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<document id="78BABDA0A730E3E4BEB5C78B5AE63E0B" ID-CLB-Dataset="2947" ID-DOI="10.11646/zootaxa.5082.1.2" ID-GBIF-Dataset="722c160e-2d11-4a58-83fc-d346d227f9e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783094" ID-ZooBank="795488E3-DE16-4268-8968-628C9D5E3A4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564243538" checkinUser="plazi" docAuthor="Rueda Martín, Paola A., Gibon, Francois-Marie &amp; Sganga, Julieta V." docDate="2021" docId="03DEE90BFFA94511FF0AFE94FD85FA37" docLanguage="en" docName="zootaxa.5082.1.2.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrnellus guyanensis Rueda Martín &amp; Gibon &amp; Sganga 2021, new species" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FFE79173FFAA4512FF9DFF8FFF9CFFC9" masterDocTitle="New species and records of the Neotropical genus Cyrnellus (Trichoptera: Polycentropodidae) and the re-establishment of C. minimus and C. marginalis" masterLastPageNumber="29" masterPageNumber="15" pageNumber="18" updateTime="1699442671739" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4FE19095941219943DDCA634458C9EF2">New species and records of the Neotropical genus Cyrnellus (Trichoptera: Polycentropodidae) and the re-establishment of C. minimus and C. marginalis</mods:title>
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<mods:name id="F821E55C48852DADDFEEBC4D60E3C117" type="personal">
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<mods:namePart id="4FCC19143F4B46DEFE8184F20ED1967B">Rueda Martín, Paola A.</mods:namePart>
<mods:affiliation id="BD7006CC6B72F4459E61E6317C0A9487">Instituto de Biodiversidad Neotropical, CONICET Facultad de Ciencias Naturales e Instituto Miguel Lillo, Miguel Lillo 205, San Miguel de Tucumán, Tucumán, Argentina</mods:affiliation>
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<mods:namePart id="E841C516897BA4190D5E4C1F6C1607E7">Gibon, Francois-Marie</mods:namePart>
<mods:affiliation id="7BC8FB046CFFBEEDD2291B5269F02052">CBGP, IRD, CIRAD, INRAE, Montpellier SupAgro, Univ. Montpellier, Montpellier, France.</mods:affiliation>
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<mods:namePart id="B16602CB796625F082AE35986EA71B6D">Sganga, Julieta V.</mods:namePart>
<mods:nameIdentifier id="5E37A84A670603DE00B7D0566D8B1F6E" type="ORCID">0000-0003-1210-5937</mods:nameIdentifier>
<mods:affiliation id="5E5EB8E79CAAC6573D3BB4189724E299">Departamento de Biodiversidad y Biología Experimental, Facultad de Ciencias Exactas y Naturales (UBA). CABA, Argentina. jsganga @ gmail. com; https: // orcid. org / 0000 - 0003 - 1210 - 5937</mods:affiliation>
<mods:nameIdentifier id="12EAEBB56DCAA2AC6AFCC5001B6D163E" type="email">jsganga@gmail.com</mods:nameIdentifier>
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<mods:title id="EE1085C49C6F0603B163627961A60A02">Zootaxa</mods:title>
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<mods:date id="4CBB007DE1A7F19E08E5D165381204A3">2021</mods:date>
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<mods:number id="3E599ED541917B9CBD2786FCBB85F862">2021-12-15</mods:number>
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<treatment id="03DEE90BFFA94511FF0AFE94FD85FA37" ID-DOI="http://doi.org/10.5281/zenodo.5783108" ID-GBIF-Taxon="191647283" ID-Zenodo-Dep="5783108" LSID="urn:lsid:plazi:treatment:03DEE90BFFA94511FF0AFE94FD85FA37" httpUri="http://treatment.plazi.org/id/03DEE90BFFA94511FF0AFE94FD85FA37" lastPageNumber="18" pageId="3" pageNumber="18">
<subSubSection id="C36D0B96FFA94511FF0AFE94FDAEFEFF" box="[151,562,283,310]" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="8BC8581DFFA94511FF0AFE94FDAEFEFF" blockId="3.[151,562,283,346]" box="[151,562,283,310]" pageId="3" pageNumber="18">
<heading id="D080EF71FFA94511FF0AFE94FDAEFEFF" bold="true" box="[151,562,283,310]" fontSize="11" level="1" pageId="3" pageNumber="18" reason="1">
<emphasis id="B903840FFFA94511FF0AFE94FDAEFEFF" bold="true" box="[151,562,283,310]" pageId="3" pageNumber="18">
<taxonomicName id="4C77239EFFA94511FF0AFE94FE07FEFF" authority="Rueda Martín &amp; Gibon &amp; Sganga, 2021" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" box="[151,411,283,310]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis" status="sp. nov.">
<emphasis id="B903840FFFA94511FF0AFE94FE07FEFF" bold="true" box="[151,411,283,310]" italics="true" pageId="3" pageNumber="18">Cyrnellus guyanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2303974FFA94511FE3EFE93FDAEFEFF" box="[419,562,284,310]" pageId="3" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36D0B96FFA94511FF0AFECFFEBDFE93" box="[151,289,320,346]" pageId="3" pageNumber="18" type="description">
<paragraph id="8BC8581DFFA94511FF0AFECFFEBDFE93" blockId="3.[151,562,283,346]" box="[151,289,320,346]" pageId="3" pageNumber="18">
<figureCitation id="134C4498FFA94511FF0AFECFFEBDFE93" box="[151,289,320,346]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Figs 2A–2D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF0AFE07FEF9FCC2" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="8BC8581DFFA94511FF0AFE07FEF9FCC2" blockId="3.[151,1437,392,1534]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF0AFE07FE96FE6B" bold="true" box="[151,266,392,418]" pageId="3" pageNumber="18">Diagnosis</emphasis>
.
<taxonomicName id="4C77239EFFA94511FE8AFE07FD9AFE6B" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" box="[279,518,392,418]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="B903840FFFA94511FE8AFE07FD9AFE6B" box="[279,518,392,418]" italics="true" pageId="3" pageNumber="18">Cyrnellus guyanensis</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="4C77239EFFA94511FCEDFE07FBA8FE6B" box="[880,1076,392,418]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B903840FFFA94511FCEDFE07FC41FE6B" box="[880,989,392,418]" italics="true" pageId="3" pageNumber="18">Cyrnellus</emphasis>
species
</taxonomicName>
by the following character combination: The apex of each inferior appendage is as wide as the median part (
<figureCitation id="134C4498FFA94511FC67FE23FBCFFE0F" box="[1018,1107,428,454]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FBC2FE22FBE9FE0F" box="[1119,1141,429,454]" italics="true" pageId="3" pageNumber="18">ia</emphasis>
), with two subapicomesal spines dorsoventrally flat (depressed,
<figureCitation id="134C4498FFA94511FDA3FE5FFD09FE23" box="[574,661,464,490]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2B</figureCitation>
,
<emphasis id="B903840FFFA94511FD3DFE5EFD5EFE23" box="[672,706,465,490]" italics="true" pageId="3" pageNumber="18">asI</emphasis>
,
<emphasis id="B903840FFFA94511FD4DFE5EFD60FE23" box="[720,764,465,490]" italics="true" pageId="3" pageNumber="18">asII</emphasis>
), and lacking an internal lobe; the anterior arm of the dorsal phallic sclerite is thick (
<figureCitation id="134C4498FFA94511FE02FE7BFDB6FDC7" box="[415,554,500,526]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Figs 2C, 2D</figureCitation>
,
<emphasis id="B903840FFFA94511FDA9FE7BFDD1FDC7" box="[564,589,500,526]" italics="true" pageId="3" pageNumber="18">ds</emphasis>
), the posterior arms are divergent, short and slender (
<figureCitation id="134C4498FFA94511FB00FE7BFB69FDC7" box="[1181,1269,500,526]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2D</figureCitation>
,
<emphasis id="B903840FFFA94511FA9DFE7BFA85FDC7" box="[1280,1305,500,526]" italics="true" pageId="3" pageNumber="18">ds</emphasis>
).
<taxonomicName id="4C77239EFFA94511FAB2FE7BFE8FFDFB" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="B903840FFFA94511FAB2FE7BFE8FFDFB" italics="true" pageId="3" pageNumber="18">Cyrnellus guyanensis</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C77239EFFA94511FE79FD97FD56FDFB" authority="Flint 1971" authorityName="Flint" authorityYear="1971" box="[484,714,536,562]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FE79FD97FDD2FDFB" box="[484,590,536,562]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
<bibRefCitation id="EFE625ECFFA94511FDCBFD97FD56FDFB" author="Flint, O. S. Jr." box="[598,714,536,562]" pageId="3" pageNumber="18" pagination="1 - 67" refId="ref8968" refString="Flint, O. S. Jr. (1971) Studies of Neotropical caddisflies, XII: Rhyacophilidae, Glossosomatidae, Philopotamidae and Psychomyiidae from the Amazon Basin (Trichoptera). Amazoniana, 3, 1 - 67." type="journal article" year="1971">Flint 1971</bibRefCitation>
</taxonomicName>
. The difference in size is important, the length of each forewing is
<quantity id="4C8FF5F8FFA94511FF2CFDB3FF6FFD9E" box="[177,243,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="18" unit="mm" value="5.0">5 mm</quantity>
in
<taxonomicName id="4C77239EFFA94511FE8AFDB3FE1DFD9F" authorityName="Flint" authorityYear="1971" box="[279,385,572,598]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FE8AFDB3FE1DFD9F" box="[279,385,572,598]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
whereas it is 3.8 mm in
<taxonomicName id="4C77239EFFA94511FD0BFDB3FCAFFD9F" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" box="[662,819,572,598]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="B903840FFFA94511FD0BFDB3FCAFFD9F" box="[662,819,572,598]" italics="true" pageId="3" pageNumber="18">C. guyanensis</emphasis>
</taxonomicName>
. The genitalia of
<taxonomicName id="4C77239EFFA94511FC66FDB3FB04FD9F" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" box="[1019,1176,572,598]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="B903840FFFA94511FC66FDB3FB04FD9F" box="[1019,1176,572,598]" italics="true" pageId="3" pageNumber="18">C. guyanensis</emphasis>
</taxonomicName>
are distinguished from those of
<taxonomicName id="4C77239EFFA94511FF64FDEFFEF8FDB3" authorityName="Flint" authorityYear="1971" box="[249,356,608,634]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FF64FDEFFEF8FDB3" box="[249,356,608,634]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
by the apex of each inferior appendage being as wide as the median part (whereas it is apically broadened in
<taxonomicName id="4C77239EFFA94511FEAFFD0BFE00FD57" authorityName="Flint" authorityYear="1971" box="[306,412,644,670]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FEAFFD0BFE00FD57" box="[306,412,644,670]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
); the subapicomesal spines are flat (conical in
<taxonomicName id="4C77239EFFA94511FC2FFD0BFB80FD57" authorityName="Flint" authorityYear="1971" box="[946,1052,644,670]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FC2FFD0BFB80FD57" box="[946,1052,644,670]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
); an internal lobe of each inferior appendage is absent (present in
<taxonomicName id="4C77239EFFA94511FD99FD27FDF3FD0B" authorityName="Flint" authorityYear="1971" box="[516,623,680,706]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FD99FD27FDF3FD0B" box="[516,623,680,706]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
); the anterior arm of the dorsal phallic sclerite is reduced (absent in
<taxonomicName id="4C77239EFFA94511FA1EFD27FF7CFD2F" authorityName="Flint" authorityYear="1971" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FA1EFD27FF7CFD2F" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
); and the posterior arms of the dorsal phallic sclerite are divergent, short, and slender (fused and forming a tube in
<taxonomicName id="4C77239EFFA94511FF76FD7FFEC9FCC3" authorityName="Flint" authorityYear="1971" box="[235,341,752,778]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FF76FD7FFEC9FCC3" box="[235,341,752,778]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF5AFC9BFDEEFBBA" pageId="3" pageNumber="18" type="description">
<paragraph id="8BC8581DFFA94511FF5AFC9BFB56FCE7" blockId="3.[151,1437,392,1534]" box="[199,1226,788,815]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFC9BFECDFCE7" bold="true" box="[199,337,788,814]" pageId="3" pageNumber="18">Description</emphasis>
. Adult: Length of each male forewing 3.8 mm. General coloration light brown.
</paragraph>
<paragraph id="8BC8581DFFA94511FF5AFCB7FDEEFBBA" blockId="3.[151,1437,392,1534]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFCB7FEEEFC9B" bold="true" box="[199,370,824,850]" pageId="3" pageNumber="18">Male genitalia</emphasis>
. Anterior margins of sternite IX sinuous; posterolateral margins irregular, each with acute projection (
<figureCitation id="134C4498FFA94511FF6EFCD3FED0FCBF" box="[243,332,860,886]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FEC5FCD2FEF0FCBF" box="[344,364,861,886]" italics="true" pageId="3" pageNumber="18">ix</emphasis>
). Width of intermediate appendage constant from base to apex, with dorsal and ventral margins parallel (
<figureCitation id="134C4498FFA94511FF66FC0FFEC9FC53" box="[251,341,896,922]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FEFCFC0EFE19FC53" box="[353,389,897,922]" italics="true" pageId="3" pageNumber="18">iap</emphasis>
). Preanal appendages each with mesolateral process narrow at base (
<figureCitation id="134C4498FFA94511FB0DFC0FFB75FC53" box="[1168,1257,896,922]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FB6BFC0FFABCFC53" box="[1270,1312,896,922]" italics="true" pageId="3" pageNumber="18">mlp</emphasis>
); dorsolateral process subequal in length to mesolateral process, slightly curved, directed distad (
<figureCitation id="134C4498FFA94511FBECFC2BFB56FC77" box="[1137,1226,932,958]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FB4AFC2BFB67FC77" box="[1239,1275,932,958]" italics="true" pageId="3" pageNumber="18">dlp</emphasis>
); mesoventral process wide at base, gradually tapered to blunt apex (
<figureCitation id="134C4498FFA94511FD70FC47FCD9FC2B" box="[749,837,968,994]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FCCDFC46FCE3FC2B" box="[848,895,969,994]" italics="true" pageId="3" pageNumber="18">mvp</emphasis>
). Inferior appendages long, each with dorsal and ventral margins sinuous, parallel (
<figureCitation id="134C4498FFA94511FD8CFC63FDF6FBCF" box="[529,618,1004,1030]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2A</figureCitation>
,
<emphasis id="B903840FFFA94511FDE8FC62FD17FBCF" box="[629,651,1005,1030]" italics="true" pageId="3" pageNumber="18">ia</emphasis>
); apex slightly curved toward midline, subapicomesal lobe high, bearing two flat, triangular spines with broad bases at 2/3 length of inferior appendage (
<figureCitation id="134C4498FFA94511FBD9FB9FFB00FBE3" box="[1092,1180,1040,1066]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2B</figureCitation>
,
<emphasis id="B903840FFFA94511FB35FB9EFB58FBE3" box="[1192,1220,1041,1066]" italics="true" pageId="3" pageNumber="18">ap</emphasis>
,
<emphasis id="B903840FFFA94511FB4FFB9EFB68FBE3" box="[1234,1268,1041,1066]" italics="true" pageId="3" pageNumber="18">asI</emphasis>
,
<emphasis id="B903840FFFA94511FA9EFB9EFAB3FBE3" box="[1283,1327,1041,1066]" italics="true" pageId="3" pageNumber="18">asII</emphasis>
); internal lobe absent (
<figureCitation id="134C4498FFA94511FEBAFBBBFE1CFB87" box="[295,384,1076,1102]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2B</figureCitation>
). Phallus tubular; dorsal phallic sclerite Y-shaped, with anterior arm thick and posterior arms divergent, short, and slender (
<figureCitation id="134C4498FFA94511FE79FBD7FDA1FBBB" box="[484,573,1112,1138]" captionStart="FIGURE 2" captionStartId="4.[151,250,1832,1856]" captionTargetBox="[233,1349,185,1799]" captionTargetId="figure-19@4.[229,1358,181,1809]" captionTargetPageId="4" captionText="FIGURE 2. Cyrnellus spp., male genitalia. 2A–2E, Cyrnellus guyanensis sp. nov.: 2A, left lateral; 2B, right inferior appendage, ventral; 2C, phallus, left lateral; 2D, phallus, dorsal. 2E–2H, Cyrnellus mammillatus Flint 1971: 2E, left lateral; 2F, right inferior appendage, ventral; 2G, phallus, left lateral; 2H, dorsal phallic sclerite, dorsal. ap = apex of inferior an appendage; asI = subapicomesal spine I; asII = subapicomesal spine II; dlp = dorsolateral process; ds = dorsal phallic sclerite; ia = inferior appendage; iap = intermediate appendage; ix = sternite IX; mlp = mesolateral process; mvp = mesoventral process." figureDoi="http://doi.org/10.5281/zenodo.5783102" httpUri="https://zenodo.org/record/5783102/files/figure.png" pageId="3" pageNumber="18">Fig. 2D</figureCitation>
,
<emphasis id="B903840FFFA94511FDD4FBD7FDFEFBBB" box="[585,610,1112,1138]" italics="true" pageId="3" pageNumber="18">ds</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF5AFBF3FD83FACB" pageId="3" pageNumber="18" type="discussion">
<paragraph id="8BC8581DFFA94511FF5AFBF3FD83FACB" blockId="3.[151,1437,392,1534]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFBF3FED9FB5F" bold="true" box="[199,325,1148,1174]" pageId="3" pageNumber="18">Discussion</emphasis>
. This species was discovered among specimens of the USNM identified as
<taxonomicName id="4C77239EFFA94511FB0DFBF3FAD0FB5F" authorityName="Flint" authorityYear="1971" box="[1168,1356,1148,1174]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FB0DFBF3FAD0FB5F" box="[1168,1356,1148,1174]" italics="true" pageId="3" pageNumber="18">Cyrnellus ulmeri</emphasis>
</taxonomicName>
. These two species are very similar. Both have two subapicomesal spines I (
<emphasis id="B903840FFFA94511FC33FB2EFC4CFB73" box="[942,976,1185,1210]" italics="true" pageId="3" pageNumber="18">asI</emphasis>
), inserted at mid-length of each inferior appendage. Even though the genitalia of
<taxonomicName id="4C77239EFFA94511FDFBFB4BFC9FFB17" authority="Rueda Martín &amp; Gibon &amp; Sganga, 2021" authorityName="Rueda Martín &amp; Gibon &amp; Sganga" authorityYear="2021" box="[614,771,1220,1246]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="guyanensis" status="sp. nov.">
<emphasis id="B903840FFFA94511FDFBFB4BFC9FFB17" box="[614,771,1220,1246]" italics="true" pageId="3" pageNumber="18">C. guyanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2303974FFA94511FC91FB4AFCD5FB16" box="[780,841,1221,1247]" pageId="3" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C77239EFFA94511FC1DFB4BFC77FB17" authorityName="Flint" authorityYear="1971" box="[896,1003,1220,1246]" class="Insecta" family="Polycentropodidae" genus="Cyrnellus" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B903840FFFA94511FC1DFB4BFC77FB17" box="[896,1003,1220,1246]" italics="true" pageId="3" pageNumber="18">C. ulmeri</emphasis>
</taxonomicName>
are similar, some constant differences clearly distinguish the two species.
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF5AFA83FC09FA7F" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BC8581DFFA94511FF5AFA83FE0BFAA7" blockId="3.[151,1437,392,1534]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFA83FE2AFAEF" bold="true" box="[199,438,1292,1318]" pageId="3" pageNumber="18">Material examined.</emphasis>
<materialsCitation id="3B1F5240FFA94511FE5EFA83FE0BFAA7" ID-GBIF-Occurrence="3418589316" collectingDate="1983-12-17" collectorName="Earthwatch Research Expedition, P. J &amp; W. E. Steiner" country="Guyana" county="Takutu Mountains" latitude="6.25" location="Male" longLatPrecision="1305" longitude="-59.083332" municipality="Potaro District" pageId="3" pageNumber="18" specimenCount="1" typeStatus="holotype">
<emphasis id="B903840FFFA94511FE5EFA83FDA5FAEF" bold="true" box="[451,569,1292,1318]" pageId="3" pageNumber="18">
<typeStatus id="54CCE6BFFFA94511FE5EFA83FDA8FAEF" box="[451,564,1292,1318]" pageId="3" pageNumber="18" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="8EA80EC6FFA94511FDD8FA83FD1FFAEF" LSID="urn:lsid:plazi:treatment:03DEE90BFFA94511FF0AFE94FD85FA37:8EA80EC6FFA94511FDD8FA83FD1FFAEF" box="[581,643,1292,1318]" country="Guyana" county="Takutu Mountains" latitude="6.25" longLatPrecision="1305" longitude="-59.083332" municipality="Potaro District" name="Male" pageId="3" pageNumber="18">Male</location>
:
<emphasis id="B903840FFFA94511FD0EFA83FC89FAEF" bold="true" box="[659,789,1292,1318]" pageId="3" pageNumber="18">
<collectingCountry id="F360188DFFA94511FD0EFA83FC8CFAEF" box="[659,784,1292,1318]" name="Guyana" pageId="3" pageNumber="18">GUYANA</collectingCountry>
:
</emphasis>
<location id="8EA80EC6FFA94511FCBFFA83FC0CFAEF" LSID="urn:lsid:plazi:treatment:03DEE90BFFA94511FF0AFE94FD85FA37:8EA80EC6FFA94511FCBFFA83FC0CFAEF" box="[802,912,1292,1318]" country="Guyana" county="Takutu Mountains" latitude="6.25" longLatPrecision="1305" longitude="-59.083332" municipality="Potaro District" name="Mazaruni" pageId="3" pageNumber="18">Mazaruni</location>
,
<collectingMunicipality id="6BACC267FFA94511FC3DFA82FBD1FAEF" box="[928,1101,1292,1319]" pageId="3" pageNumber="18">Potaro District</collectingMunicipality>
,
<collectingCounty id="62A92091FFA94511FBC0FA83FAAEFAEF" box="[1117,1330,1292,1318]" pageId="3" pageNumber="18">Takutu Mountains</collectingCounty>
,
<geoCoordinate id="EE433EDAFFA94511FADFFA83FA0BFAEE" box="[1346,1431,1292,1319]" degrees="6" direction="north" minutes="15" orientation="latitude" pageId="3" pageNumber="18" precision="925" value="6.25">6°15´N</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFA94511FF0AFABFFF6CFA82" box="[151,240,1328,1355]" degrees="59" direction="west" minutes="5" orientation="longitude" pageId="3" pageNumber="18" precision="925" value="-59.083332">59°5´W</geoCoordinate>
,
<date id="FFC97EDDFFA94511FF66FABFFEE1FA83" box="[251,381,1328,1354]" pageId="3" pageNumber="18" value="1983-12-17">
<collectingDate id="EF8D8735FFA94511FF66FABFFEE1FA83" box="[251,381,1328,1354]" pageId="3" pageNumber="18" value="1983-12-17">17.xii.1983</collectingDate>
</date>
, at blacklight in forest clearing near streams,
<collectorName id="26823DCBFFA94511FCE8FABFFA8EFA82" box="[885,1298,1328,1355]" pageId="3" pageNumber="18">Earthwatch Research Expedition, P.J</collectorName>
. Spangler &amp;
<collectorName id="26823DCBFFA94511FF0AFADAFEB8FAA7" box="[151,292,1364,1391]" pageId="3" pageNumber="18">W.E. Steiner</collectorName>
(USNM).
</materialsCitation>
</paragraph>
<paragraph id="8BC8581DFFA94511FF5AFAF7FC09FA7F" blockId="3.[151,1437,392,1534]" pageId="3" pageNumber="18">
<materialsCitation id="3B1F5240FFA94511FF5AFAF7FD5EFA7F" ID-GBIF-Occurrence="3418589331" collectingDate="1986-10-17" collectingDateMax="1986-10-18" collectingDateMin="1986-10-17" collectorName="P. Spangler &amp; Flint" country="Brazil" county="Takutu Mountains" latitude="-3.8333333" location="Para" longLatPrecision="1307" longitude="-52.666668" municipality="Rio Oriri Camp." pageId="3" pageNumber="18" specimenCount="9" specimenCount-male="9" typeStatus="paratype">
<emphasis id="B903840FFFA94511FF5AFAF7FED4FA5B" bold="true" box="[199,328,1400,1426]" pageId="3" pageNumber="18">
<typeStatus id="54CCE6BFFFA94511FF5AFAF7FEDFFA5B" box="[199,323,1400,1426]" pageId="3" pageNumber="18" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D719394FFA94511FECDFAF7FE35FA5B" box="[336,425,1400,1426]" pageId="3" pageNumber="18" type="male">9 males</specimenCount>
, same data as holotype.
<emphasis id="B903840FFFA94511FD26FAF7FCB3FA5B" bold="true" box="[699,815,1400,1426]" pageId="3" pageNumber="18">
<collectingCountry id="F360188DFFA94511FD26FAF7FCB7FA5B" box="[699,811,1400,1426]" name="Brazil" pageId="3" pageNumber="18">BRAZIL</collectingCountry>
:
</emphasis>
<location id="8EA80EC6FFA94511FCAAFAF6FCF1FA5A" LSID="urn:lsid:plazi:treatment:03DEE90BFFA94511FF0AFE94FD85FA37:8EA80EC6FFA94511FCAAFAF6FCF1FA5A" box="[823,877,1401,1427]" country="Brazil" latitude="-3.8333333" longLatPrecision="1307" longitude="-52.666668" municipality="Rio Oriri Camp." name="Para" pageId="3" pageNumber="18">Para</location>
:
<collectingMunicipality id="6BACC267FFA94511FCE5FAF7FBAEFA5B" box="[888,1074,1400,1426]" pageId="3" pageNumber="18">Rio Oriri Camp.</collectingMunicipality>
,
<geoCoordinate id="EE433EDAFFA94511FBA3FAF7FB39FA5A" box="[1086,1189,1400,1427]" degrees="52" direction="west" minutes="40" orientation="longitude" pageId="3" pageNumber="18" precision="925" value="-52.666668">52°40´W</geoCoordinate>
,
<geoCoordinate id="EE433EDAFFA94511FB2DFAF7FA9DFA5A" box="[1200,1281,1400,1427]" degrees="3" direction="south" minutes="50" orientation="latitude" pageId="3" pageNumber="18" precision="925" value="-3.8333333">3°50´S</geoCoordinate>
, Altamira, Ca 1000Km,
<date id="FFC97EDDFFA94511FE9AFA13FE38FA7F" box="[263,420,1436,1462]" pageId="3" pageNumber="18" value="1986-10-17" valueMax="1986-10-18" valueMin="1986-10-17">
<collectingDate id="EF8D8735FFA94511FE9AFA13FE38FA7F" box="[263,420,1436,1462]" pageId="3" pageNumber="18" value="1986-10-17" valueMax="1986-10-18" valueMin="1986-10-17">17–18.x.1986</collectingDate>
</date>
,
<collectorName id="26823DCBFFA94511FE32FA12FDB2FA7F" box="[431,558,1436,1463]" pageId="3" pageNumber="18">P. Spangler</collectorName>
&amp; O.S,
<collectorName id="26823DCBFFA94511FD17FA13FD5EFA7F" box="[650,706,1436,1462]" pageId="3" pageNumber="18">Flint</collectorName>
</materialsCitation>
,
<specimenCount id="9D719394FFA94511FD50FA13FCBFFA7F" box="[717,803,1436,1462]" pageId="3" pageNumber="18" type="male">2 males</specimenCount>
(USNM).
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF5AFA4FFBC6FA13" box="[199,1114,1472,1499]" pageId="3" pageNumber="18" type="etymology">
<paragraph id="8BC8581DFFA94511FF5AFA4FFBC6FA13" blockId="3.[151,1437,392,1534]" box="[199,1114,1472,1499]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFA4FFED2FA13" bold="true" box="[199,334,1472,1498]" pageId="3" pageNumber="18">Etymology.</emphasis>
The name is a Latin adjective referring to the
<typeStatus id="54CCE6BFFFA94511FCC8FA4EFC19FA12" box="[853,901,1473,1499]" pageId="3" pageNumber="18">type</typeStatus>
country (
<collectingCountry id="F360188DFFA94511FC6DFA4FFBD1FA13" box="[1008,1101,1472,1498]" name="Guyana" pageId="3" pageNumber="18">Guyana</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D0B96FFA94511FF5AFA6BFD85FA37" box="[199,537,1508,1534]" pageId="3" pageNumber="18" type="distribution">
<paragraph id="8BC8581DFFA94511FF5AFA6BFD85FA37" blockId="3.[151,1437,392,1534]" box="[199,537,1508,1534]" pageId="3" pageNumber="18">
<emphasis id="B903840FFFA94511FF5AFA6BFEFDFA37" bold="true" box="[199,353,1508,1534]" pageId="3" pageNumber="18">Distribution.</emphasis>
<collectingCountry id="F360188DFFA94511FEFAFA6BFE32FA37" box="[359,430,1508,1534]" name="Brazil" pageId="3" pageNumber="18">Brazil</collectingCountry>
,
<collectingCountry id="F360188DFFA94511FE27FA6BFD89FA37" box="[442,533,1508,1534]" name="Guyana" pageId="3" pageNumber="18">Guyana</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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