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<document id="CA74AB18EC036155241ABBCADE38FA3B" ID-DOI="10.5281/zenodo.5324196" ID-ISSN="0253-116X" ID-Zenodo-Dep="5324196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630148591139" checkinUser="carolina" docAuthor="Assing, V." docDate="2011" docId="03DE8784EF59A425FF5FFF0FFE77FDAC" docLanguage="en" docName="LinzerbiolBeitr.2011.43.1.253-265.pdf" docOrigin="Linzer biologische Beiträge 43 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Leptusa (Neopisalia) microphthalma REITTER 1887" docType="treatment" docVersion="4" lastPageNumber="259" masterDocId="FFE7FFFCEF5DA423FFEDFFD6FF87FFD1" masterDocTitle="New species and additional records of Leptusa from the Caucasus region and the Himalaya (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="265" masterPageNumber="253" pageNumber="257" updateTime="1699164357483" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="30BE41DCEA91D297667BA326B7BB1A13" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="48934E8266E0BEB728D9FB4EF0F8CAFD">
<mods:title id="16C13784AE7293829EA4377D1AD4BA08">New species and additional records of Leptusa from the Caucasus region and the Himalaya (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="EF37DA7CE056362DD6B161C690B6A7E9">Assing, V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="96B882F4C4723B0A336444A665BE709A">text</mods:typeOfResource>
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<mods:title id="51C51C31009A1BCC094AB840B9B8C5E0">Linzer biologische Beiträge</mods:title>
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<mods:date id="B203A5BB991486D624A5DA965C9EA50F">2011</mods:date>
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<mods:number id="EF967A7084C7E188991A559E369AC834">2011-07-25</mods:number>
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<mods:number id="959265D8CE2BEDD59754FFF6136A9F91">43</mods:number>
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<mods:classification id="DD706B099A7A6E518AE808ED5CB39F64">journal article</mods:classification>
<mods:identifier id="4C682B05050F035208B6FF44C9BCF8DC" type="DOI">10.5281/zenodo.5324196</mods:identifier>
<mods:identifier id="0C2A4E9FC3266735170458296ABDFEC7" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="4A5D2CC9E6DCB125FD5B4DFC4DF82F39" type="Zenodo-Dep">5324196</mods:identifier>
<mods:identifier id="54346655FDED370287A59EC86858F688" type="ZooBank">A0546682-3521-434F-B896-D5806F286D2D</mods:identifier>
</mods:mods>
<treatment id="03DE8784EF59A425FF5FFF0FFE77FDAC" LSID="urn:lsid:plazi:treatment:03DE8784EF59A425FF5FFF0FFE77FDAC" httpUri="http://treatment.plazi.org/id/03DE8784EF59A425FF5FFF0FFE77FDAC" lastPageId="6" lastPageNumber="259" pageId="4" pageNumber="257">
<subSubSection id="C36D6519EF59A427FF5FFF0FFC7FFE5E" pageId="4" pageNumber="257" type="nomenclature">
<paragraph id="8BC83692EF59A427FF5FFF0FFCD4FF3F" blockId="4.[178,851,216,238]" box="[178,851,216,238]" pageId="4" pageNumber="257">
<heading id="D08081FEEF59A427FF5FFF0FFCD4FF3F" box="[178,851,216,238]" fontSize="8" level="2" pageId="4" pageNumber="257" reason="2">
<taxonomicName id="4C774D11EF59A427FF5FFF0FFD48FF3F" ID-CoL="722DC" authority="REITTER 1887" authorityName="REITTER" authorityYear="1887" box="[178,719,216,238]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="microphthalma" subGenus="Neopisalia">
<emphasis id="B903EA80EF59A427FF5FFF0FFD48FF3F" bold="true" box="[178,719,216,238]" pageId="4" pageNumber="257">
<emphasis id="B903EA80EF59A427FF5FFF0FFE82FF3F" bold="true" box="[178,261,217,238]" italics="true" pageId="4" pageNumber="257">Leptusa</emphasis>
(
<emphasis id="B903EA80EF59A427FEF9FF0EFE04FF3F" bold="true" box="[276,387,216,238]" italics="true" pageId="4" pageNumber="257">Neopisalia</emphasis>
)
<emphasis id="B903EA80EF59A427FE7FFF0EFDB5FF3F" bold="true" box="[402,562,216,238]" italics="true" pageId="4" pageNumber="257">microphthalma</emphasis>
<bibRefCitation id="EFE64B63EF59A427FDD5FF0EFD12FF3C" author="REITTER E" box="[568,661,216,238]" pageId="4" pageNumber="257" pagination="241 - 288" refId="ref5956" refString="REITTER E. (1887): Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Dritter Theil. - Deutsche Entomologische Zeitschrift 31: 241 - 288." type="journal article" year="1887">REITTER</bibRefCitation>
1887
</emphasis>
</taxonomicName>
(
<figureCitation id="134C2A17EF59A427FD31FF0EFCCCFF3F" box="[732,843,216,238]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Figs 15-26</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFED1FC7FFE5E" blockId="4.[178,1087,263,399]" pageId="4" pageNumber="257">
<materialsCitation id="3B1F3CCFEF59A427FF5FFED1FC80FEE3" collectingDate="1999-06-08" collectionCode="VI" collectorName="Temnolesskaia &amp; Mezmai &amp; Mezmay &amp; Smetana" country="Russia" elevation="800" location="Krasnodarskiy Kray" pageId="4" pageNumber="257" specimenCount="14">
M a t e r i a l e x a m i n e d:
<collectingCountry id="F3607602EF59A427FE1EFED1FDB7FECA" box="[499,560,263,283]" name="Russia" pageId="4" pageNumber="257">Russia</collectingCountry>
:
<location id="8EA86049EF59A427FDD4FED1FD69FECA" LSID="urn:lsid:plazi:treatment:03DE8784EF59A425FF5FFF0FFE77FDAC:8EA86049EF59A427FDD4FED1FD69FECA" box="[569,750,263,283]" country="Russia" name="Krasnodarskiy Kray" pageId="4" pageNumber="257">Krasnodarskiy Kray</location>
:
<specimenCount id="9D71FD1BEF59A427FD1AFED1FCBEFECA" box="[759,825,263,283]" pageId="4" pageNumber="257" type="generic">14 exs.</specimenCount>
,
<collectorName id="26825344EF59A427FCAFFED1FC43FECA" box="[834,964,263,283]" pageId="4" pageNumber="257">Temnolesskaia</collectorName>
near
<collectorName id="26825344EF59A427FC1AFED1FBB8FECA" box="[1015,1087,263,283]" pageId="4" pageNumber="257">Mezmai</collectorName>
[=
<collectorName id="26825344EF59A427FF1CFEC8FEC3FEE3" box="[241,324,286,306]" pageId="4" pageNumber="257">Mezmay</collectorName>
],
<quantity id="4C8F9B77EF59A427FEBBFEC8FE12FEE3" box="[342,405,286,306]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="257" unit="m" value="800.0">
<elevation id="005AD1A1EF59A427FEBBFEC8FE12FEE3" box="[342,405,286,306]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="257" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFC91052EF59A427FE4EFEC8FE7AFEE3" box="[419,509,286,306]" pageId="4" pageNumber="257" value="1999-06-08">
<collectingDate id="EF8DE9BAEF59A427FE4EFEC8FE7AFEE3" box="[419,509,286,306]" pageId="4" pageNumber="257" value="1999-06-08">
8.
<collectionCode id="ED66AE57EF59A427FE5DFEC8FE4DFEE3" box="[432,458,286,306]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="257">VI</collectionCode>
.1999
</collectingDate>
</date>
, leg.
<collectorName id="26825344EF59A427FDD8FEC8FD07FEE3" box="[565,640,286,306]" pageId="4" pageNumber="257">Smetana</collectorName>
(cSme, cAss)
</materialsCitation>
;
<materialsCitation id="3B1F3CCFEF59A427FCF8FEC8FE30FE9B" collectingDate="1999-06-07" collectionCode="VI" collectorName="Temnolesskaia &amp; Mezmai &amp; Mezmay &amp; Smetana" country="Russia" elevation="750" location="Krasnodarskiy Kray" pageId="4" pageNumber="257" specimenCount="10">
<specimenCount id="9D71FD1BEF59A427FCF8FEC8FCDCFEE3" box="[789,859,286,306]" pageId="4" pageNumber="257" type="generic">10 exs.</specimenCount>
, same data, but
<quantity id="4C8F9B77EF59A427FC16FEC8FBBEFEE3" box="[1019,1081,286,306]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="257" unit="m" value="750.0">
<elevation id="005AD1A1EF59A427FC16FEC8FBBEFEE3" box="[1019,1081,286,306]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="257" unit="m" value="750.0">750 m</elevation>
</quantity>
,
<date id="FFC91052EF59A427FF32FEE0FEBFFE9B" box="[223,312,310,330]" pageId="4" pageNumber="257" value="1999-06-07">
<collectingDate id="EF8DE9BAEF59A427FF32FEE0FEBFFE9B" box="[223,312,310,330]" pageId="4" pageNumber="257" value="1999-06-07">
7.
<collectionCode id="ED66AE57EF59A427FF00FEE0FE81FE9B" box="[237,262,310,330]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="257">VI</collectionCode>
.1999
</collectingDate>
</date>
(cSme, cAss)
</materialsCitation>
;
<materialsCitation id="3B1F3CCFEF59A427FE2EFEE0FCE9FE9B" box="[451,878,310,330]" collectingDate="1999-06-08" collectionCode="VI" collectorName="Temnolesskaia &amp; Mezmai &amp; Mezmay &amp; Smetana" country="Russia" elevation="850" location="Krasnodarskiy Kray" pageId="4" pageNumber="257" specimenCount="1">
<specimenCount id="9D71FD1BEF59A427FE2EFEE0FE75FE9B" box="[451,498,310,330]" pageId="4" pageNumber="257" type="generic">1 ex.</specimenCount>
, same data, but
<quantity id="4C8F9B77EF59A427FD6BFEE0FD45FE9B" box="[646,706,310,330]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="257" unit="m" value="850.0">
<elevation id="005AD1A1EF59A427FD6BFEE0FD45FE9B" box="[646,706,310,330]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="257" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="FFC91052EF59A427FD20FEE0FCA2FE9B" box="[717,805,310,330]" pageId="4" pageNumber="257" value="1999-06-08">
<collectingDate id="EF8DE9BAEF59A427FD20FEE0FCA2FE9B" box="[717,805,310,330]" pageId="4" pageNumber="257" value="1999-06-08">
8.
<collectionCode id="ED66AE57EF59A427FD37FEE0FD73FE9B" box="[730,756,310,330]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="257">VI</collectionCode>
.1999
</collectingDate>
</date>
(cSme)
</materialsCitation>
;
<materialsCitation id="3B1F3CCFEF59A427FC94FEE0FE70FEB0" collectingDate="1999-06-09" collectionCode="VI" collectorName="Temnolesskaia &amp; Mezmai &amp; Mezmay &amp; Smetana" country="Russia" elevation="850" location="Krasnodarskiy Kray" pageId="4" pageNumber="257" specimenCount="4">
<specimenCount id="9D71FD1BEF59A427FC94FEE0FC36FE9B" box="[889,945,310,330]" pageId="4" pageNumber="257" type="generic">4 exs.</specimenCount>
, same data, but
<quantity id="4C8F9B77EF59A427FF32FE9BFE9FFEB0" box="[223,280,333,353]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="257" unit="m" value="850.0">
<elevation id="005AD1A1EF59A427FF32FE9BFE9FFEB0" box="[223,280,333,353]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="257" unit="m" value="850.0">850 m</elevation>
</quantity>
,
<date id="FFC91052EF59A427FECFFE9BFEFCFEB0" box="[290,379,333,353]" pageId="4" pageNumber="257" value="1999-06-09">
<collectingDate id="EF8DE9BAEF59A427FECFFE9BFEFCFEB0" box="[290,379,333,353]" pageId="4" pageNumber="257" value="1999-06-09">
9.
<collectionCode id="ED66AE57EF59A427FEC2FE9BFECFFEB0" box="[303,328,333,353]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="257">VI</collectionCode>
.1999
</collectingDate>
</date>
(cSme, cAss)
</materialsCitation>
;
<specimenCount id="9D71FD1BEF59A427FDECFE9BFDC5FEB0" box="[513,578,333,353]" pageId="4" pageNumber="257" type="generic">40 exs.</specimenCount>
, Guama near Mezmai,
<quantity id="4C8F9B77EF59A427FCE2FE9BFCD3FEB0" box="[783,852,333,353]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="257" unit="m" value="1000.0">1000 m</quantity>
,
<date id="FFC91052EF59A427FCB3FE9BFC45FEB0" box="[862,962,333,353]" pageId="4" pageNumber="257" value="1999-06-10">10.VI.1999</date>
, leg. Smetana (cSme, cAss);
<materialsCitation id="3B1F3CCFEF59A427FE88FEB2FEE7FE5E" collectionCode="SDEI" collectorName="Leder &amp; Reitter" country="Russia" pageId="4" pageNumber="257" specimenCount="9">
<specimenCount id="9D71FD1BEF59A427FE88FEB2FE11FEA9" box="[357,406,356,376]" pageId="4" pageNumber="257" type="generic">1 ex.</specimenCount>
, &quot;Kuban&quot;, leg.
<collectorName id="26825344EF59A427FDCEFEB2FDD1FEA9" box="[547,598,356,376]" pageId="4" pageNumber="257">Leder</collectorName>
&amp;
<collectorName id="26825344EF59A427FD94FEB2FD33FEA9" box="[633,692,356,376]" pageId="4" pageNumber="257">Reitter</collectorName>
(
<collectionCode id="ED66AE57EF59A427FD2EFEB2FD7FFEA9" box="[707,760,356,376]" pageId="4" pageNumber="257">SDEI</collectionCode>
).
<specimenCount id="9D71FD1BEF59A427FCE4FEB2FCC5FEA9" box="[777,834,356,376]" pageId="4" pageNumber="257" type="generic">8 exs.</specimenCount>
, &quot;Circassien&quot;, leg.
<collectorName id="26825344EF59A427FC1DFEB2FBA4FEA9" box="[1008,1059,356,376]" pageId="4" pageNumber="257">Leder</collectorName>
&amp;
<collectorName id="26825344EF59A427FF32FEADFE9DFE5E" box="[223,282,379,399]" pageId="4" pageNumber="257">Reitter</collectorName>
(
<collectionCode id="ED66AE57EF59A427FEC8FEADFEDDFE5E" box="[293,346,379,399]" pageId="4" pageNumber="257">SDEI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F3CCFEF59A427FE84FEADFDC3FE5E" box="[361,580,379,399]" collectionCode="SDEI" country="Russia" pageId="4" pageNumber="257" specimenCount="1">
<specimenCount id="9D71FD1BEF59A427FE84FEADFE11FE5E" box="[361,406,379,399]" pageId="4" pageNumber="257" type="generic">1 ex.</specimenCount>
, &quot;Circassia&quot; (
<collectionCode id="ED66AE57EF59A427FDE4FEADFDB9FE5E" box="[521,574,379,399]" pageId="4" pageNumber="257">SDEI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F3CCFEF59A427FDA0FEADFC73FE5E" box="[589,1012,379,399]" collectionCode="SDEI" collectorName="Leder &amp; Reitter" country="Russia" pageId="4" pageNumber="257" specimenCount="2">
<specimenCount id="9D71FD1BEF59A427FDA0FEADFD05FE5E" box="[589,642,379,399]" pageId="4" pageNumber="257" type="generic">2 exs.</specimenCount>
, &quot;Swanetien&quot;, leg.
<collectorName id="26825344EF59A427FCC9FEADFCD0FE5E" box="[804,855,379,399]" pageId="4" pageNumber="257">Leder</collectorName>
&amp;
<collectorName id="26825344EF59A427FC9EFEADFC29FE5E" box="[883,942,379,399]" pageId="4" pageNumber="257">Reitter</collectorName>
(
<collectionCode id="ED66AE57EF59A427FC54FEADFC69FE5E" box="[953,1006,379,399]" pageId="4" pageNumber="257">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D6519EF59A425FF5FFE48FE77FDAC" lastPageId="6" lastPageNumber="259" pageId="4" pageNumber="257" type="description">
<paragraph id="8BC83692EF59A427FF5FFE48FD67FD50" blockId="4.[178,1086,414,641]" pageId="4" pageNumber="257">
C o m m e n t: The original description is based on an unspecified number of
<typeStatus id="54CC8830EF59A427FC09FE48FBBAFE62" box="[996,1085,414,435]" pageId="4" pageNumber="257" type="syntype">syntypes</typeStatus>
from &quot;Utsch-Dere in Circassien. Von Herrn Eugen König entdeckt&quot; (
<bibRefCitation id="EFE64B63EF59A427FC74FE6DFBB6FE00" author="REITTER E" box="[921,1073,443,465]" pageId="4" pageNumber="257" pagination="241 - 288" refId="ref5956" refString="REITTER E. (1887): Neue Coleopteren aus Europa, den angrenzenden Landern und Sibirien, mit Bemerkungen uber bekannte Arten. Dritter Theil. - Deutsche Entomologische Zeitschrift 31: 241 - 288." type="journal article" year="1887">REITTER 1887</bibRefCitation>
).
<bibRefCitation id="EFE64B63EF59A427FF5FFE0FFEB4FE3F" author="PACE R" box="[178,307,472,494]" pageId="4" pageNumber="257" pagination="1 - 307" refId="ref5908" refString="PACE R. (1989): Monografia del genere Leptusa KRAATZ (Coleoptera Staphylinidae). - Memorie del Museo Civico di Storia Naturale di Verona (II ° Serie), Sezione Scienze della Vita (A: Biologica) 8: 1 - 307." type="journal article" year="1989">PACE (1989)</bibRefCitation>
designated a female
<typeStatus id="54CC8830EF59A427FDFCFE0FFDE7FE3F" box="[529,608,473,494]" pageId="4" pageNumber="257" type="syntype">syntype</typeStatus>
as the
<typeStatus id="54CC8830EF59A427FD40FE0EFC8AFE3F" box="[685,781,472,494]" pageId="4" pageNumber="257" type="lectotype">lectotype</typeStatus>
. Based on the illustrations of the aedeagus of
<taxonomicName id="4C774D11EF59A427FEB5FE20FD8AFDDD" authorityName="REITTER" authorityYear="1887" box="[344,525,502,524]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="257" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="B903EA80EF59A427FEB5FE20FD8AFDDD" box="[344,525,502,524]" italics="true" pageId="4" pageNumber="257">L. microphthalma</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFE64B63EF59A427FD78FE20FC9FFDDD" author="PACE R" box="[661,792,502,524]" pageId="4" pageNumber="257" pagination="1 - 307" refId="ref5908" refString="PACE R. (1989): Monografia del genere Leptusa KRAATZ (Coleoptera Staphylinidae). - Memorie del Museo Civico di Storia Naturale di Verona (II ° Serie), Sezione Scienze della Vita (A: Biologica) 8: 1 - 307." type="journal article" year="1989">PACE (1989)</bibRefCitation>
, the recently collected material listed above was at first assumed to represent a distinct species. A subsequent comparison with old material from &quot;Circassia&quot; deposited in the collections of the SDEI, however, revealed that they are conspecific. In order to facilitate future identification, a redescription and new illustrations are provided below.
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFD47FEC1FCCA" blockId="4.[178,1086,656,795]" pageId="4" pageNumber="257">
R e d e s c r i p t i o n: Body length
<quantity id="4C8F9B77EF59A427FDA6FD46FD40FD77" box="[587,711,656,678]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="4" pageNumber="257" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.6-3.5 mm</quantity>
. Habitus as in
<figureCitation id="134C2A17EF59A427FC81FD46FC3CFD77" box="[876,955,656,678]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 15</figureCitation>
. Coloration: forebody yellowish-brown to reddish-brown; abdomen brown to dark-brown, with the posterior margins of the segments and the apex (posterior portion of segment VII, segments VIII-X) yellowish to yellowish-brown; legs and antennae dark-yellowish to yellowish-brown.
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFCFCFE7FFBFA" blockId="4.[178,1086,810,1067]" pageId="4" pageNumber="257">
Head (
<figureCitation id="134C2A17EF59A427FF15FCFCFEC4FC91" box="[248,323,810,832]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 16</figureCitation>
) weakly transverse; punctation fine and sparse, barely visible in the pronounced microsculpture (
<figureCitation id="134C2A17EF59A427FE5AFC9EFD82FC8F" box="[439,517,840,862]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 17</figureCitation>
); dorsal surface with subdued shine; eyes weakly convex, not projecting from lateral contours of head, and small, less than half the length of postocular portion in dorsal view. Antenna slender (
<figureCitation id="134C2A17EF59A427FD08FC54FCB2FC49" box="[741,821,898,920]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 18</figureCitation>
), approximately
<quantity id="4C8F9B77EF59A427FC06FC54FBB9FC49" box="[1003,1086,898,920]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="257" unit="mm" value="0.9">0.9 mm</quantity>
long; antennomeres I-III distinctly oblong and of subequal length; IV approximately as long as wide or indistinctly oblong; V as wide as long or weakly transverse; VI-X gradually increasing in width and increasingly transverse; X approximately 1.5 times as wide as long, or nearly so. Maxillary palpi moderately slender; preapical palpomere approximately 3 times as long as broad.
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFBEDFE7CFB79" blockId="4.[178,1086,1082,1192]" pageId="4" pageNumber="257">
Pronotum (
<figureCitation id="134C2A17EF59A427FECAFBECFEF2FB81" box="[295,373,1082,1104]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 16</figureCitation>
) approximately 1.25 times as wide as long and 1.20-1.25 times as wide as head; maximal width slightly before middle; posterior angles obtusely marked; punctation fine and rather sparse, barely visible in the pronounced microsculpture (
<figureCitation id="134C2A17EF59A427FBF9FBA2FF49FB79" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 19</figureCitation>
); surface with subdued shine.
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFB61FE32FA8E" blockId="4.[178,1085,1207,1375]" pageId="4" pageNumber="257">
Elytra (
<figureCitation id="134C2A17EF59A427FEECFB61FECAFB1C" box="[257,333,1207,1229]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="4" pageNumber="257">Fig. 16</figureCitation>
) short, approximately 0.75 times as long as pronotum; posterior margin distinctly sinuate near postero-lateral angles; punctation much more pronounced than that of head and pronotum; interstices more shiny than those of head and pronotum, with shallow microsculpture composed of larger meshes than that of head and pronotum. Hind wings reduced. Legs moderately slender; metatarsomere I as long as the combined length of II and III, or nearly so.
</paragraph>
<paragraph id="8BC83692EF59A427FF5FFAB9FCB2FA0D" blockId="4.[178,1085,1391,1500]" pageId="4" pageNumber="257">Abdomen 1.15-1.20 times as wide as elytra, widest at segments V/VI; tergites III-V with rather shallow and finely punctate anterior impressions; punctation fine, moderately dense on anterior and rather sparse on posterior tergites; posterior margin of tergite VII without palisade fringe; tergite VII without sexual dimorphism.</paragraph>
<caption id="DF08661AEF58A426FF5FFA73FF79F9C5" ID-DOI="http://doi.org/10.5281/zenodo.5324202" ID-Zenodo-Dep="5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="5" pageNumber="258" startId="5.[178,218,1445,1465]" targetBox="[173,1066,210,1415]" targetPageId="5">
<paragraph id="8BC83692EF58A426FF5FFA73FF79F9C5" blockId="5.[178,1086,1445,1556]" pageId="5" pageNumber="258">
<emphasis id="B903EA80EF58A426FF5FFA73FE9DFA68" bold="true" box="[178,282,1445,1465]" pageId="5" pageNumber="258">Figs 15-26</emphasis>
:
<taxonomicName id="4C774D11EF58A426FEC7FA73FDE6FA66" authority="REITTER" authorityName="REITTER" authorityYear="1887" box="[298,609,1445,1465]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="258" phylum="Arthropoda" rank="species" species="microphthalma">
<emphasis id="B903EA80EF58A426FEC7FA73FD8FFA68" box="[298,520,1445,1465]" italics="true" pageId="5" pageNumber="258">Leptusa microphthalma</emphasis>
REITTER
</taxonomicName>
: (
<emphasis id="B903EA80EF58A426FD97FA73FD16FA68" bold="true" box="[634,657,1445,1465]" pageId="5" pageNumber="258">15</emphasis>
) habitus; (
<emphasis id="B903EA80EF58A426FD11FA73FC94FA68" bold="true" box="[764,787,1445,1465]" pageId="5" pageNumber="258">16</emphasis>
) forebody; (
<emphasis id="B903EA80EF58A426FC62FA73FC21FA68" bold="true" box="[911,934,1445,1465]" pageId="5" pageNumber="258">17</emphasis>
) median dorsal portion of head; (
<emphasis id="B903EA80EF58A426FEB8FA6DFEEBFA1E" bold="true" box="[341,364,1467,1487]" pageId="5" pageNumber="258">18</emphasis>
) antenna; (
<emphasis id="B903EA80EF58A426FE3BFA6DFE6AFA1E" bold="true" box="[470,493,1467,1487]" pageId="5" pageNumber="258">19</emphasis>
) median portion of pronotum; (
<emphasis id="B903EA80EF58A426FCF8FA6DFCABFA1E" bold="true" box="[789,812,1467,1487]" pageId="5" pageNumber="258">20</emphasis>
) male tergite VIII; (
<emphasis id="B903EA80EF58A426FC06FA6DFB85FA1E" bold="true" box="[1003,1026,1467,1487]" pageId="5" pageNumber="258">21</emphasis>
) male sternite VIII; (
<emphasis id="B903EA80EF58A426FEDBFA04FEEDFA37" bold="true" box="[310,362,1490,1510]" pageId="5" pageNumber="258">22-23</emphasis>
) median lobe of aedeagus in lateral view; (
<emphasis id="B903EA80EF58A426FD1AFA04FC89FA37" bold="true" box="[759,782,1490,1510]" pageId="5" pageNumber="258">24</emphasis>
) female tergite VIII; (
<emphasis id="B903EA80EF58A426FC36FA04FC75FA37" bold="true" box="[987,1010,1490,1510]" pageId="5" pageNumber="258">25</emphasis>
) female sternite VIII; (
<emphasis id="B903EA80EF58A426FED8FA3FFECBFA2C" bold="true" box="[309,332,1513,1533]" pageId="5" pageNumber="258">26</emphasis>
) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC83692EF5BA425FF2AFF0DFC99FE52" blockId="6.[178,1085,219,387]" pageId="6" pageNumber="259">
: posterior margin of sternite VII broadly and rather weakly concave, on either side of this concavity with two very long marginal setae; posterior margin of tergite VIII indistinctly concave in the middle (
<figureCitation id="134C2A17EF5BA425FE1BFEC3FDC1FEFA" box="[502,582,277,299]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Fig. 20</figureCitation>
); sternite VIII transverse, posterior margin convexly produced in the middle (
<figureCitation id="134C2A17EF5BA425FE14FEE4FDCFFE99" box="[505,584,306,328]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Fig. 21</figureCitation>
); aedeagus ca.
<quantity id="4C8F9B77EF5BA425FD06FEE4FCCFFE99" box="[747,840,306,328]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="259" unit="mm" value="0.48">0.48 mm</quantity>
long, with pronounced additional projection of somewhat variable shape at the base of the ventral process (
<figureCitation id="134C2A17EF5BA425FBFDFE86FF76FE52" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Figs 22-23</figureCitation>
); apical lobe of paramere slender and moderately long.
</paragraph>
<paragraph id="8BC83692EF5BA425FF2AFE45FBBAFE79" blockId="6.[178,1085,402,453]" box="[199,1085,402,424]" pageId="6" pageNumber="259">
: posterior margin of tergite VIII weakly convex (
<figureCitation id="134C2A17EF5BA425FD28FE44FC94FE79" box="[709,787,402,424]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Fig. 24</figureCitation>
); posterior margin of sternite
</paragraph>
<paragraph id="8BC83692EF5BA425FF5FFE66FC37FE14" blockId="6.[178,1085,402,453]" box="[178,944,431,453]" pageId="6" pageNumber="259">
VIII somewhat produced in the middle (
<figureCitation id="134C2A17EF5BA425FDA4FE79FD13FE14" box="[585,660,431,453]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Fig. 25</figureCitation>
); spermatheca as in
<figureCitation id="134C2A17EF5BA425FC8CFE79FC37FE14" box="[865,944,431,453]" captionStart="Figs 15-26" captionStartId="5.[178,218,1445,1465]" captionTargetBox="[173,1066,210,1415]" captionTargetId="figure-91@5.[169,1076,207,1417]" captionTargetPageId="5" captionText="Figs 15-26: Leptusa microphthalma REITTER: (15) habitus; (16) forebody; (17) median dorsal portion of head; (18) antenna; (19) median portion of pronotum; (20) male tergite VIII; (21) male sternite VIII; (22-23) median lobe of aedeagus in lateral view; (24) female tergite VIII; (25) female sternite VIII; (26) spermatheca. Scale bars: 15: 1.0 mm; 16: 0.5 mm; 18, 20-25: 0.2 mm; 17, 19, 26: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5324202" httpUri="https://zenodo.org/record/5324202/files/figure.png" pageId="6" pageNumber="259">Fig. 26.</figureCitation>
</paragraph>
<paragraph id="8BC83692EF5BA425FF5FFE03FE77FDAC" blockId="6.[178,1086,469,637]" pageId="6" pageNumber="259">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The species is rather widespread in the West Caucasus region. The recently collected specimens were collected near Mezmay, at altitudes of
<quantity id="4C8F9B77EF5BA425FE1AFDD9FDFCFDF4" box="[503,635,527,549]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="6" pageNumber="259" unit="m" value="875.0" valueMax="1000.0" valueMin="750.0">750-1000 m</quantity>
, together with
<taxonomicName id="4C774D11EF5BA425FCC9FDC6FC1FFDF4" authorityName="ROUBAL" authorityYear="1917" box="[804,920,527,549]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="storkani">
<emphasis id="B903EA80EF5BA425FCC9FDC6FC1FFDF4" box="[804,920,527,549]" italics="true" pageId="6" pageNumber="259">L. storkani</emphasis>
</taxonomicName>
,
<taxonomicName id="4C774D11EF5BA425FC47FDC6FB92FDF4" authorityName="ASSING" authorityYear="2009" box="[938,1045,528,549]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="persica">
<emphasis id="B903EA80EF5BA425FC47FDC6FB92FDF4" box="[938,1045,528,549]" italics="true" pageId="6" pageNumber="259">L. persica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C774D11EF5BA425FBC5FDC6FF78FD93" baseAuthorityName="HOCHHUTH" baseAuthorityYear="1849" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="venusta">
<emphasis id="B903EA80EF5BA425FBC5FDC6FF78FD93" italics="true" pageId="6" pageNumber="259">L. venusta</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C774D11EF5BA425FED1FDFBFE2FFD92" authorityName="Assing" authorityYear="2011" box="[316,424,557,579]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="gibbera">
<emphasis id="B903EA80EF5BA425FED1FDFBFE2FFD92" box="[316,424,557,579]" italics="true" pageId="6" pageNumber="259">L. gibbera</emphasis>
</taxonomicName>
. They were sifted from deep leaf litter in mature, mixed beech and fir forests, partly near a stream, and from moist to wet leaf litter in a mature oak forest (SMETANA pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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