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<document id="07F515BFE4F8EB64BC71B0F25EA07E09" ID-CLB-Dataset="42391" ID-DOI="10.11646/zootaxa.3905.4.1" ID-GBIF-Dataset="1527fd25-cd04-459b-9914-79f4f7c6889a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244811" ID-ZooBank="7F53AA32-73B6-461A-BC93-36B3324E87BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460934395803" checkinUser="plazi" docAuthor="Komai, Tomoyuki" docDate="2015" docId="03DE87A3E2092C7CFF7DFA152AC5438E" docLanguage="en" docName="zt03905p473.pdf" docOrigin="Zootaxa 3905 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lebbeus armatus Owen 1839" docType="treatment" docVersion="9" lastPageNumber="462" masterDocId="FFE7FFDBE2082C77FFEAFFD72E4A4250" masterDocTitle="Reinstatement and redescription of Lebbeus armatus (Owen, 1839), long synonymized with L. groenlandicus (Fabricius, 1775), and description of one new species from the southwestern Sea of Okhotsk, Hokkaido, Japan (Crustacea: Decapoda: Caridea: Thoridae)" masterLastPageNumber="473" masterPageNumber="451" pageNumber="452" updateTime="1698582596382" updateUser="ExternalLinkService">
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<mods:title id="F36772EF6310AC7FAD0E98B22DFD597E">Reinstatement and redescription of Lebbeus armatus (Owen, 1839), long synonymized with L. groenlandicus (Fabricius, 1775), and description of one new species from the southwestern Sea of Okhotsk, Hokkaido, Japan (Crustacea: Decapoda: Caridea: Thoridae)</mods:title>
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<treatment id="03DE87A3E2092C7CFF7DFA152AC5438E" ID-DOI="http://doi.org/10.5281/zenodo.5695485" ID-GBIF-Taxon="119590343" ID-Zenodo-Dep="5695485" LSID="urn:lsid:plazi:treatment:03DE87A3E2092C7CFF7DFA152AC5438E" httpUri="http://treatment.plazi.org/id/03DE87A3E2092C7CFF7DFA152AC5438E" lastPageId="11" lastPageNumber="462" pageId="1" pageNumber="452">
<subSubSection id="C36D653EE2092C76FF7DFA152F4A47AE" pageId="1" pageNumber="452" type="nomenclature">
<paragraph id="8BC836B5E2092C76FF7DFA152C59478D" blockId="1.[151,531,1474,1534]" box="[151,531,1474,1501]" pageId="1" pageNumber="452">
<heading id="D08081D9E2092C76FF7DFA152C59478D" bold="true" box="[151,531,1474,1501]" fontSize="11" level="1" pageId="1" pageNumber="452" reason="1">
<taxonomicName id="4C774D36E2092C76FF7DFA152C59478D" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[151,531,1474,1501]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2092C76FF7DFA152C59478D" bold="true" box="[151,531,1474,1501]" pageId="1" pageNumber="452">
<emphasis id="B903EAA7E2092C76FF7DFA152F2C478C" bold="true" box="[151,358,1474,1500]" italics="true" pageId="1" pageNumber="452">Lebbeus armatus</emphasis>
(
<bibRefCitation id="EFE64B44E2092C76FE9CFA142C41478D" author="Owen" box="[374,523,1475,1501]" pageId="1" pageNumber="452" refString="Owen, R. (1839) Crustacea. In: Richardson, J., Beechey, F. W. &amp; Sowerby, G. B. (Eds.), The Zoology of Captain Beechey's Voyage, Compiled from the Collections and Notes Made by Captain Beechey, the Officers and Naturalist of the Expedition, during a Voyage to the Pacific and Behring's Straits Performed in His Majesty's Ship Blossom, under the Command of Captain F. W. Beechey, in the Years 1825, 26, 27, and 28. London, pp. 77 - 92. [plates. 24 - 28]" type="journal article" year="1839">Owen, 1839</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DFA322F4A47AE" blockId="1.[151,531,1474,1534]" box="[151,256,1509,1534]" pageId="1" pageNumber="452">
<figureCitation id="134C2A30E2092C76FF7DFA322F4A47AE" box="[151,256,1509,1534]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="3.[151,250,1978,2000]" captionStartId-1="4.[151,250,1335,1357]" captionStartId-2="5.[151,250,1743,1765]" captionStartId-3="6.[151,250,1953,1975]" captionStartId-4="7.[151,250,1271,1293]" captionStartId-5="9.[151,250,1110,1132]" captionTargetBox-0="[151,1436,323,1953]" captionTargetBox-1="[161,1429,204,1305]" captionTargetBox-2="[167,1428,207,1681]" captionTargetBox-3="[195,1394,204,1930]" captionTargetBox-4="[161,1424,223,1226]" captionTargetBox-5="[172,1418,182,1090]" captionTargetId-0="figure@3.[151,1436,307,1953]" captionTargetId-1="figure@4.[151,1436,193,1314]" captionTargetId-2="figure@5.[151,1436,193,1694]" captionTargetId-3="figure@6.[178,1408,193,1932]" captionTargetId-4="figure@7.[151,1436,217,1234]" captionTargetId-5="figure@9.[162,1426,175,1095]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="9" captionText-0="FIGURE 1. Lebbeus armatus (Owen, 1839), female (cl 36.7 mm), Sea of Japan off Hokkaido, CBM-ZC 7914. A, entire animal in lateral view; B, dorsum of carapace in lateral view, showing surface setation." captionText-1="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." captionText-2="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." captionText-3="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." captionText-4="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." captionText-5="FIGURE 6. Lebbeus armatus (Owen, 1839), male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C." httpUri-0="https://zenodo.org/record/244812/files/figure.png" httpUri-1="https://zenodo.org/record/244813/files/figure.png" httpUri-2="https://zenodo.org/record/244814/files/figure.png" httpUri-3="https://zenodo.org/record/244815/files/figure.png" httpUri-4="https://zenodo.org/record/244816/files/figure.png" httpUri-5="https://zenodo.org/record/244817/files/figure.png" pageId="1" pageNumber="452">Figs. 1–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE2092C76FF7DF9FA2B4C4582" pageId="1" pageNumber="452" type="reference_group">
<paragraph id="8BC836B5E2092C76FF7DF9FA2C344413" blockId="1.[151,1437,1581,2002]" box="[151,638,1581,1603]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF9FA2C344413" box="[151,638,1581,1603]" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF9FA2FBA4413" authority="Owen, 1839: 88" authorityName="Owen" authorityPageNumber="88" authorityYear="1839" box="[151,496,1581,1603]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="B903EAA7E2092C76FF7DF9FA2F0C4413" box="[151,326,1581,1603]" italics="true" pageId="1" pageNumber="452">Hippolite armata</emphasis>
<treatmentCitation id="0AD610A4E2092C76FEA6F9FA2FBA4413" author="Owen" box="[332,496,1581,1603]" page="88" pageId="1" pageNumber="452" year="1839">
<bibRefCitation id="EFE64B44E2092C76FEA6F9FA2F814413" author="Owen" box="[332,459,1581,1603]" pageId="1" pageNumber="452" refString="Owen, R. (1839) Crustacea. In: Richardson, J., Beechey, F. W. &amp; Sowerby, G. B. (Eds.), The Zoology of Captain Beechey's Voyage, Compiled from the Collections and Notes Made by Captain Beechey, the Officers and Naturalist of the Expedition, during a Voyage to the Pacific and Behring's Straits Performed in His Majesty's Ship Blossom, under the Command of Captain F. W. Beechey, in the Years 1825, 26, 27, and 28. London, pp. 77 - 92. [plates. 24 - 28]" type="journal article" year="1839">Owen, 1839</bibRefCitation>
: 88
</treatmentCitation>
</taxonomicName>
, pl. 27, fig. 2.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DF99B2CC24432" blockId="1.[151,1437,1581,2002]" box="[151,648,1612,1634]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF99B2CC24432" box="[151,648,1612,1634]" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF99B2F124432" box="[151,344,1612,1634]" class="Malacostraca" family="Hippolytidae" genus="Hippolyte" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B903EAA7E2092C76FF7DF99B2F124432" box="[151,344,1612,1634]" italics="true" pageId="1" pageNumber="452">Hippolyte aculeata</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AD610A4E2092C76FE9CF99B2C624432" author="Brandt" box="[374,552,1612,1634]" page="118" pageId="1" pageNumber="452" year="1851">
<bibRefCitation id="EFE64B44E2092C76FE9CF99B2FB34432" author="Brandt" box="[374,505,1612,1634]" pageId="1" pageNumber="452" refString="Brandt, F. (1851) Krebse. In: Middendorff, A. Th. v. (Ed.) Reise in den aussersten Norden und Osten Sibiriens wahrend der Jahre 1843 und 1844 mit allerhochster Genehmigung auf veranstaltung der Kaiserlichen Akademie der Wissenchaften zu St. Petersburg ausgefuhrt und in Verbindung mit vielen Gelehten herausgegeben, 2 (1), 79 - 148. [plates. 5, 6]" type="journal article" year="1851">Brandt 1851</bibRefCitation>
: 118
</treatmentCitation>
(in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DF9BC2FED44CF" blockId="1.[151,1437,1581,2002]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF9BC2FED44CF" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF9BC2F3544D0" box="[151,383,1642,1664]" class="Malacostraca" family="Hippolytidae" genus="Hetairus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis id="B903EAA7E2092C76FF7DF9BC2F3544D0" box="[151,383,1642,1664]" italics="true" pageId="1" pageNumber="452">Hetairus groenlandica</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AD610A4E2092C76FE76F9BD2CC244D0" author="Brashnikov" box="[412,648,1642,1664]" page="155" pageId="1" pageNumber="452" year="1907">
<bibRefCitation id="EFE64B44E2092C76FE76F9BD2C1944D0" author="Brashnikov" box="[412,595,1642,1664]" pageId="1" pageNumber="452" refString="Brashnikov, V. (1907) Material representing the fauna of the Eastern Russian Seas collected by the Schooner &quot; Storosh &quot; in 1899 - 1902. Zapiski Imperatorskoi Akademii Nauk, 20 (8), 1 - 185. [plates 1, 2] [in Russian]" type="journal article" year="1907">Brashnikov 1907</bibRefCitation>
: 155
</treatmentCitation>
, fig. 19.—
<treatmentCitation id="0AD610A4E2092C76FD1DF9BD2A1544D0" author="Derjugin" box="[759,1119,1642,1664]" page="112" pageId="1" pageNumber="452" year="1935">
<bibRefCitation id="EFE64B44E2092C76FD1DF9BD2A6144D0" author="Derjugin" box="[759,1067,1642,1664]" pageId="1" pageNumber="452" refString="Derjugin, K. M. &amp; Kobjakova, Z. I. (1935) Zur Dekapoden fauna des Japanischen Meeres. Zoologischer Anzeiger, 112, 141 - 147." type="journal article" year="1935">Derjugin &amp; Kobjakova 1935</bibRefCitation>
: 112
</treatmentCitation>
, 142.—
<treatmentCitation id="0AD610A4E2092C76FB44F9BD2BD244D0" author="Kobjakova" box="[1198,1432,1642,1664]" page="222" pageId="1" pageNumber="452" year="1936">
<bibRefCitation id="EFE64B44E2092C76FB44F9BD2B2B44D0" author="Kobjakova" box="[1198,1377,1642,1664]" pageId="1" pageNumber="452" refString="Kobjakova, Z. I. (1936) Zoogeographical review of the Decapoda fauna from the Okhotsk and Japanese Seas. Transactions of the Natural Society of Leningrad, 65, 185 - 228. [in Russian]" type="journal article" year="1936">Kobjakova 1936</bibRefCitation>
: 222
</treatmentCitation>
; 1937: 108; 1958: 227.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DF97E2DA344EE" blockId="1.[151,1437,1581,2002]" box="[151,1001,1704,1726]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF97E2DA344EE" box="[151,1001,1704,1726]" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF97E2FE644EE" box="[151,428,1704,1726]" class="Malacostraca" family="Hippolytidae" genus="Spirontocaris" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis id="B903EAA7E2092C76FF7DF97E2FE644EE" box="[151,428,1704,1726]" italics="true" pageId="1" pageNumber="452">Spirontocaris groenlandica</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AD610A4E2092C76FE21F97F2C2B44EE" author="Balss" box="[459,609,1704,1726]" page="45" pageId="1" pageNumber="452" year="1914">
<bibRefCitation id="EFE64B44E2092C76FE21F97F2C7544EE" author="Balss" box="[459,575,1704,1726]" pageId="1" pageNumber="452" refString="Balss, H. (1914) Ostasiatische Decapoden II. Die Natantia und Reptantia. Abhandlungen der Mathematisch-Physikalischen Klasse der Koniglich Baierischen Akademie der Wissenschaften, 10 (Supplement 2), 1 - 101. [plate. 1]" type="journal article" year="1914">Balss 1914</bibRefCitation>
: 45
</treatmentCitation>
.—
<treatmentCitation id="0AD610A4E2092C76FD94F97F2D6444EE" author="Yokoya" box="[638,814,1704,1726]" page="24" pageId="1" pageNumber="452" year="1933">
<bibRefCitation id="EFE64B44E2092C76FD94F97F2D4144EE" author="Yokoya" box="[638,779,1704,1726]" pageId="1" pageNumber="452" refString="Yokoya, Y. (1933) On the distribution of decapod Crustacea inhabiting the continental shelf around Japan, chiefly based upon the materials collected by S. S. &quot; Soyo Maru &quot; during the years 1923 - 1930. Journal of the College of Agriculture, Tokyo Imperial University, 12, 1 - 236." type="journal article" year="1933">Yokoya 1933</bibRefCitation>
: 24
</treatmentCitation>
.—
<treatmentCitation id="0AD610A4E2092C76FCA6F97F2DAF44EE" author="Urita" box="[844,997,1704,1726]" page="16" pageId="1" pageNumber="452" year="1942">
<bibRefCitation id="EFE64B44E2092C76FCA6F97F2DF544EE" author="Urita" box="[844,959,1704,1726]" pageId="1" pageNumber="452" refString="Urita, T. (1942) Decapod crustaceans from Saghalien, Japan. Bulletin of the Biogeographical Society of Japan, 12, 1 - 78." type="journal article" year="1942">Urita 1942</bibRefCitation>
: 16
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DF9112EBE44AB" blockId="1.[151,1437,1581,2002]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF9112EBE44AB" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF9112F32448C" box="[151,376,1734,1756]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis id="B903EAA7E2092C76FF7DF9112F32448C" box="[151,376,1734,1756]" italics="true" pageId="1" pageNumber="452">Lebbeus groenlandica</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AD610A4E2092C76FE7FF9112C37448C" author="Vinogradov" box="[405,637,1734,1756]" page="203" pageId="1" pageNumber="452" year="1950">
<bibRefCitation id="EFE64B44E2092C76FE7FF9112C01448C" author="Vinogradov" box="[405,587,1734,1756]" pageId="1" pageNumber="452" refString="Vinogradov, L. G. (1950) Classification of shrimps, prawns and crabs from Far East. Izvestia TINRO, 33, 179 - 358. [plates 1 - 53] [in Russian]" type="journal article" year="1950">Vinogradov 1950</bibRefCitation>
: 203
</treatmentCitation>
, pl. 14, fig. 53.—
<treatmentCitation id="0AD610A4E2092C76FCC4F9112D9C448C" author="Igarashi" box="[814,982,1734,1756]" page="5" pageId="1" pageNumber="452" year="1969">
<bibRefCitation id="EFE64B44E2092C76FCC4F9112DF4448C" author="Igarashi" box="[814,958,1734,1756]" pageId="1" pageNumber="452" refString="Igarashi, T. (1969) A list of marine decapod crustaceans from Hokkaido, deposited at the Fisheries Museum, Faculty of Fisheries, Hokkaido University I. Macrura. Contribution from the Fisheries Museum, Faculty of Fisheries, Hokkaido University, 11, 1 - 15. [plates. 1 - 20]" type="journal article" year="1969">Igarashi 1969</bibRefCitation>
: 5
</treatmentCitation>
, pl. 5, fig. 15.—
<treatmentCitation id="0AD610A4E2092C76FB93F9112EBA44AB" author="Adrianov" page="264" pageId="1" pageNumber="452" year="1998">
<bibRefCitation id="EFE64B44E2092C76FB93F9112BD2448C" author="Adrianov" box="[1145,1432,1734,1756]" pageId="1" pageNumber="452" refString="Adrianov, A. V. &amp; Kussakin, O. G. (1998) A Check List of Biota of the Peter the Great Bay, the Sea of Japan. Dalnauka, Vladivostok, 349 pp. [in Russian]" type="book" year="1998">Adrianov &amp; Kussakin 1998</bibRefCitation>
: 264
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC836B5E2092C76FF7DF8D32B4C4582" blockId="1.[151,1437,1581,2002]" pageId="1" pageNumber="452">
<treatmentCitationGroup id="AB67119BE2092C76FF7DF8D32B4C4582" pageId="1" pageNumber="452">
<taxonomicName id="4C774D36E2092C76FF7DF8D32FCB454A" box="[151,385,1796,1818]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2092C76FF7DF8D32FCB454A" box="[151,385,1796,1818]" italics="true" pageId="1" pageNumber="452">Lebbeus groenlandicus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AD610A4E2092C76FE75F8D32C03454A" author="Holthuis" box="[415,585,1796,1818]" page="9" pageId="1" pageNumber="452" year="1947">
<bibRefCitation id="EFE64B44E2092C76FE75F8D32C79454A" author="Holthuis" box="[415,563,1796,1818]" pageId="1" pageNumber="452" refString="Holthuis, L. B. (1947) The Hippolytidae and Rhynchocinetidae collected by the Siboga and Snellius Expeditions with remarks on other species. Siboga Expeditie, 39 a 8, 1 - 100." type="journal article" year="1947">Holthuis 1947</bibRefCitation>
: 9
</treatmentCitation>
(in part).—
<treatmentCitation id="0AD610A4E2092C76FD55F8D32D2C454A" author="Kubo" box="[703,870,1796,1818]" page="105" pageId="1" pageNumber="452" year="1960">
<bibRefCitation id="EFE64B44E2092C76FD55F8D32D7F454A" author="Kubo" box="[703,821,1796,1818]" pageId="1" pageNumber="452" refString="Kubo, I. (1960) Natantia. In: Okada, Y. K., Uchida, T. et al. (Eds.), Encyclopedia Zoologica Illustrated in Colors, Part IV. Hokuryukan Publishing Co. Ltd., Tokyo, pp. 98 - 113. [in Japanese]" type="book chapter" year="1960">
<collectingCountry id="F3607625E2092C76FD55F8D32CB2454A" box="[703,760,1796,1818]" name="Cuba" pageId="1" pageNumber="452">Kubo</collectingCountry>
1960
</bibRefCitation>
: 105
</treatmentCitation>
, pl. 52, fig. 1; 1965: 616, fig. 978.—
<treatmentCitation id="0AD610A4E2092C76FB3FF8D32EA54568" author="Miyake" page="123" pageId="1" pageNumber="452" year="1962">
<bibRefCitation id="EFE64B44E2092C76FB3FF8D32BD2454A" author="Miyake" box="[1237,1432,1796,1818]" pageId="1" pageNumber="452" refString="Miyake, S., Sakai, K. &amp; Nishikawa, S. (1962) A fauna-list of the decapod Crustacea from the coasts washed by the Tsushima Warm Current. Records of Oceanographic Works in Japan, 6, 121 - 131." type="journal article" year="1962">
Miyake
<emphasis id="B903EAA7E2092C76FAC0F8D22B17454A" box="[1322,1373,1796,1818]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
1962
</bibRefCitation>
: 123
</treatmentCitation>
.—
<treatmentCitation id="0AD610A4E2092C76FEFAF8F52C7F4568" author="Kim" box="[272,565,1826,1848]" page="203" pageId="1" pageNumber="452" year="1972">
<bibRefCitation id="EFE64B44E2092C76FEFAF8F52C4B4568" author="Kim" box="[272,513,1826,1848]" pageId="1" pageNumber="452" refString="Kim, H. S. &amp; Park, K. B. (1972) Faunal studies on the macrurans in Korea. In: Ministry of Science and Technology (Ed.), Floral Studies on Some Taxa of Plants and Faunal Studies on Some Taxa of Animals in Korea. Ministry of Science and Technology, Seoul, pp. 185 - 222. [plates 1 - 6] [in Korean]" type="book chapter" year="1972">H.S. Kim &amp; Park 1972</bibRefCitation>
: 203
</treatmentCitation>
, pl. 4, fig. 6.—
<treatmentCitation id="0AD610A4E2092C76FD3FF8F52DCB4568" author="Motoh" box="[725,897,1826,1848]" page="31" pageId="1" pageNumber="452" year="1972">
<bibRefCitation id="EFE64B44E2092C76FD3FF8F52D104568" author="Motoh" box="[725,858,1826,1848]" pageId="1" pageNumber="452" refString="Motoh, H. (1972) A faunal list of the macruran Decapoda from Nanao Bay, Ishikawa Prefecture, middle Japan. Bulletin of the Ishikawa Prefectural Marine Culture Station, 2, 29 - 83." type="journal article" year="1972">Motoh 1972</bibRefCitation>
: 31
</treatmentCitation>
, 42, pl. 9, figs. 1, 2; 2008: 18.—
<treatmentCitation id="0AD610A4E2092C76FB30F8F52BDD4568" author="Hayashi" box="[1242,1431,1826,1848]" page="17" pageId="1" pageNumber="452" year="1976">
<bibRefCitation id="EFE64B44E2092C76FB30F8F52B254568" author="Hayashi" box="[1242,1391,1826,1848]" pageId="1" pageNumber="452" refString="Hayashi, K. (1976) Review of shrimps from the Sado Island and its neighbourhood. Niigataken Seibutsu Kyoikukaishi, 11, 12 - 22. [in Japanese with English summary]" type="journal article" year="1976">Hayashi 1976</bibRefCitation>
: 17
</treatmentCitation>
; 1992a: 120; 1992b: 344 (key), fig. 232; 1992c: 439, figs. 233f,
<specimenCount id="9D71FD3CE2092C76FCB5F8962DDD4507" box="[863,919,1857,1879]" pageId="1" pageNumber="452" type="generic">234f.</specimenCount>
—
<treatmentCitation id="0AD610A4E2092C76FC59F8962AC94507" author="Kim" box="[947,1155,1857,1879]" page="143" pageId="1" pageNumber="452" year="1976">
<bibRefCitation id="EFE64B44E2092C76FC59F8962A044507" author="Kim" box="[947,1102,1857,1879]" pageId="1" pageNumber="452" refString="Kim, H. S. (1976) A checklist of Macrura (Crustacea, Decapoda) of Korea. Proceedings of the College of Natural Sciences, Seoul National University, 1, 131 - 152." type="journal article" year="1976">H.S. Kim 1976</bibRefCitation>
: 143
</treatmentCitation>
; 1977: 274, pl. 25, 26, fig. 53a–c, textfigs. 113, 116; 1985: 67.—
<treatmentCitation id="0AD610A4E2092C76FDA8F8B72CBE4526" author="Miyake" box="[578,756,1888,1910]" page="53" pageId="1" pageNumber="452" year="1982">
<bibRefCitation id="EFE64B44E2092C76FDA8F8B72C854526" author="Miyake" box="[578,719,1888,1910]" pageId="1" pageNumber="452" refString="Miyake, S. (1982) Japanese Crustacean Decapods and Stomatopods in Color, Vol. I. Hoikusha Publishing Co., Ltd., Osaka, vii + 261 pp. [56 plates] [in Japanese]" type="book" year="1982">Miyake 1982</bibRefCitation>
: 53
</treatmentCitation>
, pl. 18, fig. 3.—
<treatmentCitation id="0AD610A4E2092C76FC7FF8B72A0E4526" author="Takeda" box="[917,1092,1888,1910]" page="25" pageId="1" pageNumber="452" year="1982">
<bibRefCitation id="EFE64B44E2092C76FC7FF8B72A554526" author="Takeda" box="[917,1055,1888,1910]" pageId="1" pageNumber="452" refString="Takeda, M. (1982) Key to the Japanese and Foreign Crustaceans Fully Illustrated in Colors. Hokuryukan, Tokyo, vi + 58 pp. [(key) + 284 pp]" type="book" year="1982">Takeda 1982</bibRefCitation>
: 25
</treatmentCitation>
, fig. 73.—
<treatmentCitation id="0AD610A4E2092C76FB44F8B72B1D4526" author="Komai" box="[1198,1367,1888,1910]" page="72" pageId="1" pageNumber="452" year="1991">
<bibRefCitation id="EFE64B44E2092C76FB44F8B72B784526" author="Komai" box="[1198,1330,1888,1910]" pageId="1" pageNumber="452" refString="Komai, T. (1991) Deep-sea decapod crustaceans from the Pacific coast of eastern Hokkaido, northern Japan (Crustacea, Decapoda, Penaeidea and Caridea). Report of North Japan Sub-Commitee for Bottom Fish, Research Committee for Fishery Resources, 24, 55 - 96." type="journal article" year="1991">Komai 1991</bibRefCitation>
: 72
</treatmentCitation>
; 1994: 82; 2008: 257, fig. 21.16.—
<treatmentCitation id="0AD610A4E2092C76FE18F8A92CB845C4" author="Komai" box="[498,754,1918,1941]" page="193" pageId="1" pageNumber="452" year="1992">
<bibRefCitation id="EFE64B44E2092C76FE18F8A92CF145C4" author="Komai" box="[498,699,1918,1941]" pageId="1" pageNumber="452" refString="Komai, T., Maruyama, S. &amp; Konishi, K. (1992) A list of decapod crustaceans from Hokkaido, northern Japan. Researches on Crustacea, 21, 189 - 205." type="journal article" year="1992">
Komai
<emphasis id="B903EAA7E2092C76FDA9F8572C3145C4" box="[579,635,1918,1941]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
1992
</bibRefCitation>
: 193
</treatmentCitation>
; 2004: 122 (table 1).—
<treatmentCitation id="0AD610A4E2092C76FC03F8A92ADC45C4" author="Chace" box="[1001,1174,1918,1940]" page="51" pageId="1" pageNumber="452" year="1997">
<bibRefCitation id="EFE64B44E2092C76FC03F8A92A2545C4" author="Chace" box="[1001,1135,1918,1940]" pageId="1" pageNumber="452" refString="Chace, F. A., Jr. (1997) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 7: Families Atyidae, Eugonatonotidae, Rhynchocinetidae, Bathypalaemonellidae, Processidae, and Hippolytidae. Smithsonian Contributions to Zoology, 587, (i - v), 1 - 106." type="journal article" year="1997">Chace 1997</bibRefCitation>
: 51
</treatmentCitation>
.—
<treatmentCitation id="0AD610A4E2092C76FB5EF8A92BD245C4" author="Cha" box="[1204,1432,1918,1941]" page="112" pageId="1" pageNumber="452" year="2001">
<bibRefCitation id="EFE64B44E2092C76FB5EF8A92B2845C4" author="Cha" box="[1204,1378,1918,1941]" pageId="1" pageNumber="452" refString="Cha, H. K., Lee, J. U., Park, C. S., Baik, C. I., Hong, S. Y., Park, J. H., Lee, D. W., Choi, Y. M., Hwang, K., Kim, Z. G., Choi, K. H., Sohn, H., Sohn, M. H., Kim, D. H. &amp; Choi, J. H. (2001) Shrimps of the Korean Waters. National Fisheries Research and Development Institute, Pusan, 188 pp. [in Korean]" type="book" year="2001">
Cha
<emphasis id="B903EAA7E2092C76FB00F8572B6845C4" box="[1258,1314,1918,1941]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
2001
</bibRefCitation>
: 112
</treatmentCitation>
, unnumbered fig.—
<treatmentCitation id="0AD610A4E2092C76FE62F84A2C1A45E3" author="Sokolov" box="[392,592,1949,1971]" page="109" pageId="1" pageNumber="452" year="2001">
<bibRefCitation id="EFE64B44E2092C76FE62F84A2C5645E3" author="Sokolov" box="[392,540,1949,1971]" pageId="1" pageNumber="452" refString="Sokolov, V. I. (2001) Decapod crustaceans of the southwest Kamchatka Shelf: R / V &quot; Professor Levanidov &quot; collection in June 1996. Arthropoda Selecta, 10, 103 - 136." type="book chapter" year="2001">Sokolov 2001</bibRefCitation>
: 109
</treatmentCitation>
, fig. 7.—
<treatmentCitation id="0AD610A4E2092C76FD5AF84A2D8C45E3" author="Motoh" box="[688,966,1949,1971]" page="34" pageId="1" pageNumber="452" year="2005">
<bibRefCitation id="EFE64B44E2092C76FD5AF84A2DD545E3" author="Motoh" box="[688,927,1949,1971]" pageId="1" pageNumber="452" refString="Motoh, H. &amp; Toyota, K. (2005) Macruran Decapoda from Kyoto Prefecture, Sea of Japan. Report of the Noto Marine Center, 11, 31 - 42. [in Japanese with English abstract]" type="journal article" year="2005">Motoh &amp; Toyota 2005</bibRefCitation>
: 34
</treatmentCitation>
, fig. 3-10.—
<treatmentCitation id="0AD610A4E2092C76FBACF84A2BD245E3" author="Motoh" box="[1094,1432,1949,1971]" page="216" pageId="1" pageNumber="452" year="2009">
<bibRefCitation id="EFE64B44E2092C76FBACF84A2B2E45E3" author="Motoh" box="[1094,1380,1949,1971]" pageId="1" pageNumber="452" refString="Motoh, H. &amp; Yamaguchi, K. (2009) Macruran decapods from the Sea of Japan - II. Caridean shrimps caught with baited shrimp traps off Togi, Ishikawa Prefecture. Bulletin of the Hoshizaki Green Foundation, 12, 213 - 220. [in Japanese with English abstract]" type="journal article" year="2009">Motoh &amp; Yamaguchi 2009</bibRefCitation>
: 216
</treatmentCitation>
, fig. 3E.—
<treatmentCitation id="0AD610A4E2092C76FEC0F86B2C234582" author="De" box="[298,617,1980,2002]" page="425" pageId="1" pageNumber="452" year="2011">
<bibRefCitation id="EFE64B44E2092C76FEC0F86B2C734582" author="De" box="[298,569,1980,2002]" pageId="1" pageNumber="452" refString="De Grave, S. &amp; Fransen, C. H. J. M. (2011) Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps. Zoologische Mededelingen, 85, 195 - 588." type="journal article" year="2011">
<collectingCountry id="F3607625E2092C76FEC0F86B2F034582" box="[298,329,1980,2002]" name="South Georgia and the South Sandwich Islands" pageId="1" pageNumber="452">De</collectingCountry>
Grave &amp; Fransen 2011
</bibRefCitation>
: 425
</treatmentCitation>
(part).—
<treatmentCitation id="0AD610A4E2092C76FD21F86B2DCE4582" author="Kim" box="[715,900,1980,2002]" page="37" pageId="1" pageNumber="452" year="2012">
<bibRefCitation id="EFE64B44E2092C76FD21F86B2D2A4582" author="Kim" box="[715,864,1980,2002]" pageId="1" pageNumber="452" refString="Kim, J. N. (2012) Arthropoda: Malacostraca; Decapoda: Palaemonidae, Hippolytidae, Processidae, Pandalidae. Shrimps II. Invertebrate Fauna of Korea, Volume 21, Number 21. National Institute of Biological Resources, Ministry of Environment, Incheon, 98 pp." type="book" year="2012">J.N. Kim 2012</bibRefCitation>
: 37
</treatmentCitation>
, fig. 16, pl. 12.—
<treatmentCitation id="0AD610A4E2092C76FBD9F86B2B484582" author="Komatsu" box="[1075,1282,1980,2002]" page="190" pageId="1" pageNumber="452" year="2014">
<bibRefCitation id="EFE64B44E2092C76FBD9F86B2A9A4582" author="Komatsu" box="[1075,1232,1980,2002]" pageId="1" pageNumber="452" refString="Komatsu, H. (2014) Deep-sea decapod crustaceans from off the Japanese coast of the Sea of Japan. In: Fujita, T. (Ed.) Deep-sea Fauna of the Sea of Japan. National Museum of Nature and Science Monographs, 44, 177 - 203." type="journal article" year="2014">Komatsu 2014</bibRefCitation>
: 190
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE20A2C75FF7DFF402B4541ED" pageId="2" pageNumber="453" type="materials_examined">
<paragraph id="8BC836B5E20A2C75FF7DFF402B4541ED" blockId="2.[151,1437,151,2026]" pageId="2" pageNumber="453">
<emphasis id="B903EAA7E20A2C75FF7DFF402F3342E0" bold="true" box="[151,377,151,176]" pageId="2" pageNumber="453">Material examined</emphasis>
. Sea of Okhotsk: off Kitami Monbetsu, Hokkaido,
<geoCoordinate id="EE435072E20A2C75FC51FF4F2A5542E0" box="[955,1055,152,176]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="925" value="44.8">44°48’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FBC0FF4F2ADC42E0" box="[1066,1174,152,176]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="925" value="143.35">143°21’E</geoCoordinate>
to
<geoCoordinate id="EE435072E20A2C75FB50FF4F2B5442E0" box="[1210,1310,152,176]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="925" value="44.733334">44°44’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FAC0FF402BD242E0" box="[1322,1432,151,176]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="925" value="143.43333">143°26’E</geoCoordinate>
, 142–
<quantity id="4C8F9B50E20A2C75FF3AFF6B2F534283" box="[208,281,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="453" unit="m" value="138.0">138 m</quantity>
,
<date id="FFC91075E20A2C75FECFFF6B2C4B4284" box="[293,513,187,212]" pageId="2" pageNumber="453" value="1991-11-17">17 November 1991</date>
, RV “Hakuyo-maru”, trawl,
<specimenCount id="9D71FD3CE20A2C75FCACFF6B2DF94284" box="[838,947,187,212]" pageId="2" pageNumber="453" type="female">2 females</specimenCount>
(cl 32.0, ca. 32.0 mm), 1 ovigerous female (cl
<quantity id="4C8F9B50E20A2C75FF57FF092F6C42A7" box="[189,294,222,247]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="2" pageNumber="453" unit="mm" value="35.2">35.2 mm</quantity>
), NSMT-Cr 1585; off Kitami Monbetsu,
<geoCoordinate id="EE435072E20A2C75FD13FF092DCA42A7" box="[761,896,222,247]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="44.891335">44°53.48’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FC67FF082A5442A7" box="[909,1054,223,247]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="143.37134">143°22.28’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FBC1FF082AE742A7" box="[1067,1197,223,247]" metricMagnitude="2" metricUnit="m" metricValue="1.46" metricValueMax="1.48" metricValueMin="1.44" pageId="2" pageNumber="453" unit="m" value="146.0" valueMax="148.0" valueMin="144.0">144–148 m</quantity>
,
<date id="FFC91075E20A2C75FB50FF092BDD42A7" box="[1210,1431,222,247]" pageId="2" pageNumber="453" value="1991-11-16">16 November 1991</date>
, RV “Hakuyo-maru”, trawl,
<specimenCount id="9D71FD3CE20A2C75FE27FED42C53434B" box="[461,537,258,283]" pageId="2" pageNumber="453" type="male">1 male</specimenCount>
(cl 27.0 mm), NSMT-Cr 1598; off Kitami Monbetsu,
<geoCoordinate id="EE435072E20A2C75FB91FED42AB9434B" box="[1147,1267,259,283]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="92" value="44.81167">44°48.7’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FB14FED52B35434B" box="[1278,1407,258,283]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="92" value="143.60834">143°36.5’E</geoCoordinate>
to
<geoCoordinate id="EE435072E20A2C75FF7DFEF12F45436E" box="[151,271,294,318]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="92" value="44.746666">44°44.8’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FEF6FEF12FD5436E" box="[284,415,294,318]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="92" value="143.70667">143°42.4’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FE46FEF22C65436E" box="[428,559,293,318]" metricMagnitude="2" metricUnit="m" metricValue="1.545" metricValueMax="1.56" metricValueMin="1.53" pageId="2" pageNumber="453" unit="m" value="154.5" valueMax="156.0" valueMin="153.0">153–156 m</quantity>
,
<date id="FFC91075E20A2C75FDD7FEF22D50436E" box="[573,794,293,318]" pageId="2" pageNumber="453" value="1991-11-16">16 November 1991</date>
, RV “Hakuyo-maru”, trawl, 1 ovigerous female (cl
<quantity id="4C8F9B50E20A2C75FA81FEF12E8D4331" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="2" pageNumber="453" unit="mm" value="29.3">29.3 mm</quantity>
), NSMT-Cr 1609. Pacific side of Hokkaido: off Kushiro,
<date id="FFC91075E20A2C75FC98FE9E2A544331" box="[882,1054,329,354]" pageId="2" pageNumber="453" value="1961-03-01">March 1961, 1</date>
ovigerous female (cl 28.0 mm), NSMT-Cr 1692. Sanriku: off Miyako, Iwate Prefecture, depth not recorded,
<date id="FFC91075E20A2C75FC12FEBB2AE443D5" box="[1016,1198,364,389]" pageId="2" pageNumber="453" value="1982-12">December 1982</date>
, commercial trawler, coll. T. Komai,
<specimenCount id="9D71FD3CE20A2C75FEA2FE472FE043F9" box="[328,426,400,425]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FE3DFE472C0A43F8" box="[471,576,400,424]" metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" pageId="2" pageNumber="453" unit="mm" value="17.4">17.4 mm</quantity>
), HUMZ-C 00023; same locality,
<date id="FFC91075E20A2C75FC2BFE462A1143F8" box="[961,1115,400,425]" pageId="2" pageNumber="453" value="1987-01">January 1987</date>
, commercial trawler,
<specimenCount id="9D71FD3CE20A2C75FABAFE472BD643F9" box="[1360,1436,400,425]" pageId="2" pageNumber="453" type="male">1 male</specimenCount>
(cl 15.0 mm), HUMZ-C; Sea of
<collectingCountry id="F3607625E20A2C75FDE3FE632C07439B" box="[521,589,436,459]" name="Japan" pageId="2" pageNumber="453">Japan</collectingCountry>
: Peter the Great Bay, off Vladivostok, depth not recorded, trawl, coll.
<collectingCountry id="F3607625E20A2C75FAB4FE632B38439B" box="[1374,1394,436,459]" name="Italy" pageId="2" pageNumber="453">Y</collectingCountry>
. M. Yakovlev,
<specimenCount id="9D71FD3CE20A2C75FEF9FE002F2943A0" box="[275,355,471,496]" pageId="2" pageNumber="453" type="male">1 male</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FE70FE002C4243BF" box="[410,520,471,495]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="2" pageNumber="453" unit="mm" value="14.4">14.4 mm</quantity>
),
<specimenCount id="9D71FD3CE20A2C75FDCAFE002CD843A0" box="[544,658,471,496]" pageId="2" pageNumber="453" type="female">2 females</specimenCount>
(cl 24.0,
<quantity id="4C8F9B50E20A2C75FCE7FE002D3143BF" box="[781,891,471,495]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="2" pageNumber="453" unit="mm" value="32.3">32.3 mm</quantity>
), CBM-ZC 3821; Musashi Bank,
<geoCoordinate id="EE435072E20A2C75FAF8FE002BD243BF" box="[1298,1432,471,495]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="44.674667">44°40.48’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FF7DFE2C2F6F4043" box="[151,293,507,531]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="140.03967">140°02.38’E</geoCoordinate>
to
<geoCoordinate id="EE435072E20A2C75FEA1FE2D2F9B4043" box="[331,465,506,531]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="44.658833">44°39.53’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FE37FE2C2C254043" box="[477,623,507,531]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="140.04716">140°02.83’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FD91FE2D2CB64042" box="[635,764,506,531]" metricMagnitude="2" metricUnit="m" metricValue="2.02" metricValueMax="2.06" metricValueMin="1.98" pageId="2" pageNumber="453" unit="m" value="202.0" valueMax="206.0" valueMin="198.0">198–206 m</quantity>
,
<date id="FFC91075E20A2C75FCE3FE2C2DEB4043" box="[777,929,507,531]" pageId="2" pageNumber="453" value="2011-05-28">28 May 2011</date>
, RV “Tansei-maru”, KT 11-9 cruise, stn M2, beam trawl with
<quantity id="4C8F9B50E20A2C75FEBCFDC92FC84066" box="[342,386,542,566]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="453" unit="m" value="3.0">3 m</quantity>
span opening,
<specimenCount id="9D71FD3CE20A2C75FDC4FDC92CDA4066" box="[558,656,541,566]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FD57FDC92D524066" box="[701,792,542,566]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="2" pageNumber="453" unit="mm" value="7.8">7.8 mm</quantity>
), NSMT-Cr 22577; off Otaru, Hokkaido, ca.
<quantity id="4C8F9B50E20A2C75FAFFFDCA2BDD4066" box="[1301,1431,541,566]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="453" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150–200 m</quantity>
,
<date id="FFC91075E20A2C75FF7DFD952F71400A" box="[151,315,578,602]" pageId="2" pageNumber="453" value="2003-06-21">21 June 2003</date>
, commercial trap, coll.
<collectingCountry id="F3607625E20A2C75FDB6FD952C3A4009" box="[604,624,578,601]" name="Italy" pageId="2" pageNumber="453">Y</collectingCountry>
. Matsuzawa, 1 ovigerous female (cl
<quantity id="4C8F9B50E20A2C75FBC5FD952AD7400A" box="[1071,1181,578,602]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="2" pageNumber="453" unit="mm" value="28.4">28.4 mm</quantity>
), CBM-ZC 7356; off Hokkaido,
<quantity id="4C8F9B50E20A2C75FEFEFDB22FDC402D" box="[276,406,613,638]" metricMagnitude="2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="2" pageNumber="453" unit="m" value="225.0" valueMax="250.0" valueMin="200.0">200–250 m</quantity>
,
<date id="FFC91075E20A2C75FE49FDB22C68402D" box="[419,546,613,638]" pageId="2" pageNumber="453" value="2004-04">April 2004</date>
, commercial trap, coll.
<collectingCountry id="F3607625E20A2C75FCC4FDB12D08402D" box="[814,834,614,637]" name="Italy" pageId="2" pageNumber="453">Y</collectingCountry>
. Matsuzawa,
<specimenCount id="9D71FD3CE20A2C75FC36FDB22A74402E" box="[988,1086,613,638]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FB86FDB22A9F402D" box="[1132,1237,613,638]" metricMagnitude="-2" metricUnit="m" metricValue="3.67" pageId="2" pageNumber="453" unit="mm" value="36.7">36.7 mm</quantity>
), CBM-ZC 7914; North
<collectingCountry id="F3607625E20A2C75FF0AFD5E2F7C40F0" box="[224,310,649,672]" name="Japan" pageId="2" pageNumber="453">Yamato</collectingCountry>
Bank,
<geoCoordinate id="EE435072E20A2C75FE6CFD5E2FB440F1" box="[390,510,649,673]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="92" value="43.383335">43°23.0’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FDE1FD5F2CC440F1" box="[523,654,648,673]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="92" value="135.08333">135°05.0’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FD70FD5F2D5640F0" box="[666,796,648,673]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="453" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150–200 m</quantity>
,
<date id="FFC91075E20A2C75FCC2FD5E2D8B40F1" box="[808,961,648,673]" pageId="2" pageNumber="453" value="1996-05-30">30 May 1996</date>
, trawl, coll.
<collectingCountry id="F3607625E20A2C75FBA7FD5E2A2B40F0" box="[1101,1121,649,672]" name="Italy" pageId="2" pageNumber="453">Y</collectingCountry>
. M. Yakovlev,
<specimenCount id="9D71FD3CE20A2C75FAE1FD5E2B3D40F1" box="[1291,1399,648,673]" pageId="2" pageNumber="453" type="female">2 females</specimenCount>
(cl 18.6,
<quantity id="4C8F9B50E20A2C75FF3DFD7B2F754094" box="[215,319,684,708]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="2" pageNumber="453" unit="mm" value="19.2">19.2 mm</quantity>
), CBM-ZC 4990; off Togi, Noto Peninsula, Ishikawa Prefecture,
<date id="FFC91075E20A2C75FBC8FD7B2AF64094" box="[1058,1212,684,709]" pageId="2" pageNumber="453" value="2007-05-20">20 May 2007</date>
, commercial trap,
<specimenCount id="9D71FD3CE20A2C75FA64FD7B2E9240B8" pageId="2" pageNumber="453" type="male">2 males</specimenCount>
(cl 22.6,
<quantity id="4C8F9B50E20A2C75FEBCFD182F8F40B8" box="[342,453,719,744]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" pageId="2" pageNumber="453" unit="mm" value="24.5">24.5 mm</quantity>
),
<collectionCode id="ED66AE70E20A2C75FE35FD072C6740B7" box="[479,557,720,743]" pageId="2" pageNumber="453">IMNH</collectionCode>
; Wakasa Bay, Fukui Prefecture,
<geoCoordinate id="EE435072E20A2C75FC50FD182A0A40B8" box="[954,1088,719,744]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="36.1005">36°06.03’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FBB8FD182AAA40B8" box="[1106,1248,719,744]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="135.59067">135°35.44’E</geoCoordinate>
to
<geoCoordinate id="EE435072E20A2C75FAF8FD182BD240B8" box="[1298,1432,719,744]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="36.1105">36°06.63’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FF7DFD242F62415C" box="[151,296,755,780]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="135.61884">135°37.13’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FEDEFD242FFC415B" box="[308,438,755,780]" metricMagnitude="2" metricUnit="m" metricValue="3.37" metricValueMax="3.38" metricValueMin="3.36" pageId="2" pageNumber="453" unit="m" value="337.0" valueMax="338.0" valueMin="336.0">336–338 m</quantity>
,
<date id="FFC91075E20A2C75FE29FD242C17415B" box="[451,605,755,780]" pageId="2" pageNumber="453" value="2009-05-31">31 May 2009</date>
, RV “Tanshu-maru”, stn T 111, otter trawl,
<specimenCount id="9D71FD3CE20A2C75FBA7FD242AF0415C" box="[1101,1210,755,780]" pageId="2" pageNumber="453" type="female">2 females</specimenCount>
(cl 26.0,
<quantity id="4C8F9B50E20A2C75FACDFD242BDA415B" box="[1319,1424,755,779]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="2" pageNumber="453" unit="mm" value="28.7">28.7 mm</quantity>
), NSMT-Cr 22576; off Hamada, Shimane Prefecture,
<geoCoordinate id="EE435072E20A2C75FD34FCC12D2E417F" box="[734,868,790,815]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="35.893166">35°53.59’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FC9AFCC12A4B417F" box="[880,1025,790,815]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="131.47433">131°28.46’E</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FBE6FCC12AD8417F" box="[1036,1170,790,815]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="35.891666">35°53.50’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FB74FCC12B65417F" box="[1182,1327,790,815]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="131.44366">131°26.62’E</geoCoordinate>
, 342–
<quantity id="4C8F9B50E20A2C75FA99FCC02EE54101" metricMagnitude="2" metricUnit="m" metricValue="3.29" pageId="2" pageNumber="453" unit="m" value="329.0">329 m</quantity>
,
<date id="FFC91075E20A2C75FF56FCED2F034102" box="[188,329,826,850]" pageId="2" pageNumber="453" value="2009-05-09">9 May 2009</date>
, RV “Tanshu-maru”, otter trawl,
<specimenCount id="9D71FD3CE20A2C75FD28FCED2D6E4102" box="[706,804,825,850]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(damaged, not measured), NSMT-Cr 22574; SE of Oki Islands,
<geoCoordinate id="EE435072E20A2C75FF12FC8A2F344126" box="[248,382,861,886]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="35.913834">35°54,83’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FE67FC8A2C564126" box="[397,540,861,886]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="133.84517">133°50.71’E</geoCoordinate>
to
<geoCoordinate id="EE435072E20A2C75FDA2FC8A2C874126" box="[584,717,861,886]" direction="north" orientation="latitude" pageId="2" pageNumber="453" precision="9" value="35.888832">35°53.33’N</geoCoordinate>
,
<geoCoordinate id="EE435072E20A2C75FD37FC8A2D254126" box="[733,879,861,886]" direction="east" orientation="longitude" pageId="2" pageNumber="453" precision="9" value="133.844">133°50.64’E</geoCoordinate>
,
<quantity id="4C8F9B50E20A2C75FC94FC8A2D814125" box="[894,971,861,886]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="2" pageNumber="453" unit="m" value="206.0">206 m</quantity>
,
<date id="FFC91075E20A2C75FC30FC892A264126" box="[986,1132,862,886]" pageId="2" pageNumber="453" value="2009-06-02">2 June 2009</date>
, RV “Tanshu-maru”, otter trawl,
<specimenCount id="9D71FD3CE20A2C75FF0EFC562F7E41C9" box="[228,308,896,921]" pageId="2" pageNumber="453" type="male">1 male</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FE81FC572F9241C9" box="[363,472,896,921]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="453" unit="mm" value="30.5">30.5 mm</quantity>
),
<specimenCount id="9D71FD3CE20A2C75FE1BFC562C1D41C9" box="[497,599,896,921]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(cl 34.0 mm), NSMT-Cr 22575; NW of Hinomisaki, Izumo, Shimane Prefecture, ca.
<quantity id="4C8F9B50E20A2C75FED5FC732FC241EC" box="[319,392,932,957]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="453" unit="m" value="150.0">150 m</quantity>
, 1994, trawl,
<specimenCount id="9D71FD3CE20A2C75FDCBFC722C3D41ED" box="[545,631,932,957]" pageId="2" pageNumber="453" type="male">2 males</specimenCount>
(cl 21.5,
<quantity id="4C8F9B50E20A2C75FD08FC722D7941ED" box="[738,819,933,957]" metricMagnitude="1" metricUnit="m" metricValue="2.28" pageId="2" pageNumber="453" unit="m" value="22.8">22.8 m</quantity>
),
<specimenCount id="9D71FD3CE20A2C75FCADFC722DE241ED" box="[839,936,932,957]" pageId="2" pageNumber="453" type="female">1 female</specimenCount>
(cl
<quantity id="4C8F9B50E20A2C75FC3EFC722A7641ED" box="[980,1084,933,957]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="2" pageNumber="453" unit="mm" value="21.8">21.8 mm</quantity>
), CBM-ZC 12596.
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE20A2C7FFF2DFC102ABE4424" lastPageId="8" lastPageNumber="459" pageId="2" pageNumber="453" type="description">
<paragraph id="8BC836B5E20A2C75FF2DFC102FC84677" blockId="2.[151,1437,151,2026]" pageId="2" pageNumber="453">
<emphasis id="B903EAA7E20A2C75FF2DFC102F2741B0" bold="true" box="[199,365,967,992]" pageId="2" pageNumber="453">Redescription</emphasis>
.
<emphasis id="B903EAA7E20A2C75FE6BFC1F2C6A41B0" box="[385,544,968,992]" italics="true" pageId="2" pageNumber="453">Adult females</emphasis>
. Body (
<figureCitation id="134C2A30E20A2C75FD6DFC102C9141B0" box="[647,731,967,992]" captionStart="FIGURE 1" captionStartId="3.[151,250,1978,2000]" captionTargetBox="[151,1436,323,1953]" captionTargetId="figure@3.[151,1436,307,1953]" captionTargetPageId="3" captionText="FIGURE 1. Lebbeus armatus (Owen, 1839), female (cl 36.7 mm), Sea of Japan off Hokkaido, CBM-ZC 7914. A, entire animal in lateral view; B, dorsum of carapace in lateral view, showing surface setation." httpUri="https://zenodo.org/record/244812/files/figure.png" pageId="2" pageNumber="453">Figs. 1</figureCitation>
,
<figureCitation id="134C2A30E20A2C75FD05FC1F2CB741B0" box="[751,765,968,992]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="2" pageNumber="453">2</figureCitation>
A, B) robust; integument firm, surface with dense short pubescence on entire carapace and depressions on pleonal pleura; tips of spines or teeth frequently darkly pigmented generally.
</paragraph>
<paragraph id="8BC836B5E20A2C75FF2DFBE42CD3440F" blockId="2.[151,1437,151,2026]" pageId="2" pageNumber="453">
Rostrum (
<figureCitation id="134C2A30E20A2C75FED1FBE52FC6461A" box="[315,396,1074,1099]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="2" pageNumber="453">Figs. 2</figureCitation>
A) directed forward or slightly ascending, nearly straight or slightly curving dorsally, far overreaching distal end of antennular peduncle but not reaching distal margin of antennal scale, 0.6–1.0 times as long as carapace, with dagger-like distal part; dorsal margin armed with 2 or 3 widely spaced small teeth in proximal 0.6–0.7, followed by 4 prominent, crested postrostral teeth; ventral margin armed with 2–4 small teeth in distal 0.4, ventral lamina moderately developed, outline of ventral margin gently convex; lateral carina conspicuous in proximal 0.6. Carapace (
<figureCitation id="134C2A30E20A2C75FD8FFB342CFB46AC" box="[613,689,1251,1276]" captionStart="FIGURE 1" captionStartId="3.[151,250,1978,2000]" captionTargetBox="[151,1436,323,1953]" captionTargetId="figure@3.[151,1436,307,1953]" captionTargetPageId="3" captionText="FIGURE 1. Lebbeus armatus (Owen, 1839), female (cl 36.7 mm), Sea of Japan off Hokkaido, CBM-ZC 7914. A, entire animal in lateral view; B, dorsum of carapace in lateral view, showing surface setation." httpUri="https://zenodo.org/record/244812/files/figure.png" pageId="2" pageNumber="453">Figs. 1</figureCitation>
A, B, 2A, 3A) with high middorsal carina extending to posterior margin of carapace; postrostral teeth noticeably increasing in size anteriorly, anteriormost tooth strongest, distally overhanging rostral base, posteriormost tooth arising at 0.7 length of carapace; supraorbital tooth very strong, arising at rostral base, slightly curving dorsally in lateral view and slightly laterally in dorsal view, its dorsal margin carinate and fairly elevated (thus, surface between rostrum and supraorbital tooth forming deep channel), ventral margin slightly sinuous or convex in lateral view, merging into postrostral ridge; deep excavation just ventral to base of supraorbital tooth merging into orbit; suborbital lobe prominent, roundly subtriangular, inner margin keeled; anterolateral margin between antennal and pterygostomial teeth deeply notched at base of antennal tooth and then convex; antennal tooth very strong, far overreaching suborbital lobe, distinctly buttressed; pterygostomial tooth also strong, directed forward; postorbital region deeply depressed; branchial ridge clearly delimited, continuing to ridge supporting antennal tooth.
</paragraph>
<paragraph id="8BC836B5E20A2C74FF2DF9BE2DB2434C" blockId="2.[151,1437,151,2026]" lastBlockId="3.[151,1437,151,285]" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453">
Pleon (
<figureCitation id="134C2A30E20A2C75FEFFF9BE2F1444D2" box="[277,350,1641,1666]" captionStart="FIGURE 1" captionStartId="3.[151,250,1978,2000]" captionTargetBox="[151,1436,323,1953]" captionTargetId="figure@3.[151,1436,307,1953]" captionTargetPageId="3" captionText="FIGURE 1. Lebbeus armatus (Owen, 1839), female (cl 36.7 mm), Sea of Japan off Hokkaido, CBM-ZC 7914. A, entire animal in lateral view; B, dorsum of carapace in lateral view, showing surface setation." httpUri="https://zenodo.org/record/244812/files/figure.png" pageId="2" pageNumber="453">Figs. 1</figureCitation>
A, 2B) dorsally rounded; first to fifth pleura each with shallow but distinct depressions to receive overlapping pleura. First pleomere with anterolateral margin slightly sinuous; pleuron usually with 2 (rarely 3) strong, subequal ventral teeth (ventral margin between teeth deeply concave). Second pleomere with deep transverse groove on tergum, posterior section slightly higher than anterior section; pleuron usually with 3 teeth (rarely 4 or 5), middle tooth strong, others small, sometimes abraded in large specimens. Third pleomere with posterodorsal margin fairly produced posteriorly; pleuron usually with 3 (rarely 4) ventral teeth, second tooth strongest. Fourth pleuron usually with 2–4 (most frequently 3) ventral teeth, second tooth elongate. Fifth pleuron usually with 2 (rarely 3) strongly unequal ventral teeth, posteroventral one elongate. Sixth pleomere 2.2 times as long as fifth pleomere and 1.3 times as long as high; posterodorsal margin slightly produced and bilobed (
<figureCitation id="134C2A30E20A2C75FADCF85E2B3345F2" box="[1334,1401,1929,1954]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="2" pageNumber="453">Fig. 3</figureCitation>
C); posterolateral process terminating in slender tooth (
<figureCitation id="134C2A30E20A2C75FD03F87A2D644596" box="[745,814,1965,1990]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="2" pageNumber="453">Fig. 3</figureCitation>
D); posteroventral angle strongly flared laterally, with strong tooth (
<figureCitation id="134C2A30E20A2C75FEDAF8062F3E45BA" box="[304,372,2001,2026]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="2" pageNumber="453">Fig. 3</figureCitation>
C). Telson (
<figureCitation id="134C2A30E20A2C75FE10F8062C7745BA" box="[506,573,2001,2026]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="2" pageNumber="453">Fig. 3</figureCitation>
C) about 1.9 times as long as sixth pleomere, 3.2 times longer than greatest width, gradually tapering posteriorly; dorsal surface deeply sulcate medially, proximally with prominent protuberance bearing tuft of short setae posteriorly (
<figureCitation id="134C2A30E20B2C74FD08FF6B2D6F4284" box="[738,805,188,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="3" pageNumber="454">Fig. 3</figureCitation>
D); dorsolateral ridge distinct, with row of 6 or 7 spines on either side; posterior margin terminating in blunt to acute point, with 2 pairs of small spines (mesial pair stronger than lateral pair) and tufts of setae at base of median tooth (
<figureCitation id="134C2A30E20B2C74FC7EFED32D9C434C" box="[916,982,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="3" pageNumber="454">Fig. 3</figureCitation>
E).
</paragraph>
<caption id="DF08663DE20B2C74FF7DF86D2DF645BE" httpUri="https://zenodo.org/record/244812/files/figure.png" pageId="3" pageNumber="454" targetBox="[151,1436,323,1953]" targetPageId="3">
<paragraph id="8BC836B5E20B2C74FF7DF86D2DF645BE" blockId="3.[151,1436,1978,2030]" pageId="3" pageNumber="454">
<emphasis id="B903EAA7E20B2C74FF7DF86D2F58459F" bold="true" box="[151,274,1978,2000]" pageId="3" pageNumber="454">FIGURE 1.</emphasis>
<taxonomicName id="4C774D36E20B2C74FEF2F86D2C134580" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[280,601,1978,2000]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E20B2C74FEF2F86D2F8F4580" box="[280,453,1978,2000]" italics="true" pageId="3" pageNumber="454">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, female (cl 36.7 mm), Sea of Japan off Hokkaido, CBM-ZC 7914. A, entire animal in lateral view; B, dorsum of carapace in lateral view, showing surface setation.
</paragraph>
</caption>
<caption id="DF08663DE20C2C73FF7DFAE02B05473C" httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="4" pageNumber="455" targetBox="[161,1429,204,1305]" targetPageId="4">
<paragraph id="8BC836B5E20C2C73FF7DFAE02B05473C" blockId="4.[151,1435,1335,1388]" pageId="4" pageNumber="455">
<emphasis id="B903EAA7E20C2C73FF7DFAE02F5C471C" bold="true" box="[151,278,1335,1357]" pageId="4" pageNumber="455">FIGURE 2.</emphasis>
<taxonomicName id="4C774D36E20C2C73FEF5FAE02C21471D" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[287,619,1335,1357]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E20C2C73FEF5FAE02F9A471D" box="[287,464,1335,1357]" italics="true" pageId="4" pageNumber="455">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BC836B5E20C2C73FF2DFA432DEE479F" blockId="4.[151,1437,1427,1992]" pageId="4" pageNumber="455">
Eye (
<figureCitation id="134C2A30E20C2C73FEE3FA442F1747FC" box="[265,349,1427,1452]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="4" pageNumber="455">Figs. 2</figureCitation>
A, 3A) subpyriform with eyestalk narrowing proximally; cornea small, slightly wider than eyestalk, its maximum width 0.15 of carapace length; ocellus present.
</paragraph>
<paragraph id="8BC836B5E20C2C73FF2DFA0C2EA4449C" blockId="4.[151,1437,1427,1992]" pageId="4" pageNumber="455">
Antennular peduncle (
<figureCitation id="134C2A30E20C2C73FE2AFA0C2C4747A4" box="[448,525,1499,1524]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="4" pageNumber="455">Figs. 2</figureCitation>
A, 3A) reaching midlength of antennal scale. First segment longer than distal two segments combined, dorsodistal margin armed with 1 very strong, obliquely erect tooth; stylocerite elongate, faintly sinuous, reaching distal end of antennular peduncle, bearing small, low protuberance proximolaterally. Second segment about 0.6 length of first segment, armed with l strong dorsolateral distal tooth. Third segment short, armed with 1 strong dorsodistal tooth. Upper flagellum with thickened aesthetasc-bearing portion about 0.3 of carapace length; lower flagellum longer than upper flagellum, articles each with few short setae on distal margin.
</paragraph>
<paragraph id="8BC836B5E20C2C73FF2DF90F2B6C45F4" blockId="4.[151,1437,1427,1992]" pageId="4" pageNumber="455">
Antenna (
<figureCitation id="134C2A30E20C2C73FEDDF9002FCC44BF" box="[311,390,1751,1776]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="4" pageNumber="455">Figs. 2</figureCitation>
A, 3A, F) with basicerite bearing strong ventrolateral distal tooth directed slightly laterally; dorsolateral distal angle also produced into strong tooth distinctly shorter than ventrolateral distal tooth. Carpocerite falling short of midlength of antennal scale. Antennal scale 0.8 times as long as carapace and about 2.5 times as long as wide; lateral margin nearly straight or faintly sinuous; dorsal surface with blunt median ridge accompanied by narrow, deep groove laterally; distolateral tooth far overreaching rounded distal margin of lamella, supported by longitudinal keel extending to base of antennal scale. Flagellum slightly shorter than body.
</paragraph>
<paragraph id="8BC836B5E20C2C73FF2DF8782C584598" blockId="4.[151,1437,1427,1992]" box="[199,530,1967,1992]" pageId="4" pageNumber="455">Mouthparts typical for genus.</paragraph>
<caption id="DF08663DE20D2C72FF7DF9182FEA452F" httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="5" pageNumber="456" targetBox="[167,1428,207,1681]" targetPageId="5">
<paragraph id="8BC836B5E20D2C72FF7DF9182FEA452F" blockId="5.[151,1436,1743,1919]" pageId="5" pageNumber="456">
<emphasis id="B903EAA7E20D2C72FF7DF9182F5C44B4" bold="true" box="[151,278,1743,1765]" pageId="5" pageNumber="456">FIGURE 3.</emphasis>
<taxonomicName id="4C774D36E20D2C72FEF5F9182C2144B5" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[287,619,1743,1766]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E20D2C72FEF5F9182F9A44B6" box="[287,464,1743,1766]" italics="true" pageId="5" pageNumber="456">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E.
</paragraph>
</caption>
<caption id="DF08663DE20E2C71FF7DF8762C2E45A4" httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="6" pageNumber="457" targetBox="[195,1394,204,1930]" targetPageId="6">
<paragraph id="8BC836B5E20E2C71FF7DF8762C2E45A4" blockId="6.[151,1436,1953,2036]" pageId="6" pageNumber="457">
<emphasis id="B903EAA7E20E2C71FF7DF8762F5F45E6" bold="true" box="[151,277,1953,1975]" pageId="6" pageNumber="457">FIGURE 4.</emphasis>
<taxonomicName id="4C774D36E20E2C71FEF4F8762C2245E7" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[286,616,1953,1975]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E20E2C71FEF4F8762F8445E7" box="[286,462,1953,1975]" italics="true" pageId="6" pageNumber="457">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF08663DE20F2C70FF7DFB202D704739" httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458" targetBox="[161,1424,223,1226]" targetPageId="7">
<paragraph id="8BC836B5E20F2C70FF7DFB202D704739" blockId="7.[151,1436,1271,1385]" pageId="7" pageNumber="458">
<emphasis id="B903EAA7E20F2C70FF7DFB202F59475D" bold="true" box="[151,275,1271,1293]" pageId="7" pageNumber="458">FIGURE 5.</emphasis>
<taxonomicName id="4C774D36E20F2C70FEF1FB202C15475D" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[283,607,1271,1294]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E20F2C70FEF1FB202F83475E" box="[283,457,1271,1294]" italics="true" pageId="7" pageNumber="458">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A–D, F; 0.5 mm for E.
</paragraph>
</caption>
<paragraph id="8BC836B5E20F2C70FF2DFA412DF34547" blockId="7.[151,1437,1430,2031]" pageId="7" pageNumber="458">
Third maxilliped (
<figureCitation id="134C2A30E20F2C70FE78FA412F9E47FF" box="[402,468,1430,1455]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="7" pageNumber="458">Fig. 4</figureCitation>
A) relatively stout, reaching or slightly overreaching distal margin of antennal scale by ultimate segment. Ultimate segment about 2.2 times as long as penultimate segment (= carpus), somewhat depressed dorsoventrally, tapering to rounded distal end only in distal 0.2; dorsal, lateral and ventral surfaces partially obscured by with numerous setae, and also with minute spiniform setae; mesial surface with numerous transverse tracts of stiff setae, also grooming apparatus; distal part circumscribed by small, darkly pigmented spines (
<figureCitation id="134C2A30E20F2C70FF06F99D2F674433" box="[236,301,1610,1635]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
A). Carpus short, with scattered minute spiniform setae and tufts of short transverse rows of stiff setae on dorsal and lateral surfaces; mesial surface with numerous transverse tracts of stiff setae, forming grooming apparatus. Antepenultimate segment strongly depressed in proximal half, distal part subtrigonal in cross section; lateral surface with scattered minute spiniform setae and row of longer spiniform setae on blunt longitudinal ridge; distal margin with conspicuous tooth dorsally and stronger tooth laterally, and with row of spiniform setae on either side of dorsodistal tooth and around base of distolateral tooth (
<figureCitation id="134C2A30E20F2C70FCB9F9292DDD4547" box="[851,919,1790,1815]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
B).
</paragraph>
<paragraph id="8BC836B5E20F2C70FF2DF8F52C63450F" blockId="7.[151,1437,1430,2031]" pageId="7" pageNumber="458">Strap-like, terminally hooked epipods present on third maxilliped to third pereopod, corresponding setobranchs present on first to fourth pereopods.</paragraph>
<paragraph id="8BC836B5E20F2C7FFF2DF8BD2FD543D7" blockId="7.[151,1437,1430,2031]" lastBlockId="8.[151,1437,151,1941]" lastPageId="8" lastPageNumber="459" pageId="7" pageNumber="458">
First pereopod (
<figureCitation id="134C2A30E20F2C70FE92F8BD2FF645D3" box="[376,444,1898,1923]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="7" pageNumber="458">Fig. 4</figureCitation>
B) stout, reaching midlength of antennal scale by tip of fingers. Dactylus about 0.7 times as long as palm, terminating in 2 darkly pigmented corneous claws (
<figureCitation id="134C2A30E20F2C70FC45F8592DBE45F7" box="[943,1012,1934,1959]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
E); fixed finger terminating in single corneous claw (
<figureCitation id="134C2A30E20F2C70FEA3F8652FC6459B" box="[329,396,1970,1995]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
E); both fingers each with 1 darkly pigmented, flattened spines proximal to base of terminal claw(s) on either side (
<figureCitation id="134C2A30E20F2C70FE4DF8012FA445BF" box="[423,494,2006,2031]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
E). Palm (
<figureCitation id="134C2A30E20F2C70FD82F8012CE745BF" box="[616,685,2006,2031]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="7" pageNumber="458">Fig. 5</figureCitation>
C) subcylindrical, about twice as long as wide. Carpus widened distally, cup-like, slightly shorter than palm, lateral surface with scattered minute spiniform setae, distomesial margin with deep notch; grooming apparatus consisting of short, obliquely longitudinal row of setae on ventromesial surface of palm and several short transverse rows of setae on mesial surface of carpus (
<figureCitation id="134C2A30E2002C7FFAD8FF082B3D42A8" box="[1330,1399,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="8" pageNumber="459">Fig. 5</figureCitation>
D). Merus slightly narrowing proximally, with small tubercle proximodorsally and with several sets of minute spiniform setae on proximal half of ventral margin, dorsodistal margin slightly produced, lateral surface with numerous scattered minute spiniform setae. Ischium with blunt ventrodistal angle, ventrolateral surface with minute spiniform setae.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFE432C544380" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
Second pereopod (
<figureCitation id="134C2A30E2002C7FFE72FE432F9643FC" box="[408,476,404,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="8" pageNumber="459">Fig. 4</figureCitation>
C) slender, overreaching antennal scale by about 0.3 length of carpus. Carpus divided into 7 articles, third article longest.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFE0B2FA64198" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
Third pereopod (
<figureCitation id="134C2A30E2002C7FFE6EFE0B2F8D43A4" box="[388,455,476,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="8" pageNumber="459">Fig. 4</figureCitation>
D) overreaching antennal scale by length of dactylus; dactylus (
<figureCitation id="134C2A30E2002C7FFB79FE0B2A9F43A5" box="[1171,1237,476,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[161,1424,223,1226]" captionTargetId="figure@7.[151,1436,217,1234]" captionTargetPageId="7" captionText="FIGURE 5. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, left third maxilliped, distal part of ultimate segment, dorsal view; B, same, distal part of antepenultimate segment, lateral view; C, left first pereopod, chela, extensor view; D, same, chela and carpus, mesial view; E, same, tip of fingers; F, left third pereopod, dactylus, lateral view. Scale bars: 2 mm for A – D, F; 0.5 mm for E." httpUri="https://zenodo.org/record/244816/files/figure.png" pageId="8" pageNumber="459">Fig. 5</figureCitation>
F) 0.3–0.35 times as long as propodus, stout (about 2.5 times as long as wide), terminating in strong, darkly pigmented unguis, armed with 4–6 darkly pigmented accessory spines on flexor margin, these accessory spines noticeably increasing in size distally, distalmost spinule subconical, making tip of dactylus biunguiculate; propodus with numerous darkly pigmented spinules, arranged in irregular 3 or 4 rows, on ventral surface; carpus slightly less than half length of propodus, slightly widened distally, armed usually with 1 or 2 small spines on lateral surface; merus armed with lateroventral row of 8–15 spines extending over entire length, and 1–7 smaller additional spines not aligned with lateroventral row; ischium also with 1 small lateral spine. Fourth pereopod (
<figureCitation id="134C2A30E2002C7FFBF8FD002A1240A0" box="[1042,1112,727,752]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="8" pageNumber="459">Fig. 4</figureCitation>
E) falling far short of distal margin of antennal scale by tip of dactylus; merus with lateroventral row of 7–10 spines and 0–6 smaller additional non-aligned spines particularly on ventral surface; ischium unarmed or armed with 1 lateral spine. Fifth pereopod (
<figureCitation id="134C2A30E2002C7FFF75FC932EA9410C" box="[159,227,836,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1953,1975]" captionTargetBox="[195,1394,204,1930]" captionTargetId="figure@6.[178,1408,193,1932]" captionTargetPageId="6" captionText="FIGURE 4. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990, left thoracic appendages in lateral view. A, left third maxilliped; B, first pereopod; C, second pereopod; D, third pereopod; E, fourth pereopod; F, fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/244815/files/figure.png" pageId="8" pageNumber="459">Fig. 4</figureCitation>
F) generally similar to preceding pereopods, but merus shorter and carpus and propodus combined longer; not reaching midlength of antennal scale; propodus with brush-like grooming setae distally; carpus with 1 or 2 small spines on lateral surface; merus armed with lateroventral row of 6–9 spines and 0–3 additional smaller spines not aligned; ischium unarmed.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFC032F534640" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">Protopods of pleopods in spawning molts each with deep ventrolateral lobe distally terminating in subacute or blunt point.</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFBCB2B0D462D" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
Uropod (
<figureCitation id="134C2A30E2002C7FFEC7FBCB2F3A4664" box="[301,368,1052,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1743,1765]" captionTargetBox="[167,1428,207,1681]" captionTargetId="figure@5.[151,1436,193,1694]" captionTargetPageId="5" captionText="FIGURE 3. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, anterior part of carapace and cephalic appendages, dorsal view (setae omitted); B, suborbital lobe, dorsolateral view; C, posterior part of sixth pleomere and telson, dorsal view (setae partially omitted); D, posterior part of sixth pleomere and proximal part of telson, lateral view (setae partially omitted); E, posterior margin of telson, dorsal view; F, left antennal scale, ventral view (setae omitted); G, left uropod, dorsal view (marginal setae on rami omitted). Scale bars: 5 mm for A, C, F, G; 2 mm for B, D; 1 mm for E." httpUri="https://zenodo.org/record/244814/files/figure.png" pageId="8" pageNumber="459">Fig. 3</figureCitation>
G) with protopod bearing sharp posterodorsal tooth, posterolateral angle terminating in strong tooth. Exopod with small spine just mesial to strong posterolateral tooth; dorsal surface with distinct longitudinal carina lateral to midline. Endopod with distinct median carina and scattered minute setae on dorsal surface.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFB5E2C8746F0" blockId="8.[151,1437,151,1941]" box="[199,717,1159,1185]" pageId="8" pageNumber="459">Eggs 2.4 x 2.0 mm in eyed stage, not counted.</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFB7B2AE947CD" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
<emphasis id="B903EAA7E2002C7FFF2DFB7B2F064694" box="[199,332,1196,1220]" italics="true" pageId="8" pageNumber="459">Adult males</emphasis>
. Generally similar to females except for sexual characters and smaller body size. Outer antennular flagellum (
<figureCitation id="134C2A30E2002C7FFEF8FB182F1E46B8" box="[274,340,1231,1256]" captionStart="FIGURE 6" captionStartId="9.[151,250,1110,1132]" captionTargetBox="[172,1418,182,1090]" captionTargetId="figure@9.[162,1426,175,1095]" captionTargetPageId="9" captionText="FIGURE 6. Lebbeus armatus (Owen, 1839), male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C." httpUri="https://zenodo.org/record/244817/files/figure.png" pageId="8" pageNumber="459">Fig. 6</figureCitation>
A) with thickened aesthetasc-bearing portion about 0.5 times as long as carapace; inner flagellum elongate (
<figureCitation id="134C2A30E2002C7FFEECFB232F03475D" box="[262,329,1268,1293]" captionStart="FIGURE 6" captionStartId="9.[151,250,1110,1132]" captionTargetBox="[172,1418,182,1090]" captionTargetId="figure@9.[162,1426,175,1095]" captionTargetPageId="9" captionText="FIGURE 6. Lebbeus armatus (Owen, 1839), male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C." httpUri="https://zenodo.org/record/244817/files/figure.png" pageId="8" pageNumber="459">Fig. 6</figureCitation>
A) and stout, slightly tapering distally, 1.3–1.9 times as long as carapace; most articles wider than long. Endopod of first pleopod (
<figureCitation id="134C2A30E2002C7FFE15FAC02C0B4760" box="[511,577,1303,1328]" captionStart="FIGURE 6" captionStartId="9.[151,250,1110,1132]" captionTargetBox="[172,1418,182,1090]" captionTargetId="figure@9.[162,1426,175,1095]" captionTargetPageId="9" captionText="FIGURE 6. Lebbeus armatus (Owen, 1839), male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C." httpUri="https://zenodo.org/record/244817/files/figure.png" pageId="8" pageNumber="459">Fig. 6</figureCitation>
B) rather abruptly tapering distally to terminal appendix interna in distal 0.3; mesial margin fringed with short simple setae; lateral margin sinuous, with row of moderately long plumose setae extending from base to distal 0.3. Appendix masculina of second pleopod (
<figureCitation id="134C2A30E2002C7FFC35FA882A684728" box="[991,1058,1375,1400]" captionStart="FIGURE 6" captionStartId="9.[151,250,1110,1132]" captionTargetBox="[172,1418,182,1090]" captionTargetId="figure@9.[162,1426,175,1095]" captionTargetPageId="9" captionText="FIGURE 6. Lebbeus armatus (Owen, 1839), male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C." httpUri="https://zenodo.org/record/244817/files/figure.png" pageId="8" pageNumber="459">Fig. 6</figureCitation>
C) 0.7 length of appendix interna, rounded terminally, bearing numerous stiff setae on distal end to distal 0.6 of mesial surface.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFA702A8247B5" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">Juvenile. Generally similar to adults, but additional small spines on meri of third to fifth pereopods not differentiated; minute spiniform setae on third maxilliped and pereopods also not differentiated.</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DFA382FB34400" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
<emphasis id="B903EAA7E2002C7FFF2DFA382FD14458" bold="true" box="[199,411,1519,1544]" pageId="8" pageNumber="459">Coloration in life</emphasis>
. Carapace mottled with white and reddish brown. Pleon with brown or reddish brown transverse bands on whitish background (second to fifth pleomeres each with band). Telson whitish in anterior half, reddish brown in posterior half.
</paragraph>
<paragraph id="8BC836B5E2002C7FFF2DF98B2ABE4424" blockId="8.[151,1437,151,1941]" box="[199,1268,1628,1653]" pageId="8" pageNumber="459">
<emphasis id="B903EAA7E2002C7FFF2DF98B2EBD4425" bold="true" box="[199,247,1628,1653]" pageId="8" pageNumber="459">Size</emphasis>
. Largest male cl
<quantity id="4C8F9B50E2002C7FFE52F98B2C544424" box="[440,542,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" pageId="8" pageNumber="459" unit="mm" value="30.5">30.5 mm</quantity>
, largest female cl
<quantity id="4C8F9B50E2002C7FFD03F98B2D054424" box="[745,847,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.67" pageId="8" pageNumber="459" unit="mm" value="36.7">36.7 mm</quantity>
, ovigerous females cl 28.0–
<quantity id="4C8F9B50E2002C7FFB63F98B2ABA4424" box="[1161,1264,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="8" pageNumber="459" unit="mm" value="35.2">35.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE2002C7FFF2DF9A82D9C44EC" pageId="8" pageNumber="459" type="distribution">
<paragraph id="8BC836B5E2002C7FFF2DF9A82D9C44EC" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
<emphasis id="B903EAA7E2002C7FFF2DF9A82F1344C8" bold="true" box="[199,345,1663,1688]" pageId="8" pageNumber="459">Distribution</emphasis>
. Certain geographical range includes Kamchatka, Sea of Okhotsk, Sea of
<collectingCountry id="F3607625E2002C7FFB43F9562AA644C8" box="[1193,1260,1665,1688]" name="Japan" pageId="8" pageNumber="459">Japan</collectingCountry>
, Pacific side of northern
<collectingCountry id="F3607625E2002C7FFF16F9722F7644EC" box="[252,316,1701,1724]" name="Japan" pageId="8" pageNumber="459">Japan</collectingCountry>
extending from Hokkaido to Iwate Prefecture;
<quantity id="4C8F9B50E2002C7FFCBAF9732D9844EC" box="[848,978,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="2.41" metricValueMax="3.38" metricValueMin="1.44" pageId="8" pageNumber="459" unit="m" value="241.0" valueMax="338.0" valueMin="144.0">
144–
<specimenCount id="9D71FD3CE2002C7FFC6DF9732D9844EC" box="[903,978,1700,1724]" pageId="8" pageNumber="459" type="generic">338 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE2002C7DFF2DF9102FB9433C" lastPageId="10" lastPageNumber="461" pageId="8" pageNumber="459" type="description">
<paragraph id="8BC836B5E2002C7FFF2DF9102A6045C5" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="459">
<emphasis id="B903EAA7E2002C7FFF2DF9102F7F44B0" bold="true" box="[199,309,1735,1760]" pageId="8" pageNumber="459">Variation</emphasis>
. The number of ventral teeth on the second to fifth pleura of the pleon exhibits substantial variation not related to sex or size (
<tableCitation id="C6F5030EE2002C7FFE24F93B2C6F4554" box="[462,549,1772,1797]" captionStart="TABLE 1" captionStartId="9.[151,239,1263,1286]" captionText="TABLE 1. Variation in the number of ventral teeth of the second to fifth pleura of the pleon in Lebbeus armatus (Owen, 1839). The numbers of teeth are indicated by combination of numbers counted from both sides (only specimens with intact pair of pleura were used). In the case of asymmetrical number, the arrangement does not precisely represent the actual side of pleura, but only combination of numbers." pageId="8" pageNumber="459">Table 1</tableCitation>
), although in some cases, the absence of tooth or teeth seems to be due to secondary loss caused by damage or abrasion. The second pleura usually bear three ventral teeth, although the number of the teeth on either side varies from three to five. The third pleura have three teeth at least on one side in all specimens examined. The fourth pleura have three or more teeth in vast majority of the specimens (83.3%). The fifth pleura bear two ventral teeth at least on one side in most specimens (95.8%).
</paragraph>
<caption id="DF08663DE2012C7EFF7DFB812F4A4698" httpUri="https://zenodo.org/record/244817/files/figure.png" pageId="9" pageNumber="460" targetBox="[172,1418,182,1090]" targetPageId="9">
<paragraph id="8BC836B5E2012C7EFF7DFB812F4A4698" blockId="9.[151,1436,1110,1224]" pageId="9" pageNumber="460">
<emphasis id="B903EAA7E2012C7EFF7DFB812F58463B" bold="true" box="[151,274,1110,1132]" pageId="9" pageNumber="460">FIGURE 6.</emphasis>
<taxonomicName id="4C774D36E2012C7EFEF2FB812C10463C" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[280,602,1110,1132]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2012C7EFEF2FB812F8F463C" box="[280,453,1110,1132]" italics="true" pageId="9" pageNumber="460">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
, male (cl 21.5 mm), NW of Hinomisaki, Izumo, Sea of Japan, CBM-ZC 12596. A, anterior part of carapace and cephalic appendages, lateral view (setae partially omitted); B, endopod of left first pleopod, ventral view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 5 mm for A; 2 mm for B; 1 mm for C.
</paragraph>
</caption>
<caption id="DF08663DE2012C7EFF7DFB382CAC4738" pageId="9" pageNumber="460">
<paragraph id="8BC836B5E2012C7EFF7DFB382CAC4738" blockId="9.[151,1436,1262,1384]" pageId="9" pageNumber="460">
<emphasis id="B903EAA7E2012C7EFF7DFB382F434756" bold="true" box="[151,265,1263,1286]" pageId="9" pageNumber="460">TABLE 1.</emphasis>
Variation in the number of ventral teeth of the second to fifth pleura of the pleon in
<taxonomicName id="4C774D36E2012C7EFB65FB392E924776" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2012C7EFB65FB392B0F4756" box="[1167,1349,1262,1286]" italics="true" pageId="9" pageNumber="460">Lebbeus armatus</emphasis>
(Owen, 1839)
</taxonomicName>
. The numbers of teeth are indicated by combination of numbers counted from both sides (only specimens with intact pair of pleura were used). In the case of asymmetrical number, the arrangement does not precisely represent the actual side of pleura, but only combination of numbers.
</paragraph>
</caption>
<paragraph id="8BC836B5E2012C7EFF75FA532F6547CA" blockId="9.[159,1361,1412,1517]" box="[159,303,1412,1434]" pageId="9" pageNumber="460">Second pleura</paragraph>
<paragraph id="8BC836B5E2012C7EFF75FA792F564410" blockId="9.[159,1361,1412,1517]" lastBlockId="9.[159,1170,1578,1682]" pageId="9" pageNumber="460">Combination of number of teeth 2-3 3-3 3-4 4-5 Number of individuals 2 (8.3%) 19 (79%) 2 (8.3%) 1 (4.4%) Third pleura</paragraph>
<paragraph id="8BC836B5E2012C7EFF75F9842F6244B5" blockId="9.[159,1170,1578,1682]" lastBlockId="9.[159,1374,1743,1848]" pageId="9" pageNumber="460">Combination of number of teeth 2-3 3-3 3-4 Number of individuals 1 (4.5%) 19 (86.4%) 2 (9.1%) Fourth pleura</paragraph>
<paragraph id="8BC836B5E2012C7EFF75F92F2F5F45DA" blockId="9.[159,1374,1743,1848]" lastBlockId="9.[159,1170,1908,2013]" pageId="9" pageNumber="460">Combination of number of teeth 2-2 2-3 3-3 3-4 Number of individuals 4 (16.7%) 1 (4.2%) 14 (58.3%) 5 (20.8%) Fifth pleura</paragraph>
<paragraph id="8BC836B5E2012C7DFF75F8492FB9433C" blockId="9.[159,1170,1908,2013]" lastBlockId="10.[151,1437,152,2021]" lastPageId="10" lastPageNumber="461" pageId="9" pageNumber="460">
Combination of number of teeth 2-2 2-3 3-3 Number of individuals 20 (83.3%) 3 (12.5%) 1 (4.2%) Size-related variation is seen in the development of scattered minute spiniform setae on the surfaces of meral segments on the third maxilliped through the fifth pereopod and additional spines on the meri of the third to fifth pereopods: in juveniles, such spiniform setae or additional spines are not differentiated, but in large specimens, these setae or spines are prominent. The structure of the inner antennular flagellum exhibits noticeable sexual dimorphism as described and figured above, as in other congeneric species (e.g.,
<bibRefCitation id="EFE64B44E2022C7DFAC5FEFA2EB0433C" author="Komai" pageId="10" pageNumber="461" refString="Komai, T., Hayashi, K. &amp; Kohtsuka, H. (2004) Two new species of the shrimp genus Lebbeus White from the Sea of Japan, with redescription of Lebbeus kuboi Hayashi (Decapoda: Caridea: Hippolytidae). Crustacean Research, 33, 103 - 125." type="journal article" year="2004">
Komai
<emphasis id="B903EAA7E2022C7DFA6DFEFA2EFF433C" italics="true" pageId="10" pageNumber="461">et al.</emphasis>
2004
</bibRefCitation>
,
<bibRefCitation id="EFE64B44E2022C7DFEEFFE852F09433C" author="Komai" box="[261,323,338,364]" pageId="10" pageNumber="461" refString="Komai, T., Tsuchida, S. &amp; Segonzac, M. (2012) Records of species of the hippolytid genus Lebbeus White, 1847 (Crustacea: Decapoda: Caridea) from hydrothermal vents in the Pacific Ocean, with descriptions of three new species. Zootaxa, 3241, 35 - 63." type="journal article" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="EFE64B44E2022C7DFEBAFE852FAC433C" author="Komai" box="[336,486,338,364]" pageId="10" pageNumber="461" refString="Komai, T. (2013) A new species of the hippolytid shrimp genus Lebbeus (Crustacea: Decapoda: Caridea) from lower bathyal zone in the Izu Islands, central Japan. Natural History Research, 12, 81 - 89." type="journal article" year="2013">Komai 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36D653EE2022C7CFF2DFEA02AC5438E" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461" type="discussion">
<paragraph id="8BC836B5E2022C7DFF2DFEA02EAA46F8" blockId="10.[151,1437,152,2021]" pageId="10" pageNumber="461">
<emphasis id="B903EAA7E2022C7DFF2DFEA02F7E43C0" bold="true" box="[199,308,375,400]" pageId="10" pageNumber="461">Remarks</emphasis>
.
<taxonomicName id="4C774D36E2022C7DFEA0FEAF2C5843C0" box="[330,530,376,400]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFEA0FEAF2C5843C0" box="[330,530,376,400]" italics="true" pageId="10" pageNumber="461">Lebbeus armatus</emphasis>
</taxonomicName>
shares with
<taxonomicName id="4C774D36E2022C7DFD52FEAE2D3743C0" box="[696,893,376,400]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFD52FEAE2D3743C0" box="[696,893,376,400]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D36E2022C7DFC2EFEAE2A2E43C0" box="[964,1124,376,400]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">
<emphasis id="B903EAA7E2022C7DFC2EFEAE2A2E43C0" box="[964,1124,376,400]" italics="true" pageId="10" pageNumber="461">L. magnificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057DCE2022C7DFB9EFEA02AF143DF" box="[1140,1211,375,400]" pageId="10" pageNumber="461" rank="species">
<emphasis id="B903EAA7E2022C7DFB9EFEA02AF943C0" bold="true" box="[1140,1203,375,400]" pageId="10" pageNumber="461">n. sp</emphasis>
.
</taxonomicNameLabel>
several diagnostic characters, including the strong postrostral, antennal and pterygostomial teeth on the carapace, the suborbital lobe with a carinate inner margin, the presence of conspicuous ventral tooth/teeth on the first to fifth pleura, the prominent, laterally flared posteroventral teeth on the sixth pleomere, the presence of a prominent proximal protuberance on the dorsal surface of the telson, the possession of a strong tooth at the dorsolateral distal angle of the antennal basicerite, and the presence of a conspicuous tooth on the posterodorsal margin of the uropodal endopod. No other congeneric species have these characteristics, as far as known. Nevertheless,
<taxonomicName id="4C774D36E2022C7DFAEBFD862B354038" box="[1281,1407,593,616]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFAEBFD862B354038" box="[1281,1407,593,616]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
is easily distinguished from these two species by many characters, including the possession of a dense covering of short setae on the carapace, extending to postrostral teeth, and on depressions of pleura of the second to sixth pleomeres, the extremely strong, crested postrostral teeth noticeably increasing in the size anteriorly (the first tooth is strongest and produced beyond the second tooth), the clearly delimited branchial ridge on the carapace, the usual possession of at least three pleural ventral teeth on the second to fourth pleomeres, the presence of lateral spines on the carpi of the third and fourth pereopods, and the presence of additional spines, not aligned with lateroventral row, on the meri of the third to fifth pereopods. These characters appear autapomorphic to
<taxonomicName id="4C774D36E2022C7DFB4CFC9A2B6B4134" box="[1190,1313,845,868]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFB4CFC9A2B6B4134" box="[1190,1313,845,868]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
within the genus. In the latter two species, the carapace and pleon bear only sparse setae (in particular, postrostral teeth are glabrous); the postrostral teeth are generally weaker than in
<taxonomicName id="4C774D36E2022C7DFCD6FC422DFD41FC" box="[828,951,917,940]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFCD6FC422DFD41FC" box="[828,951,917,940]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
, of which the first tooth is weaker than the second tooth; the branchial ridge on the carapace is obsolescent; the second to fourth pleomeres are armed with one (second) or at most two (third and fourth; of which the anterior tooth, if present, is minute to tiny) pleural ventral teeth; the carpi of the third and fourth pereopods are unarmed; there are no additional spines on the meri of the third to fifth pereopods. From
<taxonomicName id="4C774D36E2022C7DFE53FBF22C32466C" box="[441,632,1060,1084]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFE53FBF22C32466C" box="[441,632,1060,1084]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C774D36E2022C7DFD63FBF22D4F466C" box="[649,773,1061,1084]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFD63FBF22D4F466C" box="[649,773,1061,1084]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
further differs in the more numerous meral spines on the third to fifth pereopods (for details, see
<tableCitation id="C6F5030EE2022C7DFDBEFB902CE34630" box="[596,681,1095,1120]" captionStart="TABLE 2" captionStartId="11.[151,239,260,283]" captionTargetBox="[159,1429,498,1910]" captionTargetPageId="11" captionText="TABLE 2. Differentiating characters among L. groenlandicus (Fabricius, 1775), L. armatus (Owen, 1839) and L. magnificus n. sp." httpUri="http://table.plazi.org/id/DF08663DE2032C7CFF7DFED32F6B436E" pageId="10" pageNumber="461" tableUuid="DF08663DE2032C7CFF7DFED32F6B436E">Table 2</tableCitation>
). From
<taxonomicName id="4C774D36E2022C7DFCE8FB9E2DD14630" box="[770,923,1096,1120]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">
<emphasis id="B903EAA7E2022C7DFCE8FB9E2DD14630" box="[770,923,1096,1120]" italics="true" pageId="10" pageNumber="461">L. magnificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057DCE2022C7DFC48FB902DAA460F" box="[930,992,1095,1120]" pageId="10" pageNumber="461" rank="species">
<emphasis id="B903EAA7E2022C7DFC48FB902D914630" bold="true" box="[930,987,1095,1120]" pageId="10" pageNumber="461">n. sp</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C774D36E2022C7DFC05FB9E2A234630" box="[1007,1129,1097,1120]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFC05FB9E2A234630" box="[1007,1129,1097,1120]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
differs in the possession of two pleural ventral teeth on the fifth pleomere (versus usually three) and the less stout inner antennular flagellum in males.
</paragraph>
<paragraph id="8BC836B5E2022C7DFF2DFB632C2547F5" blockId="10.[151,1437,152,2021]" pageId="10" pageNumber="461">
The material studied by
<bibRefCitation id="EFE64B44E2022C7DFE3DFB632C26469C" author="Owen" box="[471,620,1204,1228]" pageId="10" pageNumber="461" refString="Owen, R. (1839) Crustacea. In: Richardson, J., Beechey, F. W. &amp; Sowerby, G. B. (Eds.), The Zoology of Captain Beechey's Voyage, Compiled from the Collections and Notes Made by Captain Beechey, the Officers and Naturalist of the Expedition, during a Voyage to the Pacific and Behring's Straits Performed in His Majesty's Ship Blossom, under the Command of Captain F. W. Beechey, in the Years 1825, 26, 27, and 28. London, pp. 77 - 92. [plates. 24 - 28]" type="journal article" year="1839">Owen (1839)</bibRefCitation>
is no longer extant (cf.
<bibRefCitation id="EFE64B44E2022C7DFC9DFB632A16469C" author="Kim" box="[887,1116,1204,1229]" pageId="10" pageNumber="461" refString="Kim, J. N. &amp; Komai, T. (2002) Sclerocrangon gasuyebi Yokoya synonymized with Sclerocrangon salebrosa (Owen) (Decapoda, Caridea, Crangonidae). Crustaceana, 74, 1335 - 1347. http: // dx. doi. org / 10.1163 / 15685400152885273" type="journal article" year="2002">Kim &amp; Komai 2002</bibRefCitation>
; Sammy
<collectingCountry id="F3607625E2022C7DFB29FB622AAE469C" box="[1219,1252,1205,1228]" name="South Georgia and the South Sandwich Islands" pageId="10" pageNumber="461">De</collectingCountry>
Grave, personal communication). Nevertheless, Owen’s (1839) original description and figure of
<taxonomicName id="4C774D36E2022C7DFBCCFB0F2AA246A0" box="[1062,1256,1240,1264]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="B903EAA7E2022C7DFBCCFB0F2AA246A0" box="[1062,1256,1240,1264]" italics="true" pageId="10" pageNumber="461">Hippolite armata</emphasis>
</taxonomicName>
clearly indicate the presence of three teeth on the ventral margins of the second to fourth pleonal pleura. Furthermore, not specifically mentioned in the text, the given figure (pl. 27, fig. 2) clearly shows the postrostral teeth noticeably becoming stronger anteriorly with the first tooth strongest, and the presence of a clearly delimited branchial ridge on the carapace. Consequently, I have little hesitation to resurrect
<taxonomicName id="4C774D36E2022C7DFC7FFABE2A5B47D0" box="[917,1041,1385,1408]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFC7FFABE2A5B47D0" box="[917,1041,1385,1408]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
as a distinct species and refer the present East Asian material to the species.
</paragraph>
<paragraph id="8BC836B5E2022C7DFF2DFA782F114560" blockId="10.[151,1437,152,2021]" pageId="10" pageNumber="461">
There are many records of
<taxonomicName id="4C774D36E2022C7DFDCAFA662CAF4798" box="[544,741,1456,1480]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFDCAFA662CAF4798" box="[544,741,1456,1480]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
from East Asian waters (see synonymy), but references accompanied with figures or photos all matches
<taxonomicName id="4C774D36E2022C7DFD10FA022D3447BC" box="[762,894,1493,1516]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFD10FA022D3447BC" box="[762,894,1493,1516]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
. Therefore, these East Asian records of
<taxonomicName id="4C774D36E2022C7DFA6FFA022F7C4440" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFA6FFA022F7C4440" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
are all referred to
<taxonomicName id="4C774D36E2022C7DFDFBFA2E2CC74440" box="[529,653,1529,1552]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFDFBFA2E2CC74440" box="[529,653,1529,1552]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
. On the other hand, further study is needed to establish the specific identity of the records of
<taxonomicName id="4C774D36E2022C7DFE58F9CA2C254464" box="[434,623,1564,1588]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFE58F9CA2C254464" box="[434,623,1564,1588]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
from the Bering Sea to the west coast of North
<collectingCountry id="F3607625E2022C7DFB6CF9CB2AAC4465" box="[1158,1254,1564,1589]" name="United States of America" pageId="10" pageNumber="461">America</collectingCountry>
(
<bibRefCitation id="EFE64B44E2022C7DFB1FF9CB2BDD4464" author="Rathbun" box="[1269,1431,1564,1589]" pageId="10" pageNumber="461" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition, 10, 1 - 190. [plates 1 - 10]" type="journal article" year="1904">Rathbun 1904</bibRefCitation>
;
<bibRefCitation id="EFE64B44E2022C7DFF7DF9E82F6D4408" author="Butler" box="[151,295,1599,1624]" pageId="10" pageNumber="461" refString="Butler, T. H. (1980) Shrimps of the Pacific coast of Canada. Canadian Bulletin of Fisheries and Aquatic Sciences, 202, (i - ix), 1 - 280." type="journal article" year="1980">Butler 1980</bibRefCitation>
).
<bibRefCitation id="EFE64B44E2022C7DFEABF9E82F954408" author="Butler" box="[321,479,1599,1624]" pageId="10" pageNumber="461" refString="Butler, T. H. (1980) Shrimps of the Pacific coast of Canada. Canadian Bulletin of Fisheries and Aquatic Sciences, 202, (i - ix), 1 - 280." type="journal article" year="1980">Butler (1980)</bibRefCitation>
provided a rather detailed description and figure using specimens from British
<collectingCountry id="F3607625E2022C7DFF7DF9B32F42442D" box="[151,264,1636,1661]" name="United States of America" pageId="10" pageNumber="461">Columbia</collectingCountry>
,
<collectingCountry id="F3607625E2022C7DFEF9F9B32F21442D" box="[275,363,1636,1661]" name="Canada" pageId="10" pageNumber="461">Canada</collectingCountry>
. According to his account, the Canadian specimens agree with
<taxonomicName id="4C774D36E2022C7DFBC4F9B22AE2442C" box="[1070,1192,1637,1660]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFBC4F9B22AE2442C" box="[1070,1192,1637,1660]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
in the setose surfaces of the carapace and of the pleura of the pleon, and the postrostral teeth noticeably becoming stronger anteriorly, but still differs from
<taxonomicName id="4C774D36E2022C7DFEBEF97A2F844494" box="[340,462,1709,1732]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2022C7DFEBEF97A2F844494" box="[340,462,1709,1732]" italics="true" pageId="10" pageNumber="461">L. armatus</emphasis>
</taxonomicName>
in the fewer pleural ventral teeth of the third and fourth pleomeres and the fewer meral spines on the third to fifth pereopods (seven to nine in the third, six to eight in the fourth, and six to eight in the fifth). It is highly likely that the northeastern Pacific population represents another, third species closely related to
<taxonomicName id="4C774D36E2022C7DFF7DF8CE2F194560" box="[151,339,1816,1840]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFF7DF8CE2F194560" box="[151,339,1816,1840]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC836B5E2022C7CFF2DF8EB2A2C4284" blockId="10.[151,1437,152,2021]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="462" pageId="10" pageNumber="461">
Comments on
<taxonomicName id="4C774D36E2022C7DFE86F8EB2C7E4504" box="[364,564,1852,1876]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B903EAA7E2022C7DFE86F8EB2C7E4504" box="[364,564,1852,1876]" italics="true" pageId="10" pageNumber="461">Hippolite cornuta</emphasis>
</taxonomicName>
are noteworthy.
<bibRefCitation id="EFE64B44E2022C7DFD1EF8EA2DDF4504" author="Kroyer" box="[756,917,1852,1877]" pageId="10" pageNumber="461" refString="Kroyer, H. (1842) Monografisk Fremstilling af Slaegten Hippolyte's nordiske Arter. Med Bidrag til Decapodernes Udiklingshistorie. Kongelige Danske Videnskabernes Selskabs Naturvidenskebelig og Mathematisk Afhandeling, 9, 209 - 360. [plates. 1 - 6]" type="journal article" year="1842">Krøyer (1842)</bibRefCitation>
and
<bibRefCitation id="EFE64B44E2022C7DFC24F8EB2A274505" author="Brandt" box="[974,1133,1852,1877]" pageId="10" pageNumber="461" refString="Brandt, F. (1851) Krebse. In: Middendorff, A. Th. v. (Ed.) Reise in den aussersten Norden und Osten Sibiriens wahrend der Jahre 1843 und 1844 mit allerhochster Genehmigung auf veranstaltung der Kaiserlichen Akademie der Wissenchaften zu St. Petersburg ausgefuhrt und in Verbindung mit vielen Gelehten herausgegeben, 2 (1), 79 - 148. [plates. 5, 6]" type="journal article" year="1851">Brandt (1851)</bibRefCitation>
concluded that
<taxonomicName id="4C774D36E2022C7DFAC8F8EA2BD64504" box="[1314,1436,1853,1876]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B903EAA7E2022C7DFAC8F8EA2BD64504" box="[1314,1436,1853,1876]" italics="true" pageId="10" pageNumber="461">H. cornuta</emphasis>
</taxonomicName>
represented a male of
<taxonomicName id="4C774D36E2022C7DFE73F8B72C394528" box="[409,627,1888,1912]" class="Malacostraca" family="Hippolytidae" genus="Hippolyte" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis id="B903EAA7E2022C7DFE73F8B72C394528" box="[409,627,1888,1912]" italics="true" pageId="10" pageNumber="461">Hippolyte aculeata</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C774D36E2022C7DFD75F8B62D144528" box="[671,862,1888,1912]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFD75F8B62D144528" box="[671,862,1888,1912]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
). In the descriptive text,
<bibRefCitation id="EFE64B44E2022C7DFB6CF8B72B574528" author="Owen" box="[1158,1309,1888,1912]" pageId="10" pageNumber="461" refString="Owen, R. (1839) Crustacea. In: Richardson, J., Beechey, F. W. &amp; Sowerby, G. B. (Eds.), The Zoology of Captain Beechey's Voyage, Compiled from the Collections and Notes Made by Captain Beechey, the Officers and Naturalist of the Expedition, during a Voyage to the Pacific and Behring's Straits Performed in His Majesty's Ship Blossom, under the Command of Captain F. W. Beechey, in the Years 1825, 26, 27, and 28. London, pp. 77 - 92. [plates. 24 - 28]" type="journal article" year="1839">Owen (1839)</bibRefCitation>
stated that “The abdominal segments terminate inferiorly in two spines, of which the posterior is the strongest;”, although the given figure seems to show the presence of only one tooth in the second pleuron and of two teeth on the first, third to fifth pleuron.
<collectingCountry id="F3607625E2022C7DFEB8F81B2F3B45B4" box="[338,369,1996,2020]" name="American Samoa" pageId="10" pageNumber="461">As</collectingCountry>
shown in this study, males of
<taxonomicName id="4C774D36E2022C7DFD26F81A2DC045B4" box="[716,906,1996,2020]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2022C7DFD26F81A2DC045B4" box="[716,906,1996,2020]" italics="true" pageId="10" pageNumber="461">L. groenlandicus</emphasis>
</taxonomicName>
can have two ventral teeth in the first, third to fifth pleura and only one tooth in the second pleuron. The sampling locality of the
<typeStatus id="54CC8817E2032C7CFB8BFF4F2ADB42E0" box="[1121,1169,152,176]" pageId="11" pageNumber="462">type</typeStatus>
of
<taxonomicName id="4C774D36E2032C7CFB56FF4E2B7342E0" box="[1212,1337,153,176]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B903EAA7E2032C7CFB56FF4E2B7342E0" box="[1212,1337,153,176]" italics="true" pageId="11" pageNumber="462">H. cornuta</emphasis>
</taxonomicName>
was not indicated. The synonymy of
<taxonomicName id="4C774D36E2032C7CFE32FF6A2C1B4284" box="[472,593,189,212]" class="Malacostraca" family="Hippolytidae" genus="Hippolite" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="B903EAA7E2032C7CFE32FF6A2C1B4284" box="[472,593,189,212]" italics="true" pageId="11" pageNumber="462">H. cornuta</emphasis>
</taxonomicName>
with
<taxonomicName id="4C774D36E2032C7CFD7BFF6A2D044284" box="[657,846,188,212]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2032C7CFD7BFF6A2D044284" box="[657,846,188,212]" italics="true" pageId="11" pageNumber="462">L. groenlandicus</emphasis>
</taxonomicName>
is maintained at present.
</paragraph>
<caption id="DF08663DE2032C7CFF7DFED32F6B436E" ID-Table-UUID="DF08663DE2032C7CFF7DFED32F6B436E" httpUri="http://table.plazi.org/id/DF08663DE2032C7CFF7DFED32F6B436E" pageId="11" pageNumber="462" targetBox="[159,1429,498,1910]" targetIsTable="true" targetPageId="11">
<paragraph id="8BC836B5E2032C7CFF7DFED32F6B436E" blockId="11.[151,1436,260,318]" pageId="11" pageNumber="462">
<emphasis id="B903EAA7E2032C7CFF7DFED32F41434B" bold="true" box="[151,267,260,283]" pageId="11" pageNumber="462">TABLE 2.</emphasis>
Differentiating characters among
<taxonomicName id="4C774D36E2032C7CFD96FED22DBE434B" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[636,1012,260,284]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="groenlandicus">
<emphasis id="B903EAA7E2032C7CFD96FED22D7B434C" box="[636,817,260,284]" italics="true" pageId="11" pageNumber="462">L. groenlandicus</emphasis>
(Fabricius, 1775)
</taxonomicName>
,
<taxonomicName id="4C774D36E2032C7CFBEAFED22B58434B" authority="Owen, 1839" authorityName="Owen" authorityYear="1839" box="[1024,1298,260,284]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B903EAA7E2032C7CFBEAFED22A3E434C" box="[1024,1140,261,284]" italics="true" pageId="11" pageNumber="462">L. armatus</emphasis>
(Owen, 1839)
</taxonomicName>
and
<taxonomicName id="4C774D36E2032C7CFAA2FED22E94436E" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">
<emphasis id="B903EAA7E2032C7CFAA2FED22E94436E" italics="true" pageId="11" pageNumber="462">L. magnificus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057DCE2032C7CFF0EFEF02F6B436E" box="[228,289,295,318]" pageId="11" pageNumber="462" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BC836B5E2032C7CFF75FE8D2B6F4320" blockId="11.[159,1393,346,478]" box="[159,1317,346,368]" pageId="11" pageNumber="462">
Characters/species
<emphasis id="B903EAA7E2032C7CFE2AFE8C2AAF4320" box="[448,1253,346,368]" italics="true" pageId="11" pageNumber="462">
<taxonomicName id="4C774D36E2032C7CFE2AFE8C2C214320" box="[448,619,346,368]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="groenlandicus">L. groenlandicus</taxonomicName>
<taxonomicName id="4C774D36E2032C7CFCE7FE8C2D314320" box="[781,891,347,368]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="armatus">L. armatus</taxonomicName>
<taxonomicName id="4C774D36E2032C7CFBB6FE8C2AAF4320" box="[1116,1253,346,368]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="magnificus" status="sp. nov.">L. magnificus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23057DCE2032C7CFB06FE8C2B6F4320" box="[1260,1317,347,368]" pageId="11" pageNumber="462" rank="species">
<emphasis id="B903EAA7E2032C7CFB06FE8C2B6F4320" bold="true" box="[1260,1317,347,368]" pageId="11" pageNumber="462">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BC836B5E2032C7CFF75FE5D2B7343F0" blockId="11.[159,1393,346,478]" box="[159,1337,394,416]" pageId="11" pageNumber="462">Rostral shape dagger-like distal part dagger-like distal part dagger-like distal part</paragraph>
<paragraph id="8BC836B5E2032C7CFE2AFE7E2AC5438E" blockId="11.[159,1393,346,478]" pageId="11" pageNumber="462">
relatively small (
<figureCitation id="134C2A30E2032C7CFD80FE7E2CEC4390" box="[618,678,425,448]" captionStart="FIGURE 7" captionStartId="12.[151,250,1803,1825]" captionTargetBox="[161,1424,201,1768]" captionTargetId="figure@12.[151,1436,193,1781]" captionTargetPageId="12" captionText="FIGURE 7. Lebbeus groenlandicus (Fabricius, 1775), female (cl 16.4 mm), W of Greenland, HUMZ-C 1396. A, carapace and cephalic appendages, lateral view; B, pleon, lateral view (setae omitted); C, distal part of right antennal scale, dorsal view; D, left third pereopod, lateral view. Scale bars: 5 mm for A, B, D; 2 mm for C." httpUri="https://zenodo.org/record/244818/files/figure.png" pageId="11" pageNumber="462">Fig. 7</figureCitation>
A) relatively large (
<figureCitation id="134C2A30E2032C7CFC59FE7E2DB343EF" box="[947,1017,425,447]" captionStart="FIGURE 2" captionStartId="4.[151,250,1335,1357]" captionTargetBox="[161,1429,204,1305]" captionTargetId="figure@4.[151,1436,193,1314]" captionTargetPageId="4" captionText="FIGURE 2. Lebbeus armatus (Owen, 1839), female (cl 19.2 mm), North Yamato Bank, Sea of Japan, CBM-ZC 4990. A, carapace and cephalic appendages, lateral view (setae omitted); B, pleon, lateral view (setae omitted). Scale bar: 5 mm." httpUri="https://zenodo.org/record/244813/files/figure.png" pageId="11" pageNumber="462">Figs. 2</figureCitation>
A, 6A) relatively small (
<figureCitation id="134C2A30E2032C7CFAECFE7E2B1243EF" box="[1286,1368,425,447]" captionStart="FIGURE 10" captionStartId="15.[151,250,1851,1873]" captionTargetBox="[214,1365,206,1825]" captionTargetId="figure@15.[203,1383,193,1836]" captionTargetPageId="15" captionText="FIGURE 10. Lebbeus magnificus n. sp., holotype, ovigerous female (cl 24.0 mm), Kitami Yamato Bank, southwestern Sea of Okhotsk, CBM-ZC 12597. A, anterior part of carapace and cephalic appendages, right lateral view (setae partially omitted); B, same, dorsal view (marginal setae on antennal scales omitted; right stylocerite damaged); C, tergite of second pleomere, lateral view; D, third to sixth pleomeres, lateral view; E, posterior part of sixth pleomere, telson, and left uropod, dorsal view (marginal setae on uropodal rami omitted); F, posterior margin of telson, dorsal view (right spines broken off). Scale bars: 5 mm for A – E; 1 mm for F." httpUri="https://zenodo.org/record/244821/files/figure.png" pageId="11" pageNumber="462">Figs. 10</figureCitation>
A, 13A)
</paragraph>
</subSubSection>
</treatment>
</document>
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