasis id="B93FEA888C7F7604FF2EA1D0E142EBAA" bold="true" box="[151,382,221,247]" italics="true" pageId="1" pageNumber="61">Salmoneus pinguis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20C57F38C7F7604FE3DA1D0E1DEEBAA" box="[388,482,221,247]" pageId="1" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF4369A8C7F7604FF2EA1F2E0C3EA4A" blockId="1.[151,482,221,279]" box="[151,255,255,279]" pageId="1" pageNumber="61">
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<figureCitation id="13702A1F8C7F7604FF2EA1F2E0C3EA4A" box="[151,255,255,279]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="1.[151,250,1811,1834]" captionStartId-1="3.[151,250,1905,1928]" captionStartId-2="4.[151,250,1774,1797]" captionStartId-3="5.[151,250,1853,1876]" captionStartId-4="6.[151,250,1044,1067]" captionStartId-5="6.[151,250,1641,1664]" captionTargetBox-0="[159,1429,1173,1779]" captionTargetBox-1="[230,1357,206,1874]" captionTargetBox-2="[182,1422,228,1694]" captionTargetBox-3="[239,1337,202,1815]" captionTargetBox-4="[151,1435,193,1022]" captionTargetBox-5="[159,1424,1156,1602]" captionTargetId-0="figure@1.[151,1436,1161,1791]" captionTargetId-1="figure@3.[211,1374,193,1901]" captionTargetId-2="figure@4.[151,1436,193,1723]" captionTargetId-3="figure@5.[218,1369,193,1832]" captionTargetId-4="figure@6.[151,1436,193,1023]" captionTargetId-5="figure@6.[151,1436,1147,1619]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionText-0="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." captionText-1="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." captionText-2="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." captionText-3="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." captionText-4="FIGURE 5. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, habitus and colour in life. Photograph courtesy of Ryo Minemizu." captionText-5="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/281469/files/figure.png" httpUri-1="https://zenodo.org/record/281470/files/figure.png" httpUri-2="https://zenodo.org/record/281471/files/figure.png" httpUri-3="https://zenodo.org/record/281472/files/figure.png" httpUri-4="https://zenodo.org/record/281473/files/figure.png" httpUri-5="https://zenodo.org/record/281474/files/figure.png" pageId="1" pageNumber="61">Figs. 1–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35165118C7F7604FF2EA045E4BFEAB2" pageId="1" pageNumber="61" type="materials_examined">
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<paragraph id="8BF4369A8C7F7604FF2EA045E53CEAD9" blockId="1.[151,1437,327,1108]" pageId="1" pageNumber="61">
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<emphasis id="B93FEA888C7F7604FF2EA045E17FEA3D" bold="true" box="[151,323,327,352]" pageId="1" pageNumber="61">
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<typeStatus id="54F088388C7F7604FF2EA045E0ECEA3D" box="[151,208,328,352]" pageId="1" pageNumber="61">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54F088388C7F7604FEF4A04AE186EA02" box="[333,442,327,351]" pageId="1" pageNumber="61" type="holotype">Holotype</typeStatus>
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: ovigerous specimen (cl
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<quantity id="4CB39B7F8C7F7604FD6AA04AE312EA02" box="[723,814,327,351]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="61" unit="mm" value="5.7">5.7 mm</quantity>
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),
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<collectingCountry id="F35C760A8C7F7604FCF8A045E3B8EA02" box="[833,900,328,351]" name="Japan" pageId="1" pageNumber="61">Japan</collectingCountry>
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, Ryukyu Archipelago, Okinawa Island, Manza, depth
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<quantity id="4CB39B7F8C7F7604FF65A061E135EAD9" box="[220,265,364,388]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="61" unit="m" value="2.0">2 m</quantity>
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, cracking of coral rocks with a hammer,
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<date id="FFF5105A8C7F7604FD6BA061E36AEAD9" box="[722,854,364,388]" pageId="1" pageNumber="61" value="2011-07-01">1 July 2011</date>
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, coll. N. Shirakawa, CBM-ZC 10666.
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</paragraph>
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<paragraph id="8BF4369A8C7F7604FF70A082E4BFEAB2" blockId="1.[151,1437,327,1108]" pageId="1" pageNumber="61">
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<emphasis id="B93FEA888C7F7604FF70A082E203EAF5" bold="true" box="[201,575,399,424]" pageId="1" pageNumber="61">Additional material examined.</emphasis>
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1 non-ovigerous specimen (cl 3.0 mm),
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<collectingCountry id="F35C760A8C7F7604FB9FA082E49BEAFA" box="[1062,1191,399,423]" name="Philippines" pageId="1" pageNumber="61">Philippines</collectingCountry>
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, Bohol Sea, Panglao Island, Station B8, Napaling,
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<geoCoordinate id="EE7F505D8C7F7604FE5BA0B9E274EA91" box="[482,584,436,460]" direction="north" orientation="latitude" pageId="1" pageNumber="61" precision="92" value="9.618333">9°37.1'N</geoCoordinate>
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,
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<geoCoordinate id="EE7F505D8C7F7604FDEDA0B9E2EEEA91" box="[596,722,436,460]" direction="east" orientation="longitude" pageId="1" pageNumber="61" precision="92" value="123.76833">123°46.1'E</geoCoordinate>
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, depth
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<quantity id="4CB39B7F8C7F7604FC9AA0B9E36CEA91" box="[803,848,436,460]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="61" unit="m" value="3.0">3 m</quantity>
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, subtidal reef platform, dead coral and rubble brushing,
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<date id="FFF5105A8C7F7604FF70A0D5E16FEAB2" box="[201,339,471,495]" pageId="1" pageNumber="61" value="2004-06-07">7 June 2004</date>
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, coll. Panglao 2004 Marine Biodiversity Project, MNHN-IU-2010-5564.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35165118C7F7602FF70A0F6E309EADA" lastPageId="7" lastPageNumber="67" pageId="1" pageNumber="61" type="description">
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<paragraph id="8BF4369A8C7F7604FF70A0F6E44FE99A" blockId="1.[151,1437,327,1108]" pageId="1" pageNumber="61">
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<emphasis id="B93FEA888C7F7604FF70A0F6E168E949" bold="true" box="[201,340,507,532]" pageId="1" pageNumber="61">Description</emphasis>
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(based on
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<typeStatus id="54F088388C7F7604FE6CA0F1E200E949" box="[469,572,508,532]" pageId="1" pageNumber="61" type="holotype">holotype</typeStatus>
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). Body stout, bulky, plump, with relatively soft integuments. Carapace glabrous, conspicuously domed dorsally, steeply sloping anteriorly into rostrum; anterolateral region without distinct suture, however, with shallow notch on anterolateral margin dorsal to rounded pterygostomial angle (
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<figureCitation id="13702A1F8C7F7604FB47A349E503E901" box="[1278,1343,580,604]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61">Fig. 1</figureCitation>
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); posterior margin with deep cardiac notch. Rostrum very short, broadly triangular, with rounded tip, reaching distal margin of second article of antennular peduncle, strongly deflexed; dorsal surface smooth, without a distinct carina; ventral surface flattened, unarmed (
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<figureCitation id="13702A1F8C7F7604FD9DA3A2E24EE99A" box="[548,626,687,711]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61">Figs. 1</figureCitation>
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,
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<figureCitation id="13702A1F8C7F7604FD39A3A2E2B2E99A" box="[640,654,687,711]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">2</figureCitation>
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A, C). Orbital teeth absent (
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<figureCitation id="13702A1F8C7F7604FC7DA3A2E42EE99A" box="[964,1042,687,711]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61">Figs. 1</figureCitation>
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,
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<figureCitation id="13702A1F8C7F7604FB99A3A2E412E99A" box="[1056,1070,687,711]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">2</figureCitation>
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A, C).
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</paragraph>
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<paragraph id="8BF4369A8C7F7604FF70A3D9E2DAE8B5" blockId="1.[151,1437,327,1108]" pageId="1" pageNumber="61">
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Pleura of first four abdominal somites enlarged, rounded posteroventrally; fourth pleuron greatly overlapping fifth pleuron, latter with rounded posteroventral angle (
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<figureCitation id="13702A1F8C7F7604FCB0A3FAE36BE852" box="[777,855,759,783]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61">Figs. 1</figureCitation>
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,
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<figureCitation id="13702A1F8C7F7604FCDFA3FAE349E852" box="[870,885,759,783]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">2</figureCitation>
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A); sixth somite without distinct articulated flap, with very faint suture only (
|
||
<figureCitation id="13702A1F8C7F7604FE65A211E21EE869" box="[476,546,796,820]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">Fig. 2</figureCitation>
|
||
D); preanal plate rounded. Telson relatively broad, widest at base, posteriorly tapering; dorsal surface with two pairs of minute spiniform setae inserted at some distance from lateral margin, located at about 0.63 and 0.82 telson length, respectively; posterior margin approximately half as long as anterior margin, with moderately deep, U-shaped, median incision, two pairs of long spiniform setae on either side of median incision (mesial ones slightly longer and stouter than lateral ones), and one pair of long stiff setae arising from lateral margins of median incision (
|
||
<figureCitation id="13702A1F8C7F7604FDDBA2DDE299E8B5" box="[610,677,976,1000]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">Fig. 2</figureCitation>
|
||
B, E).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7F7604FF70A2F9E0D8EF09" blockId="1.[151,1437,327,1108]" pageId="1" pageNumber="61">
|
||
Eyestalks stout, short, concealed in dorsal view, fully visible in lateral view (
|
||
<figureCitation id="13702A1F8C7F7604FB9DA2F9E44DEF51" box="[1060,1137,1012,1036]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13702A1F8C7F7604FBC7A2F9E4B0EF51" box="[1150,1164,1012,1036]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">2</figureCitation>
|
||
A, C); cornea somewhat reduced, occupying only dorsolateral part of eyestalk (
|
||
<figureCitation id="13702A1F8C7F7604FCA1A515E35CEF6D" box="[792,864,1048,1072]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="1" pageNumber="61">Fig. 2</figureCitation>
|
||
F). Epistomial sclerites without conspicuous processes.
|
||
</paragraph>
|
||
<caption id="DF3466128C7F7604FF2EA61EE118EC3A" httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="1" pageNumber="61" targetBox="[159,1429,1173,1779]" targetPageId="1">
|
||
<paragraph id="8BF4369A8C7F7604FF2EA61EE118EC3A" blockId="1.[151,1436,1811,1895]" pageId="1" pageNumber="61">
|
||
<emphasis id="B93FEA888C7F7604FF2EA61EE131EC74" bold="true" box="[151,269,1811,1834]" pageId="1" pageNumber="61">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C7F7604FEA5A619E1E0EC77" box="[284,476,1812,1834]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="61" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C7F7604FEA5A619E1E0EC77" box="[284,476,1812,1834]" italics="true" pageId="1" pageNumber="61">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C7F7604FE5DA619E20BEC74" box="[484,567,1812,1833]" pageId="1" pageNumber="61" rank="species">
|
||
<emphasis id="B93FEA888C7F7604FE5DA619E20BEC74" bold="true" box="[484,567,1812,1833]" pageId="1" pageNumber="61">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4369A8C7F7607FF70A682E456EA62" blockId="1.[151,1436,1935,2032]" lastBlockId="2.[151,1437,151,2012]" lastPageId="2" lastPageNumber="62" pageId="1" pageNumber="61">
|
||
Antennular peduncles widely separated basally, short, stout, with second article slightly wider than long; first article without distinct tooth on ventromesial carina; stylocerite with subacute tip reaching distal margin of second article; lateral flagellum biramous, fused portion short, with two or three very short articles; accessory ramus consisting of four articles with aesthetascs; both flagella setose (
|
||
<figureCitation id="13702A1F8C7C7607FC86A19AE3BEEBF2" box="[831,898,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="2" pageNumber="62">Fig. 2</figureCitation>
|
||
C). Antenna with basicerite ending in short subacute tooth distolaterally, with prominent, blunt process distodorsally; scaphocerite more rounded than ovate, short, 1.4 times longer than wide, reaching distal margin of third article of antennular peduncle; distal margin of blade broad, rounded, reaching well beyond small, subacute, distolateral tooth; carpocerite very short, stout, barely reaching mid-length of scaphocerite; flagellum conspicuously setose (
|
||
<figureCitation id="13702A1F8C7C7607FC1FA02AE3CFEA62" box="[934,1011,295,319]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="2" pageNumber="62">Figs. 2</figureCitation>
|
||
A, C; 3A).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7C7607FF70A041E40AE9B2" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="62">
|
||
Mouthparts fairly typical for genus. Mandible with moderately strong molar process bearing setal rows on dorsal surface; incisor process strong, distal margin with seven teeth, latter unequal in size; palp flattened, with oval distal article (
|
||
<figureCitation id="13702A1F8C7C7607FE8AA099E149EAF1" box="[307,373,404,428]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
B–E). Maxillule with slender, inwardly curved coxal endite; basial endite curved, armed with two rows of slender spinules on distal margin; endopod strongly curved, ventral lobe bearing one stiff apical seta, dorsal lobe broad, rounded, with smaller setae (
|
||
<figureCitation id="13702A1F8C7C7607FD10A0D1E2D6EAA9" box="[681,746,476,500]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
F, G). Maxilla comparatively small; coxal endite not reaching basial endite; basial endite divided into two subequal, subquadrate lobes; endopod stout, tapering distally; scaphognathite with short, rounded posterior lobe (
|
||
<figureCitation id="13702A1F8C7C7607FD3BA329E2FAE961" box="[642,710,548,572]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
H). First maxilliped with a few short setae on mesial margin of coxal endite; basial endite distally rounded; endopod biarticulated, consisting of broad basal article and slender, distally tapering distal article; exopod with narrow caridean lobe, flagellum well developed; epipod moderately large, subquadrate (
|
||
<figureCitation id="13702A1F8C7C7607FECAA382E189E9FA" box="[371,437,655,679]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">
|
||
Fig.
|
||
<date id="FFF5105A8C7C7607FE1EA382E189E9FA" box="[423,437,655,679]" pageId="2" pageNumber="62">3</date>
|
||
</figureCitation>
|
||
I, J). Second maxilliped with prominently produced mesial margin of coxa; propodus and dactylus apparently partly fused (articulation distinct only dorsally); exopodal flagellum well developed, with one long, stiff seta on lateral margin proximal to mid-length; epipod suboval (
|
||
<figureCitation id="13702A1F8C7C7607FC76A3DAE42DE9B2" box="[975,1041,727,751]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
K).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7C7607FF70A3F1E2DEE89A" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="62">
|
||
Third maxilliped slender, falling short of distal margin of antennal scale when fully extended; coxa with ovate lateral plate above mastigobranch; antepenultimate article longer than penultimate and ultimate articles combined; penultimate article about 2.5 times as long as wide, slightly widening distally, with two transverse rows of serrulate setae distomesially; ultimate article tapering distally, with transverse rows of thick serrulate setae mesially, apex with at least one small spiniform seta; exopod falling short of distal margin of antepenultimate article (
|
||
<figureCitation id="13702A1F8C7C7607FAADA281E56BE8F9" box="[1300,1367,908,932]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
A–C); arthrobranch well developed (not drawn in
|
||
<figureCitation id="13702A1F8C7C7607FDC5A2A2E282E89A" box="[636,702,943,967]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7C7607FF70A2D9E2CDEEC1" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="62">
|
||
Chelipeds short relative to body and other pereopods (
|
||
<figureCitation id="13702A1F8C7C7607FC96A2D9E34EE8B1" box="[815,882,980,1004]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="2" pageNumber="62">Fig. 1</figureCitation>
|
||
), slightly unequal in size, asymmetrical in shape (
|
||
<figureCitation id="13702A1F8C7C7607FF26A2FAE0DEEF52" box="[159,226,1015,1039]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
D, F); minor (right) cheliped about 0.9 length of major (left) cheliped, with stouter ischium and merus, comparatively longer carpus, and significantly smaller and shorter chela (
|
||
<figureCitation id="13702A1F8C7C7607FC24A511E3E3EF69" box="[925,991,1052,1076]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
F, G); both chelipeds capable of folding under cephalothorax when not in use (therefore mostly invisible in
|
||
<figureCitation id="13702A1F8C7C7607FC29A532E3E8EF0A" box="[912,980,1087,1111]" captionStart="FIGURE 5" captionStartId="6.[151,250,1044,1067]" captionTargetBox="[151,1435,193,1022]" captionTargetId="figure@6.[151,1436,193,1023]" captionTargetPageId="6" captionText="FIGURE 5. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, habitus and colour in life. Photograph courtesy of Ryo Minemizu." httpUri="https://zenodo.org/record/281473/files/figure.png" pageId="2" pageNumber="62">Fig. 5</figureCitation>
|
||
). Major cheliped reaching beyond distal margin of antennal scale, slender; ischium not widening distally, unarmed on ventrolateral surface; merus about 5.2 times as long as wide, smooth, compressed laterally; carpus short, cup-shaped, about 0.3 merus length; chela somewhat compressed, smooth, mesial margin slightly concave; palm with deep proximal constriction; fingers distinctly longer than palm, terminating in blunt, slightly bulbous, curving and crossing tips; cutting edges of both dactylus and pollex with row of low, blunt teeth (
|
||
<figureCitation id="13702A1F8C7C7607FDECA5F9E2A4EE51" box="[597,664,1268,1292]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
D, E). Minor cheliped with ischium lacking spiniform seta on ventrolateral surface; merus about 3.5 times as long as wide, smooth, dorsal margin slightly convex, ventral surface shallowly sulcate in distal half; carpus elongate, slightly shorter than merus, distally widening; chela smooth, simple, with slightly swollen palm; fingers moderately stout, in length subequal to palm; cutting edges smooth, unarmed; tips slightly curved, not strongly crossing (
|
||
<figureCitation id="13702A1F8C7C7607FDCCA489E28DEEC1" box="[629,689,1412,1436]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig 4</figureCitation>
|
||
F, G).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7C7607FF70A4A5E2F2EC11" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="62">
|
||
Second pereopod slender; ischium without spiniform setae; merus slightly longer than ischium; carpus fourarticulated (but see under Variation), ratio of carpal articles (from proximal to distal) approximately equal to: 2.5: 0.4: 0.5: 1.0; chela simple, slightly longer than distal-most carpal article (
|
||
<figureCitation id="13702A1F8C7C7607FC77A4FDE42DED55" box="[974,1041,1520,1544]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
H). Third pereopod relatively stout, compressed laterally; ischium with two tiny spiniform setae on ventrolateral surface; merus about 3.2 times as long as wide, unarmed, with stiff setae along on dorsal margin; carpus more slender than merus, about 0.8 merus length, with stiff setae on dorsal surface; propodus with one spiniform seta on ventral margin distal to mid-length and one pair of stouter, longer spiniform setae on distoventral margin, dorsal margin with rows of stiff setae; dactylus about 0.4 times as long as propodus, simple, moderately slender, slightly curving distally, unguis clearly demarcated (
|
||
<figureCitation id="13702A1F8C7C7607FAD7A7A9E09BEDBD" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">
|
||
Fig.
|
||
<date id="FFF5105A8C7C7607FF2EA7C5E09BEDBD" box="[151,167,1736,1760]" pageId="2" pageNumber="62">4</date>
|
||
</figureCitation>
|
||
I, J). Fourth pereopod generally very similar to third pereopod (
|
||
<figureCitation id="13702A1F8C7C7607FC3CA7C5E3F0EDBD" box="[901,972,1736,1760]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
K). Fifth pereopod in length subequal to third pereopod, however, with longer carpus and propodus combined; ischium unarmed; merus in length subequal to carpus; propodus with two spiniform setae and about five rows of short, serrulate setae in distal 0.4, distoventral margin with two slender spiniform setae (
|
||
<figureCitation id="13702A1F8C7C7607FDD0A639E291EC11" box="[617,685,1844,1868]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="2" pageNumber="62">Fig. 4</figureCitation>
|
||
L).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C7C7607FF70A655E101EC81" blockId="2.[151,1437,151,2012]" pageId="2" pageNumber="62">
|
||
Second pleopod with appendix masculina about half-length of appendix interna, its apex and mesial margin being furnished with stiff setae (
|
||
<figureCitation id="13702A1F8C7C7607FDB8A671E27FECC9" box="[513,579,1916,1940]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
L). Uropod with lateral lobe of protopod unarmed; endopod and exopod narrowly ovoid; diaeresis on exopod not distinct; exopod with minute distolateral tooth adjacent to slender spiniform seta (
|
||
<figureCitation id="13702A1F8C7C7607FF6BA6C9E12FEC81" box="[210,275,1988,2012]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="2" pageNumber="62">Fig. 3</figureCitation>
|
||
M).
|
||
</paragraph>
|
||
<caption id="DF3466128C7D7606FF2EA67CE31DECBE" httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="3" pageNumber="63" targetBox="[230,1357,206,1874]" targetPageId="3">
|
||
<paragraph id="8BF4369A8C7D7606FF2EA67CE31DECBE" blockId="3.[151,1436,1905,2019]" pageId="3" pageNumber="63">
|
||
<emphasis id="B93FEA888C7D7606FF2EA67CE137ECDA" bold="true" box="[151,267,1905,1928]" pageId="3" pageNumber="63">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C7D7606FEA1A67CE1EAECDA" box="[280,470,1905,1927]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C7D7606FEA1A67CE1EAECDA" box="[280,470,1905,1927]" italics="true" pageId="3" pageNumber="63">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C7D7606FE64A67FE211ECDA" box="[477,557,1906,1927]" pageId="3" pageNumber="63" rank="species">
|
||
<emphasis id="B93FEA888C7D7606FE64A67FE211ECDA" bold="true" box="[477,557,1906,1927]" pageId="3" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D–F.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466128C7A7601FF2EA7E3E487EC22" httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="4" pageNumber="64" targetBox="[182,1422,228,1694]" targetPageId="4">
|
||
<paragraph id="8BF4369A8C7A7601FF2EA7E3E487EC22" blockId="4.[151,1436,1774,1919]" pageId="4" pageNumber="64">
|
||
<emphasis id="B93FEA888C7A7601FF2EA7E3E131EC59" bold="true" box="[151,269,1774,1797]" pageId="4" pageNumber="64">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C7A7601FEA5A7E2E1E0EC58" box="[284,476,1775,1797]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="64" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C7A7601FEA5A7E2E1E0EC58" box="[284,476,1775,1797]" italics="true" pageId="4" pageNumber="64">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C7A7601FE5DA7E2E20BEC59" box="[484,567,1775,1796]" pageId="4" pageNumber="64" rank="species">
|
||
<emphasis id="B93FEA888C7A7601FE5DA7E2E20BEC59" bold="true" box="[484,567,1775,1796]" pageId="4" pageNumber="64">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466128C7B7600FF2EA630E11FECB1" httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="5" pageNumber="65" targetBox="[239,1337,202,1815]" targetPageId="5">
|
||
<paragraph id="8BF4369A8C7B7600FF2EA630E11FECB1" blockId="5.[151,1436,1853,2028]" pageId="5" pageNumber="65">
|
||
<emphasis id="B93FEA888C7B7600FF2EA630E137EC0F" bold="true" box="[151,267,1853,1876]" pageId="5" pageNumber="65">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C7B7600FEA1A630E1EAEC0E" box="[280,470,1853,1875]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="65" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C7B7600FEA1A630E1EAEC0E" box="[280,470,1853,1875]" italics="true" pageId="5" pageNumber="65">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C7B7600FE64A633E211EC0E" box="[477,557,1854,1875]" pageId="5" pageNumber="65" rank="species">
|
||
<emphasis id="B93FEA888C7B7600FE64A633E211EC0E" bold="true" box="[477,557,1854,1875]" pageId="5" pageNumber="65">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466128C787603FF2EA519E30EEF14" httpUri="https://zenodo.org/record/281473/files/figure.png" pageId="6" pageNumber="66" targetBox="[151,1435,193,1022]" targetPageId="6">
|
||
<paragraph id="8BF4369A8C787603FF2EA519E30EEF14" blockId="6.[151,1436,1044,1097]" pageId="6" pageNumber="66">
|
||
<emphasis id="B93FEA888C787603FF2EA519E131EF77" bold="true" box="[151,269,1044,1067]" pageId="6" pageNumber="66">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C787603FEA5A518E1E0EF76" box="[284,476,1045,1067]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C787603FEA5A518E1E0EF76" box="[284,476,1045,1067]" italics="true" pageId="6" pageNumber="66">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C787603FE5DA518E20BEF77" box="[484,567,1045,1066]" pageId="6" pageNumber="66" rank="species">
|
||
<emphasis id="B93FEA888C787603FE5DA518E20BEF77" bold="true" box="[484,567,1045,1066]" pageId="6" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, habitus and colour in life. Photograph courtesy of Ryo Minemizu.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3466128C787603FF2EA764E5B6EDE1" httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="6" pageNumber="66" targetBox="[159,1424,1156,1602]" targetPageId="6">
|
||
<paragraph id="8BF4369A8C787603FF2EA764E5B6EDE1" blockId="6.[151,1436,1641,1724]" pageId="6" pageNumber="66">
|
||
<emphasis id="B93FEA888C787603FF2EA764E12DED23" bold="true" box="[151,273,1641,1664]" pageId="6" pageNumber="66">FIGURE 6</emphasis>
|
||
.
|
||
<taxonomicName id="4C4B4D198C787603FE9BA764E1DAED22" box="[290,486,1641,1663]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="66" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C787603FE9BA764E1DAED22" box="[290,486,1641,1663]" italics="true" pageId="6" pageNumber="66">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C787603FE48A767E27AED22" box="[497,582,1642,1663]" pageId="6" pageNumber="66" rank="species">
|
||
<emphasis id="B93FEA888C787603FE48A767E27AED22" bold="true" box="[497,582,1642,1663]" pageId="6" pageNumber="66">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU-2010-5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF4369A8C787603FF70A7E4E246EC5F" blockId="6.[151,1436,1769,2009]" box="[201,634,1769,1794]" pageId="6" pageNumber="66">Gill-exopod formula typical for genus.</paragraph>
|
||
<paragraph id="8BF4369A8C787602FF70A600E309EADA" blockId="6.[151,1436,1769,2009]" lastBlockId="7.[151,1437,151,1508]" lastPageId="7" lastPageNumber="67" pageId="6" pageNumber="66">
|
||
<emphasis id="B93FEA888C787603FF70A600E153EC7B" bold="true" box="[201,367,1805,1830]" pageId="6" pageNumber="66">Colour in life.</emphasis>
|
||
Semitransparent, with yellowish tinge; ovary and eggs bright yellow-orange (
|
||
<figureCitation id="13702A1F8C787603FB61A600E520EC78" box="[1240,1308,1805,1829]" captionStart="FIGURE 5" captionStartId="6.[151,250,1044,1067]" captionTargetBox="[151,1435,193,1022]" captionTargetId="figure@6.[151,1436,193,1023]" captionTargetPageId="6" captionText="FIGURE 5. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, habitus and colour in life. Photograph courtesy of Ryo Minemizu." httpUri="https://zenodo.org/record/281473/files/figure.png" pageId="6" pageNumber="66">Fig. 5</figureCitation>
|
||
).
|
||
<emphasis id="B93FEA888C787603FF70A63DE102EC14" bold="true" box="[201,318,1840,1865]" pageId="6" pageNumber="66">Variation.</emphasis>
|
||
The apparently post-ovigerous specimen from Panglao is much smaller than the
|
||
<typeStatus id="54F088388C787603FB5EA63CE575EC14" box="[1255,1353,1841,1865]" pageId="6" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
(cl 3.0 mm compared to cl
|
||
<quantity id="4CB39B7F8C787603FECDA658E1F6EC30" box="[372,458,1877,1901]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="66" unit="mm" value="5.7">5.7 mm</quantity>
|
||
in the
|
||
<typeStatus id="54F088388C787603FDACA658E245EC30" box="[533,633,1877,1901]" pageId="6" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
) and differs from the latter by the more triangular rostrum (
|
||
<figureCitation id="13702A1F8C787603FAB4A658E572EC30" box="[1293,1358,1877,1901]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="6" pageNumber="66">Fig. 6</figureCitation>
|
||
A), the eyestalks being more concealed in lateral view (
|
||
<figureCitation id="13702A1F8C787603FD0EA674E2C7ECCC" box="[695,763,1913,1937]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="6" pageNumber="66">Fig. 6</figureCitation>
|
||
B), the second pereopods with a five-articulated carpus (vs. four-articulated in the
|
||
<typeStatus id="54F088388C787603FE36A690E1CFECE8" box="[399,499,1949,1973]" pageId="6" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="13702A1F8C787603FDB0A690E270ECE8" box="[521,588,1949,1973]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="6" pageNumber="66">Fig. 6</figureCitation>
|
||
D), and the third pereopod with a somewhat more slender dactylus, and with two spiniform setae on the ventral margin of the propodus (vs. one in the
|
||
<typeStatus id="54F088388C787603FC6AA6CCE40BEC84" box="[979,1079,1985,2009]" pageId="6" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="13702A1F8C787603FBF7A6CCE4AEEC84" box="[1102,1170,1985,2009]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="6" pageNumber="66">Fig. 6</figureCitation>
|
||
E). In all other features, this specimen agrees well with the
|
||
<typeStatus id="54F088388C797602FDA4A19AE2BEEBF2" box="[541,642,151,175]" pageId="7" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
, including all of the new species’ diagnostic features, such as the stout, plump body; the general shape of the frontal margin of the carapace (including the absence of orbital teeth); the general shape and proportions of the antennules and antennae, including their setose flagella (
|
||
<figureCitation id="13702A1F8C797602FB08A1D2E4C9EBAA" box="[1201,1269,223,247]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="7" pageNumber="67">Fig. 6</figureCitation>
|
||
A); the general shape and details of the major cheliped, including the very diagnostic blunt fingertips of the major chela (
|
||
<figureCitation id="13702A1F8C797602FA8FA009E545EA41" box="[1334,1401,260,284]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="7" pageNumber="67">Fig. 6</figureCitation>
|
||
C); the proportions and ischial armature of the third pereopod (
|
||
<figureCitation id="13702A1F8C797602FC8DA02AE344EA62" box="[820,888,295,319]" captionStart="FIGURE 6" captionStartId="6.[151,250,1641,1664]" captionTargetBox="[159,1424,1156,1602]" captionTargetId="figure@6.[151,1436,1147,1619]" captionTargetPageId="6" captionText="FIGURE 6. Salmoneus pinguis sp. nov., non-type, post-ovigerous specimen (cl 3.0 mm) from Panglao, the Philippines, MNHN-IU- 2010 - 5564, A, frontal margin of carapace and frontal appendages, dorsal view; B, frontal margin of carapace, lateral view; C, major chela, lateral view; D, second pereopod, lateral view; E, third pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281474/files/figure.png" pageId="7" pageNumber="67">Fig. 6</figureCitation>
|
||
E), and the general shape and dorsal armature of the telson. Therefore, the Panglao specimen is tentatively assigned to
|
||
<taxonomicName id="4C4B4D198C797602FC05A040E410EA39" box="[956,1068,333,356]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FC05A040E410EA39" box="[956,1068,333,356]" italics="true" pageId="7" pageNumber="67">S. pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FB8FA041E4AEEA39" box="[1078,1170,332,356]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FB8FA041E4AEEA39" bold="true" box="[1078,1170,332,356]" pageId="7" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, awaiting collection of more specimens in both the
|
||
<collectingCountry id="F35C760A8C797602FE6BA062E26CEADA" box="[466,592,367,391]" name="Philippines" pageId="7" pageNumber="67">Philippines</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="F35C760A8C797602FD54A07DE30DEADA" box="[749,817,368,391]" name="Japan" pageId="7" pageNumber="67">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165118C797602FF70A099E0E2EA92" pageId="7" pageNumber="67" type="etymology">
|
||
<paragraph id="8BF4369A8C797602FF70A099E0E2EA92" blockId="7.[151,1437,151,1508]" pageId="7" pageNumber="67">
|
||
<emphasis id="B93FEA888C797602FF70A099E173EAF1" bold="true" box="[201,335,404,428]" pageId="7" pageNumber="67">Etymology.</emphasis>
|
||
From the Latin word “
|
||
<taxonomicName id="4C4B4D198C797602FDF6A099E294EAF1" box="[591,680,404,428]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis">pinguis</taxonomicName>
|
||
”, meaning bulky, plump, referring to the new species’ general body shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165118C797602FF70A0D1E3F9EAA9" box="[201,965,475,500]" pageId="7" pageNumber="67" type="materials_examined">
|
||
<paragraph id="8BF4369A8C797602FF70A0D1E3F9EAA9" blockId="7.[151,1437,151,1508]" box="[201,965,475,500]" pageId="7" pageNumber="67">
|
||
<emphasis id="B93FEA888C797602FF70A0D1E159EAA9" bold="true" box="[201,357,475,500]" pageId="7" pageNumber="67">
|
||
<typeStatus id="54F088388C797602FF70A0D1E13EEAA9" box="[201,258,476,500]" pageId="7" pageNumber="67">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Manza, Okinawa Island, Ryukyu Archipelago,
|
||
<collectingCountry id="F35C760A8C797602FCC4A0D0E3FDEAA9" box="[893,961,477,500]" name="Japan" pageId="7" pageNumber="67">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165118C797602FF70A0F2E4F9E9FA" pageId="7" pageNumber="67" type="distribution">
|
||
<paragraph id="8BF4369A8C797602FF70A0F2E311E961" blockId="7.[151,1437,151,1508]" pageId="7" pageNumber="67">
|
||
<emphasis id="B93FEA888C797602FF70A0F2E15EE945" bold="true" box="[201,354,511,536]" pageId="7" pageNumber="67">Distribution.</emphasis>
|
||
Tropical northwestern Pacific: presently known only from the Ryukyu Archipelago,
|
||
<collectingCountry id="F35C760A8C797602FAEDA30DE5ABE94A" box="[1364,1431,512,535]" name="Japan" pageId="7" pageNumber="67">Japan</collectingCountry>
|
||
; possibly also in the Bohol Sea, the
|
||
<collectingCountry id="F35C760A8C797602FD99A329E2A2E961" box="[544,670,548,572]" name="Philippines" pageId="7" pageNumber="67">Philippines</collectingCountry>
|
||
(see above).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C797602FF70A34AE4F9E9FA" blockId="7.[151,1437,151,1508]" pageId="7" pageNumber="67">
|
||
<emphasis id="B93FEA888C797602FF70A34AE110E93D" bold="true" box="[201,300,583,608]" pageId="7" pageNumber="67">Habitat.</emphasis>
|
||
The
|
||
<typeStatus id="54F088388C797602FED4A34AE1F3E902" box="[365,463,583,607]" pageId="7" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
was extracted from a crevice in large coral rock broken apart with a hammer. No association with other invertebrates was noted in the field (N. Shirakawa and R. Minemizu, pers. comm.). The nontype specimen was collected by brushing coral rubble lifted from about
|
||
<quantity id="4CB39B7F8C797602FC02A382E3D5E9FA" box="[955,1001,655,679]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="67" unit="m" value="3.0">3 m</quantity>
|
||
, on a reef platform.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35165118C797602FF70A3B9E159EEB9" pageId="7" pageNumber="67" type="discussion">
|
||
<paragraph id="8BF4369A8C797602FF70A3B9E292EF99" blockId="7.[151,1437,151,1508]" pageId="7" pageNumber="67">
|
||
<emphasis id="B93FEA888C797602FF70A3B9E101E991" bold="true" box="[201,317,692,716]" pageId="7" pageNumber="67">Remarks.</emphasis>
|
||
In spite of its morphological distinctiveness,
|
||
<taxonomicName id="4C4B4D198C797602FC8EA3B8E398E991" box="[823,932,693,716]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FC8EA3B8E398E991" box="[823,932,693,716]" italics="true" pageId="7" pageNumber="67">S. pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FC12A3B9E438E991" box="[939,1028,692,716]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FC12A3B9E438E991" bold="true" box="[939,1028,692,716]" pageId="7" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is clearly a species of
|
||
<taxonomicName id="4C4B4D198C797602FB47A3B9E54BE991" box="[1278,1399,692,716]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA888C797602FB47A3B9E54BE991" box="[1278,1399,692,716]" italics="true" pageId="7" pageNumber="67">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
, as indicated by the possession of an appendix masculine in ovigerous specimens, the presence of a median notch on the posterior margin of the telson, the major chela with a deep proximal constriction on the palm and with serrate finger cutting edges, all features typical to
|
||
<taxonomicName id="4C4B4D198C797602FDC8A22DE2D6E865" box="[625,746,800,824]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA888C797602FDC8A22DE2D6E865" box="[625,746,800,824]" italics="true" pageId="7" pageNumber="67">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDA4B6B8C797602FD41A212E3D4E86A" author="Anker" box="[760,1000,799,823]" pageId="7" pageNumber="67" refString="Anker, A. & Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker & Marin 2006</bibRefCitation>
|
||
; Anker et al. 2006).
|
||
<taxonomicName id="4C4B4D198C797602FB73A22DE5A7E865" box="[1226,1435,800,824]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FB73A22DE5A7E865" box="[1226,1435,800,824]" italics="true" pageId="7" pageNumber="67">Salmoneus pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FF2EA249E0CCE801" box="[151,240,836,860]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FF2EA249E0CCE801" bold="true" box="[151,240,836,860]" pageId="7" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is presently the only species in the genus combining the carapace strongly domed or swollen dorsally (
|
||
<figureCitation id="13702A1F8C797602FAD7A249E098E822" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="7" pageNumber="67">Fig. 1</figureCitation>
|
||
); the frontal margin of the carapace broadly triangular, with a rounded rostrum and no trace of orbital teeth (
|
||
<figureCitation id="13702A1F8C797602FAD6A26AE09AE8F9" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="7" pageNumber="67">Fig. 2</figureCitation>
|
||
C); the second abdominal pleuron conspicuously enlarged (
|
||
<figureCitation id="13702A1F8C797602FC84A281E343E8F9" box="[829,895,908,932]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="7" pageNumber="67">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13702A1F8C797602FC34A281E3A0E8F9" box="[909,924,908,932]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="7" pageNumber="67">2</figureCitation>
|
||
A); the fourth pleuron greatly overlapping the fifth pleuron (
|
||
<figureCitation id="13702A1F8C797602FE81A2A2E140E89A" box="[312,380,943,967]" captionStart="FIGURE 1" captionStartId="1.[151,250,1811,1834]" captionTargetBox="[159,1429,1173,1779]" captionTargetId="figure@1.[151,1436,1161,1791]" captionTargetPageId="1" captionText="FIGURE 1. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, entire animal in lateral view (note: lateral surface of carapace somewhat curled up, probably caused by water absorption during preservation)." httpUri="https://zenodo.org/record/281469/files/figure.png" pageId="7" pageNumber="67">Fig. 1</figureCitation>
|
||
); the antennular and antennal flagella densely covered with long setae (
|
||
<figureCitation id="13702A1F8C797602FB02A2A2E53DE89A" box="[1211,1281,943,967]" captionStart="FIGURE 2" captionStartId="3.[151,250,1905,1928]" captionTargetBox="[230,1357,206,1874]" captionTargetId="figure@3.[211,1374,193,1901]" captionTargetPageId="3" captionText="FIGURE 2. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, entire animal, dorsal view; B, fourth to sixth abdominal somites, telson and uropods, perpendicular dorsal view; C, frontal margin and antennae, anterodorsal view; D, fifth and sixth abdominal somites, lateral view (right side); E, telson, dorsal view; F, eyes, frontal view. Scale bars: 2 mm for A, B; 1 mm for C, D – F." httpUri="https://zenodo.org/record/281470/files/figure.png" pageId="7" pageNumber="67">Fig. 2</figureCitation>
|
||
A); the major cheliped with distally blunt, slightly bulbous fingertips (
|
||
<figureCitation id="13702A1F8C797602FCA5A2D9E35DE8B1" box="[796,865,980,1004]" captionStart="FIGURE 4" captionStartId="5.[151,250,1853,1876]" captionTargetBox="[239,1337,202,1815]" captionTargetId="figure@5.[218,1369,193,1832]" captionTargetPageId="5" captionText="FIGURE 4. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666. A, left third maxilliped, lateral view; B, same, distal part of ultimate segment, mesial view; C, same, coxal lateral plate, dorsal view; D, left first pereopod (cheliped), lateral view; E, same, chela and carpus, extensor view; F, right first pereopod (cheliped), lateral view; G, same, chela, extensor view; H, left second pereopod, lateral view; I, left third pereopod, lateral view; J, same, dactylus and distal part of propodus, lateral view; K, left fourth pereopod, lateral view; L, left fifth pereopod, lateral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281472/files/figure.png" pageId="7" pageNumber="67">Fig. 4</figureCitation>
|
||
D, E); and the uropodal exopod without a distinct diaeresis (
|
||
<figureCitation id="13702A1F8C797602FEB3A2FAE172EF52" box="[266,334,1015,1039]" captionStart="FIGURE 3" captionStartId="4.[151,250,1774,1797]" captionTargetBox="[182,1422,228,1694]" captionTargetId="figure@4.[151,1436,193,1723]" captionTargetPageId="4" captionText="FIGURE 3. Salmoneus pinguis sp. nov., holotype, ovigerous specimen (cl 5.7 mm) from Okinawa, Japan, CBM-ZC 10666, left appendages. A, antenna, ventral view (distal part of flagellum omitted); B, mandible, outer view; C, same, inner view; D, same, incisor process, oblique outer view; E, same, distal article of palp; F, maxillule, outer view; G, same, distal part of endopod, inner view; H, maxilla, outer view; I, first maxilliped, outer view; J, same, endopod, inner view; K, second maxilliped, outer view; L. endopod of second pleopod, dorsal view; M. left uropod, dorsal view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/281471/files/figure.png" pageId="7" pageNumber="67">Fig. 3</figureCitation>
|
||
M). The orbital teeth are absent or greatly reduced in two other species of
|
||
<taxonomicName id="4C4B4D198C797602FB15A2F5E519EF4D" box="[1196,1317,1016,1040]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA888C797602FB15A2F5E519EF4D" box="[1196,1317,1016,1040]" italics="true" pageId="7" pageNumber="67">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C4B4D198C797602FA8FA2F5E112EF69" authority="Komai, 2009" authorityName="Komai" authorityYear="2009" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="brucei">
|
||
<emphasis id="B93FEA888C797602FA8FA2F5E5A0EF4D" box="[1334,1436,1016,1040]" italics="true" pageId="7" pageNumber="67">S. brucei</emphasis>
|
||
<bibRefCitation id="EFDA4B6B8C797602FF2EA511E112EF69" author="Komai" box="[151,302,1052,1076]" pageId="7" pageNumber="67" refString="Komai, T. (2009) A new species of the alpheid shrimp genus Salmoneus (Decapoda, Caridea) from the Ryukyu Islands, Japan, associated with a callianassid ghost shrimp (Decapoda, Thalassinidea). Crustaceana, 82, 869 - 880." type="journal article" year="2009">Komai, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D198C797602FED4A510E2B3EF69" authority="Anker, 2010" authorityName="Anker" authorityYear="2010" box="[365,655,1052,1076]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="degravei">
|
||
<emphasis id="B93FEA888C797602FED4A510E1CCEF69" box="[365,496,1052,1076]" italics="true" pageId="7" pageNumber="67">S. degravei</emphasis>
|
||
<bibRefCitation id="EFDA4B6B8C797602FE42A511E2B3EF69" author="Anker" box="[507,655,1052,1076]" pageId="7" pageNumber="67" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
; however, both differ from
|
||
<taxonomicName id="4C4B4D198C797602FC61A510E470EF69" box="[984,1100,1053,1076]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FC61A510E470EF69" box="[984,1100,1053,1076]" italics="true" pageId="7" pageNumber="67">S. pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FBEEA511E489EF69" box="[1111,1205,1052,1076]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FBEEA511E489EF69" bold="true" box="[1111,1205,1052,1076]" pageId="7" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by numerous other morphological characters and ecology (
|
||
<bibRefCitation id="EFDA4B6B8C797602FDECA532E2DAEF0A" author="Komai" box="[597,742,1087,1111]" pageId="7" pageNumber="67" refString="Komai, T. (2009) A new species of the alpheid shrimp genus Salmoneus (Decapoda, Caridea) from the Ryukyu Islands, Japan, associated with a callianassid ghost shrimp (Decapoda, Thalassinidea). Crustaceana, 82, 869 - 880." type="journal article" year="2009">Komai 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFDA4B6B8C797602FD4AA532E341EF0A" author="Anker" box="[755,893,1087,1111]" pageId="7" pageNumber="67" refString="Anker, A. (2010) The shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda, Alpheidae) in the tropical western Atlantic, with description of five new species. Zootaxa, 2372, 177 - 205." type="journal article" year="2010">Anker 2010</bibRefCitation>
|
||
), and are not closely related to the new species. The carapace of
|
||
<taxonomicName id="4C4B4D198C797602FEEFA568E280EF21" authority="Miyake & Miya, 1966" authorityName="Miyake & Miya" authorityYear="1966" box="[342,700,1124,1148]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="babai">
|
||
<emphasis id="B93FEA888C797602FEEFA568E188EF21" box="[342,436,1124,1148]" italics="true" pageId="7" pageNumber="67">S. babai</emphasis>
|
||
<bibRefCitation id="EFDA4B6B8C797602FE05A569E280EF21" author="Miyake" box="[444,700,1124,1148]" pageId="7" pageNumber="67" refString="Miyake, S. & Miya, Y. (1966) On a new species and a new record of alpheid shrimps from Japan. Journal of the Faculty of Agriculture, Kyushu University, 14, 133 - 141." type="journal article" year="1966">Miyake & Miya, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
is somewhat domed, but this species has sharp orbital teeth and additionally differs from
|
||
<taxonomicName id="4C4B4D198C797602FE17A585E220EFFD" box="[430,540,1160,1184]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FE17A585E220EFFD" box="[430,540,1160,1184]" italics="true" pageId="7" pageNumber="67">S. pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FD9AA585E247EFFD" box="[547,635,1160,1184]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FD9AA585E248EFFD" bold="true" box="[547,628,1160,1184]" pageId="7" pageNumber="67">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by the shape of the major cheliped and the much stouter dactylus on the third to fifth pereopods (
|
||
<bibRefCitation id="EFDA4B6B8C797602FE11A5A1E29DEF99" author="Miyake" box="[424,673,1196,1220]" pageId="7" pageNumber="67" refString="Miyake, S. & Miya, Y. (1966) On a new species and a new record of alpheid shrimps from Japan. Journal of the Faculty of Agriculture, Kyushu University, 14, 133 - 141." type="journal article" year="1966">Miyake & Miya 1966</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF4369A8C797602FF70A5DDE159EEB9" blockId="7.[151,1437,151,1508]" pageId="7" pageNumber="67">
|
||
The great number of autapomorphic characters of
|
||
<taxonomicName id="4C4B4D198C797602FD46A5DDE351EFB5" box="[767,877,1232,1256]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="pinguis" status="sp. nov.">
|
||
<emphasis id="B93FEA888C797602FD46A5DDE351EFB5" box="[767,877,1232,1256]" italics="true" pageId="7" pageNumber="67">S. pinguis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20C57F38C797602FCCCA5DDE3F3EFB5" box="[885,975,1232,1256]" pageId="7" pageNumber="67" rank="species">
|
||
<emphasis id="B93FEA888C797602FCCCA5DDE3F3EFB5" bold="true" box="[885,975,1232,1256]" pageId="7" pageNumber="67">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
obscures its affinities and makes it difficult to place in one of the previously established species groups of
|
||
<taxonomicName id="4C4B4D198C797602FC37A5F9E529EE51" authority="Anker & Marin 2006" authorityName="Anker & Marin" authorityYear="2006" box="[910,1301,1268,1292]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA888C797602FC37A5F9E43BEE51" box="[910,1031,1268,1292]" italics="true" pageId="7" pageNumber="67">Salmoneus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6B8C797602FBAEA5F9E531EE51" author="Anker" box="[1047,1293,1268,1292]" pageId="7" pageNumber="67" refString="Anker, A. & Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker & Marin 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. As pointed out by
|
||
<bibRefCitation id="EFDA4B6B8C797602FF53A415E1AFEE6D" author="Anker" box="[234,403,1304,1328]" pageId="7" pageNumber="67" refString="Anker, A. (2011 a) Three new species of the alpheid shrimp genus Salmoneus Holthuis, 1955 (Crustacea, Decapoda) from the tropical western Pacific. Zootaxa, 2839, 67 - 84." type="journal article" year="2011" yearSuffix="a">Anker (2011a)</bibRefCitation>
|
||
, with the rapidly increasing number of species and new character combinations, some of these species groups are becoming of rather limited usage. The intra-generic structure of
|
||
<taxonomicName id="4C4B4D198C797602FBCEA431E4CCEE09" box="[1143,1264,1340,1364]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93FEA888C797602FBCEA431E4CCEE09" box="[1143,1264,1340,1364]" italics="true" pageId="7" pageNumber="67">Salmoneus</emphasis>
|
||
</taxonomicName>
|
||
will need to be assessed with a comprehensive phylogenetic analysis based on both morphological and molecular data. However, such an analysis would only make sense after redescription of some poorly described species (e.g.,
|
||
<taxonomicName id="4C4B4D198C797602FAA9A488E16DEE9D" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="latirostris">
|
||
<emphasis id="B93FEA888C797602FAA9A488E5A7EEC1" box="[1296,1435,1412,1436]" italics="true" pageId="7" pageNumber="67">S. latirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDA4B6B8C797602FF27A4A5E174EE9D" author="Coutiere" box="[158,328,1448,1472]" pageId="7" pageNumber="67" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4B4D198C797602FE30A4A5E310EE9D" authority="Banner & Banner, 1966" authorityName="Banner & Banner" authorityYear="1966" box="[393,812,1448,1472]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="67" phylum="Arthropoda" rank="species" species="tafaongae">
|
||
<emphasis id="B93FEA888C797602FE30A4A5E22AEE9D" box="[393,534,1448,1472]" italics="true" pageId="7" pageNumber="67">S. tafaongae</emphasis>
|
||
<bibRefCitation id="EFDA4B6B8C797602FDA7A4A4E310EE9D" author="Banner" box="[542,812,1448,1472]" pageId="7" pageNumber="67" refString="Banner, A. H. & Banner, D. M. (1966) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part X. Collections from Fiji, Tonga and Samoa. Pacific Science, 20, 145 - 188." type="journal article" year="1966">Banner & Banner, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
) and description of at least 10 new taxa worldwide (A. Anker, pers. obs.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |