330 lines
36 KiB
XML
330 lines
36 KiB
XML
<document id="616AF901852FA67E9EF14F8F128612DA" ID-DOI="10.11646/zootaxa.3722.4.1" ID-GBIF-Dataset="c092e547-503a-41ef-a855-4a12b5d776c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="223434" ID-ZooBank="3A461275-7565-4F6E-9785-12C45E12992E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460692161620" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2013" docId="03DE87B0FFC1FFCCCADAB146FB881C24" docLanguage="en" docName="zt03722p438.pdf" docOrigin="Zootaxa 3722 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrphonothrus kimberleyi Colloff, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="408" masterDocId="FFE7FFC8FFC4FFCBCA4DB328FF871A47" masterDocTitle="Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region" masterLastPageNumber="438" masterPageNumber="401" pageNumber="406" updateTime="1698351897157" updateUser="plazi">
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<mods:mods id="85B950AF67DF12BAA0F5E53EF47A17E7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B83923D9B5F1E094AAD5FFB9C0BB3AF7">
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<mods:title id="BAE1131B46F89CACA4A16DC806DCCDC7">Species-groups and biogeography of the oribatid mite family Malaconothridae (Oribatida: Malaconothroidea), with new species from the south-western Pacific region</mods:title>
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</mods:titleInfo>
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<mods:name id="886B35EE2E5EF6DE1899007960E734A7" type="personal">
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<mods:role id="91230544FEDBEC17D6FED492C362BA8A">
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<mods:roleTerm id="494374CC83DA4F80909DF3BA6B4E4C80">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4A1583809C36844F340E5BEA31237123">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="66685A2032D851CDACB5EE252CF12C8D">text</mods:typeOfResource>
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<mods:titleInfo id="2C12B274382D6EDCD8CAC9B536DA4F99">
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<mods:title id="6FFC4CF81F414ACD92E2839B832687E3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="56EB2796D729B860DE727A21737A2306">
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<mods:date id="E215BBCB8F21529E77209F3852B3157B">2013</mods:date>
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<mods:detail id="9CA2679869951EC7C07AA43C8F1C801C" type="volume">
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<mods:number id="F7912828DFFEA8CBBB5A60DAF635B4BA">3722</mods:number>
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</mods:detail>
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<mods:detail id="C236597E5674AF98CB5FF92BA06865E9" type="issue">
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<mods:number id="C136BA357ED84CF860FFB042104A98DE">4</mods:number>
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<mods:start id="77A7744C83D31A23A3A80D5AE4CFBDD6">401</mods:start>
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<mods:classification id="93CDDC1DDCFA8F0FCBD4ED87427282C6">journal article</mods:classification>
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<mods:identifier id="913AAABADA2E85BAC890E9FF1D90C9C8" type="DOI">10.11646/zootaxa.3722.4.1</mods:identifier>
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<mods:identifier id="167774FADD45C674701C586341E32C8D" type="GBIF-Dataset">c092e547-503a-41ef-a855-4a12b5d776c4</mods:identifier>
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<mods:identifier id="09892052FF9F579E3044731B0A26B5C2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="34289C3EED0517EFA9DC64DB67925952" type="Zenodo-Dep">223434</mods:identifier>
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<mods:identifier id="C15A2D1B6A9A33E482985D0C4EFE96F3" type="ZooBank">3A461275-7565-4F6E-9785-12C45E12992E</mods:identifier>
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</mods:mods>
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<treatment id="03DE87B0FFC1FFCCCADAB146FB881C24" ID-DOI="http://doi.org/10.5281/zenodo.6157846" ID-GBIF-Taxon="119580502" ID-Zenodo-Dep="6157846" LSID="urn:lsid:plazi:treatment:03DE87B0FFC1FFCCCADAB146FB881C24" httpUri="http://treatment.plazi.org/id/03DE87B0FFC1FFCCCADAB146FB881C24" lastPageId="7" lastPageNumber="408" pageId="5" pageNumber="406">
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<subSubSection id="C36D652DFFC1FFCECADAB146FAD91951" pageId="5" pageNumber="406" type="nomenclature">
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<paragraph id="8BC836A6FFC1FFCECADAB146FDB018CF" blockId="5.[151,567,622,682]" box="[151,567,622,648]" pageId="5" pageNumber="406">
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<heading id="D08081CAFFC1FFCECADAB146FDB018CF" bold="true" box="[151,567,622,648]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
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<emphasis id="B903EAB4FFC1FFCECADAB146FDB018CF" bold="true" box="[151,567,622,648]" pageId="5" pageNumber="406">
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<taxonomicName id="4C774D25FFC1FFCECADAB146FE5518CF" ID-CoL="7X63Q" box="[151,466,622,648]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="kimberleyi" status="sp. nov.">
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<emphasis id="B903EAB4FFC1FFCECADAB146FE5518CF" bold="true" box="[151,466,622,648]" italics="true" pageId="5" pageNumber="406">Tyrphonothrus kimberleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057CFFFC1FFCECB97B146FDB018CF" box="[474,567,622,648]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC836A6FFC1FFCECADAB1B9FE8118ED" blockId="5.[151,567,622,682]" box="[151,262,657,682]" pageId="5" pageNumber="406">
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(
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<figureCitation id="134C2A23FFC1FFCECAD2B1B9FF6418ED" box="[159,227,657,682]" captionStart="FIGURE 3" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[166,1424,847,1858]" captionTargetId="figure@5.[165,1426,842,1864]" captionTargetPageId="5" captionText="FIGURE 3. Tyrphonothrus kimberleyi sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="5" pageNumber="406">Fig. 3</figureCitation>
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,
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<figureCitation id="134C2A23FFC1FFCECAA2B1BAFF7918ED" box="[239,254,658,682]" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="5" pageNumber="406">4</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC836A6FFC1FFCECADAB1F1FAD91951" blockId="5.[151,1436,729,790]" pageId="5" pageNumber="406">
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<emphasis id="B903EAB4FFC1FFCECADAB1F1FEA518B5" bold="true" box="[151,290,729,754]" pageId="5" pageNumber="406">Dimensions</emphasis>
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.
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<typeStatus id="54CC8804FFC1FFCECB7FB1F1FE1D18B5" box="[306,410,729,754]" pageId="5" pageNumber="406" type="holotype">Holotype</typeStatus>
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female length 495 Μm, breadth 290 Μm;
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<typeStatus id="54CC8804FFC1FFCEC931B1F2FC6018B5" box="[892,999,730,754]" pageId="5" pageNumber="406" type="paratype">paratypes</typeStatus>
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(
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<emphasis id="B903EAB4FFC1FFCEC9B6B1F2FB8E18B6" box="[1019,1033,730,753]" italics="true" pageId="5" pageNumber="406">n</emphasis>
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= 10) mean length 502 Μm (range 470–542 Μm); mean breadth 282 Μm (range 272–293 Μm). Ratio of prodorsum to total length: 0.3 (
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<typeStatus id="54CC8804FFC1FFCECEA7B1D5FAD51951" box="[1258,1362,765,790]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF08662EFFC1FFCECADAB440FB851D39" box="[151,1026,1896,1918]" httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="5" pageNumber="406" targetBox="[166,1424,847,1858]" targetPageId="5">
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<paragraph id="8BC836A6FFC1FFCECADAB440FB851D39" blockId="5.[151,1026,1896,1918]" box="[151,1026,1896,1918]" pageId="5" pageNumber="406">
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<emphasis id="B903EAB4FFC1FFCECADAB440FE951D3A" bold="true" box="[151,274,1896,1918]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
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<taxonomicName id="4C774D25FFC1FFCECB54B440FD981D39" box="[281,543,1896,1918]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="kimberleyi" status="sp. nov.">
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<emphasis id="B903EAB4FFC1FFCECB54B440FD981D39" box="[281,543,1896,1918]" italics="true" pageId="5" pageNumber="406">Tyrphonothrus kimberleyi</emphasis>
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</taxonomicName>
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<emphasis id="B903EAB4FFC1FFCEC868B440FDF21D3A" bold="true" box="[549,629,1896,1917]" pageId="5" pageNumber="406">
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<taxonomicNameLabel id="A23057CFFFC1FFCEC868B440FDF21D3A" box="[549,629,1896,1917]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype female; a) dorsal; b) ventral.
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</paragraph>
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</caption>
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<subSubSection id="C36D652DFFC1FFCCCA8AB482FAF51E66" lastPageId="7" lastPageNumber="408" pageId="5" pageNumber="406" type="description">
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<paragraph id="8BC836A6FFC1FFCECA8AB482FA1A1DA0" blockId="5.[151,1437,1962,2023]" pageId="5" pageNumber="406">
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<emphasis id="B903EAB4FFC1FFCECA8AB482FE991D84" bold="true" box="[199,286,1962,1987]" pageId="5" pageNumber="406">Female</emphasis>
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.
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<emphasis id="B903EAB4FFC1FFCECB66B482FE2F1D84" box="[299,424,1962,1987]" italics="true" pageId="5" pageNumber="406">Prodorsum</emphasis>
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. Rostrum rounded, smooth, porose (
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<figureCitation id="134C2A23FFC1FFCEC973B482FC071D84" box="[830,896,1962,1987]" captionStart="FIGURE 3" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[166,1424,847,1858]" captionTargetId="figure@5.[165,1426,842,1864]" captionTargetPageId="5" captionText="FIGURE 3. Tyrphonothrus kimberleyi sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="5" pageNumber="406">Fig. 3</figureCitation>
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a). Rostral seta (
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<emphasis id="B903EAB4FFC1FFCECE78B484FBCB1D84" box="[1077,1100,1964,1987]" italics="true" pageId="5" pageNumber="406">ro</emphasis>
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) smooth, curved, flagelliform apically, 42 Μm long, shorter than mutual distance, emerging from alveolus on slight transverse indentation.
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</paragraph>
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<paragraph id="8BC836A6FFC2FFCDCADAB3BFFD141AB0" blockId="6.[151,1436,151,248]" pageId="6" pageNumber="407">
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Lamellar seta (
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<emphasis id="B903EAB4FFC2FFCDCB0DB3BFFED31AF7" box="[320,340,151,176]" italics="true" pageId="6" pageNumber="407">le</emphasis>
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) smooth, curved, flagelliform apically, 60 Μm long, as long as their mutual distance, enclosed by incurved, S-shaped lateral prodorsal ridge. Interlamellar seta (
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<emphasis id="B903EAB4FFC2FFCDC92DB395FCF11A93" box="[864,886,189,212]" italics="true" pageId="6" pageNumber="407">in</emphasis>
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) smooth, thin, extremely long (128 Μm), 7–8 × longer than seta
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<emphasis id="B903EAB4FFC2FFCDCB02B3C9FEEF1ABF" box="[335,360,225,248]" italics="true" pageId="6" pageNumber="407">ex</emphasis>
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. Prodorsum finely porose.
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</paragraph>
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<caption id="DF08662EFFC2FFCDCADAB4AAFE3E1DF0" httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="6" pageNumber="407" targetBox="[160,1426,315,1894]" targetPageId="6">
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<paragraph id="8BC836A6FFC2FFCDCADAB4AAFE3E1DF0" blockId="6.[151,1436,1922,1975]" pageId="6" pageNumber="407">
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<emphasis id="B903EAB4FFC2FFCDCADAB4AAFE951DD0" bold="true" box="[151,274,1922,1944]" pageId="6" pageNumber="407">FIGURE 4.</emphasis>
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<taxonomicName id="4C774D25FFC2FFCDCB55B4AAFD991DDF" box="[280,542,1922,1944]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="kimberleyi" status="sp. nov.">
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<emphasis id="B903EAB4FFC2FFCDCB55B4AAFD991DDF" box="[280,542,1922,1944]" italics="true" pageId="6" pageNumber="407">Tyrphonothrus kimberleyi</emphasis>
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</taxonomicName>
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<emphasis id="B903EAB4FFC2FFCDC869B4ABFDF31DDF" bold="true" box="[548,628,1923,1944]" pageId="6" pageNumber="407">
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<taxonomicNameLabel id="A23057CFFFC2FFCDC869B4ABFDF31DDF" box="[548,628,1923,1944]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV.
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</paragraph>
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</caption>
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<paragraph id="8BC836A6FFC3FFCCCA8AB3B1FE371BD3" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
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<emphasis id="B903EAB4FFC3FFCCCA8AB3B1FEC51AF7" box="[199,322,153,176]" italics="true" pageId="7" pageNumber="408">Notogaster</emphasis>
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. Anterior notogastral margin straight, transverse. Humeral region obtusely angled (
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<figureCitation id="134C2A23FFC3FFCCCEB6B3BFFAB81AF7" box="[1275,1343,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[166,1424,847,1858]" captionTargetId="figure@5.[165,1426,842,1864]" captionTargetPageId="5" captionText="FIGURE 3. Tyrphonothrus kimberleyi sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="7" pageNumber="408">Fig. 3</figureCitation>
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a). With 15 pairs of smooth, thin, setiform setae. Setae
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<emphasis id="B903EAB4FFC3FFCCC8E6B395FD301A93" box="[683,695,189,212]" italics="true" pageId="7" pageNumber="408">e</emphasis>
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2,
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<emphasis id="B903EAB4FFC3FFCCC886B394FD5E1A92" box="[715,729,188,213]" italics="true" pageId="7" pageNumber="408">h</emphasis>
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2 and
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<emphasis id="B903EAB4FFC3FFCCC956B395FCAE1A93" box="[795,809,189,212]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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2 much longer (55–67 Μm long) than others (13–35 Μm long). Setae
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<emphasis id="B903EAB4FFC3FFCCCB6EB3CCFEB61ABC" box="[291,305,228,251]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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3 markedly anterior of
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<emphasis id="B903EAB4FFC3FFCCC861B3CCFDBD1ABC" box="[556,570,228,251]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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2;
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<emphasis id="B903EAB4FFC3FFCCC81CB3CCFDD81ABC" box="[593,607,228,251]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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3 30 Μm long, shorter than distance to
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<emphasis id="B903EAB4FFC3FFCCCE4AB3CCFB921ABC" box="[1031,1045,228,251]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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2;
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<emphasis id="B903EAB4FFC3FFCCCE66B3CCFBBE1ABC" box="[1067,1081,228,251]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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2 on squat tubercle, 55 Μm long. Median longitudinal ridges absent but with vestiges of posterior M-shaped ridge in the form of semicircular indentations anterior of
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<emphasis id="B903EAB4FFC3FFCCCBEBB206FE331B02" box="[422,436,302,325]" italics="true" pageId="7" pageNumber="408">p</emphasis>
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1 and posterolateral to
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<emphasis id="B903EAB4FFC3FFCCC8FEB205FD461B01" box="[691,705,301,326]" italics="true" pageId="7" pageNumber="408">h</emphasis>
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2. Margins narrowing anteriorly, broadest in caudal region. Ratio of length to breadth 1.3. Caudal margin broadly rounded, indented at level of seta
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<emphasis id="B903EAB4FFC3FFCCCE04B27CFBD01B2A" box="[1097,1111,340,365]" italics="true" pageId="7" pageNumber="408">h</emphasis>
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2. Notogaster densely porose interspersed with foveae.
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</paragraph>
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<paragraph id="8BC836A6FFC3FFCCCA8AB288FBF8180F" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
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<emphasis id="B903EAB4FFC3FFCCCA8AB288FED11BF0" box="[199,342,416,439]" italics="true" pageId="7" pageNumber="408">Coxisternum</emphasis>
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. Epimeral setal formula 2-1-3-3. Epimeral setae smooth, setiform, minute (
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<figureCitation id="134C2A23FFC3FFCCCE84B2B7FA881BFF" box="[1225,1295,415,440]" captionStart="FIGURE 3" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[166,1424,847,1858]" captionTargetId="figure@5.[165,1426,842,1864]" captionTargetPageId="5" captionText="FIGURE 3. Tyrphonothrus kimberleyi sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="7" pageNumber="408">Fig. 3</figureCitation>
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b). Setae on epimeral plates
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<date id="FFC91066FFC3FFCCCB04B2ECFE5C1B9C" box="[329,475,451,476]" pageId="7" pageNumber="408" value="2012-01-08" valueMax="2012-02-08" valueMin="2012-01-08">I and II 8–12</date>
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Μm long; setae
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<emphasis id="B903EAB4FFC3FFCCC8C3B2EBFD2D1B9B" box="[654,682,451,476]" italics="true" pageId="7" pageNumber="408">3b</emphasis>
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,
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<emphasis id="B903EAB4FFC3FFCCC8F5B2EBFD551B9C" box="[696,722,451,475]" italics="true" pageId="7" pageNumber="408">3c</emphasis>
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and
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<emphasis id="B903EAB4FFC3FFCCC944B2EBFCA21B9C" box="[777,805,451,475]" italics="true" pageId="7" pageNumber="408">4a</emphasis>
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–
|
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<emphasis id="B903EAB4FFC3FFCCC97EB2ECFCB81B9C" box="[819,831,452,475]" italics="true" pageId="7" pageNumber="408">c</emphasis>
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longer than others (15–20 Μm);
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<emphasis id="B903EAB4FFC3FFCCCEE1B2EBFB4E1B9B" box="[1196,1225,451,476]" italics="true" pageId="7" pageNumber="408">3b</emphasis>
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in lateral position;
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<emphasis id="B903EAB4FFC3FFCCCADAB2CFFF341847" box="[151,179,487,512]" italics="true" pageId="7" pageNumber="408">4b</emphasis>
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in lateral position. Epimeral plates I separated medially, rounded anterolaterally. Epimeral plates II acute posterolaterally, partly fused medially. Epimeral plates III and IV separated medially. Apodeme IV, as far as posterolateral acetabulum, straight, transverse. Epimeral plate IV with blunt lateral spine.
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</paragraph>
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<paragraph id="8BC836A6FFC3FFCCCA8AB17BFBC418B9" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB17BFE0C182C" box="[199,395,595,620]" italics="true" pageId="7" pageNumber="408">Anogenital Region</emphasis>
|
||
. With seven pairs of smooth genital setae,
|
||
<emphasis id="B903EAB4FFC3FFCCC91BB17CFCE4182C" box="[854,867,596,619]" italics="true" pageId="7" pageNumber="408">g</emphasis>
|
||
1-3 subequal in length (12–17 Μm), closely spaced,
|
||
<emphasis id="B903EAB4FFC3FFCCCF31B17CFA0E182C" box="[1404,1417,596,619]" italics="true" pageId="7" pageNumber="408">g</emphasis>
|
||
4-7 longer (23–31 Μm), spaced more broadly apart (
|
||
<figureCitation id="134C2A23FFC3FFCCC8D6B152FD5D18D4" box="[667,730,634,659]" captionStart="FIGURE 3" captionStartId="5.[151,250,1896,1918]" captionTargetBox="[166,1424,847,1858]" captionTargetId="figure@5.[165,1426,842,1864]" captionTargetPageId="5" captionText="FIGURE 3. Tyrphonothrus kimberleyi sp. nov., holotype female; a) dorsal; b) ventral." httpUri="https://zenodo.org/record/223437/files/figure.png" pageId="7" pageNumber="408">Fig. 3</figureCitation>
|
||
b). Genital plates sub-rectangular; each plate 117 Μm long, 55 Μm broad, with well-developed median carina. Posterior margin of genital plate transverse, overlapping adanal plate. Anterior margin of adanal plate transverse, with sclerotised transverse region. Each adanal plate 127 Μm long, 47 Μm broad; three pairs of smooth, setiform setae, 17–27 Μm long. Two pairs of minute anal setae 5 Μm long.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FFCCCA8AB023FAF51E66" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB023FF7E1965" box="[199,249,779,802]" italics="true" pageId="7" pageNumber="408">Legs</emphasis>
|
||
. Pre-tarsi homotridactylous; claws smooth (
|
||
<figureCitation id="134C2A23FFC3FFCCC883B022FC881965" box="[718,783,778,803]" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
). Tarsi long, thin: ratio of lengths to breadths: I 4.1; II 4.1; III 4.3; IV 5.3. Setal formula: Leg
|
||
<date id="FFC91066FFC3FFCCCB90B007FD8A1901" box="[477,525,813,838]" pageId="7" pageNumber="408">I 1-5</date>
|
||
-4(1)-3(1)-11(3); Leg
|
||
<date id="FFC91066FFC3FFCCC8B9B007FCAB1901" box="[756,812,813,838]" pageId="7" pageNumber="408">II 1-5</date>
|
||
-4(1)-4(1)-10(1); Leg
|
||
<date id="FFC91066FFC3FFCCCE58B007FBDB1901" box="[1045,1116,814,838]" pageId="7" pageNumber="408">III 2-2</date>
|
||
-1-2(1)-10; Leg
|
||
<date id="FFC91066FFC3FFCCCF4FB007FAC11901" box="[1282,1350,814,838]" pageId="7" pageNumber="408">IV 1-2</date>
|
||
-1-2-10. Setae of Femur I barbed, setae of other segments on Leg I smooth except
|
||
<emphasis id="B903EAB4FFC3FFCCC9D0B07BFC37192D" box="[925,944,851,874]" italics="true" pageId="7" pageNumber="408">tc</emphasis>
|
||
,
|
||
<emphasis id="B903EAB4FFC3FFCCC9F1B07BFC4D192D" box="[956,970,851,874]" italics="true" pageId="7" pageNumber="408">u</emphasis>
|
||
and
|
||
<emphasis id="B903EAB4FFC3FFCCC9B0B07BFB93192D" box="[1021,1044,851,874]" italics="true" pageId="7" pageNumber="408">pv</emphasis>
|
||
. Proral setae (
|
||
<emphasis id="B903EAB4FFC3FFCCCEE6B07BFB3F192D" box="[1195,1208,851,874]" italics="true" pageId="7" pageNumber="408">p</emphasis>
|
||
) short, broad basally; those on Tarsi I flagelliform apically, others spiniform. Primiventral setae (
|
||
<emphasis id="B903EAB4FFC3FFCCC9E3B05FFC4019C9" box="[942,967,887,910]" italics="true" pageId="7" pageNumber="408">pv</emphasis>
|
||
) on Tarsus I heavily barbed, expanded (
|
||
<figureCitation id="134C2A23FFC3FFCCCF3CB05DFF2119F4" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
b). Solenidia ω1-3 clustered. Famulus (ε) spiniform, peg-like, 10 Μm long. Fastigial setae on Tarsus I heteromorphic: both long, curved but
|
||
<emphasis id="B903EAB4FFC3FFCCCB1FB0E9FEE7199E" box="[338,352,961,985]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
’ thicker and longer than
|
||
<emphasis id="B903EAB4FFC3FFCCC83AB0E9FD02199E" box="[631,645,961,985]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
”. Setae
|
||
<emphasis id="B903EAB4FFC3FFCCC8AEB0E9FD76199E" box="[739,753,961,985]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
” and
|
||
<emphasis id="B903EAB4FFC3FFCCC97BB0E9FCC3199E" box="[822,836,961,985]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
’ on tarsi II similarly shaped, both sparingly barbed,
|
||
<emphasis id="B903EAB4FFC3FFCCCFC8B0E9FA14199E" box="[1413,1427,961,985]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
’ thicker and longer (
|
||
<figureCitation id="134C2A23FFC3FFCCCB24B0CCFE2D19BA" box="[361,426,996,1021]" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
c); those on tarsi III heteromorphic;
|
||
<emphasis id="B903EAB4FFC3FFCCC963B0CDFCBB19BA" box="[814,828,997,1021]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
” long, thin, straight,
|
||
<emphasis id="B903EAB4FFC3FFCCCE6FB0CDFBB719BA" box="[1058,1072,997,1021]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
’ thicker and curved; both smooth (
|
||
<figureCitation id="134C2A23FFC3FFCCCAD2B720FF591E66" box="[159,222,1032,1057]" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
d). Tarsus IV with seta
|
||
<emphasis id="B903EAB4FFC3FFCCCB9CB721FE581E66" box="[465,479,1033,1057]" italics="true" pageId="7" pageNumber="408">ft</emphasis>
|
||
” curved, pointed, smooth; seta
|
||
<emphasis id="B903EAB4FFC3FFCCC961B722FCB01E66" box="[812,823,1034,1057]" italics="true" pageId="7" pageNumber="408">s</emphasis>
|
||
smooth, primiventral setae sparingly barbed (
|
||
<figureCitation id="134C2A23FFC3FFCCCF55B720FADF1E66" box="[1304,1368,1032,1057]" captionStart="FIGURE 4" captionStartId="6.[151,250,1922,1944]" captionTargetBox="[160,1426,315,1894]" captionTargetId="figure@6.[151,1435,301,1901]" captionTargetPageId="6" captionText="FIGURE 4. Tyrphonothrus kimberleyi sp. nov., left legs, left-hand view; a) Leg I; b) detail of primiventral setae on Tarsus I; c) Leg II; d) Leg III; e) Leg IV." httpUri="https://zenodo.org/record/223438/files/figure.png" pageId="7" pageNumber="408">Fig. 4</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC3FFCCCA8AB704FAAE1ECA" pageId="7" pageNumber="408" type="materials_examined">
|
||
<paragraph id="8BC836A6FFC3FFCCCA8AB704FAAE1ECA" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB704FE331E02" bold="true" box="[199,436,1068,1093]" pageId="7" pageNumber="408">Material examined.</emphasis>
|
||
Adult female
|
||
<typeStatus id="54CC8804FFC3FFCCC813B704FD471E02" box="[606,704,1068,1093]" pageId="7" pageNumber="408" type="holotype">holotype</typeStatus>
|
||
and 15
|
||
<typeStatus id="54CC8804FFC3FFCCC969B705FC151E02" box="[804,914,1069,1093]" pageId="7" pageNumber="408" type="paratype">paratypes</typeStatus>
|
||
, moss, Corin Forest, Namadji National Park, Australian Capital Territory,
|
||
<geoCoordinate id="EE435061FFC3FFCCCBA7B778FDEB1E2E" box="[490,620,1104,1129]" direction="south" orientation="latitude" pageId="7" pageNumber="408" precision="15" value="-35.51389">35°30'50"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE435061FFC3FFCCC830B778FC891E2E" box="[637,782,1104,1129]" direction="east" orientation="longitude" pageId="7" pageNumber="408" precision="15" value="148.9189">148°55'08"E</geoCoordinate>
|
||
,
|
||
<quantity id="4C8F9B43FFC3FFCCC952B778FCF81E2F" box="[799,895,1104,1129]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="408" unit="m" value="1250.0">1250 m</quantity>
|
||
., coll. M.J. Colloff,
|
||
<date id="FFC91066FFC3FFCCCE39B778FB681E2E" box="[1140,1263,1104,1129]" pageId="7" pageNumber="408" value="1995-06-21">21.vi.1995</date>
|
||
.
|
||
<typeStatus id="54CC8804FFC3FFCCCEB2B778FAE01E2E" box="[1279,1383,1104,1129]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54CC8804FFC3FFCCCADAB75DFE851ECA" box="[151,258,1141,1165]" pageId="7" pageNumber="408" type="paratype">paratypes</typeStatus>
|
||
deposited in the Australian National Insect Collection, CSIRO Ecosystem Sciences, Canberra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC3FFCCCA8AB7B0FBEB1EF7" box="[199,1132,1176,1201]" pageId="7" pageNumber="408" type="etymology">
|
||
<paragraph id="8BC836A6FFC3FFCCCA8AB7B0FBEB1EF7" blockId="7.[151,1437,151,1636]" box="[199,1132,1176,1201]" pageId="7" pageNumber="408">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB7B0FEC91EF6" bold="true" box="[199,334,1176,1201]" pageId="7" pageNumber="408">Etymology.</emphasis>
|
||
This species is named after my eldest step-son, Kimberley Garth Dent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D652DFFC3FFCCCA8AB794FB881C24" pageId="7" pageNumber="408" type="discussion">
|
||
<paragraph id="8BC836A6FFC3FFCCCA8AB794FA161F2F" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB794FEB31E92" bold="true" box="[199,308,1212,1237]" pageId="7" pageNumber="408">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C774D25FFC3FFCCCB0AB794FDE91E92" box="[327,622,1212,1237]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="kimberleyi" status="sp. nov.">
|
||
<emphasis id="B903EAB4FFC3FFCCCB0AB794FDE91E92" box="[327,622,1212,1237]" italics="true" pageId="7" pageNumber="408">Tyrphonothrus kimberleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23057CFFFC3FFCCC836B794FD5D1E93" box="[635,730,1212,1237]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B903EAB4FFC3FFCCC836B794FD551E92" bold="true" box="[635,722,1212,1237]" pageId="7" pageNumber="408">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from other members of the genus by the following combination of characters: 1) the long, thin interlamellar setae; 2) the straight, transverse anterior notogastral margin; 3) the foveolate notogaster; 4) the vestigial, posterior notogastral ridge in the form of semicircular indentations; 5) the extremely short setae
|
||
<emphasis id="B903EAB4FFC3FFCCC834B602FD001F06" box="[633,647,1322,1345]" italics="true" pageId="7" pageNumber="408">p</emphasis>
|
||
1; 6) the epimeral setal formula 2-1-3-3; 7) the seven pairs of genital setae; 8) the two pairs of anal setae; 9) the sclerotised transverse region on the anterior margin of the adanal plate.
|
||
</paragraph>
|
||
<paragraph id="8BC836A6FFC3FFCCCA8AB65BFB881C24" blockId="7.[151,1437,151,1636]" pageId="7" pageNumber="408">
|
||
<taxonomicName id="4C774D25FFC3FFCCCA8AB65BFE6D1FCB" box="[199,490,1395,1420]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="kimberleyi" status="sp. nov.">
|
||
<emphasis id="B903EAB4FFC3FFCCCA8AB65BFE6D1FCB" box="[199,490,1395,1420]" italics="true" pageId="7" pageNumber="408">Tyrphonothrus kimberleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23057CFFFC3FFCCCBBFB65BFDCA1FCC" box="[498,589,1395,1420]" pageId="7" pageNumber="408" rank="species">
|
||
<emphasis id="B903EAB4FFC3FFCCCBBFB65BFDC11FCB" bold="true" box="[498,582,1395,1420]" pageId="7" pageNumber="408">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
shares with
|
||
<taxonomicName id="4C774D25FFC3FFCCC893B65DFB791FCB" authority="Fain, Lambrechts & Wauthy 1990" authorityName="Fain, Lambrechts & Wauthy" authorityYear="1990" box="[734,1278,1395,1420]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="aquatilis">
|
||
<emphasis id="B903EAB4FFC3FFCCC893B65DFCDA1FCB" box="[734,861,1395,1420]" italics="true" pageId="7" pageNumber="408">T. aquatilis</emphasis>
|
||
(Fain, Lambrechts & Wauthy 1990)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D25FFC3FFCCCF41B65DFEE21FF7" authority="Willmann, 1931" authorityName="Willmann" authorityYear="1931" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B903EAB4FFC3FFCCCF41B65DFA1B1FCB" box="[1292,1436,1395,1420]" italics="true" pageId="7" pageNumber="408">T. foveolatus</emphasis>
|
||
(Willmann, 1931)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D25FFC3FFCCCB3EB6B0FD1A1FF7" authority="Michael, 1888" authorityName="Michael" authorityYear="1888" box="[371,669,1431,1456]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B903EAB4FFC3FFCCCB3EB6B0FE5C1FF7" box="[371,475,1431,1456]" italics="true" pageId="7" pageNumber="408">T. glaber</emphasis>
|
||
(Michael, 1888)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D25FFC3FFCCC897B6B0FC711FF7" authority="Berlese, 1910" authorityName="Berlese" authorityYear="1910" box="[730,1014,1431,1456]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="maior">
|
||
<emphasis id="B903EAB4FFC3FFCCC897B6B0FCBB1FE8" box="[730,828,1432,1455]" italics="true" pageId="7" pageNumber="408">T. maior</emphasis>
|
||
(Berlese, 1910)
|
||
</taxonomicName>
|
||
seven pairs of setae and reduced or vestigial elements of the longitudinal and posterior M-shaped ridges, foveolate notogastral cuticle and setae
|
||
<emphasis id="B903EAB4FFC3FFCCCF2FB695FAFA1F93" box="[1378,1405,1469,1492]" italics="true" pageId="7" pageNumber="408">pv</emphasis>
|
||
of Tarsus I heavily barbed and expanded (expect in
|
||
<taxonomicName id="4C774D25FFC3FFCCC8A8B6C8FCFE1FBF" box="[741,889,1503,1528]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B903EAB4FFC3FFCCC8A8B6C8FCFE1FBF" box="[741,889,1503,1528]" italics="true" pageId="7" pageNumber="408">T. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
for which setae
|
||
<emphasis id="B903EAB4FFC3FFCCCE0AB6C8FBE51FB0" box="[1095,1122,1504,1527]" italics="true" pageId="7" pageNumber="408">pv</emphasis>
|
||
have not been described).
|
||
<taxonomicName id="4C774D25FFC3FFCCCADAB52BFE3F1C5B" box="[151,440,1539,1564]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="kimberleyi">
|
||
<emphasis id="B903EAB4FFC3FFCCCADAB52BFE3F1C5B" box="[151,440,1539,1564]" italics="true" pageId="7" pageNumber="408">Tyrphonothrus kimberleyi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C774D25FFC3FFCCC806B52DFD4F1C5B" box="[587,712,1539,1564]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="aquatilis">
|
||
<emphasis id="B903EAB4FFC3FFCCC806B52DFD4F1C5B" box="[587,712,1539,1564]" italics="true" pageId="7" pageNumber="408">T. aquatilis</emphasis>
|
||
</taxonomicName>
|
||
in that the latter species has the shorter notogastral setae barbed.
|
||
<taxonomicName id="4C774D25FFC3FFCCCADAB50FFE3D1C07" box="[151,442,1575,1600]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="kimberleyi">
|
||
<emphasis id="B903EAB4FFC3FFCCCADAB50FFE3D1C07" box="[151,442,1575,1600]" italics="true" pageId="7" pageNumber="408">Tyrphonothrus kimberleyi</emphasis>
|
||
</taxonomicName>
|
||
has much longer interlamellar and exobothridial setae than
|
||
<taxonomicName id="4C774D25FFC3FFCCCE26B500FB561C07" box="[1131,1233,1575,1600]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="glaber">
|
||
<emphasis id="B903EAB4FFC3FFCCCE26B500FB561C07" box="[1131,1233,1575,1600]" italics="true" pageId="7" pageNumber="408">T. glaber</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D25FFC3FFCCCF46B500FA1C1C07" box="[1291,1435,1575,1600]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="foveolatus">
|
||
<emphasis id="B903EAB4FFC3FFCCCF46B500FA1C1C07" box="[1291,1435,1575,1600]" italics="true" pageId="7" pageNumber="408">T. foveolatus</emphasis>
|
||
</taxonomicName>
|
||
and much longer adanal and anal setae and shorter lamellar setae than
|
||
<taxonomicName id="4C774D25FFC3FFCCC9E6B565FB8F1C23" box="[939,1032,1613,1636]" class="Arachnida" family="Malaconothridae" genus="Tyrphonothrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="maior">
|
||
<emphasis id="B903EAB4FFC3FFCCC9E6B565FB8F1C23" box="[939,1032,1613,1636]" italics="true" pageId="7" pageNumber="408">T. maior</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |