treatments-xml/data/03/DF/87/03DF87DAD151FFB111A5F91F49BAF8AD.xml

<document id="09ACFAED48AAC416611CF427121DFA6F" ID-CLB-Dataset="45982" ID-DOI="10.5281/zenodo.279150" ID-GBIF-Dataset="f3ff3539-17c9-42f6-9d00-6555b1bfbe54" ID-ISSN="1175-5326" ID-Zenodo-Dep="279150" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460409746054" checkinUser="plazi" docAuthor="Miko, Jan Mourek Ladislav &amp; Bernini, Fabio" docDate="2011" docId="03DF87DAD151FFB111A5F91F49BAF8AD" docLanguage="en" docName="zt03099p042.pdf" docOrigin="Zootaxa 3099" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metabelba (Pateribelba) romandiolae Sellnick 1943" docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FFE6FFA2D140FFAB1132FFBB4971FF9C" masterDocTitle="Taxonomy of European Damaeidae (Acari: Oribatida) IV. Partial revision of Metabelba Grandjean, 1936 with proposal of one new subgenus, one new species and redescriptions of two known species" masterLastPageNumber="42" masterPageNumber="1" pageNumber="18" updateTime="1698300307928" updateUser="ExternalLinkService">
<mods:mods id="7F9DADD5C77B2E4A71CDB8D2373067EA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4DE2DB18624ED29D6CF71531FA65566C">
<mods:title id="49EE188DEB84417C14372F98258EF84D">Taxonomy of European Damaeidae (Acari: Oribatida) IV. Partial revision of Metabelba Grandjean, 1936 with proposal of one new subgenus, one new species and redescriptions of two known species</mods:title>
</mods:titleInfo>
<mods:name id="7941B7D113684B111622318E129EDD63" type="personal">
<mods:role id="C67E1147E6A31ED949EF2F001EA3246C">
<mods:roleTerm id="C070325CF76B66E95D1638DAA38807B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E66407F7D59ED8339EA6F8E59285BCB">Miko, Jan Mourek Ladislav</mods:namePart>
</mods:name>
<mods:name id="9676FA3F8B11CDEB935B4634C588ECFD" type="personal">
<mods:role id="8B7DA084742BAB868F62CB30D335875F">
<mods:roleTerm id="BA0939D3A0FAECBD7ED3B792BC11DA5D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="746D920C59258137EE853AAD362657EA">Bernini, Fabio</mods:namePart>
</mods:name>
<mods:typeOfResource id="6FB0911E81404D532317F3883739A531">text</mods:typeOfResource>
<mods:relatedItem id="BC208390E2C0EEFE3DBF4E1DB32D310A" type="host">
<mods:titleInfo id="C5B3B7F05904324DDDE886A670880ABE">
<mods:title id="717F646C294F367A1094E95B6D9721DA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FEEC2042DEEAF62AC3B6815157FFAE09">
<mods:date id="CC835CBC3FCFA011C1104D8347CE9C46">2011</mods:date>
<mods:detail id="815B31DD5188EF6D69383F1A464984DC" type="volume">
<mods:number id="659D8309F1347DE104CFB8752808DBFB">3099</mods:number>
</mods:detail>
<mods:extent id="E59CF84034CEDAA0871E00A7E92AC4E7" unit="page">
<mods:start id="547188B2A72C0D110A90235109A82DFC">1</mods:start>
<mods:end id="042B5BD0735D0B3724CC01145F17A20D">42</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B2E69618BF4D96936D60E905705989E0">journal article</mods:classification>
<mods:identifier id="1D7B3F42C8D8E45E3737EFDFB70F6EC9" type="CLB-Dataset">45982</mods:identifier>
<mods:identifier id="4185CF91D3C070368A5F5DD711D92DA9" type="DOI">10.5281/zenodo.279150</mods:identifier>
<mods:identifier id="79E81A599A215AA8E0CAF854499FEFC6" type="GBIF-Dataset">f3ff3539-17c9-42f6-9d00-6555b1bfbe54</mods:identifier>
<mods:identifier id="2A323C21BC5CBF300611A0F2CB7D2D4F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1E6EEAD6A0927FA0BF66BF80A018F38A" type="Zenodo-Dep">279150</mods:identifier>
</mods:mods>
<treatment id="03DF87DAD151FFB111A5F91F49BAF8AD" ID-DOI="http://doi.org/10.5281/zenodo.6194470" ID-GBIF-Taxon="119423034" ID-Zenodo-Dep="6194470" LSID="urn:lsid:plazi:treatment:03DF87DAD151FFB111A5F91F49BAF8AD" httpUri="http://treatment.plazi.org/id/03DF87DAD151FFB111A5F91F49BAF8AD" lastPageId="26" lastPageNumber="27" pageId="17" pageNumber="18">
<subSubSection id="C36C6547D151FFBA11A5F91F4BB3F883" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BC936CCD151FFBA11A5F91F4A53F921" blockId="17.[151,802,1699,1726]" box="[151,802,1699,1726]" pageId="17" pageNumber="18">
<heading id="D08181A0D151FFBA11A5F91F4A53F921" bold="true" box="[151,802,1699,1726]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C764D4FD151FFBA11A5F91F4A53F921" ID-CoL="8TBWG" authority="Sellnick, 1943" authorityName="Sellnick" authorityYear="1943" box="[151,802,1699,1726]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="romandiolae" subGenus="Pateribelba">
<emphasis id="B902EADED151FFBA11A5F91F4A53F921" bold="true" box="[151,802,1699,1726]" pageId="17" pageNumber="18">
<emphasis id="B902EADED151FFBA11A5F91F4B2AF922" bold="true" box="[151,603,1700,1726]" italics="true" pageId="17" pageNumber="18">Metabelba (Pateribelba) romandiolae</emphasis>
(
<bibRefCitation id="EFE74B3DD151FFBA1358F9184A6BF921" author="Sellnick" box="[618,794,1699,1725]" pageId="17" pageNumber="18" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick, 1943</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC936CCD151FFBA11A5F9514BB3F883" blockId="17.[151,1436,1770,1824]" pageId="17" pageNumber="18">
<emphasis id="B902EADED151FFBA11A5F951487CF89D" bold="true" box="[151,269,1770,1793]" pageId="17" pageNumber="18">Synonymy.</emphasis>
<taxonomicName id="4C764D4FD151FFBA102BF9504BD8F89C" authority="Sellnick, 1943" authorityName="Sellnick" authorityYear="1943" box="[281,681,1770,1792]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED151FFBA102BF9504B72F89C" box="[281,515,1770,1792]" italics="true" pageId="17" pageNumber="18">Damaeus romandiolae</emphasis>
<bibRefCitation id="EFE74B3DD151FFBA133CF9514BD8F89C" author="Sellnick" box="[526,681,1770,1792]" pageId="17" pageNumber="18" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick, 1943</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="4C764D4FD151FFBA1385F9514D49F89C" authority="Mihelcic, 1963" authorityName="Mihelcic" authorityYear="1963" box="[695,1080,1770,1792]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED151FFBA1385F9514AFDF89C" box="[695,908,1770,1792]" italics="true" pageId="17" pageNumber="18">Metabelba gladiator</emphasis>
<bibRefCitation id="EFE74B3DD151FFBA12A5F9514D49F89C" author="Mihelcic" box="[919,1080,1770,1792]" pageId="17" pageNumber="18" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič, 1963</bibRefCitation>
</taxonomicName>
. 
<taxonomicName id="4C764D4FD151FFBA1574F951487FF8BC" authority="Bernini (1982)" authorityName="Bernini" authorityYear="1982" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED151FFBA1574F9514C4CF89C" box="[1094,1341,1770,1792]" italics="true" pageId="17" pageNumber="18">Metabelba romandiolae</emphasis>
: 
<bibRefCitation id="EFE74B3DD151FFBA1462F951487FF8BC" author="Bernini" pageId="17" pageNumber="18" refString="Bernini, F. (1982) Notulae Oribatologicae XXVII. Contributo alla conoscenza degli Oribatei della Val di Farma (Toscana meridionale). Redia, 65, 377 - 405." type="journal article" year="1982">Bernini (1982)</bibRefCitation>
</taxonomicName>
; 
<bibRefCitation id="EFE74B3DD151FFBA102AF8B2489BF8BC" box="[280,490,1801,1824]" pageId="17" pageNumber="18" refString="Bernini, F., Avanzati, A. M. &amp; Bernini, S. (1987) Notulae Oribatologicae XXXVII. Gli Acari Oribatei del Massiccio del Pollino (Italia Meridionale): aspetti faunistici e biogeografici. Biogeographia. Lavori della societa italiana di biogeografia. Nuova serie, 10, 379 - 488." type="journal article">
Bernini 
<emphasis id="B902EADED151FFBA1058F8B148ECF883" box="[362,413,1801,1823]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(1987)
</bibRefCitation>
; 
<bibRefCitation id="EFE74B3DD151FFBA10C6F8B24BB3F883" box="[500,706,1801,1823]" pageId="17" pageNumber="18" refString="Bernini, F., Arcidiacono, R. &amp; Migliorini, M. (1996) Notulae Oribatologicae LXII. Contributo alla conoscenza degli Acari Oribatei dei Monti Nebrodi e dei Peloritani (Sicilia Nordorientale). Animalia, 21, 131 - 172." type="journal article">
Bernini 
<emphasis id="B902EADED151FFBA1374F8B14B08F883" box="[582,633,1801,1823]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
(1996)
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36C6547D151FFB911A5F8F04817FEA3" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8BC936CCD151FFB911A5F8F04817FEA3" blockId="17.[151,1436,1867,2036]" lastBlockId="18.[151,1437,151,2012]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B902EADED151FFBA11A5F8F04860F8F8" bold="true" box="[151,273,1867,1892]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Propodolateral apophysis 
<emphasis id="B902EADED151FFBA130DF8F64B21F8F8" box="[575,592,1869,1892]" italics="true" pageId="17" pageNumber="18">P</emphasis>
absent. Single pair of anterior centrodorsal tubercles (
<emphasis id="B902EADED151FFBA158EF8F64DAFF8F8" box="[1212,1246,1869,1892]" italics="true" pageId="17" pageNumber="18">Da</emphasis>
) present on prodorsum posteromedial from insertions of interlamellar setae; posterior centrodorsal (
<emphasis id="B902EADED151FFBA1569F8CA4D0CF814" box="[1115,1149,1905,1928]" italics="true" pageId="17" pageNumber="18">Dp</emphasis>
), postbothridial (
<emphasis id="B902EADED151FFBA1476F8CA4C12F814" box="[1348,1379,1905,1928]" italics="true" pageId="17" pageNumber="18">Ba</emphasis>
, 
<emphasis id="B902EADED151FFBA1446F8CA4CE2F814" box="[1396,1427,1905,1928]" italics="true" pageId="17" pageNumber="18">Bp</emphasis>
) and lateral (
<emphasis id="B902EADED151FFBA102EF82E4848F830" box="[284,313,1941,1964]" italics="true" pageId="17" pageNumber="18">La</emphasis>
) tubercles absent. Parastigmatic apophyses (
<emphasis id="B902EADED151FFBA121DF82E4A05F830" box="[815,884,1941,1964]" italics="true" pageId="17" pageNumber="18">Sa, Sp</emphasis>
) slender, spiniform, with convergent tips. Sensillus (
<emphasis id="B902EADED151FFBA11F5F80249ACF84C" box="[199,221,1977,2000]" italics="true" pageId="17" pageNumber="18">ss</emphasis>
) very long, thin, flagellate, with minute barbs in middle part, basal part and flagellate tip smooth. Interlamellar seta (
<emphasis id="B902EADED151FFBA11CFF8664862F868" box="[253,275,2013,2036]" italics="true" pageId="17" pageNumber="18">in</emphasis>
) longer than ½ sensillus length, thin, attenuate, covered with minute barbs. Notogastral setae of the series 
<emphasis id="B902EADED152FFB911D3FF234850FF2C" box="[225,289,152,176]" italics="true" pageId="18" pageNumber="19">c1-h3</emphasis>
always longer than the mutual distance between the insertions of the two respective setae in pair; strong, erect, directed in radial pattern; darkly pigmented with hyaline bases and faintly serrate tips. Leg setae, except on distal parts of tarsi, of unique shape: broadly flattened basally with long attenuate tip, sparsely covered with minute spines. Discidium spiniform, curved posterolaterad, well developed. Cerotegument with long filamentous excrescences.
</paragraph>
</subSubSection>
<subSubSection id="C36C6547D152FFB911F5FEF74DDBFA90" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BC936CCD152FFB911F5FEF74D2DFCAB" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F5FEF74885FEF8" bold="true" box="[199,500,331,356]" pageId="18" pageNumber="19">
<typeStatus id="54CD886ED152FFB911F5FEF74873FEF8" box="[199,258,332,356]" pageId="18" pageNumber="19">Type</typeStatus>
material examined.
</emphasis>
Collection of Pietro Zangheri, Museo Civico di Storia Naturale, Verona: slide n. 17897 (box 42) containing two specimens, labelled as “ 
<taxonomicName id="4C764D4FD152FFB9122EFECB4D2EFE1B" authority="Romandiolae Selln." authorityName="Romandiolae Selln." box="[796,1119,367,392]" class="Arachnida" family="Damaeidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADED152FFB9122EFECB4D65FE14" box="[796,1044,368,392]" italics="true" pageId="18" pageNumber="19">Oribata Romandiolae</emphasis>
Selln.
</taxonomicName>
<emphasis id="B902EADED152FFB9155AFECB4DDBFE14" bold="true" box="[1128,1194,368,392]" pageId="18" pageNumber="19">n. sp.</emphasis>
”. The localization is “Poggio Orticai terr.[
<emphasis id="B902EADED152FFB910B6FE2E48A6FE30" box="[388,471,404,428]" italics="true" pageId="18" pageNumber="19">iccio di</emphasis>
] faggete, 
<date id="FFC8100CD152FFB9137AFE2F4BDFFE30" box="[584,686,404,428]" pageId="18" pageNumber="19" value="1929-05-05">5.V.1929</date>
, racc. Zangheri, det. Sellnick”. The two specimens are prepared by Zangheri and embedded “in Faure Andrè, 
<date id="FFC8100CD152FFB91348FE034B82FE53" box="[634,755,439,463]" pageId="18" pageNumber="19" value="1946-11">Nov. 1946</date>
” as registered on a separate label on the slide. These specimens were apparently used in the original description of the species by 
<bibRefCitation id="EFE74B3DD152FFB91287FE674D13FE68" author="Sellnick" box="[949,1122,476,500]" pageId="18" pageNumber="19" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick (1943)</bibRefCitation>
and therefore are to be considered as 
<typeStatus id="54CD886ED152FFB91023FDBA4801FD84" box="[273,368,513,536]" pageId="18" pageNumber="19" type="syntype">
<emphasis id="B902EADED152FFB91023FDBA4801FD84" box="[273,368,513,536]" italics="true" pageId="18" pageNumber="19">syntypes</emphasis>
</typeStatus>
according to article 72.4.1 of 
<collectionCode id="ED67AE09D152FFB913FFFE444A7FFD8B" box="[717,782,511,535]" pageId="18" pageNumber="19">ICZN</collectionCode>
(International Commission on Zoological Nomenclature, 1999). The dorsally placed specimen, even if crushed (
<figureCitation id="134D2A49D152FFB913CAFD9F4A37FDA0" box="[760,838,548,572]" captionStart="FIGURE 11" captionStartId="21.[151,250,1827,1850]" captionTargetBox="[151,1404,289,1792]" captionTargetId="figure@21.[151,1436,259,1807]" captionTargetPageId="21" captionText="FIGURE 11. Metabelba romandiolae. — lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; Belba setiger &quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E." httpUri="https://zenodo.org/record/279161/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
A), is complete and 
<bibRefCitation id="EFE74B3DD152FFB91515FD9F4DBDFDA0" author="Bernini" box="[1063,1228,548,572]" pageId="18" pageNumber="19" refString="Bernini, F. (1982) Notulae Oribatologicae XXVII. Contributo alla conoscenza degli Oribatei della Val di Farma (Toscana meridionale). Redia, 65, 377 - 405." type="journal article" year="1982">Bernini (1982)</bibRefCitation>
has stated it as the 
<typeStatus id="54CD886ED152FFB911A5FDF2499FFDFC" box="[151,238,585,608]" pageId="18" pageNumber="19" type="neotype">
<emphasis id="B902EADED152FFB911A5FDF2499FFDFC" box="[151,238,585,608]" italics="true" pageId="18" pageNumber="19">neotype</emphasis>
</typeStatus>
. The second specimen in the slide is ventrally placed, but completely embedded in cerotegument and in a dirty medium and was utilized only for measurements. According to article 74.1 of 
<collectionCode id="ED67AE09D152FFB91504FDD74D06FD18" box="[1078,1143,620,644]" pageId="18" pageNumber="19">ICZN</collectionCode>
(op. cit.) we designate the first, dorsally placed, specimen as the 
<typeStatus id="54CD886ED152FFB91373FD2B4BD7FD34" box="[577,678,656,680]" pageId="18" pageNumber="19" type="lectotype">
<emphasis id="B902EADED152FFB91373FD2B4BD7FD34" box="[577,678,656,680]" italics="true" pageId="18" pageNumber="19">lectotype</emphasis>
</typeStatus>
, and the second, ventrally placed one as the 
<typeStatus id="54CD886ED152FFB915A5FD2B4C40FD34" box="[1175,1329,656,680]" pageId="18" pageNumber="19" type="paralectotype">
<emphasis id="B902EADED152FFB915A5FD2B4C40FD34" box="[1175,1329,656,680]" italics="true" pageId="18" pageNumber="19">paralectotype</emphasis>
</typeStatus>
. Presumably, the other two specimens of the 
<typeStatus id="54CD886ED152FFB9136FFD0E4BFCFD51" box="[605,653,693,717]" pageId="18" pageNumber="19">type</typeStatus>
series mentioned by 
<bibRefCitation id="EFE74B3DD152FFB912A6FD0F4D36FD50" author="Sellnick" box="[916,1095,692,716]" pageId="18" pageNumber="19" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick (1943)</bibRefCitation>
in original description were destroyed by the fire of the Sellnick’s house in Königsberg (now Kaliningrad, 
<collectingCountry id="F361765CD152FFB91514FD6C4D03FD73" box="[1062,1138,727,751]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
) during the World War II (
<bibRefCitation id="EFE74B3DD152FFB911ADFD47484CFC88" author="Bernini" box="[159,317,764,788]" pageId="18" pageNumber="19" refString="Bernini, F. (1982) Notulae Oribatologicae XXVII. Contributo alla conoscenza degli Oribatei della Val di Farma (Toscana meridionale). Redia, 65, 377 - 405." type="journal article" year="1982">Bernini, 1982</bibRefCitation>
; H. Dastych, 2009—pers. comm.). The Sellnick’s collection in the Zoologisches Institut und Zoologisches Museum Universität Hamburg contains only the material collected after 1950.
</paragraph>
<paragraph id="8BC936CCD152FFB911F7FCFF4BB6FB93" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
Collection of Franz Mihelčič, Tiroler Landesmuseum Ferdinandeum, Innsbruck: Slide MIH 1-42 labelled as &quot; 
<taxonomicName id="4C764D4FD152FFB911A5FCD3480EFC1C" box="[151,383,872,896]" class="Arachnida" family="Damaeidae" genus="Metabalba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gladiator" status="sp. nov.">
<emphasis id="B902EADED152FFB911A5FCD3480EFC1C" box="[151,383,872,896]" italics="true" pageId="18" pageNumber="19">Metabalba gladiator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A5D152FFB910B4FCD348B4FC1C" box="[390,453,872,896]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B902EADED152FFB910B4FCD348B4FC1C" bold="true" box="[390,453,872,896]" pageId="18" pageNumber="19">n. sp.</emphasis>
</taxonomicNameLabel>
Mih. 
<typeStatus id="54CD886ED152FFB9133BFCDC4B37FCE3" box="[521,582,871,895]" pageId="18" pageNumber="19">Type</typeStatus>
“. The locality is hardly legible; according to 
<bibRefCitation id="EFE74B3DD152FFB9150CFCDC4D8FFCE3" author="Totschnig" box="[1086,1278,871,895]" pageId="18" pageNumber="19" refString="Totschnig, U. (2001) Die Hornmilbensammlung (Acari, Oribatida) Franz Mihelcic im Tiroler Landesmuseum Ferdinandeum, Innsbruck. Veroffentlichungen des Tiroler Landesmuseum Ferdinandeum, 81, 205 - 240." type="journal article" year="2001">Totschnig (2001)</bibRefCitation>
it is “St. Veit, Jauntal, Kärnten, A, Boden, Moos, trocken 
<date id="FFC8100CD152FFB913BEFC374B8AFC38" box="[652,763,908,932]" pageId="18" pageNumber="19" value="1962-07">Juli 1962</date>
.” The specimen is crushed (see 
<figureCitation id="134D2A49D152FFB9155FFC374DC2FC38" box="[1133,1203,908,932]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
), but complete. It is embedded probably in a medium based on gum arabic and mounted between two cover slides, which are attached to a piece of pasteboard with central hole. The remaining three specimens mentioned in the original description (
<bibRefCitation id="EFE74B3DD152FFB911ACFC4C483AFB93" author="Mihelcic" box="[158,331,1015,1039]" pageId="18" pageNumber="19" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič, 1963</bibRefCitation>
) were not found in the collection.
</paragraph>
<paragraph id="8BC936CCD152FFB911F5FBA74B50FB58" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F5FBA74B6BFBA8" bold="true" box="[199,538,1051,1076]" pageId="18" pageNumber="19">Further material examined.</emphasis>
Collection of Fabio Bernini, Department of Evolutionary Biology, Siena: six specimens (two of which given to the Collection of J. Mourek, Dept. of Zoology, Faculty of Science, Charles University Prague, 
<collectingCountry id="F361765CD152FFB91077FBDF4889FBE0" box="[325,504,1124,1148]" name="Czech Republic" pageId="18" pageNumber="19">Czech Republic</collectingCountry>
) labelled as 
<taxonomicName id="4C764D4FD152FFB913B0FBDF4AFFFBE0" box="[642,910,1124,1148]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED152FFB913B0FBDF4AFFFBE0" box="[642,910,1124,1148]" italics="true" pageId="18" pageNumber="19">Metabelba romandiolae</emphasis>
</taxonomicName>
and collected by Prof. R. Dallai in the forest of 
<taxonomicName id="4C764D4FD152FFB911A5FB32486DFB3C" box="[151,284,1160,1184]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="B902EADED152FFB911A5FB32486DFB3C" box="[151,284,1160,1184]" italics="true" pageId="18" pageNumber="19">Picea abies</emphasis>
</taxonomicName>
near the village of S. Benedetto in Alpe, about 
<quantity id="4C8E9B29D152FFB91261FB3C4AECFB3C" box="[851,925,1159,1184]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="19" unit="m" value="900.0">900 m</quantity>
a.s.l., 
<quantity id="4C8E9B29D152FFB912DFFB3C4D46FB03" box="[1005,1079,1159,1183]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="10.0">10 km</quantity>
from the 
<typeStatus id="54CD886ED152FFB9159DFB324DAEFB3D" box="[1199,1247,1161,1185]" pageId="18" pageNumber="19">type</typeStatus>
locality, Poggio Orticai, Tosco-Emiliano Apennine.
</paragraph>
<paragraph id="8BC936CCD152FFB911F7FB744DDBFA90" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F7FB744B63FB74" bold="true" box="[197,530,1231,1256]" pageId="18" pageNumber="19">Collection of Carl Wilmann</emphasis>
, Zoologische Staatssammlung München: slide labelled as &quot; 
<taxonomicName id="4C764D4FD152FFB9159FFB744CE5FB7B" authority="Kulcz." authorityName="Kulcz." box="[1197,1428,1231,1255]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="setiger">Belba setiger Kulcz.</taxonomicName>
/ Abs. Det C. Willmann&quot;, 
<specimenCount id="9D70FD45D152FFB91099FB4F4B5BFA90" box="[427,554,1268,1292]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
(sex unknown), locality and date of collection not given.
</paragraph>
</subSubSection>
<subSubSection id="C36C6547D152FFB311F5FAAC4D8AF80B" lastPageId="24" lastPageNumber="25" pageId="18" pageNumber="19" type="description">
<paragraph id="8BC936CCD152FFB911F5FAAC4833F9E8" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F5FAAC48BDFAAC" bold="true" box="[199,460,1303,1328]" pageId="18" pageNumber="19">Redescription (adult).</emphasis>
<figureCitation id="134D2A49D152FFB910E1FAAC4B3CFAB3" box="[467,589,1303,1327]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStart-5="FIGURE 14" captionStart-6="FIGURE 15" captionStart-7="FIGURE 16" captionStartId-0="19.[151,250,1536,1559]" captionStartId-1="20.[151,250,1399,1422]" captionStartId-2="21.[151,250,1827,1850]" captionStartId-3="22.[151,250,1777,1800]" captionStartId-4="23.[151,250,1894,1917]" captionStartId-5="24.[151,250,1591,1614]" captionStartId-6="25.[151,250,1910,1933]" captionStartId-7="26.[151,250,1473,1496]" captionTargetBox-0="[167,1419,356,1448]" captionTargetBox-1="[181,1421,181,1364]" captionTargetBox-2="[151,1404,289,1792]" captionTargetBox-3="[152,1432,249,1776]" captionTargetBox-4="[204,1428,277,1854]" captionTargetBox-5="[180,1428,211,1553]" captionTargetBox-6="[187,1419,554,1870]" captionTargetBox-7="[151,1436,721,1451]" captionTargetId-0="figure@19.[151,1436,305,1515]" captionTargetId-1="figure@20.[151,1436,159,1394]" captionTargetId-2="figure@21.[151,1436,259,1807]" captionTargetId-3="figure@22.[151,1436,193,1788]" captionTargetId-4="figure@23.[168,1453,268,1871]" captionTargetId-5="figure@24.[151,1436,193,1570]" captionTargetId-6="figure@25.[151,1436,517,1889]" captionTargetId-7="figure@26.[151,1436,721,1452]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="23" captionTargetPageId-5="24" captionTargetPageId-6="25" captionTargetPageId-7="26" captionText-0="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." captionText-1="FIGURE 10. Metabelba romandiolae. A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — details of some notogastral setae. Cerotegument not drawn." captionText-2="FIGURE 11. Metabelba romandiolae. — lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; Belba setiger &quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E." captionText-3="FIGURE 12. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym). A—dorsal view. Legs only partly drawn, leg setae and cerotegument not drawn. B—exobothridial seta in detail." captionText-4="FIGURE 13. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym). A) ventral view in total. Legs only partly drawn, cerotegument and setae of legs and pedipalps not drawn; B) detail of epimeral seta 4 c." captionText-5="FIGURE 14. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg I and II. A) left trochanter and femur I, antiaxial view; B) left genu, tibia and tarsus I, antiaxial view; C) left femur II, paraxial and slightly ventral view; D) left genu and tibia II, paraxial view; E) left tarsus II, paraxial view." captionText-6="FIGURE 15. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg III and IV. A) right genu, tibia and tarsus I, antiaxial view; B) left trochanter and femur III, antiaxial and slightly ventral view; C) left genu, tibia and tarsus IV, antiaxial and slightly ventral view; D) left trochanter and femur IV, antiaxial and slightly ventral view." captionText-7="FIGURE 16. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym), light micrographs, Nomarski DIC. A) detail of genu III with typical leg setae; B) cerotegument on tibia IV. Scale bars: 100 Μm." httpUri-0="https://zenodo.org/record/279159/files/figure.png" httpUri-1="https://zenodo.org/record/279160/files/figure.png" httpUri-2="https://zenodo.org/record/279161/files/figure.png" httpUri-3="https://zenodo.org/record/279162/files/figure.png" httpUri-4="https://zenodo.org/record/279163/files/figure.png" httpUri-5="https://zenodo.org/record/279164/files/figure.png" httpUri-6="https://zenodo.org/record/279165/files/figure.png" httpUri-7="https://zenodo.org/record/279166/files/figure.png" pageId="18" pageNumber="19">Figs. 9–16</figureCitation>
. 
<emphasis id="B902EADED152FFB9136AFAA24B95FAAC" box="[600,740,1305,1328]" italics="true" pageId="18" pageNumber="19">Dimensions.</emphasis>
The crushed 
<typeStatus id="54CD886ED152FFB9124EFAAC4A90FAAC" box="[892,993,1303,1328]" pageId="18" pageNumber="19" type="lectotype">
<emphasis id="B902EADED152FFB9124EFAAC4A90FAAC" bold="true" box="[892,993,1303,1328]" italics="true" pageId="18" pageNumber="19">lectotype</emphasis>
</typeStatus>
from Zangheri’s collection (
<figureCitation id="134D2A49D152FFB91411FAAC4C1BFAB3" box="[1315,1386,1303,1327]" captionStart="FIGURE 11" captionStartId="21.[151,250,1827,1850]" captionTargetBox="[151,1404,289,1792]" captionTargetId="figure@21.[151,1436,259,1807]" captionTargetPageId="21" captionText="FIGURE 11. Metabelba romandiolae. — lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; Belba setiger &quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E." httpUri="https://zenodo.org/record/279161/files/figure.png" pageId="18" pageNumber="19">Fig 11</figureCitation>
AB) could not be measured accurately, thus we have measured the second specimen present in the same slide placed ventrally: total length about 640 μm and notogastral width 420 μm. Similarily the 
<typeStatus id="54CD886ED152FFB91564FADB4DF5FAEB" box="[1110,1156,1376,1399]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB91564FADB4DF5FAEB" bold="true" box="[1110,1156,1376,1399]" italics="true" pageId="18" pageNumber="19">type</emphasis>
</typeStatus>
specimen of 
<taxonomicName id="4C764D4FD152FFB91416FADB498EFA00" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED152FFB91416FADB498EFA00" italics="true" pageId="18" pageNumber="19">Metabelba gladiator</emphasis>
</taxonomicName>
from Mihelčič’s collection (
<figureCitation id="134D2A49D152FFB91377FA3F4BD2FA00" box="[581,675,1412,1436]" captionStart="FIGURE 10" captionStartId="20.[151,250,1399,1422]" captionTargetBox="[181,1421,181,1364]" captionTargetId="figure@20.[151,1436,159,1394]" captionTargetPageId="20" captionText="FIGURE 10. Metabelba romandiolae. A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — details of some notogastral setae. Cerotegument not drawn." httpUri="https://zenodo.org/record/279160/files/figure.png" pageId="18" pageNumber="19">Figs. 10</figureCitation>
B, 12–16) is crushed, ventral length approximately 585 μm. Specimen from Willmann’s collection labelled as &quot; 
<taxonomicName id="4C764D4FD152FFB913A4FA134A5AFA5C" box="[662,811,1448,1472]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="setiger">
<emphasis id="B902EADED152FFB913A4FA134A5AFA5C" box="[662,811,1448,1472]" italics="true" pageId="18" pageNumber="19">Belba setiger</emphasis>
</taxonomicName>
&quot; (
<figureCitation id="134D2A49D152FFB91274FA1C4AE5FA23" box="[838,916,1447,1471]" captionStart="FIGURE 11" captionStartId="21.[151,250,1827,1850]" captionTargetBox="[151,1404,289,1792]" captionTargetId="figure@21.[151,1436,259,1807]" captionTargetPageId="21" captionText="FIGURE 11. Metabelba romandiolae. — lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; Belba setiger &quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E." httpUri="https://zenodo.org/record/279161/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
CDE): ventral length measured in dorsal view 584 μm, maximum notogastral width 408 μm. In the original description of 
<taxonomicName id="4C764D4FD152FFB912CCFA764DFDFA78" box="[1022,1164,1484,1508]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED152FFB912CCFA764DFDFA78" box="[1022,1164,1484,1508]" italics="true" pageId="18" pageNumber="19">M. gladiator</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFE74B3DD152FFB91588FA704C00FA78" author="Mihelcic" box="[1210,1393,1483,1508]" pageId="18" pageNumber="19" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič (1963)</bibRefCitation>
the body length ranged from 680 to 720 μm (mean calculated by us 708 μm) and the maximum notogastral width 410– 430 μm (mean calculated by us 421 μm); n = 4. Ventral length of specimens of 
<taxonomicName id="4C764D4FD152FFB91526F9AE4DB9F9B0" box="[1044,1224,1556,1580]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED152FFB91526F9AE4DB9F9B0" box="[1044,1224,1556,1580]" italics="true" pageId="18" pageNumber="19">M. romandiolae</emphasis>
</taxonomicName>
from S. Benedetto in Alpe was 568–576 (572) μm, maximum notogastral width 408–440 (424) μm; n = 2. Detailed measurements of legs in Table 1.
</paragraph>
<paragraph id="8BC936CCD152FFB911F7F93A4DF6F898" blockId="18.[151,1437,151,2012]" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F7F93A483BF904" box="[197,330,1665,1688]" italics="true" pageId="18" pageNumber="19">Integument.</emphasis>
Body colour medium brown. Cerotegument with long &quot;wool-like&quot; filamentous excrescences (
<figureCitation id="134D2A49D152FFB9145DF9C449C3F920" captionStart="FIGURE 11" captionStartId="21.[151,250,1827,1850]" captionTargetBox="[151,1404,289,1792]" captionTargetId="figure@21.[151,1436,259,1807]" captionTargetPageId="21" captionText="FIGURE 11. Metabelba romandiolae. — lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; Belba setiger &quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E." httpUri="https://zenodo.org/record/279161/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
E, 16AB); excrescences shorter on legs, longer in sejugal area, on epimeres and around notogastral setae. Underlying cuticle mostly smooth, but faintly granular on apophyses. Gastronotic exuviae (scalps) not present in the studied specimens and not noted in original descriptions (
<bibRefCitation id="EFE74B3DD152FFB91229F9574AB0F898" author="Sellnick" box="[795,961,1772,1796]" pageId="18" pageNumber="19" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick, 1943</bibRefCitation>
; 
<bibRefCitation id="EFE74B3DD152FFB912FFF9504D0BF898" author="Mihelcic" box="[973,1146,1771,1796]" pageId="18" pageNumber="19" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič, 1963</bibRefCitation>
).
</paragraph>
<paragraph id="8BC936CCD152FFB811F7F8AB4B79FE80" blockId="18.[151,1437,151,2012]" lastBlockId="19.[151,1436,151,284]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B902EADED152FFB911F7F8AB4838F8B4" box="[197,329,1808,1832]" italics="true" pageId="18" pageNumber="19">Prodorsum.</emphasis>
<figureCitation id="134D2A49D152FFB91063F8B448EEF8BB" box="[337,415,1807,1831]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="18" pageNumber="19">Figs. 9</figureCitation>
A, 12A. Prodorsum of roughly triangular shape. Anterior centrodorsal tubercle (
<emphasis id="B902EADED152FFB91402F8AA4C23F8B4" box="[1328,1362,1809,1832]" italics="true" pageId="18" pageNumber="19">Da</emphasis>
) present, posterior centrodorsal tubercle (
<emphasis id="B902EADED152FFB91303F88E4B22F8D0" box="[561,595,1845,1868]" italics="true" pageId="18" pageNumber="19">Dp</emphasis>
) absent, but cuticle slightly thickened in normal position of 
<emphasis id="B902EADED152FFB91432F88E4C53F8D0" box="[1280,1314,1845,1868]" italics="true" pageId="18" pageNumber="19">Dp</emphasis>
; postbothridial (
<emphasis id="B902EADED152FFB911DBF8E2484DF8EC" box="[233,316,1881,1904]" italics="true" pageId="18" pageNumber="19">Ba, Bp</emphasis>
) and lateral (
<emphasis id="B902EADED152FFB910DAF8E24B74F8EC" box="[488,517,1881,1904]" italics="true" pageId="18" pageNumber="19">La</emphasis>
) tubercles absent. Parastigmatic apophyses (
<emphasis id="B902EADED152FFB9152DF8E34D1DF8F3" box="[1055,1132,1880,1903]" italics="true" pageId="18" pageNumber="19">Sa, Sp</emphasis>
) slender, spiniform, with convergent tips. Rostral (
<emphasis id="B902EADED152FFB9108DF8C648A7F808" box="[447,470,1917,1940]" italics="true" pageId="18" pageNumber="19">ro</emphasis>
) and lamellar (
<emphasis id="B902EADED152FFB913A2F8C74BD5F808" box="[656,676,1916,1940]" italics="true" pageId="18" pageNumber="19">le</emphasis>
) setae thin long, of approximately equal length, strongly curved inwards. Seta 
<emphasis id="B902EADED152FFB91007F81A483DF824" box="[309,332,1953,1976]" italics="true" pageId="18" pageNumber="19">ro</emphasis>
thin, almost hyaline, with few minute barbs on outer curvature. Seta 
<emphasis id="B902EADED152FFB91565F81B4D1DF824" box="[1111,1132,1952,1976]" italics="true" pageId="18" pageNumber="19">le</emphasis>
thicker and darker than 
<emphasis id="B902EADED152FFB9144FF81A4CEBF824" box="[1405,1434,1953,1976]" italics="true" pageId="18" pageNumber="19">ro,</emphasis>
with dense short barbs on outer curvature. Interlamellar seta (
<emphasis id="B902EADED152FFB91279F87E4A10F840" box="[843,865,1989,2012]" italics="true" pageId="18" pageNumber="19">in</emphasis>
) arising on minute apophysis, comparatively long, slightly longer than half of sensillus length, dark, densely covered with short barbs, flagellate. Exobothridial seta (
<emphasis id="B902EADED153FFB81192FF0649C8FF48" box="[160,185,189,212]" italics="true" pageId="19" pageNumber="20">ex</emphasis>
) comparatively long, thin, covered with short barbs on outer curvature (
<figureCitation id="134D2A49D153FFB81532FF074D25FF48" box="[1024,1108,188,212]" captionStart="FIGURE 10" captionStartId="20.[151,250,1399,1422]" captionTargetBox="[181,1421,181,1364]" captionTargetId="figure@20.[151,1436,159,1394]" captionTargetPageId="20" captionText="FIGURE 10. Metabelba romandiolae. A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — details of some notogastral setae. Cerotegument not drawn." httpUri="https://zenodo.org/record/279160/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
A, 12B). Sensillus (
<emphasis id="B902EADED153FFB81408FF064C21FF48" box="[1338,1360,189,212]" italics="true" pageId="19" pageNumber="20">ss</emphasis>
) long, thin, flagellate; covered with short barbs in middle 1/3, basal part and flagellate tip smooth. Bothridial funnel of regular shape, normal for family.
</paragraph>
<caption id="DF096644D153FFB811A5F9BB4AEFF9A9" httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="19" pageNumber="20" targetBox="[167,1419,356,1448]" targetPageId="19">
<paragraph id="8BC936CCD153FFB811A5F9BB4AEFF9A9" blockId="19.[151,1436,1536,1589]" pageId="19" pageNumber="20">
<emphasis id="B902EADED153FFB811A5F9BB4862F989" bold="true" box="[151,275,1536,1559]" pageId="19" pageNumber="20">FIGURE 9.</emphasis>
<taxonomicName id="4C764D4FD153FFB81029F9BB4B7FF98A" box="[283,526,1536,1558]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED153FFB81029F9BB4B7FF98A" box="[283,526,1536,1558]" italics="true" pageId="19" pageNumber="20">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn.
</paragraph>
</caption>
<paragraph id="8BC936CCD153FFB811F7F9E64852F8B4" blockId="19.[151,1436,1628,1976]" pageId="19" pageNumber="20">
<emphasis id="B902EADED153FFB811F7F9E64837F9E8" box="[197,326,1629,1652]" italics="true" pageId="19" pageNumber="20">Notogaster.</emphasis>
<figureCitation id="134D2A49D153FFB8107FF9E748EAF9E8" box="[333,411,1628,1652]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="19" pageNumber="20">Figs. 9</figureCitation>
A, 10AB. Notogaster of roughly circular outline in dorsal view. Notogastral setae of series 
<emphasis id="B902EADED153FFB811A5F93B49A4F904" box="[151,213,1664,1688]" italics="true" pageId="19" pageNumber="20">c1-c2</emphasis>
, 
<emphasis id="B902EADED153FFB811D7F93B486BF904" box="[229,282,1664,1688]" italics="true" pageId="19" pageNumber="20">la -lp</emphasis>
and 
<emphasis id="B902EADED153FFB81066F93B48E7F904" box="[340,406,1664,1688]" italics="true" pageId="19" pageNumber="20">h1-h3</emphasis>
longer than mutual distances within respective setal pairs, strong, erect, directed in radial pattern. Setae darkly pigmented with hyaline bases and faintly serrate tips, the &quot;teeth&quot; directed backwards (
<figureCitation id="134D2A49D153FFB8145DF91F49C4F97C" captionStart="FIGURE 10" captionStartId="20.[151,250,1399,1422]" captionTargetBox="[181,1421,181,1364]" captionTargetId="figure@20.[151,1436,159,1394]" captionTargetPageId="20" captionText="FIGURE 10. Metabelba romandiolae. A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — details of some notogastral setae. Cerotegument not drawn." httpUri="https://zenodo.org/record/279160/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
B). Seta 
<emphasis id="B902EADED153FFB81024F9734840F97C" box="[278,305,1736,1760]" italics="true" pageId="19" pageNumber="20">c1</emphasis>
shorter than 
<emphasis id="B902EADED153FFB810F5F9734B77F97C" box="[455,518,1736,1760]" italics="true" pageId="19" pageNumber="20">c2-h3</emphasis>
. Setae 
<emphasis id="B902EADED153FFB81368F9734BC0F97C" box="[602,689,1736,1760]" italics="true" pageId="19" pageNumber="20">ps1-ps3</emphasis>
comparatively long, thin, with row of short barbs on outer curvature, 
<emphasis id="B902EADED153FFB811E2F9574986F898" box="[208,247,1772,1796]" italics="true" pageId="19" pageNumber="20">ps1</emphasis>
&gt; 
<emphasis id="B902EADED153FFB81026F957484AF898" box="[276,315,1772,1796]" italics="true" pageId="19" pageNumber="20">ps2</emphasis>
&gt; 
<emphasis id="B902EADED153FFB8106BF957480EF898" box="[345,383,1772,1796]" italics="true" pageId="19" pageNumber="20">ps3</emphasis>
, seta 
<emphasis id="B902EADED153FFB810F2F9574897F898" box="[448,486,1772,1796]" italics="true" pageId="19" pageNumber="20">ps1</emphasis>
very long with attenuate fragile tip (
<figureCitation id="134D2A49D153FFB8124DF9574ABEF898" box="[895,975,1772,1796]" captionStart="FIGURE 10" captionStartId="20.[151,250,1399,1422]" captionTargetBox="[181,1421,181,1364]" captionTargetId="figure@20.[151,1436,159,1394]" captionTargetPageId="20" captionText="FIGURE 10. Metabelba romandiolae. A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — details of some notogastral setae. Cerotegument not drawn." httpUri="https://zenodo.org/record/279160/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
AB), projecting distinctly more posteriad than seta 
<emphasis id="B902EADED153FFB81032F8AB486DF8B4" box="[256,284,1808,1832]" italics="true" pageId="19" pageNumber="20">h1</emphasis>
.
</paragraph>
<paragraph id="8BC936CCD153FFB811F7F88F4B29F824" blockId="19.[151,1436,1628,1976]" pageId="19" pageNumber="20">
<emphasis id="B902EADED153FFB811F7F88F482DF8D0" box="[197,348,1844,1868]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C764D4FD153FFB811F7F88F4829F8D0" box="[197,344,1844,1868]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="19" pageNumber="20" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
<figureCitation id="134D2A49D153FFB81056F88F48D0F8D0" box="[356,417,1844,1868]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="19" pageNumber="20">Fig 9</figureCitation>
B, 13A. Subcapitulum normal for family: diarthric, with three pairs of thin and comparatively long setae (
<emphasis id="B902EADED153FFB81056F8E24809F8EC" box="[356,376,1881,1904]" italics="true" pageId="19" pageNumber="20">m</emphasis>
&gt; 
<emphasis id="B902EADED153FFB810ABF8E348D6F8EC" box="[409,423,1880,1904]" italics="true" pageId="19" pageNumber="20">h</emphasis>
&gt; 
<emphasis id="B902EADED153FFB810FAF8E248A7F8EC" box="[456,470,1881,1904]" italics="true" pageId="19" pageNumber="20">a</emphasis>
) and two pairs of setiform processes (
<emphasis id="B902EADED153FFB812BFF8E34AB8F8EC" box="[909,969,1880,1904]" italics="true" pageId="19" pageNumber="20">or1,2</emphasis>
). Setae 
<emphasis id="B902EADED153FFB8151AF8E24D2BF8EC" box="[1064,1114,1880,1904]" italics="true" pageId="19" pageNumber="20">m, h</emphasis>
, 
<emphasis id="B902EADED153FFB8155BF8E24D06F8EC" box="[1129,1143,1881,1904]" italics="true" pageId="19" pageNumber="20">a</emphasis>
covered with few minute barbs on outer curvature. Setation of chelicera not discernible, setal formula of pedipalp (from trochanter to tarsus, solenidion in parenthesis): 0-2-1-3-9(1).
</paragraph>
<caption id="DF096644D154FFBF11A5FACC484CFA56" httpUri="https://zenodo.org/record/279160/files/figure.png" pageId="20" pageNumber="21" targetBox="[181,1421,181,1364]" targetPageId="20">
<paragraph id="8BC936CCD154FFBF11A5FACC484CFA56" blockId="20.[151,1436,1399,1482]" pageId="20" pageNumber="21">
<emphasis id="B902EADED154FFBF11A5FACC486EFA10" bold="true" box="[151,287,1399,1422]" pageId="20" pageNumber="21">FIGURE 10.</emphasis>
<emphasis id="B902EADED154FFBF1017FACC4B51FA11" box="[293,544,1399,1421]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C764D4FD154FFBF1017FACC4B6DFA11" box="[293,540,1399,1421]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="romandiolae">Metabelba romandiolae</taxonomicName>
.
</emphasis>
A—specimen from S. Benedetto in Alpe, partial lateral view; B) specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED154FFBF131EFA2D4A78FA30" box="[556,777,1430,1452]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C764D4FD154FFBF131EFA2D4A70FA30" box="[556,769,1430,1452]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym)—details of some notogastral setae. Cerotegument not drawn.
</paragraph>
</caption>
<paragraph id="8BC936CCD154FFBF11F5FA4D4D59F927" blockId="20.[151,1437,1526,2036]" pageId="20" pageNumber="21">
<emphasis id="B902EADED154FFBF11F5FA4D48F7F992" box="[199,390,1526,1550]" italics="true" pageId="20" pageNumber="21">Epimeral region.</emphasis>
<figureCitation id="134D2A49D154FFBF10BFFA4D48A0F992" box="[397,465,1526,1550]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
B, 13A. Medial pit 
<emphasis id="B902EADED154FFBF139EFA4C4BB7F992" box="[684,710,1527,1550]" italics="true" pageId="20" pageNumber="21">cp</emphasis>
on coxisternum I not developed Propodoventral (
<emphasis id="B902EADED154FFBF15CBFA4D4C57F992" box="[1273,1318,1526,1550]" italics="true" pageId="20" pageNumber="21">E2a</emphasis>
, 
<emphasis id="B902EADED154FFBF1407FA4D4C13F992" box="[1333,1378,1526,1550]" italics="true" pageId="20" pageNumber="21">E2p</emphasis>
) and ventrosejugal (
<emphasis id="B902EADED154FFBF1073F9A14828F9AD" box="[321,345,1562,1585]" italics="true" pageId="20" pageNumber="21">Va</emphasis>
, 
<emphasis id="B902EADED154FFBF105DF9A148FFF9AD" box="[367,398,1562,1585]" italics="true" pageId="20" pageNumber="21">Vp</emphasis>
) enantiophyses absent. Apodeme I well developed in lateral parts and weakly defined in medial part, apodemes 2 confluent, ventrosejugal furrow complete. Epimeral setal formula (I to IV): 3-1-3-4, normal for 
<taxonomicName id="4C764D4FD154FFBF11C6F9E5481DF9EA" box="[244,364,1630,1654]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADED154FFBF11C6F9E5481DF9EA" box="[244,364,1630,1654]" italics="true" pageId="20" pageNumber="21">Metabelba</emphasis>
</taxonomicName>
, rarely 3-1-4-4 (
<figureCitation id="134D2A49D154FFBF131CF9E54B02F9EA" box="[558,627,1630,1654]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
B). Epimeral setae comparatively long, smooth or with several minute indistinct barbs. All epimeral setae at least slightly flattened and broadened with long attenuate tips, similar to leg setae; broadening especially well developed in setae 
<emphasis id="B902EADED154FFBF13D4F91F4A5AF920" box="[742,811,1700,1724]" italics="true" pageId="20" pageNumber="21">3a,b,c</emphasis>
and 
<emphasis id="B902EADED154FFBF1253F91F4AD7F920" box="[865,934,1700,1724]" italics="true" pageId="20" pageNumber="21">4b,c,d</emphasis>
(
<figureCitation id="134D2A49D154FFBF1287F9184D77F927" box="[949,1030,1699,1723]" captionStart="FIGURE 13" captionStartId="23.[151,250,1894,1917]" captionTargetBox="[204,1428,277,1854]" captionTargetId="figure@23.[168,1453,268,1871]" captionTargetPageId="23" captionText="FIGURE 13. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym). A) ventral view in total. Legs only partly drawn, cerotegument and setae of legs and pedipalps not drawn; B) detail of epimeral seta 4 c." httpUri="https://zenodo.org/record/279163/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
B).
</paragraph>
<paragraph id="8BC936CCD154FFBF11F7F97D49AFF8DA" blockId="20.[151,1437,1526,2036]" pageId="20" pageNumber="21">
<emphasis id="B902EADED154FFBF11F7F97D48E0F942" box="[197,401,1734,1758]" italics="true" pageId="20" pageNumber="21">Anogenital region</emphasis>
. 
<figureCitation id="134D2A49D154FFBF1093F97D4894F942" box="[417,485,1734,1758]" captionStart="FIGURE 9" captionStartId="19.[151,250,1536,1559]" captionTargetBox="[167,1419,356,1448]" captionTargetId="figure@19.[151,1436,305,1515]" captionTargetPageId="19" captionText="FIGURE 9. Metabelba romandiolae — specimen from S. Benedetto in Alpe. A—dorsal view, arrows point to light spots; Bventral view. Legs only partly drawn, leg setae and cerotegument not drawn." httpUri="https://zenodo.org/record/279159/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
B, 13A. Discidium (
<emphasis id="B902EADED154FFBF13FFF97D4B92F942" box="[717,739,1734,1758]" italics="true" pageId="20" pageNumber="21">di</emphasis>
) spiniform, well developed, curved posterolaterad. Setal formulas of anogenital region normal for the 
<taxonomicName id="4C764D4FD154FFBF134DF9534B8CF89C" box="[639,765,1768,1792]" class="Arachnida" family="Damaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Damaeidae</taxonomicName>
(
<bibRefCitation id="EFE74B3DD154FFBF123CF9534AB8F89C" author="Grandjean" box="[782,969,1768,1792]" pageId="20" pageNumber="21" refString="Grandjean, F. (1960) Damaeus arvernensis n. sp. (Oribate). Acarologia, 2, 250 - 275." type="journal article" year="1960">Grandjean 1960</bibRefCitation>
; 
<bibRefCitation id="EFE74B3DD154FFBF12E5F9524D08F89C" author="Norton" box="[983,1145,1768,1792]" pageId="20" pageNumber="21" refString="Norton, R. A. (1977 b) The genus Damaeus Koch (Acarina: Oribatei) in the United States. Acarologia, 19, 331 - 353." type="journal article" year="1977" yearSuffix="b">Norton 1977b</bibRefCitation>
), identical as in 
<taxonomicName id="4C764D4FD154FFBF140AF95149A3F8B8" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="denscanis" status="sp. nov.">
<emphasis id="B902EADED154FFBF140AF95149A3F8B8" italics="true" pageId="20" pageNumber="21">M. denscanis</emphasis>
</taxonomicName>
<emphasis id="B902EADED154FFBF11EAF8B74866F8B8" bold="true" box="[216,279,1804,1828]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="A23157A5D154FFBF11EAF8B74863F8B8" box="[216,274,1804,1828]" pageId="20" pageNumber="21" rank="species">n.sp.</taxonomicNameLabel>
,
</emphasis>
see above. All anogenital setae slightly flattened and broadened. Lyrifissure 
<emphasis id="B902EADED154FFBF1542F8B74DE5F8B8" box="[1136,1172,1804,1828]" italics="true" pageId="20" pageNumber="21">iad</emphasis>
oblique, divergent posteriad.
</paragraph>
<paragraph id="8BC936CCD154FFBE11F7F8E94DCFFF69" blockId="20.[151,1437,1526,2036]" lastBlockId="21.[151,1435,151,246]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B902EADED154FFBF11F7F8E94870F8F5" box="[197,257,1874,1897]" italics="true" pageId="20" pageNumber="21">Legs.</emphasis>
<figureCitation id="134D2A49D154FFBF103AF8EB4814F8F4" box="[264,357,1872,1896]" captionStart="FIGURE 14" captionStartId="24.[151,250,1591,1614]" captionTargetBox="[180,1428,211,1553]" captionTargetId="figure@24.[151,1436,193,1570]" captionTargetPageId="24" captionText="FIGURE 14. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg I and II. A) left trochanter and femur I, antiaxial view; B) left genu, tibia and tarsus I, antiaxial view; C) left femur II, paraxial and slightly ventral view; D) left genu and tibia II, paraxial view; E) left tarsus II, paraxial view." httpUri="https://zenodo.org/record/279164/files/figure.png" pageId="20" pageNumber="21">Figs. 14</figureCitation>
, 
<figureCitation id="134D2A49D154FFBF1043F8EB48E1F8F4" box="[369,400,1872,1896]" captionStart="FIGURE 15" captionStartId="25.[151,250,1910,1933]" captionTargetBox="[187,1419,554,1870]" captionTargetId="figure@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 15. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg III and IV. A) right genu, tibia and tarsus I, antiaxial view; B) left trochanter and femur III, antiaxial and slightly ventral view; C) left genu, tibia and tarsus IV, antiaxial and slightly ventral view; D) left trochanter and femur IV, antiaxial and slightly ventral view." httpUri="https://zenodo.org/record/279165/files/figure.png" pageId="20" pageNumber="21">15</figureCitation>
. Legs monodactyl, moderately long, segments distinctly swollen in distal part (moniliform). Leg IV distinctly longer than body, leg I and III approximately as long as body, leg II slightly shorter than body (detailed measurements of leg segments in Table 1). Leg setae, except those on distal parts of tarsi, of unique conspicuous shape: broadly flattened in basal part, with long attenuate tip, sparsely covered with minute spines on outer surface. Leg setal formula identical with those of 
<taxonomicName id="4C764D4FD154FFBF1220F8674DC0F86F" authority="Strenzke, 1953" authorityName="Strenzke" authorityYear="1953" box="[786,1201,2011,2036]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="pulverosa">
<emphasis id="B902EADED154FFBF1220F8674A8EF868" box="[786,1023,2012,2036]" italics="true" pageId="20" pageNumber="21">Metabelba pulverosa</emphasis>
<bibRefCitation id="EFE74B3DD154FFBF153AF8604DC0F86F" author="Strenzke" box="[1032,1201,2011,2035]" pageId="20" pageNumber="21" refString="Hammen, L. van der &amp; Strenzke, K. (1953) A partial revision of the genus Metabelba Grandjean (Oribatei, Acari). Zoologische Mededelingen, 32 (14), 141 - 154." type="journal article" year="1953">Strenzke, 1953</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="EFE74B3DD154FFBF15DDF8674CFEF86F" author="Norton" box="[1263,1423,2011,2035]" pageId="20" pageNumber="21" refString="Norton, R. A. (1977 a) A review of J. Grandjean's system of leg chaetotaxy in the Oribatei and its application to Damaeidae. In: Dindal, D. L. (Ed.), Biology of oribatid mites, 33, SUNY College of Environmental Science and Forestry, Syracuse, New York, 33 - 62." type="journal article" year="1977" yearSuffix="a">Norton 1977a</bibRefCitation>
), and 
<taxonomicName id="4C764D4FD155FFBE11F5FF22482AFF2C" box="[199,347,152,176]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="denscanis" status="sp. nov.">
<emphasis id="B902EADED155FFBE11F5FF22482AFF2C" box="[199,347,152,176]" italics="true" pageId="21" pageNumber="22">M. denscanis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157A5D155FFBE1051FF2348D2FF2C" box="[355,419,152,176]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B902EADED155FFBE1051FF2348D2FF2C" bold="true" box="[355,419,152,176]" pageId="21" pageNumber="22">n. sp.</emphasis>
</taxonomicNameLabel>
(see above). Solenidia ω 
<emphasis id="B902EADED155FFBE13F0FF234B94FF2C" box="[706,741,152,176]" italics="true" pageId="21" pageNumber="22">1,2</emphasis>
on tarsus I, φ 
<emphasis id="B902EADED155FFBE12B1FF234AE0FF2C" box="[899,913,152,176]" italics="true" pageId="21" pageNumber="22">1</emphasis>
on tibia I, ω 
<emphasis id="B902EADED155FFBE1516FF234D37FF2C" box="[1060,1094,152,176]" italics="true" pageId="21" pageNumber="22">1,2</emphasis>
on tarsus II and φ on tibia IV long, tactile; φ 
<emphasis id="B902EADED155FFBE1008FF004839FF4F" box="[314,328,187,211]" italics="true" pageId="21" pageNumber="22">1</emphasis>
on tibia I about 
<date id="FFC8100CD155FFBE1330FF014B6FFF4E" box="[514,542,186,210]" pageId="21" pageNumber="22">3x</date>
longer than φ 
<emphasis id="B902EADED155FFBE138EFF004BBBFF4F" box="[700,714,187,211]" italics="true" pageId="21" pageNumber="22">2</emphasis>
; φ on tibia IV more than 
<date id="FFC8100CD155FFBE12D8FF014D74FF4E" box="[1002,1029,186,210]" pageId="21" pageNumber="22">2x</date>
longer than tibia itself. Solenidia of tibiae II and III and genua I, II and III of approximately same length as their respective setae 
<emphasis id="B902EADED155FFBE159BFF664DC6FF69" box="[1193,1207,221,245]" italics="true" pageId="21" pageNumber="22">d</emphasis>
.
</paragraph>
<caption id="DF096644D155FFBE11A5F8984875F80A" httpUri="https://zenodo.org/record/279161/files/figure.png" pageId="21" pageNumber="22" targetBox="[151,1404,289,1792]" targetPageId="21">
<paragraph id="8BC936CCD155FFBE11A5F8984875F80A" blockId="21.[151,1436,1827,1942]" pageId="21" pageNumber="22">
<emphasis id="B902EADED155FFBE11A5F898486EF8A5" bold="true" box="[151,287,1827,1850]" pageId="21" pageNumber="22">FIGURE 11.</emphasis>
<emphasis id="B902EADED155FFBE1017F89F4B51F8A6" box="[293,544,1828,1850]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C764D4FD155FFBE1017F89F4B6CF8A6" box="[293,541,1828,1850]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="romandiolae">Metabelba romandiolae</taxonomicName>
.
</emphasis>
—lectotype, coll. of P. Zangheri, drawn by F. Bernini (A, B); specimen from coll. of C. Willmann, labelled as &quot; 
<taxonomicName id="4C764D4FD155FFBE10B8F8F94B63F8C4" box="[394,530,1858,1880]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="setiger">
<emphasis id="B902EADED155FFBE10B8F8F94B63F8C4" box="[394,530,1858,1880]" italics="true" pageId="21" pageNumber="22">Belba setiger</emphasis>
</taxonomicName>
&quot;, drawn by L. Miko (C, D, E). A—dorsolateral view; B—tibia, genu and distal part of femur IV; C—trochanter and femur III; D—trochanter IV; E—dorsal view. Cerotegument not drawn in A, B and partly drawn in C, D, E.
</paragraph>
</caption>
<caption id="DF096644D156FFBD11A5F94A49A7F8D8" httpUri="https://zenodo.org/record/279162/files/figure.png" pageId="22" pageNumber="23" targetBox="[152,1432,249,1776]" targetPageId="22">
<paragraph id="8BC936CCD156FFBD11A5F94A49A7F8D8" blockId="22.[151,1436,1777,1860]" pageId="22" pageNumber="23">
<emphasis id="B902EADED156FFBD11A5F94A4853F89B" bold="true" box="[151,290,1777,1800]" pageId="22" pageNumber="23">FIGURE 12.</emphasis>
<taxonomicName id="4C764D4FD156FFBD1019F94A4B50F89B" box="[299,545,1777,1799]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED156FFBD1019F94A4B50F89B" box="[299,545,1777,1799]" italics="true" pageId="22" pageNumber="23">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED156FFBD158FF94A4CEDF89B" box="[1213,1436,1777,1799]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C764D4FD156FFBD158FF94A4CE5F89B" box="[1213,1428,1777,1799]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym). A—dorsal view. Legs only partly drawn, leg setae and cerotegument not drawn. B—exobothridial seta in detail.
</paragraph>
</caption>
<paragraph id="8BC936CCD156FFBD11F5F8D748BEF85B" blockId="22.[151,1436,1900,1991]" pageId="22" pageNumber="23">
The two specimens from Zangheri’ collection present variations in the leg setation: in 
<typeStatus id="54CD886ED156FFBD15BCF8D74D82F819" box="[1166,1267,1900,1925]" pageId="22" pageNumber="23" type="lectotype">
<emphasis id="B902EADED156FFBD15BCF8D74D82F819" bold="true" box="[1166,1267,1900,1925]" italics="true" pageId="22" pageNumber="23">lectotype</emphasis>
</typeStatus>
femur I with 9 setae on the right and 10 on the left; genua II with 4 on the right and 5 on the left; tibia III with 5 setae in the 
<typeStatus id="54CD886ED156FFBD146EF83649B7F85A" pageId="22" pageNumber="23" type="lectotype">
<emphasis id="B902EADED156FFBD146EF83649B7F85A" bold="true" italics="true" pageId="22" pageNumber="23">lectotype</emphasis>
</typeStatus>
and 
<quantity id="4C8E9B29D156FFBD11CEF8144856F85B" box="[252,295,1967,1991]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="22" pageNumber="23" unit="in" value="4.0">4 in</quantity>
<typeStatus id="54CD886ED156FFBD101FF81548B6F85B" box="[301,455,1966,1991]" pageId="22" pageNumber="23" type="paralectotype">
<emphasis id="B902EADED156FFBD101FF81548B6F85B" bold="true" box="[301,455,1966,1991]" italics="true" pageId="22" pageNumber="23">paralectotype</emphasis>
</typeStatus>
.
</paragraph>
<paragraph id="8BC936CCD157FFBC11F5FF2C4D06FE88" blockId="23.[151,1436,151,276]" pageId="23" pageNumber="24">
<emphasis id="B902EADED157FFBC11F5FF2C4BE7FF2C" bold="true" box="[199,662,151,176]" pageId="23" pageNumber="24">Geographical distribution and ecology.</emphasis>
<taxonomicName id="4C764D4FD157FFBC13AFFF224A21FF2C" box="[669,848,152,176]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED157FFBC13AFFF224A21FF2C" box="[669,848,152,176]" italics="true" pageId="23" pageNumber="24">M. romandiolae</emphasis>
</taxonomicName>
was collected originally by Pietro 
<bibRefCitation id="EFE74B3DD157FFBC15E4FF2C4986FF4D" author="Zangheri" pageId="23" pageNumber="24" refString="Zangheri, P. (1966) Ord. Acari. Sottord. Oribatei. pp. 611 - 634. In: &quot; Repertorio sistematico e topografico della Flora e Fauna vivente e fossile della Romagna. &quot;. Museo Civico di Storia naturale di Verona, Memorie fuori serie n. 1, 1966 - 1970, vol. 1 - 5, pp. 2174." type="journal article" year="1966">
Zangheri (
<bibRefCitation id="EFE74B3DD157FFBC147BFF2C4986FF4D" author="Zangheri" pageId="23" pageNumber="24" refString="Zangheri, P. (1966) Ord. Acari. Sottord. Oribatei. pp. 611 - 634. In: &quot; Repertorio sistematico e topografico della Flora e Fauna vivente e fossile della Romagna. &quot;. Museo Civico di Storia naturale di Verona, Memorie fuori serie n. 1, 1966 - 1970, vol. 1 - 5, pp. 2174." type="journal article" year="1966" yearSuffix="r">Zangheri 1966</bibRefCitation>
</bibRefCitation>
, as 
<emphasis id="B902EADED157FFBC1010FF0148D2FF4D" box="[290,419,186,209]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="4C764D4FD157FFBC1010FF0148FDFF4D" box="[290,396,186,209]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Damaeus</taxonomicName>
r.
</emphasis>
), a well-known naturalist from Romagna (
<bibRefCitation id="EFE74B3DD157FFBC12B7FF024D7DFF4D" author="Ruffo" box="[901,1036,185,209]" pageId="23" pageNumber="24" refString="Ruffo, S. (1987) Ricordo di Pietro Zangheri (1889 - 1983). Biogeographia. Lavori della societa italiana di biogeografia. Nuova serie, 10, 823 - 826." type="journal article" year="1987">Ruffo 1987</bibRefCitation>
), on Poggio Orticai (about 
<quantity id="4C8E9B29D157FFBC140DFF024CE6FF4D" box="[1343,1431,185,209]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1000.0">1000 m</quantity>
, Tosco-Emiliano Apennine, at present in the territory of the Italian National Park of Foreste Casentinesi) in humus of 
<taxonomicName id="4C764D4FD157FFBC1187FF464814FE88" box="[181,357,252,276]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="B902EADED157FFBC1187FF464814FE88" box="[181,357,252,276]" italics="true" pageId="23" pageNumber="24">Fagus sylvatica</emphasis>
</taxonomicName>
, and later determined and described by Max 
<bibRefCitation id="EFE74B3DD157FFBC126DFF474D03FE88" box="[863,1138,252,276]" pageId="23" pageNumber="24" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article">Sellnick (Sellnick 1943)</bibRefCitation>
.
</paragraph>
<caption id="DF096644D157FFBC11A5F8DD48EAF826" httpUri="https://zenodo.org/record/279163/files/figure.png" pageId="23" pageNumber="24" targetBox="[204,1428,277,1854]" targetPageId="23">
<paragraph id="8BC936CCD157FFBC11A5F8DD48EAF826" blockId="23.[151,1436,1894,1978]" pageId="23" pageNumber="24">
<emphasis id="B902EADED157FFBC11A5F8DD4853F8E0" bold="true" box="[151,290,1894,1917]" pageId="23" pageNumber="24">FIGURE 13.</emphasis>
<taxonomicName id="4C764D4FD157FFBC1019F8DC4B50F8E1" box="[299,545,1895,1917]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED157FFBC1019F8DC4B50F8E1" box="[299,545,1895,1917]" italics="true" pageId="23" pageNumber="24">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED157FFBC158FF8DC4CEDF8E1" box="[1213,1436,1895,1917]" italics="true" pageId="23" pageNumber="24">
<taxonomicName id="4C764D4FD157FFBC158FF8DC4CE5F8E1" box="[1213,1428,1895,1917]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym). A) ventral view in total. Legs only partly drawn, cerotegument and setae of legs and pedipalps not drawn; B) detail of epimeral seta 
<emphasis id="B902EADED157FFBC104FF81F48E4F826" box="[381,405,1956,1978]" italics="true" pageId="23" pageNumber="24">4c</emphasis>
.
</paragraph>
</caption>
<caption id="DF096644D158FFB311A5F98C4C37F916" httpUri="https://zenodo.org/record/279164/files/figure.png" pageId="24" pageNumber="25" targetBox="[180,1428,211,1553]" targetPageId="24">
<paragraph id="8BC936CCD158FFB311A5F98C4C37F916" blockId="24.[151,1436,1591,1674]" pageId="24" pageNumber="25">
<emphasis id="B902EADED158FFB311A5F98C4853F9D0" bold="true" box="[151,290,1591,1614]" pageId="24" pageNumber="25">FIGURE 14.</emphasis>
<taxonomicName id="4C764D4FD158FFB31019F98C4B50F9D1" box="[299,545,1591,1613]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED158FFB31019F98C4B50F9D1" box="[299,545,1591,1613]" italics="true" pageId="24" pageNumber="25">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED158FFB3158FF98C4CEDF9D1" box="[1213,1436,1591,1613]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C764D4FD158FFB3158FF98C4CE5F9D1" box="[1213,1428,1591,1613]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym)—leg I and II. A) left trochanter and femur I, antiaxial view; B) left genu, tibia and tarsus I, antiaxial view; C) left femur II, paraxial and slightly ventral view; D) left genu and tibia II, paraxial view; E) left tarsus II, paraxial view.
</paragraph>
</caption>
<paragraph id="8BC936CCD158FFB311F5F9084BF6F8B2" blockId="24.[151,1436,1713,2015]" pageId="24" pageNumber="25">
Twenty years later, 
<bibRefCitation id="EFE74B3DD158FFB31098F90A4B10F956" author="Mihelcic" box="[426,609,1713,1738]" pageId="24" pageNumber="25" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič (1963)</bibRefCitation>
collected and described this species as 
<taxonomicName id="4C764D4FD158FFB3151FF9084DCBF956" box="[1069,1210,1714,1738]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED158FFB3151FF9084DCBF956" box="[1069,1210,1714,1738]" italics="true" pageId="24" pageNumber="25">M. gladiator</emphasis>
</taxonomicName>
. We found no published record of 
<taxonomicName id="4C764D4FD158FFB31065F9684B31F977" box="[343,576,1747,1771]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED158FFB31065F9684B31F977" box="[343,576,1747,1771]" italics="true" pageId="24" pageNumber="25">Metabelba gladiator</emphasis>
</taxonomicName>
besides the 
<typeStatus id="54CD886ED158FFB313E6F96F4A75F970" box="[724,772,1748,1772]" pageId="24" pageNumber="25">type</typeStatus>
locality, Sankt Veit im Jauntal in Kärnten, South 
<collectingCountry id="F361765CD158FFB31478F9684CEDF977" box="[1354,1436,1747,1771]" name="Austria" pageId="24" pageNumber="25">Austria</collectingCountry>
(
<bibRefCitation id="EFE74B3DD158FFB311ACF9484837F890" author="Mihelcic" box="[158,326,1779,1804]" pageId="24" pageNumber="25" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič 1963</bibRefCitation>
), a village located at approximately 
<quantity id="4C8E9B29D158FFB313EFF94F4A2DF890" box="[733,860,1780,1804]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="5.6" metricValueMin="5.4" pageId="24" pageNumber="25" unit="m" value="550.0" valueMax="560.0" valueMin="540.0">540–560 m</quantity>
a.s.l. 
<bibRefCitation id="EFE74B3DD158FFB312ADF9484D25F890" author="Mihelcic" box="[927,1108,1779,1804]" pageId="24" pageNumber="25" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič (1963)</bibRefCitation>
found the specimens in open landscape and forest margins, all dry stands.
</paragraph>
<paragraph id="8BC936CCD158FFB311F7F88C4D8AF80B" blockId="24.[151,1436,1713,2015]" pageId="24" pageNumber="25">
The specimen from Willmann’s collection may relate to the record of one individual of &quot; 
<taxonomicName id="4C764D4FD158FFB31433F88C4984F8EF" authority="Kulcz." authorityName="Kulcz." class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="setiger">
<emphasis id="B902EADED158FFB31433F88C4CEDF8D3" box="[1281,1436,1847,1871]" italics="true" pageId="24" pageNumber="25">Belba setiger</emphasis>
(Kulcz.)
</taxonomicName>
&quot; from Moravian Karst (Moravia, 
<collectingCountry id="F361765CD158FFB313B2F8E04A4BF8EF" box="[640,826,1883,1907]" name="Czech Republic" pageId="24" pageNumber="25">Czech Republic</collectingCountry>
), published by 
<bibRefCitation id="EFE74B3DD158FFB312DEF8E04DC3F8EF" author="Willmann" box="[1004,1202,1883,1907]" pageId="24" pageNumber="25" refString="Willmann, C. (1954) Mahrische Acari, hauptsachlich aus dem Gebiete des mahrischen Karstes, gesammelt von Prof. Dr. K. Absolon, Brunn. C eskoslovenska parasitologie, 1, 213 - 272." type="journal article" year="1954">Willmann (1954)</bibRefCitation>
. It was collected by Prof. Karel Absolon in 1900 from moss in the cave of Michalova jeskynĕ (about 
<quantity id="4C8E9B29D158FFB31510F8C44DD0F80B" box="[1058,1185,1919,1943]" metricMagnitude="2" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="24" pageNumber="25" unit="m" value="445.0" valueMax="450.0" valueMin="440.0">440–450 m</quantity>
a. s. l.).
</paragraph>
</subSubSection>
<subSubSection id="C36C6547D158FFB211F7F81F48F4FE53" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="8BC936CCD158FFB211F7F81F4829FF48" blockId="24.[151,1436,1713,2015]" lastBlockId="25.[151,1436,151,463]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<taxonomicName id="4C764D4FD158FFB311F7F81F4808F827" ID-CoL="8TBWG" box="[197,377,1955,1979]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED158FFB311F7F81F4808F827" box="[197,377,1955,1979]" italics="true" pageId="24" pageNumber="25">M. romandiolae</emphasis>
</taxonomicName>
was collected in further Italian localities: Farma valley (
<quantity id="4C8E9B29D158FFB312C9F8184D11F827" box="[1019,1120,1955,1979]" metricMagnitude="2" metricUnit="m" metricValue="1.51" metricValueMax="3.0" metricValueMin="0.02" pageId="24" pageNumber="25" unit="m" value="151.0" valueMax="300.0" valueMin="2.0">2–300 m</quantity>
a.s.l.), near Siena (southern Tuscany) and Altopiano del Cansiglio (Mt. Pizzoc, 
<quantity id="4C8E9B29D158FFB313C3F87C4A3BF843" box="[753,842,1991,2015]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="m" value="1200.0">1200 m</quantity>
a.s.l., Treviso, Venetian Pre-Alps) (
<bibRefCitation id="EFE74B3DD158FFB315DEF87C4CE1F843" author="Bernini" box="[1260,1424,1991,2015]" pageId="24" pageNumber="25" refString="Bernini, F. (1982) Notulae Oribatologicae XXVII. Contributo alla conoscenza degli Oribatei della Val di Farma (Toscana meridionale). Redia, 65, 377 - 405." type="journal article" year="1982">Bernini, 1982</bibRefCitation>
), Pollino Massif (Southern 
<collectingCountry id="F361765CD159FFB2108AFF2C489DFF33" box="[440,492,151,175]" name="Italy" pageId="25" pageNumber="26">Italy</collectingCountry>
, Calabria) (
<bibRefCitation id="EFE74B3DD159FFB2135CFF2C4A3AFF33" author="Bernini" box="[622,843,151,176]" pageId="25" pageNumber="26" refString="Bernini, F., Avanzati, A. M. &amp; Bernini, S. (1987) Notulae Oribatologicae XXXVII. Gli Acari Oribatei del Massiccio del Pollino (Italia Meridionale): aspetti faunistici e biogeografici. Biogeographia. Lavori della societa italiana di biogeografia. Nuova serie, 10, 379 - 488." type="journal article" year="1987">
Bernini 
<emphasis id="B902EADED159FFB213F9FF224A75FF2C" box="[715,772,152,176]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
, 1987
</bibRefCitation>
) and Nebrodi Mountains (north-eastern Sicily) (
<bibRefCitation id="EFE74B3DD159FFB2145BFF2C483AFF48" author="Bernini" pageId="25" pageNumber="26" refString="Bernini, F., Arcidiacono, R. &amp; Migliorini, M. (1996) Notulae Oribatologicae LXII. Contributo alla conoscenza degli Acari Oribatei dei Monti Nebrodi e dei Peloritani (Sicilia Nordorientale). Animalia, 21, 131 - 172." type="journal article" year="1996">
Bernini 
<emphasis id="B902EADED159FFB211FBFF064873FF48" box="[201,258,188,212]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
, 1996
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC936CCD159FFB211F7FF6448F4FE53" blockId="25.[151,1436,151,463]" pageId="25" pageNumber="26">
This species occurs mainly in the Italian peninsula reaching in the south to Sicily Island, but large populations are found very rarely. Despite these southern localizations, 
<taxonomicName id="4C764D4FD159FFB21203FEBE4A95FE80" ID-CoL="8TBWG" box="[817,996,260,284]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED159FFB21203FEBE4A95FE80" box="[817,996,260,284]" italics="true" pageId="25" pageNumber="26">M. romandiolae</emphasis>
</taxonomicName>
is considered to be a mesophilous species generally inhabiting montane sites all characterized by humus of 
<taxonomicName id="4C764D4FD159FFB21282FE924D10FEDC" ID-CoL="3DSK5" box="[944,1121,296,320]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="species" species="sylvatica">
<emphasis id="B902EADED159FFB21282FE924D10FEDC" box="[944,1121,296,320]" italics="true" pageId="25" pageNumber="26">Fagus sylvatica</emphasis>
</taxonomicName>
, and by elevations over the thousand meters. The unique exception is the Farma Valley, a site located at 
<quantity id="4C8E9B29D159FFB212DAFEF74D3BFEF8" box="[1000,1098,332,356]" metricMagnitude="2" metricUnit="m" metricValue="1.51" metricValueMax="3.0" metricValueMin="0.02" pageId="25" pageNumber="26" unit="m" value="151.0" valueMax="300.0" valueMin="2.0">2–300 m</quantity>
a.s.l., but retained as a “mesic oasis”, almost an “island”, colonized during the past glaciations by montane elements (
<bibRefCitation id="EFE74B3DD159FFB2154FFED44C2AFE1B" author="Bernini" box="[1149,1371,367,392]" pageId="25" pageNumber="26" refString="Bernini, F., Avanzati, A. M., Baratti, M. &amp; Migliorini, M. (1995) Oribatid mites (Acari Oribatida) of the Farma Valley (Southern Tuscany). Notulae Oribatologicae LXV. Redia, 78, 45 - 129." type="journal article" year="1995">
Bernini 
<emphasis id="B902EADED159FFB215EEFECA4C67FE14" box="[1244,1302,368,392]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
1995
</bibRefCitation>
). The putative record of this species from a cave in Moravian Karst presented above (
<bibRefCitation id="EFE74B3DD159FFB21510FE2F4DA6FE30" author="Willmann" box="[1058,1239,404,428]" pageId="25" pageNumber="26" refString="Willmann, C. (1954) Mahrische Acari, hauptsachlich aus dem Gebiete des mahrischen Karstes, gesammelt von Prof. Dr. K. Absolon, Brunn. C eskoslovenska parasitologie, 1, 213 - 272." type="journal article" year="1954">Willmann 1954</bibRefCitation>
), might represent another glacial relict.
</paragraph>
</subSubSection>
<caption id="DF096644D159FFB211A5F8CD48BEF874" httpUri="https://zenodo.org/record/279165/files/figure.png" pageId="25" pageNumber="26" targetBox="[187,1419,554,1870]" targetPageId="25">
<paragraph id="8BC936CCD159FFB211A5F8CD48BEF874" blockId="25.[151,1436,1910,2024]" pageId="25" pageNumber="26">
<emphasis id="B902EADED159FFB211A5F8CD4853F817" bold="true" box="[151,290,1910,1933]" pageId="25" pageNumber="26">FIGURE 15.</emphasis>
<taxonomicName id="4C764D4FD159FFB21019F8CD4B50F810" box="[299,545,1910,1932]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED159FFB21019F8CD4B50F810" box="[299,545,1910,1932]" italics="true" pageId="25" pageNumber="26">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED159FFB2158FF8CD4CEDF810" box="[1213,1436,1910,1932]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="4C764D4FD159FFB2158FF8CD4CE5F810" box="[1213,1428,1910,1932]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym)—leg III and IV. A) right genu, tibia and tarsus I, antiaxial view; B) left trochanter and femur III, antiaxial and slightly ventral view; C) left genu, tibia and tarsus IV, antiaxial and slightly ventral view; D) left trochanter and femur IV, antiaxial and slightly ventral view.
</paragraph>
</caption>
<subSubSection id="C36C6547D15AFFB111F5FF2349BAF8AD" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BC936CCD15AFFB111F5FF234AE2FF48" blockId="26.[151,1436,151,679]" pageId="26" pageNumber="27">
<emphasis id="B902EADED15AFFB111F5FF23484AFF2C" bold="true" box="[199,315,152,176]" pageId="26" pageNumber="27">Remarks.</emphasis>
<taxonomicName id="4C764D4FD15AFFB11070FF224887FF2C" box="[322,502,152,176]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED15AFFB11070FF224887FF2C" box="[322,502,152,176]" italics="true" pageId="26" pageNumber="27">M. romandiolae</emphasis>
</taxonomicName>
can be easily recognized by the unique shape of its leg setae. Moreover, the tips of the notogastral setae are finely serrated with minute backward teeth.
</paragraph>
<paragraph id="8BC936CCD15AFFB111F7FF644CF3FEA3" blockId="26.[151,1436,151,679]" pageId="26" pageNumber="27">
The original description of 
<taxonomicName id="4C764D4FD15AFFB11337FF5A4A0FFF6B" authority="Sellnick, 1943" authorityName="Sellnick" authorityYear="1943" box="[517,894,223,248]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED15AFFB11337FF5A4BCDFF64" box="[517,700,224,248]" italics="true" pageId="26" pageNumber="27">M. romandiolae</emphasis>
(
<bibRefCitation id="EFE74B3DD15AFFB113FCFF644A04FF6B" author="Sellnick" box="[718,885,223,247]" pageId="26" pageNumber="27" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick, 1943</bibRefCitation>
)
</taxonomicName>
is insufficient and was not illustrated; 
<bibRefCitation id="EFE74B3DD15AFFB1147AFF644990FE80" author="Bernini" pageId="26" pageNumber="27" refString="Bernini, F. (1982) Notulae Oribatologicae XXVII. Contributo alla conoscenza degli Oribatei della Val di Farma (Toscana meridionale). Redia, 65, 377 - 405." type="journal article" year="1982">Bernini (1982)</bibRefCitation>
designated the 
<typeStatus id="54CD886ED15AFFB110A2FEBE4896FE80" box="[400,487,261,284]" pageId="26" pageNumber="27" type="neotype">
<emphasis id="B902EADED15AFFB110A2FEBE4896FE80" box="[400,487,261,284]" italics="true" pageId="26" pageNumber="27">neotype</emphasis>
</typeStatus>
(now designated as the 
<typeStatus id="54CD886ED15AFFB113C7FEBF4A2BFE80" box="[757,858,260,284]" pageId="26" pageNumber="27" type="lectotype">
<emphasis id="B902EADED15AFFB113C7FEBF4A2BFE80" box="[757,858,260,284]" italics="true" pageId="26" pageNumber="27">lectotype</emphasis>
</typeStatus>
by us) of this species on the specimens returned by Sellnick to Pietro Zangheri and belonging to the 
<typeStatus id="54CD886ED15AFFB1138AFE934B99FEDC" box="[696,744,296,320]" pageId="26" pageNumber="27">type</typeStatus>
series of 
<bibRefCitation id="EFE74B3DD15AFFB11267FE9C4D75FEA3" author="Sellnick" box="[853,1028,295,319]" pageId="26" pageNumber="27" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick (1943)</bibRefCitation>
, but did not make a redescription.
</paragraph>
<paragraph id="8BC936CCD15AFFB111F7FEF74D1EFD3B" blockId="26.[151,1436,151,679]" pageId="26" pageNumber="27">
The original description of 
<taxonomicName id="4C764D4FD15AFFB11333FEF64BE1FEF8" box="[513,656,332,356]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED15AFFB11333FEF64BE1FEF8" box="[513,656,332,356]" italics="true" pageId="26" pageNumber="27">M. gladiator</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFE74B3DD15AFFB1138DFEF04A06FEF8" author="Mihelcic" box="[703,887,331,356]" pageId="26" pageNumber="27" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič (1963)</bibRefCitation>
is comparatively detailed and sufficiently illustrated, but some important characters were described incorrectly, as we revealed with detailed study of the 
<typeStatus id="54CD886ED15AFFB1145EFECB4CEDFE14" box="[1388,1436,368,392]" pageId="26" pageNumber="27">type</typeStatus>
specimen. Namely, he described and illustrated the species with two pairs of centrodorsal tubercles (
<emphasis id="B902EADED15AFFB115C1FE2E4C64FE30" box="[1267,1301,405,428]" italics="true" pageId="26" pageNumber="27">Da</emphasis>
, 
<emphasis id="B902EADED15AFFB11411FE2E4C34FE30" box="[1315,1349,405,428]" italics="true" pageId="26" pageNumber="27">Dp</emphasis>
), but in fact only 
<emphasis id="B902EADED15AFFB11036FE024856FE4C" box="[260,295,441,464]" italics="true" pageId="26" pageNumber="27">Da</emphasis>
is present. He presented the following incorrect leg setal formulas (solenidia and famulus were not distinguished from normal setae): leg I (from femur to tarsus) 
<date id="FFC8100CD15AFFB11261FE674AC9FE68" box="[851,952,476,500]" pageId="26" pageNumber="27" value="1918-03-07" valueMax="1918-06-07" valueMin="1918-03-07">7-3-6-18</date>
; leg II (from femur to tarsus) 
<date id="FFC8100CD15AFFB1143BFE674C1FFE68" box="[1289,1390,476,500]" pageId="26" pageNumber="27" value="1916-03-07" valueMax="1916-05-07" valueMin="1916-03-07">7-3-5-16</date>
; leg III (trochanter to tarsus) 
<date id="FFC8100CD15AFFB1109EFE444B56FD8B" box="[428,551,511,535]" pageId="26" pageNumber="27" value="1911-04-03" valueMax="1911-05-06" valueMin="1911-04-03">3-6-4-5-11</date>
; leg IV (trochanter to tarsus) 
<date id="FFC8100CD15AFFB11246FE444A9EFD8B" box="[884,1007,511,535]" pageId="26" pageNumber="27" value="1911-03-03" valueMax="1911-05-06" valueMin="1911-03-03">3-6-3-5-11</date>
. The generally lower numbers on particular segments might raise doubts about the generic placement of 
<taxonomicName id="4C764D4FD15AFFB112A0FD9E4D6EFDA0" box="[914,1055,548,572]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED15AFFB112A0FD9E4D6EFDA0" box="[914,1055,548,572]" italics="true" pageId="26" pageNumber="27">M. gladiator</emphasis>
</taxonomicName>
, but in fact the leg setation of the 
<typeStatus id="54CD886ED15AFFB111A5FDF349B6FDFC" box="[151,199,584,608]" pageId="26" pageNumber="27">type</typeStatus>
specimen is normal for 
<taxonomicName id="4C764D4FD15AFFB110E8FDF34B23FDFC" box="[474,594,584,608]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B902EADED15AFFB110E8FDF34B23FDFC" box="[474,594,584,608]" italics="true" pageId="26" pageNumber="27">Metabelba</emphasis>
</taxonomicName>
(see above and 
<figureCitation id="134D2A49D15AFFB1123BFDFC4A17FDC3" box="[777,870,583,607]" captionStart="FIGURE 14" captionStartId="24.[151,250,1591,1614]" captionTargetBox="[180,1428,211,1553]" captionTargetId="figure@24.[151,1436,193,1570]" captionTargetPageId="24" captionText="FIGURE 14. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg I and II. A) left trochanter and femur I, antiaxial view; B) left genu, tibia and tarsus I, antiaxial view; C) left femur II, paraxial and slightly ventral view; D) left genu and tibia II, paraxial view; E) left tarsus II, paraxial view." httpUri="https://zenodo.org/record/279164/files/figure.png" pageId="26" pageNumber="27">Figs. 14</figureCitation>
, 
<figureCitation id="134D2A49D15AFFB11241FDFC4AE3FDC3" box="[883,914,583,607]" captionStart="FIGURE 15" captionStartId="25.[151,250,1910,1933]" captionTargetBox="[187,1419,554,1870]" captionTargetId="figure@25.[151,1436,517,1889]" captionTargetPageId="25" captionText="FIGURE 15. Metabelba romandiolae — specimen from collection of F. Mihelčič labelled as &quot; type of Metabelba gladiator &quot; (junior synonym) — leg III and IV. A) right genu, tibia and tarsus I, antiaxial view; B) left trochanter and femur III, antiaxial and slightly ventral view; C) left genu, tibia and tarsus IV, antiaxial and slightly ventral view; D) left trochanter and femur IV, antiaxial and slightly ventral view." httpUri="https://zenodo.org/record/279165/files/figure.png" pageId="26" pageNumber="27">15</figureCitation>
). Parastigmatic apophyses (
<emphasis id="B902EADED15AFFB115FEFDF34D99FDC3" box="[1228,1256,584,607]" italics="true" pageId="26" pageNumber="27">Sa</emphasis>
, 
<emphasis id="B902EADED15AFFB115C5FDF34C62FDC3" box="[1271,1299,584,607]" italics="true" pageId="26" pageNumber="27">Sp</emphasis>
) are neither illustrated nor described in the text. Mihelčič also did not describe the filamentous cerotegument and only stated that the body surface is smooth or “covered by debris” (&quot;Schmutz&quot; in 
<collectingCountry id="F361765CD15AFFB11294FD344D71FD3B" box="[934,1024,655,679]" name="Germany" pageId="26" pageNumber="27">German</collectingCountry>
original).
</paragraph>
<caption id="DF096644D15AFFB111A5FA7A4813F98B" httpUri="https://zenodo.org/record/279166/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1436,721,1451]" targetPageId="26">
<paragraph id="8BC936CCD15AFFB111A5FA7A4813F98B" blockId="26.[151,1437,1473,1559]" pageId="26" pageNumber="27">
<emphasis id="B902EADED15AFFB111A5FA7A4853FA4B" bold="true" box="[151,290,1473,1496]" pageId="26" pageNumber="27">FIGURE 16.</emphasis>
<taxonomicName id="4C764D4FD15AFFB11019FA7A4B50FA4B" box="[299,545,1473,1495]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED15AFFB11019FA7A4B50FA4B" box="[299,545,1473,1495]" italics="true" pageId="26" pageNumber="27">Metabelba romandiolae</emphasis>
</taxonomicName>
—specimen from collection of F. Mihelčič labelled as &quot;type of 
<emphasis id="B902EADED15AFFB1158FFA7A4CEDFA4B" box="[1213,1436,1473,1495]" italics="true" pageId="26" pageNumber="27">
<taxonomicName id="4C764D4FD15AFFB1158FFA7A4CE5FA4B" box="[1213,1428,1473,1495]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gladiator">Metabelba gladiator</taxonomicName>
&quot;
</emphasis>
(junior synonym), light micrographs, Nomarski DIC. A) detail of genu III with typical leg setae; B) cerotegument on tibia IV. Scale bars: 100 Μm.
</paragraph>
</caption>
<paragraph id="8BC936CCD15AFFB111F7F9FA48EAF959" blockId="26.[151,1436,1601,1841]" pageId="26" pageNumber="27">
<bibRefCitation id="EFE74B3DD15AFFB111F7F9FA482FF9C5" author="Subias" box="[197,350,1601,1625]" pageId="26" pageNumber="27" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1748 - 2002). Graellsia, 60, 3 - 305." type="journal article" year="2004">Subías (2004</bibRefCitation>
, 
<bibRefCitation id="EFE74B3DD15AFFB1105FF9FA48D6F9C5" author="Subias" box="[365,423,1601,1625]" pageId="26" pageNumber="27" refString="Subias, L. (2008, 2009, 2011) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (Excepto fosiles). 540 pp. The last version is available from: http: // www. ucm. es / info / zoo / Artropodos / Catalogo. pdf [last changes in February 2011; cited 24. 8. 2011]" type="book" year="2008">2008</bibRefCitation>
, 2009) was the first author who listed 
<taxonomicName id="4C764D4FD15AFFB11242F9F94D70F9C5" box="[880,1025,1601,1625]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED15AFFB11242F9F94D70F9C5" box="[880,1025,1601,1625]" italics="true" pageId="26" pageNumber="27">M. gladiator</emphasis>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="4C764D4FD15AFFB11416F9FA48AEF9E1" authority="Sellnick 1943" authorityName="Sellnick" authorityYear="1943" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED15AFFB11416F9FA4854F9E2" italics="true" pageId="26" pageNumber="27">Metabelba romandiolae</emphasis>
(
<bibRefCitation id="EFE74B3DD15AFFB11007F9DE48A6F9E1" author="Sellnick" box="[309,471,1637,1661]" pageId="26" pageNumber="27" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick 1943</bibRefCitation>
)
</taxonomicName>
. He provided no reasons or reference to previous works, but kindly informed us (L. Subías pers. comm., 2010) that his assumption was based on 
<bibRefCitation id="EFE74B3DD15AFFB11278F9324D43F93D" box="[842,1074,1673,1697]" pageId="26" pageNumber="27" refString="Bernini, F., Avanzati, A. M. &amp; Bernini, S. (1987) Notulae Oribatologicae XXXVII. Gli Acari Oribatei del Massiccio del Pollino (Italia Meridionale): aspetti faunistici e biogeografici. Biogeographia. Lavori della societa italiana di biogeografia. Nuova serie, 10, 379 - 488." type="journal article">
Bernini 
<emphasis id="B902EADED15AFFB11297F9314AAFF93D" box="[933,990,1673,1697]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(1987)
</bibRefCitation>
, who hypothesized that the species might be identical.
</paragraph>
<paragraph id="8BC936CCD15AFFB111F7F96949BAF8AD" blockId="26.[151,1436,1601,1841]" pageId="26" pageNumber="27">
In the present paper we analyzed the 
<typeStatus id="54CD886ED15AFFB11345F9694BD6F976" box="[631,679,1746,1770]" pageId="26" pageNumber="27">type</typeStatus>
material of both names and provided a modern redescription. We consider the synonymy of 
<taxonomicName id="4C764D4FD15AFFB110F6F94C4A66F891" authority="Mihelcic, 1963" authorityName="Mihelcic" authorityYear="1963" box="[452,791,1780,1806]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="gladiator">
<emphasis id="B902EADED15AFFB110F6F94C4B23F892" box="[452,594,1782,1806]" italics="true" pageId="26" pageNumber="27">M. gladiator</emphasis>
(
<bibRefCitation id="EFE74B3DD15AFFB11350F94F4A7EF891" author="Mihelcic" box="[610,783,1780,1805]" pageId="26" pageNumber="27" refString="Mihelcic, F. (1963) Beitrag zur Kenntnis der Oribatidenfauna Karntens (Oribatei, Acarina) Zoologischer Anzeiger, 170, 230 - 240." type="journal article" year="1963">Mihelčič, 1963</bibRefCitation>
)
</taxonomicName>
with respect to 
<taxonomicName id="4C764D4FD15AFFB112FCF94C4C31F891" authority="Sellnick, 1943" authorityName="Sellnick" authorityYear="1943" box="[974,1344,1781,1806]" class="Arachnida" family="Damaeidae" genus="Metabelba" kingdom="Animalia" order="Sarcoptiformes" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="romandiolae">
<emphasis id="B902EADED15AFFB112FCF94C4DF3F891" box="[974,1154,1781,1806]" italics="true" pageId="26" pageNumber="27">M. romandiolae</emphasis>
(
<bibRefCitation id="EFE74B3DD15AFFB115A0F94E4C49F891" author="Sellnick" box="[1170,1336,1781,1805]" pageId="26" pageNumber="27" refString="Sellnick, M. (1943) Einige neue Milben aus der Romagna. Bollettino della Societa Entomologica Italiana 75, 22 - 26." type="journal article" year="1943">Sellnick, 1943</bibRefCitation>
)
</taxonomicName>
conclusive.
</paragraph>
</subSubSection>
</treatment>
</document>