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<document id="FCEC0E5C7B410FE090E7CEA15BF8B70A" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FF9AFFC8FF43FE5FFE69FB62" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus borbonicus Pers. Bourbon. Legit 1806" docType="treatment" docVersion="4" lastPageNumber="22" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="22" updateTime="1699527950928" updateUser="plazi">
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<mods:title id="6105552786D98310BCC02D10B3B585D6">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
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<mods:namePart id="11E7F884B779D8F5AA28611C3E28221D">Beek, Abraham Van De</mods:namePart>
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<mods:title id="78DD1FCD8D9B9F256E3CFB9AD0483AFD">Phytotaxa</mods:title>
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<mods:date id="82D32490740BF1F986DCB87B6CF9FA18">2021</mods:date>
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<mods:identifier id="9B8F636151597212E1DC270F018B361F" type="DOI">10.11646/phytotaxa.515.1.1</mods:identifier>
<mods:identifier id="7EDFEC78FAF113CA3D4C895FF64A426A" type="ISSN">1179-3163</mods:identifier>
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<subSubSection id="C36DD4DBFF9AFFC8FF43FE5FFDC1FE4A" box="[136,599,446,473]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BC88750FF9AFFC8FF43FE5FFDC1FE4A" blockId="21.[136,599,446,473]" box="[136,599,446,473]" pageId="21" pageNumber="22">
<emphasis id="B9035B42FF9AFFC8FF43FE5FFF09FE4B" bold="true" box="[136,159,446,472]" pageId="21" pageNumber="22">b.</emphasis>
<taxonomicName id="4C77FCD3FF9AFFC8FF6DFE5FFDC1FE4A" ID-CoL="4TJLW" authority="Persoon (1806: 51)" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[166,599,446,473]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FF6DFE5FFEEEFE4A" bold="true" box="[166,376,446,473]" italics="true" pageId="21" pageNumber="22">Rubus borbonicus</emphasis>
<bibRefCitation id="EFE6FAA1FF9AFFC8FEB4FE5EFDC1FE4A" author="Persoon, C. H." box="[383,599,446,473]" pageId="21" pageNumber="22" refId="ref48715" refString="Persoon, C. H. (1806) Synopsis plantarum, seu Enchiridium botanicum, complectens enumerationem systematicam specierum hucusque cognitarum. Parisiis Lutetiorum: C. F. Cramerum, 656 pp. https: // doi. org / 10.5962 / bhl. title. 638" type="book" year="1806">Persoon (1806: 51)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF9AFFC8FF43FDEBFAD6FD8C" box="[136,1344,522,544]" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="8BC88750FF9AFFC8FF43FDEBFAD6FD8C" blockId="21.[136,1451,522,616]" box="[136,1344,522,544]" pageId="21" pageNumber="22">
<emphasis id="B9035B42FF9AFFC8FF43FDEBFEC7FDB3" box="[136,337,522,544]" italics="true" pageId="21" pageNumber="22">Homotypic synonym</emphasis>
:—
<taxonomicName id="4C77FCD3FF9AFFC8FEBBFDEBFC01FDB3" ID-CoL="4TLWD" authority="Bory de Saint-Vincent (1804: 375)" authorityName="Bory de Saint-Vincent" authorityPageNumber="375" authorityYear="1804" box="[368,919,522,544]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B9035B42FF9AFFC8FEBBFDEBFDB2FDB3" box="[368,548,522,544]" italics="true" pageId="21" pageNumber="22">Rubus tomentosus</emphasis>
<bibRefCitation id="EFE6FAA1FF9AFFC8FDFBFDEBFC01FDB3" author="Bory de Saint-Vincent, J. B. G. M." box="[560,919,522,544]" pageId="21" pageNumber="22" refId="ref46429" refString="Bory de Saint-Vincent, J. B. G. M. (1804) Voyages dans les quatre principales iles des mers d'Afrique, Vol. 2. F. Buisson, Paris, 431 pp. [https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 1047374. texteImage]" type="book" year="1804">Bory de Saint-Vincent (1804: 375)</bibRefCitation>
</taxonomicName>
, non
<bibRefCitation id="EFE6FAA1FF9AFFC8FC1CFDEBFB20FD8C" author="Borkhausen, M. B." box="[983,1206,522,544]" pageId="21" pageNumber="22" pagination="1 - 34" refId="ref46395" refString="Borkhausen, M. B. (1794) Beytrage zur deutschen Flora, nebst Beobachtungen einiger seltener Pflanzen. R ˆ mer's Neues Magazin fur die Botanik 1: 1 - 34." type="journal article" year="1794">Borkhausen (1794: 2)</bibRefCitation>
,
<emphasis id="B9035B42FF9AFFC8FB0DFDEAFAD6FD8C" box="[1222,1344,522,544]" italics="true" pageId="21" pageNumber="22">nom. illegit.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF9AFFC8FA80FDEBFDB1FDFB" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BC88750FF9AFFC8FA80FDEBFDB1FDFB" blockId="21.[136,1451,522,616]" pageId="21" pageNumber="22">
<materialsCitation id="3B1F8D0DFF9AFFC8FA80FDEBFDB5FDFB" collectionCode="P" country="Réunion" location="REUNION" pageId="21" pageNumber="22" specimenCode="P02579816" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54CC39F2FF9AFFC8FA80FDEBFA3DFD8C" box="[1355,1451,522,543]" pageId="21" pageNumber="22" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F360C7C0FF9AFFC8FE48FDCFFE62FDD6" box="[387,500,558,581]" name="Réunion" pageId="21" pageNumber="22">RÉUNION</collectingCountry>
.
<taxonomicName id="4C77FCD3FF9AFFC8FDCFFDCFFC04FDD7" authority="Pers. Bourbon. Legit" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[516,914,558,580]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">Rubus borbonicus Pers. Bourbon. Legit</taxonomicName>
Bory (
<specimenCode id="DBD12F2BFF9AFFC8FC2BFDCFFBD9FDD7" box="[992,1103,558,580]" pageId="21" pageNumber="22">P02579816</specimenCode>
[https://science.mnhn.fr/institution/ mnhn/collection/p/item/p02579816])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF9AFFC8FF43FD77FE69FB62" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BC88750FF9AFFC8FF43FD77FE67FD6A" blockId="21.[136,1452,662,1265]" pageId="21" pageNumber="22">
<emphasis id="B9035B42FF9AFFC8FF43FD77FF42FD23" bold="true" box="[136,212,662,688]" pageId="21" pageNumber="22">Notes:</emphasis>
<bibRefCitation id="EFE6FAA1FF9AFFC8FF3AFD76FE09FD23" author="Persoon, C. H." box="[241,415,662,689]" pageId="21" pageNumber="22" refId="ref48715" refString="Persoon, C. H. (1806) Synopsis plantarum, seu Enchiridium botanicum, complectens enumerationem systematicam specierum hucusque cognitarum. Parisiis Lutetiorum: C. F. Cramerum, 656 pp. https: // doi. org / 10.5962 / bhl. title. 638" type="book" year="1806">Persoon (1806)</bibRefCitation>
, in the protologue of
<taxonomicName id="4C77FCD3FF9AFFC8FD58FD76FCA7FD23" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[659,817,662,688]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FD58FD76FCA7FD23" box="[659,817,662,688]" italics="true" pageId="21" pageNumber="22">R. borbonicus</emphasis>
</taxonomicName>
, explicitly refers to
<taxonomicName id="4C77FCD3FF9AFFC8FBDDFD76FB62FD22" authority="Bory" authorityName="Bory de Saint-Vincent" authorityYear="1804" box="[1046,1268,663,689]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B9035B42FF9AFFC8FBDDFD76FB22FD23" box="[1046,1204,663,688]" italics="true" pageId="21" pageNumber="22">R. tomentosus</emphasis>
Bory
</taxonomicName>
as identical and the details of the description are borrowed from Borys publication. So
<taxonomicName id="4C77FCD3FF9AFFC8FC60FD5AFBDEFD47" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[939,1096,698,724]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FC60FD5AFBDEFD47" box="[939,1096,698,724]" italics="true" pageId="21" pageNumber="22">R. borbonicus</emphasis>
</taxonomicName>
is a homotypic synonym of the illegitimate
<taxonomicName id="4C77FCD3FF9AFFC8FEC4FD3EFE67FD6A" authority="Bory." authorityName="Bory de Saint-Vincent" authorityYear="1804" box="[271,497,735,761]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B9035B42FF9AFFC8FEC4FD3EFE3AFD6B" box="[271,428,735,760]" italics="true" pageId="21" pageNumber="22">R. tomentosus</emphasis>
Bory.
</taxonomicName>
</paragraph>
<paragraph id="8BC88750FF9AFFC8FF76FCE3FF5DFC43" blockId="21.[136,1452,662,1265]" pageId="21" pageNumber="22">
There are
<specimenCount id="9D714CD9FF9AFFC8FEFEFCE2FE48FC8F" box="[309,478,770,797]" pageId="21" pageNumber="22" type="generic">two specimens</specimenCount>
collected by Bory in P. On the label of the first one is written:
<taxonomicName id="4C77FCD3FF9AFFC8FB06FCE3FA35FC8F" authorityName="Bory de Saint-Vincent" authorityYear="1804" box="[1229,1443,770,796]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="tomentosus">Rubus tomentosus</taxonomicName>
/ nob. var./ en arivant a la/ plaine des cafres/ a Bourbon (P02520892). It could not be selected as
<typeStatus id="54CC39F2FF9AFFC8FB1BFCC6FA96FCD2" box="[1232,1280,807,833]" pageId="21" pageNumber="22">type</typeStatus>
because of the addition var.; the combination nob. and var. suggests that Bory already acknowledged a species
<taxonomicName id="4C77FCD3FF9AFFC8FB2AFCAAFAEBFCF7" authorityName="Bory de Saint-Vincent" authorityYear="1804" box="[1249,1405,843,868]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="tomentosus">
<emphasis id="B9035B42FF9AFFC8FB2AFCAAFAEBFCF7" box="[1249,1405,843,868]" italics="true" pageId="21" pageNumber="22">R. tomentosus</emphasis>
</taxonomicName>
and considered this plant as a variety. It has stronger curved prickles, narrow leaves, an open inflorescence, shorter ovate sepals, but on the other hand very clearly the conspicuous serrature of
<taxonomicName id="4C77FCD3FF9AFFC8FC6EFC72FBD5FC3F" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[933,1091,914,940]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FC6EFC72FBD5FC3F" box="[933,1091,914,940]" italics="true" pageId="21" pageNumber="22">R. borbonicus</emphasis>
</taxonomicName>
. It may be a
<taxonomicName id="4C77FCD3FF9AFFC8FB13FC73FADAFC3F" box="[1240,1356,914,940]" form="from" pageId="21" pageNumber="22" rank="form">form from</taxonomicName>
a sunny place.
</paragraph>
<paragraph id="8BC88750FF9AFFC8FF76FC3BFEB6FBF3" blockId="21.[136,1452,662,1265]" pageId="21" pageNumber="22">
The other sample collected by Bory consists of some broken off leaflets and a piece of an inflorescence stem (P02579816). Richard wrote on the label
<taxonomicName id="4C77FCD3FF9AFFC8FDAAFC1FFBB3FB8B" authority="Pers. Bourbon. Legit" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[609,1061,1022,1049]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">Rubus borbonicus Pers. Bourbon. Legit</taxonomicName>
Bory. Though it is not written by Bory, it is clear that the specimen was collected by him, and because no other specimen is available it was selected as the
<typeStatus id="54CC39F2FF9AFFC8FF7AFBA7FE8DFBF3" box="[177,283,1094,1120]" pageId="21" pageNumber="22" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC88750FF9AFFC8FF76FB8BFE69FB62" blockId="21.[136,1452,662,1265]" pageId="21" pageNumber="22">
The non-flowering branch (either a very young primocane or a secondary shoot) of the
<typeStatus id="54CC39F2FF9AFFC8FB7FFB8AFB72FB16" box="[1204,1252,1131,1157]" pageId="21" pageNumber="22">type</typeStatus>
of
<taxonomicName id="4C77FCD3FF9AFFC8FAC5FB8AFA19FB17" authorityName="Poiret" authorityYear="1804" box="[1294,1423,1130,1156]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="apetalus">
<emphasis id="B9035B42FF9AFFC8FAC5FB8AFA19FB17" box="[1294,1423,1130,1156]" italics="true" pageId="21" pageNumber="22">R. apetalus</emphasis>
</taxonomicName>
is not very characteristic. The inflorescence of the
<typeStatus id="54CC39F2FF9AFFC8FD7BFB6EFD76FB3A" box="[688,736,1167,1193]" pageId="21" pageNumber="22">type</typeStatus>
is more characteristic and there is no reason to not identify
<taxonomicName id="4C77FCD3FF9AFFC8FA58FB6EFE90FB5F" authorityName="Pers. Bourbon. Legit" authorityYear="1806" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FA58FB6EFE90FB5F" italics="true" pageId="21" pageNumber="22">R. borbonicus</emphasis>
</taxonomicName>
with it. Earlier authors (except
<bibRefCitation id="EFE6FAA1FF9AFFC8FDA6FB53FCBCFB5F" author="Gustafsson, C. E." box="[621,810,1202,1228]" pageId="21" pageNumber="22" pagination="1 - 68" refId="ref47689" refString="Gustafsson, C. E. (1934) Rubi Africani. Arkiv f ˆ r Botanik 26 A (7): 1 - 68." type="journal article" year="1934">Gustafsson 1934</bibRefCitation>
) drew the same conclusion. Consequently
<taxonomicName id="4C77FCD3FF9AFFC8FAC5FB52FA3DFB5F" authorityName="Pers. Bourbon. Legit" authorityYear="1806" box="[1294,1451,1202,1228]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="borbonicus">
<emphasis id="B9035B42FF9AFFC8FAC5FB52FA3DFB5F" box="[1294,1451,1202,1228]" italics="true" pageId="21" pageNumber="22">R. borbonicus</emphasis>
</taxonomicName>
is a later synonym of
<taxonomicName id="4C77FCD3FF9AFFC8FEB1FB36FE6EFB63" authorityName="Poiret" authorityYear="1804" box="[378,504,1238,1264]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="apetalus">
<emphasis id="B9035B42FF9AFFC8FEB1FB36FE6EFB63" box="[378,504,1238,1264]" italics="true" pageId="21" pageNumber="22">R. apetalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>