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<document id="BA328FE12D9D8DBB080FC405A5232C5A" ID-DOI="10.1206/816.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5401238" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1630334480974" checkinUser="felipe" docAuthor="Herman, Lee H." docDate="2013" docId="03DF87947D5ED12EFD5253BEFCA90798" docLanguage="en" docName="B375.pdf" docOrigin="Bulletin of the American Museum of Natural History 2013 (375)" docSource="http://www.bioone.org/doi/abs/10.1206/816.1" docStyle="DocumentStyle:3507F1A1389FCD0DFF4CB4E0ACF2D98A.3:BulAmeMusNatHis.2011-.journal_article.1cover.type2" docStyleId="3507F1A1389FCD0DFF4CB4E0ACF2D98A" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type2" docStyleVersion="3" docTitle="Oedichirus distortus Herman 2013, new species" docType="treatment" docVersion="4" lastPageNumber="73" masterDocId="FFE6FFEC7D18D167FFB6536AFFC2023A" masterDocTitle="Revision Of The New World Species Of Oedichirus (Coleoptera: Staphylinidae: Paederinae: Pinophilini: Procirrina)" masterLastPageNumber="137" masterPageNumber="1" pageNumber="70" updateTime="1699232470662" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EB90D5D11EF5C31C7CC33B30BEE42A08">Revision Of The New World Species Of Oedichirus (Coleoptera: Staphylinidae: Paederinae: Pinophilini: Procirrina)</mods:title>
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<treatment id="03DF87947D5ED12EFD5253BEFCA90798" LSID="urn:lsid:plazi:treatment:03DF87947D5ED12EFD5253BEFCA90798" httpUri="http://treatment.plazi.org/id/03DF87947D5ED12EFD5253BEFCA90798" lastPageId="73" lastPageNumber="73" pageId="70" pageNumber="70">
<subSubSection id="C36C65097D5ED121FD5253BEFBDA0332" pageId="70" pageNumber="70" type="nomenclature">
<paragraph id="8BC936827D5ED121FD5253BEFB9D02D1" blockId="70.[740,1119,212,265]" box="[740,1119,212,235]" pageId="70" pageNumber="70">
<heading id="D08181EE7D5ED121FD5253BEFB9D02D1" box="[740,1119,212,235]" centered="true" fontSize="9" level="2" pageId="70" pageNumber="70" reason="2">
<taxonomicName id="4C764D017D5ED121FD5253BEFC0B02D1" ID-CoL="48VTQ" authority="Herman, 2013" authorityName="Herman" authorityYear="2013" box="[740,969,212,235]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus" status="sp. nov.">
<emphasis id="B902EA907D5ED121FD5253BEFC0B02D1" bold="true" box="[740,969,212,235]" italics="true" pageId="70" pageNumber="70">Oedichirus distortus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23157EB7D5ED121FC6F53BFFB9D02D1" box="[985,1119,213,235]" pageId="70" pageNumber="70" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BC936827D5ED121FC9A5399FBDA0332" blockId="70.[740,1119,212,265]" box="[812,1048,243,265]" pageId="70" pageNumber="70">
<heading id="D08181EE7D5ED121FC9A5399FBDA0332" box="[812,1048,243,265]" centered="true" fontSize="9" level="2" pageId="70" pageNumber="70" reason="2">
<figureCitation id="134D2A077D5ED121FC9A5399FC690332" box="[812,939,243,265]" captionStart="Figs" captionStartId="37.[150,198,1630,1649]" captionTargetBox="[132,1222,211,1597]" captionTargetId="figure-1@37.[131,1224,210,1600]" captionTargetPageId="37" captionText="Figs. 3940. Oedichirus species. Distribution. 39. Mesoamerica and the Caribbean. 40. South America. (Species described prior to present work preceded with *. Symbols ³ or ♀ signify species known only by the sex indicated. Open circle denotes multiple collecting sites.)" figureDoi="http://doi.org/10.5281/zenodo.5401272" httpUri="https://zenodo.org/record/5401272/files/figure.png" pageId="70" pageNumber="70">Figures 40</figureCitation>
,
<figureCitation id="134D2A077D5ED121FC0E5399FBDA0332" box="[952,1048,243,264]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="71.[150,198,1529,1548]" captionStartId-1="72.[252,290,1198,1217]" captionTargetBox-0="[131,1224,212,1499]" captionTargetBox-1="[96,1189,211,1167]" captionTargetId-0="figure-51@71.[131,1224,212,1499]" captionTargetId-1="figure-196@72.[96,1189,211,1168]" captionTargetPageId-0="71" captionTargetPageId-1="72" captionText-0="Figs. 112117. Oedichirus distortus. 112. Aedeagus, right lateral. 113. Aedeagus, ventral. 114. Sternum VIII, male. 115. Sternum VIII, setae removed, male. 116. Segment IX, ventral, female. 117. Sternum IX, male." captionText-1="Fig. 118. Oedichirus distortus. Segment IX, vulvar plate, enlarged (CLSM)." figureDoi-0="http://doi.org/10.5281/zenodo.5401308" figureDoi-1="http://doi.org/10.5281/zenodo.5401310" httpUri-0="https://zenodo.org/record/5401308/files/figure.png" httpUri-1="https://zenodo.org/record/5401310/files/figure.png" pageId="70" pageNumber="70">112118</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D5ED121FD2D524BFB980123" pageId="70" pageNumber="70" type="materials_examined">
<paragraph id="8BC936827D5ED121FD2D524BFB8F0380" blockId="70.[667,1194,288,793]" pageId="70" pageNumber="70">
TYPE MATERIAL:
<typeStatus id="54CD88207D5ED121FCD8524AFC110309" box="[878,979,288,307]" pageId="70" pageNumber="70" type="holotype">Holotype</typeStatus>
. Male.
<collectingCountry id="F36176127D5ED121FBF0524AFB670309" box="[1094,1189,288,307]" name="Ecuador" pageId="70" pageNumber="70">Ecuador</collectingCountry>
:
<collectingRegion id="49B2F8607D5ED121FD035256FCCA0375" box="[693,776,316,335]" country="Ecuador" name="Pastaza" pageId="70" pageNumber="70">Pastaza</collectingRegion>
. Ashuara indian village on R. Macuma nr. R. Morona. VII:1116:1971. B. Malkin./ night sweeping along forest trail/
<typeStatus id="54CD88207D5ED121FBFE521BFB6B03BE" box="[1096,1193,369,388]" pageId="70" pageNumber="70" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C764D017D5ED121FD0352E1FC3C03A5" authority="Herman." authorityName="Herman." box="[693,1022,395,415]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FD0352E1FC4903A5" box="[693,907,395,415]" italics="true" pageId="70" pageNumber="70">Oedichirus distortus</emphasis>
Herman.
</taxonomicName>
Deposited in the Field Museum of Natural History.
</paragraph>
<paragraph id="8BC936827D5ED121FD2D52A9FBED007B" blockId="70.[667,1194,288,793]" pageId="70" pageNumber="70">
<materialsCitation id="3B1E3CDF7D5ED121FD2D52A9FB9F03CA" collectionCode="FMNH, AMNH" pageId="70" pageNumber="70" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<typeStatus id="54CD88207D5ED121FD2D52A9FCCD03EF" box="[667,783,451,470]" pageId="70" pageNumber="70" type="paratype">PARATYPES</typeStatus>
: One male,
<specimenCount id="9D70FD0B7D5ED121FC2752A9FBD303EF" box="[913,1041,450,470]" pageId="70" pageNumber="70" type="female">two females</specimenCount>
: Same data as holotype (
<specimenCount id="9D70FD0B7D5ED121FC9252B4FC4C03CA" box="[804,910,477,497]" pageId="70" pageNumber="70" type="female">2 females</specimenCount>
,
<collectionCode id="ED67AE477D5ED121FC2B52B4FC3203CB" box="[925,1008,478,497]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="70" pageNumber="70" type="Museum">FMNH</collectionCode>
,
<collectionCode id="ED67AE477D5ED121FBB652B7FB9403CA" box="[1024,1110,477,496]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="70" pageNumber="70">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1E3CDF7D5ED121FBDB52B4FBED007B" collectingDate="1982-06-02" collectionCode="CNC" collectorName="Campana Cocha, E. &amp; H. Frania" country="Ecuador" elevation="335" location="Misahualli" pageId="70" pageNumber="70" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="B902EA907D5ED121FBDB52B4FD150031" bold="true" pageId="70" pageNumber="70">
<collectingCountry id="F36176127D5ED121FBDB52B4FD150031" name="Ecuador" pageId="70" pageNumber="70">Ecuador</collectingCountry>
</emphasis>
:
<emphasis id="B902EA907D5ED121FD535293FCDE0031" box="[741,796,505,523]" italics="true" pageId="70" pageNumber="70">
<collectingRegion id="49B2F8607D5ED121FD535293FCDE0031" box="[741,796,505,523]" country="Ecuador" name="Napo" pageId="70" pageNumber="70">Napo</collectingRegion>
</emphasis>
,
<collectorName id="268353547D5ED121FC9F5292FC3F0036" box="[809,1021,504,524]" pageId="70" pageNumber="70">Campana Cocha, E.</collectorName>
pto.
<location id="8EA960597D5ED121FB855292FB670031" LSID="urn:lsid:plazi:treatment:03DF87947D5ED12EFD5253BEFCA90798:8EA960597D5ED121FB855292FB670031" box="[1075,1189,504,523]" country="Ecuador" name="Misahualli" pageId="70" pageNumber="70" stateProvince="Napo">Misahualli</location>
,
<date id="FFC810427D5ED121FD035179FCDE001C" box="[693,796,531,550]" pageId="70" pageNumber="70" value="1982-06-02">
<collectingDate id="EF8CE9AA7D5ED121FD035179FCDE001C" box="[693,796,531,550]" pageId="70" pageNumber="70" value="1982-06-02">2.VI.1982</collectingDate>
</date>
,
<collectorName id="268353547D5ED121FC9E5179FC49001C" box="[808,907,531,550]" pageId="70" pageNumber="70">H.Frania</collectorName>
,
<quantity id="4C8E9B677D5ED121FC215179FC2A001C" box="[919,1000,531,550]" metricMagnitude="2" metricUnit="m" metricValue="3.3528" pageId="70" pageNumber="70" unit="ft" value="1100.0">
<elevation id="005BD1B17D5ED121FC215179FC2A001C" box="[919,1000,531,550]" metricMagnitude="2" metricUnit="m" metricValue="3.3528" pageId="70" pageNumber="70" unit="ft" value="1100.0">1100 ft.</elevation>
</quantity>
, palm leaf litter (
<specimenCount id="9D70FD0B7D5ED121FB285179FD2B007B" pageId="70" pageNumber="70" type="male">1 male</specimenCount>
,
<collectionCode id="ED67AE477D5ED121FD405144FCF3007B" box="[758,817,558,577]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="70" pageNumber="70">CNC</collectionCode>
). (See Localities below.)
</materialsCitation>
</paragraph>
<paragraph id="8BC936827D5ED121FD2D5120FB980123" blockId="70.[667,1194,288,793]" pageId="70" pageNumber="70">
TYPE LOCALITY:
<emphasis id="B902EA907D5ED121FCFF5120FC5C0067" bold="true" box="[841,926,586,605]" pageId="70" pageNumber="70">
<collectingCountry id="F36176127D5ED121FCFF5120FC5C0067" box="[841,926,586,605]" name="Ecuador" pageId="70" pageNumber="70">Ecuador</collectingCountry>
</emphasis>
:
<emphasis id="B902EA907D5ED121FC1D5120FC3E0066" box="[939,1020,586,604]" italics="true" pageId="70" pageNumber="70">
<collectingRegion id="49B2F8607D5ED121FC1D5120FC3E0066" box="[939,1020,586,604]" country="Ecuador" name="Pastaza" pageId="70" pageNumber="70">Pastaza</collectingRegion>
</emphasis>
: Ashuara indian village on Rio Macuma near Rio Morona. (This locality was not found and neither river seems to be in
<collectingRegion id="49B2F8607D5ED121FD4F51F1FC720097" box="[761,944,666,686]" country="Ecuador" name="Pastaza" pageId="70" pageNumber="70">Pastaza province</collectingRegion>
. Using the 1:1,000,000 maps for Ecuador from the American Geographical Society for Hispanic America both rivers were found in
<collectingRegion id="49B2F8607D5ED121FC3C5186FB6700C4" box="[906,1189,747,767]" country="Ecuador" name="Morona-Santiago" pageId="70" pageNumber="70">Morona Santiago province</collectingRegion>
, but do not seem to be near each other.)
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D5ED121FD0E505BFB6B0435" pageId="70" pageNumber="70" type="diagnosis">
<paragraph id="8BC936827D5ED121FD0E505BFB5F0185" blockId="70.[667,1194,817,1759]" pageId="70" pageNumber="70">
DIAGNOSIS: The unique form of sternum VIII and its long, sinuate, subapical combs (fig. 114) and the form of the aedeagus (fig. 113) will separate the males of
<taxonomicName id="4C764D017D5ED121FB3A50E6FD3F0185" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FB3A50E6FD3F0185" italics="true" pageId="70" pageNumber="70">O. distortus</emphasis>
</taxonomicName>
from all others in the New World.
</paragraph>
<paragraph id="8BC936827D5ED121FD0E50ADFB6B0435" blockId="70.[667,1194,817,1759]" pageId="70" pageNumber="70">
Among the New World females of
<taxonomicName id="4C764D017D5ED121FBDC50ACFD1C01C1" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA907D5ED121FBDC50ACFD1C01C1" italics="true" pageId="70" pageNumber="70">Oedichirus</emphasis>
</taxonomicName>
, the anteroventral angles of tergum IX are fused in
<emphasis id="B902EA907D5ED121FCF25769FCB6060C" italics="true" pageId="70" pageNumber="70">
<taxonomicName id="4C764D017D5ED121FCF25769FC130623" box="[836,977,1026,1049]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="dilophus">O. dilophus</taxonomicName>
,
<taxonomicName id="4C764D017D5ED121FC505769FBBA0623" box="[998,1144,1026,1049]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">O. distortus</taxonomicName>
,
<taxonomicName id="4C764D017D5ED121FB3A5769FD32060C" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="lunatus">O. lunatus</taxonomicName>
,
<taxonomicName id="4C764D017D5ED121FD48574AFCB2060C" box="[766,880,1055,1078]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="ohausi">O. ohausi</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C764D017D5ED121FC03574AFBF9060F" box="[949,1083,1056,1077]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B902EA907D5ED121FC03574AFBF9060F" box="[949,1083,1056,1077]" italics="true" pageId="70" pageNumber="70">O. procerus</emphasis>
</taxonomicName>
. Tergum VIII of only
<taxonomicName id="4C764D017D5ED121FC815754FC7D066E" box="[823,959,1085,1108]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FC815754FC7D066E" box="[823,959,1085,1108]" italics="true" pageId="70" pageNumber="70">O. distortus</emphasis>
</taxonomicName>
has a median lobe, although small, on the posterior margin and the legs of
<taxonomicName id="4C764D017D5ED121FCAA5713FC6106B5" box="[796,931,1144,1167]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FCAA5713FC6106B5" box="[796,931,1144,1167]" italics="true" pageId="70" pageNumber="70">O. distortus</emphasis>
</taxonomicName>
are bicolored, but are unicolorous for the other four species. The median gonocoxal plate is present anteriad of the vulvar plate for
<taxonomicName id="4C764D017D5ED121FC6657BBFBA206DD" box="[976,1120,1232,1255]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FC6657BBFBA206DD" box="[976,1120,1232,1255]" italics="true" pageId="70" pageNumber="70">O. distortus</emphasis>
</taxonomicName>
(figs. 116, 118),
<taxonomicName id="4C764D017D5ED121FC935785FC60073F" box="[805,930,1262,1285]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B902EA907D5ED121FC935785FC60073F" box="[805,930,1262,1285]" italics="true" pageId="70" pageNumber="70">O. lunatus</emphasis>
</taxonomicName>
(fig. 171),
<taxonomicName id="4C764D017D5ED121FB825785FB68073F" box="[1076,1194,1262,1285]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="ohausi">
<emphasis id="B902EA907D5ED121FB825785FB68073F" box="[1076,1194,1262,1285]" italics="true" pageId="70" pageNumber="70">O. ohausi</emphasis>
</taxonomicName>
(fig. 186), and
<taxonomicName id="4C764D017D5ED121FCEB5666FC2E0718" box="[861,1004,1292,1314]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B902EA907D5ED121FCEB5666FC2E0718" box="[861,1004,1292,1314]" italics="true" pageId="70" pageNumber="70">O. procerus</emphasis>
</taxonomicName>
(fig. 197), but absent for
<taxonomicName id="4C764D017D5ED121FC9B5640FC7B077A" box="[813,953,1321,1344]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="dilophus">
<emphasis id="B902EA907D5ED121FC9B5640FC7B077A" box="[813,953,1321,1344]" italics="true" pageId="70" pageNumber="70">O. dilophus</emphasis>
</taxonomicName>
(fig. 108). For
<taxonomicName id="4C764D017D5ED121FB3A5640FD260764" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="ohausi">
<emphasis id="B902EA907D5ED121FB3A5640FD260764" italics="true" pageId="70" pageNumber="70">O. ohausi</emphasis>
</taxonomicName>
the median gonocoxal plate anteriad of the vulvar plate is fused to the transverse anteroventral strap of tergum IX (figs. 186, 187), but both sclerites are separate for
<taxonomicName id="4C764D017D5ED121FB3A56CAFD3F07EE" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="70" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5ED121FB3A56CAFD3F07EE" italics="true" pageId="70" pageNumber="70">O. distortus</emphasis>
</taxonomicName>
(figs. 116, 118). The adornment of the vulvar plate differs for each of the five species (cf. fig. 118 to figs. 110, 171, 187, 198).
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D5ED12EFD0E557FFB680099" lastPageId="73" lastPageNumber="73" pageId="70" pageNumber="70" type="description">
<paragraph id="8BC936827D5ED121FD0E557FFBB20499" blockId="70.[667,1194,817,1759]" pageId="70" pageNumber="70">
DESCRIPTION: Length:
<quantity id="4C8E9B677D5ED121FC48557DFB670416" box="[1022,1189,1559,1580]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.8" metricValueMin="8.2" pageId="70" pageNumber="70" unit="mm" value="9.0" valueMax="9.8" valueMin="8.2">8.29.8 mm</quantity>
. Length of head:
<quantity id="4C8E9B677D5ED121FCE9555FFC290470" box="[863,1003,1589,1610]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="70" pageNumber="70" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.91.1 mm</quantity>
. Width of head: 1.0
<quantity id="4C8E9B677D5ED121FD7B5538FCEC0452" box="[717,814,1618,1640]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="70" pageNumber="70" unit="mm" value="1.1">1.1 mm</quantity>
. Pronotal length:
<quantity id="4C8E9B677D5ED121FBA55538FB670452" box="[1043,1189,1618,1640]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="70" pageNumber="70" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
. Pronotal width:
<quantity id="4C8E9B677D5ED121FCD7551AFC3204BF" box="[865,1008,1648,1669]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="70" pageNumber="70" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
. Elytral length:
<quantity id="4C8E9B677D5ED121FD2D55E7FCEA0499" box="[667,808,1677,1699]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="70" pageNumber="70" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
. Elytral width:
<quantity id="4C8E9B677D5ED121FC6955E7FBAE0499" box="[991,1132,1677,1699]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="70" pageNumber="70" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
.
</paragraph>
<paragraph id="8BC936827D5ED120FD0E55C1FC520499" blockId="70.[667,1194,817,1759]" lastBlockId="71.[701,1228,1647,1759]" lastPageId="71" lastPageNumber="71" pageId="70" pageNumber="70">Body bicolored. Head and pronotum black; elytra reddish brown; abdominal segments III to V reddish brown with darker infusions basally; segments VI to VIII dark reddish brown with paler reddish-brown infusions. Legs bicolored, pale reddish brown to yellowish brown with dark reddish brown femorotibial spot.</paragraph>
<caption id="DF09660A7D5FD120FF205693FF21047B" ID-DOI="http://doi.org/10.5281/zenodo.5401308" ID-Zenodo-Dep="5401308" httpUri="https://zenodo.org/record/5401308/files/figure.png" pageId="71" pageNumber="71" startId="71.[150,198,1529,1548]" targetBox="[131,1224,212,1499]" targetPageId="71">
<paragraph id="8BC936827D5FD120FF205693FF21047B" blockId="71.[128,1228,1528,1601]" pageId="71" pageNumber="71">
Figs. 112117.
<taxonomicName id="4C764D017D5FD120FEF35692FDCD0436" box="[325,527,1528,1548]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="71" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D5FD120FEF35692FDCD0436" box="[325,527,1528,1548]" italics="true" pageId="71" pageNumber="71">Oedichirus distortus</emphasis>
</taxonomicName>
.
<emphasis id="B902EA907D5FD120FDAA5693FD8A0436" bold="true" box="[540,584,1529,1548]" pageId="71" pageNumber="71">112.</emphasis>
Aedeagus, right lateral.
<emphasis id="B902EA907D5FD120FCFE5693FCB10436" bold="true" box="[840,883,1529,1548]" pageId="71" pageNumber="71">113.</emphasis>
Aedeagus, ventral.
<emphasis id="B902EA907D5FD120FBF75693FBAE0436" bold="true" box="[1089,1132,1529,1548]" pageId="71" pageNumber="71">114.</emphasis>
Sternum VIII, male.
<emphasis id="B902EA907D5FD120FEB4557EFEEF041D" bold="true" box="[258,301,1556,1575]" pageId="71" pageNumber="71">115.</emphasis>
Sternum VIII, setae removed, male.
<emphasis id="B902EA907D5FD120FD70557EFD30041D" bold="true" box="[710,754,1556,1575]" pageId="71" pageNumber="71">116.</emphasis>
Segment IX, ventral, female.
<emphasis id="B902EA907D5FD120FB8B557EFBAA041D" bold="true" box="[1085,1128,1556,1575]" pageId="71" pageNumber="71">117.</emphasis>
Sternum IX, male.
</paragraph>
</caption>
<paragraph id="8BC936827D5FD12FFD6D55C1FED4078E" blockId="71.[701,1228,1647,1759]" lastBlockId="72.[93,619,1259,1759]" lastPageId="72" lastPageNumber="72" pageId="71" pageNumber="71">Head with length and width equal to or about a fifth wider than long (HW/HL: 1.01.2). Frontoclypeal ridge complete, not separated medially. Dorsal surface without V-shaped depression; surface polished and coarsely punctate; punctation dense, less dense basally and laterally. Labrum quadridentate; surface without tubercle near submedial denticle.</paragraph>
<caption id="DF09660A7D50D12FFF4A57C4FBE006FB" ID-DOI="http://doi.org/10.5281/zenodo.5401310" ID-Zenodo-Dep="5401310" box="[252,1058,1198,1218]" httpUri="https://zenodo.org/record/5401310/files/figure.png" pageId="72" pageNumber="72" startId="72.[252,290,1198,1217]" targetBox="[96,1189,211,1167]" targetPageId="72">
<paragraph id="8BC936827D50D12FFF4A57C4FBE006FB" blockId="72.[252,1058,1198,1218]" box="[252,1058,1198,1218]" pageId="72" pageNumber="72">
Fig. 118.
<taxonomicName id="4C764D017D50D12FFEC757C4FDFE06F8" box="[369,572,1198,1218]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="72" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D50D12FFEC757C4FDFE06F8" box="[369,572,1198,1218]" italics="true" pageId="72" pageNumber="72">Oedichirus distortus</emphasis>
</taxonomicName>
. Segment IX, vulvar plate, enlarged (CLSM).
</paragraph>
</caption>
<paragraph id="8BC936827D50D12FFFCC56D7FEF30498" blockId="72.[93,619,1259,1759]" pageId="72" pageNumber="72">Pronotum about a fifth longer than wide (PL/PW: 1.2). Pronotum polished, with scattered clusters of moderately dense to dense, coarse punctation; surface with moderately deep, submedial, punctate groove on basal two thirds. Elytra length about a tenth less than width (EW/EL: 1.1); surface flat and coarsely punctate.</paragraph>
<paragraph id="8BC936827D50D12FFFCC55C1FCC104E7" blockId="72.[93,619,1259,1759]" lastBlockId="72.[667,1194,1259,1757]" pageId="72" pageNumber="72">Abdominal segments III to VI coarsely punctate, VII and VIII less coarsely punctate; III to VI with transverse, subapical row of irregularly arranged punctures. Segment III without paratergite; paratergal carina extending to near middle of segment, but poorly developed beyond spiracle. Tergum III without median point extending from transverse basal ridge. Tergum VIII with small, median, trianguloid lobe on posterior margin; posterior margin broadly emarginate; transverse basal ridge irregularly sinuate and without median point. Tergum IX with lateroapical process about twice as long as midbasal length (LLaP/L9 5 1.92.0), slightly to moderately bent ventrally, and approximately parallel to other process; ventromedial margin without posteriorly directed spur (cf. fig. 158).</paragraph>
<paragraph id="8BC936827D51D12EFF2A53B9FE600624" blockId="73.[128,654,211,1759]" pageId="73" pageNumber="73">MALE: Sterna VI and VII unmodified. Sternum VIII (figs. 114, 115) with moderately deep, asymmetrical emargination; emargination about one sixth of length of sternum, wider than deep, with wide and broadly rounded base, with right margin longer and steeper than left, and with basal margin and part of lateral margins membranous; surface adjacent to emargination with transverse comb of long, spiniform setae (fig. 114); comb curved anteriorly on left side and posteriorly on right; surface of sternum with anteriorly curved ridge at base of comb on left side and with posteriorly curved ridge at base of comb on right side (fig. 115); surface tumescent at proximal base of both left and right curves of transverse ridge (fig. 115); transverse basal ridge broadly sinuate, separated medially, and with weak, apically open, median point (fig. 114). Tergum IX with small, lobed process on anterior margin of anteroventral angle. Sternum IX strongly asymmetrial (fig. 117); anterior margin wide and broadly emarginate; posterior margin narrow, rounded, and bluntly pointed; lateral margin tapered apically to pointed apex; right margin broadly rounded on basal third, then strongly sinuate to apex, left margin strongly sinuate to apex.</paragraph>
<paragraph id="8BC936827D51D12EFF2B574DFEB3042B" blockId="73.[128,654,211,1759]" pageId="73" pageNumber="73">Aedeagus asymmetrical (figs. 112, 113). Ventral sclerite with short, broad, apically acute lobe on right side of apex and long, basally broad, tapering, apically acute lobe on right side of apex (fig. 113); left lobe much longer than right (fig. 113), with diagonally transverse, membranous depression extending from left basal margin of left lobe to near right lateral margin (fig. 113), with robust, small tumescence near base of right lobe at right lateral margin, with small, tapered, apically rounded lobe on left side just distad of middle (fig. 113), and without apicoventral process (cf. fig. 99) extending from posterior margin. Parameres short, slender, tapered, sinuate, and free of median lobe except at base (figs. 112, 113).</paragraph>
<paragraph id="8BC936827D51D12EFF2A557DFB680099" blockId="73.[128,654,211,1759]" lastBlockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">FEMALE: Sternum VIII with transverse basal ridge broadly sinuate and curved anteriorly at middle and without median point; posterior margin with wide, slight, microserrate, median emargination. Tergum IX with anteroventral angles fused medially to form wide strap (figs. 116, 118); strap with shallow groove at point of fusion. Median gonocoxal plate anteriad of vulvar plate reduced to narrow strap (figs. 116, 118) with anterior margin broadly rounded with small median lobe; gonocoxal plate posteriad of vulvar plate gradually tapered to irregularly microsinuate posterior margin (fig. 116). Vulvar plate embedded midanteriorly in gonocoxal plate (fig. 116). Anterior vulvar lobe transverse and extending across anterior margin of posterior vulvar lobe; surface slightly wrinkled (fig. 118). Posterior vulvar lobe transverse, larger than anterior lobe; surface with ornamentation near right lateral side. Vulva crescent shaped and orientated at slight angle to longitudinal axis (fig. 118).</paragraph>
</subSubSection>
<subSubSection id="C36C65097D51D12EFD6C51C3FCB401DD" pageId="73" pageNumber="73" type="etymology">
<paragraph id="8BC936827D51D12EFD6C51C3FBAE0123" blockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">
ETYMOLOGY: The name is from the Latin
<emphasis id="B902EA907D51D12EFD0B51ADFCE400E4" box="[701,806,711,734]" italics="true" pageId="73" pageNumber="73">
<taxonomicName id="4C764D017D51D12EFD0B51ADFCE000E4" box="[701,802,711,734]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="73" phylum="Arthropoda" rank="species" species="distortus">distortus</taxonomicName>
,
</emphasis>
distorted or misshapened, and refers to the strongly asymmetrical posterior margin of sternum VIII of the male.
</paragraph>
<paragraph id="8BC936827D51D12EFD6C5075FCB401DD" blockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">
LOCALITIES: Puerto Misahualli,
<collectingCountry id="F36176127D51D12EFBD7504BFB05010D" box="[1121,1223,801,823]" name="Ecuador" pageId="73" pageNumber="73">Ecuador</collectingCountry>
, is at the mouth of Río Misahualli at
<quantity id="4C8E9B677D51D12EFB375055FB0E016E" box="[1153,1228,831,852]" metricMagnitude="2" metricUnit="m" metricValue="4.31" pageId="73" pageNumber="73" unit="m" value="431.0">431 m</quantity>
elevation at
<geoCoordinate id="EE4250457D51D12EFCFC5036FC67014B" box="[842,933,859,882]" degrees="01" direction="south" minutes="029" orientation="latitude" pageId="73" pageNumber="73" precision="925" value="-1.4833333">
01
<emphasis id="B902EA907D51D12EFCD35036FCB2014B" bold="true" box="[869,880,860,881]" pageId="73" pageNumber="73">°</emphasis>
029S
</geoCoordinate>
, 77
<emphasis id="B902EA907D51D12EFC7A5036FC15014B" bold="true" box="[972,983,860,881]" pageId="73" pageNumber="73">°</emphasis>
409W (
<bibRefCitation id="EFE74B737D51D12EFB925036FD2801B5" author="Paynter, R. A." pageId="73" pageNumber="73" refId="ref77340" refString="Paynter, R. A. 1993. Ornithological gazetteer of Ecuador, xi + 247 pp. Cambridge, MA: Harvard University." type="book" year="1993">Paynter, 1993: 128</bibRefCitation>
). The actual locality is Campana Cocha, east of Puerto Misahualli; that site was not found, but is presumably on the east side of Río Misahualli.
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D51D12EFD6C5087FC390619" pageId="73" pageNumber="73" type="distribution">
<paragraph id="8BC936827D51D12EFD6C5087FC390619" blockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">
DISTRIBUTION:
<collectingCountry id="F36176127D51D12EFC145085FBC4063F" box="[930,1030,1007,1029]" name="Ecuador" pageId="73" pageNumber="73">Ecuador</collectingCountry>
in
<collectingRegion id="49B2F8607D51D12EFBF1509AFB4B063F" box="[1095,1161,1008,1029]" country="Ecuador" name="Napo" pageId="73" pageNumber="73">Napo</collectingRegion>
and
<collectingRegion id="49B2F8607D51D12EFD0B5767FCD40618" box="[701,790,1037,1058]" country="Ecuador" name="Pastaza" pageId="73" pageNumber="73">Pastaza</collectingRegion>
provinces (fig. 40).
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D51D12EFD6C5743FCA90798" pageId="73" pageNumber="73" type="discussion">
<paragraph id="8BC936827D51D12EFD6C5743FB4A075C" blockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">
REMARKS: The
<specimenCount id="9D70FD0B7D51D12EFC205741FBD4067B" box="[918,1046,1067,1089]" pageId="73" pageNumber="73" type="female">two female</specimenCount>
<typeStatus id="54CD88207D51D12EFBAB5741FB4C067A" box="[1053,1166,1067,1088]" pageId="73" pageNumber="73" type="paratype">paratypes</typeStatus>
were collected together with the
<typeStatus id="54CD88207D51D12EFC435722FB990664" box="[1013,1115,1096,1118]" pageId="73" pageNumber="73" type="holotype">holotype</typeStatus>
male and all three have dense coarse pronotal punctation, lateroapical processes of tergum IX that are about twice as long as the midbasal length of the tergum, and a small, triangular, median lobe on the broadly emarginate posterior margin of tergum VIII. The triangular lobe is more poorly developed on the male
<typeStatus id="54CD88207D51D12EFCB45659FCAB0772" box="[770,873,1331,1352]" pageId="73" pageNumber="73" type="paratype">paratype</typeStatus>
from Puerto Misahualli and the pronotal punctation is more dense.
</paragraph>
<paragraph id="8BC936827D51D12EFD6D5604FCA90798" blockId="73.[701,1228,213,1442]" pageId="73" pageNumber="73">
This species is the first record of the genus from
<collectingCountry id="F36176127D51D12EFCB656E6FCA50798" box="[768,871,1420,1442]" name="Ecuador" pageId="73" pageNumber="73">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>