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<document id="2E42573C8A7A50A947BEA32A44C4FA0D" ID-DOI="10.1111/j.1096-3642.2011.00787.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5407240" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630347082768" checkinUser="marcus" docAuthor="Hooker, Jerry J. &amp; Russell, Donald E." docDate="2012" docId="03DE8792FFBB6566FF31FB71FC0BFC07" docLanguage="en" docName="j.1096-3642.2011.00787.x.pdf" docOrigin="Zoological Journal of the Linnean Society 164 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00787.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Thryptodon brailloni Hooker &amp; Russell 2012, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="899" masterDocId="FFE7FFEAFF92654DFFF1FFB0FF80FF80" masterDocTitle="Early Palaeogene Louisinidae (Macroscelidea, Mammalia), their relationships and north European diversity" masterLastPageNumber="936" masterPageNumber="856" pageNumber="897" updateTime="1699238364316" updateUser="plazi">
<mods:mods id="B5ADE355C7CFD3BB3144F9FEDAECB92C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EFCEF5AA61CC911B9FF9268D15C12E97">Early Palaeogene Louisinidae (Macroscelidea, Mammalia), their relationships and north European diversity</mods:title>
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<mods:namePart id="04DEC5B9E7F94CB487E3357833EFB63E">Hooker, Jerry J.</mods:namePart>
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<mods:namePart id="6A5F0C1CA60026BEC61683B37AE964FF">Russell, Donald E.</mods:namePart>
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<mods:title id="5B00C19C17F9448AA4621FAE4CA410E6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="769715CDF0093C02B3A7DF7D9BCA7A23">2012</mods:date>
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<mods:number id="AF69FBBFECD36822228B0C3586C813FB">2012-03-20</mods:number>
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<mods:number id="D05B9D4AE61AB88864D66A50C5DCE851">164</mods:number>
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<mods:start id="FD13F60528ED559E9D5F92DA0E1E5217">856</mods:start>
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<mods:url id="545064CCBF6A28D58B9F34676D4BAAC5">http://dx.doi.org/10.1111/j.1096-3642.2011.00787.x</mods:url>
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<mods:classification id="468A9932E4BF4AE98BCE66933A57BE85">journal article</mods:classification>
<mods:identifier id="1DB5A0296CFB382136AF5C5E31060206" type="DOI">10.1111/j.1096-3642.2011.00787.x</mods:identifier>
<mods:identifier id="DB7483922D42440433152E5241F434F3" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="59E524127F745635162A0B5B2AE8EFE5" type="Zenodo-Dep">5407240</mods:identifier>
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<treatment id="03DE8792FFBB6566FF31FB71FC0BFC07" LSID="urn:lsid:plazi:treatment:03DE8792FFBB6566FF31FB71FC0BFC07" httpUri="http://treatment.plazi.org/id/03DE8792FFBB6566FF31FB71FC0BFC07" lastPageId="43" lastPageNumber="899" pageId="41" pageNumber="897">
<subSubSection id="C36D650FFFBB6564FF31FB71FD77FB7D" pageId="41" pageNumber="897" type="nomenclature">
<paragraph id="8BC83684FFBB6564FF31FB71FD46FB57" blockId="41.[192,710,1217,1239]" box="[192,710,1217,1239]" pageId="41" pageNumber="897">
<heading id="D08081E8FFBB6564FF31FB71FD46FB57" box="[192,710,1217,1239]" centered="true" fontSize="9" level="2" pageId="41" pageNumber="897" reason="4">
<taxonomicName id="4C774D07FFBB6564FF31FB71FE69FB55" authority="Hooker &amp; Russell, 2012" authorityName="Hooker &amp; Russell" authorityYear="2012" box="[192,489,1217,1238]" class="Mammalia" family="Louisinidae" genus="Thryptodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="41" pageNumber="899" phylum="Chordata" rank="species" species="brailloni" status="gen. et sp. nov.">THRYPTODON BRAILLONI</taxonomicName>
<emphasis id="B903EA96FFBB6564FE02FB74FDD8FB56" bold="true" box="[499,600,1217,1238]" pageId="41" pageNumber="897">
<taxonomicNameLabel id="A23057EDFFBB6564FE02FB74FDD8FB56" box="[499,600,1217,1238]" pageId="41" pageNumber="897" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="134C2A01FFBB6564FD9AFB71FD3EFB57" box="[619,702,1217,1239]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">FIG. 20</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BC83684FFBB6564FF61FB57FD77FB7D" blockId="41.[144,759,1255,1369]" box="[144,759,1255,1277]" pageId="41" pageNumber="897">vp 1964 Hyopsodontidé?; Russell, pp. 247–248, pl. 16,</paragraph>
</subSubSection>
<subSubSection id="C36D650FFFBB6566FF59FAB6FC0BFC07" lastPageId="43" lastPageNumber="899" pageId="41" pageNumber="897" type="description">
<paragraph id="8BC83684FFBB6564FF59FAB6FD76FABA" blockId="41.[144,759,1255,1369]" pageId="41" pageNumber="897">fig. 6b. vp 1967 Hyopsodontidé, genre indéterminé; Russell,</paragraph>
<paragraph id="8BC83684FFBB6564FF59FAF3FD29FAD9" blockId="41.[144,759,1255,1369]" box="[168,681,1347,1369]" pageId="41" pageNumber="897">Louis &amp; Poirier, p. 853, pl. 21, fig. 4 (right).</paragraph>
<paragraph id="8BC83684FFBB6564FF61FA39FE65FA3D" blockId="41.[144,758,1417,1469]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FF61FA39FE8DFA1E" box="[144,269,1417,1438]" italics="true" pageId="41" pageNumber="897">Etymology:</emphasis>
After the late Dr Jean Braillon, for finding key specimens of the species.
</paragraph>
<paragraph id="8BC83684FFBB6564FF61FA5EFDFEF983" blockId="41.[144,638,1516,1539]" box="[144,638,1516,1539]" pageId="41" pageNumber="897">
<materialsCitation id="3B1F3CD9FFBB6564FF61FA5EFDFEF983" box="[144,638,1516,1539]" collectionCode="MNHN" pageId="41" pageNumber="897" specimenCount="1" typeStatus="holotype">
<emphasis id="B903EA96FFBB6564FF61FA5EFF7AF983" box="[144,250,1518,1539]" italics="true" pageId="41" pageNumber="897">
<typeStatus id="54CC8826FFBB6564FF61FA5EFF74F983" box="[144,244,1518,1539]" pageId="41" pageNumber="897" type="holotype">Holotype</typeStatus>
:
</emphasis>
LM
<superScript id="7C029BCCFFBB6564FEDAFA5CFEB3FA78" attach="left" box="[299,307,1516,1528]" fontSize="5" pageId="41" pageNumber="897">2</superScript>
,
<collectionCode id="ED66AE41FFBB6564FEB5FA5DFE1DF983" box="[324,413,1517,1539]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="897">MNHN</collectionCode>
.F.CR-1268, Cernay.
</materialsCitation>
</paragraph>
<paragraph id="8BC83684FFBB6564FF61F984FDEEF906" blockId="41.[144,759,1586,1732]" pageId="41" pageNumber="897">
<materialsCitation id="3B1F3CD9FFBB6564FF61F984FEC9F9BE" box="[144,329,1586,1609]" collectionCode="RM" pageId="41" pageNumber="897" specimenCode="RM2" specimenCount="1" typeStatus="paratype">
<emphasis id="B903EA96FFBB6564FF61F984FE87F9C9" box="[144,263,1588,1609]" italics="true" pageId="41" pageNumber="897">
<typeStatus id="54CC8826FFBB6564FF61F984FE81F9C9" box="[144,257,1588,1609]" pageId="41" pageNumber="897" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCode id="DBD19EFFFFBB6564FEE9F984FEC9F9BE" box="[280,329,1586,1609]" collectionCode="RM" country="Canada" name="McGill University, Redpath Museum" pageId="41" pageNumber="897" type="Museum">
RM
<superScript id="7C029BCCFFBB6564FEB0F982FEC9F9BE" attach="left" box="[321,329,1586,1598]" fontSize="5" pageId="41" pageNumber="897">2</superScript>
</specimenCode>
</materialsCitation>
, MNHN.F.CR-311-L;
<materialsCitation id="3B1F3CD9FFBB6564FDA2F984FD04F9CA" box="[595,644,1588,1610]" collectionCode="RM" pageId="41" pageNumber="897" specimenCode="RM1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD19EFFFFBB6564FDA2F984FD04F9CA" box="[595,644,1588,1610]" collectionCode="RM" country="Canada" name="McGill University, Redpath Museum" pageId="41" pageNumber="897" type="Museum">
RM
<subScript id="17F334C1FFBB6564FD8DF98EFD04F9CA" attach="left" box="[636,644,1598,1610]" fontSize="5" pageId="41" pageNumber="897">1</subScript>
</specimenCode>
</materialsCitation>
, MNHN. F.CR-55-Bn; LM
<subScript id="17F334C1FFBB6564FEA9F9ECFEE0F9E8" attach="left" box="[344,352,1628,1640]" fontSize="5" pageId="41" pageNumber="897">2</subScript>
, MNHN.F.CR-317-L; two
<materialsCitation id="3B1F3CD9FFBB6564FD42F9E3FD64F9E8" box="[691,740,1619,1640]" collectionCode="RM" pageId="41" pageNumber="897" specimenCode="RM2" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBD19EFFFFBB6564FD42F9E3FD64F9E8" box="[691,740,1619,1640]" collectionCode="RM" country="Canada" name="McGill University, Redpath Museum" pageId="41" pageNumber="897" type="Museum">
RM
<subScript id="17F334C1FFBB6564FD2DF9ECFD64F9E8" attach="both" box="[732,740,1628,1640]" fontSize="5" pageId="41" pageNumber="897">2</subScript>
</specimenCode>
</materialsCitation>
s, MNHN.F.CR-90-Bn, CR-63-V; all Cernay.
</paragraph>
<paragraph id="8BC83684FFBB6564FF59F93FFF78F943" blockId="41.[144,759,1586,1732]" pageId="41" pageNumber="897">
Right M
<superScript id="7C029BCCFFBB6564FEF7F93DFE8EF919" attach="left" box="[262,270,1677,1689]" fontSize="5" pageId="41" pageNumber="897">1</superScript>
, MNHN.F.I-681; RM
<subScript id="17F334C1FFBB6564FE0EF929FD87F925" attach="left" box="[511,519,1689,1701]" fontSize="5" pageId="41" pageNumber="897">2</subScript>
, MNHN.F.I-685; bed 5, Berru.
</paragraph>
<paragraph id="8BC83684FFBB6564FF61F944FD31F8A8" blockId="41.[144,758,1780,1833]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FF61F944FE6CF88A" box="[144,492,1780,1802]" italics="true" pageId="41" pageNumber="897">Tentatively referred specimens:</emphasis>
LP
<subScript id="17F334C1FFBB6564FDE5F94EFD9CF88A" attach="left" box="[532,540,1790,1802]" fontSize="5" pageId="41" pageNumber="897">3</subScript>
, MNHN.F.CR-170- Ph; Cernay. RP
<subScript id="17F334C1FFBB6564FEB2F8ADFECBF8A9" attach="left" box="[323,331,1821,1833]" fontSize="5" pageId="41" pageNumber="897">3</subScript>
, MNHN.F.I-691; bed 5, Berru.
</paragraph>
<paragraph id="8BC83684FFBB6564FF61F8E9FDFBF8EE" blockId="41.[144,635,1881,1903]" box="[144,635,1881,1903]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FF61F8E9FEE0F8EE" box="[144,352,1881,1902]" italics="true" pageId="41" pageNumber="897">Cast in MNHN.F:</emphasis>
RP
<subScript id="17F334C1FFBB6564FE7DF8D3FE14F8EF" attach="left" box="[396,404,1891,1903]" fontSize="5" pageId="41" pageNumber="897">3</subScript>
, CR-64-Ro; Cernay.
</paragraph>
<paragraph id="8BC83684FFBB6564FCD7FF76FBDFFE99" blockId="41.[806,1421,198,281]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FCD7FF76FBAAFF5B" box="[806,1066,198,219]" italics="true" pageId="41" pageNumber="897">
<taxonomicName id="4C774D07FFBB6564FCD7FF76FCCFFF5B" box="[806,847,198,219]" class="Insecta" family="Tortricidae" genus="Age" kingdom="Animalia" order="Lepidoptera" pageId="41" pageNumber="897" phylum="Arthropoda" rank="genus">Age</taxonomicName>
and distribution:
</emphasis>
Sables de Châlons-sur-Vesle supérieurs, late Thanetian, Late Paleocene, Cernay and Berru (bed 5),
<collectingCountry id="F3607614FFBB6564FBF9FEB4FBDBFE99" box="[1032,1115,260,281]" name="France" pageId="41" pageNumber="897">France</collectingCountry>
.
</paragraph>
<paragraph id="8BC83684FFBB6564FCD7FEF1FB48FED6" blockId="41.[806,1224,320,342]" box="[806,1224,320,342]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FCD7FEF1FC1DFED6" box="[806,925,321,342]" italics="true" pageId="41" pageNumber="897">Diagnosis:</emphasis>
As for genus, monotypic.
</paragraph>
<paragraph id="8BC83684FFBB6564FCD7FECEFC27FE13" blockId="41.[806,1422,382,1444]" box="[806,935,382,403]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FCD7FECEFC27FE13" box="[806,935,382,403]" italics="true" pageId="41" pageNumber="897">Description</emphasis>
</paragraph>
<paragraph id="8BC83684FFBB6564FCD7FE2DFAABFBB5" blockId="41.[806,1422,382,1444]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FCD7FE2DFCDCFE32" box="[806,860,410,434]" italics="true" pageId="41" pageNumber="897">
M
<superScript id="7C029BCCFFBB6564FCCCFE2AFCD5FE26" attach="left" box="[829,853,410,422]" fontSize="5" pageId="41" pageNumber="897">1–2</superScript>
:
</emphasis>
What are interpreted here as M
<superScript id="7C029BCCFFBB6564FB1EFE2AFB77FE26" attach="left" box="[1263,1271,410,422]" fontSize="5" pageId="41" pageNumber="897">1</superScript>
and M
<superScript id="7C029BCCFFBB6564FAA2FE2AFADBFE26" attach="left" box="[1363,1371,410,422]" fontSize="5" pageId="41" pageNumber="897">2</superScript>
are similar in morphology (
<figureCitation id="134C2A01FFBB6564FBC7FE0BFB3AFE51" box="[1078,1210,443,465]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">Fig. 20A–C</figureCitation>
). Both tooth
<typeStatus id="54CC8826FFBB6564FAA1FE0BFA0DFE50" box="[1360,1421,443,464]" pageId="41" pageNumber="897">types</typeStatus>
are wider than long (
<tableCitation id="C6F5033FFFBB6564FBB4FE69FB19FE6F" box="[1093,1177,473,495]" captionStart="Table 7" captionStartId="43.[144,208,198,217]" captionText="Table 7. Length (L) and width (W) measurements in millimetres of Thryptodon brailloni gen. et sp. nov., Cernay and Berru. Widths of trigonids (W1) and talonids (W2) of lower molars are given separately" pageId="41" pageNumber="897">Table 7</tableCitation>
) and basically tribosphenic, with a moderate-sized hypocone that is smaller than the protocone. There is thus a welldeveloped trigon and no sign of any crest joining it to the hypocone. Main crests, except preparacrista, are present but weak and there is overall moderate bunodonty. The preparaconule crista almost reaches the lingual end of the paracingulum, which is confluent with the precingulum, ectocingulum, metacingulum, and postcingulum, which embrace the tooth uninterrupted on three sides, but not lingually. The paraconule is present but a little variable in size and position. The metaconule is also consistently present. Cristae are essentially absent from the metaconule, but there is a very thin low postparaconule crista on all three teeth. Also consistent are the relative sizes of the paracone and metacone, the latter being smaller than the former, although the degree of difference varies. Representation of a parastyle is negligible. The smallest bears an entostyle (
<figureCitation id="134C2A01FFBB6564FB5CFBAFFA95FBB5" box="[1197,1301,1055,1077]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">Fig. 20A</figureCitation>
<figureCitation id="134C2A01FFBB6564FAE4FBAFFA9EFBB5" box="[1301,1310,1055,1077]" captionStart="Figure 1" captionStartId="1.[144,223,1185,1204]" captionTargetBox="[312,1251,203,1150]" captionTargetId="figure-102@1.[303,1263,195,1156]" captionTargetPageId="1" captionText="Figure 1. Dental terminology. A, B, upper molar; C, D, lower molar, all shown as from left side in occlusal view. Mesial is to left; distal is to right; buccal in uppers is above and in lowers below; lingual in uppers is below and in lowers above. A, C, primitive morphotype; B, D, derived morphotype. Some cusps and crests shown in A and C are assumed for B and D, respectively. Some taxa combine some features of the different morphotypes. Abbreviations: co, cristid obliqua; dclm, distal cingulum; ecf, ectoflexus; ecfd, ectoflexid; eclm, ectocingulum; ectsd, ectostylid; entd, entoconid; entf, entoflexus; entld, entoconulid; ents, entostyle; hyd, hypoconid; hyfd, hypoflexid; hyld, hypoconulid; hyp, hypocone; hyphd, hypolophid; mbstd, mesiobuccal stylid; mclm, mesial cingulum; mes, mesostyle; mesd, mesoconid; mestd, mesostylid; met, metacone; metclm, metacingulum; metd, metaconid; metle, metaconule; metph, metaloph; mets, metastyle; pacd, paracristid; paclm, paracingulum; pad, paraconid; par, paracone; parle, paraconule; pas, parastyle; pocld, postcingulid; poclm, postcingulum; pof, postflexus; poentcd, postentocristid; pohycd, posthypocristid; pomec, postmetacrista; pometcd, postmetacristid; pometlc, postmetaconule crista; popac, postparacrista; poparlc, postparaconule crista; poprc, postprotocrista; precld, precingulid; preclm, precingulum; preentcd, pre-entocristid; pref, preflexus; premec, premetacrista; premetcd, premetacristid; premetlc, premetaconule crista; prepac, preparacrista; preparlc, preparaconule crista; preprc, preprotocrista; procd, protocristid; prot, protocone; protd, protoconid; prs, protostyle; prtph, protoloph; prtphd, protolophid; tb, trigon basin; tgb, trigonid basin; tlb, talonid basin. Tooth abbreviations in text: I, incisor; C, canine; P, premolar; M, molar; DP, deciduous premolar." figureDoi="http://doi.org/10.5281/zenodo.5407242" httpUri="https://zenodo.org/record/5407242/files/figure.png" pageId="41" pageNumber="897">1</figureCitation>
).
</paragraph>
<paragraph id="8BC83684FFBB6564FCCEFB8DFBA0FA23" blockId="41.[806,1422,382,1444]" pageId="41" pageNumber="897">
The smallest tooth (
<figureCitation id="134C2A01FFBB6564FBD9FB8DFB09FBD3" box="[1064,1161,1085,1107]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">Fig. 20A</figureCitation>
) has less difference in size between the paracone and metacone and these cusps are also better separated than on the other two teeth. There is a slight mesial parastylar bulge in the outline, the precingulum ends lingually in a small protostyle, and there is a small cusp on the postcingulum just buccal of the hypocone, all features lacking in the two larger teeth. The two larger teeth (
<figureCitation id="134C2A01FFBB6564FCDFFA82FC36FAC8" box="[814,950,1330,1352]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">Fig. 20B, C</figureCitation>
) are slightly shorter and broader and their buccal and lingual cusps are more occlusally convergent. The smallest tooth is interpreted as M
<superScript id="7C029BCCFFBB6564FA8FFADDFA06FAF9" attach="left" box="[1406,1414,1389,1401]" fontSize="5" pageId="41" pageNumber="897">1</superScript>
, the larger two as M
<superScript id="7C029BCCFFBB6564FBE0FA3CFB99FA18" attach="right" box="[1041,1049,1420,1432]" fontSize="5" pageId="41" pageNumber="897">2</superScript>
.
</paragraph>
<paragraph id="8BC83684FFBB6564FCD7FA7CFA0DF8EE" blockId="41.[806,1421,1483,1903]" pageId="41" pageNumber="897">
<emphasis id="B903EA96FFBB6564FCD7FA7CFCDCFA61" box="[806,860,1484,1505]" italics="true" pageId="41" pageNumber="897">
M
<subScript id="17F334C1FFBB6564FCCCFA65FCD5FA61" attach="left" box="[829,853,1493,1505]" fontSize="5" pageId="41" pageNumber="897">1–2</subScript>
:
</emphasis>
These four teeth are moderately bunodont like the uppers and display slight exodaenodonty as separate lobate bases to the trigonid and talonid (
<figureCitation id="134C2A01FFBB6564FAD7F9B8FCB9F9BC" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="41" pageNumber="897">Fig. 20E, F</figureCitation>
). The protoconid and metaconid are larger than the hypoconid and entoconid, but the difference varies in degree. The trigonid is longer and wider than the talonid, but this difference also varies in degree (
<tableCitation id="C6F5033FFFBB6564FCDFF911FC03F937" box="[814,899,1697,1719]" captionStart="Table 7" captionStartId="43.[144,208,198,217]" captionText="Table 7. Length (L) and width (W) measurements in millimetres of Thryptodon brailloni gen. et sp. nov., Cernay and Berru. Widths of trigonids (W1) and talonids (W2) of lower molars are given separately" pageId="41" pageNumber="897">Table 7</tableCitation>
). There is a distinct, small hypoconulid, but the paraconid is very small to absent. The protocristid is moderately notched. The protoconid bulges lingually, almost filling the trigonid basin. There is a blunt premetacristid that does not join the paraconid or its position at the end of the paracristid. The cristid obliqua is fairly strong, dipping steeply mesially at
</paragraph>
<caption id="DF08660CFFB86567FF55F97DFE6FF8F3" ID-DOI="http://doi.org/10.5281/zenodo.5407268" ID-Zenodo-Dep="5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="42" pageNumber="898" startId="42.[164,243,1741,1760]" targetBox="[164,1445,193,1711]" targetPageId="42">
<paragraph id="8BC83684FFB86567FF55F97DFE6FF8F3" blockId="42.[164,1441,1741,1907]" pageId="42" pageNumber="898">
<emphasis id="B903EA96FFB86567FF55F97DFE99F960" bold="true" box="[164,281,1741,1760]" pageId="42" pageNumber="898">Figure 20.</emphasis>
Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of
<taxonomicName id="4C774D07FFB86567FB80F97DFAE9F960" authority="Hooker &amp; Russell, 2012" authorityName="Hooker &amp; Russell" authorityYear="2012" box="[1137,1385,1741,1760]" class="Mammalia" family="Louisinidae" genus="Thryptodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="42" pageNumber="898" phylum="Chordata" rank="species" species="brailloni" status="gen. et sp. nov.">Thryptodon brailloni</taxonomicName>
<taxonomicNameLabel id="A23057EDFFB86567FA80F97EFEA2F97E" pageId="42" pageNumber="898" rank="species">
<emphasis id="B903EA96FFB86567FA80F97EFA21F961" bold="true" box="[1393,1441,1742,1761]" pageId="42" pageNumber="898">gen.</emphasis>
et
<emphasis id="B903EA96FFB86567FF34F95BFEA2F97E" bold="true" box="[197,290,1771,1790]" pageId="42" pageNumber="898">sp. nov.</emphasis>
</taxonomicNameLabel>
, late Thanetian, France. A, RM
<superScript id="7C029BCCFFB86567FD7EF957FD16F973" attach="left" box="[655,662,1767,1779]" fontSize="5" pageId="42" pageNumber="898">1</superScript>
(reversed), bed 5, Berru (MNHN.F.I-681). B, RM
<superScript id="7C029BCCFFB86567FB35F957FB4BF973" attach="left" box="[1220,1227,1767,1779]" fontSize="5" pageId="42" pageNumber="898">2</superScript>
(reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM
<superScript id="7C029BCCFFB86567FDDFF8B4FDB5F890" attach="right" box="[558,565,1796,1808]" fontSize="5" pageId="42" pageNumber="898">2</superScript>
, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E,
<subScript id="17F334C1FFB86567FC45F8A0FC3BF89C" attach="left" box="[948,955,1808,1820]" fontSize="5" pageId="42" pageNumber="898">3</subScript>
RM
<subScript id="17F334C1FFB86567FF38F89DFF50F8B9" attach="left" box="[201,208,1837,1849]" fontSize="5" pageId="42" pageNumber="898">1</subScript>
(reversed), Cernay (MNHN.F.CR-55-Bn). F, RM
<subScript id="17F334C1FFB86567FD0DF89DFC83F8B9" attach="left" box="[764,771,1837,1849]" fontSize="5" pageId="42" pageNumber="898">2</subScript>
(reversed), Cernay (MNHN.F.CR-90-Bn). G, LP
<subScript id="17F334C1FFB86567FADDF89DFAB3F8B9" attach="left" box="[1324,1331,1837,1849]" fontSize="5" pageId="42" pageNumber="898">3</subScript>
?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar
<emphasis id="B903EA96FFB86567FE6BF8EFFE26F8F3" bold="true" box="[410,422,1887,1907]" pageId="42" pageNumber="898">=</emphasis>
2 mm.
</paragraph>
</caption>
<caption id="DF08660CFFB96566FF61FF76FDCEFEB2" pageId="43" pageNumber="899" startId="43.[144,208,198,217]">
<paragraph id="8BC83684FFB96566FF61FF76FDCEFEB2" blockId="43.[144,758,198,307]" pageId="43" pageNumber="899">
<emphasis id="B903EA96FFB96566FF61FF76FF69FF59" bold="true" box="[144,233,198,217]" pageId="43" pageNumber="899">Table 7.</emphasis>
Length (L) and width (W) measurements in millimetres of
<taxonomicName id="4C774D07FFB96566FEEBFF53FD9AFF76" authority="Hooker &amp; Russell, 2012" authorityName="Hooker &amp; Russell" authorityYear="2012" box="[282,538,227,246]" class="Mammalia" family="Louisinidae" genus="Thryptodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="brailloni" status="gen. et sp. nov.">Thryptodon brailloni</taxonomicName>
<taxonomicNameLabel id="A23057EDFFB96566FDDAFF54FD70FF77" box="[555,752,228,247]" pageId="43" pageNumber="899" rank="species">
<emphasis id="B903EA96FFB96566FDDAFF54FDDBFF77" bold="true" box="[555,603,228,247]" pageId="43" pageNumber="899">gen.</emphasis>
et
<emphasis id="B903EA96FFB96566FD61FF54FD70FF77" bold="true" box="[656,752,228,247]" pageId="43" pageNumber="899">sp. nov.</emphasis>
</taxonomicNameLabel>
, Cernay and Berru. Widths of trigonids (W
<subScript id="17F334C1FFB96566FDA8FEB8FDE0FE94" attach="left" box="[601,608,264,276]" fontSize="5" pageId="43" pageNumber="899">1</subScript>
) and talonids (W
<subScript id="17F334C1FFB96566FF5CFE97FF34FEB3" attach="left" box="[173,180,295,307]" fontSize="5" pageId="43" pageNumber="899">2</subScript>
) of lower molars are given separately
</paragraph>
</caption>
<paragraph id="8BC83684FFB96566FF61FEEAFD67FD68" pageId="43" pageNumber="899">
<table id="F977C424FFB99AB2FF61FEE9FD76FD69" box="[144,758,345,745]" colsContinueFrom="16.[164,778,345,950]" gridcols="5" gridrows="13" pageId="43" pageNumber="899">
<tr id="354734C6FFB99AB2FF61FEE9FD76FEED" box="[144,758,345,365]" gridrow="0" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FEE9FEA1FEED" box="[144,289,345,365]" gridcol="0" gridrow="0" pageId="43" pageNumber="899">Measurement</th>
<th id="76965DBAFFB99AB2FEAAFEE9FE15FEED" box="[347,405,345,365]" gridcol="1" gridrow="0" pageId="43" pageNumber="899">Tooth</th>
<th id="76965DBAFFB99AB2FE3EFEE9FE61FEED" box="[463,481,345,365]" gridcol="2" gridrow="0" pageId="43" pageNumber="899">
<emphasis id="B903EA96FFB96566FE3EFEEAFE61FEEC" box="[463,481,346,364]" italics="true" pageId="43" pageNumber="899">N</emphasis>
</th>
<th id="76965DBAFFB99AB2FDEDFEE9FD01FEED" box="[540,641,345,365]" gridcol="3" gridrow="0" pageId="43" pageNumber="899">OR</th>
<th id="76965DBAFFB99AB2FD4AFEE9FD76FEED" box="[699,758,345,365]" gridcol="4" gridrow="0" pageId="43" pageNumber="899">Mean</th>
</tr>
<tr id="354734C6FFB99AB2FF61FE3FFD76FE27" box="[144,758,399,423]" gridrow="1" pageId="43" pageNumber="899" rowspan-4="1">
<th id="76965DBAFFB99AB2FF61FE3FFEA1FE27" box="[144,289,399,423]" gridcol="0" gridrow="1" pageId="43" pageNumber="899">L</th>
<td id="76965DBAFFB99AB2FEAAFE3FFE15FE27" box="[347,405,399,423]" gridcol="1" gridrow="1" pageId="43" pageNumber="899">M1</td>
<td id="76965DBAFFB99AB2FE3EFE3FFE61FE27" box="[463,481,399,423]" gridcol="2" gridrow="1" pageId="43" pageNumber="899">1</td>
<td id="76965DBAFFB99AB2FDEDFE3FFD01FE27" box="[540,641,399,423]" gridcol="3" gridrow="1" pageId="43" pageNumber="899">3.07</td>
</tr>
<tr id="354734C6FFB99AB2FF61FE1DFD76FE44" box="[144,758,429,452]" gridrow="2" pageId="43" pageNumber="899" rowspan-4="1">
<th id="76965DBAFFB99AB2FF61FE1DFEA1FE44" box="[144,289,429,452]" gridcol="0" gridrow="2" pageId="43" pageNumber="899">W</th>
<td id="76965DBAFFB99AB2FEAAFE1DFE15FE44" box="[347,405,429,452]" gridcol="1" gridrow="2" pageId="43" pageNumber="899">M1</td>
<td id="76965DBAFFB99AB2FE3EFE1DFE61FE44" box="[463,481,429,452]" gridcol="2" gridrow="2" pageId="43" pageNumber="899">1</td>
<td id="76965DBAFFB99AB2FDEDFE1DFD01FE44" box="[540,641,429,452]" gridcol="3" gridrow="2" pageId="43" pageNumber="899">4.08</td>
</tr>
<tr id="354734C6FFB99AB2FF61FE7AFD76FE61" box="[144,758,458,481]" gridrow="3" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FE7AFEA1FE61" box="[144,289,458,481]" gridcol="0" gridrow="3" pageId="43" pageNumber="899">L</th>
<td id="76965DBAFFB99AB2FEAAFE7AFE15FE61" box="[347,405,458,481]" gridcol="1" gridrow="3" pageId="43" pageNumber="899">M2</td>
<td id="76965DBAFFB99AB2FE3EFE7AFE61FE61" box="[463,481,458,481]" gridcol="2" gridrow="3" pageId="43" pageNumber="899">2</td>
<td id="76965DBAFFB99AB2FDEDFE7AFD01FE61" box="[540,641,458,481]" gridcol="3" gridrow="3" pageId="43" pageNumber="899">3.29–3.41</td>
<td id="76965DBAFFB99AB2FD4AFE7AFD76FE61" box="[699,758,458,481]" gridcol="4" gridrow="3" pageId="43" pageNumber="899">3.33</td>
</tr>
<tr id="354734C6FFB99AB2FF61FE57FD76FE7E" box="[144,758,487,510]" gridrow="4" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FE57FEA1FE7E" box="[144,289,487,510]" gridcol="0" gridrow="4" pageId="43" pageNumber="899">W</th>
<td id="76965DBAFFB99AB2FEAAFE57FE15FE7E" box="[347,405,487,510]" gridcol="1" gridrow="4" pageId="43" pageNumber="899">M2</td>
<td id="76965DBAFFB99AB2FE3EFE57FE61FE7E" box="[463,481,487,510]" gridcol="2" gridrow="4" pageId="43" pageNumber="899">2</td>
<td id="76965DBAFFB99AB2FDEDFE57FD01FE7E" box="[540,641,487,510]" gridcol="3" gridrow="4" pageId="43" pageNumber="899">4.44–5.20</td>
<td id="76965DBAFFB99AB2FD4AFE57FD76FE7E" box="[699,758,487,510]" gridcol="4" gridrow="4" pageId="43" pageNumber="899">4.82</td>
</tr>
<tr id="354734C6FFB99AB2FF61FDB8FD76FD9C" box="[144,758,520,540]" gridrow="5" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FDB8FEA1FD9C" box="[144,289,520,540]" gridcol="0" gridrow="5" pageId="43" pageNumber="899">L</th>
<td id="76965DBAFFB99AB2FEAAFDB8FE15FD9C" box="[347,405,520,540]" gridcol="1" gridrow="5" pageId="43" pageNumber="899">P3?</td>
<td id="76965DBAFFB99AB2FE3EFDB8FE61FD9C" box="[463,481,520,540]" gridcol="2" gridrow="5" pageId="43" pageNumber="899">3</td>
<td id="76965DBAFFB99AB2FDEDFDB8FD01FD9C" box="[540,641,520,540]" gridcol="3" gridrow="5" pageId="43" pageNumber="899">3.32–3.44</td>
<td id="76965DBAFFB99AB2FD4AFDB8FD76FD9C" box="[699,758,520,540]" gridcol="4" gridrow="5" pageId="43" pageNumber="899">3.40</td>
</tr>
<tr id="354734C6FFB99AB2FF61FD95FD76FDB9" box="[144,758,549,569]" gridrow="6" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FD95FEA1FDB9" box="[144,289,549,569]" gridcol="0" gridrow="6" pageId="43" pageNumber="899">W</th>
<td id="76965DBAFFB99AB2FEAAFD95FE15FDB9" box="[347,405,549,569]" gridcol="1" gridrow="6" pageId="43" pageNumber="899">P3?</td>
<td id="76965DBAFFB99AB2FE3EFD95FE61FDB9" box="[463,481,549,569]" gridcol="2" gridrow="6" pageId="43" pageNumber="899">3</td>
<td id="76965DBAFFB99AB2FDEDFD95FD01FDB9" box="[540,641,549,569]" gridcol="3" gridrow="6" pageId="43" pageNumber="899">1.85–2.05</td>
<td id="76965DBAFFB99AB2FD4AFD95FD76FDB9" box="[699,758,549,569]" gridcol="4" gridrow="6" pageId="43" pageNumber="899">1.98</td>
</tr>
<tr id="354734C6FFB99AB2FF61FDF3FD76FDD6" box="[144,758,579,598]" gridrow="7" pageId="43" pageNumber="899" rowspan-4="1">
<th id="76965DBAFFB99AB2FF61FDF3FEA1FDD6" box="[144,289,579,598]" gridcol="0" gridrow="7" pageId="43" pageNumber="899">L</th>
<td id="76965DBAFFB99AB2FEAAFDF3FE15FDD6" box="[347,405,579,598]" gridcol="1" gridrow="7" pageId="43" pageNumber="899">M1</td>
<td id="76965DBAFFB99AB2FE3EFDF3FE61FDD6" box="[463,481,579,598]" gridcol="2" gridrow="7" pageId="43" pageNumber="899">1</td>
<td id="76965DBAFFB99AB2FDEDFDF3FD01FDD6" box="[540,641,579,598]" gridcol="3" gridrow="7" pageId="43" pageNumber="899">3.75</td>
</tr>
<tr id="354734C6FFB99AB2FF61FDD0FD76FDF4" box="[144,758,608,628]" gridrow="8" pageId="43" pageNumber="899" rowspan-4="1">
<th id="76965DBAFFB99AB2FF61FDD0FEA1FDF4" box="[144,289,608,628]" gridcol="0" gridrow="8" pageId="43" pageNumber="899">W1</th>
<td id="76965DBAFFB99AB2FEAAFDD0FE15FDF4" box="[347,405,608,628]" gridcol="1" gridrow="8" pageId="43" pageNumber="899">M1</td>
<td id="76965DBAFFB99AB2FE3EFDD0FE61FDF4" box="[463,481,608,628]" gridcol="2" gridrow="8" pageId="43" pageNumber="899">1</td>
<td id="76965DBAFFB99AB2FDEDFDD0FD01FDF4" box="[540,641,608,628]" gridcol="3" gridrow="8" pageId="43" pageNumber="899">2.96</td>
</tr>
<tr id="354734C6FFB99AB2FF61FDCDFD76FD11" box="[144,758,637,657]" gridrow="9" pageId="43" pageNumber="899" rowspan-4="1">
<th id="76965DBAFFB99AB2FF61FDCDFEA1FD11" box="[144,289,637,657]" gridcol="0" gridrow="9" pageId="43" pageNumber="899">W2</th>
<td id="76965DBAFFB99AB2FEAAFDCDFE15FD11" box="[347,405,637,657]" gridcol="1" gridrow="9" pageId="43" pageNumber="899">M1</td>
<td id="76965DBAFFB99AB2FE3EFDCDFE61FD11" box="[463,481,637,657]" gridcol="2" gridrow="9" pageId="43" pageNumber="899">1</td>
<td id="76965DBAFFB99AB2FDEDFDCDFD01FD11" box="[540,641,637,657]" gridcol="3" gridrow="9" pageId="43" pageNumber="899">2.91</td>
</tr>
<tr id="354734C6FFB99AB2FF61FD2AFD76FD2E" box="[144,758,666,686]" gridrow="10" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FD2AFEA1FD2E" box="[144,289,666,686]" gridcol="0" gridrow="10" pageId="43" pageNumber="899">L</th>
<td id="76965DBAFFB99AB2FEAAFD2AFE15FD2E" box="[347,405,666,686]" gridcol="1" gridrow="10" pageId="43" pageNumber="899">M2</td>
<td id="76965DBAFFB99AB2FE3EFD2AFE61FD2E" box="[463,481,666,686]" gridcol="2" gridrow="10" pageId="43" pageNumber="899">3</td>
<td id="76965DBAFFB99AB2FDEDFD2AFD01FD2E" box="[540,641,666,686]" gridcol="3" gridrow="10" pageId="43" pageNumber="899">3.56–3.90</td>
<td id="76965DBAFFB99AB2FD4AFD2AFD76FD2E" box="[699,758,666,686]" gridcol="4" gridrow="10" pageId="43" pageNumber="899">3.74</td>
</tr>
<tr id="354734C6FFB99AB2FF61FD08FD76FD4B" box="[144,758,696,715]" gridrow="11" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FD08FEA1FD4B" box="[144,289,696,715]" gridcol="0" gridrow="11" pageId="43" pageNumber="899">W1</th>
<td id="76965DBAFFB99AB2FEAAFD08FE15FD4B" box="[347,405,696,715]" gridcol="1" gridrow="11" pageId="43" pageNumber="899">M2</td>
<td id="76965DBAFFB99AB2FE3EFD08FE61FD4B" box="[463,481,696,715]" gridcol="2" gridrow="11" pageId="43" pageNumber="899">3</td>
<td id="76965DBAFFB99AB2FDEDFD08FD01FD4B" box="[540,641,696,715]" gridcol="3" gridrow="11" pageId="43" pageNumber="899">3.05–3.34</td>
<td id="76965DBAFFB99AB2FD4AFD08FD76FD4B" box="[699,758,696,715]" gridcol="4" gridrow="11" pageId="43" pageNumber="899">3.15</td>
</tr>
<tr id="354734C6FFB99AB2FF61FD65FD76FD69" box="[144,758,725,745]" gridrow="12" pageId="43" pageNumber="899">
<th id="76965DBAFFB99AB2FF61FD65FEA1FD69" box="[144,289,725,745]" gridcol="0" gridrow="12" pageId="43" pageNumber="899">W2</th>
<td id="76965DBAFFB99AB2FEAAFD65FE15FD69" box="[347,405,725,745]" gridcol="1" gridrow="12" pageId="43" pageNumber="899">M2</td>
<td id="76965DBAFFB99AB2FE3EFD65FE61FD69" box="[463,481,725,745]" gridcol="2" gridrow="12" pageId="43" pageNumber="899">3</td>
<td id="76965DBAFFB99AB2FDEDFD65FD01FD69" box="[540,641,725,745]" gridcol="3" gridrow="12" pageId="43" pageNumber="899">2.97–3.14</td>
<td id="76965DBAFFB99AB2FD4AFD65FD76FD69" box="[699,758,725,745]" gridcol="4" gridrow="12" pageId="43" pageNumber="899">3.04</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC83684FFB96566FF61FCA0FEE9FCA3" blockId="43.[144,361,784,803]" box="[144,361,784,803]" pageId="43" pageNumber="899">OR, observed range.</paragraph>
<paragraph id="8BC83684FFB96566FF61FCEDFECDFB8C" blockId="43.[144,759,861,1679]" pageId="43" pageNumber="899">first then, after a notch, rises some way up the back of the trigonid; the attachment position is slightly buccal of the midline. There is a pre-entocristid that dips down to a shallow talonid notch. A faint entoconulid is sometimes present. The precingulid is sharp but short.</paragraph>
<paragraph id="8BC83684FFB96566FF59FBA5FD37F90E" blockId="43.[144,759,861,1679]" pageId="43" pageNumber="899">
Three of the four teeth have a talonid basin about one third of the length of the trigonid and the hypoconid and entoconid are about half the height of the protoconid and metaconid (
<figureCitation id="134C2A01FFB96566FE3BFBC1FDADFB07" box="[458,557,1137,1159]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="43" pageNumber="899">Fig. 20F</figureCitation>
). The level of the talonid notch is about half the height of the metaconid in the least worn of these three. The fourth tooth has moderate tip wear, but all the main cusps are distinctly separated (
<figureCitation id="134C2A01FFB96566FE39FB5BFDACFA81" box="[456,556,1259,1281]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="43" pageNumber="899">Fig. 20E</figureCitation>
). The talonid is nearly half as long as the trigonid and nearly as wide. There is less difference in height between the protoconid and metaconid on the one hand and the hypoconid and entoconid on the other. There is a tiny paraconid, lacking from the only other tooth with sufficiently light wear to have left the area unscathed. The talonid notch is also at a lower level. This tooth occludes well with the M
<superScript id="7C029BCCFFB96566FDF8FA6EFD91FA6A" attach="left" box="[521,529,1502,1514]" fontSize="5" pageId="43" pageNumber="899">1</superScript>
and is interpreted as M
<subScript id="17F334C1FFB96566FF21F9B9FF58F995" attach="left" box="[208,216,1545,1557]" fontSize="5" pageId="43" pageNumber="899">1</subScript>
. The other three, of which the least worn (MNHN.F.CR-90-Bn:
<figureCitation id="134C2A01FFB96566FE76F9ADFE66F9B3" box="[391,486,1565,1587]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="43" pageNumber="899">Fig. 20F</figureCitation>
) occludes well with the M
<superScript id="7C029BCCFFB96566FF56F98AFF2FF9C6" attach="both" box="[167,175,1594,1606]" fontSize="5" pageId="43" pageNumber="899">2</superScript>
s (because of the shorter proportions and greater buccal and lingual cusp convergence, which reduces the width of the trigon), are interpreted as M
<subScript id="17F334C1FFB96566FD59F933FD30F90F" attach="left" box="[680,688,1667,1679]" fontSize="5" pageId="43" pageNumber="899">2</subScript>
.
</paragraph>
<paragraph id="8BC83684FFB96566FF61F970FC10FD26" blockId="43.[144,759,1728,1903]" lastBlockId="43.[806,1422,198,678]" pageId="43" pageNumber="899">
<emphasis id="B903EA96FFB96566FF61F970FF3AF955" box="[144,186,1728,1750]" italics="true" pageId="43" pageNumber="899">
P
<subScript id="17F334C1FFB96566FF51F97AFF28F956" attach="left" box="[160,168,1738,1750]" fontSize="5" pageId="43" pageNumber="899">3</subScript>
?:
</emphasis>
Three simple premolars may represent this locus. MNHN.F.I-691 is from the right side and is
<quantity id="4C8F9B61FFB96566FD63F96FFD77F974" box="[658,759,1759,1780]" metricMagnitude="-3" metricUnit="m" metricValue="3.43" pageId="43" pageNumber="899" unit="mm" value="3.43">3.43 mm</quantity>
long by
<quantity id="4C8F9B61FFB96566FF1CF94DFED2F892" box="[237,338,1789,1811]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="43" pageNumber="899" unit="mm" value="1.85">1.85 mm</quantity>
wide (
<figureCitation id="134C2A01FFB96566FE53F94DFD88F893" box="[418,520,1789,1811]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="43" pageNumber="899">Fig. 20D</figureCitation>
). Another right (CR- 64-Ro) is
<quantity id="4C8F9B61FFB96566FF0EF8ACFEE4F8B1" box="[255,356,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="3.32" pageId="43" pageNumber="899" unit="mm" value="3.32">3.32 mm</quantity>
long by
<quantity id="4C8F9B61FFB96566FE3AF8ACFDB0F8B1" box="[459,560,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="2.04" pageId="43" pageNumber="899" unit="mm" value="2.04">2.04 mm</quantity>
wide. The protoconid is dominant, with a fairly low, lingually positioned paraconid and a small, scarcely basined talonid with a single cusp that is not transversely elongate. Weak crests descend the protoconid mesially and distally, the former steeper than the latter. Two exodaenodont lobes protrude over the anterior and posterior roots, but to a lesser extent than in P
<subScript id="17F334C1FFB96566FAB9FEFAFAD0FED6" attach="left" box="[1352,1360,330,342]" fontSize="5" pageId="43" pageNumber="899">3</subScript>
of
<taxonomicName id="4C774D07FFB96566FA87FEF1FCE8FEF4" authorityName="Hooker &amp; Russell" authorityYear="2012" class="Mammalia" family="Louisinidae" genus="Louisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="marci">
<emphasis id="B903EA96FFB96566FA87FEF1FCE8FEF4" italics="true" pageId="43" pageNumber="899">L. marci</emphasis>
</taxonomicName>
. The broad pattern is similar to P
<subScript id="17F334C1FFB96566FB1DFED9FB74FEF5" attach="right" box="[1260,1268,361,373]" fontSize="5" pageId="43" pageNumber="899">3</subScript>
s of the latter species, but the paraconid is much lower, like
<taxonomicName id="4C774D07FFB96566FAACFECEFC6BFE31" authorityName="Hooker &amp; Russell" authorityYear="2012" baseAuthorityName="Russell" baseAuthorityYear="1964" class="Mammalia" family="Louisinidae" genus="Prolouisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="atavella">
<emphasis id="B903EA96FFB96566FAACFECEFC6BFE31" italics="true" pageId="43" pageNumber="899">Prolouisina atavella</emphasis>
</taxonomicName>
, and overall crown height is less. The left tooth (MNHN.F.CR-170-Ph) is more worn than MNHN.F.I-691, with slightly greater relative width (
<quantity id="4C8F9B61FFB96566FC86FE48FC5CFD8D" box="[887,988,504,525]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="43" pageNumber="899" unit="mm" value="3.44">3.44 mm</quantity>
long by
<quantity id="4C8F9B61FFB96566FBCFFE48FB22FD8D" box="[1086,1186,504,526]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="43" pageNumber="899" unit="mm" value="2.05">2.05 mm</quantity>
wide) and a distally procumbent talonid (
<figureCitation id="134C2A01FFB96566FBEBFDA6FBFFFDAD" box="[1050,1151,534,557]" captionStart="Figure 20" captionStartId="42.[164,243,1741,1760]" captionTargetBox="[164,1445,193,1711]" captionTargetId="figure-0@42.[164,1445,193,1711]" captionTargetPageId="42" captionText="Figure 20. Scanning electron micrographs of gold-palladium coated epoxy casts of teeth of Thryptodon brailloni gen. et sp. nov., late Thanetian, France. A, RM1 (reversed), bed 5, Berru (MNHN.F.I-681). B, RM2 (reversed), Cernay (MNHN.F.CR-311-L). C, holotype LM2, Cernay (MNHN.F.CR-1268). D, RP? (reversed), bed 5, Berru (MNHN.F.I-691). E, 3 RM1 (reversed), Cernay (MNHN.F.CR-55-Bn). F, RM2 (reversed), Cernay (MNHN.F.CR-90-Bn). G, LP3?, Cernay (MNHN.F.CR-170-Ph). Views are: buccal (A2, B2, D2, E2, F2, G1), occlusal (A1, B1, C, D1, E1, F1, G2), and lingual (D3, E3, F3, G3). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5407268" httpUri="https://zenodo.org/record/5407268/files/figure.png" pageId="43" pageNumber="899">Fig. 20G</figureCitation>
), but is otherwise very similar. The large size and weak exodaenodonty, but otherwise similar structure to
<taxonomicName id="4C774D07FFB96566FB36FDE4FAACFDE9" authorityName="Hooker &amp; Russell" authorityYear="2012" box="[1223,1324,596,617]" class="Mammalia" family="Louisinidae" genus="Louisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="marci">
<emphasis id="B903EA96FFB96566FB36FDE4FAACFDE9" box="[1223,1324,596,617]" italics="true" pageId="43" pageNumber="899">L. marci</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D07FFB96566FA9FFDE4FC01FD07" authorityName="Hooker &amp; Russell" authorityYear="2012" baseAuthorityName="Russell" baseAuthorityYear="1964" class="Mammalia" family="Louisinidae" genus="Prolouisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="atavella">
<emphasis id="B903EA96FFB96566FA9FFDE4FC01FD07" italics="true" pageId="43" pageNumber="899">Pr. atavella</emphasis>
</taxonomicName>
suggests that these teeth may belong to
<taxonomicName id="4C774D07FFB96566FA97FDC2FC09FD26" authorityName="Hooker &amp; Russell" authorityYear="2012" class="Mammalia" family="Louisinidae" genus="Thryptodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="species" species="brailloni">
<emphasis id="B903EA96FFB96566FA97FDC2FC09FD26" italics="true" pageId="43" pageNumber="899">Th. brailloni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC83684FFB96566FCD7FD68FC0BFC07" blockId="43.[806,1421,728,903]" pageId="43" pageNumber="899">
<emphasis id="B903EA96FFB96566FCD7FD68FC27FD6D" box="[806,935,728,749]" italics="true" pageId="43" pageNumber="899">Discussion:</emphasis>
This monotypic genus is clearly distinct from
<taxonomicName id="4C774D07FFB96566FC9AFD46FC6FFC8B" authorityName="Hooker &amp; Russell" authorityYear="2012" box="[875,1007,758,779]" class="Mammalia" family="Louisinidae" genus="Prolouisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="genus">
<emphasis id="B903EA96FFB96566FC9AFD46FC6FFC8B" box="[875,1007,758,779]" italics="true" pageId="43" pageNumber="899">Prolouisina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C774D07FFB96566FBF4FD47FBE9FC8C" authorityName="Russell" authorityYear="1964" box="[1029,1129,759,780]" class="Mammalia" family="Louisinidae" genus="Louisina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Macroscelidea" pageId="43" pageNumber="899" phylum="Chordata" rank="genus">
<emphasis id="B903EA96FFB96566FBF4FD47FBE9FC8C" box="[1029,1129,759,780]" italics="true" pageId="43" pageNumber="899">Louisina</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C774D07FFB96566FB4BFD46FAD0FC8B" authorityName="Kretzoi &amp; Kretzoi" authorityYear="2000" box="[1210,1360,758,779]" class="Mammalia" family="Hyopsodontidae" genus="Teilhardimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Condylarthra" pageId="43" pageNumber="899" phylum="Chordata" rank="genus">
<emphasis id="B903EA96FFB96566FB4BFD46FAD0FC8B" box="[1210,1360,758,779]" italics="true" pageId="43" pageNumber="899">Teilhardimys</emphasis>
</taxonomicName>
, but more closely related to these genera than to other louisinids, sharing with them exodaenodonty and a lower molar precingulid that is short and rises mesially.
</paragraph>
</subSubSection>
</treatment>
</document>
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