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<document id="A6F6A822465085EBA2F20EB893B8204E" ID-DOI="10.5852/ejt.2024.919.2407" ID-ISSN="2118-9773" ID-Zenodo-Dep="10559520" ID-ZooBank="72B7907F-F901-4DDD-B2DD-63D03253837E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706085863884" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03DE87B0FFF1FFBEFDCBFE0BA37AFCB9" docLanguage="en" docName="EJT.2024.919.1-87.pdf" docOrigin="European Journal of Taxonomy 919 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2407/10617" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gergithoides olivaceus Constant &amp; Pham 2024, sp. nov." docType="treatment" docUuid="EC8ABF4A-A058-4E0A-B635-235CC5FFE995" docUuidSource="ZooBank" docVersion="2" lastPageNumber="11" masterDocId="FFE7FFC8FFF7FFB4FFABFFD4A706FFDA" masterDocTitle="Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)" masterLastPageNumber="87" masterPageNumber="1" pageNumber="7" updateTime="1706086549316" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6CAF8157900DE99D91F2BDC4741AEDEE">Sixteen issid planthopper species in one day in Dong Son-Ky Thuong Nature Reserve in North Vietnam: Eight new species, one new genus and additional new records (Hemiptera: Fulgoromorpha: Issidae)</mods:title>
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<mods:namePart id="97AFC30B99E30CBDC5AE52A36755AE0D">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="581AE70EFB6E32B11473CCC52966720D">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="550AC389C25356A5B3E9F877B58124E4" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="5A3E5143FC4646405F1BD9DE030D9C6C">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="E5E2BC93460DE51E1ADE00BBB0B64BE4">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Street, Hanoi, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="999D3E3ECEEEDC473F63D9E3E89ACE2C" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="BD12EC72150F568A8C4B4643C26E3EA5">European Journal of Taxonomy</mods:title>
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<mods:date id="E34DE04AF05ADF2AEC9A5B6FAB1C28EF">2024</mods:date>
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<subSubSection id="C36D652DFFF1FFB2FDCBFE0BA3CAFDC7" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BC836A6FFF1FFB2FDCBFE0BA4D2FE20" blockId="6.[360,1228,479,575]" box="[608,980,479,506]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FDCBFE0BA4D2FE20" box="[608,980,479,506]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="4C774D25FFF1FFB2FDCBFE0BA474FE20" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[608,882,479,506]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFF1FFB2FDCBFE0BA474FE20" bold="true" box="[608,882,479,506]" italics="true" pageId="6" pageNumber="7">Gergithoides olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFF1FFB2FCD1FE34A4D2FE20" box="[890,980,480,506]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
<uuid id="FFD10C73FFF1FFB2FD2EFDD6A3CAFDC7" box="[645,1228,514,541]" pageId="6" pageNumber="7">EC8ABF4A-A058-4E0A-B635-235CC5FFE995</uuid>
</uri>
</paragraph>
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<subSubSection id="C36D652DFFF1FFB2FD73FDF1A45DFDE5" box="[728,859,549,575]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BC836A6FFF1FFB2FD73FDF1A45DFDE5" blockId="6.[360,1228,479,575]" box="[728,859,549,575]" pageId="6" pageNumber="7">
<figureCitation id="134C2A23FFF1FFB2FD73FDF1A427FDE5" box="[728,801,549,575]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="6" pageNumber="7">Figs 1</figureCitation>
,
<figureCitation id="134C2A23FFF1FFB2FC84FDF1A45DFDE5" box="[815,859,549,575]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1928,1954]" captionStartId-1="9.[189,232,1892,1918]" captionTargetBox-0="[249,1338,273,1882]" captionTargetBox-1="[258,1320,276,1852]" captionTargetId-0="figure-18@8.[242,1346,265,1888]" captionTargetId-1="figure-12@9.[254,1333,265,1852]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." captionText-1="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.10559526" figureDoi-1="http://doi.org/10.5281/zenodo.10559532" httpUri-0="https://zenodo.org/record/10559526/files/figure.png" httpUri-1="https://zenodo.org/record/10559532/files/figure.png" pageId="6" pageNumber="7">34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFF1FFB2FF16FDB1A4B8FA99" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BC836A6FFF1FFB2FF16FDB1A630FDA5" blockId="6.[189,310,613,639]" box="[189,310,613,639]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FF16FDB1A630FDA5" bold="true" box="[189,310,613,639]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B903EAB4FFF1FFB2FF16FDB1A630FDA5" bold="true" box="[189,310,613,639]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FD40A42AFD74" blockId="6.[189,1399,660,964]" box="[189,812,660,687]" pageId="6" pageNumber="7">
<taxonomicName id="4C774D25FFF1FFB2FF16FD40A6CDFD74" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,459,660,686]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFF1FFB2FF16FD40A6CDFD74" box="[189,459,660,686]" italics="true" pageId="6" pageNumber="7">Gergithoides olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFF1FFB2FE79FD41A52BFD75" box="[466,557,661,687]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
can be recognized by
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FD63A5F6FCE3" blockId="6.[189,1399,660,964]" pageId="6" pageNumber="7">
(1) the shape of the lateroventral bispinose process of the periandrium with anterior spine strongly elongate cephalad, weakly curved, then sinuate in distal portion and pointed apically and posterior spine directed posterodorsad along basal half, then strongly but evenly recurved ventrad and tapering to pointed apex (
<emphasis id="B903EAB4FFF1FFB2FD8CFCCBA54DFCE3" box="[551,587,799,825]" italics="true" pageId="6" pageNumber="7">lvp</emphasis>
<figureCitation id="134C2A23FFF1FFB2FDC3FCCBA5E5FCE3" box="[616,739,799,825]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="6" pageNumber="7">Fig. 4EF</figureCitation>
);
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FC96A663FCA4" blockId="6.[189,1399,660,964]" pageId="6" pageNumber="7">
(2) the ventral lobe of the periandrium with dorsal margin broadly rounded, nearly truncate (
<emphasis id="B903EAB4FFF1FFB2FAECFC95A25AFC81" box="[1351,1372,833,859]" italics="true" pageId="6" pageNumber="7">vl</emphasis>
<figureCitation id="134C2A23FFF1FFB2FF52FCB0A651FCA4" box="[249,343,868,894]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="6" pageNumber="7">Fig. 4G</figureCitation>
);
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FC53A66AFC1E" blockId="6.[189,1399,660,964]" pageId="6" pageNumber="7">
(3) the median posterior processes of periandrium pointed apically and laterally emarginate subapically (
<figureCitation id="134C2A23FFF1FFB2FEABFC7DA659FC1E" box="[256,351,937,964]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="6" pageNumber="7">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FC3EA6CFFBDE" blockId="6.[189,457,1002,1028]" box="[189,457,1002,1028]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FF16FC3EA6CFFBDE" bold="true" box="[189,457,1002,1028]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B903EAB4FFF1FFB2FF16FC3EA6CFFBDE" bold="true" box="[189,457,1002,1028]" pageId="6" pageNumber="7">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FBCCA3FAFBA2" blockId="6.[189,1399,1048,1145]" pageId="6" pageNumber="7">
The closest species, sharing the character of a bispinose lateroventral process of the periandrium with anterior spine elongate and posterior spine curved, are
<taxonomicName id="4C774D25FFF1FFB2FC32FBEFA35EFB8F" authorityName="Schumacher" authorityYear="1915" box="[921,1112,1083,1109]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carinatifrons">
<emphasis id="B903EAB4FFF1FFB2FC32FBEFA35EFB8F" box="[921,1112,1083,1109]" italics="true" pageId="6" pageNumber="7">G. carinatifrons</emphasis>
</taxonomicName>
described from
<collectingCountry id="F3607636FFF1FFB2FA8AFBEFA271FB8F" box="[1313,1399,1083,1109]" name="Taiwan" pageId="6" pageNumber="7">Taiwan</collectingCountry>
(
<bibRefCitation id="EFE64B57FFF1FFB2FF6EFB8AA69DFBA3" author="Schumacher F." box="[197,411,1118,1145]" pageId="6" pageNumber="7" pagination="71 - 134" refId="ref35587" refString="Schumacher F. 1915. Der gegenwartige Stand unserer Kenntnis von der Homopteren-Fauna der Insel Formosa. Mitteilungen aus dem Zoologischen Museum in Berlin 8 (1): 71 - 134. Available from https: // www. biodiversitylibrary. org / page / 30243518 [accessed 21 Dec. 2023]." type="journal article" year="1915">Schumacher 1915</bibRefCitation>
) and
<taxonomicName id="4C774D25FFF1FFB2FE76FB8AA572FBA2" authorityName="Wang &amp; Che" authorityYear="2003" box="[477,628,1118,1144]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B903EAB4FFF1FFB2FE76FB8AA572FBA2" box="[477,628,1118,1144]" italics="true" pageId="6" pageNumber="7">G. undulatus</emphasis>
</taxonomicName>
described from Hainan and Guangxi (
<bibRefCitation id="EFE64B57FFF1FFB2FB9DFB8AA3F6FBA2" author="Che Y. - L. &amp; Wang Y. - L. &amp; Chou I." box="[1078,1264,1118,1144]" pageId="6" pageNumber="7" pagination="102 - 108" refId="ref32807" refString="Che Y. - L., Wang Y. - L. &amp; Chou I. 2003. Taxonomic study on the genus Gergithoides Schumacher (Homoptera: Fulgoroidea: Issidae). Entomotaxonomia 25 (2): 102 - 108. [In Chinese.]" type="journal article" year="2003">
Che
<emphasis id="B903EAB4FFF1FFB2FBC5FB8BA3A7FBA2" box="[1134,1185,1118,1144]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2003
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FB4BA4B8FA99" blockId="6.[189,1398,1182,1348]" pageId="6" pageNumber="7">
From both
<taxonomicName id="4C774D25FFF1FFB2FEEFFB4BA507FB63" authorityName="Schumacher" authorityYear="1915" box="[324,513,1183,1209]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carinatifrons">
<emphasis id="B903EAB4FFF1FFB2FEEFFB4BA507FB63" box="[324,513,1183,1209]" italics="true" pageId="6" pageNumber="7">G. carinatifrons</emphasis>
</taxonomicName>
(illustrations in
<bibRefCitation id="EFE64B57FFF1FFB2FD6DFB4BA4B6FB63" author="Chan M. - L. &amp; Yang C. - T." box="[710,944,1183,1209]" pageId="6" pageNumber="7" refId="ref32694" refString="Chan M. - L. &amp; Yang C. - T. 1994. Issidae of Taiwan: (Homoptera: Fulgoroidea). National Chung Hsing University, Department of Entomology, Taichung, Taiwan." type="book" year="1994">Chan &amp; Yang 1994</bibRefCitation>
: fig. 5) and
<taxonomicName id="4C774D25FFF1FFB2FBEBFB4BA3DCFB63" authorityName="Wang &amp; Che" authorityYear="2003" box="[1088,1242,1183,1209]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B903EAB4FFF1FFB2FBEBFB4BA3DCFB63" box="[1088,1242,1183,1209]" italics="true" pageId="6" pageNumber="7">G. undulatus</emphasis>
</taxonomicName>
(illustrations in
<bibRefCitation id="EFE64B57FFF1FFB2FF4BFB15A6A6FB01" author="Che Y. - L. &amp; Wang Y. - L. &amp; Chou I." box="[224,416,1217,1243]" pageId="6" pageNumber="7" pagination="102 - 108" refId="ref32807" refString="Che Y. - L., Wang Y. - L. &amp; Chou I. 2003. Taxonomic study on the genus Gergithoides Schumacher (Homoptera: Fulgoroidea: Issidae). Entomotaxonomia 25 (2): 102 - 108. [In Chinese.]" type="journal article" year="2003">
Che
<emphasis id="B903EAB4FFF1FFB2FEB3FB16A651FB01" box="[280,343,1217,1243]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2003
</bibRefCitation>
: figs 16),
<taxonomicName id="4C774D25FFF1FFB2FD80FB16A5C4FB01" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[555,706,1217,1244]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFF1FFB2FD80FB16A5C4FB01" box="[555,706,1217,1244]" italics="true" pageId="6" pageNumber="7">G. olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFF1FFB2FD67FB16A42CFB06" box="[716,810,1218,1244]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
can be separated by its apically sinuate anterior spine of the lateroventral process of the periandrium (evenly curved dorsad in
<taxonomicName id="4C774D25FFF1FFB2FB2AFB30A247FB24" authorityName="Schumacher" authorityYear="1915" box="[1153,1345,1252,1278]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carinatifrons">
<emphasis id="B903EAB4FFF1FFB2FB2AFB30A247FB24" box="[1153,1345,1252,1278]" italics="true" pageId="6" pageNumber="7">G. carinatifrons</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D25FFF1FFB2FF16FAD3A651FAFB" authorityName="Wang &amp; Che" authorityYear="2003" box="[189,343,1287,1313]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B903EAB4FFF1FFB2FF16FAD3A651FAFB" box="[189,343,1287,1313]" italics="true" pageId="6" pageNumber="7">G. undulatus</emphasis>
</taxonomicName>
) and the evenly rounded hook formed by the posterior spine of the periandrium process (strongly, abruptly reflexed in
<taxonomicName id="4C774D25FFF1FFB2FD8AFAFEA5DBFA99" authorityName="Schumacher" authorityYear="1915" box="[545,733,1321,1348]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="carinatifrons">
<emphasis id="B903EAB4FFF1FFB2FD8AFAFEA5DBFA99" box="[545,733,1321,1348]" italics="true" pageId="6" pageNumber="7">G. carinatifrons</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D25FFF1FFB2FCBDFAFEA4A8FA99" authorityName="Wang &amp; Che" authorityYear="2003" box="[790,942,1321,1348]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undulatus">
<emphasis id="B903EAB4FFF1FFB2FCBDFAFEA4A8FA99" box="[790,942,1321,1348]" italics="true" pageId="6" pageNumber="7">G. undulatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFF1FFB2FF16FABEA56CFA0F" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BC836A6FFF1FFB2FF16FABEA642FA5E" blockId="6.[189,324,1386,1412]" box="[189,324,1386,1412]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FF16FABEA642FA5E" bold="true" box="[189,324,1386,1412]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B903EAB4FFF1FFB2FF16FABEA642FA5E" bold="true" box="[189,324,1386,1412]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16FA4CA56CFA0F" blockId="6.[189,1399,1432,1493]" pageId="6" pageNumber="7">
The species epithet
<taxonomicName id="4C774D25FFF1FFB2FE03FA4CA511FA68" authorityName="Constant &amp; Pham" authorityYear="2024" box="[424,535,1432,1458]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="olivaceus">
<emphasis id="B903EAB4FFF1FFB2FE03FA4CA511FA68" box="[424,535,1432,1458]" italics="true" pageId="6" pageNumber="7">olivaceus</emphasis>
</taxonomicName>
is a Latin adjective meaning olive-green. It refers to the general colour of the specimens of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFF1FFB2FF16FA28A692F8E9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BC836A6FFF1FFB2FF16FA28A66AF9CF" blockId="6.[189,364,1531,1558]" box="[189,364,1531,1558]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FF16FA28A66AF9CF" bold="true" box="[189,364,1531,1558]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B903EAB4FFF1FFB2FF16FA28A66AF9CF" bold="true" box="[189,364,1531,1558]" pageId="6" pageNumber="7">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF41F9FEA65AF99E" blockId="6.[189,1399,1578,1709]" box="[234,348,1578,1604]" pageId="6" pageNumber="7">
<emphasis id="B903EAB4FFF1FFB2FF41F9FEA65AF99E" bold="true" box="[234,348,1578,1604]" pageId="6" pageNumber="7">
<typeStatus id="54CC8804FFF1FFB2FF41F9FEA65AF99E" box="[234,348,1578,1604]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16F999A330F976" blockId="6.[189,1399,1578,1709]" pageId="6" pageNumber="7">
<materialsCitation id="3B1F3CFBFFF1FFB2FF16F999A330F976" collectingDate="2022-08-27" collectionCode="RBINS" collectorName="Project &amp; J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Quang Ninh" typeStatus="holotype">
<collectingCountry id="F3607636FFF1FFB2FF16F999A642F9BD" box="[189,324,1613,1639]" name="Vietnam" pageId="6" pageNumber="7">VIETNAM</collectingCountry>
<specimenCount id="9D71FD2FFFF1FFB2FECBF999A670F9BD" box="[352,374,1613,1639]" count="1" pageId="6" pageNumber="7" type="male"></specimenCount>
(dissected, right hind wing mounted,
<figureCitation id="134C2A23FFF1FFB2FC97F998A4A1F9BD" box="[828,935,1612,1639]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1928,1954]" captionStartId-1="9.[189,232,1892,1918]" captionTargetBox-0="[249,1338,273,1882]" captionTargetBox-1="[258,1320,276,1852]" captionTargetId-0="figure-18@8.[242,1346,265,1888]" captionTargetId-1="figure-12@9.[254,1333,265,1852]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." captionText-1="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.10559526" figureDoi-1="http://doi.org/10.5281/zenodo.10559532" httpUri-0="https://zenodo.org/record/10559526/files/figure.png" httpUri-1="https://zenodo.org/record/10559532/files/figure.png" pageId="6" pageNumber="7">Figs 34</figureCitation>
);
<collectingRegion id="49B3F844FFF1FFB2FC15F999A3C2F9BC" box="[958,1220,1612,1639]" country="Vietnam" name="Quang Ninh" pageId="6" pageNumber="7">Quang Ninh Province</collectingRegion>
,
<collectingMunicipality id="6BACACDCFFF1FFB2FB7AF999A214F9BD" box="[1233,1298,1613,1639]" pageId="6" pageNumber="7">Dong</collectingMunicipality>
Son-Ky
<location id="8EA8607DFFF1FFB2FF16F9BBA6DEF950" LSID="urn:lsid:plazi:treatment:03DE87B0FFF1FFBEFDCBFE0BA37AFCB9:8EA8607DFFF1FFB2FF16F9BBA6DEF950" box="[189,472,1647,1674]" country="Vietnam" latitude="21.141388" longLatPrecision="21" longitude="107.08139" municipality="Dong" name="Thuong Nature Reserve" pageId="6" pageNumber="7" stateProvince="Quang Ninh">Thuong Nature Reserve</location>
;
<geoCoordinate id="EE435061FFF1FFB2FE4FF9BAA57FF950" box="[484,633,1646,1674]" degrees="21" direction="north" minutes="08" orientation="latitude" pageId="6" pageNumber="7" precision="15" seconds="29" value="21.141388">21°0829″ N</geoCoordinate>
,
<geoCoordinate id="EE435061FFF1FFB2FD2EF9BAA420F950" box="[645,806,1646,1674]" degrees="107" direction="east" minutes="04" orientation="longitude" pageId="6" pageNumber="7" precision="15" seconds="53" value="107.08139">107°0453″ E</geoCoordinate>
;
<date id="FFC91066FFF1FFB2FC99F9BBA4D5F953" box="[818,979,1647,1673]" pageId="6" pageNumber="7" value="2022-08-27">
<collectingDate id="EF8DE98EFFF1FFB2FC99F9BBA4D5F953" box="[818,979,1647,1673]" pageId="6" pageNumber="7" value="2022-08-27">27 Aug. 2022</collectingDate>
</date>
;
<elevation id="005AD195FFF1FFB2FC74F9BBA36FF953" box="[991,1129,1647,1674]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="m" value="550.0">
<quantity id="4C8F9B43FFF1FFB2FC74F9BBA32FF950" box="[991,1065,1647,1674]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
; secondary forest; GTI
<collectorName id="26825370FFF1FFB2FF16F946A612F976" box="[189,276,1682,1708]" pageId="6" pageNumber="7">Project</collectorName>
;
<collectorName id="26825370FFF1FFB2FE8BF946A6A3F976" box="[288,421,1682,1708]" pageId="6" pageNumber="7">J. Constant</collectorName>
,
<collectorName id="26825370FFF1FFB2FE1AF946A528F976" box="[433,558,1682,1708]" pageId="6" pageNumber="7">J. Bresseel</collectorName>
and
<collectorName id="26825370FFF1FFB2FDCCF946A5FEF976" box="[615,760,1682,1708]" pageId="6" pageNumber="7">L. Semeraro</collectorName>
leg.; I.G.: 34.518;
<collectionCode id="ED66AE63FFF1FFB2FC7CF946A337F976" box="[983,1073,1682,1708]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="6" pageNumber="7" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF41F907A661F937" blockId="6.[189,1398,1747,1843]" box="[234,359,1747,1773]" pageId="6" pageNumber="7">
<emphasis id="B903EAB4FFF1FFB2FF41F907A661F937" bold="true" box="[234,359,1747,1773]" pageId="6" pageNumber="7">
<typeStatus id="54CC8804FFF1FFB2FF41F907A661F937" box="[234,359,1747,1773]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16F922A692F8E9" blockId="6.[189,1398,1747,1843]" pageId="6" pageNumber="7">
<materialsCitation id="3B1F3CFBFFF1FFB2FF16F922A4CDF8D5" box="[189,971,1781,1808]" collectingDate="2022-08-27" collectionCode="RBINS" collectorName="Project &amp; J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Quang Ninh" typeStatus="paratype">
<collectingCountry id="F3607636FFF1FFB2FF16F922A642F8CA" box="[189,324,1782,1808]" name="Vietnam" pageId="6" pageNumber="7">VIETNAM</collectingCountry>
<specimenCount id="9D71FD2FFFF1FFB2FECEF921A693F8CA" box="[357,405,1781,1808]" count="1" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED66AE63FFF1FFB2FCDEF921A4CDF8D5" box="[885,971,1781,1807]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="6" pageNumber="7" type="Museum">RBINS</collectionCode>
</materialsCitation>
<materialsCitation id="3B1F3CFBFFF1FFB2FC47F921A697F8E9" collectingDate="2022-08-27" collectionCode="VNMN" collectorName="Project &amp; J. Constant &amp; J. Bresseel &amp; L. Semeraro" country="Vietnam" elevation="550" latitude="21.141388" location="Thuong Nature Reserve" longLatPrecision="21" longitude="107.08139" municipality="Dong" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Quang Ninh" typeStatus="paratype">
<specimenCount id="9D71FD2FFFF1FFB2FC47F921A31AF8CA" box="[1004,1052,1781,1808]" count="1" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED66AE63FFF1FFB2FE98F8CDA697F8E9" box="[307,401,1817,1843]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="6" pageNumber="7">VNMN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFF1FFBEFF16F88DA6FCFDBA" lastPageId="10" lastPageNumber="11" pageId="6" pageNumber="7" type="description">
<paragraph id="8BC836A6FFF1FFB2FF16F88DA648F8A9" blockId="6.[189,334,1881,1907]" box="[189,334,1881,1907]" pageId="6" pageNumber="7">
<heading id="D08081CAFFF1FFB2FF16F88DA648F8A9" bold="true" box="[189,334,1881,1907]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B903EAB4FFF1FFB2FF16F88DA648F8A9" bold="true" box="[189,334,1881,1907]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFF1FFB2FF16F853A400F81E" blockId="6.[189,1399,1927,1988]" pageId="6" pageNumber="7">
<smallCapsWord id="8D2EA07AFFF1FFB2FF16F853A668F845" baselines="1946,1946" box="[189,366,1927,1953]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="6" pageNumber="7">MEASUREMENTS</smallCapsWord>
<smallCapsWord id="8D2EA07AFFF1FFB2FEDCF859A6A2F87A" baselines="1947" box="[375,420,1933,1952]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="6" pageNumber="7">AND</smallCapsWord>
<smallCapsWord id="8D2EA07AFFF1FFB2FE07F858A6F3F845" baselines="1946" box="[428,501,1932,1951]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="ratios" pageId="6" pageNumber="7">RATIOS</smallCapsWord>
. LT:
<specimenCount id="9D71FD2FFFF1FFB2FD9FF853A54CF87B" box="[564,586,1927,1953]" count="1" pageId="6" pageNumber="7" type="male"></specimenCount>
(n = 1):
<quantity id="4C8F9B43FFF1FFB2FD1FF853A414F87B" box="[692,786,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="7" unit="mm" value="5.6">5.6 mm</quantity>
,
<specimenCount id="9D71FD2FFFF1FFB2FCB5F853A432F87B" box="[798,820,1927,1953]" count="1" pageId="6" pageNumber="7" type="female"></specimenCount>
(n = 2): 6.0
<quantity id="4C8F9B43FFF1FFB2FC79F853A336F87B" box="[978,1072,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="7" unit="mm" value="6.4">6.4 mm</quantity>
; LT/BB = 1.34; LTg/BTg = 1.67; LW/BW = 1.9; BV/LV = 0.8; LF/BF = 1.27.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16FEDDA442FDE6" blockId="7.[187,1400,265,572]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16FEDDA7FAFEF8" baselines="284,285" box="[189,252,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="7" pageNumber="8">HEAD</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FEA4FEDDA694FEF9" box="[271,402,265,291]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3AD</figureCitation>
). Vertex slightly longer in midline than broad, brown with margins carinate; anterior margin convex, posterior one concave and lateral converging anteriorly; disc excavate with obsolete tubercles. Side of head yellowish brown. Frons elongate and rugulose, dark reddish brown; median carina yellowish, narrowly margined with a black line, extending from dorsal margin down to frontoclypeal suture; row of yellowish tubercles on black background along dorsal and lateral margins extending to level of base of eyes; lateral margins yellowish under eyes, the yellowish line marginated by inner thin black line. Clypeus black-brown with basilateral yellowish spot and black on sides and apex, elevated medially. Labium dark brown with last segment longer than broad, and shorter than penultimate. Scape short, ring-shaped, black; pedicel bulbous, dark brown.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16FDB1A6A1FC42" blockId="7.[189,1400,612,920]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16FDB1A61FFDA7" baselines="632,632" box="[189,281,613,639]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="7" pageNumber="8">THORAX</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FE84FDB0A6B1FDA4" box="[303,439,612,639]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3AD</figureCitation>
). Variegated olivaceous brown. Pronotum very short, about half the length of mesonotum in midline, with median carina; anterior and posterior margins carinate; disc rugulose, concave with impressed point on each side of median carina; row of yellowish tubercles along anterior margin extending on paranotal fields to level of base of eyes; another oblique row of 3 weak yellowish tubercles on each side of disc, parallel to anterior margin. Mesonotum short, slightly coriaceous with yellowish median carina and obsolete impressed point on each side of disc; transverse carina along anterior margin yellowish; 24 yellowish blunt tubercles grouped at each external angle; yellowish suffused marking on each side of scutellum; apex of scutellum elevated and marked with dark brown. Tegulae olivaceous.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16FC14A538FBBC" blockId="7.[189,1400,959,1126]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16FC14A624FC02" baselines="979,979" box="[189,290,960,986]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Tegmina" pageId="7" pageNumber="8">TEGMINA</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FE9FFC6BA6B2FC03" box="[308,436,959,986]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3A, C</figureCitation>
). Strongly convex; slightly longer than broad when taken together in dorsal view; slightly concave at basocostal angle; subcoriaceous with dense reticulum of slightly raised veins and veinlets; mains veins barely distinct basally; yellowish brown with brown to black irregular markings, especially along inner margin of cells, making the insect look olivaceous from a distance; small darker spot in middle of sutural margin.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16FB5AA46FFACB" blockId="7.[188,1399,1165,1297]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16FB5AA7F1FB7C" baselines="1185,1185" box="[189,247,1166,1192]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="7" pageNumber="8">HIND</smallCapsWord>
<smallCapsWord id="8D2EA07AFFF0FFB3FF55FB47A644FB7C" baselines="1185" box="[254,322,1171,1190]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="wings" pageId="7" pageNumber="8">WINGS</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FEF8FB59A6ABFB7D" box="[339,429,1165,1192]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3E</figureCitation>
). Brown, with costal half suffused with blackish, except basicostal angle, unilobed, with veins darker than cells; elongate, slightly shorter than tegmina. Venation reticulate with main longitudinal veins distinct basally (C, ScP+R, MP, CuA); numerous cross-veinlets. Anal area obsolete. Costal and cubital margins sinuate, distal margin rounded.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16FAEDA613FA05" blockId="7.[186,1401,1336,1503]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16FAEDA7F0FA8B" baselines="1356,1356" box="[189,246,1337,1363]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="7" pageNumber="8">LEGS</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FEAFFAECA682FA88" box="[260,388,1336,1363]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3AD</figureCitation>
). Elongate and slender, brown with trochanters, base of femora, line along externoventral margin of femora, black; black-brown line on ventral carinae of tibiae; base and apex of tibiae, apex of tarsi and apex of metatibial spines black-brown; pro- and mesofemora with pale yellow tubercles. Posterior tibiae with 2 lateral spines on apical ⅓ and 6 apical spines. Metatibiotarsal formula: (2) 6 / 89 / 2.
</paragraph>
<paragraph id="8BC836A6FFF0FFB3FF16F9D3A6A8F99E" blockId="7.[189,1398,1542,1604]" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16F9D3A62AF9C5" baselines="1562,1562" box="[189,300,1543,1569]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="7" pageNumber="8">ABDOMEN</smallCapsWord>
(
<figureCitation id="134C2A23FFF0FFB3FE94F9D2A69DF9FB" box="[319,411,1542,1569]" captionStart="Fig" captionStartId="8.[189,232,1928,1954]" captionTargetBox="[249,1338,273,1882]" captionTargetId="figure-18@8.[242,1346,265,1888]" captionTargetPageId="8" captionText="Fig. 3. Gergithoides olivaceus sp. nov., dissected holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Habitus, ventral view. C. Habitus, lateral view. D. Habitus, perpendicular view of frons. E. Right hind wing." figureDoi="http://doi.org/10.5281/zenodo.10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
). Brown with middle and posterior area darker, often with irregular paler markings on sides of sternites.
</paragraph>
<paragraph id="8BC836A6FFF0FFBEFF16F9B8A6FCFDBA" blockId="7.[186,1398,1643,2022]" lastBlockId="10.[189,1399,265,608]" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<smallCapsWord id="8D2EA07AFFF0FFB3FF16F9B8A7F9F95E" baselines="1663,1663" box="[189,255,1644,1670]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="7" pageNumber="8">MALE</smallCapsWord>
<smallCapsWord id="8D2EA07AFFF0FFB3FEADF9A5A681F95E" baselines="1663" box="[262,391,1649,1668]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="terminalia" pageId="7" pageNumber="8">TERMINALIA</smallCapsWord>
. Pygofer (
<emphasis id="B903EAB4FFF0FFB3FDA9F9B8A527F95F" box="[514,545,1644,1669]" italics="true" pageId="7" pageNumber="8">Py</emphasis>
) 2.2 times as high as broad in lateral view, with posterior margin strongly sinuate, rounded along dorsal ⅔ (
<figureCitation id="134C2A23FFF0FFB3FDF2F95AA5D9F972" box="[601,735,1678,1704]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="7" pageNumber="8">Fig. 4AB</figureCitation>
); ventral margin rounded in lateral view (
<figureCitation id="134C2A23FFF0FFB3FB4FF95AA26CF972" box="[1252,1386,1678,1704]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="7" pageNumber="8">Fig. 4AB</figureCitation>
). Gonostyli (
<emphasis id="B903EAB4FFF0FFB3FEE8F966A65EF916" box="[323,344,1714,1740]" italics="true" pageId="7" pageNumber="8">G</emphasis>
<figureCitation id="134C2A23FFF0FFB3FED3F965A51DF916" box="[376,539,1713,1740]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="7" pageNumber="8">Fig. 4AB, D</figureCitation>
) elongate in lateral view, with ventral margin broadly rounded in lateral view, posterior margin more narrowly rounded in lateral view, dorsal margin oblique basally to neck of capitulum, more posteriorly forming a straight angle with neck of capitulum; capitulum (
<emphasis id="B903EAB4FFF0FFB3FB1BF92DA3CAF8C8" box="[1200,1228,1785,1810]" italics="true" pageId="7" pageNumber="8">ca</emphasis>
) with elongate neck, curved dorsocephalad in lateral view, mesad in posterior view, with small blunt lateral projection in middle of outer margin; apical portion anteroposteriorly laminate, subrectangular in caudal view, bearing one dorsal tooth, one tooth at inner angle and one tooth pointing anteroventrally at anterobasal, followed posterad by a lateral lamina curved ventrad looking like an additional tooth in caudal view.Anal tube (
<emphasis id="B903EAB4FFF0FFB3FEAAF87CA624F818" box="[257,290,1960,1986]" italics="true" pageId="7" pageNumber="8">An</emphasis>
<figureCitation id="134C2A23FFF0FFB3FEEAF87CA6C6F818" box="[321,448,1960,1986]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="7" pageNumber="8">Fig. 4AC</figureCitation>
) nearly as broad as long in midline in dorsal view, dorsoventrally flattened with sides sinuate, diverging towards apex, apical margin weakly bisinuate in dorsal view; in lateral view, anal tube curved ventrally; apicolateral angles roundly projecting lateroventrally. Aedeagus strongly curved posterodorsally, rather simple (
<figureCitation id="134C2A23FFFDFFBEFD29FEF8A5FEFE9C" box="[642,760,300,326]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="10" pageNumber="11">Fig. 4EI</figureCitation>
). Periandrium with lateroventral process (
<emphasis id="B903EAB4FFFDFFBEFB4CFEF8A20DFE9C" box="[1255,1291,300,326]" italics="true" pageId="10" pageNumber="11">lvp</emphasis>
) at basal half showing 2 spines; anterior spine strongly elongate cephalad, weakly curved, more or less parallel to ventral margin of aedeagus, sinuate in distal portion and pointed apically; posterior spine directed posterodorsad, forming a nearly right angle with anterior spine, slightly widening along basal half, then strongly recurved ventrad and tapering to pointed apex (
<figureCitation id="134C2A23FFFDFFBEFCFBFE6DA4CAFE09" box="[848,972,441,467]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="10" pageNumber="11">Fig. 4EF</figureCitation>
); periandrium with laminate process reflexed cephalad on each side in distal portion, with right one slightly more developed and projecting slightly further laterally than left one (
<figureCitation id="134C2A23FFFDFFBEFD2EFE2BA413FDC3" box="[645,789,511,537]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="10" pageNumber="11">Fig. 4EF, I</figureCitation>
) and 2 median processes pointed apically and with lateral margin sinuate anteapically (
<figureCitation id="134C2A23FFFDFFBEFDC9FDF6A5C6FDE7" box="[610,704,546,573]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="10" pageNumber="11">Fig. 4G</figureCitation>
). Ventral lobe of periandrium (
<emphasis id="B903EAB4FFFDFFBEFB81FDF6A339FDE6" box="[1066,1087,546,572]" italics="true" pageId="10" pageNumber="11">vl</emphasis>
) broad with apical margin roundly truncate (
<figureCitation id="134C2A23FFFDFFBEFE25FD92A6EBFDBA" box="[398,493,582,608]" captionStart="Fig" captionStartId="9.[189,232,1892,1918]" captionTargetBox="[258,1320,276,1852]" captionTargetId="figure-12@9.[254,1333,265,1852]" captionTargetPageId="9" captionText="Fig. 4. Gergithoides olivaceus sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Anterolateral view, showing apex of capitulum. C. Dorsal view. D. Caudal view. EI. Aedeagus and connective. E. Left lateral view. F. Right lateral view. G. Posteroventral view. H. Ventral view. I. Anterodorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="10" pageNumber="11">Fig. 4G</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF08662EFFFFFFBCFF16F85CA63DF832" ID-DOI="http://doi.org/10.5281/zenodo.10559526" ID-Zenodo-Dep="10559526" httpUri="https://zenodo.org/record/10559526/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1928,1954]" targetBox="[249,1338,273,1882]" targetPageId="8">
<paragraph id="8BC836A6FFFFFFBCFF16F85CA63DF832" blockId="8.[189,1399,1928,2024]" pageId="8" pageNumber="9">
<emphasis id="B903EAB4FFFFFFBCFF16F85CA616F878" bold="true" box="[189,272,1928,1954]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
<taxonomicName id="4C774D25FFFFFFBCFEB5F85CA534F878" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[286,562,1928,1954]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFFFFFBCFEB5F85CA534F878" box="[286,562,1928,1954]" italics="true" pageId="8" pageNumber="9">Gergithoides olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFFFFFBCFD94F85CA5A4F878" box="[575,674,1928,1954]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, dissected holotype, ♂ (RBINS).
<emphasis id="B903EAB4FFFFFFBCFBE2F85CA358F878" bold="true" box="[1097,1118,1928,1954]" pageId="8" pageNumber="9">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B903EAB4FFFFFFBCFF16F87FA7D7F81F" bold="true" box="[189,209,1963,1989]" pageId="8" pageNumber="9">B</emphasis>
. Habitus, ventral view.
<emphasis id="B903EAB4FFFFFFBCFE59F87FA501F81F" bold="true" box="[498,519,1963,1989]" pageId="8" pageNumber="9">C</emphasis>
. Habitus, lateral view.
<emphasis id="B903EAB4FFFFFFBCFC8BF87FA433F81F" bold="true" box="[800,821,1963,1989]" pageId="8" pageNumber="9">D</emphasis>
. Habitus, perpendicular view of frons.
<emphasis id="B903EAB4FFFFFFBCFABBF87FA222F81F" bold="true" box="[1296,1316,1963,1989]" pageId="8" pageNumber="9">E</emphasis>
. Right hind wing.
</paragraph>
</caption>
<caption id="DF08662EFFFEFFBDFF16F8B0A271F832" ID-DOI="http://doi.org/10.5281/zenodo.10559532" ID-Zenodo-Dep="10559532" httpUri="https://zenodo.org/record/10559532/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1892,1918]" targetBox="[258,1320,276,1852]" targetPageId="9">
<paragraph id="8BC836A6FFFEFFBDFF16F8B0A271F832" blockId="9.[189,1399,1892,2025]" pageId="9" pageNumber="10">
<emphasis id="B903EAB4FFFEFFBDFF16F8B0A609F8A5" bold="true" box="[189,271,1892,1919]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
<taxonomicName id="4C774D25FFFEFFBDFEB1F8B1A52BF8A5" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[282,557,1893,1919]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFFEFFBDFEB1F8B1A52BF8A5" box="[282,557,1893,1919]" italics="true" pageId="9" pageNumber="10">Gergithoides olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFFEFFBDFD92F8B1A59CF8A5" box="[569,666,1893,1919]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B903EAB4FFFEFFBDFBE6F8B0A380F8A4" bold="true" box="[1101,1158,1892,1918]" pageId="9" pageNumber="10">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B903EAB4FFFEFFBDFEDDF85CA68DF878" bold="true" box="[374,395,1928,1954]" pageId="9" pageNumber="10">A</emphasis>
. Left lateral view.
<emphasis id="B903EAB4FFFEFFBDFD2BF85CA592F878" bold="true" box="[640,660,1928,1954]" pageId="9" pageNumber="10">B</emphasis>
. Anterolateral view, showing apex of capitulum.
<emphasis id="B903EAB4FFFEFFBDFB54F85CA212F878" bold="true" box="[1279,1300,1928,1954]" pageId="9" pageNumber="10">C</emphasis>
. Dorsal view.
<emphasis id="B903EAB4FFFEFFBDFEA3F87FA61BF81F" bold="true" box="[264,285,1963,1989]" pageId="9" pageNumber="10">D</emphasis>
. Caudal view.
<emphasis id="B903EAB4FFFEFFBDFE76F87FA50CF81F" bold="true" box="[477,522,1963,1989]" pageId="9" pageNumber="10">EI</emphasis>
. Aedeagus and connective.
<emphasis id="B903EAB4FFFEFFBDFCCCF87FA47DF81F" bold="true" box="[871,891,1963,1989]" pageId="9" pageNumber="10">E</emphasis>
. Left lateral view.
<emphasis id="B903EAB4FFFEFFBDFBC5F87FA386F81F" bold="true" box="[1134,1152,1963,1989]" pageId="9" pageNumber="10">F</emphasis>
. Right lateral view.
<emphasis id="B903EAB4FFFEFFBDFF16F81AA7D2F832" bold="true" box="[189,212,1998,2024]" pageId="9" pageNumber="10">G</emphasis>
. Posteroventral view.
<emphasis id="B903EAB4FFFEFFBDFE7FF81AA6EDF832" bold="true" box="[468,491,1998,2024]" pageId="9" pageNumber="10">H</emphasis>
. Ventral view.
<emphasis id="B903EAB4FFFEFFBDFD3CF81AA5A4F832" bold="true" box="[663,674,1998,2024]" pageId="9" pageNumber="10">I</emphasis>
. Anterodorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<subSubSection id="C36D652DFFFDFFBEFF16FD53A48CFD29" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="8BC836A6FFFDFFBEFF16FD53A61AFD7B" blockId="10.[189,284,647,673]" box="[189,284,647,673]" pageId="10" pageNumber="11">
<heading id="D08081CAFFFDFFBEFF16FD53A61AFD7B" bold="true" box="[189,284,647,673]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B903EAB4FFFDFFBEFF16FD53A61AFD7B" bold="true" box="[189,284,647,673]" pageId="10" pageNumber="11">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFFDFFBEFF16FD62A48CFD29" blockId="10.[189,1398,694,756]" pageId="10" pageNumber="11">
<taxonomicName id="4C774D25FFFDFFBEFF16FD62A6D6FD0A" authority="Constant &amp; Pham, 2024" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,464,694,720]" class="Insecta" family="Issidae" genus="Gergithoides" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="olivaceus" status="sp. nov.">
<emphasis id="B903EAB4FFFDFFBEFF16FD62A6D6FD0A" box="[189,464,694,720]" italics="true" pageId="10" pageNumber="11">Gergithoides olivaceus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057CFFFFDFFBEFE77FD62A53AFD0A" box="[476,572,694,720]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
was collected in August on lower vegetation and bushes, in moist evergreen tropical forest at about
<quantity id="4C8F9B43FFFDFFBEFDECFD0DA594FD29" box="[583,658,729,756]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="11" unit="m" value="550.0">550 m</quantity>
in altitude (
<figureCitation id="134C2A23FFFDFFBEFCB4FD0DA47CFD29" box="[799,890,729,755]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="10" pageNumber="11">Fig. 1B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D652DFFFDFFBEFF16FCCEA37AFCB9" box="[189,1148,794,867]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BC836A6FFFDFFBEFF16FCCEA650FCEE" blockId="10.[189,342,794,820]" box="[189,342,794,820]" pageId="10" pageNumber="11">
<heading id="D08081CAFFFDFFBEFF16FCCEA650FCEE" bold="true" box="[189,342,794,820]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B903EAB4FFFDFFBEFF16FCCEA650FCEE" bold="true" box="[189,342,794,820]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836A6FFFDFFBEFF16FC9DA37AFCB9" blockId="10.[189,1148,841,867]" box="[189,1148,841,867]" pageId="10" pageNumber="11">
<collectingCountry id="F3607636FFFDFFBEFF16FC9DA623FCB9" box="[189,293,841,867]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
:
<collectingRegion id="49B3F844FFFDFFBEFE9BFC9DA534FCB9" box="[304,562,841,867]" country="Vietnam" name="Quang Ninh" pageId="10" pageNumber="11">Quang Ninh Province</collectingRegion>
, Dong Son-Ky Thuong Nature Reserve (
<figureCitation id="134C2A23FFFDFFBEFBBAFC9DA368FCB9" box="[1041,1134,841,867]" captionStart="Fig" captionStartId="2.[189,232,1998,2024]" captionTargetBox="[218,1369,265,1958]" captionTargetId="figure-18@2.[218,1369,265,1958]" captionTargetPageId="2" captionText="Fig. 1. Dong Son-Ky Thuong Nature Reserve. A. Topographic map. B. Sampled habitat." figureDoi="http://doi.org/10.5281/zenodo.10559522" httpUri="https://zenodo.org/record/10559522/files/figure.png" pageId="10" pageNumber="11">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>