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<document id="D86E3CCCA1726FD042ADD08E6BB7B47E" ID-DOI="10.5281/zenodo.191273" ID-GBIF-Dataset="d2982f3f-a5f5-46ff-bbd8-a1e186245a89" ID-ISSN="1175-5326" ID-Zenodo-Dep="191273" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461125997720" checkinUser="plazi" docAuthor="Smetana, Aleš" docDate="2009" docId="03DFE029FFAEF856FF194D364D1EF932" docLanguage="en" docName="zt02286p030.pdf" docOrigin="Zootaxa 2286" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ocypus (Pseudocypus) helleni J. Muller 1926" docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="FFE69851FFAAF850FF8E4A524F65FF8E" masterDocTitle="Contributions to the knowledge of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 21. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant &amp; Rey, 1876. Section 4" masterLastPageNumber="30" masterPageNumber="1" pageNumber="5" updateTime="1698596766005" updateUser="plazi">
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<mods:title id="3C2E587FD6873AA5569D754A9B36EA12">Contributions to the knowledge of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part 21. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant &amp; Rey, 1876. Section 4</mods:title>
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<mods:namePart id="6E57DAD97111B46FC307F0F155FC4975">Smetana, Aleš</mods:namePart>
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<mods:date id="917D8F84E91C0CE1099DE89B963486AE">2009</mods:date>
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<mods:identifier id="38808370C99141D9EA79D6C08965AB9B" type="DOI">10.5281/zenodo.191273</mods:identifier>
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<treatment id="03DFE029FFAEF856FF194D364D1EF932" ID-DOI="http://doi.org/10.5281/zenodo.6217654" ID-GBIF-Taxon="119626385" ID-Zenodo-Dep="6217654" LSID="urn:lsid:plazi:treatment:03DFE029FFAEF856FF194D364D1EF932" httpUri="http://treatment.plazi.org/id/03DFE029FFAEF856FF194D364D1EF932" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C36C02B4FFAEF854FF194D364E37F82E" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BC9513FFFAEF854FF194D364DB3F8F3" blockId="4.[151,726,1891,1952]" box="[151,726,1891,1918]" pageId="4" pageNumber="5">
<heading id="D081E653FFAEF854FF194D364DB3F8F3" bold="true" box="[151,726,1891,1918]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<taxonomicName id="4C762ABCFFAEF854FF194D364DB3F8F3" ID-CoL="6SHHG" authority="J. Muller, 1926" authorityName="J. Muller" authorityYear="1926" box="[151,726,1891,1918]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="helleni" subGenus="Pseudocypus">
<emphasis id="B9028D2DFFAEF854FF194D364DB3F8F3" bold="true" box="[151,726,1891,1918]" pageId="4" pageNumber="5">
<emphasis id="B9028D2DFFAEF854FF194D364F94F8F0" bold="true" box="[151,241,1892,1918]" italics="true" pageId="4" pageNumber="5">Ocypus</emphasis>
(
<emphasis id="B9028D2DFFAEF854FE8F4D364EF8F8F0" bold="true" box="[257,413,1892,1918]" italics="true" pageId="4" pageNumber="5">Pseudocypus</emphasis>
)
<emphasis id="B9028D2DFFAEF854FE214D364D67F8F0" bold="true" box="[431,514,1892,1918]" italics="true" pageId="4" pageNumber="5">helleni</emphasis>
(J. Müller, 1926)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC9513FFFAEF854FF194DD44E37F82E" blockId="4.[151,726,1891,1952]" box="[151,338,1926,1952]" pageId="4" pageNumber="5">
(
<figureCitation id="134D4DBAFFAEF854FF114DD44E7AF82E" box="[159,287,1926,1952]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="4" pageNumber="5">Figs. 8–14</figureCitation>
,
<figureCitation id="134D4DBAFFAEF854FEA54DD44E2FF82E" box="[299,330,1926,1952]" captionStart="FIGURES 68 – 69" captionStartId="26.[151,269,1684,1708]" captionText="FIGURES 68 – 69. Habitus of O. helleni (68), O. fuscatus (69)." pageId="4" pageNumber="5">68</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36C02B4FFAEF855FF194D864AE1FF41" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
<paragraph id="8BC9513FFFAEF854FF194D864C4AF865" blockId="4.[151,815,2004,2027]" box="[151,815,2004,2027]" pageId="4" pageNumber="5">
<taxonomicName id="4C762ABCFFAEF854FF194D864EEFF865" authority="J. Muller, 1926" authorityName="J. Muller" authorityYear="1926" box="[151,394,2004,2027]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFAEF854FF194D864F85F865" box="[151,224,2004,2027]" italics="true" pageId="4" pageNumber="5">helleni</emphasis>
<bibRefCitation id="EFE72CCEFFAEF854FF684D864EEFF865" author="Muller" box="[230,394,2004,2027]" pageId="4" pageNumber="29" refString="Muller, J. (1926 a) Quarto contributo alla conoscenza del genere Staphylinus L. Bolletino della Societa Entomologica Italiana, 58, 27 - 32." type="journal article" year="1926" yearSuffix="a">J. Müller, 1926</bibRefCitation>
</taxonomicName>
a, 30 (
<taxonomicName id="4C762ABCFFAEF854FE454D864D23F865" box="[459,582,2004,2027]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAEF854FE454D864D23F865" box="[459,582,2004,2027]" italics="true" pageId="4" pageNumber="5">Staphylinus</emphasis>
</taxonomicName>
; subg.
<taxonomicName id="4C762ABCFFAEF854FD1A4D864C7AF865" box="[660,799,2004,2027]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAEF854FD1A4D864C7AF865" box="[660,799,2004,2027]" italics="true" pageId="4" pageNumber="5">Pseudocypus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC9513FFFAFF855FF494ACA4AE1FF41" blockId="5.[199,1435,152,207]" pageId="5" pageNumber="6">
<bibRefCitation id="EFE72CCEFFAFF855FF494ACA4EB3FF21" author="Smetana" box="[199,470,152,175]" pageId="5" pageNumber="30" refString="Smetana, A. &amp; Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88." type="journal article" year="2000">Smetana &amp; Davies, 2000</bibRefCitation>
, 40 (
<taxonomicName id="4C762ABCFFAFF855FD854ACA4D3EFF21" box="[523,603,152,175]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FD854ACA4D3EFF21" box="[523,603,152,175]" italics="true" pageId="5" pageNumber="6">Ocypus</emphasis>
</taxonomicName>
; subg.
<taxonomicName id="4C762ABCFFAFF855FD244ACA4C50FF21" box="[682,821,152,175]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FD244ACA4C50FF21" box="[682,821,152,175]" italics="true" pageId="5" pageNumber="6">Pseudocypus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFE72CCEFFAFF855FCC24ACA4C8DFF21" author="Herman" box="[844,1000,152,175]" pageId="5" pageNumber="28" refString="Herman, L. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millenium. VI. Staphylinine group (part 3). Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina). Xantholinini; Staphylinidae incertae sedis. Fossils. Protactinae. Bulletin of the American Museum of natural History, 265, 3021 - 3084." type="journal article" year="2001">Herman, 2001</bibRefCitation>
, 3382 (
<taxonomicName id="4C762ABCFFAFF855FBB74ACA4BECFF21" box="[1081,1161,152,175]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FBB74ACA4BECFF21" box="[1081,1161,152,175]" italics="true" pageId="5" pageNumber="6">Ocypus</emphasis>
</taxonomicName>
, complete synonymy and secondary references included);
<bibRefCitation id="EFE72CCEFFAFF855FDAE4AEA4DA7FF41" author="Smetana" box="[544,706,184,207]" pageId="5" pageNumber="29" refString="Smetana, A. (2004) Staphylininae. pp. 624 - 698. In: Lobl, I. &amp; Smetana, A. (Editors). Catalogue of Palaearctic Coleoptera. Vol. 2. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
, 677 (
<taxonomicName id="4C762ABCFFAFF855FC8A4AEA4C31FF41" box="[772,852,184,207]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FC8A4AEA4C31FF41" box="[772,852,184,207]" italics="true" pageId="5" pageNumber="6">Ocypus</emphasis>
</taxonomicName>
; subg.
<taxonomicName id="4C762ABCFFAFF855FC2F4AEA4B49FF41" box="[929,1068,184,207]" class="Insecta" family="Staphylinidae" genus="Pseudocypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FC2F4AEA4B49FF41" box="[929,1068,184,207]" italics="true" pageId="5" pageNumber="6">Pseudocypus</emphasis>
</taxonomicName>
, complete synonymy included).
</paragraph>
</subSubSection>
<subSubSection id="C36C02B4FFAFF856FF194AA54B9BFC39" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BC9513FFFAFF855FF194AA54BEAFDC9" blockId="5.[151,1437,247,583]" pageId="5" pageNumber="6">
<emphasis id="B9028D2DFFAFF855FF194AA54E36FE9F" bold="true" box="[151,339,247,273]" pageId="5" pageNumber="6">
<typeStatus id="54CDEF9DFFAFF855FF194AA54FB3FE9F" box="[151,214,247,273]" pageId="5" pageNumber="6">Type</typeStatus>
material.
</emphasis>
<bibRefCitation id="EFE72CCEFFAFF855FED24AAA4D21FE9C" author="Muller" box="[348,580,248,274]" pageId="5" pageNumber="29" refString="Muller, J. (1926 a) Quarto contributo alla conoscenza del genere Staphylinus L. Bolletino della Societa Entomologica Italiana, 58, 27 - 32." type="journal article" year="1926" yearSuffix="a">Müller (1926a: 30)</bibRefCitation>
described the species from two specimens, one male from “Aschabad (Transcaspia, leg. Ahnger)”, and one female from “Kopet Dagh (Transcaspia, leg. Ahnger)”. The specimens were expected to be deposited in the collection of the Zoological Museum, University of Helsinki,
<collectingCountry id="F36111AFFFAFF855FAB44B174AFCFED1" box="[1338,1433,325,351]" name="Finland" pageId="5" pageNumber="6">Finland</collectingCountry>
. The collection of the Zoological Museum of the University of Helsinki indeed contains two specimens under
<taxonomicName id="4C762ABCFFAFF855FF194BC04E75FE25" box="[151,272,402,427]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFAFF855FF194BC04E75FE25" box="[151,272,402,427]" italics="true" pageId="5" pageNumber="6">O. helleni</emphasis>
</taxonomicName>
, but only the first one (a female) belongs to the original series. It is labeled as follows: “Kopet Dagh” / ”Ahnger” / ”248&quot; / ”
<typeStatus id="54CDEF9DFFAFF855FE604BEB4D5EFE5D" box="[494,571,441,467]" pageId="5" pageNumber="6">Typus</typeStatus>
” [red label] / ”
<taxonomicName id="4C762ABCFFAFF855FD624BEB4C26FE5C" box="[748,835,441,466]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="27" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFAFF855FD624BEB4C26FE5C" box="[748,835,441,466]" italics="true" pageId="5" pageNumber="6">Hélleni</emphasis>
</taxonomicName>
m. det. J. Müller” / ”Mus. Zool. H:fors Spec. Typ. No 1381
<taxonomicName id="4C762ABCFFAFF855FE864BB24D3BFE74" authority="Helleni J. Mull." authorityName="Helleni J. Mull." box="[264,606,480,506]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9028D2DFFAFF855FE864BB24E93FE77" box="[264,502,480,505]" italics="true" pageId="5" pageNumber="6">Staphylinus Helleni</emphasis>
J. Müll.
</taxonomicName>
” The second specimen (a male) is labeled as follows: “Mar.Casp.” / ”Stenroos” / ”414&quot;. Both specimens were dissected; the female genital segment, and the male genital segment and the aedoeagus were glued to a small plate and attached to the respective specimen.
</paragraph>
<caption id="DF0901B7FFAFF855FF194D214E04F87D" httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="5" pageNumber="6" targetBox="[159,1416,642,1861]" targetPageId="5">
<paragraph id="8BC9513FFFAFF855FF194D214E04F87D" blockId="5.[151,1437,1907,2035]" pageId="5" pageNumber="6">
<emphasis id="B9028D2DFFAFF855FF194D214E2AF805" bold="true" box="[151,335,1907,1931]" pageId="5" pageNumber="6">FIGURES 8–14</emphasis>
. Details of
<emphasis id="B9028D2DFFAFF855FE5A4D264DE7F805" box="[468,642,1908,1931]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C762ABCFFAFF855FE5A4D264D1BF805" box="[468,638,1908,1931]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="helleni">Ocypus helleni</taxonomicName>
.
</emphasis>
8—tergite 10 of male genital segment (1.3 mm); 9—sternite 9 of male genital segment (2.0 mm); 10—aedoeagus, ventral view (2.0 mm); 11—aedoeagus, lateral view (2.0 mm); 12— underside of paramere (1.45 mm); 13—apical portion of underside of paramere; 14—tergite 10 of female genital segment (1.5 mm).
</paragraph>
</caption>
<paragraph id="8BC9513FFFACF856FF484ACA4C45FEA8" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
Ahnger’s label “248&quot; refers to the following information in his notebook: “Kopet Dagh, Mergen Ulja,
<date id="FFC877FFFFACF856FF194AEC4E25FF56" box="[151,320,190,216]" pageId="6" pageNumber="7" value="1901-06-24">June 24, 1901</date>
&quot;. Stenroos’ label “414&quot; refers to the information in his notebook: “Petrovsk vid västra stranden av Kaspiska havet”. Makhachkala in Daghestan,
<collectingCountry id="F36111AFFFACF856FC844AB74C07FF71" box="[778,866,229,255]" name="Russia" pageId="6" pageNumber="7">Russia</collectingCountry>
, is the present name of the place. The above information was made available to me by Jyrki Muona.
</paragraph>
<paragraph id="8BC9513FFFACF856FF484B604BF2FD0C" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
In order to preserve stability of nomenclature of the group, it is necessary to designate a
<typeStatus id="54CDEF9DFFACF856FB6C4B604A2BFEC2" box="[1250,1358,306,332]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
for
<taxonomicName id="4C762ABCFFACF856FAF14B604F8DFEFC" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FAF14B604AF1FEC5" box="[1407,1428,306,331]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FF194B0B4F8DFEFC" box="[151,232,345,370]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
</taxonomicName>
. Since the whereabouts of the second (male) specimen of the original series are unknown, and since the shape of tergite 10 of the female genital segment of
<taxonomicName id="4C762ABCFFACF856FCCE4BD24CDCFE17" box="[832,953,384,410]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FCCE4BD24C30FE17" box="[832,853,384,409]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FCE84BD24CDCFE17" box="[870,953,384,409]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
</taxonomicName>
is characteristic and markedly different from that of
<taxonomicName id="4C762ABCFFACF856FEA24BF54EADFE4E" box="[300,456,422,448]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FEA24BF54E24FE4E" box="[300,321,423,448]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FEDE4BF54EADFE4E" box="[336,456,423,448]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
, I designate the female specimen of the original series, bearing labels listed above, as the
<typeStatus id="54CDEF9DFFACF856FF6E4B9F4E2EFE69" box="[224,331,461,487]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C762ABCFFACF856FEFF4B9F4E86FE68" box="[369,483,461,487]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FEFF4B9F4EE3FE68" box="[369,390,461,486]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FE1A4B9F4E86FE68" box="[404,483,461,486]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
</taxonomicName>
. The label “
<typeStatus id="54CDEF9DFFACF856FDFE4B9F4D8DFE69" box="[624,744,461,487]" pageId="6" pageNumber="7" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C762ABCFFACF856FD604B9F4BE3FE69" authority="J. Muller, 1926" authorityName="J. Muller" authorityYear="1926" box="[750,1158,461,487]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FD604B9F4CA8FE68" box="[750,973,461,486]" italics="true" pageId="6" pageNumber="7">Staphylinus helleni</emphasis>
J. Müller, 1926
</taxonomicName>
, A. Smetana des. 2009&quot; was attached to the specimen. The specimen is in reasonably good condition, only segments one and two of the left antenna and segments 1–4 of the right antenna are present, and the left maxillary palpus and the claws of the right hind leg are missing.
<taxonomicName id="4C762ABCFFACF856FDA848134C67FDD4" box="[550,770,577,602]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FDA848134C67FDD4" box="[550,770,577,602]" italics="true" pageId="6" pageNumber="7">Ocypus picipennis</emphasis>
</taxonomicName>
is not known to occur in Kopet Dagh at present, but its occurrence there, as the only other representative of the
<taxonomicName id="4C762ABCFFACF856FCA6483A4CC4FD0F" box="[808,929,616,641]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FCA6483A4CC4FD0F" box="[808,929,616,641]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
-lineage, is probable.
</paragraph>
<paragraph id="8BC9513FFFACF856FF4848DC4D30FD78" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
The male specimen in the collection of the Zoological Museum, University of Helsinki, that is not part of the original series (see above), belongs to
<taxonomicName id="4C762ABCFFACF856FD4A48E74BF4FD41" authority="J. Muller, 1926" authorityName="J. Muller" authorityYear="1926" box="[708,1169,693,719]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caucasicus">
<emphasis id="B9028D2DFFACF856FD4A48E74CD9FD40" box="[708,956,693,718]" italics="true" pageId="6" pageNumber="7">Ocypus caucasicus</emphasis>
<bibRefCitation id="EFE72CCEFFACF856FC4B48E74BF4FD41" author="Muller" box="[965,1169,693,719]" pageId="6" pageNumber="29" refString="Muller, J. (1926 b) Quinto contributo alla conoscenza del genere Staphylinus L. Bolletino della Societa Entomologica Italiana, 58, 41 - 48." type="journal article" year="1926" yearSuffix="b">J. Müller, 1926</bibRefCitation>
</taxonomicName>
b. My corresponding determination label was attached to it.
</paragraph>
<paragraph id="8BC9513FFFACF856FF4849504B9BFC39" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
<emphasis id="B9028D2DFFACF856FF4849504DC0FC92" bold="true" box="[198,677,770,796]" pageId="6" pageNumber="7">
Additional material studied.
<collectingCountry id="F36111AFFFACF856FDB449504DC5FC92" box="[570,672,770,796]" name="China" pageId="6" pageNumber="7">CHINA</collectingCountry>
:
</emphasis>
During my work on this paper I had an opportunity to study several specimens (including males) of
<taxonomicName id="4C762ABCFFACF856FDE7497B4DB8FCCC" box="[617,733,809,835]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FDE7497B4D1BFCCC" box="[617,638,809,834]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FD03497B4DB8FCCC" box="[653,733,809,834]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
</taxonomicName>
from Sichuan. Unfortunately, due to a computer glitch and/ or my computer fear, I managed to loose these records after the specimens were returned to the loaning institution, and I was not able to recover the data. At any rate,
<taxonomicName id="4C762ABCFFACF856FC0049254B63FC1E" box="[910,1030,886,912]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FC0049254CC6FC1E" box="[910,931,887,912]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FC3A49254B63FC1E" box="[948,1030,887,912]" italics="true" pageId="6" pageNumber="7">helleni</emphasis>
</taxonomicName>
does occur on the territory of the
<collectingCountry id="F36111AFFFACF856FF1949CF4EBDFC39" box="[151,472,925,951]" name="China" pageId="6" pageNumber="7">People’s Republic of China</collectingCountry>
and I am sure that additional specimens will become available soon.
</paragraph>
</subSubSection>
<subSubSection id="C36C02B4FFACF856FF4849914D1EF932" pageId="6" pageNumber="7" type="description">
<paragraph id="8BC9513FFFACF856FF4849914C1EFB8A" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
<emphasis id="B9028D2DFFACF856FF4849914E3AFC53" bold="true" box="[198,351,963,989]" pageId="6" pageNumber="7">Description.</emphasis>
In all character states quite similar to
<taxonomicName id="4C762ABCFFACF856FCB549964CBEFC53" box="[827,987,964,990]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FCB549964C35FC53" box="[827,848,964,989]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FCED49964CBEFC53" box="[867,987,964,989]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
and different only by the differently shaped aedoeagus and tergite 10 of the female genital segment.
</paragraph>
<paragraph id="8BC9513FFFACF856FF484E434A6DFB63" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
Male. Sternite 8 with shallow, obtusely triangular medioapical emargination, similar to that of
<taxonomicName id="4C762ABCFFACF856FAF14E434E72FBDF" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FAF14E434AF1FBA4" box="[1407,1428,1041,1066]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FF194E6A4E72FBDF" box="[151,279,1080,1105]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
. Genital segment with tergite 10 similar to that of
<taxonomicName id="4C762ABCFFACF856FC194E6A4B5BFBDF" box="[919,1086,1080,1106]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FC194E6A4CC9FBDF" box="[919,940,1080,1105]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FC304E6A4B5BFBDF" box="[958,1086,1080,1105]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
, but on average shorter and narrower (
<figureCitation id="134D4DBAFFACF856FE9F4E0C4E3DFBF6" box="[273,344,1118,1144]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
); sternite 9 similar to that of
<taxonomicName id="4C762ABCFFACF856FD3F4E0D4C28FBF6" box="[689,845,1118,1144]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FD3F4E0D4DA3FBF6" box="[689,710,1119,1144]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FD5B4E0D4C28FBF6" box="[725,845,1119,1144]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
, (
<figureCitation id="134D4DBAFFACF856FCE84E0C4CC8FBF6" box="[870,941,1118,1144]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
, sternite artificially flattened). Aedoeagus (
<figureCitation id="134D4DBAFFACF856FF114ED74E49FB11" box="[159,300,1157,1183]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">Figs. 10–13</figureCitation>
) with median lobe anteriorly extended into sharp apex on side adjacent to paramere. Paramere in ventral view narrow, gradually narrowed anteriad into sharp apex not quite, or about, reaching apex of median lobe; underside of paramere lacking sensory peg setae, apical setae minute, situated as in
<figureCitation id="134D4DBAFFACF856FB204E814A60FB63" box="[1198,1285,1235,1261]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">Fig. 13</figureCitation>
.
</paragraph>
<paragraph id="8BC9513FFFACF856FF194EAB4A3FFA9D" blockId="6.[151,1437,152,1724]" box="[151,1370,1273,1299]" pageId="6" pageNumber="7">
Female. Tergite 10 of genital segment relatively small, apical portion short, with obtuse apex (
<figureCitation id="134D4DBAFFACF856FB7B4EAB4A29FA9D" box="[1269,1356,1273,1299]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BC9513FFFACF856FF484F724CF3FAB4" blockId="6.[151,1437,152,1724]" box="[198,918,1312,1338]" pageId="6" pageNumber="7">Body length 11.0–17.0 mm (see Recognition and comments).</paragraph>
<paragraph id="8BC9513FFFACF856FF484F144B60FA20" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
<emphasis id="B9028D2DFFACF856FF484F144D72FAEE" bold="true" box="[198,535,1350,1376]" pageId="6" pageNumber="7">Geographical distribution.</emphasis>
<taxonomicName id="4C762ABCFFACF856FDAD4F154DB2FAEE" box="[547,727,1351,1376]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FDAD4F154DB2FAEE" box="[547,727,1351,1376]" italics="true" pageId="6" pageNumber="7">Ocypus helleni</emphasis>
</taxonomicName>
is widely distributed eastward from
<collectingCountry id="F36111AFFFACF856FB2E4F144A62FAEE" box="[1184,1287,1350,1376]" name="Armenia" pageId="6" pageNumber="7">Armenia</collectingCountry>
and eastern
<collectingCountry id="F36111AFFFACF856FF194F3F4F88FA09" box="[151,237,1389,1415]" name="Turkey" pageId="6" pageNumber="7">Turkey</collectingCountry>
, through
<collectingCountry id="F36111AFFFACF856FED04F3F4EFAFA09" box="[350,415,1389,1415]" name="Syria" pageId="6" pageNumber="7">Syria</collectingCountry>
,
<collectingCountry id="F36111AFFFACF856FE224F3F4EBAFA09" box="[428,479,1389,1415]" name="Iran" pageId="6" pageNumber="7">Iran</collectingCountry>
,
<collectingCountry id="F36111AFFFACF856FE654F3F4DF7FA09" box="[491,658,1389,1415]" name="Turkmenistan" pageId="6" pageNumber="7">Turkmenistan</collectingCountry>
,
<collectingCountry id="F36111AFFFACF856FD114F3F4C4DFA09" box="[671,808,1389,1415]" name="Uzbekistan" pageId="6" pageNumber="7">Uzbekistan</collectingCountry>
,
<collectingCountry id="F36111AFFFACF856FCBA4F3F4CADFA09" box="[820,968,1389,1415]" name="Afghanistan" pageId="6" pageNumber="7">Afghanistan</collectingCountry>
, northern
<collectingCountry id="F36111AFFFACF856FBCF4F3F4BCCFA09" box="[1089,1193,1389,1415]" name="Pakistan" pageId="6" pageNumber="7">Pakistan</collectingCountry>
, Kashmir, Himashal Pradesh, Uttarakhand and
<collectingCountry id="F36111AFFFACF856FE424FC64D77FA20" box="[460,530,1428,1454]" name="Nepal" pageId="6" pageNumber="7">Nepal</collectingCountry>
all the way to Sichuan in mainland
<collectingCountry id="F36111AFFFACF856FC394FC64B64FA20" box="[951,1025,1428,1454]" name="China" pageId="6" pageNumber="7">China</collectingCountry>
.
</paragraph>
<paragraph id="8BC9513FFFACF856FF484FE84A07FA75" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
<emphasis id="B9028D2DFFACF856FF484FE84E35FA5A" bold="true" box="[198,336,1466,1492]" pageId="6" pageNumber="7">Bionomics.</emphasis>
<taxonomicName id="4C762ABCFFACF856FED54FE94D6AFA5A" box="[347,527,1467,1492]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FED54FE94D6AFA5A" box="[347,527,1467,1492]" italics="true" pageId="6" pageNumber="7">Ocypus helleni</emphasis>
</taxonomicName>
seems to frequent similar habitats as
<taxonomicName id="4C762ABCFFACF856FC6D4FE94BE1FA5A" box="[995,1156,1467,1493]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FC6D4FE94C9DFA5A" box="[995,1016,1467,1492]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FB854FE94BE1FA5A" box="[1035,1156,1467,1492]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
, with the more or less pronounced preference of drier habitats. No habitat data are available for the Chinese specimens studied.
</paragraph>
<paragraph id="8BC9513FFFACF856FF484C554D1EF932" blockId="6.[151,1437,152,1724]" pageId="6" pageNumber="7">
<emphasis id="B9028D2DFFACF856FF484C554D44F9AF" bold="true" box="[198,545,1543,1569]" pageId="6" pageNumber="7">Recognition and comments.</emphasis>
<taxonomicName id="4C762ABCFFACF856FDA74C5A4DBDF9AF" box="[553,728,1544,1569]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FDA74C5A4DBDF9AF" box="[553,728,1544,1569]" italics="true" pageId="6" pageNumber="7">Ocypus helleni</emphasis>
</taxonomicName>
displays similar variability described for
<taxonomicName id="4C762ABCFFACF856FB4D4C5A4A06F9AF" box="[1219,1379,1544,1570]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FB4D4C5A4BBDF9AF" box="[1219,1240,1544,1569]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FB694C5A4A06F9AF" box="[1255,1379,1544,1569]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
. The smallest specimen known to me, from around Sonamarg,
<quantity id="4C8EFCDAFFACF856FCD24C7C4B6DF9C6" box="[860,1032,1582,1608]" metricMagnitude="3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="6" pageNumber="7" unit="m" value="2750.0" valueMax="2900.0" valueMin="2600.0">2600–2900 m</quantity>
, Kashmir, is only 11.0 mm long.
<taxonomicName id="4C762ABCFFACF856FF194C074E2BF9E0" box="[151,334,1621,1646]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="helleni">
<emphasis id="B9028D2DFFACF856FF194C074E2BF9E0" box="[151,334,1621,1646]" italics="true" pageId="6" pageNumber="7">Ocypus helleni</emphasis>
</taxonomicName>
may be confused, within the mainland
<collectingCountry id="F36111AFFFACF856FCB64C074CE4F9E1" box="[824,897,1621,1647]" name="China" pageId="6" pageNumber="7">China</collectingCountry>
fauna, only with
<taxonomicName id="4C762ABCFFACF856FBD24C074A65F9E0" box="[1116,1280,1621,1647]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="picipennis">
<emphasis id="B9028D2DFFACF856FBD24C074B14F9E0" box="[1116,1137,1621,1646]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="B9028D2DFFACF856FB0C4C074A65F9E0" box="[1154,1280,1621,1646]" italics="true" pageId="6" pageNumber="7">picipennis</emphasis>
</taxonomicName>
, but the two species may be easily distinguished by the shapes of the aedoeagi (see
<figureCitation id="134D4DBAFFACF856FC584C2E4B42F918" box="[982,1063,1660,1686]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,271,1672,1696]" captionTargetBox="[151,1407,421,1645]" captionTargetId="figure@3.[151,1436,421,1649]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Details of Ocypus picipennis. 1 — tergite 10 of male genital segment (1.7 mm); 2 — sternite 9 of male genital segment (2.45 mm); 3 — aedoeagus, ventral view (2.6 mm); 4 — aedoeagus, lateral view (2.6 mm); 5 — underside of paramere (2.0 mm); 6 — apical portion of underside of paramere; 7 — tergite 10 of female genital segment (1.75 mm)." httpUri="https://zenodo.org/record/191274/files/figure.png" pageId="6" pageNumber="7">Figs. 3</figureCitation>
,
<figureCitation id="134D4DBAFFACF856FBBB4C2E4B30F918" box="[1077,1109,1660,1686]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">10</figureCitation>
), and those of tergites 10 of the female genital segments (
<figureCitation id="134D4DBAFFACF856FE634CF04D5AF932" box="[493,575,1698,1724]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,271,1672,1696]" captionTargetBox="[151,1407,421,1645]" captionTargetId="figure@3.[151,1436,421,1649]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Details of Ocypus picipennis. 1 — tergite 10 of male genital segment (1.7 mm); 2 — sternite 9 of male genital segment (2.45 mm); 3 — aedoeagus, ventral view (2.6 mm); 4 — aedoeagus, lateral view (2.6 mm); 5 — underside of paramere (2.0 mm); 6 — apical portion of underside of paramere; 7 — tergite 10 of female genital segment (1.75 mm)." httpUri="https://zenodo.org/record/191274/files/figure.png" pageId="6" pageNumber="7">Figs. 7</figureCitation>
,
<figureCitation id="134D4DBAFFACF856FDC24CF04D09F932" box="[588,620,1698,1724]" captionStart="FIGURES 8 – 14" captionStartId="5.[151,273,1907,1931]" captionTargetBox="[159,1416,642,1861]" captionTargetId="figure@5.[151,1436,633,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 14. Details of Ocypus helleni. 8 — tergite 10 of male genital segment (1.3 mm); 9 — sternite 9 of male genital segment (2.0 mm); 10 — aedoeagus, ventral view (2.0 mm); 11 — aedoeagus, lateral view (2.0 mm); 12 — underside of paramere (1.45 mm); 13 — apical portion of underside of paramere; 14 — tergite 10 of female genital segment (1.5 mm)." httpUri="https://zenodo.org/record/191275/files/figure.png" pageId="6" pageNumber="7">14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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