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<document ID-DOI="10.1111/j.1096-3642.2004.00121.x" ID-GBIF-Dataset="b6188841-cdc6-42fa-bb29-ac39d113dd72" ID-ISSN="0024-4082" ID-Zenodo-Dep="4687299" checkinTime="1618408847792" checkinUser="carolina" docAuthor="Dzik, Jerzy, Ivantsov, Andrey Yu. &amp; Deulin, Yuriy V." docDate="2004" docId="03DFC827FFE4FFC1FF20F083FEDAFE2B" docLanguage="en" docName="ZoolJLinnSoc.142.83-90.pdf" docOrigin="Zoological Journal of the Linnean Society 142 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00121.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.1:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="1" docTitle="Pechoracaris ACULICAUDA 2004, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="85" masterDocId="FFE6B05FFFE6FFC2FFC2F13BFFE3FF84" masterDocTitle="Oldest shrimp and associated phyllocarid from the Lower Devonian of northern Russia" masterLastPageNumber="90" masterPageNumber="83" pageNumber="85" updateTime="1638552323672" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Oldest shrimp and associated phyllocarid from the Lower Devonian of northern Russia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dzik, Jerzy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ivantsov, Andrey Yu.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deulin, Yuriy V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2004</mods:date>
<mods:detail type="pubDate">
<mods:number>2004-09-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>142</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>83</mods:start>
<mods:end>90</mods:end>
</mods:extent>
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<mods:location>
<mods:url>https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2004.00121.x</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1111/j.1096-3642.2004.00121.x</mods:identifier>
<mods:identifier type="GBIF-Dataset">b6188841-cdc6-42fa-bb29-ac39d113dd72</mods:identifier>
<mods:identifier type="ISSN">0024-4082</mods:identifier>
<mods:identifier type="Zenodo-Dep">4687299</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5700759" ID-GBIF-Taxon="190875568" ID-Zenodo-Dep="5700759" LSID="urn:lsid:plazi:treatment:03DFC827FFE4FFC1FF20F083FEDAFE2B" httpUri="http://treatment.plazi.org/id/03DFC827FFE4FFC1FF20F083FEDAFE2B" lastPageId="3" lastPageNumber="85" pageId="2" pageNumber="85">
<materialsCitation ID-GBIF-Occurrence="3413075301" pageId="2" pageNumber="85" typeStatus="holotype">
<subSubSection box="[226,676,440,464]" pageId="2" pageNumber="85" type="nomenclature">
<paragraph blockId="2.[226,676,440,496]" box="[226,676,440,464]" pageId="2" pageNumber="85">
<heading box="[226,676,440,464]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="85" reason="4">
<taxonomicName authority="ACULICAUDA" authorityName="ACULICAUDA" authorityYear="2004" box="[226,568,440,464]" class="Malacostraca" genus="Pechoracaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeostraca" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus" status="sp. nov.">PECHORACARIS ACULICAUDA</taxonomicName>
<emphasis bold="true" box="[576,676,440,464]" pageId="2" pageNumber="85">
<taxonomicNameLabel box="[576,676,440,464]" pageId="2" pageNumber="85" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[377,525,472,496]" pageId="2" pageNumber="85" type="description">
<paragraph blockId="2.[226,676,440,496]" box="[377,525,472,496]" pageId="2" pageNumber="85">
(
<figureCitation box="[385,462,472,496]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="2" pageNumber="85">FIGS 1</figureCitation>
,
<figureCitation box="[477,517,472,496]" captionStart="Figure 5" captionStartId="6.[143,221,1883,1902]" captionTargetBox="[154,1410,920,1827]" captionTargetId="figure-89@6.[162,1411,914,1848]" captionTargetPageId="6" captionText="Figure 5. Restoration of (A) Pechoracaris aculicauda sp. nov. and (B) Archangeliphausia spinosa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4687309" httpUri="https://zenodo.org/record/4687309/files/figure.png" pageId="2" pageNumber="85">5 A</figureCitation>
)
</paragraph>
</subSubSection>
<subSection lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="multiple">
<subSubSection pageId="2" pageNumber="85" type="materials_examined">
<paragraph blockId="2.[143,524,521,543]" box="[143,524,521,543]" pageId="2" pageNumber="85">
<heading box="[143,524,521,543]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="85" reason="8">
<emphasis box="[143,249,521,542]" italics="true" pageId="2" pageNumber="85">
<typeStatus box="[143,243,521,542]" pageId="2" pageNumber="85">Holotype</typeStatus>
:
</emphasis>
PIN 4983/21b (
<figureCitation box="[432,515,521,543]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="2" pageNumber="85">Fig. 1D</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="2.[143,759,567,681]" pageId="2" pageNumber="85">
<emphasis box="[143,441,567,588]" italics="true" pageId="2" pageNumber="85">
<typeStatus box="[143,196,567,588]" pageId="2" pageNumber="85">Type</typeStatus>
horizon and locality:
</emphasis>
Dark-grey claystone from between 4255.0 and
<quantity box="[380,487,597,619]" metricMagnitude="3" metricUnit="m" metricValue="4.2627" pageId="2" pageNumber="85" unit="m" value="4262.7">4262.7 m</quantity>
, Early Devonian (Lochkovian?). Borehole Medynskoye
<quantity box="[545,601,628,650]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="85" unit="in" value="1.0">1 in</quantity>
the Timan-Pechora region of polar
<collectingCountry box="[418,499,659,681]" name="Russia" pageId="2" pageNumber="85">Russia</collectingCountry>
.
</paragraph>
<paragraph blockId="2.[143,758,705,757]" pageId="2" pageNumber="85">
<emphasis box="[143,248,705,726]" italics="true" pageId="2" pageNumber="85">Material:</emphasis>
Thirty relatively complete specimens and many more fragments.
</paragraph>
</subSubSection>
<subSubSection box="[143,468,781,803]" pageId="2" pageNumber="85" type="diagnosis">
<paragraph blockId="2.[143,468,781,803]" box="[143,468,781,803]" pageId="2" pageNumber="85">
<emphasis box="[143,262,782,803]" italics="true" pageId="2" pageNumber="85">Diagnosis:</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="85" type="etymology">
<paragraph blockId="2.[807,1423,195,247]" pageId="2" pageNumber="85">
<emphasis box="[807,933,195,216]" italics="true" pageId="2" pageNumber="85">Etymology:</emphasis>
Derived from Latin
<emphasis box="[1204,1255,195,216]" italics="true" pageId="2" pageNumber="85">acus</emphasis>
(needle) and
<emphasis box="[807,876,225,246]" italics="true" pageId="2" pageNumber="85">cauda</emphasis>
(tail), referring to the form of the telson.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="85" type="materials_examined">
<paragraph blockId="2.[807,1423,272,325]" pageId="2" pageNumber="85">
<emphasis box="[807,912,272,293]" italics="true" pageId="2" pageNumber="85">Material:</emphasis>
Sixty relatively well preserved specimens, mostly with abdomen.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="description">
<paragraph blockId="2.[807,1423,352,803]" lastBlockId="3.[156,772,195,431]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
<emphasis box="[807,943,353,374]" italics="true" pageId="2" pageNumber="85">Description:</emphasis>
The carapace lacks any hinge or separate dorsal plates, as visible in slightly obliquely compressed specimens. The lower margin frequently shows a narrow strengthening belt, probably a little thicker than the rest of the cuticle, perhaps representing a doublure. In specimens PIN 4983/20 and 4983/7, minute sparsely distributed denticles are recognizable near the posterior end of the carapace at its margin. Two somewhat more prominent spines arm the ventro-posterior lobe of the carapace in PIN 4983/1d. The anterior end of the carapace narrows parabolically and is a little pointed, as shown by PIN 4983/8a. The carapace length in the studied material ranges from
<quantity box="[1063,1149,751,773]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="85" unit="mm" value="2.5">2.5 mm</quantity>
to
<quantity box="[1190,1275,751,772]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="2" pageNumber="85" unit="mm" value="4.7">4.7 mm</quantity>
(
<figureCitation box="[1293,1374,751,773]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="2" pageNumber="85">Fig. 1B</figureCitation>
). In most specimens dark crushed mandibles are visible across the compressed carapace near its anterior end (
<figureCitation box="[164,244,225,247]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="3" pageNumber="86">Fig. 1F</figureCitation>
). Appendages of the abdomen are transformed into paired sharp spines (
<figureCitation box="[589,673,256,278]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="3" pageNumber="86">Fig. 1D</figureCitation>
), somewhat longer than their segments. The caudal spine is definitely much longer than the carapace (
<figureCitation box="[681,763,317,339]" captionStart="Figure 1" captionStartId="2.[143,221,1736,1755]" captionTargetBox="[147,1417,857,1703]" captionTargetId="figure-373@2.[147,1418,857,1704]" captionTargetPageId="2" captionText="Figure 1. Pechoracaris aculicauda sp. nov. from the Medynskoye 1 borehole, Timan-Pechora region, polar Russia, depth 4255.04262.7 m, Lower Devonian (Lochkovian?); flattened specimens in laminated claystone. A, carapace with abdomen and partially preserved spine (PIN 4983/21a). B, slightly obliquely compressed specimen (PIN 4983/31a). C, specimen with well preserved dorsal margin of the carapace (PIN 4983/10). D, holotype showing spinose appearance of abdominal appendages (PIN 4983/21b). E, partially preserved specimen with complete abdominal spine (PIN 4983/7). F, specimen with clearly visible crushed mandibles (PIN 4983/31b)." figureDoi="http://doi.org/10.5281/zenodo.4687301" httpUri="https://zenodo.org/record/4687301/files/figure.png" pageId="3" pageNumber="86">Fig. 1E</figureCitation>
) but its exact length is difficult to trace in the fossils, the spine being either exfoliated or hidden in the sediment.
</paragraph>
</subSubSection>
</subSection>
</materialsCitation>
</treatment>
</document>