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<document id="F7BF5F8A8ADC1C59C1269F4120B4F1C2" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FFB5FFE6FF43FBA7FEFFF8D4" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus fultus A. Beek 2021, nothosp. nov." docType="treatment" docVersion="5" lastPageNumber="60" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="59" updateTime="1687457424327" updateUser="ExternalLinkService">
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<mods:title id="2F874D554D6AAC71CB612ED16F09D5C1">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
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<mods:namePart id="77C0A728CFB1F9920CD683B755BF5E10">Beek, Abraham Van De</mods:namePart>
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<mods:title id="CCEA8E01948E17646E84E690A1D61014">Phytotaxa</mods:title>
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<mods:part id="F439CC96AA1D242B113BC9BB5CC2BB2E">
<mods:date id="576CB60DC32491A5999C1B42FA5D365B">2021</mods:date>
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<mods:classification id="E592C1FFD29408E4A29369D233DF5D29">journal article</mods:classification>
<mods:identifier id="FFD22069E5F2FE29C184E1DDA0DA9C98" type="DOI">10.11646/phytotaxa.515.1.1</mods:identifier>
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<treatment id="03DE3646FFB5FFE6FF43FBA7FEFFF8D4" ID-DOI="http://doi.org/10.5281/zenodo.8071594" ID-Zenodo-Dep="8071594" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4" httpUri="http://treatment.plazi.org/id/03DE3646FFB5FFE6FF43FBA7FEFFF8D4" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
<subSubSection id="C36DD4DBFFB5FFE7FF43FBA7FDC3FBF2" box="[136,597,1094,1121]" pageId="58" pageNumber="59" type="nomenclature">
<paragraph id="8BC88750FFB5FFE7FF43FBA7FDC3FBF2" blockId="58.[136,1452,1094,1192]" box="[136,597,1094,1121]" pageId="58" pageNumber="59">
<emphasis id="B9035B42FFB5FFE7FF43FBA7FF3DFBF3" bold="true" box="[136,171,1094,1120]" pageId="58" pageNumber="59">33.</emphasis>
<taxonomicName id="4C77FCD3FFB5FFE7FF78FBA7FE23FBF3" ID-CoL="8RTD9" authority="A.Beek" authorityName="A. Beek" authorityYear="2021" box="[179,437,1094,1121]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" isHybrid="true" kingdom="Plantae" order="Rosales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="fultus" status="nothosp. nov.">
<emphasis id="B9035B42FFB5FFE7FF78FBA7FECFFBF2" bold="true" box="[179,345,1094,1121]" italics="true" pageId="58" pageNumber="59">Rubus ×fultus</emphasis>
A.Beek
</taxonomicName>
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<emphasis id="B9035B42FFB5FFE7FE76FBA7FDD8FBF3" box="[445,590,1094,1120]" italics="true" pageId="58" pageNumber="59">nothosp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB5FFE7FD96FBA7FCF2FBF3" box="[605,868,1094,1121]" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BC88750FFB5FFE7FD96FBA7FCF2FBF3" blockId="58.[136,1452,1094,1192]" box="[605,868,1094,1121]" pageId="58" pageNumber="59">
(
<taxonomicName id="4C77FCD3FFB5FFE7FDACFBA6FD5BFBF3" authorityName="Weihe &amp; Nees" authorityYear="1822" box="[615,717,1094,1120]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B9035B42FFB5FFE7FDACFBA6FD5BFBF3" box="[615,717,1094,1120]" italics="true" pageId="58" pageNumber="59">R. affinis</emphasis>
</taxonomicName>
<taxonomicName id="4C77FCD3FFB5FFE7FD1DFBA6FCD8FBF3" box="[726,846,1095,1121]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" isHybrid="true" kingdom="Plantae" order="Rosales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="pinnatus">
×
<emphasis id="B9035B42FFB5FFE7FD25FBA6FCD8FBF3" box="[750,846,1095,1120]" italics="true" pageId="58" pageNumber="59">pinnatus</emphasis>
</taxonomicName>
?)
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB5FFE7FCA7FBA7FF32FB3B" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BC88750FFB5FFE7FCA7FBA7FF32FB3B" blockId="58.[136,1452,1094,1192]" pageId="58" pageNumber="59">
<materialsCitation id="3B1F8D0DFFB5FFE7FCA7FBA7FF32FB3B" collectingDate="2018-10-17" collectionCode="L" collectorName="Beek" country="South Africa" location="Uniondale" municipality="Knysna" pageId="58" pageNumber="59" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
Type:—
<collectingCountry id="F360C7C0FFB5FFE7FC0CFBA7FB00FBF3" box="[967,1174,1094,1120]" name="South Africa" pageId="58" pageNumber="59">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FFB5FFE7FB69FBA6FAD5FBF3" box="[1186,1347,1094,1121]" country="South Africa" name="Western Cape" pageId="58" pageNumber="59">Western Cape</collectingRegion>
:
<collectingMunicipality id="6BAC1D2AFFB5FFE7FA84FBA6FA31FBF2" box="[1359,1447,1095,1121]" pageId="58" pageNumber="59">Knysna</collectingMunicipality>
, road to
<location id="8EA8D18BFFB5FFE7FF2BFB8BFECEFB17" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4:8EA8D18BFFB5FFE7FF2BFB8BFECEFB17" box="[224,344,1130,1156]" country="South Africa" municipality="Knysna" name="Uniondale" pageId="58" pageNumber="59" stateProvince="Western Cape">Uniondale</location>
, at the end of the town,
<date id="FFC9A190FFB5FFE7FDA7FB8BFCB8FB17" box="[620,814,1130,1156]" pageId="58" pageNumber="59" value="2018-10-17">
<collectingDate id="EF8D5878FFB5FFE7FDA7FB8BFCB8FB17" box="[620,814,1130,1156]" pageId="58" pageNumber="59" value="2018-10-17">17 October 2018</collectingDate>
</date>
,
<emphasis id="B9035B42FFB5FFE7FCF1FB8BFC74FB17" box="[826,994,1130,1156]" italics="true" pageId="58" pageNumber="59">
<collectorName id="2682E286FFB5FFE7FCF1FB8BFCE6FB17" box="[826,880,1130,1156]" pageId="58" pageNumber="59">Beek</collectorName>
2018.102
</emphasis>
(
<typeStatus id="54CC39F2FFB5FFE7FC39FB8BFBC0FB17" box="[1010,1110,1130,1156]" pageId="58" pageNumber="59">holotype</typeStatus>
<collectionCode id="ED661F95FFB5FFE7FB95FB8AFBE4FB16" box="[1118,1138,1131,1157]" pageId="58" pageNumber="59">L</collectionCode>
; isotype NBG). (
<figureCitation id="134C9BD5FFB5FFE7FAF8FB8BFF0DFB3B" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Figs 25A B</figureCitation>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB5FFE7FF43FB37FE27FAC8" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8BC88750FFB5FFE7FF43FB37FE27FAC8" blockId="58.[136,1452,1238,2040]" pageId="58" pageNumber="59">
<emphasis id="B9035B42FFB5FFE7FF43FB37FE93FB63" bold="true" box="[136,261,1238,1264]" pageId="58" pageNumber="59">Diagnosis:</emphasis>
—Leaves with strong anthocyanin colouring when young, digitate-pinnate 4- to 5-foliolate, abaxially with only few short hairs on main veins; inflorescence large, broad, with only few simple leaves at base, with numerous strong hooked prickles, thinly hairy; petals
<quantity id="4C8F2AB5FFB5FFE7FDBDFAFFFD67FAAB" box="[630,753,1310,1336]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="58" pageNumber="59" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
long, light pink to almost white, broad ovate to suborbicular, longer than sepals. Sterile.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB5FFE7FF76FA84FCBDF847" pageId="58" pageNumber="59" type="description">
<paragraph id="8BC88750FFB5FFE7FF76FA84FCBDF847" blockId="58.[136,1452,1238,2040]" pageId="58" pageNumber="59">
<emphasis id="B9035B42FFB5FFE7FF76FA84FEAEFAED" box="[189,312,1381,1406]" italics="true" pageId="58" pageNumber="59">Primocane</emphasis>
(
<figureCitation id="134C9BD5FFB5FFE7FE83FA85FE59FAEC" box="[328,463,1380,1407]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Fig. 25CD</figureCitation>
) arching, diam.
<quantity id="4C8F2AB5FFB5FFE7FD4EFA85FD72FAED" box="[645,740,1380,1407]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="58" pageNumber="59" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
, (bluntly) angular, eglandular or with few very scattered sessile glands, glabrous; prickles 410 per
<quantity id="4C8F2AB5FFB5FFE7FDD7FA66FDC1FA31" box="[540,599,1415,1442]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="58" pageNumber="59" unit="cm" value="5.0">5 cm</quantity>
, from
<quantity id="4C8F2AB5FFB5FFE7FD69FA66FC96FA31" box="[674,768,1415,1442]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="58" pageNumber="59" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
wide base compressed, curved, up to
<quantity id="4C8F2AB5FFB5FFE7FB64FA66FA9BFA31" box="[1199,1293,1415,1442]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="58" pageNumber="59" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long; stipules
<quantity id="4C8F2AB5FFB5FFE7FF43FA4BFF61FA56" box="[136,247,1450,1477]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="58" pageNumber="59" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">813 mm</quantity>
, linear, with some hairs and short stipitate glands.
<emphasis id="B9035B42FFB5FFE7FCF0FA4AFC1FFA57" box="[827,905,1451,1476]" italics="true" pageId="58" pageNumber="59">Leaves</emphasis>
with strong anthocyanin colouring when young (
<figureCitation id="134C9BD5FFB5FFE7FF5BFA2CFF60FA7B" box="[144,246,1485,1512]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Fig. 25E</figureCitation>
), digitate-pinnate 4- to 5-foliolate, sometimes intermixed with trifoliolate one, or with extra leaflet at base, adaxially glabrous, abaxially (
<figureCitation id="134C9BD5FFB5FFE7FE14FA10FDD4F998" box="[479,578,1520,1547]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Fig. 25F</figureCitation>
) with only few short hairs on main veins; serrature moderate, rather regular, with somewhat obtuse triangular mucronate teeth, straight; petiole
<quantity id="4C8F2AB5FFB5FFE7FC5EF9F5FC7AF9BD" box="[917,1004,1556,1582]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="58" pageNumber="59" unit="cm" value="4.5" valueMax="6.0" valueMin="3.0">36 cm</quantity>
long, with some hairs, few subsessile glands, 69 curved or hooked prickles; basal leaflets
<quantity id="4C8F2AB5FFB5FFE7FD18F9D6FCB9F9C1" box="[723,815,1591,1618]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="58" pageNumber="59" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
stalked; central leaflet from rounded or slightly attenuate base
<quantity id="4C8F2AB5FFB5FFE7FF09F9BBFEABF9E6" box="[194,317,1626,1653]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" metricValueMax="9.3" metricValueMin="5.9" pageId="58" pageNumber="59" unit="mm" value="76.0" valueMax="93.0" valueMin="59.0">5993 mm</quantity>
long, in main
<taxonomicName id="4C77FCD3FFB5FFE7FE2DF9BBFD1CF9E7" box="[486,650,1626,1652]" form="elliptical" pageId="58" pageNumber="59" rank="form">form elliptical</taxonomicName>
, tending to (ob-)ovate or rhombic, gradually attenuate into rather long tip, widthlength index (0.50)0.600.67(0.78), length of petiolule (11)1629% of length of leaflet.
<emphasis id="B9035B42FFB5FFE7FAFCF99CFF41F928" italics="true" pageId="58" pageNumber="59">Flowering branch</emphasis>
(
<figureCitation id="134C9BD5FFB5FFE7FF2DF940FEE2F928" box="[230,372,1696,1723]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Figs 25GH</figureCitation>
) angular, in basal part (almost) glabrous, apically with some long hairs and (sub-)sessile glands; prickles 45 per
<quantity id="4C8F2AB5FFB5FFE7FE8DF925FE17F94D" box="[326,385,1732,1759]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="58" pageNumber="59" unit="cm" value="5.0">5 cm</quantity>
, from
<quantity id="4C8F2AB5FFB5FFE7FE07F925FDBCF94D" box="[460,554,1732,1759]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="58" pageNumber="59" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
broad base, strongly curved or hooked, up to
<quantity id="4C8F2AB5FFB5FFE7FBFFF925FB04F94D" box="[1076,1170,1732,1759]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="58" pageNumber="59" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
long.
<emphasis id="B9035B42FFB5FFE7FB11F924FABEF94D" box="[1242,1320,1733,1758]" italics="true" pageId="58" pageNumber="59">Leaves</emphasis>
3-foliolate, apical ones simple; hairs and serrature as for primocane leaves.
<emphasis id="B9035B42FFB5FFE7FCA9F906FC6EF892" box="[866,1016,1767,1793]" italics="true" pageId="58" pageNumber="59">Inflorescence</emphasis>
(
<figureCitation id="134C9BD5FFB5FFE7FBC2F906FBF3F891" box="[1033,1125,1767,1794]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Fig. 25I</figureCitation>
) large, broad, with only few simple leaves at base, with ascendant peduncles, numerous strong hooked prickles, thinly hairy; pedicels up to
<quantity id="4C8F2AB5FFB5FFE7FA9CF8EBFA31F8B6" box="[1367,1447,1802,1829]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="58" pageNumber="59" unit="mm" value="15.0">15 mm</quantity>
, thinly tomentose and with appressed long hairs, with 510 curved pricklets and few sessile glands.
<emphasis id="B9035B42FFB5FFE7FB30F8CCFAC1F8D4" box="[1275,1367,1837,1863]" italics="true" pageId="58" pageNumber="59">Flowers</emphasis>
: sepals loosely reflexed to patent, 34 ×
<quantity id="4C8F2AB5FFB5FFE7FE31F8B0FDF0F8F8" box="[506,614,1873,1899]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="58" pageNumber="59" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">712 mm</quantity>
, gradually attenuate into long tip, short greyish green tomentose and hairy, with some sessile glands; petals (
<figureCitation id="134C9BD5FFB5FFE7FDCFF895FDF4F81C" box="[516,610,1908,1935]" captionStart="FIGURE 25" captionStartId="59.[136,229,1451,1473]" captionTargetBox="[151,1436,190,1426]" captionTargetId="figure-17@59.[151,1436,190,1426]" captionTargetPageId="59" captionText="FIGURE 25. Rubus ×fultus. A. Holotype, primocane. B. Holotype, inflorescence. C. Primocane with leaves. D. Primocane. E. Young leaves. F. Leaf abaxially. G. Flowering branch. H. Flowering branch. I. Inflorescence. J. Flowers. Photographs:A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="58" pageNumber="59">Fig. 25J</figureCitation>
)
<quantity id="4C8F2AB5FFB5FFE7FDB8F895FD7BF81D" box="[627,749,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="58" pageNumber="59" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">1013 mm</quantity>
long, light pink to almost white, broad ovate to suborbicular, sometimes with emarginate tip, longer than sepals; stamens longer than styles; anthers glabrous; styles red; ovaries almost glabrous or with short hairs; receptacle hairy. Sterile.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB5FFE7FF76F83FFE0CF86B" box="[189,410,2014,2040]" pageId="58" pageNumber="59" type="biology_ecology">
<paragraph id="8BC88750FFB5FFE7FF76F83FFE0CF86B" blockId="58.[136,1452,1238,2040]" box="[189,410,2014,2040]" pageId="58" pageNumber="59">
<emphasis id="B9035B42FFB5FFE7FF76F83FFEB0F86B" bold="true" box="[189,294,2014,2040]" pageId="58" pageNumber="59">Ecology:</emphasis>
—Forests.
</paragraph>
</subSubSection>
<caption id="DF08D7D8FFB4FFE6FF43FA4AFACEFA76" ID-DOI="http://doi.org/10.5281/zenodo.8061205" ID-Zenodo-Dep="8061205" httpUri="https://zenodo.org/record/8061205/files/figure.png" pageId="59" pageNumber="60" startId="59.[136,229,1451,1473]" targetBox="[151,1436,190,1426]" targetPageId="59" targetType="figure">
<paragraph id="8BC88750FFB4FFE6FF43FA4AFACEFA76" blockId="59.[136,1451,1451,1509]" pageId="59" pageNumber="60">
<emphasis id="B9035B42FFB4FFE6FF43FA4AFE9DFA53" bold="true" box="[136,267,1451,1473]" pageId="59" pageNumber="60">FIGURE 25.</emphasis>
<taxonomicName id="4C77FCD3FFB4FFE6FED8FA4AFE0DFA53" authorityName="A. Beek" authorityYear="2021" box="[275,411,1451,1472]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" isHybrid="true" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="fultus">
<emphasis id="B9035B42FFB4FFE6FED8FA4AFE0DFA53" box="[275,411,1451,1472]" italics="true" pageId="59" pageNumber="60">Rubus ×fultus</emphasis>
</taxonomicName>
.
<emphasis id="B9035B42FFB4FFE6FE62FA4AFE57FA53" bold="true" box="[425,449,1451,1472]" pageId="59" pageNumber="60">A.</emphasis>
Holotype, primocane.
<emphasis id="B9035B42FFB4FFE6FD69FA4AFD2EFA53" bold="true" box="[674,696,1451,1472]" pageId="59" pageNumber="60">B.</emphasis>
Holotype, inflorescence.
<emphasis id="B9035B42FFB4FFE6FC7FFA4AFC5DFA52" bold="true" box="[948,971,1451,1473]" pageId="59" pageNumber="60">C.</emphasis>
Primocane with leaves.
<emphasis id="B9035B42FFB4FFE6FB76FA4AFB42FA53" bold="true" box="[1213,1236,1451,1472]" pageId="59" pageNumber="60">D.</emphasis>
Primocane.
<emphasis id="B9035B42FFB4FFE6FA9BFA4AFAF0FA53" bold="true" box="[1360,1382,1451,1472]" pageId="59" pageNumber="60">E.</emphasis>
Young leaves.
<emphasis id="B9035B42FFB4FFE6FF1BFA2EFF74FA77" bold="true" box="[208,226,1487,1508]" pageId="59" pageNumber="60">F.</emphasis>
Leaf abaxially.
<emphasis id="B9035B42FFB4FFE6FEB6FA2EFE00FA77" bold="true" box="[381,406,1487,1509]" pageId="59" pageNumber="60">G.</emphasis>
Flowering branch.
<emphasis id="B9035B42FFB4FFE6FD9AFA2EFDFDFA77" bold="true" box="[593,619,1487,1508]" pageId="59" pageNumber="60">H.</emphasis>
Flowering branch.
<emphasis id="B9035B42FFB4FFE6FCEDFA2EFCA3FA77" bold="true" box="[806,821,1487,1508]" pageId="59" pageNumber="60">I.</emphasis>
Inflorescence.
<emphasis id="B9035B42FFB4FFE6FC02FA2EFC4DFA77" bold="true" box="[969,987,1487,1508]" pageId="59" pageNumber="60">J.</emphasis>
Flowers. Photographs: A. van de Beek.
</paragraph>
</caption>
<subSubSection id="C36DD4DBFFB4FFE6FF76F9D4FE92F9E1" pageId="59" pageNumber="60" type="distribution">
<paragraph id="8BC88750FFB4FFE6FF76F9D4FE92F9E1" blockId="59.[136,1452,1589,1863]" pageId="59" pageNumber="60">
<emphasis id="B9035B42FFB4FFE6FF76F9D4FECFF9DC" bold="true" box="[189,345,1589,1615]" pageId="59" pageNumber="60">Distribution:</emphasis>
—On scattered localities in the
<collectingRegion id="49B349B2FFB4FFE6FD74F9D7FCC8F9DC" box="[703,862,1589,1616]" country="South Africa" name="Western Cape" pageId="59" pageNumber="60">Western Cape</collectingRegion>
,
<collectingCountry id="F360C7C0FFB4FFE6FCA0F9D4FC6BF9DC" box="[875,1021,1589,1615]" name="South Africa" pageId="59" pageNumber="60">South Africa</collectingCountry>
; much along the road from Knysna to Uniondale.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB4FFE6FF76F99AFCD0F94F" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="8BC88750FFB4FFE6FF76F99AFCD0F94F" blockId="59.[136,1452,1589,1863]" pageId="59" pageNumber="60">
<materialsCitation id="3B1F8D0DFFB4FFE6FF76F99AFD8CF92A" collectingDate="2018-10-12" collectionCode="L" country="South Africa" county="Road" location="Houtbaai" municipality="Kirstenbos" pageId="59" pageNumber="60" specimenCount="1" stateProvince="Western Cape">
<emphasis id="B9035B42FFB4FFE6FF76F99AFD7AF906" bold="true" box="[189,748,1659,1686]" pageId="59" pageNumber="60">
Specimens:—
<collectingCountry id="F360C7C0FFB4FFE6FEAAF99AFDAEF906" box="[353,568,1659,1685]" name="South Africa" pageId="59" pageNumber="60">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FFB4FFE6FD88F99DFD7AF906" box="[579,748,1659,1686]" country="South Africa" name="Western Cape" pageId="59" pageNumber="60">Western Cape</collectingRegion>
</emphasis>
:
<collectingCounty id="62A9FFDCFFB4FFE6FD37F99AFCA1F906" box="[764,823,1659,1685]" pageId="59" pageNumber="60">Road</collectingCounty>
from
<collectingMunicipality id="6BAC1D2AFFB4FFE6FCB7F99AFC63F906" box="[892,1013,1659,1685]" pageId="59" pageNumber="60">Kirstenbos</collectingMunicipality>
to
<location id="8EA8D18BFFB4FFE6FBD1F99AFB12F906" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4:8EA8D18BFFB4FFE6FBD1F99AFB12F906" box="[1050,1156,1659,1685]" country="South Africa" county="Road" municipality="Kirstenbos" name="Houtbaai" pageId="59" pageNumber="60" stateProvince="Western Cape">Houtbaai</location>
, corner of road to
<location id="8EA8D18BFFB4FFE6FA99F99AFA31F906" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4:8EA8D18BFFB4FFE6FA99F99AFA31F906" box="[1362,1447,1659,1685]" country="South Africa" county="Road" municipality="Kirstenbos" name="Alphen" pageId="59" pageNumber="60" stateProvince="Western Cape">Alphen</location>
,
<date id="FFC9A190FFB4FFE6FF43F97EFEDEF92A" box="[136,328,1695,1721]" pageId="59" pageNumber="60" value="2018-10-12">
<collectingDate id="EF8D5878FFB4FFE6FF43F97EFEDEF92A" box="[136,328,1695,1721]" pageId="59" pageNumber="60" value="2018-10-12">12 October 2018</collectingDate>
</date>
,
<emphasis id="B9035B42FFB4FFE6FE98F97EFE7AF92A" box="[339,492,1695,1721]" italics="true" pageId="59" pageNumber="60">Beek 2018.86</emphasis>
(
<collectionCode id="ED661F95FFB4FFE6FE37F97EFD87F92A" box="[508,529,1695,1721]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="59" pageNumber="60" type="Herbarium">L</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFB4FFE6FDEDF97EFBACF92A" box="[550,1082,1695,1721]" collectingDate="1976-06-08" collectionCode="PRE" country="South Africa" location="Kirstenbosch" pageId="59" pageNumber="60" specimenCount="1" stateProvince="Western Cape">
<location id="8EA8D18BFFB4FFE6FDEDF97EFD2BF92A" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4:8EA8D18BFFB4FFE6FDEDF97EFD2BF92A" box="[550,701,1695,1721]" country="South Africa" name="Kirstenbosch" pageId="59" pageNumber="60" stateProvince="Western Cape">Kirstenbosch</location>
,
<date id="FFC9A190FFB4FFE6FD03F97EFCC5F92A" box="[712,851,1695,1721]" pageId="59" pageNumber="60" value="1976-06-08">
<collectingDate id="EF8D5878FFB4FFE6FD03F97EFCC5F92A" box="[712,851,1695,1721]" pageId="59" pageNumber="60" value="1976-06-08">8 June 1976</collectingDate>
</date>
,
<emphasis id="B9035B42FFB4FFE6FC95F97EFC7CF92A" box="[862,1002,1695,1721]" italics="true" pageId="59" pageNumber="60">Stirton 5842</emphasis>
(
<collectionCode id="ED661F95FFB4FFE6FC32F97EFBA4F92A" box="[1017,1074,1695,1721]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="59" pageNumber="60" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFB4FFE6FB8DF97EFCD4F94F" collectingDate="2018-10-17" collectionCode="L" country="South Africa" location="Uniondale" municipality="Knysna" pageId="59" pageNumber="60" specimenCount="1" stateProvince="Western Cape">
<collectingMunicipality id="6BAC1D2AFFB4FFE6FB8DF97EFB08F92A" box="[1094,1182,1695,1721]" pageId="59" pageNumber="60">Knysna</collectingMunicipality>
, road to
<location id="8EA8D18BFFB4FFE6FACBF97EFAEEF92A" LSID="urn:lsid:plazi:treatment:03DE3646FFB5FFE6FF43FBA7FEFFF8D4:8EA8D18BFFB4FFE6FACBF97EFAEEF92A" box="[1280,1400,1695,1721]" country="South Africa" municipality="Knysna" name="Uniondale" pageId="59" pageNumber="60" stateProvince="Western Cape">Uniondale</location>
, just after transition to gravel,
<date id="FFC9A190FFB4FFE6FE6FF923FDF5F94F" box="[420,611,1730,1756]" pageId="59" pageNumber="60" value="2018-10-17">
<collectingDate id="EF8D5878FFB4FFE6FE6FF923FDF5F94F" box="[420,611,1730,1756]" pageId="59" pageNumber="60" value="2018-10-17">17 October 2018</collectingDate>
</date>
,
<emphasis id="B9035B42FFB4FFE6FDA5F923FC82F94F" box="[622,788,1730,1756]" italics="true" pageId="59" pageNumber="60">Beek 2018.101</emphasis>
(
<collectionCode id="ED661F95FFB4FFE6FCEFF923FCAFF94F" box="[804,825,1730,1756]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="59" pageNumber="60" type="Herbarium">L</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFB4FFE6FF76F904FEFFF8D4" pageId="59" pageNumber="60" type="discussion">
<paragraph id="8BC88750FFB4FFE6FF76F904FEFFF8D4" blockId="59.[136,1452,1589,1863]" pageId="59" pageNumber="60">
<emphasis id="B9035B42FFB4FFE6FF76F904FF69F96C" bold="true" box="[189,255,1765,1791]" pageId="59" pageNumber="60">Note:</emphasis>
<taxonomicName id="4C77FCD3FFB4FFE6FED0F904FE2EF96C" authorityName="A. Beek" authorityYear="2021" box="[283,440,1765,1791]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" isHybrid="true" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="fultus">
<emphasis id="B9035B42FFB4FFE6FED0F904FE2EF96C" box="[283,440,1765,1791]" italics="true" pageId="59" pageNumber="60">Rubus ×fultus</emphasis>
</taxonomicName>
is most probably a hybrid of
<taxonomicName id="4C77FCD3FFB4FFE6FD34F907FCE8F96C" box="[767,894,1766,1791]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FFB4FFE6FD34F907FCE8F96C" box="[767,894,1766,1791]" italics="true" pageId="59" pageNumber="60">R. pinnatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C77FCD3FFB4FFE6FC7FF907FB8EF96C" authorityName="Weihe &amp; Nees" authorityYear="1822" box="[948,1048,1765,1791]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B9035B42FFB4FFE6FC7FF907FB8EF96C" box="[948,1048,1765,1791]" italics="true" pageId="59" pageNumber="60">R. affinis</emphasis>
</taxonomicName>
. It has the colour of
<taxonomicName id="4C77FCD3FFB4FFE6FB36F907FAEAF96C" box="[1277,1404,1766,1791]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FFB4FFE6FB36F907FAEAF96C" box="[1277,1404,1766,1791]" italics="true" pageId="59" pageNumber="60">R. pinnatus</emphasis>
</taxonomicName>
and the leaves have abaxially only few hairs; the inflorescence is also more glabrous and the prickles are more slender than with
<taxonomicName id="4C77FCD3FFB4FFE6FF0AF8CCFEF4F8D5" authorityName="A. Beek" authorityYear="2021" box="[193,354,1836,1862]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" isHybrid="true" kingdom="Plantae" order="Rosales" pageId="59" pageNumber="60" phylum="Tracheophyta" rank="species" species="bergianus">
<emphasis id="B9035B42FFB4FFE6FF0AF8CCFEF4F8D5" box="[193,354,1836,1862]" italics="true" pageId="59" pageNumber="60">R. ×bergianus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>