197 lines
25 KiB
XML
197 lines
25 KiB
XML
<document id="3865E7E33BB6E8097877BA702A7A3FDB" ID-CLB-Dataset="29069" ID-DOI="10.11646/zootaxa.4457.2.4" ID-GBIF-Dataset="1645ea42-4c26-466d-9be8-c22bf3c12c45" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457760" ID-ZooBank="486B0557-DA8D-4042-A69B-1AAAEFFFE915" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539266480787" checkinUser="plazi" docAuthor="Wagner, Rüdiger" docDate="2018" docId="03DE8784FFC5105329B0F9E1FBC518CA" docLanguage="en" docName="zootaxa.4457.2.4.pdf" docOrigin="Zootaxa 4457 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theischingerus imbricatus Wagner 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="291" masterDocId="FFE7FFFCFFC110552927FFE2FFC61A05" masterDocTitle="Remarkable new Australian mothflies (Diptera, Psychodidae)" masterLastPageNumber="295" masterPageNumber="285" pageNumber="289" updateTime="1698576920796" updateUser="ExternalLinkService">
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<mods:titleInfo id="CD1BC0D61053E23AF6A05FEF8ED1BDDB">
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<mods:title id="F3CDB79E8427F4C330AA1E2D6CCBDEF0">Remarkable new Australian mothflies (Diptera, Psychodidae)</mods:title>
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<mods:name id="34ABF6F7CF85AE4442FF5F10254B8778" type="personal">
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<mods:namePart id="96D247C318CA9C7E059DA55BF910A5C7">Wagner, Rüdiger</mods:namePart>
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<mods:title id="6592BA84E6319C2897B048A4EFE5A46A">Zootaxa</mods:title>
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<mods:part id="E0483083435E910A365569C343B17359">
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<mods:date id="1262D7B8076ACFD8FA31927AC9F897CD">2018</mods:date>
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<mods:number id="F13D2F0D2F7D96FDB1F8AFE9238DD9D4">2018-08-08</mods:number>
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<mods:number id="4E623B0F3E770DD668D6FD0987753B61">4457</mods:number>
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<mods:identifier id="4BA20DF70BDC941960F61909E71E5B1B" type="DOI">10.11646/zootaxa.4457.2.4</mods:identifier>
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<treatment id="03DE8784FFC5105329B0F9E1FBC518CA" ID-DOI="http://doi.org/10.5281/zenodo.5975758" ID-GBIF-Taxon="148692084" ID-Zenodo-Dep="5975758" LSID="urn:lsid:plazi:treatment:03DE8784FFC5105329B0F9E1FBC518CA" httpUri="http://treatment.plazi.org/id/03DE8784FFC5105329B0F9E1FBC518CA" lastPageId="6" lastPageNumber="291" pageId="4" pageNumber="289">
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<subSubSection id="C36D6519FFC5105129B0F9E1FED81C3B" pageId="4" pageNumber="289" type="nomenclature">
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<paragraph id="8BC83692FFC5105129B0F9E1FDF91C18" blockId="4.[151,575,1539,1598]" box="[151,575,1539,1565]" pageId="4" pageNumber="289">
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<heading id="D08081FEFFC5105129B0F9E1FDF91C18" bold="true" box="[151,575,1539,1565]" fontSize="11" level="1" pageId="4" pageNumber="289" reason="1">
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<emphasis id="B903EA80FFC5105129B0F9E1FDF91C18" bold="true" box="[151,575,1539,1565]" pageId="4" pageNumber="289">
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<taxonomicName id="4C774D11FFC5105129B0F9E1FE1F1C18" ID-CoL="8FDXH" authority="Wagner, 2018" authorityName="Wagner" authorityYear="2018" box="[151,473,1539,1565]" class="Insecta" family="Psychodidae" genus="Theischingerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="imbricatus" status="sp. nov.">
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<emphasis id="B903EA80FFC5105129B0F9E1FE1F1C18" bold="true" box="[151,473,1539,1565]" italics="true" pageId="4" pageNumber="289">Theischingerus imbricatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057FBFFC5105128C6F9E1FDF91C18" box="[481,575,1539,1565]" pageId="4" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC83692FFC5105129B0F9C7FED81C3B" blockId="4.[151,575,1539,1598]" box="[151,286,1573,1598]" pageId="4" pageNumber="289">
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(
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<figureCitation id="134C2A17FFC5105129B8F9C7FED01C3B" box="[159,278,1573,1598]" captionStart="FIGURE 3" captionStartId="5.[151,250,1747,1769]" captionTargetBox="[175,1407,520,1665]" captionTargetId="figure@5.[150,1436,498,1726]" captionTargetPageId="5" captionText="FIGURE 3. Theischingerus imbricatus sp. nov. a. wing; b. wing base with expandable organ; c. genitalia, ventral view; d. aedeagus with parameral bridge. [Scale: a 1.0 mm, b–d 0.1 mm]" httpUri="https://zenodo.org/record/1457766/files/figure.png" pageId="4" pageNumber="289">Figs 3 a–d</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6519FFC5105129B0F98FFADB1C83" box="[151,1309,1645,1670]" pageId="4" pageNumber="289" type="etymology">
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<paragraph id="8BC83692FFC5105129B0F98FFADB1C83" blockId="4.[151,1436,1645,2030]" box="[151,1309,1645,1670]" pageId="4" pageNumber="289">
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<emphasis id="B903EA80FFC5105129B0F98FFED81C83" bold="true" box="[151,286,1645,1670]" pageId="4" pageNumber="289">Etymology.</emphasis>
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From latin
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<emphasis id="B903EA80FFC510512886F98FFDD91C83" box="[417,543,1645,1670]" italics="true" pageId="4" pageNumber="289">
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<taxonomicName id="4C774D11FFC510512886F98FFDDA1C83" authorityName="Wagner" authorityYear="2018" box="[417,540,1645,1670]" class="Insecta" family="Psychodidae" genus="Theischingerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="imbricatus">imbricatus</taxonomicName>
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,
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</emphasis>
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meaning “tiled”, referring to the median surface of the hypandrium.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6519FFC5105129E0F973FD5F1CCB" pageId="4" pageNumber="289" type="diagnosis">
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<paragraph id="8BC83692FFC5105129E0F973FD5F1CCB" blockId="4.[151,1436,1645,2030]" pageId="4" pageNumber="289">
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<emphasis id="B903EA80FFC5105129E0F973FE871CAF" bold="true" box="[199,321,1681,1706]" pageId="4" pageNumber="289">Diagnosis.</emphasis>
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<taxonomicName id="4C774D11FFC51051286DF973FDB31CAF" authority="Wagner, 2018" authorityName="Wagner" authorityYear="2018" box="[330,629,1681,1706]" class="Insecta" family="Psychodidae" genus="Theischingerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="289" phylum="Arthropoda" rank="species" species="imbricatus" status="sp. nov.">
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<emphasis id="B903EA80FFC51051286DF973FDB31CAF" box="[330,629,1681,1706]" italics="true" pageId="4" pageNumber="289">Theischingerus imbricatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057FBFFC510512B5AF973FD1F1CAF" box="[637,729,1681,1706]" pageId="4" pageNumber="289" rank="species">
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<emphasis id="B903EA80FFC510512B5AF973FD1F1CAF" bold="true" box="[637,729,1681,1706]" pageId="4" pageNumber="289">sp. nov.</emphasis>
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</taxonomicNameLabel>
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is distinguished from its congeners by the tiled surface of the broadened median section of the hypandrium.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6519FFC5105129E0F93BFAB51CF7" box="[199,1395,1753,1778]" pageId="4" pageNumber="289" type="materials_examined">
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<paragraph id="8BC83692FFC5105129E0F93BFAB51CF7" blockId="4.[151,1436,1645,2030]" box="[199,1395,1753,1778]" pageId="4" pageNumber="289">
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<materialsCitation id="3B1F3CCFFFC5105129E0F93BFAB51CF7" ID-GBIF-Occurrence="1917076156" box="[199,1395,1753,1778]" collectingDate="1986-10-13" collectorName="Theischinger." country="Australia" location="Macquarie Pass" pageId="4" pageNumber="289" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales" typeStatus="holotype">
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<emphasis id="B903EA80FFC5105129E0F93BFEF01CF7" bold="true" box="[199,310,1753,1778]" pageId="4" pageNumber="289">Material.</emphasis>
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<typeStatus id="54CC8830FFC51051281BF93BFE621CF7" box="[316,420,1753,1778]" pageId="4" pageNumber="289" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D71FD1BFFC51051288BF938FE091CF7" box="[428,463,1754,1778]" pageId="4" pageNumber="289" type="male">1♂</specimenCount>
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<collectingCountry id="F3607602FFC5105128F1F93BFD871CF7" box="[470,577,1753,1778]" name="Australia" pageId="4" pageNumber="289">Australia</collectingCountry>
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:
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<collectingRegion id="49B3F870FFC510512B6BF938FCD11CF7" box="[588,791,1753,1778]" country="Australia" name="New South Wales" pageId="4" pageNumber="289">
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New
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<collectingRegion id="49B3F870FFC510512BA0F93BFCD11CF7" box="[647,791,1753,1778]" inferredBySuffix="true" pageId="4" pageNumber="289">
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<collectingRegion id="49B3F870FFC510512BA0F93BFD0E1CF7" box="[647,712,1753,1778]" country="Tanzania" name="Kusini Unguja" pageId="4" pageNumber="289">South</collectingRegion>
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Wales
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</collectingRegion>
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</collectingRegion>
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,
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<location id="8EA86049FFC510512A05F93BFC131CF4" LSID="urn:lsid:plazi:treatment:03DE8784FFC5105329B0F9E1FBC518CA:8EA86049FFC510512A05F93BFC131CF4" box="[802,981,1753,1778]" country="Australia" name="Macquarie Pass" pageId="4" pageNumber="289" stateProvince="New South Wales">Macquarie Pass</location>
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,
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<collectingDate id="EF8DE9BAFFC510512AC7F938FB591CF7" box="[992,1183,1753,1778]" pageId="4" pageNumber="289" value="1986-10-13">
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<date id="FFC91052FFC510512AC7F938FB591CF7" box="[992,1183,1753,1778]" pageId="4" pageNumber="289" value="1986-10-13">13 October 1986</date>
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</collectingDate>
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, leg.
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<collectorName id="26825344FFC510512DFCF93BFAB51CF7" box="[1243,1395,1753,1778]" pageId="4" pageNumber="289">Theischinger.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6519FFC5105329E0F91FFB2E1B18" lastPageId="6" lastPageNumber="291" pageId="4" pageNumber="289" type="description">
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<paragraph id="8BC83692FFC5105029E0F91FFD001AAA" blockId="4.[151,1436,1645,2030]" lastBlockId="5.[151,1437,151,446]" lastPageId="5" lastPageNumber="290" pageId="4" pageNumber="289">
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<emphasis id="B903EA80FFC5105129E0F91FFE9F1D13" bold="true" box="[199,345,1789,1814]" pageId="4" pageNumber="289">Description.</emphasis>
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Eyes reniform, tight around antennae bases, interocular suture deep U-shaped, distance between the eyes 0.5 facet diameter; antenna with scape, pedicel and 14 flagellomeres; scape increasing in width distally, twice as long as its greatest width, pedicel spherical, diameter smaller than apical width of scape; flagellomeres bottle-shaped, all with a pair of digitate ascoids twice as long as a flagellomere, basal flagellomere with neck as long as bulb, subsequent segments with neck up to twice as long as bulb. Relative length of antennal articles: 80- 29-53-53-57-59-60-60-58-57-60-55-57-57-51-47; absolute length: 0.21-0.08-0.14-0.14-0.15-0.16-0.16-0.16-0.15- 0.15-0.16-0.14-0.15-0.15-0.13-
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<quantity id="4C8F9B77FFC5105128D5F837FD9B1DE8" box="[498,605,2005,2030]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="289" unit="mm" value="0.12">0.12 mm</quantity>
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. Four palpus segments, terminal segment flexible; relative length: 42-78- 78-119; absolute length: 0.11-0.21-
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<quantity id="4C8F9B77FFC410502B07FF7AFD041AAA" box="[544,706,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" metricValueMax="3.1" metricValueMin="2.1" pageId="5" pageNumber="290" unit="mm" value="0.26" valueMax="0.31" valueMin="0.21">0.21-0.31 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BC83692FFC4105029E0FF5EFC1C1BBE" blockId="5.[151,1437,151,446]" pageId="5" pageNumber="290">
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Wing broad with C basally curved out, Sc terminates in C at the end of the outcurve, R1 runs parallel to C and ends in it in distal third of the wing; an elongate digitate ‘sac’ (translucent probably flexible with apical apertures) protrudes from base of Rs; split-up of Rs unusual in
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<taxonomicName id="4C774D11FFC410502BCDFEE4FCBC1B1A" authorityName="Rondani" authorityYear="1858" box="[746,890,262,287]" class="Insecta" family="Psychodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="290" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
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, with fork R2/ R3+4 at level of Sc ending, R2 runs parallel to R1 and terminates into C distally of R1; R3/R4 fork at middle of wing; R5 straight terminates short behind wing tip; fork M1/2 basally of fork R3/4; M2 and R2 slightly thicker in short distance (wider with denser setae bases); jugum rhomboid with setose tip pointing distally; CuA long curved, terminates basally of fork M1/
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<quantity id="4C8F9B77FFC410502DC5FE68FAC21B91" box="[1250,1284,379,408]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="290" unit="in" value="2.0">2 in</quantity>
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C; CuP short distally curved. Cross veins r s -r5, r-m1+2, and m1+2-m3 present but very faint.
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</paragraph>
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<caption id="DF08661AFFC4105029B0F931FCE71D02" httpUri="https://zenodo.org/record/1457766/files/figure.png" pageId="5" pageNumber="290" startId="5.[151,250,1747,1769]" targetBox="[175,1407,520,1665]" targetPageId="5">
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<paragraph id="8BC83692FFC4105029B0F931FCE71D02" blockId="5.[151,1436,1747,1799]" pageId="5" pageNumber="290">
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||
<emphasis id="B903EA80FFC4105029B0F931FED31CEC" bold="true" box="[151,277,1747,1769]" pageId="5" pageNumber="290">FIGURE 3.</emphasis>
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<taxonomicName id="4C774D11FFC410502839F931FDE81CEC" authority="Wagner, 2018" authorityName="Wagner" authorityYear="2018" box="[286,558,1747,1769]" class="Insecta" family="Psychodidae" genus="Theischingerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="290" phylum="Arthropoda" rank="species" species="imbricatus" status="sp. nov.">
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<emphasis id="B903EA80FFC410502839F931FDE81CEC" box="[286,558,1747,1769]" italics="true" pageId="5" pageNumber="290">Theischingerus imbricatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057FBFFC410502B11F931FD4F1CEC" box="[566,649,1747,1769]" pageId="5" pageNumber="290" rank="species">
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<emphasis id="B903EA80FFC410502B11F931FD4F1CEC" bold="true" box="[566,649,1747,1769]" pageId="5" pageNumber="290">sp. nov.</emphasis>
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</taxonomicNameLabel>
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a. wing; b. wing base with expandable organ; c. genitalia, ventral view; d. aedeagus with parameral bridge. [Scale: a 1.0 mm, b–d 0.1 mm]
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</paragraph>
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</caption>
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<paragraph id="8BC83692FFC4105329E0F8D6FB2E1B18" blockId="5.[151,1437,1844,2013]" lastBlockId="6.[151,1437,151,719]" lastPageId="6" lastPageNumber="291" pageId="5" pageNumber="290">Abdomen with 8 segments, genitalia inverted; hypandrium broadened in the middle, almost rectangular, surface structure apically as covered with tiles. Gonocoxites tubular, slightly bent, more than twice as long as wide, dorsally connected by a laterally thin, medially expanded parameral bridge; expansion elongate, distally short bilobed, reaching almost the tip of the aedeagus. Gonostyli thin scythe shaped with sharp tip. Aedeagus is a single sclerite as long as gonocoxites; from the thin base it permanently increases in width developing lateral lobes with apically converging tips. Distiphallus sclerites probably missing or fused with basiphallus. Epandrium subquadrate with 1 transverse foramen; proctiger with a large triangular setose hypoproct and a small rhomboid setose epiproct. Surstyli more than twice as long as epandrium, slightly curved, laterally with a line of longer setae at about equal distance, apically with about 15 probably striated retinacula. Subepandial plate widely rectangular.</paragraph>
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</subSubSection>
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<subSubSection id="C36D6519FFC7105329E0FEC5FBC518CA" pageId="6" pageNumber="291" type="discussion">
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<paragraph id="8BC83692FFC7105329E0FEC5FDF51BF2" blockId="6.[151,1437,151,719]" pageId="6" pageNumber="291">
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<emphasis id="B903EA80FFC7105329E0FEC5FE891B45" bold="true" box="[199,335,295,320]" pageId="6" pageNumber="291">Comments.</emphasis>
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The most striking features of
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<taxonomicName id="4C774D11FFC710532B83FEC5FC891B45" authority="Wagner, 2018" authorityName="Wagner" authorityYear="2018" box="[676,847,295,320]" class="Insecta" family="Psychodidae" genus="Theischingerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="289" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B903EA80FFC710532B83FEC5FC891B45" box="[676,847,295,320]" italics="true" pageId="6" pageNumber="291">Theischingerus</emphasis>
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</taxonomicName>
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males are the slight increase of the subcostal region of the wing, a flexible scent organ at the base of R1, and the unusual furcation of Rs so that only R5 is between radial and medial forks. The gonocoxite bears 2–3 long setae basally on inner side; parameral bridge long, thin and distally split. The construction of the aedeagus is comparatively simple, it is only a single elongate pear shaped sclerite. A sclerotized connection of basiphallus and parameral bridge (like the furca in Palearctic
|
||
<taxonomicName id="4C774D11FFC710532DC5FE58FAAB1BD6" box="[1250,1389,442,467]" class="Insecta" family="Psychodidae" genus="Vaillantodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA80FFC710532DC5FE58FAAB1BD6" box="[1250,1389,442,467]" italics="true" pageId="6" pageNumber="291">Vaillantodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D11FFC7105329B0FE3CFEC61BF2" authorityName="Vaillant" authorityYear="1972" box="[151,256,478,503]" class="Insecta" family="Psychodidae" genus="Jungiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA80FFC7105329B0FE3CFEC61BF2" box="[151,256,478,503]" italics="true" pageId="6" pageNumber="291">Jungiella</emphasis>
|
||
</taxonomicName>
|
||
species) is not discernable.
|
||
</paragraph>
|
||
<paragraph id="8BC83692FFC7105329E0FDE0FBC518CA" blockId="6.[151,1437,151,719]" pageId="6" pageNumber="291">
|
||
Extension of the costal wing area is rare in
|
||
<taxonomicName id="4C774D11FFC710532B80FDE0FCFE181E" authorityName="Newman" authorityYear="1834" box="[679,824,514,539]" class="Insecta" family="Psychodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
|
||
; it is known form the extant
|
||
<taxonomicName id="4C774D11FFC710532D52FDE0FEBF183A" authority="Wagner & Andersen, 2007" authorityName="Wagner & Andersen" authorityYear="2007" class="Insecta" family="Psychodidae" genus="Threticus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="saetheri">
|
||
<emphasis id="B903EA80FFC710532D52FDE0FAF9181E" box="[1141,1343,514,539]" italics="true" pageId="6" pageNumber="291">Threticus saetheri</emphasis>
|
||
<bibRefCitation id="EFE64B63FFC710532C62FDE1FEBF183A" pageId="6" pageNumber="291" refId="ref5862" refString="Wagner, R. & Andersen, T. (2007) Psychodidae (Diptera: Nematocera) from the West Usambara Mountains, Tanzania. In: Andersen, T. (Ed.), Contributions to the Systematics and Ecology of Aquatic Diptera-A Tribute to Ole A. Saether. The Caddis Press, Columbus, OH, pp. 287 - 307." type="book chapter">Wagner & Andersen, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3607602FFC7105328B5FDC4FE3D183A" box="[402,507,550,575]" name="Tanzania" pageId="6" pageNumber="291">Tanzania</collectingCountry>
|
||
, tribus
|
||
<taxonomicName id="4C774D11FFC710532B46FDC4FD24183A" authorityName="Newman" authorityYear="1834" box="[609,738,550,575]" class="Insecta" family="Psychodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="tribe" tribe="Psychodini">Psychodini</taxonomicName>
|
||
), and two species from Caribbean and Mexican Amber (
|
||
<taxonomicName id="4C774D11FFC710532987FDA9FD981866" authority="Wagner, 2001" authorityName="Wagner" authorityYear="2001" box="[160,606,586,611]" class="Insecta" family="Psychodidae" genus="Succinarisemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="scheveni">
|
||
<emphasis id="B903EA80FFC710532987FDA9FE7F1866" box="[160,441,586,611]" italics="true" pageId="6" pageNumber="291">Succinarisemus scheveni</emphasis>
|
||
<bibRefCitation id="EFE64B63FFC7105328E5FDA9FD981866" box="[450,606,587,611]" pageId="6" pageNumber="291" refId="ref5671" refString="Wagner, R. (2001) A remarkable new species and genus of moth-flies (Diptera, Psychodidae) from Dominican amber. Studia Dipterologica, 8, 423 - 426." type="journal article">Wagner, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D11FFC710532BBFFDA9FBCF1866" authority="Coty et al., 2013" authorityName="Coty et al." authorityYear="2013" box="[664,1033,586,611]" class="Insecta" family="Psychodidae" genus="Succinarisemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="totolapensis">
|
||
<emphasis id="B903EA80FFC710532BBFFDA9FCF81866" box="[664,830,586,611]" italics="true" pageId="6" pageNumber="291">S. totolapensis</emphasis>
|
||
<bibRefCitation id="EFE64B63FFC710532A61FDA9FBCF1866" box="[838,1033,586,611]" pageId="6" pageNumber="291" refId="ref4712" refString="Coty, D., Salame, Y., Nel, A. & Azar, D. (2013) A new psychodid fly from Mexican amber (Diptera; Psychodidae). In: Azar, D., Engel, M. S., Jarzembowski, E., Krogmann, L., Nel, A. & Santiago-Blay, J. (Eds.), Insect Evolution in an Amberiferous and Stone Alphabet, Brill, Leiden / Boston, pp. 13 - 26. https: // doi. org / 10.1163 / 9789004210714 _ 003" type="book chapter">
|
||
Coty
|
||
<emphasis id="B903EA80FFC710532AA2FDAEFC711866" box="[901,951,586,611]" italics="true" pageId="6" pageNumber="291">et al</emphasis>
|
||
., 2013
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
; tribus
|
||
<taxonomicName id="4C774D11FFC710532D78FDA8FB121866" authorityName="Enderlein" authorityYear="1937" box="[1119,1236,586,611]" class="Insecta" family="Psychodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
|
||
). Some European
|
||
<taxonomicName id="4C774D11FFC7105329B0FD8CFE891882" authority="Walker" authorityName="Walker" authorityYear="1856" box="[151,335,622,647]" class="Insecta" family="Psychodidae" genus="Ulomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA80FFC7105329B0FD8CFF3E1882" box="[151,248,622,647]" italics="true" pageId="6" pageNumber="291">Ulomyia</emphasis>
|
||
Walker
|
||
</taxonomicName>
|
||
species show morphological variation of the wing area, partly related to probable ‘scent organs’ (e.g.
|
||
<bibRefCitation id="EFE64B63FFC7105329F5FD70FE9318AE" box="[210,341,658,683]" pageId="6" pageNumber="291" refId="ref5244" refString="Elger, M. (1981) The ultrastructure of special epidermal organs in Psychodidae (Diptera). I. Appendages on mesothorax of male Ulomyia fuliginosa (Meigen). Acta zoological Stockholm, 62, 1 - 16. https: // doi. org / 10.1111 / j. 1463 - 6395.1981. tb 00612. x" type="journal article">Elger 1981</bibRefCitation>
|
||
). The mentioned species belong to different tribes and thus it is evident that broadening of the basal costal area has developed independently in several lines in
|
||
<taxonomicName id="4C774D11FFC710532A4AFD54FC3818CA" authorityName="Newman" authorityYear="1834" box="[877,1022,694,719]" class="Insecta" family="Psychodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="291" phylum="Arthropoda" rank="family">Psychodidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |